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<document id="5D65565920DA09347A6D4BD0475BED33" ID-CLB-Dataset="42386" ID-DOI="10.11646/zootaxa.3905.4.3" ID-GBIF-Dataset="a0918429-1be8-4836-a9b1-17589de87b0f" ID-ISSN="1175-5326" ID-Zenodo-Dep="242883" ID-ZooBank="489B8CE1-6AEF-4D31-8CB6-4CD94B81CBE6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460934794706" checkinUser="plazi" docAuthor="Zhang, Feng" docDate="2015" docId="03FB87EAFF96FFE9FF1EFD1CFC60FF08" docLanguage="en" docName="zt03905p499.pdf" docOrigin="Zootaxa 3905 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Willowsia cassagnaui Zhang, 2015, sp. nov." docType="treatment" docVersion="10" lastPageNumber="494" masterDocId="FFC2FF92FF97FFEFFF89FFA9FFE0FFDD" masterDocTitle="Some Willowsia from Nepal and Vietnam (Collembola: Entomobryidae) and description of one new species" masterLastPageNumber="499" masterPageNumber="489" pageNumber="490" updateTime="1698582648178" updateUser="ExternalLinkService">
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<mods:title id="C179C8527DF96A6EF6BC8BE51A9C981F">Some Willowsia from Nepal and Vietnam (Collembola: Entomobryidae) and description of one new species</mods:title>
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<mods:namePart id="50D11FACF131ECABB3F10D03110279A5">Zhang, Feng</mods:namePart>
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<mods:date id="35711C2D030FA23D6DD584CC8839EF46">2015</mods:date>
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<treatment id="03FB87EAFF96FFE9FF1EFD1CFC60FF08" ID-DOI="http://doi.org/10.5281/zenodo.5689648" ID-GBIF-Taxon="119590459" ID-Zenodo-Dep="5689648" LSID="urn:lsid:plazi:treatment:03FB87EAFF96FFE9FF1EFD1CFC60FF08" httpUri="http://treatment.plazi.org/id/03FB87EAFF96FFE9FF1EFD1CFC60FF08" lastPageId="6" lastPageNumber="494" pageId="1" pageNumber="490">
<subSubSection id="C3486577FF96FFEEFF1EFD1CFD89FC79" pageId="1" pageNumber="490" type="nomenclature">
<paragraph id="8BED36FCFF96FFEEFF1EFD1CFDE0FD12" blockId="1.[151,512,693,753]" box="[151,512,693,719]" pageId="1" pageNumber="490">
<heading id="D0A58190FF96FFEEFF1EFD1CFDE0FD12" bold="true" box="[151,512,693,719]" fontSize="11" level="1" pageId="1" pageNumber="490" reason="1">
<emphasis id="B926EAEEFF96FFEEFF1EFD1CFDE0FD12" bold="true" box="[151,512,693,719]" pageId="1" pageNumber="490">
<taxonomicName id="4C524D7FFF96FFEEFF1EFD1CFE7BFD12" box="[151,411,693,719]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="1" pageNumber="490" phylum="Arthropoda" rank="species" species="cassagnaui" status="sp. nov.">
<emphasis id="B926EAEEFF96FFEEFF1EFD1CFE7BFD12" bold="true" box="[151,411,693,719]" italics="true" pageId="1" pageNumber="490">Willowsia cassagnaui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2155795FF96FFEEFE2BFD1CFDE0FD12" box="[418,512,693,719]" pageId="1" pageNumber="490" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BED36FCFF96FFEEFF1EFD71FEE8FD2C" blockId="1.[151,512,693,753]" box="[151,264,728,753]" pageId="1" pageNumber="490">
<figureCitation id="13692A79FF96FFEEFF1EFD71FEE8FD2C" box="[151,264,728,753]" captionStart-0="FIGURES 1 12. W" captionStart-1="FIGURES 13 15. W" captionStartId-0="2.[151,264,1908,1931]" captionStartId-1="3.[151,264,772,795]" captionTargetBox-0="[198,1369,198,1864]" captionTargetBox-1="[158,1427,204,728]" captionTargetId-0="figure@2.[189,1376,193,1872]" captionTargetId-1="figure@3.[150,1436,193,737]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 12. W. cassagnaui sp. nov.: 1, habitus; 2, scales on terga; 3, scales on manubrium; 4, labrum; 5, lateral process of labial palp; 6, maxillary outer lobe; 7, labial chaetae; 8, dorsal cephalic chaetotaxy; 9, chaetotaxy of Th. II Abd. III; 10, trochanteral organ; 11, hind claw; 12, chaetotaxy of Abd. IV. Larger circles, macrochaetae; smaller circles, microchaetae; circle with a slash, pseudopore; intersectional lines, bothriotricha." captionText-1="FIGURES 13 15. W. cassagnaui sp. nov.: 13, anterior face of ventral tube; 14, posterior face and lateral flap of ventral tube; 15, mucro." httpUri-0="https://zenodo.org/record/242884/files/figure.png" httpUri-1="https://zenodo.org/record/242885/files/figure.png" pageId="1" pageNumber="490">Figs 115</figureCitation>
</paragraph>
<paragraph id="8BED36FCFF96FFEEFF1EFC89FD89FC79" blockId="1.[151,1437,800,1977]" pageId="1" pageNumber="490">
<emphasis id="B926EAEEFF96FFEEFF1EFC89FF03FCE4" bold="true" box="[151,227,800,825]" pageId="1" pageNumber="490">
<typeStatus id="54E9885EFF96FFEEFF1EFC89FF3FFCE4" box="[151,223,800,825]" pageId="1" pageNumber="490">Types</typeStatus>
.
</emphasis>
<typeStatus id="54E9885EFF96FFEEFF79FC89FE88FCE4" box="[240,360,800,825]" pageId="1" pageNumber="490" type="holotype">Holotypes</typeStatus>
: female on slide,
<collectingCountry id="F345766CFF96FFEEFDCAFC89FD6BFCE4" box="[579,651,800,825]" name="Nepal" pageId="1" pageNumber="490">Nepal</collectingCountry>
: Machapuchare, alt.
<quantity id="4CAA9B19FF96FFEEFC0AFC89FC01FCE5" box="[899,993,800,824]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="490" unit="m" value="2200.0">2200 m</quantity>
, above Pokhara, forest of large trees (
<taxonomicName id="4C524D7FFF96FFEEFF29FCEDFDD8FC81" authority="Sm. In Rees" authorityName="Sm. In Rees" box="[160,568,836,860]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="490" phylum="Tracheophyta" rank="species" species="semicarpifolia">
<emphasis id="B926EAEEFF96FFEEFF29FCEDFE47FC81" box="[160,423,836,860]" italics="true" pageId="1" pageNumber="490">Quercus semicarpifolia</emphasis>
Sm. In Rees
</taxonomicName>
,
<taxonomicName id="4C524D7FFF96FFEEFDCAFCEDFD98FC86" box="[579,632,836,859]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="1" pageNumber="490" phylum="Tracheophyta" rank="genus">
<emphasis id="B926EAEEFF96FFEEFDCAFCEDFD98FC86" box="[579,632,836,859]" italics="true" pageId="1" pageNumber="490">Acer</emphasis>
</taxonomicName>
sp., rhododendrons, bamboos etc.), dry moss on a branch,
<date id="FFEC103CFF96FFEEFA82FCEDFA9DFC80" box="[1291,1405,836,861]" pageId="1" pageNumber="490" value="1976-10-26">26.x.1976</date>
, P. Cassagnau leg. (# Nep. 76.51).
<typeStatus id="54E9885EFF96FFEEFE72FCC0FD8CFC5C" box="[507,620,873,897]" pageId="1" pageNumber="490" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D54FD75FF96FFEEFDFEFCC1FD39FC5C" box="[631,729,872,897]" pageId="1" pageNumber="490" type="female">4 female</specimenCount>
and 4 young specimens on slides, same data as
<typeStatus id="54E9885EFF96FFEEFB73FCC1FA80FC5C" box="[1274,1376,872,897]" pageId="1" pageNumber="490" type="holotype">holotype</typeStatus>
. Two
<typeStatus id="54E9885EFF96FFEEFF1EFC24FEE2FC78" box="[151,258,909,933]" pageId="1" pageNumber="490" type="paratype">paratypes</typeStatus>
in NJAU and others in MNHN.
</paragraph>
</subSubSection>
<subSubSection id="C3486577FF96FFEEFF4EFC19FB8FF890" pageId="1" pageNumber="490" type="description">
<paragraph id="8BED36FCFF96FFEEFF4EFC19FD7BFC15" blockId="1.[151,1437,800,1977]" box="[199,667,944,969]" pageId="1" pageNumber="490">
<emphasis id="B926EAEEFF96FFEEFF4EFC19FEB9FC14" bold="true" box="[199,345,944,969]" pageId="1" pageNumber="490">Description.</emphasis>
Body length up to
<quantity id="4CAA9B19FF96FFEEFDA6FC19FD76FC15" box="[559,662,944,968]" metricMagnitude="-3" metricUnit="m" metricValue="1.82" pageId="1" pageNumber="490" unit="mm" value="1.82">1.82 mm</quantity>
.
</paragraph>
<paragraph id="8BED36FCFF96FFEEFF4EFC7DFCBCFB18" blockId="1.[151,1437,800,1977]" pageId="1" pageNumber="490">
Ground colour pale yellow in alcohol. Blue pigment forming irregular patterns on body. Ant. I IV gradually becoming darker from anterior to posterior. A transverse band between the eye patches and a small lateral patch behind each eye patch. Lateral patches present from Th. II to Abd. VI. A pair of lighter small patches on median Th. III and Abd. I. Two transverse bands on posterior Abd. II and III. A pair of large median and small posterior patches on Abd. IV. Legs slightly pigmented. Furcula pale (
<figureCitation id="13692A79FF96FFEEFCCEFBCDFC6BFBA1" box="[839,907,1124,1149]" captionStart="FIGURES 1 12. W" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[198,1369,198,1864]" captionTargetId="figure@2.[189,1376,193,1872]" captionTargetPageId="2" captionText="FIGURES 1 12. W. cassagnaui sp. nov.: 1, habitus; 2, scales on terga; 3, scales on manubrium; 4, labrum; 5, lateral process of labial palp; 6, maxillary outer lobe; 7, labial chaetae; 8, dorsal cephalic chaetotaxy; 9, chaetotaxy of Th. II Abd. III; 10, trochanteral organ; 11, hind claw; 12, chaetotaxy of Abd. IV. Larger circles, macrochaetae; smaller circles, microchaetae; circle with a slash, pseudopore; intersectional lines, bothriotricha." httpUri="https://zenodo.org/record/242884/files/figure.png" pageId="1" pageNumber="490">Fig. 1</figureCitation>
). Scales on body pointed, striate heavily with basal ribs longer than distal ones (
<figureCitation id="13692A79FF96FFEEFDACFB21FD8CFB7D" box="[549,620,1160,1185]" captionStart="FIGURES 1 12. W" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[198,1369,198,1864]" captionTargetId="figure@2.[189,1376,193,1872]" captionTargetPageId="2" captionText="FIGURES 1 12. W. cassagnaui sp. nov.: 1, habitus; 2, scales on terga; 3, scales on manubrium; 4, labrum; 5, lateral process of labial palp; 6, maxillary outer lobe; 7, labial chaetae; 8, dorsal cephalic chaetotaxy; 9, chaetotaxy of Th. II Abd. III; 10, trochanteral organ; 11, hind claw; 12, chaetotaxy of Abd. IV. Larger circles, macrochaetae; smaller circles, microchaetae; circle with a slash, pseudopore; intersectional lines, bothriotricha." httpUri="https://zenodo.org/record/242884/files/figure.png" pageId="1" pageNumber="490">Fig. 2</figureCitation>
); scales on ventral manubrium obviously narrower than those on terga (
<figureCitation id="13692A79FF96FFEEFF16FB05FF04FB19" box="[159,228,1196,1221]" captionStart="FIGURES 1 12. W" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[198,1369,198,1864]" captionTargetId="figure@2.[189,1376,193,1872]" captionTargetPageId="2" captionText="FIGURES 1 12. W. cassagnaui sp. nov.: 1, habitus; 2, scales on terga; 3, scales on manubrium; 4, labrum; 5, lateral process of labial palp; 6, maxillary outer lobe; 7, labial chaetae; 8, dorsal cephalic chaetotaxy; 9, chaetotaxy of Th. II Abd. III; 10, trochanteral organ; 11, hind claw; 12, chaetotaxy of Abd. IV. Larger circles, macrochaetae; smaller circles, microchaetae; circle with a slash, pseudopore; intersectional lines, bothriotricha." httpUri="https://zenodo.org/record/242884/files/figure.png" pageId="1" pageNumber="490">Fig. 3</figureCitation>
). Scales absent on antennae, legs, ventral tube and dens.
</paragraph>
<paragraph id="8BED36FCFF96FFEEFF4EFB78FF11FA41" blockId="1.[151,1437,800,1977]" pageId="1" pageNumber="490">
Antenna twice as long as cephalic diagonal. Antennal segments ratio as I: II: III: IV = 1: 1.9: 1.8: 2.9. Ant. III organ not clearly seen. Antennal apical bulb bilobed. Labral margin with 4 rounded papillae. Labral intrusion slightly curved (
<figureCitation id="13692A79FF96FFEEFED0FAB1FE44FAED" box="[345,420,1304,1329]" captionStart="FIGURES 1 12. W" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[198,1369,198,1864]" captionTargetId="figure@2.[189,1376,193,1872]" captionTargetPageId="2" captionText="FIGURES 1 12. W. cassagnaui sp. nov.: 1, habitus; 2, scales on terga; 3, scales on manubrium; 4, labrum; 5, lateral process of labial palp; 6, maxillary outer lobe; 7, labial chaetae; 8, dorsal cephalic chaetotaxy; 9, chaetotaxy of Th. II Abd. III; 10, trochanteral organ; 11, hind claw; 12, chaetotaxy of Abd. IV. Larger circles, macrochaetae; smaller circles, microchaetae; circle with a slash, pseudopore; intersectional lines, bothriotricha." httpUri="https://zenodo.org/record/242884/files/figure.png" pageId="1" pageNumber="490">Fig. 4</figureCitation>
). Tip of lateral process of labial palp nearly reaching apex of labial papilla (
<figureCitation id="13692A79FF96FFEEFACDFAB1FA6EFAEC" box="[1348,1422,1304,1329]" captionStart="FIGURES 1 12. W" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[198,1369,198,1864]" captionTargetId="figure@2.[189,1376,193,1872]" captionTargetPageId="2" captionText="FIGURES 1 12. W. cassagnaui sp. nov.: 1, habitus; 2, scales on terga; 3, scales on manubrium; 4, labrum; 5, lateral process of labial palp; 6, maxillary outer lobe; 7, labial chaetae; 8, dorsal cephalic chaetotaxy; 9, chaetotaxy of Th. II Abd. III; 10, trochanteral organ; 11, hind claw; 12, chaetotaxy of Abd. IV. Larger circles, macrochaetae; smaller circles, microchaetae; circle with a slash, pseudopore; intersectional lines, bothriotricha." httpUri="https://zenodo.org/record/242884/files/figure.png" pageId="1" pageNumber="490">Fig. 5</figureCitation>
). Subapical chaeta of maxillary outer lobe slightly stronger than apical one; 3 hairs on sublobal plate (
<figureCitation id="13692A79FF96FFEEFB72FA95FADFFA88" box="[1275,1343,1340,1365]" captionStart="FIGURES 1 12. W" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[198,1369,198,1864]" captionTargetId="figure@2.[189,1376,193,1872]" captionTargetPageId="2" captionText="FIGURES 1 12. W. cassagnaui sp. nov.: 1, habitus; 2, scales on terga; 3, scales on manubrium; 4, labrum; 5, lateral process of labial palp; 6, maxillary outer lobe; 7, labial chaetae; 8, dorsal cephalic chaetotaxy; 9, chaetotaxy of Th. II Abd. III; 10, trochanteral organ; 11, hind claw; 12, chaetotaxy of Abd. IV. Larger circles, macrochaetae; smaller circles, microchaetae; circle with a slash, pseudopore; intersectional lines, bothriotricha." httpUri="https://zenodo.org/record/242884/files/figure.png" pageId="1" pageNumber="490">Fig. 6</figureCitation>
). Labial chaeta R 0.40.5 x the length of M (
<figureCitation id="13692A79FF96FFEEFDA7FAC9FD92FAA5" box="[558,626,1376,1401]" captionStart="FIGURES 1 12. W" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[198,1369,198,1864]" captionTargetId="figure@2.[189,1376,193,1872]" captionTargetPageId="2" captionText="FIGURES 1 12. W. cassagnaui sp. nov.: 1, habitus; 2, scales on terga; 3, scales on manubrium; 4, labrum; 5, lateral process of labial palp; 6, maxillary outer lobe; 7, labial chaetae; 8, dorsal cephalic chaetotaxy; 9, chaetotaxy of Th. II Abd. III; 10, trochanteral organ; 11, hind claw; 12, chaetotaxy of Abd. IV. Larger circles, macrochaetae; smaller circles, microchaetae; circle with a slash, pseudopore; intersectional lines, bothriotricha." httpUri="https://zenodo.org/record/242884/files/figure.png" pageId="1" pageNumber="490">Fig. 7</figureCitation>
). Dorsal cephalic chaetotaxy with 6 antennal (An) and 5 sutural (S) mac (
<figureCitation id="13692A79FF96FFEEFF16FA2DFF04FA41" box="[159,228,1412,1437]" captionStart="FIGURES 1 12. W" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[198,1369,198,1864]" captionTargetId="figure@2.[189,1376,193,1872]" captionTargetPageId="2" captionText="FIGURES 1 12. W. cassagnaui sp. nov.: 1, habitus; 2, scales on terga; 3, scales on manubrium; 4, labrum; 5, lateral process of labial palp; 6, maxillary outer lobe; 7, labial chaetae; 8, dorsal cephalic chaetotaxy; 9, chaetotaxy of Th. II Abd. III; 10, trochanteral organ; 11, hind claw; 12, chaetotaxy of Abd. IV. Larger circles, macrochaetae; smaller circles, microchaetae; circle with a slash, pseudopore; intersectional lines, bothriotricha." httpUri="https://zenodo.org/record/242884/files/figure.png" pageId="1" pageNumber="490">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8BED36FCFF96FFEEFF4EFA01FC9AF98D" blockId="1.[151,1437,800,1977]" pageId="1" pageNumber="490">
Th. II with 10 posterior mac; median mac absent; p46 and p5pi as mac. Th. III with 11 mac (
<figureCitation id="13692A79FF96FFEEFB56FA01FAC4FA1D" box="[1247,1316,1448,1473]" captionStart="FIGURES 1 12. W" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[198,1369,198,1864]" captionTargetId="figure@2.[189,1376,193,1872]" captionTargetPageId="2" captionText="FIGURES 1 12. W. cassagnaui sp. nov.: 1, habitus; 2, scales on terga; 3, scales on manubrium; 4, labrum; 5, lateral process of labial palp; 6, maxillary outer lobe; 7, labial chaetae; 8, dorsal cephalic chaetotaxy; 9, chaetotaxy of Th. II Abd. III; 10, trochanteral organ; 11, hind claw; 12, chaetotaxy of Abd. IV. Larger circles, macrochaetae; smaller circles, microchaetae; circle with a slash, pseudopore; intersectional lines, bothriotricha." httpUri="https://zenodo.org/record/242884/files/figure.png" pageId="1" pageNumber="490">Fig. 9</figureCitation>
). Thoracic S-chaetae not clearly seen. Metatrochanteral organ with about 14 smooth spiny chaetae;
<quantity id="4CAA9B19FF96FFEEFBF4FA65FB57FA38" box="[1149,1207,1484,1509]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="490" unit="in" value="10.0">10 in</quantity>
arms, V-shaped and 4 between arms (
<figureCitation id="13692A79FF96FFEEFED0FA59FE4DF9D5" box="[345,429,1520,1545]" captionStart="FIGURES 1 12. W" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[198,1369,198,1864]" captionTargetId="figure@2.[189,1376,193,1872]" captionTargetPageId="2" captionText="FIGURES 1 12. W. cassagnaui sp. nov.: 1, habitus; 2, scales on terga; 3, scales on manubrium; 4, labrum; 5, lateral process of labial palp; 6, maxillary outer lobe; 7, labial chaetae; 8, dorsal cephalic chaetotaxy; 9, chaetotaxy of Th. II Abd. III; 10, trochanteral organ; 11, hind claw; 12, chaetotaxy of Abd. IV. Larger circles, macrochaetae; smaller circles, microchaetae; circle with a slash, pseudopore; intersectional lines, bothriotricha." httpUri="https://zenodo.org/record/242884/files/figure.png" pageId="1" pageNumber="490">Fig. 10</figureCitation>
). Unguis with 4 inner teeth; basal inner pair with tip reaching 0.45 distance from base of ventral edge, median one at 0.71 and distal one 0.87 distance from base. Unguiculus lanceolate with outer edge slightly serrate. Tenent hair subequal to unguis in length (
<figureCitation id="13692A79FF96FFEEFC92F991FC8DF98D" box="[795,877,1592,1617]" captionStart="FIGURES 1 12. W" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[198,1369,198,1864]" captionTargetId="figure@2.[189,1376,193,1872]" captionTargetPageId="2" captionText="FIGURES 1 12. W. cassagnaui sp. nov.: 1, habitus; 2, scales on terga; 3, scales on manubrium; 4, labrum; 5, lateral process of labial palp; 6, maxillary outer lobe; 7, labial chaetae; 8, dorsal cephalic chaetotaxy; 9, chaetotaxy of Th. II Abd. III; 10, trochanteral organ; 11, hind claw; 12, chaetotaxy of Abd. IV. Larger circles, macrochaetae; smaller circles, microchaetae; circle with a slash, pseudopore; intersectional lines, bothriotricha." httpUri="https://zenodo.org/record/242884/files/figure.png" pageId="1" pageNumber="490">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="8BED36FCFF96FFEEFF4EF9F5FB8FF890" blockId="1.[151,1437,800,1977]" pageId="1" pageNumber="490">
Abd. IV four times Abd. III in length along dorsal midline. Abd. I with 3 (m24) mac and 2 S-chaetae; ms internal to sens. Abd. II with 3 (a2, m3, m3e) central, 1 (m5) lateral mac and 2 sens. Abd. III with 2 (a2, m3) central, 3 (am6, pm6, p6) lateral mac, 1 ms and 2 sens. Abd. IV with 7 central mac and about 12 sens; homology of anterior three mac unclear; 3 lateral sens obviously shorter than others (
<figureCitation id="13692A79FF96FFEEFC40F961FBFEF93D" box="[969,1054,1736,1761]" captionStart="FIGURES 1 12. W" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[198,1369,198,1864]" captionTargetId="figure@2.[189,1376,193,1872]" captionTargetPageId="2" captionText="FIGURES 1 12. W. cassagnaui sp. nov.: 1, habitus; 2, scales on terga; 3, scales on manubrium; 4, labrum; 5, lateral process of labial palp; 6, maxillary outer lobe; 7, labial chaetae; 8, dorsal cephalic chaetotaxy; 9, chaetotaxy of Th. II Abd. III; 10, trochanteral organ; 11, hind claw; 12, chaetotaxy of Abd. IV. Larger circles, macrochaetae; smaller circles, microchaetae; circle with a slash, pseudopore; intersectional lines, bothriotricha." httpUri="https://zenodo.org/record/242884/files/figure.png" pageId="1" pageNumber="490">Fig. 12</figureCitation>
). Accessory mic of bothriotrichal complexes on Abd. II IV unmodified. Ventral tube anteriorly with about 14 ciliate chaetae on each side (
<figureCitation id="13692A79FF96FFEEFAB2F945FA6EF8D9" box="[1339,1422,1772,1797]" captionStart="FIGURES 13 15. W" captionStartId="3.[151,264,772,795]" captionTargetBox="[158,1427,204,728]" captionTargetId="figure@3.[150,1436,193,737]" captionTargetPageId="3" captionText="FIGURES 13 15. W. cassagnaui sp. nov.: 13, anterior face of ventral tube; 14, posterior face and lateral flap of ventral tube; 15, mucro." httpUri="https://zenodo.org/record/242885/files/figure.png" pageId="1" pageNumber="490">Fig. 13</figureCitation>
), 2 of them larger than others; posteriorly with 11 chaetae, apical two smooth; each lateral flap with 7 smooth and 1 ciliate chaetae (
<figureCitation id="13692A79FF96FFEEFECFF89DFE79F891" box="[326,409,1844,1869]" captionStart="FIGURES 13 15. W" captionStartId="3.[151,264,772,795]" captionTargetBox="[158,1427,204,728]" captionTargetId="figure@3.[150,1436,193,737]" captionTargetPageId="3" captionText="FIGURES 13 15. W. cassagnaui sp. nov.: 13, anterior face of ventral tube; 14, posterior face and lateral flap of ventral tube; 15, mucro." httpUri="https://zenodo.org/record/242885/files/figure.png" pageId="1" pageNumber="490">Fig. 14</figureCitation>
). Distal smooth part of dens 1.92.5 x mucro in length (
<figureCitation id="13692A79FF96FFEEFB87F89DFB81F890" box="[1038,1121,1844,1869]" captionStart="FIGURES 13 15. W" captionStartId="3.[151,264,772,795]" captionTargetBox="[158,1427,204,728]" captionTargetId="figure@3.[150,1436,193,737]" captionTargetPageId="3" captionText="FIGURES 13 15. W. cassagnaui sp. nov.: 13, anterior face of ventral tube; 14, posterior face and lateral flap of ventral tube; 15, mucro." httpUri="https://zenodo.org/record/242885/files/figure.png" pageId="1" pageNumber="490">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3486577FF96FFEEFF4EF8F1FB43F8AC" box="[199,1187,1880,1905]" pageId="1" pageNumber="490" type="etymology">
<paragraph id="8BED36FCFF96FFEEFF4EF8F1FB43F8AC" blockId="1.[151,1437,800,1977]" box="[199,1187,1880,1905]" pageId="1" pageNumber="490">
<emphasis id="B926EAEEFF96FFEEFF4EF8F1FEAEF8AC" bold="true" box="[199,334,1880,1905]" pageId="1" pageNumber="490">Etymology.</emphasis>
The new species is named after P. Cassagnau, who collected the specimens.
</paragraph>
</subSubSection>
<subSubSection id="C3486577FF96FFECFF4EF8D5FE0AFBEF" lastPageId="3" lastPageNumber="492" pageId="1" pageNumber="490" type="discussion">
<paragraph id="8BED36FCFF96FFECFF4EF8D5FE0AFBEF" blockId="1.[151,1437,800,1977]" lastBlockId="3.[151,1436,870,1075]" lastPageId="3" lastPageNumber="492" pageId="1" pageNumber="490">
<emphasis id="B926EAEEFF96FFEEFF4EF8D5FEDBF848" bold="true" box="[199,315,1916,1941]" pageId="1" pageNumber="490">Remarks.</emphasis>
<taxonomicName id="4C524D7FFF96FFEEFECBF8D5FDD7F849" box="[322,567,1916,1940]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="1" pageNumber="490" phylum="Arthropoda" rank="species" species="cassagnaui" status="sp. nov.">
<emphasis id="B926EAEEFF96FFEEFECBF8D5FDD7F849" box="[322,567,1916,1940]" italics="true" pageId="1" pageNumber="490">Willowsia cassagnaui</emphasis>
</taxonomicName>
<emphasis id="B926EAEEFF96FFEEFDB7F8D5FD78F848" bold="true" box="[574,664,1916,1941]" pageId="1" pageNumber="490">
<taxonomicNameLabel id="A2155795FF96FFEEFDB7F8D5FD78F848" box="[574,664,1916,1941]" pageId="1" pageNumber="490" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="4C524D7FFF96FFEEFCA4F8D4FC1CF849" authority="Yosii, 1971" authorityName="Yosii" authorityYear="1971" box="[813,1020,1916,1941]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="1" pageNumber="490" phylum="Arthropoda" rank="species" species="ieti">
<emphasis id="B926EAEEFF96FFEEFCA4F8D4FC95F849" box="[813,885,1917,1940]" italics="true" pageId="1" pageNumber="490">W. ieti</emphasis>
<bibRefCitation id="EFC34B0DFF96FFEEFCF5F8D5FC1CF849" author="Yosii" box="[892,1020,1916,1941]" pageId="1" pageNumber="490" refString="Yosii, R. (1971) Collembola of Khumbu Himal. Khumbu Himal, 4, 80 - 130." type="journal article" year="1971">
Yosii,
<quantity id="4CAA9B19FF96FFEEFC4DF8D5FC1CF849" box="[964,1020,1916,1940]" metricMagnitude="1" metricUnit="m" metricValue="5.00634" pageId="1" pageNumber="490" unit="in" value="1971.0">1971</quantity>
</bibRefCitation>
</taxonomicName>
in most features. It differs from it in colour pattern, lateral process of labial palp longer with tip nearly reaching apex of labial papilla, slightly curved labral intrusion, fewer chaetae on trochanteral organ (14 vs&gt;20), lanceolate unguiculus, presence of mac m2 on Abd. I, and absence of scales on Ant. I (
<tableCitation id="C6D00347FF94FFECFDE2FC23FD22FC7E" box="[619,706,906,931]" captionStart="TABLE 1" captionStartId="3.[151,239,1119,1142]" captionTargetBox="[151,1180,1170,1789]" captionTargetPageId="3" captionText="TABLE 1. Comparison between W. cassagnaui sp. nov. and W. ieti." httpUri="http://table.plazi.org/id/DF2D6674FF94FFECFF1EFBF6FC8CFBAB" pageId="3" pageNumber="492" tableUuid="DF2D6674FF94FFECFF1EFBF6FC8CFBAB">Table 1</tableCitation>
). Lateral stripes, transverse band on Abd. II, and paired lighter small patches on median Th. III, Abd. I and Abd. IV are present in
<taxonomicName id="4C524D7FFF94FFECFC0AFC06FBC3FC1B" box="[899,1059,943,966]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="492" phylum="Arthropoda" rank="species" species="cassagnaui" status="sp. nov.">
<emphasis id="B926EAEEFF94FFECFC0AFC06FBC3FC1B" box="[899,1059,943,966]" italics="true" pageId="3" pageNumber="492">W. cassagnaui</emphasis>
</taxonomicName>
<emphasis id="B926EAEEFF94FFECFBA3FC07FB63FC1A" bold="true" box="[1066,1155,942,967]" pageId="3" pageNumber="492">
<taxonomicNameLabel id="A2155795FF94FFECFBA3FC07FB63FC1A" box="[1066,1155,942,967]" pageId="3" pageNumber="492" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but are absent in
<taxonomicName id="4C524D7FFF94FFECFAC6FC06FA73FC1B" box="[1359,1427,943,966]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="492" phylum="Arthropoda" rank="species" species="ieti">
<emphasis id="B926EAEEFF94FFECFAC6FC06FA73FC1B" box="[1359,1427,943,966]" italics="true" pageId="3" pageNumber="492">W. ieti</emphasis>
</taxonomicName>
, which possesses a pair of small spots on Abd. II. The slightly curved labral intrusion of the new species is rarely seen in entomobryids. The shape of the unguiculus is also a characteristic separating them, as it is lanceolate in new species and truncate in
<taxonomicName id="4C524D7FFF94FFECFE15FBB2FE02FBEF" box="[412,482,1051,1074]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="492" phylum="Arthropoda" rank="species" species="ieti">
<emphasis id="B926EAEEFF94FFECFE15FBB2FE02FBEF" box="[412,482,1051,1074]" italics="true" pageId="3" pageNumber="492">W. ieti</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF2D6674FF95FFEDFF1EF8DDFD12F83A" httpUri="https://zenodo.org/record/242884/files/figure.png" pageId="2" pageNumber="491" targetBox="[198,1369,198,1864]" targetPageId="2">
<paragraph id="8BED36FCFF95FFEDFF1EF8DDFD12F83A" blockId="2.[151,1436,1908,2023]" pageId="2" pageNumber="491">
<emphasis id="B926EAEEFF95FFEDFF1EF8DDFEA6F857" bold="true" box="[151,326,1908,1931]" pageId="2" pageNumber="491">FIGURES 112.</emphasis>
<taxonomicName id="4C524D7FFF95FFEDFEC5F8DFFE3EF856" box="[332,478,1910,1931]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="2" pageNumber="491" phylum="Arthropoda" rank="species" species="cassagnaui" status="sp. nov.">
<emphasis id="B926EAEEFF95FFEDFEC5F8DFFE3EF856" box="[332,478,1910,1931]" italics="true" pageId="2" pageNumber="491">W. cassagnaui</emphasis>
</taxonomicName>
<emphasis id="B926EAEEFF95FFEDFE6DF8DCFDD3F857" bold="true" box="[484,563,1909,1930]" pageId="2" pageNumber="491">
<taxonomicNameLabel id="A2155795FF95FFEDFE6DF8DCFDD3F857" box="[484,563,1909,1930]" pageId="2" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 1, habitus; 2, scales on terga; 3, scales on manubrium; 4, labrum; 5, lateral process of labial palp; 6, maxillary outer lobe; 7, labial chaetae; 8, dorsal cephalic chaetotaxy; 9, chaetotaxy of Th. IIAbd. III; 10, trochanteral organ; 11, hind claw; 12, chaetotaxy of Abd. IV. Larger circles, macrochaetae; smaller circles, microchaetae; circle with a slash, pseudopore; intersectional lines, bothriotricha.
</paragraph>
</caption>
<caption id="DF2D6674FF94FFECFF1EFCADFEE5FCE4" httpUri="https://zenodo.org/record/242885/files/figure.png" pageId="3" pageNumber="492" targetBox="[158,1427,204,728]" targetPageId="3">
<paragraph id="8BED36FCFF94FFECFF1EFCADFEE5FCE4" blockId="3.[151,1436,772,825]" pageId="3" pageNumber="492">
<emphasis id="B926EAEEFF94FFECFF1EFCADFEB4FCC7" bold="true" box="[151,340,772,795]" pageId="3" pageNumber="492">FIGURES 1315.</emphasis>
<taxonomicName id="4C524D7FFF94FFECFED2FCAFFE0DFCC6" box="[347,493,774,795]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="492" phylum="Arthropoda" rank="species" species="cassagnaui" status="sp. nov.">
<emphasis id="B926EAEEFF94FFECFED2FCAFFE0DFCC6" box="[347,493,774,795]" italics="true" pageId="3" pageNumber="492">W. cassagnaui</emphasis>
</taxonomicName>
<emphasis id="B926EAEEFF94FFECFE7DFCACFDA6FCC7" bold="true" box="[500,582,773,794]" pageId="3" pageNumber="492">
<taxonomicNameLabel id="A2155795FF94FFECFE7DFCACFDA6FCC7" box="[500,582,773,794]" pageId="3" pageNumber="492" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 13, anterior face of ventral tube; 14, posterior face and lateral flap of ventral tube; 15, mucro.
</paragraph>
</caption>
<caption id="DF2D6674FF94FFECFF1EFBF6FC8CFBAB" ID-Table-UUID="DF2D6674FF94FFECFF1EFBF6FC8CFBAB" box="[151,876,1118,1143]" httpUri="http://table.plazi.org/id/DF2D6674FF94FFECFF1EFBF6FC8CFBAB" pageId="3" pageNumber="492" targetBox="[151,1180,1170,1789]" targetIsTable="true" targetPageId="3">
<paragraph id="8BED36FCFF94FFECFF1EFBF6FC8CFBAB" blockId="3.[151,876,1118,1143]" box="[151,876,1118,1143]" pageId="3" pageNumber="492">
<emphasis id="B926EAEEFF94FFECFF1EFBF6FEEAFBAB" bold="true" box="[151,266,1119,1142]" pageId="3" pageNumber="492">TABLE 1.</emphasis>
Comparison between
<taxonomicName id="4C524D7FFF94FFECFE70FBC9FD72FBAA" box="[505,658,1120,1143]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="492" phylum="Arthropoda" rank="species" species="cassagnaui" status="sp. nov.">
<emphasis id="B926EAEEFF94FFECFE70FBC9FD72FBAA" box="[505,658,1120,1143]" italics="true" pageId="3" pageNumber="492">W. cassagnaui</emphasis>
</taxonomicName>
<emphasis id="B926EAEEFF94FFECFD10FBF6FD0EFBAB" bold="true" box="[665,750,1119,1142]" pageId="3" pageNumber="492">
<taxonomicNameLabel id="A2155795FF94FFECFD10FBF6FD0EFBAB" box="[665,750,1119,1142]" pageId="3" pageNumber="492" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C524D7FFF94FFECFCABFBC9FC83FBAA" box="[802,867,1120,1143]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="492" phylum="Arthropoda" rank="species" species="ieti">
<emphasis id="B926EAEEFF94FFECFCABFBC9FC83FBAA" box="[802,867,1120,1143]" italics="true" pageId="3" pageNumber="492">W. ieti</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BED36FCFF94FFECFF16FB3BFEF4F920" pageId="3" pageNumber="492">
<table id="F952C45CFF940010FF1EFB3BFB7CF920" box="[151,1180,1170,1789]" gridcols="3" gridrows="11" pageId="3" pageNumber="492">
<tr id="356234BEFF940010FF1EFB3BFB7CFB75" box="[151,1180,1170,1192]" gridrow="0" pageId="3" pageNumber="492">
<th id="76B35DC2FF940010FF1EFB3BFDE0FB75" box="[151,512,1170,1192]" gridcol="0" gridrow="0" pageId="3" pageNumber="492">Characters</th>
<th id="76B35DC2FF940010FD0AFB3BFC8CFB75" box="[643,876,1170,1192]" gridcol="1" gridrow="0" pageId="3" pageNumber="492">
<taxonomicName id="4C524D7FFF94FFECFD0DFB3AFCF5FB75" box="[644,789,1171,1192]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="492" phylum="Arthropoda" rank="species" species="cassagnaui" status="sp. nov.">
<emphasis id="B926EAEEFF94FFECFD0DFB3AFCF5FB75" box="[644,789,1171,1192]" italics="true" pageId="3" pageNumber="492">W. cassagnaui</emphasis>
</taxonomicName>
<emphasis id="B926EAEEFF94FFECFC92FB3BFC8CFB7A" bold="true" box="[795,876,1170,1191]" pageId="3" pageNumber="492">
<taxonomicNameLabel id="A2155795FF94FFECFC92FB3BFC8CFB7A" box="[795,876,1170,1191]" pageId="3" pageNumber="492" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76B35DC2FF940010FC7DFB3BFB7CFB75" box="[1012,1180,1170,1192]" gridcol="2" gridrow="0" pageId="3" pageNumber="492">
<taxonomicName id="4C524D7FFF94FFECFC7DFB3AFBD4FB75" box="[1012,1076,1171,1192]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="492" phylum="Arthropoda" rank="species" species="ieti">
<emphasis id="B926EAEEFF94FFECFC7DFB3AFBD4FB75" box="[1012,1076,1171,1192]" italics="true" pageId="3" pageNumber="492">W. ieti</emphasis>
</taxonomicName>
.
</th>
</tr>
<tr id="356234BEFF940010FF1EFB15FB7CFB0F" box="[151,1180,1212,1234]" gridrow="1" pageId="3" pageNumber="492">
<th id="76B35DC2FF940010FF1EFB15FDE0FB0F" box="[151,512,1212,1234]" gridcol="0" gridrow="1" pageId="3" pageNumber="492">Labral papillae</th>
<td id="76B35DC2FF940010FD0AFB15FC8CFB0F" box="[643,876,1212,1234]" gridcol="1" gridrow="1" pageId="3" pageNumber="492">rounded</td>
<td id="76B35DC2FF940010FC7DFB15FB7CFB0F" box="[1012,1180,1212,1234]" gridcol="2" gridrow="1" pageId="3" pageNumber="492">conical</td>
</tr>
<tr id="356234BEFF940010FF1EFB4CFB7CFB26" box="[151,1180,1253,1275]" gridrow="2" pageId="3" pageNumber="492">
<th id="76B35DC2FF940010FF1EFB4CFDE0FB26" box="[151,512,1253,1275]" gridcol="0" gridrow="2" pageId="3" pageNumber="492">Labral intrusion</th>
<td id="76B35DC2FF940010FD0AFB4CFC8CFB26" box="[643,876,1253,1275]" gridcol="1" gridrow="2" pageId="3" pageNumber="492">slightly curved</td>
<td id="76B35DC2FF940010FC7DFB4CFB7CFB26" box="[1012,1180,1253,1275]" gridcol="2" gridrow="2" pageId="3" pageNumber="492">U-shaped</td>
</tr>
<tr id="356234BEFF940010FF1EFAA7FB7CFAF9" box="[151,1180,1294,1316]" gridrow="3" pageId="3" pageNumber="492">
<th id="76B35DC2FF940010FF1EFAA7FDE0FAF9" box="[151,512,1294,1316]" gridcol="0" gridrow="3" pageId="3" pageNumber="492">Ratio of R/M on labium</th>
<td id="76B35DC2FF940010FD0AFAA7FC8CFAF9" box="[643,876,1294,1316]" gridcol="1" gridrow="3" pageId="3" pageNumber="492">0.40.5</td>
<td id="76B35DC2FF940010FC7DFAA7FB7CFAF9" box="[1012,1180,1294,1316]" gridcol="2" gridrow="3" pageId="3" pageNumber="492">0.7</td>
</tr>
<tr id="356234BEFF940010FF1EFA9EFB7CFA90" box="[151,1180,1335,1357]" gridrow="4" pageId="3" pageNumber="492">
<th id="76B35DC2FF940010FF1EFA9EFDE0FA90" box="[151,512,1335,1357]" gridcol="0" gridrow="4" pageId="3" pageNumber="492">Lateral process of labial palp</th>
<td id="76B35DC2FF940010FD0AFA9EFC8CFA90" box="[643,876,1335,1357]" gridcol="1" gridrow="4" pageId="3" pageNumber="492">long</td>
<td id="76B35DC2FF940010FC7DFA9EFB7CFA90" box="[1012,1180,1335,1357]" gridcol="2" gridrow="4" pageId="3" pageNumber="492">short</td>
</tr>
<tr id="356234BEFF940010FF1EFAC8FB7CFAAA" box="[151,1180,1377,1399]" gridrow="5" pageId="3" pageNumber="492">
<th id="76B35DC2FF940010FF1EFAC8FDE0FAAA" box="[151,512,1377,1399]" gridcol="0" gridrow="5" pageId="3" pageNumber="492">Chaetae on metatrochanteral organ</th>
<td id="76B35DC2FF940010FD0AFAC8FC8CFAAA" box="[643,876,1377,1399]" gridcol="1" gridrow="5" pageId="3" pageNumber="492">about 14</td>
<td id="76B35DC2FF940010FC7DFAC8FB7CFAAA" box="[1012,1180,1377,1399]" gridcol="2" gridrow="5" pageId="3" pageNumber="492">&gt;20</td>
</tr>
<tr id="356234BEFF940010FF1EFA23FB7CFA7D" box="[151,1180,1418,1440]" gridrow="6" pageId="3" pageNumber="492">
<th id="76B35DC2FF940010FF1EFA23FDE0FA7D" box="[151,512,1418,1440]" gridcol="0" gridrow="6" pageId="3" pageNumber="492">Scales on Ant. I</th>
<td id="76B35DC2FF940010FD0AFA23FC8CFA7D" box="[643,876,1418,1440]" gridcol="1" gridrow="6" pageId="3" pageNumber="492">absent</td>
<td id="76B35DC2FF940010FC7DFA23FB7CFA7D" box="[1012,1180,1418,1440]" gridcol="2" gridrow="6" pageId="3" pageNumber="492">present</td>
</tr>
<tr id="356234BEFF940010FF1EFA1DFB7CFA17" box="[151,1180,1460,1482]" gridrow="7" pageId="3" pageNumber="492">
<th id="76B35DC2FF940010FF1EFA1DFDE0FA17" box="[151,512,1460,1482]" gridcol="0" gridrow="7" pageId="3" pageNumber="492">Unguiculus</th>
<td id="76B35DC2FF940010FD0AFA1DFC8CFA17" box="[643,876,1460,1482]" gridcol="1" gridrow="7" pageId="3" pageNumber="492">lanceolate</td>
<td id="76B35DC2FF940010FC7DFA1DFB7CFA17" box="[1012,1180,1460,1482]" gridcol="2" gridrow="7" pageId="3" pageNumber="492">truncate</td>
</tr>
<tr id="356234BEFF940010FF1EFA74FB7CFA2E" box="[151,1180,1501,1523]" gridrow="8" pageId="3" pageNumber="492">
<th id="76B35DC2FF940010FF1EFA74FDE0FA2E" box="[151,512,1501,1523]" gridcol="0" gridrow="8" pageId="3" pageNumber="492">Anterior face of ventral tube</th>
<td id="76B35DC2FF940010FD0AFA74FC8CFA2E" box="[643,876,1501,1523]" gridcol="1" gridrow="8" pageId="3" pageNumber="492">about 14</td>
<td id="76B35DC2FF940010FC7DFA74FB7CFA2E" box="[1012,1180,1501,1523]" gridcol="2" gridrow="8" pageId="3" pageNumber="492">about 22</td>
</tr>
<tr id="356234BEFF940010FF1EF9AFFB7CF987" box="[151,1180,1542,1626]" gridrow="9" pageId="3" pageNumber="492">
<th id="76B35DC2FF940010FF1EF9AFFDE0F987" box="[151,512,1542,1626]" gridcol="0" gridrow="9" pageId="3" pageNumber="492">Tergal chaetotaxy Abd. I Abd. IV</th>
<td id="76B35DC2FF940010FD0AF9AFFC8CF987" box="[643,876,1542,1626]" gridcol="1" gridrow="9" pageId="3" pageNumber="492">3+3 mac 7+7 central</td>
<td id="76B35DC2FF940010FC7DF9AFFB7CF987" box="[1012,1180,1542,1626]" gridcol="2" gridrow="9" pageId="3" pageNumber="492">2+2 mac 78+78 central</td>
</tr>
<tr id="356234BEFF940010FF1EF96BFB7CF920" box="[151,1180,1730,1789]" gridrow="10" pageId="3" pageNumber="492" rowspan-1="1" rowspan-2="1">
<th id="76B35DC2FF940010FF1EF96BFDE0F920" box="[151,512,1730,1789]" gridcol="0" gridrow="10" pageId="3" pageNumber="492">
<taxonomicName id="4C524D7FFF94FFECFF1EF96BFE27F901" authority="Yosii, 1971" authorityName="Yosii" authorityYear="1971" box="[151,455,1730,1756]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="492" phylum="Arthropoda" rank="species" species="ieti">
<emphasis id="B926EAEEFF94FFECFF1EF96BFE27F901" bold="true" box="[151,455,1730,1756]" pageId="3" pageNumber="492">
<emphasis id="B926EAEEFF94FFECFF1EF96BFEDAF901" bold="true" box="[151,314,1730,1756]" italics="true" pageId="3" pageNumber="492">Willowsia ieti</emphasis>
Yosii, 1971
</emphasis>
</taxonomicName>
Figs 1625
</th>
</tr>
</table>
</paragraph>
<subSubSection id="C3486577FF94FFEAFF1EF885FC91F8F1" lastPageId="5" lastPageNumber="494" pageId="3" pageNumber="492" type="materials_examined">
<paragraph id="8BED36FCFF94FFECFF1EF885FC33F898" blockId="3.[151,1436,1836,2006]" box="[151,979,1836,1861]" pageId="3" pageNumber="492">
<emphasis id="B926EAEEFF94FFECFF1EF885FED5F898" bold="true" box="[151,309,1836,1861]" pageId="3" pageNumber="492">
<typeStatus id="54E9885EFF94FFECFF1EF885FF32F898" box="[151,210,1836,1861]" pageId="3" pageNumber="492">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F345766CFF94FFECFEB5F885FE64F898" box="[316,388,1836,1861]" name="Nepal" pageId="3" pageNumber="492">Nepal</collectingCountry>
: Khumbu Himal, Yaral near Pangpoche, alt.
<quantity id="4CAA9B19FF94FFECFCFEF884FC2FF898" box="[887,975,1837,1861]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="492" unit="m" value="4100.0">4100 m</quantity>
.
</paragraph>
<paragraph id="8BED36FCFF94FFECFF4EF8F8FACBF808" blockId="3.[151,1436,1836,2006]" pageId="3" pageNumber="492">
<emphasis id="B926EAEEFF94FFECFF4EF8F8FE26F8B7" bold="true" box="[199,454,1873,1898]" pageId="3" pageNumber="492">Specimens examined.</emphasis>
<specimenCount id="9D54FD75FF94FFECFE45F8F8FDD7F8B7" box="[460,567,1873,1898]" pageId="3" pageNumber="492" type="female">4 females</specimenCount>
on slides,
<collectingCountry id="F345766CFF94FFECFD27F8F8FD15F8B7" box="[686,757,1873,1898]" name="Nepal" pageId="3" pageNumber="492">Nepal</collectingCountry>
: Machapuchare, alt.
<quantity id="4CAA9B19FF94FFECFC52F8F8FBD1F8B4" box="[987,1073,1873,1897]" metricMagnitude="3" metricUnit="m" metricValue="2.7399999999999998" pageId="3" pageNumber="492" unit="m" value="2740.0">2740 m</quantity>
, level site with large trees (oaks, rhododendrons, bamboos), moss and lichen on a branch,
<date id="FFEC103CFF94FFECFC9DF8DCFC67F853" box="[788,903,1909,1934]" pageId="3" pageNumber="492" value="1976-10-25">25.x.1976</date>
, P. Cassagnau leg. (# Nep. 76.40);
<specimenCount id="9D54FD75FF94FFECFA84F8DCFA99F853" box="[1293,1401,1909,1934]" pageId="3" pageNumber="492" type="female">8 females</specimenCount>
on slides,
<collectingCountry id="F345766CFF94FFECFF6DF830FECBF86F" box="[228,299,1945,1970]" name="Nepal" pageId="3" pageNumber="492">Nepal</collectingCountry>
: Gosaikund, alt.
<quantity id="4CAA9B19FF94FFECFE6FF830FDDCF86C" box="[486,572,1945,1970]" metricMagnitude="3" metricUnit="m" metricValue="3.66" pageId="3" pageNumber="492" unit="m" value="3660.0">3660 m</quantity>
, upper edge of the fir forest and juniper heath, moss and liverworts on a rock, moderately humid,
<date id="FFEC103CFF94FFECFEF8F814FE09F80B" box="[369,489,1981,2006]" pageId="3" pageNumber="492" value="1978-11-20">20.xi.1978</date>
(# Nep. 78.59). Eight specimens on slides in
<collectionCode id="ED43AE39FF94FFECFC6EF814FBCFF808" box="[999,1071,1981,2005]" pageId="3" pageNumber="492">NJAU</collectionCode>
and others in
<collectionCode id="ED43AE39FF94FFECFB47F817FAC7F808" LSID="urn:lsid:biocol.org:col:34988" box="[1230,1319,1982,2005]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="492">MNHN</collectionCode>
.
</paragraph>
<caption id="DF2D6674FF93FFEBFF1EF836FE74F82E" httpUri="https://zenodo.org/record/242886/files/figure.png" pageId="4" pageNumber="493" targetBox="[181,1418,194,1919]" targetPageId="4">
<paragraph id="8BED36FCFF93FFEBFF1EF836FE74F82E" blockId="4.[151,1436,1951,2035]" pageId="4" pageNumber="493">
<emphasis id="B926EAEEFF93FFEBFF1EF836FEB5F868" bold="true" box="[151,341,1951,1974]" pageId="4" pageNumber="493">FIGURES 1624.</emphasis>
<taxonomicName id="4C524D7FFF93FFEBFED2F808FE7BF86B" box="[347,411,1953,1974]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="ieti">
<emphasis id="B926EAEEFF93FFEBFED2F808FE7BF86B" box="[347,411,1953,1974]" italics="true" pageId="4" pageNumber="493">W. ieti</emphasis>
</taxonomicName>
: 16, scales on terga; 17, scales on manubrium; 18, lateral process of labial palp; 19, maxillary outer lobe; 20, dorsal cephalic chaetotaxy; 21, trochanteral organ; 22, anterior face of ventral tube; 23, thoracic chaetotaxy; 24, chaetotaxy of Abd. IIII.
</paragraph>
</caption>
<caption id="DF2D6674FF92FFEAFF1EFAEDFDB9FA87" box="[151,601,1348,1370]" pageId="5" pageNumber="494">
<paragraph id="8BED36FCFF92FFEAFF1EFAEDFDB9FA87" blockId="5.[151,601,1348,1370]" box="[151,601,1348,1370]" pageId="5" pageNumber="494">
<emphasis id="B926EAEEFF92FFEAFF1EFAEDFEC0FA84" bold="true" box="[151,288,1348,1369]" pageId="5" pageNumber="494">FIGURE 25.</emphasis>
<taxonomicName id="4C524D7FFF92FFEAFEACFAECFE85FA87" box="[293,357,1349,1370]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="5" pageNumber="494" phylum="Arthropoda" rank="species" species="ieti">
<emphasis id="B926EAEEFF92FFEAFEACFAECFE85FA87" box="[293,357,1349,1370]" italics="true" pageId="5" pageNumber="494">W. ieti</emphasis>
</taxonomicName>
: chaetotaxy of Abd. IV.
</paragraph>
</caption>
<paragraph id="8BED36FCFF92FFEAFF4EFA2EFC91F8F1" blockId="5.[151,1437,1415,2016]" pageId="5" pageNumber="494">
<emphasis id="B926EAEEFF92FFEAFF4EFA2EFDB7FA7D" bold="true" box="[199,599,1415,1440]" pageId="5" pageNumber="494">Additions to original description.</emphasis>
Body scales heavily striate with pointed tips (
<figureCitation id="13692A79FF92FFEAFBE1FA2EFB5EFA7D" box="[1128,1214,1415,1440]" captionStart="FIGURES 16 24. W" captionStartId="4.[151,264,1951,1974]" captionTargetBox="[181,1418,194,1919]" captionTargetId="figure@4.[163,1424,193,1931]" captionTargetPageId="4" captionText="FIGURES 16 24. W. ieti: 16, scales on terga; 17, scales on manubrium; 18, lateral process of labial palp; 19, maxillary outer lobe; 20, dorsal cephalic chaetotaxy; 21, trochanteral organ; 22, anterior face of ventral tube; 23, thoracic chaetotaxy; 24, chaetotaxy of Abd. I III." httpUri="https://zenodo.org/record/242886/files/figure.png" pageId="5" pageNumber="494">Fig. 16</figureCitation>
); present on Ant. I, body and ventral side of manubrium; scales on manubrium narrower than those on body, some of them extremely elongate (
<figureCitation id="13692A79FF92FFEAFE81FA66FEBFFA3A" box="[264,351,1487,1512]" captionStart="FIGURES 16 24. W" captionStartId="4.[151,264,1951,1974]" captionTargetBox="[181,1418,194,1919]" captionTargetId="figure@4.[163,1424,193,1931]" captionTargetPageId="4" captionText="FIGURES 16 24. W. ieti: 16, scales on terga; 17, scales on manubrium; 18, lateral process of labial palp; 19, maxillary outer lobe; 20, dorsal cephalic chaetotaxy; 21, trochanteral organ; 22, anterior face of ventral tube; 23, thoracic chaetotaxy; 24, chaetotaxy of Abd. I III." httpUri="https://zenodo.org/record/242886/files/figure.png" pageId="5" pageNumber="494">Fig. 17</figureCitation>
). Tip of lateral process of labial palp not reaching apex of labial papilla (
<figureCitation id="13692A79FF92FFEAFB36FA66FAF6FA3A" box="[1215,1302,1487,1512]" captionStart="FIGURES 16 24. W" captionStartId="4.[151,264,1951,1974]" captionTargetBox="[181,1418,194,1919]" captionTargetId="figure@4.[163,1424,193,1931]" captionTargetPageId="4" captionText="FIGURES 16 24. W. ieti: 16, scales on terga; 17, scales on manubrium; 18, lateral process of labial palp; 19, maxillary outer lobe; 20, dorsal cephalic chaetotaxy; 21, trochanteral organ; 22, anterior face of ventral tube; 23, thoracic chaetotaxy; 24, chaetotaxy of Abd. I III." httpUri="https://zenodo.org/record/242886/files/figure.png" pageId="5" pageNumber="494">Fig. 18</figureCitation>
). Subapical chaeta of maxillary outer lobe stronger than apical one; 3 hairs on sublobal plate (
<figureCitation id="13692A79FF92FFEAFBA8FA5AFB94F9D6" box="[1057,1140,1523,1548]" captionStart="FIGURES 16 24. W" captionStartId="4.[151,264,1951,1974]" captionTargetBox="[181,1418,194,1919]" captionTargetId="figure@4.[163,1424,193,1931]" captionTargetPageId="4" captionText="FIGURES 16 24. W. ieti: 16, scales on terga; 17, scales on manubrium; 18, lateral process of labial palp; 19, maxillary outer lobe; 20, dorsal cephalic chaetotaxy; 21, trochanteral organ; 22, anterior face of ventral tube; 23, thoracic chaetotaxy; 24, chaetotaxy of Abd. I III." httpUri="https://zenodo.org/record/242886/files/figure.png" pageId="5" pageNumber="494">Fig. 19</figureCitation>
). Labial chaeta R 0.7 x the length of M. Dorsal cephalic chaetotaxy as in
<figureCitation id="13692A79FF92FFEAFD39F9BEFCE5F9F2" box="[688,773,1559,1584]" captionStart="FIGURES 16 24. W" captionStartId="4.[151,264,1951,1974]" captionTargetBox="[181,1418,194,1919]" captionTargetId="figure@4.[163,1424,193,1931]" captionTargetPageId="4" captionText="FIGURES 16 24. W. ieti: 16, scales on terga; 17, scales on manubrium; 18, lateral process of labial palp; 19, maxillary outer lobe; 20, dorsal cephalic chaetotaxy; 21, trochanteral organ; 22, anterior face of ventral tube; 23, thoracic chaetotaxy; 24, chaetotaxy of Abd. I III." httpUri="https://zenodo.org/record/242886/files/figure.png" pageId="5" pageNumber="494">Fig. 20</figureCitation>
. Metatrochanteral organ with more than 20 smooth spiny chaetae (
<figureCitation id="13692A79FF92FFEAFF75F992FEB2F98E" box="[252,338,1595,1620]" captionStart="FIGURES 16 24. W" captionStartId="4.[151,264,1951,1974]" captionTargetBox="[181,1418,194,1919]" captionTargetId="figure@4.[163,1424,193,1931]" captionTargetPageId="4" captionText="FIGURES 16 24. W. ieti: 16, scales on terga; 17, scales on manubrium; 18, lateral process of labial palp; 19, maxillary outer lobe; 20, dorsal cephalic chaetotaxy; 21, trochanteral organ; 22, anterior face of ventral tube; 23, thoracic chaetotaxy; 24, chaetotaxy of Abd. I III." httpUri="https://zenodo.org/record/242886/files/figure.png" pageId="5" pageNumber="494">Fig. 21</figureCitation>
). Ventral tube anteriorly with about 22 ciliate chaetae on each side, 2 of them much larger than others (
<figureCitation id="13692A79FF92FFEAFF63F9F6FEDFF9AA" box="[234,319,1631,1656]" captionStart="FIGURES 16 24. W" captionStartId="4.[151,264,1951,1974]" captionTargetBox="[181,1418,194,1919]" captionTargetId="figure@4.[163,1424,193,1931]" captionTargetPageId="4" captionText="FIGURES 16 24. W. ieti: 16, scales on terga; 17, scales on manubrium; 18, lateral process of labial palp; 19, maxillary outer lobe; 20, dorsal cephalic chaetotaxy; 21, trochanteral organ; 22, anterior face of ventral tube; 23, thoracic chaetotaxy; 24, chaetotaxy of Abd. I III." httpUri="https://zenodo.org/record/242886/files/figure.png" pageId="5" pageNumber="494">Fig. 22</figureCitation>
). Manubrial plaque with about 6 ciliate chaetae; pseudopores not clearly seen. Distal smooth part of dens about twice mucro in length. Th. II with 10 posterior mac; median mac absent. Th. III with 11 mac (
<figureCitation id="13692A79FF92FFEAFAB2F92AFA6FF946" box="[1339,1423,1667,1692]" captionStart="FIGURES 16 24. W" captionStartId="4.[151,264,1951,1974]" captionTargetBox="[181,1418,194,1919]" captionTargetId="figure@4.[163,1424,193,1931]" captionTargetPageId="4" captionText="FIGURES 16 24. W. ieti: 16, scales on terga; 17, scales on manubrium; 18, lateral process of labial palp; 19, maxillary outer lobe; 20, dorsal cephalic chaetotaxy; 21, trochanteral organ; 22, anterior face of ventral tube; 23, thoracic chaetotaxy; 24, chaetotaxy of Abd. I III." httpUri="https://zenodo.org/record/242886/files/figure.png" pageId="5" pageNumber="494">Fig. 23</figureCitation>
). Abd. I with 2 (m34) mac. Abd. II with 3 (a2, m3, m3e) central and 1 (m5) lateral mac. Abd. III with 2 central (a2, m3) and 3 (am6, pm6, p6) lateral mac (
<figureCitation id="13692A79FF92FFEAFDD0F962FD46F939" box="[601,678,1739,1764]" captionStart="FIGURES 16 24. W" captionStartId="4.[151,264,1951,1974]" captionTargetBox="[181,1418,194,1919]" captionTargetId="figure@4.[163,1424,193,1931]" captionTargetPageId="4" captionText="FIGURES 16 24. W. ieti: 16, scales on terga; 17, scales on manubrium; 18, lateral process of labial palp; 19, maxillary outer lobe; 20, dorsal cephalic chaetotaxy; 21, trochanteral organ; 22, anterior face of ventral tube; 23, thoracic chaetotaxy; 24, chaetotaxy of Abd. I III." httpUri="https://zenodo.org/record/242886/files/figure.png" pageId="5" pageNumber="494">Fig.24</figureCitation>
). Abd. IV with 78 central and about 14 lateral mac; homology of anterior four central mac unclear, the second mac close to midline often as a mesochaeta (
<figureCitation id="13692A79FF92FFEAFB04F946FB01F8D5" box="[1165,1249,1775,1800]" captionStart="FIGURE 25. W" captionStartId="5.[151,250,1348,1369]" captionText="FIGURE 25. W. ieti: chaetotaxy of Abd. IV." pageId="5" pageNumber="494">Fig. 25</figureCitation>
). Accessory mic of bothriotrichal complexes on Abd. II IV unmodified or absent.
</paragraph>
</subSubSection>
<subSubSection id="C3486577FF92FFE9FF4EF89EFC60FF08" lastPageId="6" lastPageNumber="495" pageId="5" pageNumber="494" type="discussion">
<paragraph id="8BED36FCFF92FFE9FF4EF89EFC60FF08" blockId="5.[151,1437,1415,2016]" lastBlockId="6.[151,1436,151,213]" lastPageId="6" lastPageNumber="495" pageId="5" pageNumber="494">
<emphasis id="B926EAEEFF92FFEAFF4EF89EFEDBF88D" bold="true" box="[199,315,1847,1872]" pageId="5" pageNumber="494">Remarks.</emphasis>
The
<typeStatus id="54E9885EFF92FFEAFEF1F891FE48F88D" box="[376,424,1848,1872]" pageId="5" pageNumber="494">type</typeStatus>
specimens in Natural History Museum, Geneva, could not be studied as permission was not given to borrow it. The
<typeStatus id="54E9885EFF92FFEAFE43F8F5FE1AF8A9" box="[458,506,1884,1908]" pageId="5" pageNumber="494">type</typeStatus>
locality (Yaral) is located between the two new sites for this species recorded here. The Gosaikund record is
<quantity id="4CAA9B19FF92FFEAFE3FF8D6FDECF845" box="[438,524,1919,1944]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="494" unit="km" value="100.0">100 km</quantity>
directly east of the
<typeStatus id="54E9885EFF92FFEAFD66F829FCFFF845" box="[751,799,1920,1944]" pageId="5" pageNumber="494">type</typeStatus>
locality, Machapuchare about
<quantity id="4CAA9B19FF92FFEAFBF2F8D6FB24F845" box="[1147,1220,1919,1944]" metricMagnitude="4" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="494" unit="km" value="75.0">75 km</quantity>
west of Yaral. Our specimens from these two new populations comply well with Yosiis (1971) original description, particularly in colour and tergal chaetotaxy (Th. IIAbd. III), so that they are designated as
<taxonomicName id="4C524D7FFF92FFEAFB8CF861FB6FF83D" authority="Yosii" authorityName="Yosii" box="[1029,1167,1991,2016]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="5" pageNumber="494" phylum="Arthropoda" rank="species" species="ieti">
<emphasis id="B926EAEEFF92FFEAFB8CF861FBADF802" box="[1029,1101,1992,2015]" italics="true" pageId="5" pageNumber="494">W. ieti</emphasis>
Yosii
</taxonomicName>
here. We added further details of mouthpart, chaetae on dorsal and ventral head, ventral tube and total body chaetotaxy. For differences between
<taxonomicName id="4C524D7FFF91FFE9FF72FF14FEA1FF09" box="[251,321,189,212]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="ieti">
<emphasis id="B926EAEEFF91FFE9FF72FF14FEA1FF09" box="[251,321,189,212]" italics="true" pageId="6" pageNumber="495">W. ieti</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D7FFF91FFE9FEF1FF14FDF9FF09" box="[376,537,189,212]" class="Entognatha" family="Entomobryidae" genus="Willowsia" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="cassagnaui" status="sp. nov.">
<emphasis id="B926EAEEFF91FFE9FEF1FF14FDF9FF09" box="[376,537,189,212]" italics="true" pageId="6" pageNumber="495">W. cassagnaui</emphasis>
</taxonomicName>
<emphasis id="B926EAEEFF91FFE9FDA9FF15FD99FF08" bold="true" box="[544,633,188,213]" pageId="6" pageNumber="495">
<taxonomicNameLabel id="A2155795FF91FFE9FDA9FF15FD99FF08" box="[544,633,188,213]" pageId="6" pageNumber="495" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
see the remarks above.
</paragraph>
</subSubSection>
</treatment>
</document>