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<mods:title id="D73C5C46FED484AEBA36C07A6C7123DE">On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the Southwest Pacific (Decapoda: Brachyura: Pinnotheridae)</mods:title>
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<mods:namePart id="AD372C90A2E6D422614AE0603D2985EB">Komai, Tomoyuki</mods:namePart>
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<mods:namePart id="CD8020F6507CDB30F91CF9B0AB845873">Kawai, Kei</mods:namePart>
<mods:affiliation id="0164E8F07167B7B5D2431D4BA1818C71">International Center for Island Studies, Kagoshima University, Korimoto 1 - 21 - 24, Kagoshima, 890 - 8580.</mods:affiliation>
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<mods:namePart id="2F75EF5AF9FEB9CC7BC1D58F8C2B8827">Ng, Peter K. L.</mods:namePart>
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<mods:affiliation id="C6B01F0195AB15AFAB9DE7FE2259EC81">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore. peterng @ nus. edu. sg; https: // orcid. org / 0000 - 0001 - 5946 - 0608</mods:affiliation>
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<taxonomicName id="5CBC4D4C950C426F73A2FF50FDCFC2D8" ID-CoL="84GPX" authority="(Dana, 1852)" authorityName="Komai &amp; Kawai &amp; Ng" authorityYear="2020" baseAuthorityName="Dana" baseAuthorityYear="1852" box="[151,539,151,178]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="223" phylum="Arthropoda" rank="species" species="obesus" status="comb. nov.">
<emphasis id="A9C8EADD950C426F73A2FF50FEA0C2DB" bold="true" box="[151,372,151,178]" italics="true" pageId="2" pageNumber="223">Arcotheres obesus</emphasis>
(
<bibRefCitation id="FF2D4B3E950C426F72B6FF50FDC7C2D8" author="Dana, J. D." box="[387,531,151,177]" pageId="2" pageNumber="223" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana, 1852</bibRefCitation>
)
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<taxonomicNameLabel id="B2FB57A6950C426F7117FF50FD59C2D8" box="[546,653,151,177]" pageId="2" pageNumber="223" rank="species">n. comb.</taxonomicNameLabel>
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<subSubSection id="D3A66544950C426F73A2FF7CFEC6C2BF" box="[151,274,187,214]" pageId="2" pageNumber="223" type="description">
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(
<figureCitation id="03872A4A950C426F73AAFF7CFEDDC2BF" box="[159,265,187,214]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="2.[151,250,1747,1771]" captionStartId-1="3.[151,250,1941,1965]" captionStartId-2="4.[151,250,1848,1872]" captionStartId-3="6.[151,250,1862,1886]" captionStartId-4="8.[151,250,1867,1891]" captionStartId-5="10.[151,250,1791,1815]" captionTargetBox-0="[160,1425,839,1716]" captionTargetBox-1="[151,1435,181,1917]" captionTargetBox-2="[162,1423,247,1813]" captionTargetBox-3="[226,1364,190,1827]" captionTargetBox-4="[157,1424,499,1834]" captionTargetBox-5="[166,1424,193,1760]" captionTargetId-0="figure-247@2.[151,1436,833,1724]" captionTargetId-1="figure-17@3.[151,1435,181,1917]" captionTargetId-2="figure-46@4.[151,1436,237,1824]" captionTargetId-3="figure-20@6.[216,1371,181,1838]" captionTargetId-4="figure-186@8.[151,1436,489,1844]" captionTargetId-5="figure-20@10.[151,1436,181,1768]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="6" captionTargetPageId-4="8" captionTargetPageId-5="10" captionText-0="FIGURE 1. Arcotheres obesus (Dana, 1852). A, “male”, overall habitus in dorsal view; B, C, E, F, female, carapace, dorsal view; D, “male”, frontal view showing epistome, antennae, antennules and eyes; E, maxillipeds 3; F, left maxilliped 3; G, outer view of right chela; H, “male” pleon, outer view. After Dana (1855: pl. 24 fig. 3ai)." captionText-1="FIGURE 2. Arcotheres obesus (Dana, 1852). A, E, female (5.9×8.3 mm), ZRC 2019.1876; BD, female (6.9×8.9 mm), ZRC 2019.1876. A, B, overall habitus, dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view." captionText-2="FIGURE 3. Arcotheres obesus (Dana, 1852). AD, neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945; E, F, female (6.9×8.9 mm), ZRC 2019.1876; G, H, female (5.9×8.3 mm), ZRC 2019.1876. A, overall habitus, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, pleomere 6 and telson, ventral view; E, G, left maxilliped 3, outer view; F, H, left maxilliped 3, inner view." captionText-3="FIGURE 4. Arcotheres obesus (Dana, 1852), neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945. A, right chela, outer view; B, same, inner view; C, chela and carpus of right cheliped, dorsal view; D, distal part of fixed finger of right chela, ventral view; E, carpus and merus of right cheliped, ventral view; F, right P2, dorsal view; G, same dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (shorter side), dorsal view; K, same, dactylus; L, left P4 (longer side), dorsal view; M, same dactylus; N, right P5, dorsal view; O, same dactylus." captionText-4="FIGURE 5. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, habitus, dorsal view; B, left MXP3, outer view; C, left thoracic sternum, ventral view; D, pleon, telson and exposed part of sternite 8 (left), ventral view; E, left G1, outer view; F, same, inner view; G, left G2, outer view; H, same, different view." captionText-5="FIGURE 6. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, left chela, outer view; B, same, inner view; C, chela and carpus of left cheliped, dorsal view; D, merus of left cheliped, ventral view; E, left P2, dorsal view; F, same, dactylus; G, left P3, dorsal view; H, same, dactylus; I, left P4 (longer side), dorsal view; J, same, dactylus; K, right P4 (shorter side), dorsal view; L, same, dactylus and propodus, dorsal (perpendicular) view; M, right P5, dorsal view; N, same, dactylus." figureDoi-0="http://doi.org/10.5281/zenodo.4401463" figureDoi-1="http://doi.org/10.5281/zenodo.4401467" figureDoi-2="http://doi.org/10.5281/zenodo.4401469" figureDoi-3="http://doi.org/10.5281/zenodo.4401471" figureDoi-4="http://doi.org/10.5281/zenodo.4401473" figureDoi-5="http://doi.org/10.5281/zenodo.4401475" httpUri-0="https://zenodo.org/record/4401463/files/figure.png" httpUri-1="https://zenodo.org/record/4401467/files/figure.png" httpUri-2="https://zenodo.org/record/4401469/files/figure.png" httpUri-3="https://zenodo.org/record/4401471/files/figure.png" httpUri-4="https://zenodo.org/record/4401473/files/figure.png" httpUri-5="https://zenodo.org/record/4401475/files/figure.png" pageId="2" pageNumber="223">Figs. 16</figureCitation>
)
</paragraph>
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<treatmentCitation id="1A1D10DE950C426F73A2FEC7FE2DC371" author="Dana, J. D." box="[151,505,255,280]" page="380" pageId="2" pageNumber="223" year="1852">
<taxonomicName id="5CBC4D4C950C426F73A2FEC7FE2DC371" ID-CoL="84H4Q" authority="Dana, 1852: 380" authorityName="Dana" authorityPageNumber="380" authorityYear="1852" box="[151,505,255,280]" class="Malacostraca" family="Pinnotheridae" genus="Pinnothera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="223" phylum="Arthropoda" rank="species" species="obesa">
<emphasis id="A9C8EADD950C426F73A2FEC7FE9FC371" box="[151,331,256,280]" italics="true" pageId="2" pageNumber="223">Pinnothera obesa</emphasis>
<bibRefCitation id="FF2D4B3E950C426F7264FEC7FE2DC371" author="Dana, J. D." box="[337,505,255,280]" pageId="2" pageNumber="223" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana, 1852: 380</bibRefCitation>
</taxonomicName>
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.—
<bibRefCitation id="FF2D4B3E950C426F7122FEC7FD5FC37E" author="Dana, J. D." box="[535,651,255,280]" pageId="2" pageNumber="223" refId="ref13137" refString="Dana, J. D. (1855) Atlas. Crustacea in United States Exploring Expedition during the years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 14. C. Sherman, Philadelphia, 27 pp., 96 pls." type="book" year="1855">Dana 1855</bibRefCitation>
: pl. 24 fig. 3.—
<treatmentCitation id="1A1D10DE950C426F7010FEC7FBD1C371" author="Woodward, H." box="[805,1029,255,280]" page="177" pageId="2" pageNumber="223" year="1886">
<bibRefCitation id="FF2D4B3E950C426F7010FEC7FBD1C371" author="Woodward, H." box="[805,1029,255,280]" pageId="2" pageNumber="223" pagination="176 - 177" refId="ref15438" refString="Woodward, H. (1886) Animals commensal or parasitic in Meleagrina margaritifera Lin. Proceedings of the Zoological Society of London, 1886 (2), 176 - 177." type="journal article" year="1886">Woodward 1886: 177</bibRefCitation>
</treatmentCitation>
(list).
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</paragraph>
<treatmentCitationGroup id="BBAC11E1950C426F73A2FED9FDA9C33C" pageId="2" pageNumber="223">
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<taxonomicName id="5CBC4D4C950C426F73A2FED9FE88C35F" authorityName=", Dana" authorityYear="1852" box="[151,348,286,310]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="223" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD950C426F73A2FED9FE88C35F" box="[151,348,286,310]" italics="true" pageId="2" pageNumber="223">Pinnotheres obesus</emphasis>
</taxonomicName>
.—
<treatmentCitation id="1A1D10DE950C426F724EFED9FD18C35F" author="Silas, E. G. &amp; Alagarswami, K." box="[379,716,286,310]" page="1204" pageId="2" pageNumber="223" year="1967">
<bibRefCitation id="FF2D4B3E950C426F724EFED9FD18C35F" author="Silas, E. G. &amp; Alagarswami, K." box="[379,716,286,310]" pageId="2" pageNumber="223" pagination="1161 - 1227" refId="ref14962" refString="Silas, E. G. &amp; Alagarswami, K. (1967) On an instance of parasitisation by the pea-crab (Pinnotheres sp.) on the backwater clam [Meretrix casta (Chemnitz)] from India, with a review of the work on systematics, ecology, biology and etiology of pea crabs of the genus Pinnotheres, Latreille, 1802. Proceedings of the Symposium on Crustacea held at Ernakulam, 1965. Symposium Series 2. Marine Biological Association of India, Tamil Nadu, pp. 1161 - 1227." type="book chapter" year="1967">Silas &amp; Alagarswami 1967: 1204</bibRefCitation>
</treatmentCitation>
(list).—
<treatmentCitation id="1A1D10DE950C426F702BFED9FBDDC35F" author="Schmitt, W. L. &amp; Mccain, J. C. &amp; Davidson, E." box="[798,1033,286,310]" page="60" pageId="2" pageNumber="223" year="1973">
<bibRefCitation id="FF2D4B3E950C426F702BFED9FBDDC35F" author="Schmitt, W. L. &amp; Mccain, J. C. &amp; Davidson, E." box="[798,1033,286,310]" pageId="2" pageNumber="223" refId="ref14912" refString="Schmitt, W. L., Mccain, J. C. &amp; Davidson, E. (1973) Decapoda I. Brachyura I. Fam. Pinnotheridae. Crustaceorum Catalogus. Vol. 3. Dr. W. Junk, Den Haag, 160 pp." type="book" year="1973">
Schmitt
<emphasis id="A9C8EADD950C426F7042FED8FC70C35F" box="[887,932,286,310]" italics="true" pageId="2" pageNumber="223">et al</emphasis>
. 1973: 60
</bibRefCitation>
</treatmentCitation>
(part) (list and discussion).
</paragraph>
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<emphasis id="A9C8EADD950C426F73A2FEFAFEB7C33C" box="[151,355,317,341]" italics="true" pageId="2" pageNumber="223">
<taxonomicName id="5CBC4D4C950C426F73A2FEFAFE8BC33C" authorityName=", Dana" authorityYear="1852" box="[151,351,317,341]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="223" phylum="Arthropoda" rank="species" species="obesus">Pinnotheres obesus</taxonomicName>
.
</emphasis>
<treatmentCitation id="1A1D10DE950C426F7249FEFAFD94C33C" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[380,576,317,341]" page="250" pageId="2" pageNumber="223" year="2008">
<bibRefCitation id="FF2D4B3E950C426F7249FEFAFD94C33C" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[380,576,317,341]" pageId="2" pageNumber="223" pagination="1 - 286" refId="ref14697" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="A9C8EADD950C426F7294FEF9FE1AC33C" box="[417,462,317,341]" italics="true" pageId="2" pageNumber="223">et al</emphasis>
. 2008: 250
</bibRefCitation>
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(list).
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<paragraph id="9B0336CF950C426F73A2FE44FB92C3A8" blockId="2.[151,1437,387,810]" pageId="2" pageNumber="223">
<emphasis id="A9C8EADD950C426F73A2FE44FE90C3F4" bold="true" box="[151,324,387,413]" pageId="2" pageNumber="223">Type material.</emphasis>
<materialsCitation id="2BD43C92950C426F727CFE43FB92C3A8" ID-GBIF-Occurrence="3019935317" collectingDate="2019-02-22" collectionCode="V" collectorName="K. Kawai." country="Fiji" location="Kumi Village" municipality="Levu" pageId="2" pageNumber="223" specimenCode="CBM-ZC 15945" specimenCount="1" typeStatus="neotype">
<typeStatus id="4407886D950C426F727CFE43FE78C3F7" box="[329,428,388,414]" pageId="2" pageNumber="223" type="neotype">Neotype</typeStatus>
:
<specimenCode id="CB1A9EB4950C426F7283FE44FDA4C3F7" box="[438,624,387,414]" collectionCode="CBM-ZC" pageId="2" pageNumber="223">CBM-ZC 15945</specimenCode>
, female of post-hard stage
<collectionCode id="FDADAE0A950C426F70A2FE43FC7FC3F7" box="[919,939,388,414]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="223" type="Museum">V</collectionCode>
(4.4×
<quantity id="5C449B2A950C426F70D9FE44FB91C3F7" box="[1004,1093,387,414]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="223" unit="mm" value="5.5">5.5 mm</quantity>
),
<location id="9E636014950C426F7763FE44FB3BC3F4" LSID="urn:lsid:plazi:treatment:131587D9950C426473A2FF50FDEFC518:9E636014950C426F7763FE44FB3BC3F4" box="[1110,1263,387,413]" country="Fiji" municipality="Levu" name="Kumi Village" pageId="2" pageNumber="223">Kumi Village</location>
, Viti
<collectingMunicipality id="7B67ACB5950C426F761DFE43FAB0C3F7" box="[1320,1380,388,414]" pageId="2" pageNumber="223">Levu</collectingMunicipality>
,
<collectingCountry id="E3AB765F950C426F765BFE44FA4CC3F4" box="[1390,1432,387,413]" name="Fiji" pageId="2" pageNumber="223">Fiji</collectingCountry>
, intertidal, associated with
<taxonomicName id="5CBC4D4C950C426F7288FE60FD6CC3A8" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[445,696,423,449]" class="Bivalvia" family="Veneridae" genus="Gafrarium" kingdom="Animalia" order="Veneroida" pageId="2" pageNumber="223" phylum="Mollusca" rank="species" species="pectinatum">
<emphasis id="A9C8EADD950C426F7288FE60FD6CC3A8" box="[445,696,423,449]" italics="true" pageId="2" pageNumber="223">Gafrarium pectinatum</emphasis>
</taxonomicName>
,
<date id="EF02100F950C426F71F3FE60FC44C3A8" box="[710,912,423,449]" pageId="2" pageNumber="223" value="2019-02-22">
<collectingDate id="FF46E9E7950C426F71F3FE60FC44C3A8" box="[710,912,423,449]" pageId="2" pageNumber="223" value="2019-02-22">22 February 2019</collectingDate>
</date>
, coll.
<collectorName id="36495319950C426F70E6FE6FFB92C3A8" box="[979,1094,423,450]" pageId="2" pageNumber="223">K. Kawai.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="9B0336CF950C426F73F2FE0CFC30C088" blockId="2.[151,1437,387,810]" pageId="2" pageNumber="223">
<materialsCitation id="2BD43C92950C426F73F2FE0CFB2DC060" ID-GBIF-Occurrence="3019935306" collectionCode="V" collectorName="Females" country="Fiji" location="Kumi Village" pageId="2" pageNumber="223" specimenCode="CBM-ZC 15946" specimenCount="5" specimenCount-female="3" specimenCount-male="2">
<emphasis id="A9C8EADD950C426F73F2FE0CFE2FC38C" bold="true" box="[199,507,459,485]" pageId="2" pageNumber="223">Other material examined.</emphasis>
<collectingCountry id="E3AB765F950C426F7134FE0CFDF8C38C" box="[513,556,459,485]" name="Fiji" pageId="2" pageNumber="223">Fiji</collectingCountry>
.
<collectorName id="36495319950C426F7100FE0CFD46C38C" box="[565,658,459,485]" pageId="2" pageNumber="223">Females</collectorName>
are all in the post-hard stage
<collectionCode id="FDADAE0A950C426F70ECFE0BFC39C38F" box="[985,1005,460,486]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="223" type="Museum">V</collectionCode>
.
<specimenCode id="CB1A9EB4950C426F70CDFE0CFB66C38F" box="[1016,1202,459,486]" collectionCode="CBM-ZC" pageId="2" pageNumber="223">CBM-ZC 15946</specimenCode>
,
<specimenCount id="8DBAFD46950C426F7788FE0CFAFCC38C" box="[1213,1320,459,485]" pageId="2" pageNumber="223" type="female">3 females</specimenCount>
(4.2×5.3 5.1×
<quantity id="5C449B2A950C426F73FFFE28FEF1C063" box="[202,293,495,522]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="2" pageNumber="223" unit="mm" value="6.1">6.1 mm</quantity>
), 1 ovigerous female (4.8×
<quantity id="5C449B2A950C426F7167FE28FD7AC063" box="[594,686,495,522]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="2" pageNumber="223" unit="mm" value="5.4">5.4 mm</quantity>
),
<specimenCount id="8DBAFD46950C426F71F4FE28FCCCC060" box="[705,792,495,521]" pageId="2" pageNumber="223" type="male">2 males</specimenCount>
(3.6×ca
<quantity id="5C449B2A950C426F7049FE28FC01C063" box="[892,981,495,522]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="223" unit="mm" value="4.2">4.2 mm</quantity>
, damaged),
<location id="9E636014950C426F776BFE28FB2DC060" LSID="urn:lsid:plazi:treatment:131587D9950C426473A2FF50FDEFC518:9E636014950C426F776BFE28FB2DC060" box="[1118,1273,495,521]" country="Fiji" name="Kumi Village" pageId="2" pageNumber="223">Kumi Village</location>
</materialsCitation>
,
<materialsCitation id="2BD43C92950C426F7631FE28FBDBC044" ID-GBIF-Occurrence="3019935324" collectingDate="2010-08-16" collectorName="Levu &amp; K. Kawai" country="Fiji" location="Viti" pageId="2" pageNumber="223" specimenCount="1">
<collectingCountry id="E3AB765F950C426F7631FE28FAFAC060" box="[1284,1326,495,521]" name="Fiji" pageId="2" pageNumber="223">Viti</collectingCountry>
<collectorName id="36495319950C426F7600FE37FAA5C063" box="[1333,1393,496,522]" pageId="2" pageNumber="223">Levu</collectorName>
, intertidal, associated with
<taxonomicName id="5CBC4D4C950C426F7290FDD4FD4BC044" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[421,671,531,557]" class="Bivalvia" family="Veneridae" genus="Gafrarium" kingdom="Animalia" order="Veneroida" pageId="2" pageNumber="223" phylum="Mollusca" rank="species" species="pectinatum">
<emphasis id="A9C8EADD950C426F7290FDD4FD4BC044" box="[421,671,531,557]" italics="true" pageId="2" pageNumber="223">Gafrarium pectinatum</emphasis>
</taxonomicName>
,
<date id="EF02100F950C426F7199FDD4FC8BC044" box="[684,863,531,557]" pageId="2" pageNumber="223" value="2010-08-16">
<collectingDate id="FF46E9E7950C426F7199FDD4FC8BC044" box="[684,863,531,557]" pageId="2" pageNumber="223" value="2010-08-16">16 August 2010</collectingDate>
</date>
, coll.
<collectorName id="36495319950C426F7094FDD3FBDBC044" box="[929,1039,531,558]" pageId="2" pageNumber="223">K. Kawai</collectorName>
</materialsCitation>
;
<materialsCitation id="2BD43C92950C426F772CFDD4FD1CC01C" ID-GBIF-Occurrence="3019935313" collectionCode="ZRC" country="Fiji" location="Waiganake Village" pageId="2" pageNumber="223" specimenCode="CBM-ZC 15947, 15948, ZRC 2019.1876" specimenCount="6" specimenCount-female="4" specimenCount-juvenile="2">
<specimenCode id="CB1A9EB4950C426F772CFDD4FB00C047" box="[1049,1236,531,558]" collectionCode="CBM-ZC" pageId="2" pageNumber="223">CBM-ZC 15947</specimenCode>
,
<specimenCode id="CB1A9EB4950C426F77EBFDD4FAF3C047" box="[1246,1319,531,558]" collectionCode="CBM-ZC" pageId="2" pageNumber="223">15948</specimenCode>
,
<specimenCount id="8DBAFD46950C426F7604FDD4FA48C044" box="[1329,1436,531,557]" pageId="2" pageNumber="223" type="female">2 females</specimenCount>
(4.0×5.3, 7.2×
<quantity id="5C449B2A950C426F7200FDF0FE5BC03B" box="[309,399,567,594]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="2" pageNumber="223" unit="mm" value="9.2">9.2 mm</quantity>
),
<specimenCode id="CB1A9EB4950C426F7295FDF0FD83C038" box="[416,599,567,593]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="223">ZRC 2019.1876</specimenCode>
,
<specimenCount id="8DBAFD46950C426F7155FDF0FD1FC038" box="[608,715,567,593]" pageId="2" pageNumber="223" type="female">2 females</specimenCount>
(6.2×8.8, 6.9×
<quantity id="5C449B2A950C426F7045FDF0FC1DC03B" box="[880,969,567,594]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="2" pageNumber="223" unit="mm" value="8.9">8.9 mm</quantity>
),
<specimenCount id="8DBAFD46950C426F70EFFDF0FB9CC038" box="[986,1096,567,593]" pageId="2" pageNumber="223" type="juvenile">2 juvenile</specimenCount>
males (gonopods not differentiated) (1.7×1.9, 2.2×
<quantity id="5C449B2A950C426F72B3FD9CFE35C01F" box="[390,481,603,630]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="223" unit="mm" value="2.6">2.6 mm</quantity>
),
<location id="9E636014950C426F72C1FD9CFD1CC01C" LSID="urn:lsid:plazi:treatment:131587D9950C426473A2FF50FDEFC518:9E636014950C426F72C1FD9CFD1CC01C" box="[500,712,603,629]" country="Fiji" name="Waiganake Village" pageId="2" pageNumber="223">Waiganake Village</location>
</materialsCitation>
,
<materialsCitation id="2BD43C92950C426F71E0FD9CFE0AC0F0" ID-GBIF-Occurrence="3019935319" collectingDate="2011-08-22" collectorName="Levu &amp; K. Kawai" country="Fiji" location="Viti" pageId="2" pageNumber="223" specimenCount="1">
<collectingCountry id="E3AB765F950C426F71E0FD9CFD2BC01C" box="[725,767,603,629]" name="Fiji" pageId="2" pageNumber="223">Viti</collectingCountry>
<collectorName id="36495319950C426F7032FD9BFC97C01F" box="[775,835,604,630]" pageId="2" pageNumber="223">Levu</collectorName>
, intertidal, associated with
<taxonomicName id="5CBC4D4C950C426F7743FD9CFAA5C01C" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1142,1393,603,629]" class="Bivalvia" family="Veneridae" genus="Gafrarium" kingdom="Animalia" order="Veneroida" pageId="2" pageNumber="223" phylum="Mollusca" rank="species" species="pectinatum">
<emphasis id="A9C8EADD950C426F7743FD9CFAA5C01C" box="[1142,1393,603,629]" italics="true" pageId="2" pageNumber="223">Gafrarium pectinatum</emphasis>
</taxonomicName>
,
<date id="EF02100F950C426F76B5FD9CFEFEC0F0" pageId="2" pageNumber="223" value="2011-08-22">
<collectingDate id="FF46E9E7950C426F76B5FD9CFEFEC0F0" pageId="2" pageNumber="223" value="2011-08-22">22 August 2011</collectingDate>
</date>
, coll.
<collectorName id="36495319950C426F725BFD47FE0AC0F0" box="[366,478,639,666]" pageId="2" pageNumber="223">K. Kawai</collectorName>
</materialsCitation>
;
<materialsCitation id="2BD43C92950C426F72DCFDB8FD5BC0D4" ID-GBIF-Occurrence="3019935303" collectingDate="2019-02-22" collectionCode="V" collectorName="K. Kawai." country="Fiji" location="Kumi Village" municipality="Levu" pageId="2" pageNumber="223" specimenCode="CBM-ZC 15949-15951, 3" specimenCount="1">
<specimenCode id="CB1A9EB4950C426F72DCFDB8FD2DC0F3" box="[489,761,639,666]" collectionCode="CBM-ZC" pageId="2" pageNumber="223">CBM-ZC 1594915951</specimenCode>
,
<specimenCode id="CB1A9EB4950C426F7030FDB8FCC7C0F0" box="[773,787,639,665]" collectionCode="CBM-ZC" pageId="2" pageNumber="223">3</specimenCode>
ovigerous females (5.4×6.7, 5.6×6.6, 4.6×
<quantity id="5C449B2A950C426F77C1FDB8FA9BC0F3" box="[1268,1359,639,666]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="2" pageNumber="223" unit="mm" value="5.9">5.9 mm</quantity>
), same data as neotype, associated with
<taxonomicName id="5CBC4D4C950C426F713FFD63FD5BC0D4" baseAuthorityName="Roding" baseAuthorityYear="1798" box="[522,655,675,702]" class="Bivalvia" family="Veneridae" genus="Gafrarium" kingdom="Animalia" order="Veneroida" pageId="2" pageNumber="223" phylum="Mollusca" rank="species" species="tumidum">
<emphasis id="A9C8EADD950C426F713FFD63FD5BC0D4" box="[522,655,675,702]" italics="true" pageId="2" pageNumber="223">G. tumidum</emphasis>
</taxonomicName>
</materialsCitation>
;
<materialsCitation id="2BD43C92950C426F71AAFD64FE35C088" ID-GBIF-Occurrence="3019935321" collectionCode="ZRC" country="Fiji" pageId="2" pageNumber="223" specimenCode="ZRC 2020.0006, 1" specimenCount="1">
<specimenCode id="CB1A9EB4950C426F71AAFD64FC83C0D4" box="[671,855,675,701]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="223">ZRC 2020.0006</specimenCode>
,
<specimenCode id="CB1A9EB4950C426F7056FD64FCA5C0D4" box="[867,881,675,701]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="223">1</specimenCode>
ovigerous female (4.9×
<quantity id="5C449B2A950C426F774AFD64FB0FC0D7" box="[1151,1243,675,702]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="2" pageNumber="223" unit="mm" value="6.2">6.2 mm</quantity>
), associated with
<taxonomicName id="5CBC4D4C950C426F73A2FD00FE35C088" authority=", Telau" authorityName="Telau" box="[151,481,711,738]" class="Bivalvia" family="Veneridae" genus="Gafrarium" kingdom="Animalia" order="Veneroida" pageId="2" pageNumber="223" phylum="Mollusca" rank="species" species="pectinatum">
<emphasis id="A9C8EADD950C426F73A2FD00FE46C088" box="[151,402,711,737]" italics="true" pageId="2" pageNumber="223">Gafrarium pectinatum</emphasis>
, Telau
</taxonomicName>
</materialsCitation>
,
<materialsCitation id="2BD43C92950C426F72DEFD00FC30C088" ID-GBIF-Occurrence="3019935318" box="[491,996,711,738]" collectingDate="2019-02-22" collectorName="Levu &amp; K. Kawai." country="Fiji" location="Viti" pageId="2" pageNumber="223" specimenCount="1">
<collectingCountry id="E3AB765F950C426F72DEFD00FDC1C088" box="[491,533,711,737]" name="Fiji" pageId="2" pageNumber="223">Viti</collectingCountry>
<collectorName id="36495319950C426F7129FD0FFD8CC08B" box="[540,600,712,738]" pageId="2" pageNumber="223">Levu</collectorName>
,
<date id="EF02100F950C426F7151FD00FCFAC088" box="[612,814,711,737]" pageId="2" pageNumber="223" value="2019-02-22">
<collectingDate id="FF46E9E7950C426F7151FD00FCFAC088" box="[612,814,711,737]" pageId="2" pageNumber="223" value="2019-02-22">22 February 2019</collectingDate>
</date>
, coll.
<collectorName id="36495319950C426F7044FD0FFC30C088" box="[881,996,711,738]" pageId="2" pageNumber="223">K. Kawai.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="9B0336CF950C426F73F2FD2CFD72C140" blockId="2.[151,1437,387,810]" pageId="2" pageNumber="223">
<materialsCitation id="2BD43C92950C426F73F2FD2CFD76C140" ID-GBIF-Occurrence="3019935312" collectingDate="1969-05-07" collectionCode="ZRC" country="Malaysia" location="Pulau Tinggi" municipality="Peninsular" pageId="2" pageNumber="223" specimenCode="ZRC 2000.2281" specimenCount="2" specimenCount-female="2" stateProvince="Johor">
<collectingMunicipality id="7B67ACB5950C426F73F2FD2CFEE9C16C" box="[199,317,747,773]" pageId="2" pageNumber="223">Peninsular</collectingMunicipality>
<collectingCountry id="E3AB765F950C426F7270FD2CFE7BC16C" box="[325,431,747,773]" name="Malaysia" pageId="2" pageNumber="223">Malaysia</collectingCountry>
.
<specimenCode id="CB1A9EB4950C426F728EFD2CFDA7C16C" box="[443,627,747,773]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="223">ZRC 2000.2281</specimenCode>
,
<specimenCount id="8DBAFD46950C426F714AFD2CFD38C16C" box="[639,748,747,773]" pageId="2" pageNumber="223" type="female">2 females</specimenCount>
(5.4×7.2, 4.7×
<quantity id="5C449B2A950C426F70A0FD2CFC24C16F" box="[917,1008,747,774]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="2" pageNumber="223" unit="mm" value="5.7">5.7 mm</quantity>
),
<location id="9E636014950C426F7731FD2CFB4CC16C" LSID="urn:lsid:plazi:treatment:131587D9950C426473A2FF50FDEFC518:9E636014950C426F7731FD2CFB4CC16C" box="[1028,1176,747,773]" country="Malaysia" municipality="Peninsular" name="Pulau Tinggi" pageId="2" pageNumber="223" stateProvince="Johor">Pulau Tinggi</location>
,
<collectingRegion id="5978F82D950C426F7796FD2CFB37C16C" box="[1187,1251,747,773]" country="Malaysia" name="Johor" pageId="2" pageNumber="223">Johor</collectingRegion>
, associated with
<taxonomicName id="5CBC4D4C950C426F73A2FCC8FDD8C140" authority="(Holten, 1802)" baseAuthorityName="Holten" baseAuthorityYear="1802" box="[151,524,783,809]" class="Bivalvia" family="Veneridae" genus="Gafrarium" kingdom="Animalia" order="Veneroida" pageId="2" pageNumber="223" phylum="Mollusca" rank="species" species="dispar">
<emphasis id="A9C8EADD950C426F73A2FCC8FE8AC140" box="[151,350,783,809]" italics="true" pageId="2" pageNumber="223">Gafrarium dispar</emphasis>
(
<bibRefCitation id="FF2D4B3E950C426F7259FCC8FDD0C140" author="Holten, H. S." box="[364,516,783,809]" pageId="2" pageNumber="223" refId="ref13550" refString="Holten, H. S. (1802) Enumeratio Systematica Conchyliae Beat. J. H. Chemnitzii Quondam Ecclesiae Zebaothu Havniae Pastoris, Plurium Societatum Sodalis p. p. quae Publica Auctione Venduntur. K. H. Seidelini, Copenhagen, vii + 88 pp." type="book" year="1802">Holten, 1802</bibRefCitation>
)
</taxonomicName>
,
<date id="EF02100F950C426F7122FCD7FD76C140" box="[535,674,783,810]" pageId="2" pageNumber="223" value="1969-05-07">
<collectingDate id="FF46E9E7950C426F7122FCD7FD76C140" box="[535,674,783,810]" pageId="2" pageNumber="223" value="1969-05-07">7 May 1969</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="CFC36647950C426F73A2F914FC3AC55A" ID-DOI="http://doi.org/10.5281/zenodo.4401463" ID-Zenodo-Dep="4401463" httpUri="https://zenodo.org/record/4401463/files/figure.png" pageId="2" pageNumber="223" startId="2.[151,250,1747,1771]" targetBox="[160,1425,839,1716]" targetPageId="2">
<paragraph id="9B0336CF950C426F73A2F914FC3AC55A" blockId="2.[151,1436,1747,1844]" pageId="2" pageNumber="223">
<emphasis id="A9C8EADD950C426F73A2F914FEC1C482" bold="true" box="[151,277,1747,1771]" pageId="2" pageNumber="223">FIGURE 1.</emphasis>
<taxonomicName id="5CBC4D4C950C426F7228F914FDBEC482" authority="(Dana, 1852)" authorityName="Komai &amp; Kawai &amp; Ng" authorityYear="2020" baseAuthorityName="Dana" baseAuthorityYear="1852" box="[285,618,1747,1772]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="223" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD950C426F7228F914FE03C482" box="[285,471,1747,1771]" italics="true" pageId="2" pageNumber="223">Arcotheres obesus</emphasis>
(
<bibRefCitation id="FF2D4B3E950C426F72D2F913FDB7C482" author="Dana, J. D." box="[487,611,1747,1772]" pageId="2" pageNumber="223" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana, 1852</bibRefCitation>
)
</taxonomicName>
. A, “male”, overall habitus in dorsal view; B, C, E, F, female, carapace, dorsal view; D, “male”, frontal view showing epistome, antennae, antennules and eyes; E, maxillipeds 3; F, left maxilliped 3; G, outer view of right chela; H, “male” pleon, outer view. After
<bibRefCitation id="FF2D4B3E950C426F71F3F8DBFC95C55A" author="Dana, J. D." box="[710,833,1819,1843]" pageId="2" pageNumber="223" refId="ref13137" refString="Dana, J. D. (1855) Atlas. Crustacea in United States Exploring Expedition during the years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 14. C. Sherman, Philadelphia, 27 pp., 96 pls." type="book" year="1855">Dana (1855</bibRefCitation>
: pl. 24 fig. 3ai).
</paragraph>
</caption>
<subSubSection id="D3A66544950C426A73F2F8A6FEF3C184" lastPageId="7" lastPageNumber="228" pageId="2" pageNumber="223" type="description">
<paragraph id="9B0336CF950C426F73F2F8A6FCFDC5AD" blockId="2.[151,1436,1889,2024]" pageId="2" pageNumber="223">
<emphasis id="A9C8EADD950C426F73F2F8A6FEA1C512" bold="true" box="[199,373,1889,1915]" pageId="2" pageNumber="223">Redescription.</emphasis>
<emphasis id="A9C8EADD950C426F724BF8A5FD7CC515" box="[382,680,1890,1916]" italics="true" pageId="2" pageNumber="223">
<typeStatus id="4407886D950C426F724BF8A5FE0EC512" box="[382,474,1890,1915]" pageId="2" pageNumber="223" type="neotype">Neotype</typeStatus>
(stage V female).
</emphasis>
Carapace (
<figureCitation id="03872A4A950C426F701EF8A5FCAFC515" box="[811,891,1890,1916]" captionStart="FIGURE 2" captionStartId="3.[151,250,1941,1965]" captionTargetBox="[151,1435,181,1917]" captionTargetId="figure-17@3.[151,1435,181,1917]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres obesus (Dana, 1852). A, E, female (5.9×8.3 mm), ZRC 2019.1876; BD, female (6.9×8.9 mm), ZRC 2019.1876. A, B, overall habitus, dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401467" httpUri="https://zenodo.org/record/4401467/files/figure.png" pageId="2" pageNumber="223">Figs. 2</figureCitation>
AD, 3A) roundly hexagonal, 1.25 times wider than long; dorsal surface gently convex, smooth, glabrous; lateral regions smooth, glabrous; front truncate, weakly projecting anteriorly beyond orbits, margin nearly straight.
</paragraph>
<paragraph id="9B0336CF950C426F73F2F809FBBBC581" blockId="2.[151,1436,1889,2024]" box="[199,1135,1998,2024]" pageId="2" pageNumber="223">
Eyes (
<figureCitation id="03872A4A950C426F7238F809FE8EC581" box="[269,346,1998,2024]" captionStart="FIGURE 2" captionStartId="3.[151,250,1941,1965]" captionTargetBox="[151,1435,181,1917]" captionTargetId="figure-17@3.[151,1435,181,1917]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres obesus (Dana, 1852). A, E, female (5.9×8.3 mm), ZRC 2019.1876; BD, female (6.9×8.9 mm), ZRC 2019.1876. A, B, overall habitus, dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401467" httpUri="https://zenodo.org/record/4401467/files/figure.png" pageId="2" pageNumber="223">Figs. 2</figureCitation>
AD, 3A, B) small, not visible in dorsal view, completely filling orbit.
</paragraph>
<caption id="CFC36647950D426E73A2F852FF1AC59C" ID-DOI="http://doi.org/10.5281/zenodo.4401467" ID-Zenodo-Dep="4401467" httpUri="https://zenodo.org/record/4401467/files/figure.png" pageId="3" pageNumber="224" startId="3.[151,250,1941,1965]" targetBox="[151,1435,181,1917]" targetPageId="3">
<paragraph id="9B0336CF950D426E73A2F852FF1AC59C" blockId="3.[151,1436,1941,2037]" pageId="3" pageNumber="224">
<emphasis id="A9C8EADD950D426E73A2F852FEC0C5C4" bold="true" box="[151,276,1941,1965]" pageId="3" pageNumber="224">FIGURE 2.</emphasis>
<taxonomicName id="5CBC4D4C950D426E722EF852FDB3C5C4" authority="(Dana, 1852)" authorityName="Komai &amp; Kawai &amp; Ng" authorityYear="2020" baseAuthorityName="Dana" baseAuthorityYear="1852" box="[283,615,1941,1965]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="224" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD950D426E722EF852FE01C5C4" box="[283,469,1941,1965]" italics="true" pageId="3" pageNumber="224">Arcotheres obesus</emphasis>
(
<bibRefCitation id="FF2D4B3E950D426E72D6F852FD8BC5C4" author="Dana, J. D." box="[483,607,1941,1965]" pageId="3" pageNumber="224" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana, 1852</bibRefCitation>
)
</taxonomicName>
. A, E, female (5.9×8.3 mm), ZRC 2019.1876; BD, female (6.9×8.9 mm), ZRC 2019.1876. A, B, overall habitus, dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view.
</paragraph>
</caption>
<paragraph id="9B0336CF950A426973F2FF50FC16C2BC" blockId="4.[151,1436,151,213]" pageId="4" pageNumber="225">
Antennular peduncle (
<figureCitation id="03872A4A950A426972F4FF50FDC3C2D8" box="[449,535,151,177]" captionStart="FIGURE 3" captionStartId="4.[151,250,1848,1872]" captionTargetBox="[162,1423,247,1813]" captionTargetId="figure-46@4.[151,1436,237,1824]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres obesus (Dana, 1852). AD, neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945; E, F, female (6.9×8.9 mm), ZRC 2019.1876; G, H, female (5.9×8.3 mm), ZRC 2019.1876. A, overall habitus, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, pleomere 6 and telson, ventral view; E, G, left maxilliped 3, outer view; F, H, left maxilliped 3, inner view." figureDoi="http://doi.org/10.5281/zenodo.4401469" httpUri="https://zenodo.org/record/4401469/files/figure.png" pageId="4" pageNumber="225">Fig. 3B</figureCitation>
) folded obliquely into antennular fossae. Antenna (
<figureCitation id="03872A4A950A42697762FF50FB78C2D8" box="[1111,1196,151,177]" captionStart="FIGURE 3" captionStartId="4.[151,250,1848,1872]" captionTargetBox="[162,1423,247,1813]" captionTargetId="figure-46@4.[151,1436,237,1824]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres obesus (Dana, 1852). AD, neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945; E, F, female (6.9×8.9 mm), ZRC 2019.1876; G, H, female (5.9×8.3 mm), ZRC 2019.1876. A, overall habitus, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, pleomere 6 and telson, ventral view; E, G, left maxilliped 3, outer view; F, H, left maxilliped 3, inner view." figureDoi="http://doi.org/10.5281/zenodo.4401469" httpUri="https://zenodo.org/record/4401469/files/figure.png" pageId="4" pageNumber="225">Fig. 3B</figureCitation>
) extending along lateral margin of front; article 2 subtrapezoidal, slightly narrowing distally.
</paragraph>
<caption id="CFC36647950A426973A2F8FFFC4FC5D5" ID-DOI="http://doi.org/10.5281/zenodo.4401469" ID-Zenodo-Dep="4401469" httpUri="https://zenodo.org/record/4401469/files/figure.png" pageId="4" pageNumber="225" startId="4.[151,250,1848,1872]" targetBox="[162,1423,247,1813]" targetPageId="4">
<paragraph id="9B0336CF950A426973A2F8FFFC4FC5D5" blockId="4.[151,1437,1848,1980]" pageId="4" pageNumber="225">
<emphasis id="A9C8EADD950A426973A2F8FFFEC7C539" bold="true" box="[151,275,1848,1872]" pageId="4" pageNumber="225">FIGURE 3.</emphasis>
<taxonomicName id="5CBC4D4C950A4269722CF8FFFDB5C539" authority="(Dana, 1852)" authorityName="Komai &amp; Kawai &amp; Ng" authorityYear="2020" baseAuthorityName="Dana" baseAuthorityYear="1852" box="[281,609,1848,1872]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="225" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD950A4269722CF8FFFE06C539" box="[281,466,1848,1872]" italics="true" pageId="4" pageNumber="225">Arcotheres obesus</emphasis>
(
<bibRefCitation id="FF2D4B3E950A426972EAF8FFFD8DC539" author="Dana, J. D." box="[479,601,1848,1872]" pageId="4" pageNumber="225" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana, 1852</bibRefCitation>
)
</taxonomicName>
. AD, neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945; E, F, female (6.9×8.9 mm), ZRC 2019.1876; G, H, female (5.9×8.3 mm), ZRC 2019.1876. A, overall habitus, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, pleomere 6 and telson, ventral view; E, G, left maxilliped 3, outer view; F, H, left maxilliped 3, inner view.
</paragraph>
</caption>
<paragraph id="9B0336CF950B426873F2FF50FE3EC328" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="226">
Maxilliped 3 (
<figureCitation id="03872A4A950B42687251FF50FE6EC2D8" box="[356,442,151,177]" captionStart="FIGURE 3" captionStartId="4.[151,250,1848,1872]" captionTargetBox="[162,1423,247,1813]" captionTargetId="figure-46@4.[151,1436,237,1824]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres obesus (Dana, 1852). AD, neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945; E, F, female (6.9×8.9 mm), ZRC 2019.1876; G, H, female (5.9×8.3 mm), ZRC 2019.1876. A, overall habitus, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, pleomere 6 and telson, ventral view; E, G, left maxilliped 3, outer view; F, H, left maxilliped 3, inner view." figureDoi="http://doi.org/10.5281/zenodo.4401469" httpUri="https://zenodo.org/record/4401469/files/figure.png" pageId="5" pageNumber="226">Fig. 3C</figureCitation>
, EH) outer surface glabrous; propodus about 2.8 times as long as high, moderately stout, distinctly longer than carpus; dactylus short, digitiform, inserted submedially on flexor margin of propodus, tip not reaching propodal apex; ischiomerus completely fused, relatively long, distinctly longer than wide, mesial margin rounded at widest point; exopod slender, about half length of ischiomerus (not illustrated for
<typeStatus id="4407886D950B426877A8FEC4FB2DC374" box="[1181,1273,259,285]" pageId="5" pageNumber="226" type="neotype">neotype</typeStatus>
, but illustrated for non-type female,
<figureCitation id="03872A4A950B426872B1FEE0FE09C328" box="[388,477,295,321]" captionStart="FIGURE 3" captionStartId="4.[151,250,1848,1872]" captionTargetBox="[162,1423,247,1813]" captionTargetId="figure-46@4.[151,1436,237,1824]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres obesus (Dana, 1852). AD, neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945; E, F, female (6.9×8.9 mm), ZRC 2019.1876; G, H, female (5.9×8.3 mm), ZRC 2019.1876. A, overall habitus, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, pleomere 6 and telson, ventral view; E, G, left maxilliped 3, outer view; F, H, left maxilliped 3, inner view." figureDoi="http://doi.org/10.5281/zenodo.4401469" httpUri="https://zenodo.org/record/4401469/files/figure.png" pageId="5" pageNumber="226">Fig. 3G</figureCitation>
).
</paragraph>
<paragraph id="9B0336CF950B426873F2FE8CFAD2C0EC" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="226">
Chelipeds (
<figureCitation id="03872A4A950B42687270FE8CFE7CC30C" box="[325,424,331,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,1848,1872]" captionTargetBox="[162,1423,247,1813]" captionTargetId="figure-46@4.[151,1436,237,1824]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres obesus (Dana, 1852). AD, neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945; E, F, female (6.9×8.9 mm), ZRC 2019.1876; G, H, female (5.9×8.3 mm), ZRC 2019.1876. A, overall habitus, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, pleomere 6 and telson, ventral view; E, G, left maxilliped 3, outer view; F, H, left maxilliped 3, inner view." figureDoi="http://doi.org/10.5281/zenodo.4401469" httpUri="https://zenodo.org/record/4401469/files/figure.png" pageId="5" pageNumber="226">Figs. 3A</figureCitation>
,
<figureCitation id="03872A4A950B42687286FE8CFE15C30C" box="[435,449,331,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1862,1886]" captionTargetBox="[226,1364,190,1827]" captionTargetId="figure-20@6.[216,1371,181,1838]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres obesus (Dana, 1852), neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945. A, right chela, outer view; B, same, inner view; C, chela and carpus of right cheliped, dorsal view; D, distal part of fixed finger of right chela, ventral view; E, carpus and merus of right cheliped, ventral view; F, right P2, dorsal view; G, same dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (shorter side), dorsal view; K, same, dactylus; L, left P4 (longer side), dorsal view; M, same dactylus; N, right P5, dorsal view; O, same dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401471" httpUri="https://zenodo.org/record/4401471/files/figure.png" pageId="5" pageNumber="226">4</figureCitation>
AE) short, symmetrical. Dactylus terminating in hooked, acute tip; upper margin rounded, gently convex; occlusal margin nearly straight, with 1 prominent, proximally directed, triangular tooth proximally and row of minute spiniform setae in distal 0.3. Palm slightly widened distally, 1.6 times as long as high; outer surface glabrous; inner surface almost glabrous, but with longitudinal row of short setae adjacent to lower margin, extending onto fixed finger, becoming multiple rows. Fixed finger terminating in hooked, acute tip, bearing short longitudinal row of stiff setae on inner surface distally (
<figureCitation id="03872A4A950B42687034FE38FC8EC070" box="[769,858,511,537]" captionStart="FIGURE 2" captionStartId="3.[151,250,1941,1965]" captionTargetBox="[151,1435,181,1917]" captionTargetId="figure-17@3.[151,1435,181,1917]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres obesus (Dana, 1852). A, E, female (5.9×8.3 mm), ZRC 2019.1876; BD, female (6.9×8.9 mm), ZRC 2019.1876. A, B, overall habitus, dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401467" httpUri="https://zenodo.org/record/4401467/files/figure.png" pageId="5" pageNumber="226">Fig. 2D</figureCitation>
); occlusal margin nearly straight, with 1 small tooth proximally, hiatus proximal to occlusal tooth accommodating occlusal tooth on dactylus, and with comb-like row of minute spiniform setae distal to tooth. Carpus short, subcylindrical, glabrous. Merus slightly curved inwards, slightly compressed laterally, outer surface convex, glabrous; inner surface with scattered short setae.
</paragraph>
<paragraph id="9B0336CF950B426873F2FD48FE09C6AC" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="226">
P2, P3 and P5 symmetrical, P4 asymmetrical; propodus to merus subcylindrical; pereopod 4 longest; relative length of merus P 4&gt;P3&gt;P2&gt;P5. P2 (
<figureCitation id="03872A4A950B4268710EFD74FD74C0A4" box="[571,672,691,717]" captionStart="FIGURE 3" captionStartId="4.[151,250,1848,1872]" captionTargetBox="[162,1423,247,1813]" captionTargetId="figure-46@4.[151,1436,237,1824]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres obesus (Dana, 1852). AD, neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945; E, F, female (6.9×8.9 mm), ZRC 2019.1876; G, H, female (5.9×8.3 mm), ZRC 2019.1876. A, overall habitus, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, pleomere 6 and telson, ventral view; E, G, left maxilliped 3, outer view; F, H, left maxilliped 3, inner view." figureDoi="http://doi.org/10.5281/zenodo.4401469" httpUri="https://zenodo.org/record/4401469/files/figure.png" pageId="5" pageNumber="226">Figs. 3A</figureCitation>
,
<figureCitation id="03872A4A950B42687198FD74FD38C0A4" box="[685,748,691,717]" captionStart="FIGURE 4" captionStartId="6.[151,250,1862,1886]" captionTargetBox="[226,1364,190,1827]" captionTargetId="figure-20@6.[216,1371,181,1838]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres obesus (Dana, 1852), neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945. A, right chela, outer view; B, same, inner view; C, chela and carpus of right cheliped, dorsal view; D, distal part of fixed finger of right chela, ventral view; E, carpus and merus of right cheliped, ventral view; F, right P2, dorsal view; G, same dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (shorter side), dorsal view; K, same, dactylus; L, left P4 (longer side), dorsal view; M, same dactylus; N, right P5, dorsal view; O, same dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401471" httpUri="https://zenodo.org/record/4401471/files/figure.png" pageId="5" pageNumber="226">4F, G</figureCitation>
) moderately long, slender, entirely glabrous; dactylus gently curved, 0.6 times as long as propodus; carpus curved at base; merus almost straight. Pereopod 3 (
<figureCitation id="03872A4A950B426877D2FD10FA9FC098" box="[1255,1355,727,753]" captionStart="FIGURE 2" captionStartId="3.[151,250,1941,1965]" captionTargetBox="[151,1435,181,1917]" captionTargetId="figure-17@3.[151,1435,181,1917]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres obesus (Dana, 1852). A, E, female (5.9×8.3 mm), ZRC 2019.1876; BD, female (6.9×8.9 mm), ZRC 2019.1876. A, B, overall habitus, dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401467" httpUri="https://zenodo.org/record/4401467/files/figure.png" pageId="5" pageNumber="226">Figs. 2A</figureCitation>
,
<figureCitation id="03872A4A950B4268766DFD10FA47C098" box="[1368,1427,727,753]" captionStart="FIGURE 3" captionStartId="4.[151,250,1848,1872]" captionTargetBox="[162,1423,247,1813]" captionTargetId="figure-46@4.[151,1436,237,1824]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres obesus (Dana, 1852). AD, neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945; E, F, female (6.9×8.9 mm), ZRC 2019.1876; G, H, female (5.9×8.3 mm), ZRC 2019.1876. A, overall habitus, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, pleomere 6 and telson, ventral view; E, G, left maxilliped 3, outer view; F, H, left maxilliped 3, inner view." figureDoi="http://doi.org/10.5281/zenodo.4401469" httpUri="https://zenodo.org/record/4401469/files/figure.png" pageId="5" pageNumber="226">3H, I</figureCitation>
) moderately long, slender, entirely glabrous, slightly longer than P2, but otherwise similar to it; dactylus slightly curved, about half-length of propodus; merus faintly curved. Shorter P4 (
<figureCitation id="03872A4A950B426870F7FCD8FBF0C150" box="[962,1060,799,825]" captionStart="FIGURE 3" captionStartId="4.[151,250,1848,1872]" captionTargetBox="[162,1423,247,1813]" captionTargetId="figure-46@4.[151,1436,237,1824]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres obesus (Dana, 1852). AD, neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945; E, F, female (6.9×8.9 mm), ZRC 2019.1876; G, H, female (5.9×8.3 mm), ZRC 2019.1876. A, overall habitus, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, pleomere 6 and telson, ventral view; E, G, left maxilliped 3, outer view; F, H, left maxilliped 3, inner view." figureDoi="http://doi.org/10.5281/zenodo.4401469" httpUri="https://zenodo.org/record/4401469/files/figure.png" pageId="5" pageNumber="226">Figs. 3A</figureCitation>
,
<figureCitation id="03872A4A950B42687705FCD8FBB8C150" box="[1072,1132,799,825]" captionStart="FIGURE 4" captionStartId="6.[151,250,1862,1886]" captionTargetBox="[226,1364,190,1827]" captionTargetId="figure-20@6.[216,1371,181,1838]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres obesus (Dana, 1852), neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945. A, right chela, outer view; B, same, inner view; C, chela and carpus of right cheliped, dorsal view; D, distal part of fixed finger of right chela, ventral view; E, carpus and merus of right cheliped, ventral view; F, right P2, dorsal view; G, same dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (shorter side), dorsal view; K, same, dactylus; L, left P4 (longer side), dorsal view; M, same dactylus; N, right P5, dorsal view; O, same dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401471" httpUri="https://zenodo.org/record/4401471/files/figure.png" pageId="5" pageNumber="226">4J, K</figureCitation>
) dactylus about half-length of propodus, glabrous except for sparse short setae on flexor margin; propodus almost straight, glabrous except for row of sparse setae on flexor margin (shorter than those on right P4); carpus and merus glabrous, merus slightly curved. Longer P4 (
<figureCitation id="03872A4A950B42687243FC4CFE0DC1CC" box="[374,473,907,933]" captionStart="FIGURE 3" captionStartId="4.[151,250,1848,1872]" captionTargetBox="[162,1423,247,1813]" captionTargetId="figure-46@4.[151,1436,237,1824]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres obesus (Dana, 1852). AD, neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945; E, F, female (6.9×8.9 mm), ZRC 2019.1876; G, H, female (5.9×8.3 mm), ZRC 2019.1876. A, overall habitus, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, pleomere 6 and telson, ventral view; E, G, left maxilliped 3, outer view; F, H, left maxilliped 3, inner view." figureDoi="http://doi.org/10.5281/zenodo.4401469" httpUri="https://zenodo.org/record/4401469/files/figure.png" pageId="5" pageNumber="226">Figs. 3A</figureCitation>
,
<figureCitation id="03872A4A950B426872D0FC4CFDF8C1CC" box="[485,556,907,933]" captionStart="FIGURE 4" captionStartId="6.[151,250,1862,1886]" captionTargetBox="[226,1364,190,1827]" captionTargetId="figure-20@6.[216,1371,181,1838]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres obesus (Dana, 1852), neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945. A, right chela, outer view; B, same, inner view; C, chela and carpus of right cheliped, dorsal view; D, distal part of fixed finger of right chela, ventral view; E, carpus and merus of right cheliped, ventral view; F, right P2, dorsal view; G, same dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (shorter side), dorsal view; K, same, dactylus; L, left P4 (longer side), dorsal view; M, same dactylus; N, right P5, dorsal view; O, same dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401471" httpUri="https://zenodo.org/record/4401471/files/figure.png" pageId="5" pageNumber="226">4L, M</figureCitation>
) about 1.2 times as long as shorter P4; dactylus about half-length of propodus, with row of sparse short setae on extensor margin, scattered very short setae on outer surface and row of mixture of short and long setae on flexor margin; propodus faintly curved, glabrous except for row of sparse long setae on flexor margin; carpus glabrous; merus slightly curved. P5 (
<figureCitation id="03872A4A950B42687004FC30FC41C678" box="[817,917,1015,1041]" captionStart="FIGURE 3" captionStartId="4.[151,250,1848,1872]" captionTargetBox="[162,1423,247,1813]" captionTargetId="figure-46@4.[151,1436,237,1824]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres obesus (Dana, 1852). AD, neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945; E, F, female (6.9×8.9 mm), ZRC 2019.1876; G, H, female (5.9×8.3 mm), ZRC 2019.1876. A, overall habitus, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, pleomere 6 and telson, ventral view; E, G, left maxilliped 3, outer view; F, H, left maxilliped 3, inner view." figureDoi="http://doi.org/10.5281/zenodo.4401469" httpUri="https://zenodo.org/record/4401469/files/figure.png" pageId="5" pageNumber="226">Figs. 3A</figureCitation>
,
<figureCitation id="03872A4A950B42687097FC30FC3CC678" box="[930,1000,1015,1041]" captionStart="FIGURE 4" captionStartId="6.[151,250,1862,1886]" captionTargetBox="[226,1364,190,1827]" captionTargetId="figure-20@6.[216,1371,181,1838]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres obesus (Dana, 1852), neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945. A, right chela, outer view; B, same, inner view; C, chela and carpus of right cheliped, dorsal view; D, distal part of fixed finger of right chela, ventral view; E, carpus and merus of right cheliped, ventral view; F, right P2, dorsal view; G, same dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (shorter side), dorsal view; K, same, dactylus; L, left P4 (longer side), dorsal view; M, same dactylus; N, right P5, dorsal view; O, same dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401471" httpUri="https://zenodo.org/record/4401471/files/figure.png" pageId="5" pageNumber="226">4N, O</figureCitation>
) dactylus longest, 0.8 times as long as propodus, approximately 6 times as long as wide, generally gently arcuate but with substantial curvature near base; extensor margin and upper and lower surfaces glabrous, but flexor margin with row of moderately long setae in distal 0.6, more closely spaced distally, not extending to tip, no comb-like rows of minute spiniform setae; propodus almost glabrous except for few setae on flexor margin; carpus glabrous, substantially curved; merus slightly narrowing proximally, glabrous.
</paragraph>
<paragraph id="9B0336CF950B426873F2FB08FD04C764" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="226">
Pleon (
<figureCitation id="03872A4A950B42687223FB08FE44C680" box="[278,400,1231,1257]" captionStart="FIGURE 3" captionStartId="4.[151,250,1848,1872]" captionTargetBox="[162,1423,247,1813]" captionTargetId="figure-46@4.[151,1436,237,1824]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres obesus (Dana, 1852). AD, neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945; E, F, female (6.9×8.9 mm), ZRC 2019.1876; G, H, female (5.9×8.3 mm), ZRC 2019.1876. A, overall habitus, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, pleomere 6 and telson, ventral view; E, G, left maxilliped 3, outer view; F, H, left maxilliped 3, inner view." figureDoi="http://doi.org/10.5281/zenodo.4401469" httpUri="https://zenodo.org/record/4401469/files/figure.png" pageId="5" pageNumber="226">Fig. 3A, D</figureCitation>
) extending to buccal region, covering bases of ambulatory legs; telson (
<figureCitation id="03872A4A950B42687783FB08FAD9C680" box="[1206,1293,1231,1257]" captionStart="FIGURE 3" captionStartId="4.[151,250,1848,1872]" captionTargetBox="[162,1423,247,1813]" captionTargetId="figure-46@4.[151,1436,237,1824]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres obesus (Dana, 1852). AD, neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945; E, F, female (6.9×8.9 mm), ZRC 2019.1876; G, H, female (5.9×8.3 mm), ZRC 2019.1876. A, overall habitus, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, pleomere 6 and telson, ventral view; E, G, left maxilliped 3, outer view; F, H, left maxilliped 3, inner view." figureDoi="http://doi.org/10.5281/zenodo.4401469" httpUri="https://zenodo.org/record/4401469/files/figure.png" pageId="5" pageNumber="226">Fig. 3D</figureCitation>
) very broad, partially recessed into distal margin of pleomere 6.
</paragraph>
<paragraph id="9B0336CF950B426873F2FAD0FBD9C758" blockId="5.[151,1437,151,2013]" box="[199,1037,1303,1329]" pageId="5" pageNumber="226">
<emphasis id="A9C8EADD950B426873F2FAD0FEB5C758" box="[199,353,1303,1329]" italics="true" pageId="5" pageNumber="226">Adult females</emphasis>
. Similar to
<typeStatus id="4407886D950B426872D0FAD0FD95C758" box="[485,577,1303,1329]" pageId="5" pageNumber="226" type="neotype">neotype</typeStatus>
. Carapace 1.21.3 times as long as wide.
</paragraph>
<paragraph id="9B0336CF950B426873F2FAFCFBB6C7F4" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="226">
<emphasis id="A9C8EADD950B426873F2FAFCFE60C73D" box="[199,436,1339,1365]" italics="true" pageId="5" pageNumber="226">Male post-hard stage</emphasis>
. Cuticle of body and appendages not strongly calcified but well chitinised. Carapace (
<figureCitation id="03872A4A950B4268765AFAFCFF6EC710" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[157,1424,499,1834]" captionTargetId="figure-186@8.[151,1436,489,1844]" captionTargetPageId="8" captionText="FIGURE 5. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, habitus, dorsal view; B, left MXP3, outer view; C, left thoracic sternum, ventral view; D, pleon, telson and exposed part of sternite 8 (left), ventral view; E, left G1, outer view; F, same, inner view; G, left G2, outer view; H, same, different view." figureDoi="http://doi.org/10.5281/zenodo.4401473" httpUri="https://zenodo.org/record/4401473/files/figure.png" pageId="5" pageNumber="226">Fig. 5A</figureCitation>
) roundly suboctagonal, 1.1 times wider than long; dorsal surface and lateral regions mostly smooth, glabrous; front weakly projecting anteriorly beyond orbits, margin nearly straight in dorsal view.
</paragraph>
<paragraph id="9B0336CF950B426873F2FA60FB4BC7A8" blockId="5.[151,1437,151,2013]" box="[199,1183,1446,1473]" pageId="5" pageNumber="226">
Eyes (
<figureCitation id="03872A4A950B42687238FA60FEB0C7A8" box="[269,356,1446,1473]" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[157,1424,499,1834]" captionTargetId="figure-186@8.[151,1436,489,1844]" captionTargetPageId="8" captionText="FIGURE 5. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, habitus, dorsal view; B, left MXP3, outer view; C, left thoracic sternum, ventral view; D, pleon, telson and exposed part of sternite 8 (left), ventral view; E, left G1, outer view; F, same, inner view; G, left G2, outer view; H, same, different view." figureDoi="http://doi.org/10.5281/zenodo.4401473" httpUri="https://zenodo.org/record/4401473/files/figure.png" pageId="5" pageNumber="226">Fig. 5A</figureCitation>
) moderately small, clearly visible in dorsal view, completely filling orbit.
</paragraph>
<paragraph id="9B0336CF950B426873F2FA0CFD3FC78C" blockId="5.[151,1437,151,2013]" box="[199,747,1483,1509]" pageId="5" pageNumber="226">
Maxilliped 3 (
<figureCitation id="03872A4A950B42687253FA0CFE68C78C" box="[358,444,1483,1509]" captionStart="FIGURE 3" captionStartId="4.[151,250,1848,1872]" captionTargetBox="[162,1423,247,1813]" captionTargetId="figure-46@4.[151,1436,237,1824]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres obesus (Dana, 1852). AD, neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945; E, F, female (6.9×8.9 mm), ZRC 2019.1876; G, H, female (5.9×8.3 mm), ZRC 2019.1876. A, overall habitus, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, pleomere 6 and telson, ventral view; E, G, left maxilliped 3, outer view; F, H, left maxilliped 3, inner view." figureDoi="http://doi.org/10.5281/zenodo.4401469" httpUri="https://zenodo.org/record/4401469/files/figure.png" pageId="5" pageNumber="226">Fig. 3B</figureCitation>
) similar to that of females.
</paragraph>
<paragraph id="9B0336CF950B426873F2FA28FB77C540" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="226">
Chelipeds (
<figureCitation id="03872A4A950B42687273FA28FE7DC460" box="[326,425,1518,1545]" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[157,1424,499,1834]" captionTargetId="figure-186@8.[151,1436,489,1844]" captionTargetPageId="8" captionText="FIGURE 5. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, habitus, dorsal view; B, left MXP3, outer view; C, left thoracic sternum, ventral view; D, pleon, telson and exposed part of sternite 8 (left), ventral view; E, left G1, outer view; F, same, inner view; G, left G2, outer view; H, same, different view." figureDoi="http://doi.org/10.5281/zenodo.4401473" httpUri="https://zenodo.org/record/4401473/files/figure.png" pageId="5" pageNumber="226">Figs. 5A</figureCitation>
,
<figureCitation id="03872A4A950B42687280FA28FE16C463" box="[437,450,1519,1546]" captionStart="FIGURE 6" captionStartId="10.[151,250,1791,1815]" captionTargetBox="[166,1424,193,1760]" captionTargetId="figure-20@10.[151,1436,181,1768]" captionTargetPageId="10" captionText="FIGURE 6. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, left chela, outer view; B, same, inner view; C, chela and carpus of left cheliped, dorsal view; D, merus of left cheliped, ventral view; E, left P2, dorsal view; F, same, dactylus; G, left P3, dorsal view; H, same, dactylus; I, left P4 (longer side), dorsal view; J, same, dactylus; K, right P4 (shorter side), dorsal view; L, same, dactylus and propodus, dorsal (perpendicular) view; M, right P5, dorsal view; N, same, dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401475" httpUri="https://zenodo.org/record/4401475/files/figure.png" pageId="5" pageNumber="226">6</figureCitation>
AD) symmetrical, proportionately larger compared with females. Dactylus 0.6 times as long as palm, terminating in hooked, acute tip; upper margin rounded, gently convex; occlusal margin with 1 conspicuous triangular tooth proximally but otherwise apparently smooth. Palm slightly widened distally, 2.2 times as long as high; outer surface glabrous; inner surface almost glabrous, but with longitudinal row of short setae adjacent to lower margin, extending onto fixed finger, becoming multiple rows. Fixed finger terminating in hooked, acute tip; occlusal margin with 1 small tooth proximal to midlength, hiatus proximal to occlusal tooth accommodating occlusal tooth on dactylus, and with comb-like row of minute spiniform setae distal to tooth. Carpus distinctly shorter than palm, widened distally, glabrous. Merus slightly curved inward, slightly compressed laterally; outer surface convex, glabrous, lower distal angle somewhat produced; inner surface with few short setae.
</paragraph>
<paragraph id="9B0336CF950B426A73F2F8F4FA57C3C4" blockId="5.[151,1437,151,2013]" lastBlockId="7.[151,1437,151,2013]" lastPageId="7" lastPageNumber="228" pageId="5" pageNumber="226">
P2, P3 and P5 symmetrical, P4 asymmetrical (left longer than right), P4 longest; relative length of merus P4&gt;P3&gt;P2&gt;P5. P2 (
<figureCitation id="03872A4A950B4268724EF890FE34C518" box="[379,480,1878,1905]" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[157,1424,499,1834]" captionTargetId="figure-186@8.[151,1436,489,1844]" captionTargetPageId="8" captionText="FIGURE 5. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, habitus, dorsal view; B, left MXP3, outer view; C, left thoracic sternum, ventral view; D, pleon, telson and exposed part of sternite 8 (left), ventral view; E, left G1, outer view; F, same, inner view; G, left G2, outer view; H, same, different view." figureDoi="http://doi.org/10.5281/zenodo.4401473" httpUri="https://zenodo.org/record/4401473/files/figure.png" pageId="5" pageNumber="226">Figs. 5A</figureCitation>
,
<figureCitation id="03872A4A950B426872D8F890FDF9C518" box="[493,557,1879,1905]" captionStart="FIGURE 6" captionStartId="10.[151,250,1791,1815]" captionTargetBox="[166,1424,193,1760]" captionTargetId="figure-20@10.[151,1436,181,1768]" captionTargetPageId="10" captionText="FIGURE 6. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, left chela, outer view; B, same, inner view; C, chela and carpus of left cheliped, dorsal view; D, merus of left cheliped, ventral view; E, left P2, dorsal view; F, same, dactylus; G, left P3, dorsal view; H, same, dactylus; I, left P4 (longer side), dorsal view; J, same, dactylus; K, right P4 (shorter side), dorsal view; L, same, dactylus and propodus, dorsal (perpendicular) view; M, right P5, dorsal view; N, same, dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401475" httpUri="https://zenodo.org/record/4401475/files/figure.png" pageId="5" pageNumber="226">6E, F</figureCitation>
) moderately long, stout, entirely glabrous; dactylus curved distally, 0.6 times as long as propodus; merus almost straight. P3 (
<figureCitation id="03872A4A950B42687186F8BCFCCCC5FC" box="[691,792,1914,1941]" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[157,1424,499,1834]" captionTargetId="figure-186@8.[151,1436,489,1844]" captionTargetPageId="8" captionText="FIGURE 5. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, habitus, dorsal view; B, left MXP3, outer view; C, left thoracic sternum, ventral view; D, pleon, telson and exposed part of sternite 8 (left), ventral view; E, left G1, outer view; F, same, inner view; G, left G2, outer view; H, same, different view." figureDoi="http://doi.org/10.5281/zenodo.4401473" httpUri="https://zenodo.org/record/4401473/files/figure.png" pageId="5" pageNumber="226">Figs. 5A</figureCitation>
,
<figureCitation id="03872A4A950B42687011F8BCFCBEC5FC" box="[804,874,1915,1941]" captionStart="FIGURE 6" captionStartId="10.[151,250,1791,1815]" captionTargetBox="[166,1424,193,1760]" captionTargetId="figure-20@10.[151,1436,181,1768]" captionTargetPageId="10" captionText="FIGURE 6. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, left chela, outer view; B, same, inner view; C, chela and carpus of left cheliped, dorsal view; D, merus of left cheliped, ventral view; E, left P2, dorsal view; F, same, dactylus; G, left P3, dorsal view; H, same, dactylus; I, left P4 (longer side), dorsal view; J, same, dactylus; K, right P4 (shorter side), dorsal view; L, same, dactylus and propodus, dorsal (perpendicular) view; M, right P5, dorsal view; N, same, dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401475" httpUri="https://zenodo.org/record/4401475/files/figure.png" pageId="5" pageNumber="226">6G, H</figureCitation>
) entirely glabrous; dactylus curved distally, about half-length of propodus; merus faintly curved. Shorter (right) P4 (
<figureCitation id="03872A4A950B426870BEF858FC24C5D0" box="[907,1008,1950,1977]" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[157,1424,499,1834]" captionTargetId="figure-186@8.[151,1436,489,1844]" captionTargetPageId="8" captionText="FIGURE 5. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, habitus, dorsal view; B, left MXP3, outer view; C, left thoracic sternum, ventral view; D, pleon, telson and exposed part of sternite 8 (left), ventral view; E, left G1, outer view; F, same, inner view; G, left G2, outer view; H, same, different view." figureDoi="http://doi.org/10.5281/zenodo.4401473" httpUri="https://zenodo.org/record/4401473/files/figure.png" pageId="5" pageNumber="226">Figs. 5A</figureCitation>
,
<figureCitation id="03872A4A950B426870CBF858FB96C5D0" box="[1022,1090,1951,1977]" captionStart="FIGURE 6" captionStartId="10.[151,250,1791,1815]" captionTargetBox="[166,1424,193,1760]" captionTargetId="figure-20@10.[151,1436,181,1768]" captionTargetPageId="10" captionText="FIGURE 6. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, left chela, outer view; B, same, inner view; C, chela and carpus of left cheliped, dorsal view; D, merus of left cheliped, ventral view; E, left P2, dorsal view; F, same, dactylus; G, left P3, dorsal view; H, same, dactylus; I, left P4 (longer side), dorsal view; J, same, dactylus; K, right P4 (shorter side), dorsal view; L, same, dactylus and propodus, dorsal (perpendicular) view; M, right P5, dorsal view; N, same, dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401475" httpUri="https://zenodo.org/record/4401475/files/figure.png" pageId="5" pageNumber="226">6K, L</figureCitation>
) dactylus about half-length of propodus, glabrous except for 4 widely spaced, long setae on flexor margin and some very short setae on extensor margin; propodus slightly arcuate, almost glabrous except for 2 long setae on distal part of flexor margin. Longer (left) P4 (
<figureCitation id="03872A4A9509426A7236FF7CFEB3C2BC" box="[259,359,186,213]" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[157,1424,499,1834]" captionTargetId="figure-186@8.[151,1436,489,1844]" captionTargetPageId="8" captionText="FIGURE 5. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, habitus, dorsal view; B, left MXP3, outer view; C, left thoracic sternum, ventral view; D, pleon, telson and exposed part of sternite 8 (left), ventral view; E, left G1, outer view; F, same, inner view; G, left G2, outer view; H, same, different view." figureDoi="http://doi.org/10.5281/zenodo.4401473" httpUri="https://zenodo.org/record/4401473/files/figure.png" pageId="7" pageNumber="228">Figs. 5A</figureCitation>
,
<figureCitation id="03872A4A9509426A7247FF7CFE70C2BC" box="[370,420,187,213]" captionStart="FIGURE 6" captionStartId="10.[151,250,1791,1815]" captionTargetBox="[166,1424,193,1760]" captionTargetId="figure-20@10.[151,1436,181,1768]" captionTargetPageId="10" captionText="FIGURE 6. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, left chela, outer view; B, same, inner view; C, chela and carpus of left cheliped, dorsal view; D, merus of left cheliped, ventral view; E, left P2, dorsal view; F, same, dactylus; G, left P3, dorsal view; H, same, dactylus; I, left P4 (longer side), dorsal view; J, same, dactylus; K, right P4 (shorter side), dorsal view; L, same, dactylus and propodus, dorsal (perpendicular) view; M, right P5, dorsal view; N, same, dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401475" httpUri="https://zenodo.org/record/4401475/files/figure.png" pageId="7" pageNumber="228">6I, J</figureCitation>
) about 1.2 times as long as shorter (right) P4; dactylus about half-length of propodus, with row of sparse short setae on extensor margin and 3 long and few short setae on flexor margin; propodus faintly curved, almost glabrous except for 3 long setae on distal 0.3 of flexor margin; carpus and glabrous, merus nearly straight. P5 (
<figureCitation id="03872A4A9509426A721CFEE0FE5AC328" box="[297,398,294,321]" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[157,1424,499,1834]" captionTargetId="figure-186@8.[151,1436,489,1844]" captionTargetPageId="8" captionText="FIGURE 5. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, habitus, dorsal view; B, left MXP3, outer view; C, left thoracic sternum, ventral view; D, pleon, telson and exposed part of sternite 8 (left), ventral view; E, left G1, outer view; F, same, inner view; G, left G2, outer view; H, same, different view." figureDoi="http://doi.org/10.5281/zenodo.4401473" httpUri="https://zenodo.org/record/4401473/files/figure.png" pageId="7" pageNumber="228">Figs. 5A</figureCitation>
,
<figureCitation id="03872A4A9509426A72AEFEE0FE33C328" box="[411,487,295,321]" captionStart="FIGURE 6" captionStartId="10.[151,250,1791,1815]" captionTargetBox="[166,1424,193,1760]" captionTargetId="figure-20@10.[151,1436,181,1768]" captionTargetPageId="10" captionText="FIGURE 6. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, left chela, outer view; B, same, inner view; C, chela and carpus of left cheliped, dorsal view; D, merus of left cheliped, ventral view; E, left P2, dorsal view; F, same, dactylus; G, left P3, dorsal view; H, same, dactylus; I, left P4 (longer side), dorsal view; J, same, dactylus; K, right P4 (shorter side), dorsal view; L, same, dactylus and propodus, dorsal (perpendicular) view; M, right P5, dorsal view; N, same, dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401475" httpUri="https://zenodo.org/record/4401475/files/figure.png" pageId="7" pageNumber="228">6M, N</figureCitation>
) shortest; dactylus distinctly longer than dactyli of P24, arcuate, 0.7 times as long as propodus, approximately 4 times as long as wide; extensor margin and upper and lower surfaces glabrous, but flexor margin with row of well-spaced, moderately long setae over entire length; propodus almost glabrous except for several setae on flexor margin; carpus glabrous, nearly straight; merus subcylindrical, straight, also glabrous.
</paragraph>
<caption id="CFC366479508426B73A2F881FD3CC587" ID-DOI="http://doi.org/10.5281/zenodo.4401471" ID-Zenodo-Dep="4401471" httpUri="https://zenodo.org/record/4401471/files/figure.png" pageId="6" pageNumber="227" startId="6.[151,250,1862,1886]" targetBox="[226,1364,190,1827]" targetPageId="6">
<paragraph id="9B0336CF9508426B73A2F881FD3CC587" blockId="6.[151,1436,1862,2030]" pageId="6" pageNumber="227">
<emphasis id="A9C8EADD9508426B73A2F881FEC1C537" bold="true" box="[151,277,1862,1886]" pageId="6" pageNumber="227">FIGURE 4.</emphasis>
<taxonomicName id="5CBC4D4C9508426B7228F881FDBCC537" authority="(Dana, 1852)" authorityName="Komai &amp; Kawai &amp; Ng" authorityYear="2020" baseAuthorityName="Dana" baseAuthorityYear="1852" box="[285,616,1862,1886]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="227" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD9508426B7228F881FE03C537" box="[285,471,1862,1886]" italics="true" pageId="6" pageNumber="227">Arcotheres obesus</emphasis>
(
<bibRefCitation id="FF2D4B3E9508426B72D3F881FDB5C537" author="Dana, J. D." box="[486,609,1862,1886]" pageId="6" pageNumber="227" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana, 1852</bibRefCitation>
)
</taxonomicName>
, neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945. A, right chela, outer view; B, same, inner view; C, chela and carpus of right cheliped, dorsal view; D, distal part of fixed finger of right chela, ventral view; E, carpus and merus of right cheliped, ventral view; F, right P2, dorsal view; G, same dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (shorter side), dorsal view; K, same, dactylus; L, left P4 (longer side), dorsal view; M, same dactylus; N, right P5, dorsal view; O, same dactylus.
</paragraph>
</caption>
<paragraph id="9B0336CF9509426A73F2FE70FB42C0C0" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="228">
Thoracic sternum (
<figureCitation id="03872A4A9509426A7296FE70FE28C3B8" box="[419,508,438,465]" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[157,1424,499,1834]" captionTargetId="figure-186@8.[151,1436,489,1844]" captionTargetPageId="8" captionText="FIGURE 5. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, habitus, dorsal view; B, left MXP3, outer view; C, left thoracic sternum, ventral view; D, pleon, telson and exposed part of sternite 8 (left), ventral view; E, left G1, outer view; F, same, inner view; G, left G2, outer view; H, same, different view." figureDoi="http://doi.org/10.5281/zenodo.4401473" httpUri="https://zenodo.org/record/4401473/files/figure.png" pageId="7" pageNumber="228">Fig. 5C</figureCitation>
) broad. Sternites 3 and 4 fused, only slit-like incision still discernible in broad concavity at base of maxilliped 3; anteromedian part produced into broadly rounded lobe, anterolateral angle also produced to rounded lobe; lateral margins concave, accommodating coxae of chelipeds; anterior surface shallowly sulcate. Exposed parts of sternites 57 nearly smooth; sternite 5 posterolaterally with distinct episternite; sternites 6 and 7 each with shallow notch on lateral margin; sternite 8 clearly visible in lateral part, with concave lateral margin. Sterno-pleonal cavity deep, defined by sharply delimited margin, narrowing anteriorly, reaching anteriorly to midlength of fused sternites 3/4; rounded press button present on sternite 5 inside of cavity.
</paragraph>
<paragraph id="9B0336CF9509426A73F2FD74FEF8C17C" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="228">
Pleon (
<figureCitation id="03872A4A9509426A722DFD74FEA6C0A4" box="[280,370,690,717]" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[157,1424,499,1834]" captionTargetId="figure-186@8.[151,1436,489,1844]" captionTargetPageId="8" captionText="FIGURE 5. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, habitus, dorsal view; B, left MXP3, outer view; C, left thoracic sternum, ventral view; D, pleon, telson and exposed part of sternite 8 (left), ventral view; E, left G1, outer view; F, same, inner view; G, left G2, outer view; H, same, different view." figureDoi="http://doi.org/10.5281/zenodo.4401473" httpUri="https://zenodo.org/record/4401473/files/figure.png" pageId="7" pageNumber="228">Fig. 5D</figureCitation>
) moderately narrow, tapering distally from pleomere 4 to telson. Pleomeres 1 and 2 as wide, longitudinally very short. Pleomere 3 oblong. Pleomeres 46 all subtrapezoidal. Telson semicircular, 1.7 times as wide as long.
</paragraph>
<paragraph id="9B0336CF9509426A73F2FCD8FC49C1E8" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="228">
G1 (
<figureCitation id="03872A4A9509426A73CDFCD8FEBDC150" box="[248,361,798,825]" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[157,1424,499,1834]" captionTargetId="figure-186@8.[151,1436,489,1844]" captionTargetPageId="8" captionText="FIGURE 5. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, habitus, dorsal view; B, left MXP3, outer view; C, left thoracic sternum, ventral view; D, pleon, telson and exposed part of sternite 8 (left), ventral view; E, left G1, outer view; F, same, inner view; G, left G2, outer view; H, same, different view." figureDoi="http://doi.org/10.5281/zenodo.4401473" httpUri="https://zenodo.org/record/4401473/files/figure.png" pageId="7" pageNumber="228">Fig. 5E, F</figureCitation>
) noticeably curved at midlength, gradually tapering, with row of sparse short setae laterally; distal part subtruncate, tip subacutely pointed. G2 (
<figureCitation id="03872A4A9509426A7185FC84FCFDC134" box="[688,809,834,861]" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[157,1424,499,1834]" captionTargetId="figure-186@8.[151,1436,489,1844]" captionTargetPageId="8" captionText="FIGURE 5. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, habitus, dorsal view; B, left MXP3, outer view; C, left thoracic sternum, ventral view; D, pleon, telson and exposed part of sternite 8 (left), ventral view; E, left G1, outer view; F, same, inner view; G, left G2, outer view; H, same, different view." figureDoi="http://doi.org/10.5281/zenodo.4401473" httpUri="https://zenodo.org/record/4401473/files/figure.png" pageId="7" pageNumber="228">Fig. 5G, H</figureCitation>
) endopod somewhat flattened, tapering distally; exopod about half-length of endopod, not tapering, terminal margin rounded.
</paragraph>
<paragraph id="9B0336CF9509426A73F2FC4DFE27C1CC" blockId="7.[151,1437,151,2013]" box="[199,499,906,933]" pageId="7" pageNumber="228">
<emphasis id="A9C8EADD9509426A73F2FC4DFE89C1CD" bold="true" box="[199,349,906,932]" pageId="7" pageNumber="228">Color in life.</emphasis>
Unavailable.
</paragraph>
<paragraph id="9B0336CF9509426A73F2FC69FEF3C184" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="228">
<emphasis id="A9C8EADD9509426A73F2FC69FF23C1A1" bold="true" box="[199,247,942,968]" pageId="7" pageNumber="228">Size</emphasis>
. Largest male in post-hard stage 2.2×
<quantity id="5C449B2A9509426A7186FC68FCC4C1A0" box="[691,784,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="7" pageNumber="228" unit="mm" value="2.6">2.6 mm</quantity>
; ovigerous females 4.6×5.95.4×
<quantity id="5C449B2A9509426A77A5FC69FB38C1A0" box="[1168,1260,942,969]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="7" pageNumber="228" unit="mm" value="6.7">6.7 mm</quantity>
; largest female 7.2×
<quantity id="5C449B2A9509426A73FCFC14FEF7C184" box="[201,291,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="7" pageNumber="228" unit="mm" value="9.2">9.2 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="D3A665449509426A73F2FC31FDF2C65C" pageId="7" pageNumber="228" type="distribution">
<paragraph id="9B0336CF9509426A73F2FC31FDF2C65C" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="228">
<emphasis id="A9C8EADD9509426A73F2FC31FEB5C679" bold="true" box="[199,353,1014,1040]" pageId="7" pageNumber="228">Distribution.</emphasis>
Presently known from
<collectingCountry id="E3AB765F9509426A7159FC30FD47C678" box="[620,659,1015,1041]" name="Fiji" pageId="7" pageNumber="228">Fiji</collectingCountry>
and Peninsular
<collectingCountry id="E3AB765F9509426A707EFC30FC61C678" box="[843,949,1015,1041]" name="Malaysia" pageId="7" pageNumber="228">Malaysia</collectingCountry>
; intertidal; associated with
<taxonomicName id="5CBC4D4C9509426A77D9FC30FF39C65D" class="Bivalvia" family="Veneridae" genus="Gafrarium" kingdom="Animalia" order="Veneroida" pageId="7" pageNumber="228" phylum="Mollusca" rank="species" species="pectinatum">
<emphasis id="A9C8EADD9509426A77D9FC30FF39C65D" italics="true" pageId="7" pageNumber="228">Gafrarium pectinatum</emphasis>
</taxonomicName>
,
<taxonomicName id="5CBC4D4C9509426A73CEFBDCFEABC65C" baseAuthorityName="Roding" baseAuthorityYear="1798" box="[251,383,1051,1077]" class="Bivalvia" family="Veneridae" genus="Gafrarium" kingdom="Animalia" order="Veneroida" pageId="7" pageNumber="228" phylum="Mollusca" rank="species" species="tumidum">
<emphasis id="A9C8EADD9509426A73CEFBDCFEABC65C" box="[251,383,1051,1077]" italics="true" pageId="7" pageNumber="228">G. tumidum</emphasis>
</taxonomicName>
and
<taxonomicName id="5CBC4D4C9509426A7280FBDCFDF4C65C" baseAuthorityName="Holten" baseAuthorityYear="1802" box="[437,544,1051,1077]" class="Bivalvia" family="Veneridae" genus="Gafrarium" kingdom="Animalia" order="Veneroida" pageId="7" pageNumber="228" phylum="Mollusca" rank="species" species="dispar">
<emphasis id="A9C8EADD9509426A7280FBDCFDF4C65C" box="[437,544,1051,1077]" italics="true" pageId="7" pageNumber="228">G. dispar</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D3A665449509426473F2FBF8FDEFC518" lastPageId="9" lastPageNumber="230" pageId="7" pageNumber="228" type="discussion">
<paragraph id="9B0336CF9509426A73F2FBF8FECEC438" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="228">
<emphasis id="A9C8EADD9509426A73F2FBF8FEEFC630" bold="true" box="[199,315,1087,1113]" pageId="7" pageNumber="228">Remarks.</emphasis>
The identity of
<taxonomicName id="5CBC4D4C9509426A72C5FBF8FD1DC630" authorityName=", Dana" authorityYear="1852" box="[496,713,1087,1113]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="228" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD9509426A72C5FBF8FD1DC630" box="[496,713,1087,1113]" italics="true" pageId="7" pageNumber="228">Pinnotheres obesus</emphasis>
</taxonomicName>
(as
<taxonomicName id="5CBC4D4C9509426A71CDFBF8FC23C630" authorityName="Dana" authorityYear="1852" box="[760,1015,1086,1113]" class="Malacostraca" family="Pinnotheridae" genus="Pinnothera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="228" phylum="Arthropoda" rank="species" species="obesa">
<emphasis id="A9C8EADD9509426A71CDFBF8FCA2C630" box="[760,886,1087,1113]" italics="true" pageId="7" pageNumber="228">Pinnothera</emphasis>
[sic]
<emphasis id="A9C8EADD9509426A7083FBF8FC23C630" box="[950,1015,1087,1113]" italics="true" pageId="7" pageNumber="228">obesa</emphasis>
</taxonomicName>
) has long been uncertain and this has been compounded by the fact that the
<typeStatus id="4407886D9509426A7177FBA4FDA6C614" box="[578,626,1123,1149]" pageId="7" pageNumber="228">type</typeStatus>
material of
<bibRefCitation id="FF2D4B3E9509426A71CFFBA4FC52C614" author="Dana, J. D." box="[762,902,1122,1149]" pageId="7" pageNumber="228" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana (1852)</bibRefCitation>
is no longer extant (see
<bibRefCitation id="FF2D4B3E9509426A77ADFBA4FACAC614" author="Evans, A. C." box="[1176,1310,1123,1149]" pageId="7" pageNumber="228" pagination="399 - 411" refId="ref13383" refString="Evans, A. C. (1967) Syntypes of Decapoda described by William Stimpson and James Dana in the collection of the British Museum (Natural History). Journal of Natural History, 1, 399 - 411. https: // doi. org / 10.1080 / 00222936700770391" type="journal article" year="1967">Evans 1967</bibRefCitation>
;
<bibRefCitation id="FF2D4B3E9509426A761FFBA4FEF4C6C8" author="Schmitt, W. L. &amp; Mccain, J. C. &amp; Davidson, E." pageId="7" pageNumber="228" refId="ref14912" refString="Schmitt, W. L., Mccain, J. C. &amp; Davidson, E. (1973) Decapoda I. Brachyura I. Fam. Pinnotheridae. Crustaceorum Catalogus. Vol. 3. Dr. W. Junk, Den Haag, 160 pp." type="book" year="1973">
Schmitt
<emphasis id="A9C8EADD9509426A76BDFBA4FF79C6C8" italics="true" pageId="7" pageNumber="228">et al</emphasis>
. 1973: 60
</bibRefCitation>
;
<bibRefCitation id="FF2D4B3E9509426A721EFB40FDE6C6C8" author="Deiss, W. A. &amp; Manning, R. B." box="[299,562,1159,1185]" pageId="7" pageNumber="228" pagination="79 - 85" refId="ref13313" refString="Deiss, W. A. &amp; Manning, R. B. (1981) The fate of the invertebrate collections of the North Pacific Exploring Expedition, 1853 - 1856. In: History in the Service of Systematics. Society for the Bibliography of Natural History, London, pp. 79 - 85. https: // doi. org / 10.3366 / anh. 1981.012" type="book chapter" year="1981">Deiss &amp; Manning 1981</bibRefCitation>
;
<bibRefCitation id="FF2D4B3E9509426A710BFB40FD30C6C8" author="Manning, R. B." box="[574,740,1159,1185]" pageId="7" pageNumber="228" pagination="125 - 177" refId="ref14101" refString="Manning, R. B. (1993) West African pinnotherid crabs, subfamily Pinnotherinae (Crustacea, Decapoda, Brachyura). Bulletin du Museum National d'Histoire Naturelle, Paris, 4 e Serie, Section A (Zoology), 15 (1 - 4), 125 - 177. https: // doi. org / 10.5962 / bhl. title. 542 y" type="journal article" year="1993">Manning 1993</bibRefCitation>
a;
<bibRefCitation id="FF2D4B3E9509426A71C9FB40FBD4C6CB" author="Manning, R. B. &amp; Reed, K. J." box="[764,1024,1159,1186]" pageId="7" pageNumber="228" pagination="283 - 293" refId="ref14170" refString="Manning, R. B. &amp; Reed, K. J. (2006) Decapod crustaceans deposited in the Zoological Museum of Copengagen by William Stimpson in 1859. Raffles Bulletin of Zoology, 54 (2), 283 - 293." type="journal article" year="2006">Manning &amp; Reed 2006</bibRefCitation>
). In establishing
<taxonomicName id="5CBC4D4C9509426A7789FB40FEB4C6AC" authority=", Dana (1852: 380)" authorityName=", Dana" authorityPageNumber="380" authorityYear="1852" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="228" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD9509426A7789FB40FA41C6C8" box="[1212,1429,1159,1185]" italics="true" pageId="7" pageNumber="228">Pinnotheres obesus</emphasis>
,
<bibRefCitation id="FF2D4B3E9509426A73A2FB6CFEB4C6AC" author="Dana, J. D." box="[151,352,1194,1221]" pageId="7" pageNumber="228" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana (1852: 380)</bibRefCitation>
</taxonomicName>
noted that he had at least one male and two females (on the basis of the captions for his plate) obtained from an unknown host or hosts from
<collectingCountry id="E3AB765F9509426A7195FB08FD1FC680" box="[672,715,1231,1257]" name="Fiji" pageId="7" pageNumber="228">Fiji</collectingCountry>
. His description, however, is too brief by modern standards, and details of the ambulatory leg structure and proportions, which are specifically diagnostic in pinnotherids, were not stated.
<bibRefCitation id="FF2D4B3E9509426A73DCFAD0FEACC758" author="Dana, J. D." box="[233,376,1302,1329]" pageId="7" pageNumber="228" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana (1852)</bibRefCitation>
noted that
<taxonomicName id="5CBC4D4C9509426A72CBFAD0FD89C758" authorityName="Dana" authorityYear="1852" box="[510,605,1303,1329]" class="Malacostraca" family="Pinnotheridae" genus="Pinnothera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="228" phylum="Arthropoda" rank="species" species="obesa">
<emphasis id="A9C8EADD9509426A72CBFAD0FD89C758" box="[510,605,1303,1329]" italics="true" pageId="7" pageNumber="228">P. obesa</emphasis>
</taxonomicName>
was close to
<taxonomicName id="5CBC4D4C9509426A71CBFAD0FA93C758" authority="Hombron &amp; Jacquinot, 1846" authorityName="Hombron &amp; Jacquinot" authorityYear="1846" box="[766,1351,1303,1329]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="228" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="A9C8EADD9509426A71CBFAD0FC26C758" box="[766,1010,1303,1329]" italics="true" pageId="7" pageNumber="228">Pinnotheres globosus</emphasis>
Hombron &amp; Jacquinot, 1846
</taxonomicName>
, but he did not elaborate in which aspects the two taxa were similar.
<bibRefCitation id="FF2D4B3E9509426A7073FAFCFC1AC73C" author="Dana, J. D." box="[838,974,1338,1365]" pageId="7" pageNumber="228" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana (1852</bibRefCitation>
: pl. 24 fig. 3a) provided an overall rather schematic figure of a specimen he believed was a male (
<figureCitation id="03872A4A9509426A7031FA98FC88C710" box="[772,860,1375,1401]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1771]" captionTargetBox="[160,1425,839,1716]" captionTargetId="figure-247@2.[151,1436,833,1724]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres obesus (Dana, 1852). A, “male”, overall habitus in dorsal view; B, C, E, F, female, carapace, dorsal view; D, “male”, frontal view showing epistome, antennae, antennules and eyes; E, maxillipeds 3; F, left maxilliped 3; G, outer view of right chela; H, “male” pleon, outer view. After Dana (1855: pl. 24 fig. 3ai)." figureDoi="http://doi.org/10.5281/zenodo.4401463" httpUri="https://zenodo.org/record/4401463/files/figure.png" pageId="7" pageNumber="228">Fig. 1A</figureCitation>
), as well as its frontal region and pleon (
<bibRefCitation id="FF2D4B3E9509426A762FFA98FA4CC710" author="Dana, J. D." box="[1306,1432,1374,1401]" pageId="7" pageNumber="228" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana 1852</bibRefCitation>
: pl. 24 fig. 3h; present
<figureCitation id="03872A4A9509426A72A7FA44FDDBC7F4" box="[402,527,1411,1437]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1771]" captionTargetBox="[160,1425,839,1716]" captionTargetId="figure-247@2.[151,1436,833,1724]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres obesus (Dana, 1852). A, “male”, overall habitus in dorsal view; B, C, E, F, female, carapace, dorsal view; D, “male”, frontal view showing epistome, antennae, antennules and eyes; E, maxillipeds 3; F, left maxilliped 3; G, outer view of right chela; H, “male” pleon, outer view. After Dana (1855: pl. 24 fig. 3ai)." figureDoi="http://doi.org/10.5281/zenodo.4401463" httpUri="https://zenodo.org/record/4401463/files/figure.png" pageId="7" pageNumber="228">Fig. 1D, H</figureCitation>
). Figures of the carapace of two female specimens (
<bibRefCitation id="FF2D4B3E9509426A776EFA44FB0FC7F4" author="Dana, J. D." box="[1115,1243,1410,1437]" pageId="7" pageNumber="228" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana 1852</bibRefCitation>
: pl. 24 fig. 3b, c; present
<figureCitation id="03872A4A9509426A73C5FA60FEBDC7A8" box="[240,361,1447,1473]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1771]" captionTargetBox="[160,1425,839,1716]" captionTargetId="figure-247@2.[151,1436,833,1724]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres obesus (Dana, 1852). A, “male”, overall habitus in dorsal view; B, C, E, F, female, carapace, dorsal view; D, “male”, frontal view showing epistome, antennae, antennules and eyes; E, maxillipeds 3; F, left maxilliped 3; G, outer view of right chela; H, “male” pleon, outer view. After Dana (1855: pl. 24 fig. 3ai)." figureDoi="http://doi.org/10.5281/zenodo.4401463" httpUri="https://zenodo.org/record/4401463/files/figure.png" pageId="7" pageNumber="228">Fig. 1B, C</figureCitation>
) and their maxilliped 3 (
<bibRefCitation id="FF2D4B3E9509426A71B6FA60FCD0C7A8" author="Dana, J. D." box="[643,772,1446,1473]" pageId="7" pageNumber="228" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana 1852</bibRefCitation>
: pl. 24 fig. 3e, f; present
<figureCitation id="03872A4A9509426A7716FA60FB43C7A8" box="[1059,1175,1447,1473]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1771]" captionTargetBox="[160,1425,839,1716]" captionTargetId="figure-247@2.[151,1436,833,1724]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres obesus (Dana, 1852). A, “male”, overall habitus in dorsal view; B, C, E, F, female, carapace, dorsal view; D, “male”, frontal view showing epistome, antennae, antennules and eyes; E, maxillipeds 3; F, left maxilliped 3; G, outer view of right chela; H, “male” pleon, outer view. After Dana (1855: pl. 24 fig. 3ai)." figureDoi="http://doi.org/10.5281/zenodo.4401463" httpUri="https://zenodo.org/record/4401463/files/figure.png" pageId="7" pageNumber="228">Fig. 1E, F</figureCitation>
) were also included. A chela was figured (
<bibRefCitation id="FF2D4B3E9509426A7259FA0CFE38C78C" author="Dana, J. D." box="[364,492,1482,1509]" pageId="7" pageNumber="228" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana 1852</bibRefCitation>
: pl. 24 fig. 3i; present
<figureCitation id="03872A4A9509426A71D2FA0CFCE9C78C" box="[743,829,1483,1509]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1771]" captionTargetBox="[160,1425,839,1716]" captionTargetId="figure-247@2.[151,1436,833,1724]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres obesus (Dana, 1852). A, “male”, overall habitus in dorsal view; B, C, E, F, female, carapace, dorsal view; D, “male”, frontal view showing epistome, antennae, antennules and eyes; E, maxillipeds 3; F, left maxilliped 3; G, outer view of right chela; H, “male” pleon, outer view. After Dana (1855: pl. 24 fig. 3ai)." figureDoi="http://doi.org/10.5281/zenodo.4401463" httpUri="https://zenodo.org/record/4401463/files/figure.png" pageId="7" pageNumber="228">Fig. 1G</figureCitation>
) but it was not specified which specimen it was from. No
<typeStatus id="4407886D9509426A73F5FA28FF24C460" box="[192,240,1519,1545]" pageId="7" pageNumber="228">type</typeStatus>
was designated. No host was mentioned and we are even unsure if the three specimens are conspecific. As noted earlier, all of Danas material is no longer extant, and as such, none of these uncertainties can now be resolved objectively.
</paragraph>
<paragraph id="9B0336CF9509426573F2F99CFBF5C3B8" blockId="7.[151,1437,151,2013]" lastBlockId="8.[151,1437,151,465]" lastPageId="8" lastPageNumber="229" pageId="7" pageNumber="228">
The adult female specimens in our series generally match the figure of
<bibRefCitation id="FF2D4B3E9509426A70DDF99CFBA1C41C" author="Dana, J. D." box="[1000,1141,1626,1653]" pageId="7" pageNumber="228" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana (1852)</bibRefCitation>
in the carapace shape and the maxilliped 3 structure (in particular, the short dactylus not reaching to the propodal apex). On the other hand, the matured male specimen in our material is substantially different from the “male” illustrated by
<bibRefCitation id="FF2D4B3E9509426A77E2F964FAB1C4D4" author="Dana, J. D." box="[1239,1381,1698,1725]" pageId="7" pageNumber="228" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana (1852)</bibRefCitation>
, particularly in the shape of the P2 and P3 dactyli (these dactyli illustrated by Dana are much more elongate and slender than in our male specimen). Nevertheless, such slender, elongate dactyli of the P2 and P
<quantity id="5C449B2A9509426A7748F92CFB73C56C" box="[1149,1191,1771,1797]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="228" unit="in" value="3.0">3 in</quantity>
males are not seen in any known species of
<taxonomicName id="5CBC4D4C9509426A72BBF8C8FDD3C540" authorityName="Manning" authorityYear="1993" box="[398,519,1807,1833]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="A9C8EADD9509426A72BBF8C8FDD3C540" box="[398,519,1807,1833]" italics="true" pageId="7" pageNumber="228">Arcotheres</emphasis>
</taxonomicName>
. Considering the close similarities in females and the locality of the type material, we believe that the specimen regarded as a male by
<bibRefCitation id="FF2D4B3E9509426A71EBF8F4FCBFC524" author="Dana, J. D." box="[734,875,1842,1869]" pageId="7" pageNumber="228" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana (1852)</bibRefCitation>
was likely a young female (early post-hard stage) instead.
<bibRefCitation id="FF2D4B3E9509426A73C3F890FE42C518" author="Dana, J. D." box="[246,406,1878,1905]" pageId="7" pageNumber="228" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Danas (1852)</bibRefCitation>
taxon, if the figures are taken at face value, can thus be assigned to
<taxonomicName id="5CBC4D4C9509426A7795F890FACDC518" authorityName="Manning" authorityYear="1993" box="[1184,1305,1879,1905]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="A9C8EADD9509426A7795F890FACDC518" box="[1184,1305,1879,1905]" italics="true" pageId="7" pageNumber="228">Arcotheres</emphasis>
</taxonomicName>
on account of the P4 and P5 being longer than the P2 and P3, the P4 and P5 dactyli being distinctly longer than those of the P2 and P3, the asymmetrical P4, and the spatulate dactylus of the maxilliped 3. Considering the many inadequacies of Danas description and figures, it can even be argued that
<taxonomicName id="5CBC4D4C9509426A707DF804FBF6C5B4" authorityName=", Dana" authorityYear="1852" box="[840,1058,1987,2013]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="228" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD9509426A707DF804FBF6C5B4" box="[840,1058,1987,2013]" italics="true" pageId="7" pageNumber="228">Pinnotheres obesus</emphasis>
</taxonomicName>
is actually conspecific with what is described here as
<taxonomicName id="5CBC4D4C95064265724EFF50FDB1C2D8" authority="Komai &amp; Kawai &amp; Ng, 2020" authorityName="Komai &amp; Kawai &amp; Ng" authorityYear="2020" box="[379,613,151,177]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
<emphasis id="A9C8EADD95064265724EFF50FDB1C2D8" box="[379,613,151,177]" italics="true" pageId="8" pageNumber="229">Arcotheres ocularius</emphasis>
</taxonomicName>
<emphasis id="A9C8EADD950642657159FF50FD78C2D8" bold="true" box="[620,684,151,177]" pageId="8" pageNumber="229">
<taxonomicNameLabel id="B2FB57A6950642657159FF50FD78C2D8" box="[620,684,151,177]" pageId="8" pageNumber="229" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, although the available evidence seems to suggest otherwise. Both species occur sympatrically in
<collectingCountry id="E3AB765F9506426572C4FF7CFDCFC2BC" box="[497,539,187,213]" name="Fiji" pageId="8" pageNumber="229">Fiji</collectingCountry>
, albeit from different hosts. In any case, as
<bibRefCitation id="FF2D4B3E950642657737FF7CFB5BC2BC" author="Dana, J. D." box="[1026,1167,186,213]" pageId="8" pageNumber="229" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana (1852)</bibRefCitation>
did not specify the host of
<taxonomicName id="5CBC4D4C950642657383FF18FE5BC290" authorityName=", Dana" authorityYear="1852" box="[182,399,223,249]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD950642657383FF18FE5BC290" box="[182,399,223,249]" italics="true" pageId="8" pageNumber="229">Pinnotheres obesus</emphasis>
</taxonomicName>
, we are not even sure if all the material came from the same bivalve species. In the interests of nomenclatural stability and with consideration of the complex state of the pinnotherid taxonomy, we hereby designate a
<typeStatus id="4407886D9506426573C2FEE0FE84C328" box="[247,336,295,321]" pageId="8" pageNumber="229" type="neotype">neotype</typeStatus>
(female of the stage V of the post-hard stage) for
<taxonomicName id="5CBC4D4C9506426570B2FEE0FB87C328" authorityName="Komai &amp; Kawai &amp; Ng" authorityYear="2020" baseAuthorityName="Dana" baseAuthorityYear="1852" box="[903,1107,295,321]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="obesus" status="comb. nov.">
<emphasis id="A9C8EADD9506426570B2FEE0FB87C328" box="[903,1107,295,321]" italics="true" pageId="8" pageNumber="229">Arcotheres obesus</emphasis>
</taxonomicName>
<emphasis id="A9C8EADD95064265776EFEE0FB16C328" bold="true" box="[1115,1218,295,321]" pageId="8" pageNumber="229">
<taxonomicNameLabel id="B2FB57A695064265776EFEE0FB16C328" box="[1115,1218,295,321]" pageId="8" pageNumber="229" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
(CBM-ZC 15945). The designated
<typeStatus id="4407886D95064265727CFE8CFE76C30C" box="[329,418,331,357]" pageId="8" pageNumber="229" type="neotype">neotype</typeStatus>
agrees with the original material relatively well. Their general carapace shape, structure of the chelipeds and legs are similar. The pleon of the specimen Dana illustrated as male (1855: pl. 24, fig. 3H; here reproduced as
<figureCitation id="03872A4A95064265720BFE54FE42C3C4" box="[318,406,403,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1771]" captionTargetBox="[160,1425,839,1716]" captionTargetId="figure-247@2.[151,1436,833,1724]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres obesus (Dana, 1852). A, “male”, overall habitus in dorsal view; B, C, E, F, female, carapace, dorsal view; D, “male”, frontal view showing epistome, antennae, antennules and eyes; E, maxillipeds 3; F, left maxilliped 3; G, outer view of right chela; H, “male” pleon, outer view. After Dana (1855: pl. 24 fig. 3ai)." figureDoi="http://doi.org/10.5281/zenodo.4401463" httpUri="https://zenodo.org/record/4401463/files/figure.png" pageId="8" pageNumber="229">Fig. 1H</figureCitation>
) is of the same general shape and proportions of the present male specimen, although the telson of our male specimen appears relatively shorter (
<figureCitation id="03872A4A950642657031FE70FCB2C3B8" box="[772,870,439,465]" captionStart="FIGURE 1" captionStartId="2.[151,250,1747,1771]" captionTargetBox="[160,1425,839,1716]" captionTargetId="figure-247@2.[151,1436,833,1724]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres obesus (Dana, 1852). A, “male”, overall habitus in dorsal view; B, C, E, F, female, carapace, dorsal view; D, “male”, frontal view showing epistome, antennae, antennules and eyes; E, maxillipeds 3; F, left maxilliped 3; G, outer view of right chela; H, “male” pleon, outer view. After Dana (1855: pl. 24 fig. 3ai)." figureDoi="http://doi.org/10.5281/zenodo.4401463" httpUri="https://zenodo.org/record/4401463/files/figure.png" pageId="8" pageNumber="229">Figs. 1H</figureCitation>
versus
<figureCitation id="03872A4A95064265708EFE70FBC0C3B8" box="[955,1044,438,465]" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[157,1424,499,1834]" captionTargetId="figure-186@8.[151,1436,489,1844]" captionTargetPageId="8" captionText="FIGURE 5. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, habitus, dorsal view; B, left MXP3, outer view; C, left thoracic sternum, ventral view; D, pleon, telson and exposed part of sternite 8 (left), ventral view; E, left G1, outer view; F, same, inner view; G, left G2, outer view; H, same, different view." figureDoi="http://doi.org/10.5281/zenodo.4401473" httpUri="https://zenodo.org/record/4401473/files/figure.png" pageId="8" pageNumber="229">Fig. 5D</figureCitation>
).
</paragraph>
<caption id="CFC366479506426573A2F88CFB69C5C2" ID-DOI="http://doi.org/10.5281/zenodo.4401473" ID-Zenodo-Dep="4401473" httpUri="https://zenodo.org/record/4401473/files/figure.png" pageId="8" pageNumber="229" startId="8.[151,250,1867,1891]" targetBox="[157,1424,499,1834]" targetPageId="8">
<paragraph id="9B0336CF9506426573A2F88CFB69C5C2" blockId="8.[151,1436,1867,1963]" pageId="8" pageNumber="229">
<emphasis id="A9C8EADD9506426573A2F88CFEC1C50A" bold="true" box="[151,277,1867,1891]" pageId="8" pageNumber="229">FIGURE 5.</emphasis>
<taxonomicName id="5CBC4D4C95064265722BF88CFDBEC50A" authority="(Dana, 1852)" authorityName="Komai &amp; Kawai &amp; Ng" authorityYear="2020" baseAuthorityName="Dana" baseAuthorityYear="1852" box="[286,618,1867,1891]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD95064265722BF88CFE0CC50A" box="[286,472,1867,1891]" italics="true" pageId="8" pageNumber="229">Arcotheres obesus</emphasis>
(
<bibRefCitation id="FF2D4B3E9506426572DDF88CFDB7C50A" author="Dana, J. D." box="[488,611,1867,1891]" pageId="8" pageNumber="229" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana, 1852</bibRefCitation>
)
</taxonomicName>
, male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, habitus, dorsal view; B, left MXP3, outer view; C, left thoracic sternum, ventral view; D, pleon, telson and exposed part of sternite 8 (left), ventral view; E, left G1, outer view; F, same, inner view; G, left G2, outer view; H, same, different view.
</paragraph>
</caption>
<paragraph id="9B0336CF9507426473F2FF50FDDBC070" blockId="9.[151,1437,150,1905]" pageId="9" pageNumber="230">
With regards to
<bibRefCitation id="FF2D4B3E95074264724FFF50FD85C2D8" author="Dana, J. D." box="[378,593,150,177]" pageId="9" pageNumber="230" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Danas (1852: 380)</bibRefCitation>
comment that
<taxonomicName id="5CBC4D4C9507426471C3FF50FC1BC2D8" authorityName=", Dana" authorityYear="1852" box="[758,975,151,177]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD9507426471C3FF50FC1BC2D8" box="[758,975,151,177]" italics="true" pageId="9" pageNumber="230">Pinnotheres obesus</emphasis>
</taxonomicName>
was close to
<taxonomicName id="5CBC4D4C950742647751FF50FE77C2BC" authority="Hombron &amp; Jacquinot, 1846" authorityName="Hombron &amp; Jacquinot" authorityYear="1846" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="A9C8EADD950742647751FF50FA81C2D8" box="[1124,1365,151,177]" italics="true" pageId="9" pageNumber="230">Pinnotheres globosus</emphasis>
Hombron &amp; Jacquinot, 1846
</taxonomicName>
, he had noted that his species possessed a weaker tooth on the cutting edge of the dactylus of the chela and the form of the maxilliped 3 was different. The two species, however, are actually morphologically different.
<taxonomicName id="5CBC4D4C950742647233FEC4FE2DC374" authorityName="Hombron &amp; Jacquinot" authorityYear="1846" box="[262,505,259,285]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="A9C8EADD950742647233FEC4FE2DC374" box="[262,505,259,285]" italics="true" pageId="9" pageNumber="230">Pinnotheres globosus</emphasis>
</taxonomicName>
was described from
<collectingCountry id="E3AB765F9507426471D3FEC4FC8CC374" box="[742,856,259,285]" name="Singapore" pageId="9" pageNumber="230">Singapore</collectingCountry>
very briefly (for precise authorship of this species, see
<bibRefCitation id="FF2D4B3E9507426473F7FEE0FE13C328" author="Clark, P. F. &amp; Crosnier, A." box="[194,455,295,321]" pageId="9" pageNumber="230" pagination="407 - 435" refId="ref12931" refString="Clark, P. F. &amp; Crosnier, A. (2000) The zoology of the Voyage au Pole Sud et dans l'Oceanie sur les corvettes l'Astrolabe et la Zelee execute par ordre du Roi pendant les annees 1837 - 1838 - 1839 - 1840 sous le commandement de M. Dumont-d'Urville (1842 - 1854): titles, volumes, plates, text, contents, proposed dates and anecdotal history of the publication. Archives Natural History, London, 27 (3), 407 - 435. https: // doi. org / 10.3366 / anh. 2000.27.3.407" type="journal article" year="2000">Clark &amp; Crosnier 2000</bibRefCitation>
;
<bibRefCitation id="FF2D4B3E9507426472E4FEE0FDA1C328" author="Holthuis, L. B." box="[465,629,295,321]" pageId="9" pageNumber="230" pagination="413 - 422" refId="ref13600" refString="Holthuis, L. B. (2002) A few notes on the authors and dates of the names of Crustacea collected by the &quot; Voyage au Pole Sud et dans l'Oceanie sur les corvettes l'Astrolabe et la Zelee &quot;. Crustaceana, 75 (3 - 4), 413 - 422. https: // doi. org / 10.1163 / 156854002760095480" type="journal article" year="2002">Holthuis 2002</bibRefCitation>
;
<bibRefCitation id="FF2D4B3E95074264714AFEE0FCF5C328" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[639,801,295,321]" pageId="9" pageNumber="230" pagination="1 - 286" refId="ref14697" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="A9C8EADD95074264719DFEE0FD0EC328" box="[680,730,295,321]" italics="true" pageId="9" pageNumber="230">et al</emphasis>
. 2008
</bibRefCitation>
) and
<bibRefCitation id="FF2D4B3E95074264706AFEE0FC3FC328" author="Dana, J. D." box="[863,1003,294,321]" pageId="9" pageNumber="230" refId="ref13078" refString="Dana, J. D. (1852) Crustacea, Part I. In: United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. Vol. 13. C. Sherman, Philadelphia, 685 pp." type="book" year="1852">Dana (1852)</bibRefCitation>
had based his observations entirely on the very schematic and inaccurate figures in Hombron &amp; Jacquinot (1846: figs. 22, 23). The identity of what is true
<taxonomicName id="5CBC4D4C9507426473A2FEA8FECCC3E0" authorityName="Hombron &amp; Jacquinot" authorityYear="1846" box="[151,280,367,393]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="A9C8EADD9507426473A2FEA8FECCC3E0" box="[151,280,367,393]" italics="true" pageId="9" pageNumber="230">P. globosus</emphasis>
</taxonomicName>
is being treated by one of the authors (PKLN) with S. T. Ahyong when a
<typeStatus id="4407886D950742647753FEA8FB6BC3E0" box="[1126,1215,367,393]" pageId="9" pageNumber="230" type="neotype">neotype</typeStatus>
will be designated. Suffice it to say,
<taxonomicName id="5CBC4D4C950742647263FE54FE02C3C4" authorityName="Hombron &amp; Jacquinot" authorityYear="1846" box="[342,470,403,429]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="A9C8EADD950742647263FE54FE02C3C4" box="[342,470,403,429]" italics="true" pageId="9" pageNumber="230">P. globosus</emphasis>
</taxonomicName>
is actually very different from
<taxonomicName id="5CBC4D4C95074264700DFE54FC74C3C4" authorityName=", Dana" authorityYear="1852" box="[824,928,403,429]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD95074264700DFE54FC74C3C4" box="[824,928,403,429]" italics="true" pageId="9" pageNumber="230">P. obesus</emphasis>
</taxonomicName>
, not only being much larger in adult size but also with less asymmetrical P4 and occupation of fan shells of the family
<taxonomicName id="5CBC4D4C9507426470DFFE70FB98C3B8" authorityName="Leach" authorityYear="1819" box="[1002,1100,439,465]" class="Bivalvia" family="Pinnidae" kingdom="Animalia" order="Pterioida" pageId="9" pageNumber="230" phylum="Mollusca" rank="family">Pinnidae</taxonomicName>
(see
<bibRefCitation id="FF2D4B3E9507426477BEFE70FAA5C3B8" author="Ng, P. K. L. &amp; Corbari, L." box="[1163,1393,439,465]" pageId="9" pageNumber="230" pagination="234 - 253" refId="ref14615" refString="Ng, P. K. L. &amp; Corbari, L. (2019) Crustacea: Crab Legacies. In: Low, M. E. Y., Pockington, K. &amp; Jusoh, W. F. A. (Eds.), Voyageurs, Explorateurs et Scientifques. The French and Natural History in Singapore. National University of Singapore and Museum national d'Histoire Naturelle, Paris, pp. 234 - 253." type="book chapter" year="2019">Ng &amp; Corbari 2019</bibRefCitation>
). In view of this, we agree with
<bibRefCitation id="FF2D4B3E9507426472FAFE1CFDB6C39C" author="Tesch, J. J." box="[463,610,475,501]" pageId="9" pageNumber="230" pagination="149 - 295" refId="ref15162" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch (1918)</bibRefCitation>
that
<taxonomicName id="5CBC4D4C9507426471AEFE1CFCCBC39C" authorityName="Hombron &amp; Jacquinot" authorityYear="1846" box="[667,799,475,501]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="A9C8EADD9507426471AEFE1CFCCBC39C" box="[667,799,475,501]" italics="true" pageId="9" pageNumber="230">A. globosus</emphasis>
</taxonomicName>
and
<taxonomicName id="5CBC4D4C950742647063FE1CFC15C39C" authorityName=", Dana" authorityYear="1852" box="[854,961,475,501]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD950742647063FE1CFC15C39C" box="[854,961,475,501]" italics="true" pageId="9" pageNumber="230">A. obesus</emphasis>
</taxonomicName>
are not synonyms, as was supposed by A.
<bibRefCitation id="FF2D4B3E9507426473A2FE38FDDFC070" author="Milne-Edwards, A." box="[151,523,511,537]" pageId="9" pageNumber="230" pagination="155 - 332" refId="ref14215" refString="Milne-Edwards, A. (1873) Recherches sur la faune carcinologique de la Nouvelle-Caledonie. Deuxieme Partie. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 9, 155 - 332, pls. 4 - 18. https: // doi. org / 10.5962 / bhl. title. 10410" type="journal article" year="1873">Milne-Edwards (1873: 318319)</bibRefCitation>
.
</paragraph>
<paragraph id="9B0336CF9507426473F2FDE4FC29C680" blockId="9.[151,1437,150,1905]" pageId="9" pageNumber="230">
<taxonomicName id="5CBC4D4C9507426473F2FDE4FE75C054" authorityName=", Dana" authorityYear="1852" box="[199,417,547,573]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD9507426473F2FDE4FE75C054" box="[199,417,547,573]" italics="true" pageId="9" pageNumber="230">Pinnotheres obesus</emphasis>
</taxonomicName>
has been mentioned or listed by various authors (see synonymy) but few authors have actually examined specimens. The record of “
<taxonomicName id="5CBC4D4C9507426471B2FD80FCB5C008" authorityName=", Dana" authorityYear="1852" box="[647,865,583,609]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD9507426471B2FD80FCB5C008" box="[647,865,583,609]" italics="true" pageId="9" pageNumber="230">Pinnotheres obesus</emphasis>
</taxonomicName>
” by
<bibRefCitation id="FF2D4B3E9507426470AFFD80FBB6C008" author="Miers, E. J." box="[922,1122,583,609]" pageId="9" pageNumber="230" pagination="304 - 317" refId="ref14269" refString="Miers, E. J. (1880) On a collection of Crustacea from the Malaysian Region. Part II. Telphusidea, Catometopa, and Oxystomata. Annals and Magazine of Natural History, Series 5, 5 (28), 304 - 317, pl. XIV. https: // doi. org / 10.1080 / 00222938009459458" type="journal article" year="1880">Miers (1880: 314</bibRefCitation>
, pl. 4 fig. 4) from
<collectingCountry id="E3AB765F950742647603FD80FA49C008" box="[1334,1437,583,609]" name="Malaysia" pageId="9" pageNumber="230">Malaysia</collectingCountry>
was referred to
<taxonomicName id="5CBC4D4C95074264727CFDACFD32C0EC" authority="Burger, 1895" authorityName="Burger" authorityYear="1895" box="[329,742,618,645]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="latissimus">
<emphasis id="A9C8EADD95074264727CFDACFD9CC0EC" box="[329,584,619,645]" italics="true" pageId="9" pageNumber="230">Pinnotheres latissimus</emphasis>
<bibRefCitation id="FF2D4B3E950742647165FDACFD32C0EC" author="Burger, O." box="[592,742,618,645]" pageId="9" pageNumber="230" pagination="361 - 390" refId="ref12768" refString="Burger, O. (1895) Ein Beitrag zur Kenntnis der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390, pls. 9, 10." type="book chapter" year="1895">Bürger, 1895</bibRefCitation>
</taxonomicName>
, with doubt by
<bibRefCitation id="FF2D4B3E9507426470ADFDACFBAEC0EF" author="Gordon, I." box="[920,1146,619,646]" pageId="9" pageNumber="230" pagination="163 - 180" refId="ref13490" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of the Linnean Society of London, Zoology, 40, 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article" year="1936">Gordon (1936: 176)</bibRefCitation>
and
<bibRefCitation id="FF2D4B3E950742647786FDACFF69C0C0" author="Schmitt, W. L. &amp; Mccain, J. C. &amp; Davidson, E." pageId="9" pageNumber="230" refId="ref14912" refString="Schmitt, W. L., Mccain, J. C. &amp; Davidson, E. (1973) Decapoda I. Brachyura I. Fam. Pinnotheridae. Crustaceorum Catalogus. Vol. 3. Dr. W. Junk, Den Haag, 160 pp." type="book" year="1973">
Schmitt
<emphasis id="A9C8EADD950742647627FDACFA90C0EC" box="[1298,1348,619,645]" italics="true" pageId="9" pageNumber="230">et al</emphasis>
. (1973: 51)
</bibRefCitation>
but we are now confident it is actually
<taxonomicName id="5CBC4D4C95074264714BFD48FC75C0C0" authority="(Burger, 1895)" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[638,929,654,681]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="A9C8EADD95074264714BFD48FD20C0C1" box="[638,756,655,681]" italics="true" pageId="9" pageNumber="230">A. exiguus</emphasis>
(
<bibRefCitation id="FF2D4B3E950742647036FD48FC4CC0C0" author="Burger, O." box="[771,920,654,681]" pageId="9" pageNumber="230" pagination="361 - 390" refId="ref12768" refString="Burger, O. (1895) Ein Beitrag zur Kenntnis der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390, pls. 9, 10." type="book chapter" year="1895">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
instead (PKLN &amp; S.T. Ahyong, unpublished data). This also applies to the record by
<bibRefCitation id="FF2D4B3E950742647155FD74FCEFC0A4" author="Walker, A. O." box="[608,827,691,717]" pageId="9" pageNumber="230" pagination="107 - 117" refId="ref15296" refString="Walker, A. O. (1887) Notes on a collection of Crustacea from Singapore. Journal of the Linnean Society of London, 20 (118), 107 - 117, pls. 6 - 9. https: // doi. org / 10.5962 / bhl. title. 115470" type="journal article" year="1887">Walker (1887: 111)</bibRefCitation>
from
<collectingCountry id="E3AB765F9507426470B7FD74FC38C0A4" box="[898,1004,691,717]" name="Malaysia" pageId="9" pageNumber="230">Malaysia</collectingCountry>
. The records of “
<taxonomicName id="5CBC4D4C950742647784FD74FACEC0A4" authorityName=", Dana" authorityYear="1852" box="[1201,1306,691,717]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD950742647784FD74FACEC0A4" box="[1201,1306,691,717]" italics="true" pageId="9" pageNumber="230">P. obesus</emphasis>
</taxonomicName>
” by
<bibRefCitation id="FF2D4B3E950742647666FD74FEA5C098" author="Fulton, S. W. &amp; Grant, F. E." pageId="9" pageNumber="230" pagination="5 - 15" refId="ref13435" refString="Fulton, S. W. &amp; Grant, F. E. (1906) Some little known Victorian decapod Crustacea with descriptions of new species No. III. Proceedings of the Royal Society of Victoria, New Series, 19 (1), 5 - 15, pls. III-V." type="journal article" year="1906">Fulton &amp; Grant (1906: 18)</bibRefCitation>
(relisted in
<bibRefCitation id="FF2D4B3E9507426472CDFD10FD28C098" author="Schmitt, W. L. &amp; Mccain, J. C. &amp; Davidson, E." box="[504,764,727,753]" pageId="9" pageNumber="230" refId="ref14912" refString="Schmitt, W. L., Mccain, J. C. &amp; Davidson, E. (1973) Decapoda I. Brachyura I. Fam. Pinnotheridae. Crustaceorum Catalogus. Vol. 3. Dr. W. Junk, Den Haag, 160 pp." type="book" year="1973">
Schmitt
<emphasis id="A9C8EADD950742647162FD10FD5CC098" box="[599,648,727,753]" italics="true" pageId="9" pageNumber="230">et al</emphasis>
. 1973: 60
</bibRefCitation>
; Davie 2002: 433) from southern
<collectingCountry id="E3AB765F95074264774DFD10FB0BC098" box="[1144,1247,727,753]" name="Australia" pageId="9" pageNumber="230">Australia</collectingCountry>
are
<taxonomicName id="5CBC4D4C95074264763AFD10FE0EC17C" authority="(Baker, 1907)" baseAuthorityName="Baker" baseAuthorityYear="1907" class="Malacostraca" family="Pinnotheridae" genus="Austrotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="holothuriensis">
<emphasis id="A9C8EADD95074264763AFD10FEE3C17C" italics="true" pageId="9" pageNumber="230">Austrotheres holothuriensis</emphasis>
(Baker, 1907)
</taxonomicName>
instead (see
<bibRefCitation id="FF2D4B3E950742647159FD3CFC91C17C" author="Ahyong, S. T." box="[620,837,762,789]" pageId="9" pageNumber="230" pagination="538 - 571" refId="ref12561" refString="Ahyong, S. T. (2018) Revision of Ostracotheres H. Milne Edwards, 1853 (Crustacea: Brachyura: Pinnotheridae). Raffles Bulletin of Zoology, 66, 538 - 571." type="journal article" year="2018">Ahyong 2018: 550</bibRefCitation>
).
<bibRefCitation id="FF2D4B3E95074264706DFD3CFB04C17C" author="Hornell, J. &amp; Southwell, T." box="[856,1232,763,789]" pageId="9" pageNumber="230" pagination="99 - 103" refId="ref13729" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103, pl. 1. https: // doi. org / 10.5962 / bhl. title. 54463" type="journal article" year="1909">Hornell &amp; Southwell (1909: 102)</bibRefCitation>
recorded “
<taxonomicName id="5CBC4D4C95074264767BFD3CFF6FC151" authorityName=", Dana" authorityYear="1852" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD95074264767BFD3CFF6FC151" italics="true" pageId="9" pageNumber="230">P. obesus</emphasis>
</taxonomicName>
” from two venerid clams,
<taxonomicName id="5CBC4D4C9507426472DDFCD8FCD3C150" authority="Schumacher, 1817" authorityName="Schumacher" authorityYear="1817" box="[488,775,799,825]" class="Bivalvia" family="Veneridae" genus="Meroe" higherTaxonomySource="CoL" kingdom="Animalia" order="Venerida" pageId="9" pageNumber="230" phylum="Mollusca" rank="genus">
<emphasis id="A9C8EADD9507426472DDFCD8FDE4C151" box="[488,560,799,824]" italics="true" pageId="9" pageNumber="230">Meroe</emphasis>
<bibRefCitation id="FF2D4B3E95074264710DFCD8FCD3C150" author="Schumacher C. F." box="[568,775,799,825]" pageId="9" pageNumber="230" refId="ref14881" refString="Schumacher C. F. (1817) Essai d'un nouveau systeme des habitations des vers testaces. Schultz, Copenhagen, iv + 288 pp., 22 pls." type="book" year="1817">Schumacher, 1817</bibRefCitation>
</taxonomicName>
(present day
<taxonomicName id="5CBC4D4C9507426470AAFCD8FBA3C150" authority="Link, 1807" authorityName="Link" authorityYear="1807" box="[927,1143,799,825]" class="Bivalvia" family="Veneridae" genus="Sunetta" kingdom="Animalia" order="Veneroida" pageId="9" pageNumber="230" phylum="Mollusca" rank="genus">
<emphasis id="A9C8EADD9507426470AAFCD8FC27C150" box="[927,1011,799,825]" italics="true" pageId="9" pageNumber="230">Sunetta</emphasis>
<bibRefCitation id="FF2D4B3E9507426470CEFCD8FBA3C150" author="Link, D. H. F." box="[1019,1143,799,825]" pageId="9" pageNumber="230" pagination="1 - 30" refId="ref13951" refString="Link, D. H. F. (1807) Beschreibung der Naturalien-Sammlung der Universitat zu Rostock. 1 - 4 Abt. [Part 1 - 4]. Adlers Erben, Rostock. 195 pp. [1 Abt. (Part 1): pp. 1 - 50, 2 Abt. (Part 2): pp. 51 - 100, 3 Abt. (Part 3): pp. 101 - 165, Abt. 4 (Part 4): pp. 1 - 30]" type="book chapter" year="1807">Link, 1807</bibRefCitation>
</taxonomicName>
) and
<taxonomicName id="5CBC4D4C950742647783FCD8FF1BC134" authority="Lamarck, 1805" authorityName="Lamarck" authorityYear="1805" class="Bivalvia" family="Arcticidae" genus="Cytheraea" higherTaxonomySource="CoL" kingdom="Animalia" order="Venerida" pageId="9" pageNumber="230" phylum="Mollusca" rank="genus">
<emphasis id="A9C8EADD950742647783FCD8FAFFC150" box="[1206,1323,799,825]" italics="true" pageId="9" pageNumber="230">Cytheraea</emphasis>
<bibRefCitation id="FF2D4B3E950742647607FCD8FF1BC134" author="Lamarck, J. B. P. A. de" pageId="9" pageNumber="230" pagination="53 - 62" refId="ref13910" refString="Lamarck, J. B. P. A. de (1805) Suite des memoires sur les fossiles des environs de Paris. Annales du Museum d'Histoire Naturelle, 7, 53 - 62 + 130 - 139." type="journal article" year="1805">Lamarck, 1805</bibRefCitation>
</taxonomicName>
(present day
<taxonomicName id="5CBC4D4C950742647250FC84FDADC134" authority="Lamarck, 1799" authorityName="Lamarck" authorityYear="1799" box="[357,633,835,861]" class="Bivalvia" family="Veneridae" genus="Meretrix" kingdom="Animalia" order="Veneroida" pageId="9" pageNumber="230" phylum="Mollusca" rank="genus">
<emphasis id="A9C8EADD950742647250FC84FE12C135" box="[357,454,835,860]" italics="true" pageId="9" pageNumber="230">Meretrix</emphasis>
<bibRefCitation id="FF2D4B3E9507426472F9FC84FDADC134" author="Lamarck, J. B. M." box="[460,633,835,861]" pageId="9" pageNumber="230" pagination="63 - 91" refId="ref13860" refString="Lamarck, J. B. M. (1799) Prodrome d'une nouvelle classification des coquilles, comprenant une redaction appropriee des caracteres generiques, et l'etablissement d'un grand nombre de genres nouveaux. Memoires de la Societe d'Histoire Naturelle de Paris, 1, 63 - 91." type="journal article" year="1799">Lamarck, 1799</bibRefCitation>
</taxonomicName>
), but given the host records, this is more likely what is today
<taxonomicName id="5CBC4D4C950742647614FC84FA42C135" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1313,1430,835,861]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="A9C8EADD950742647614FC84FA42C135" box="[1313,1430,835,861]" italics="true" pageId="9" pageNumber="230">A. exiguus</emphasis>
</taxonomicName>
.
<bibRefCitation id="FF2D4B3E9507426473A2FCA0FEFEC1E8" author="Tesch, J. J." box="[151,298,871,897]" pageId="9" pageNumber="230" pagination="149 - 295" refId="ref15162" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch (1918)</bibRefCitation>
recorded a small male specimen (3.1×3.0 mm) as “
<taxonomicName id="5CBC4D4C950742647045FCA0FC0CC1E8" authorityName=", Dana" authorityYear="1852" box="[880,984,871,897]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD950742647045FCA0FC0CC1E8" box="[880,984,871,897]" italics="true" pageId="9" pageNumber="230">P. obesus</emphasis>
</taxonomicName>
” collected from an
<taxonomicName id="5CBC4D4C95074264778DFCA0FB39C1E8" authorityName="Linnaeus" authorityYear="1758" box="[1208,1261,871,897]" class="Bivalvia" family="Arcidae" genus="Arca" kingdom="Animalia" order="Arcoida" pageId="9" pageNumber="230" phylum="Mollusca" rank="genus">
<emphasis id="A9C8EADD95074264778DFCA0FB39C1E8" box="[1208,1261,871,897]" italics="true" pageId="9" pageNumber="230">Arca</emphasis>
</taxonomicName>
shell (
<taxonomicName id="5CBC4D4C95074264760CFCA0FA41C1EB" authorityName="Lamarck" authorityYear="1809" box="[1337,1429,871,898]" class="Bivalvia" family="Arcidae" kingdom="Animalia" order="Arcoida" pageId="9" pageNumber="230" phylum="Mollusca" rank="family">Arcidae</taxonomicName>
) from Ceram in the
<collectingCountry id="E3AB765F95074264725DFC4CFE0CC1CC" box="[360,472,907,933]" name="Indonesia" pageId="9" pageNumber="230">Moluccas</collectingCountry>
,
<collectingCountry id="E3AB765F9507426472D7FC4CFD87C1CC" box="[482,595,907,933]" name="Indonesia" pageId="9" pageNumber="230">Indonesia</collectingCountry>
. This is certainly not
<taxonomicName id="5CBC4D4C950742647008FC4CFC73C1CC" authorityName=", Dana" authorityYear="1852" box="[829,935,907,933]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD950742647008FC4CFC73C1CC" box="[829,935,907,933]" italics="true" pageId="9" pageNumber="230">A. obesus</emphasis>
</taxonomicName>
or any of the species of
<taxonomicName id="5CBC4D4C950742647787FC4CFAFFC1CC" authorityName="Manning" authorityYear="1993" box="[1202,1323,907,933]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="A9C8EADD950742647787FC4CFAFFC1CC" box="[1202,1323,907,933]" italics="true" pageId="9" pageNumber="230">Arcotheres</emphasis>
</taxonomicName>
discussed here and the figured maxilliped 3 (
<bibRefCitation id="FF2D4B3E950742647113FC68FD79C1A0" author="Tesch, J. J." box="[550,685,943,969]" pageId="9" pageNumber="230" pagination="149 - 295" refId="ref15162" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch 1918</bibRefCitation>
: fig. 3a) has the dactylus very short and inserted on the propodus rather distally. The other records of “
<taxonomicName id="5CBC4D4C950742647103FC14FD76C184" authorityName=", Dana" authorityYear="1852" box="[566,674,979,1005]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD950742647103FC14FD76C184" box="[566,674,979,1005]" italics="true" pageId="9" pageNumber="230">A. obesus</emphasis>
</taxonomicName>
” from other parts of the Indo-West Pacific (see
<bibRefCitation id="FF2D4B3E95074264778BFC14FF60C67B" author="Schmitt, W. L. &amp; Mccain, J. C. &amp; Davidson, E." pageId="9" pageNumber="230" refId="ref14912" refString="Schmitt, W. L., Mccain, J. C. &amp; Davidson, E. (1973) Decapoda I. Brachyura I. Fam. Pinnotheridae. Crustaceorum Catalogus. Vol. 3. Dr. W. Junk, Den Haag, 160 pp." type="book" year="1973">
Schmitt
<emphasis id="A9C8EADD950742647628FC14FA9AC184" box="[1309,1358,979,1005]" italics="true" pageId="9" pageNumber="230">et al</emphasis>
. 1973: 60
</bibRefCitation>
) must all be rechecked to ascertain which species they actually belong to.
<bibRefCitation id="FF2D4B3E9507426470D8FC30FB29C67B" author="Schmitt, W. L. &amp; Mccain, J. C. &amp; Davidson, E." box="[1005,1277,1015,1042]" pageId="9" pageNumber="230" refId="ref14912" refString="Schmitt, W. L., Mccain, J. C. &amp; Davidson, E. (1973) Decapoda I. Brachyura I. Fam. Pinnotheridae. Crustaceorum Catalogus. Vol. 3. Dr. W. Junk, Den Haag, 160 pp." type="book" year="1973">
Schmitt
<emphasis id="A9C8EADD95074264777EFC30FBAFC678" box="[1099,1147,1015,1041]" italics="true" pageId="9" pageNumber="230">et al</emphasis>
. (1973: 60)
</bibRefCitation>
noted that “In a personal communication Dr. Isabella Gordon says that the only specimens of
<taxonomicName id="5CBC4D4C950742647716FBDCFB2AC65C" authorityName=", Dana" authorityYear="1852" box="[1059,1278,1051,1077]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD950742647716FBDCFB2AC65C" box="[1059,1278,1051,1077]" italics="true" pageId="9" pageNumber="230">Pinnotheres obesus</emphasis>
</taxonomicName>
in the British Museum are from the
<collectingCountry id="E3AB765F9507426472A7FBF8FDC2C630" box="[402,534,1087,1113]" name="Fiji" pageId="9" pageNumber="230">Fiji Islands</collectingCountry>
”. Unfortunately, we were not able to examine this non-type material; they could either be
<taxonomicName id="5CBC4D4C9507426473CAFBA4FEBEC614" authorityName=", Dana" authorityYear="1852" box="[255,362,1123,1149]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD9507426473CAFBA4FEBEC614" box="[255,362,1123,1149]" italics="true" pageId="9" pageNumber="230">A. obesus</emphasis>
</taxonomicName>
as defined here, or the new species (see below).
<bibRefCitation id="FF2D4B3E9507426470B9FBA4FBEBC614" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[908,1087,1123,1149]" pageId="9" pageNumber="230" pagination="1 - 286" refId="ref14697" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="A9C8EADD950742647080FBA4FC32C614" box="[949,998,1123,1149]" italics="true" pageId="9" pageNumber="230">et al</emphasis>
. (2008)
</bibRefCitation>
retained
<taxonomicName id="5CBC4D4C950742647793FBA4FAABC614" authorityName=", Dana" authorityYear="1852" box="[1190,1407,1123,1149]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD950742647793FBA4FAABC614" box="[1190,1407,1123,1149]" italics="true" pageId="9" pageNumber="230">Pinnotheres obesus</emphasis>
</taxonomicName>
in
<taxonomicName id="5CBC4D4C9507426473A2FB40FE76C6C8" authority="Bosc, 1802" authorityName="Bosc" authorityYear="1802" box="[151,418,1159,1185]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="A9C8EADD9507426473A2FB40FEC9C6C8" box="[151,285,1159,1185]" italics="true" pageId="9" pageNumber="230">Pinnotheres</emphasis>
<bibRefCitation id="FF2D4B3E950742647211FB40FE76C6C8" author="Bosc, L. A. G." box="[292,418,1159,1185]" pageId="9" pageNumber="230" pagination="9 - 18" refId="ref12706" refString="Bosc, L. A. G. (1802) Histoire naturelle des Crustaces, contenant leur description et leur moeurs: avec figures dessinees d'apres nature. Vol. 2. De Guilleminet, Paris, 296 pp., pls. 9 - 18. https: // doi. org / 10.5962 / bhl. title. 39831" type="book chapter" year="1802">Bosc, 1802</bibRefCitation>
</taxonomicName>
<emphasis id="A9C8EADD95074264729CFB40FE30C6C9" box="[425,484,1159,1184]" italics="true" pageId="9" pageNumber="230">
<taxonomicNameLabel id="B2FB57A695074264729CFB40FE30C6C9" box="[425,484,1159,1184]" pageId="9" pageNumber="230" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
, mainly because the detailed structure of the female ambulatory legs was not known. With the designation of the present
<typeStatus id="4407886D950742647115FB6CFDA8C6AC" box="[544,636,1195,1221]" pageId="9" pageNumber="230" type="neotype">neotype</typeStatus>
, this species clearly belongs to
<taxonomicName id="5CBC4D4C9507426470E2FB6CFAD7C6AC" authority="Manning, 1993" authorityName="Manning" authorityYear="1993" box="[983,1283,1195,1221]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="A9C8EADD9507426470E2FB6CFB84C6AC" box="[983,1104,1195,1221]" italics="true" pageId="9" pageNumber="230">Arcotheres</emphasis>
<bibRefCitation id="FF2D4B3E950742647763FB6CFAD7C6AC" author="Manning, R. B." box="[1110,1283,1195,1221]" pageId="9" pageNumber="230" pagination="125 - 177" refId="ref14101" refString="Manning, R. B. (1993) West African pinnotherid crabs, subfamily Pinnotherinae (Crustacea, Decapoda, Brachyura). Bulletin du Museum National d'Histoire Naturelle, Paris, 4 e Serie, Section A (Zoology), 15 (1 - 4), 125 - 177. https: // doi. org / 10.5962 / bhl. title. 542 y" type="journal article" year="1993">Manning, 1993</bibRefCitation>
</taxonomicName>
, as diagnosed by Campos &amp; Manning (2000) (see also
<bibRefCitation id="FF2D4B3E95074264716BFB08FC94C680" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[606,832,1231,1257]" pageId="9" pageNumber="230" pagination="191 - 226" refId="ref12598" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura. Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong &amp; Ng 2007</bibRefCitation>
;
<bibRefCitation id="FF2D4B3E950742647079FB08FC25C680" author="Ng, P. K. L. &amp; Clark, P. F. &amp; Mitra, S. &amp; Kumar, A. B." box="[844,1009,1231,1257]" pageId="9" pageNumber="230" pagination="1079 - 1097" refId="ref14467" refString="Ng, P. K. L., Clark, P. F., Mitra, S. &amp; Kumar, A. B. (2017) Arcotheres borradailei (Nobili, 1906) and Pinnotheres ridgewayi Southwell, 1911: a reassessment of characters and generic assignment of species to Arcotheres Manning, 1993 (Decapoda, Brachyura, Pinnotheridae). In: Magalhaes, C., Becker, C., Davie, P. J. F., Klimpel, S., Martinez-Arbizu, P. &amp; Sonnewald, M. (Eds.), Special Issue Dedicated to the Memory of Michael Turkay (1948 - 2015). Crustaceana, 90 (7 - 10), pp. 1079 - 1097. https: // doi. org / 10.1163 / 15685403 - 00003613" type="journal article" year="2017">
Ng
<emphasis id="A9C8EADD950742647040FB08FC73C680" box="[885,935,1231,1257]" italics="true" pageId="9" pageNumber="230">et al</emphasis>
. 2017
</bibRefCitation>
).
</paragraph>
<paragraph id="9B0336CF9507426473F2FB34FB3AC7F4" blockId="9.[151,1437,150,1905]" pageId="9" pageNumber="230">
The carapace of
<taxonomicName id="5CBC4D4C9507426472BDFB34FE21C764" authorityName=", Dana" authorityYear="1852" box="[392,501,1267,1293]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD9507426472BDFB34FE21C764" box="[392,501,1267,1293]" italics="true" pageId="9" pageNumber="230">A. obesus</emphasis>
</taxonomicName>
is superficially similar to
<taxonomicName id="5CBC4D4C950742647013FB34FC14C764" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[806,960,1267,1293]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="A9C8EADD950742647013FB34FC14C764" box="[806,960,1267,1293]" italics="true" pageId="9" pageNumber="230">A. rotundatus</emphasis>
</taxonomicName>
(known only from the
<typeStatus id="4407886D9507426477FAFB34FAE5C764" box="[1231,1329,1267,1293]" pageId="9" pageNumber="230" type="holotype">holotype</typeStatus>
from the venerid
<taxonomicName id="5CBC4D4C9507426473C7FAD0FDDEC758" authority="Schumacher, 1817" authorityName="Schumacher" authorityYear="1817" box="[242,522,1303,1329]" class="Bivalvia" family="Veneridae" genus="Circe" kingdom="Animalia" order="Veneroida" pageId="9" pageNumber="230" phylum="Mollusca" rank="genus">
<emphasis id="A9C8EADD9507426473C7FAD0FEFBC758" box="[242,303,1303,1329]" italics="true" pageId="9" pageNumber="230">Circe</emphasis>
<bibRefCitation id="FF2D4B3E950742647202FAD0FDDEC758" author="Schumacher C. F." box="[311,522,1303,1329]" pageId="9" pageNumber="230" refId="ref14881" refString="Schumacher C. F. (1817) Essai d'un nouveau systeme des habitations des vers testaces. Schultz, Copenhagen, iv + 288 pp., 22 pls." type="book" year="1817">Schumacher, 1817</bibRefCitation>
</taxonomicName>
). The maxilliped 3 dactylus of
<taxonomicName id="5CBC4D4C95074264705DFAD0FC00C758" authorityName=", Dana" authorityYear="1852" box="[872,980,1303,1329]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD95074264705DFAD0FC00C758" box="[872,980,1303,1329]" italics="true" pageId="9" pageNumber="230">A. obesus</emphasis>
</taxonomicName>
is, however, relatively more slender and shorter, never reaching the tip of the propodus (
<figureCitation id="03872A4A950742647191FAFCFD2EC73C" box="[676,762,1339,1365]" captionStart="FIGURE 3" captionStartId="4.[151,250,1848,1872]" captionTargetBox="[162,1423,247,1813]" captionTargetId="figure-46@4.[151,1436,237,1824]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres obesus (Dana, 1852). AD, neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945; E, F, female (6.9×8.9 mm), ZRC 2019.1876; G, H, female (5.9×8.3 mm), ZRC 2019.1876. A, overall habitus, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, pleomere 6 and telson, ventral view; E, G, left maxilliped 3, outer view; F, H, left maxilliped 3, inner view." figureDoi="http://doi.org/10.5281/zenodo.4401469" httpUri="https://zenodo.org/record/4401469/files/figure.png" pageId="9" pageNumber="230">Fig. 3C</figureCitation>
, EH) (versus relatively stouter and longer, reaching beyond the tip of the propodus in
<taxonomicName id="5CBC4D4C950742647284FA98FD93C710" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[433,583,1375,1401]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="A9C8EADD950742647284FA98FD93C710" box="[433,583,1375,1401]" italics="true" pageId="9" pageNumber="230">A. rotundatus</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="FF2D4B3E950742647143FA98FC82C710" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[630,854,1375,1401]" pageId="9" pageNumber="230" pagination="191 - 226" refId="ref12598" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura. Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 13I); and the female P4 is proportionately much longer (
<figureCitation id="03872A4A9507426473DBFA44FE97C7F4" box="[238,323,1411,1437]" captionStart="FIGURE 4" captionStartId="6.[151,250,1862,1886]" captionTargetBox="[226,1364,190,1827]" captionTargetId="figure-20@6.[216,1371,181,1838]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres obesus (Dana, 1852), neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945. A, right chela, outer view; B, same, inner view; C, chela and carpus of right cheliped, dorsal view; D, distal part of fixed finger of right chela, ventral view; E, carpus and merus of right cheliped, ventral view; F, right P2, dorsal view; G, same dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (shorter side), dorsal view; K, same, dactylus; L, left P4 (longer side), dorsal view; M, same dactylus; N, right P5, dorsal view; O, same dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401471" httpUri="https://zenodo.org/record/4401471/files/figure.png" pageId="9" pageNumber="230">Fig. 4L</figureCitation>
) (versus proportionately shorter in
<taxonomicName id="5CBC4D4C9507426471FAFA44FCB2C7F4" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[719,870,1411,1437]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="A9C8EADD9507426471FAFA44FCB2C7F4" box="[719,870,1411,1437]" italics="true" pageId="9" pageNumber="230">A. rotundatus</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="FF2D4B3E9507426470A2FA44FBAEC7F4" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[919,1146,1411,1437]" pageId="9" pageNumber="230" pagination="191 - 226" refId="ref12598" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura. Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 13F).
</paragraph>
<paragraph id="9B0336CF9507426473F2FA60FB99C56C" blockId="9.[151,1437,150,1905]" pageId="9" pageNumber="230">
The general carapace shape, maxilliped 3 and chelipeds of
<taxonomicName id="5CBC4D4C950742647054FA60FC18C7A8" authorityName=", Dana" authorityYear="1852" box="[865,972,1447,1473]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD950742647054FA60FC18C7A8" box="[865,972,1447,1473]" italics="true" pageId="9" pageNumber="230">A. obesus</emphasis>
</taxonomicName>
are similar to members of the
<taxonomicName id="5CBC4D4C950742647612FA60FA49C7A9" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1319,1437,1447,1473]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="A9C8EADD950742647612FA60FA49C7A9" box="[1319,1437,1447,1473]" italics="true" pageId="9" pageNumber="230">A. exiguus</emphasis>
</taxonomicName>
group (
<taxonomicName id="5CBC4D4C9507426473DCFA0CFDD9C78C" authority="(Burger, 1895)" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[233,525,1482,1509]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="A9C8EADD9507426473DCFA0CFE8BC78D" box="[233,351,1483,1509]" italics="true" pageId="9" pageNumber="230">A. exiguus</emphasis>
(
<bibRefCitation id="FF2D4B3E95074264725BFA0CFDD1C78C" author="Burger, O." box="[366,517,1482,1509]" pageId="9" pageNumber="230" pagination="361 - 390" refId="ref12768" refString="Burger, O. (1895) Ein Beitrag zur Kenntnis der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390, pls. 9, 10." type="book chapter" year="1895">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="5CBC4D4C95074264712FFA0CFCAEC78F" authority="(Gordon, 1936)" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[538,890,1483,1510]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="winckworthi">
<emphasis id="A9C8EADD95074264712FFA0CFD17C78C" box="[538,707,1483,1509]" italics="true" pageId="9" pageNumber="230">A. winckworthi</emphasis>
(
<bibRefCitation id="FF2D4B3E9507426471E6FA0CFCA6C78F" author="Gordon, I." box="[723,882,1483,1510]" pageId="9" pageNumber="230" pagination="163 - 180" refId="ref13490" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of the Linnean Society of London, Zoology, 40, 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article" year="1936">Gordon, 1936</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="5CBC4D4C950742647084FA0CFB3BC78C" authority="(Chhapgar, 1957))" authorityName="" baseAuthorityName="Chhapgar" baseAuthorityYear="1957" box="[945,1263,1482,1509]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="vicajii">
<emphasis id="A9C8EADD950742647084FA0CFBC3C78D" box="[945,1047,1483,1509]" italics="true" pageId="9" pageNumber="230">A. vicajii</emphasis>
(
<bibRefCitation id="FF2D4B3E950742647713FA0CFB08C78C" author="Chhapgar, B. F." box="[1062,1244,1482,1509]" pageId="9" pageNumber="230" pagination="503 - 549" refId="ref12890" refString="Chhapgar, B. F. (1957) On the marine crabs (Decapoda: Brachyura) of Bombay State Part II. Journal of the Bombay Natural History Society, 54 (3), 503 - 549." type="journal article" year="1957">Chhapgar, 1957</bibRefCitation>
))
</taxonomicName>
, but the female P2, P3 and P5 meri are proportionately more slender and longer (
<figureCitation id="03872A4A950742647051FA28FC13C460" box="[868,967,1519,1545]" captionStart="FIGURE 3" captionStartId="4.[151,250,1848,1872]" captionTargetBox="[162,1423,247,1813]" captionTargetId="figure-46@4.[151,1436,237,1824]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres obesus (Dana, 1852). AD, neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945; E, F, female (6.9×8.9 mm), ZRC 2019.1876; G, H, female (5.9×8.3 mm), ZRC 2019.1876. A, overall habitus, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, pleomere 6 and telson, ventral view; E, G, left maxilliped 3, outer view; F, H, left maxilliped 3, inner view." figureDoi="http://doi.org/10.5281/zenodo.4401469" httpUri="https://zenodo.org/record/4401469/files/figure.png" pageId="9" pageNumber="230">Figs. 3A</figureCitation>
,
<figureCitation id="03872A4A9507426470E7FA28FBE5C460" box="[978,1073,1519,1545]" captionStart="FIGURE 4" captionStartId="6.[151,250,1862,1886]" captionTargetBox="[226,1364,190,1827]" captionTargetId="figure-20@6.[216,1371,181,1838]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres obesus (Dana, 1852), neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945. A, right chela, outer view; B, same, inner view; C, chela and carpus of right cheliped, dorsal view; D, distal part of fixed finger of right chela, ventral view; E, carpus and merus of right cheliped, ventral view; F, right P2, dorsal view; G, same dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (shorter side), dorsal view; K, same, dactylus; L, left P4 (longer side), dorsal view; M, same dactylus; N, right P5, dorsal view; O, same dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401471" httpUri="https://zenodo.org/record/4401471/files/figure.png" pageId="9" pageNumber="230">4F, H, N</figureCitation>
) [versus P2, P3 and P5 meri proportionately shorter and stouter in the
<taxonomicName id="5CBC4D4C950742647174F9D4FD61C445" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[577,693,1555,1581]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="A9C8EADD950742647174F9D4FD61C445" box="[577,693,1555,1581]" italics="true" pageId="9" pageNumber="230">A. exiguus</emphasis>
</taxonomicName>
group; cf.
<bibRefCitation id="FF2D4B3E950742647018F9D4FBFDC444" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[813,1065,1555,1581]" pageId="9" pageNumber="230" pagination="191 - 226" refId="ref12598" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura. Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong &amp; Ng 2007: 4</bibRefCitation>
A;
<bibRefCitation id="FF2D4B3E95074264777FF9D4FB3EC444" author="Gordon, I." box="[1098,1258,1555,1581]" pageId="9" pageNumber="230" pagination="163 - 180" refId="ref13490" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of the Linnean Society of London, Zoology, 40, 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article" year="1936">Gordon, 1936</bibRefCitation>
: fig. 7a;
<bibRefCitation id="FF2D4B3E95074264767DF9D4FEA9C438" author="Antony, A. &amp; Kuttyamma, V. J." pageId="9" pageNumber="230" pagination="59 - 68" refId="ref12654" refString="Antony, A. &amp; Kuttyamma, V. J. (1971) A new species of crab Pinnotheres Latreille (Crustacea: Brachyura: Pinnotheridae) from the clam Meretrix casta (Chemnitz). Bulletin of Department of Marine Biology and Oceanography, 5, 59 - 68." type="journal article" year="1971">Antony &amp; Kuttyamma 1971</bibRefCitation>
: text-fig. 2(4);
<bibRefCitation id="FF2D4B3E950742647112F9F0FD28C438" author="Trivedi, J. N. &amp; Gosavi, S. &amp; Vachhrajani, K. D. &amp; Mitra, S. &amp; Ravinesh, R. &amp; Ng, P. K. L." box="[551,764,1591,1617]" pageId="9" pageNumber="230" pagination="496 - 508" refId="ref15197" refString="Trivedi, J. N., Gosavi, S., Vachhrajani, K. D., Mitra, S., Ravinesh, R. &amp; Ng, P. K. L. (2020) On the identities of Nepinnotheres vicajii (Chhapgar, 1957) and Arcotheres casta (Antony &amp; Kuttyamma, 1971) from western India: conspecificity and taxonomy (Decapoda, Brachyura, Pinnotheridae). Zootaxa, 4809 (3), 496 - 508. https: // doi. org / 10.11646 / zootaxa. 4809.3.4" type="journal article" year="2020">
Trivedi
<emphasis id="A9C8EADD9507426471B5F9F0FD6DC438" box="[640,697,1591,1617]" italics="true" pageId="9" pageNumber="230">et al.</emphasis>
2020
</bibRefCitation>
: figs. 4A, B, D, E, F, H, 6C, D, FK]; the longer female P4 has a dactylus that is proportionately shorter (
<figureCitation id="03872A4A9507426471AFF99CFD3BC41C" box="[666,751,1627,1653]" captionStart="FIGURE 4" captionStartId="6.[151,250,1862,1886]" captionTargetBox="[226,1364,190,1827]" captionTargetId="figure-20@6.[216,1371,181,1838]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres obesus (Dana, 1852), neotype, female of post-hard stage V (4.4×5.5 mm), CBM-ZC 15945. A, right chela, outer view; B, same, inner view; C, chela and carpus of right cheliped, dorsal view; D, distal part of fixed finger of right chela, ventral view; E, carpus and merus of right cheliped, ventral view; F, right P2, dorsal view; G, same dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (shorter side), dorsal view; K, same, dactylus; L, left P4 (longer side), dorsal view; M, same dactylus; N, right P5, dorsal view; O, same dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401471" httpUri="https://zenodo.org/record/4401471/files/figure.png" pageId="9" pageNumber="230">Fig. 4L</figureCitation>
) [versus longer female P4 dactylus proportionately longer in the
<taxonomicName id="5CBC4D4C9507426473F5F9B8FEE1C4F1" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[192,309,1663,1689]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="A9C8EADD9507426473F5F9B8FEE1C4F1" box="[192,309,1663,1689]" italics="true" pageId="9" pageNumber="230">A. exiguus</emphasis>
</taxonomicName>
group; cf.
<bibRefCitation id="FF2D4B3E95074264729BF9B8FD99C4F0" author="Gordon, I." box="[430,589,1663,1689]" pageId="9" pageNumber="230" pagination="163 - 180" refId="ref13490" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of the Linnean Society of London, Zoology, 40, 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article" year="1936">Gordon, 1936</bibRefCitation>
: fig. 7a;
<bibRefCitation id="FF2D4B3E95074264719EF9B8FC3CC4F0" author="Antony, A. &amp; Kuttyamma, V. J." box="[683,1000,1663,1689]" pageId="9" pageNumber="230" pagination="59 - 68" refId="ref12654" refString="Antony, A. &amp; Kuttyamma, V. J. (1971) A new species of crab Pinnotheres Latreille (Crustacea: Brachyura: Pinnotheridae) from the clam Meretrix casta (Chemnitz). Bulletin of Department of Marine Biology and Oceanography, 5, 59 - 68." type="journal article" year="1971">Antony &amp; Kuttyamma 1971</bibRefCitation>
: text-fig. 2(4);
<bibRefCitation id="FF2D4B3E9507426477BAF9B8FAB5C4F0" author="Trivedi, J. N. &amp; Gosavi, S. &amp; Vachhrajani, K. D. &amp; Mitra, S. &amp; Ravinesh, R. &amp; Ng, P. K. L." box="[1167,1377,1663,1689]" pageId="9" pageNumber="230" pagination="496 - 508" refId="ref15197" refString="Trivedi, J. N., Gosavi, S., Vachhrajani, K. D., Mitra, S., Ravinesh, R. &amp; Ng, P. K. L. (2020) On the identities of Nepinnotheres vicajii (Chhapgar, 1957) and Arcotheres casta (Antony &amp; Kuttyamma, 1971) from western India: conspecificity and taxonomy (Decapoda, Brachyura, Pinnotheridae). Zootaxa, 4809 (3), 496 - 508. https: // doi. org / 10.11646 / zootaxa. 4809.3.4" type="journal article" year="2020">
Trivedi
<emphasis id="A9C8EADD9507426477D2F9B8FACBC4F0" box="[1255,1311,1663,1689]" italics="true" pageId="9" pageNumber="230">et al.</emphasis>
2020
</bibRefCitation>
: figs. 4G, 6E] and the G1 is relatively stouter and gently curved, being a broad C-shaped with a subtruncate tip (
<figureCitation id="03872A4A950742647610F964FA40C4D4" box="[1317,1428,1698,1725]" captionStart="FIGURE 5" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[157,1424,499,1834]" captionTargetId="figure-186@8.[151,1436,489,1844]" captionTargetPageId="8" captionText="FIGURE 5. Arcotheres obesus (Dana, 1852), male of post-hard stage (3.6×ca 4.2 mm), CBM-ZC 15946. A, habitus, dorsal view; B, left MXP3, outer view; C, left thoracic sternum, ventral view; D, pleon, telson and exposed part of sternite 8 (left), ventral view; E, left G1, outer view; F, same, inner view; G, left G2, outer view; H, same, different view." figureDoi="http://doi.org/10.5281/zenodo.4401473" httpUri="https://zenodo.org/record/4401473/files/figure.png" pageId="9" pageNumber="230">Fig. 5E, F</figureCitation>
) [versus G1 relatively more slender, distinctly C-shaped with a short subterminal projection in the
<taxonomicName id="5CBC4D4C9507426477EDF900FA99C489" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1240,1357,1735,1761]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="A9C8EADD9507426477EDF900FA99C489" box="[1240,1357,1735,1761]" italics="true" pageId="9" pageNumber="230">A. exiguus</emphasis>
</taxonomicName>
group; cf.
<bibRefCitation id="FF2D4B3E95074264738CF92CFE2CC56C" author="Antony, A. &amp; Kuttyamma, V. J." box="[185,504,1771,1797]" pageId="9" pageNumber="230" pagination="59 - 68" refId="ref12654" refString="Antony, A. &amp; Kuttyamma, V. J. (1971) A new species of crab Pinnotheres Latreille (Crustacea: Brachyura: Pinnotheridae) from the clam Meretrix casta (Chemnitz). Bulletin of Department of Marine Biology and Oceanography, 5, 59 - 68." type="journal article" year="1971">Antony &amp; Kuttyamma 1971</bibRefCitation>
: text-fig. 1(5, 6);
<bibRefCitation id="FF2D4B3E950742647189F92CFC44C56C" author="Trivedi, J. N. &amp; Gosavi, S. &amp; Vachhrajani, K. D. &amp; Mitra, S. &amp; Ravinesh, R. &amp; Ng, P. K. L." box="[700,912,1771,1797]" pageId="9" pageNumber="230" pagination="496 - 508" refId="ref15197" refString="Trivedi, J. N., Gosavi, S., Vachhrajani, K. D., Mitra, S., Ravinesh, R. &amp; Ng, P. K. L. (2020) On the identities of Nepinnotheres vicajii (Chhapgar, 1957) and Arcotheres casta (Antony &amp; Kuttyamma, 1971) from western India: conspecificity and taxonomy (Decapoda, Brachyura, Pinnotheridae). Zootaxa, 4809 (3), 496 - 508. https: // doi. org / 10.11646 / zootaxa. 4809.3.4" type="journal article" year="2020">
Trivedi
<emphasis id="A9C8EADD950742647020F92CFC99C56C" box="[789,845,1771,1797]" italics="true" pageId="9" pageNumber="230">et al.</emphasis>
2020
</bibRefCitation>
: figs. 3D, 7H, I].
</paragraph>
<paragraph id="9B0336CF9507426473F2F8C8FDEFC518" blockId="9.[151,1437,150,1905]" pageId="9" pageNumber="230">
In addition to the good series of specimens of
<taxonomicName id="5CBC4D4C9507426471F4F8C8FCFFC540" authorityName=", Dana" authorityYear="1852" box="[705,811,1807,1833]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="230" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="A9C8EADD9507426471F4F8C8FCFFC540" box="[705,811,1807,1833]" italics="true" pageId="9" pageNumber="230">A. obesus</emphasis>
</taxonomicName>
from
<collectingCountry id="E3AB765F950742647058F8C8FC43C540" box="[877,919,1807,1833]" name="Fiji" pageId="9" pageNumber="230">Fiji</collectingCountry>
, we also examined two specimens from Peninsular
<collectingCountry id="E3AB765F9507426473E1F8F4FEEFC524" box="[212,315,1843,1869]" name="Malaysia" pageId="9" pageNumber="230">Malaysia</collectingCountry>
(ZRC 2000.2281) which are clearly referable to this species. Noteworthy is that both were collected from the reef clam
<taxonomicName id="5CBC4D4C95074264725BF890FDE1C518" baseAuthorityName="Holten" baseAuthorityYear="1802" box="[366,565,1879,1905]" class="Bivalvia" family="Veneridae" genus="Gafrarium" kingdom="Animalia" order="Veneroida" pageId="9" pageNumber="230" phylum="Mollusca" rank="species" species="dispar">
<emphasis id="A9C8EADD95074264725BF890FDE1C518" box="[366,565,1879,1905]" italics="true" pageId="9" pageNumber="230">Gafrarium dispar</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>