treatments-xml/data/03/FB/87/03FB87BDFF87C30BFF75F45FFCABBD17.xml

<document id="A47E5F0D5B022F9DFCECFD7C40319CC7" ID-CLB-Dataset="32647" ID-DOI="10.11646/zootaxa.4294.2.9" ID-GBIF-Dataset="5b03347b-4db9-473a-84c4-d653cfea6b6c" ID-ISSN="1175-5326" ID-Zenodo-Dep="831888" ID-ZooBank="7EAE9AD8-F840-40DB-843C-D1DD80568E76" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500425405979" checkinUser="plazi" docAuthor="Nadein, Konstantin S. &amp; Leschen, Richard A. B." docDate="2017" docId="03FB87BDFF87C30BFF75F45FFCABBD17" docLanguage="en" docName="zootaxa.4294.2.9.pdf" docOrigin="Zootaxa 4294 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tyrannomolpus Nadein &amp; Leschen 2017, gen. nov." docType="treatment" docUuid="F2AD6850-5644-4AAE-8ABC-86522E2266D6" docUuidSource="ZooBank" docVersion="8" lastPageNumber="278" masterDocId="FFC2FFC5FF86C30CFFE2F577FFF2BF2A" masterDocTitle="A new genus of leaf beetle (Coleoptera: Chrysomelidae: Eumolpinae) from Three Kings Islands, New Zealand" masterLastPageNumber="280" masterPageNumber="271" pageNumber="272" updateTime="1698465781638" updateUser="ExternalLinkService">
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<mods:title id="F9B0EE71EB6B1F0F7266D3804AB38148">A new genus of leaf beetle (Coleoptera: Chrysomelidae: Eumolpinae) from Three Kings Islands, New Zealand</mods:title>
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<mods:namePart id="5C35CD8F29D1863002FAB362E91C1661">Nadein, Konstantin S.</mods:namePart>
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<mods:namePart id="886A9DDF5884908F784B3EF573B14A80">Leschen, Richard A. B.</mods:namePart>
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<treatment id="03FB87BDFF87C30BFF75F45FFCABBD17" ID-DOI="http://doi.org/10.5281/zenodo.6032326" ID-GBIF-Taxon="132550483" ID-Zenodo-Dep="6032326" LSID="urn:lsid:zoobank.org:act:F2AD6850-5644-4AAE-8ABC-86522E2266D6" httpUri="http://treatment.plazi.org/id/03FB87BDFF87C30BFF75F45FFCABBD17" lastPageId="7" lastPageNumber="278" pageId="1" pageNumber="272">
<subSubSection id="C3486520FF87C30DFF75F45FFEFFBE49" pageId="1" pageNumber="272" type="nomenclature">
<paragraph id="8BED36ABFF87C30DFF75F45FFE39BE68" blockId="1.[151,459,296,355]" box="[151,459,296,322]" pageId="1" pageNumber="272">
<heading id="D0A581C7FF87C30DFF75F45FFE39BE68" bold="true" box="[151,459,296,322]" fontSize="11" level="1" pageId="1" pageNumber="272" reason="1">
<emphasis id="B926EAB9FF87C30DFF75F45FFE39BE68" bold="true" box="[151,459,296,322]" pageId="1" pageNumber="272">
<taxonomicName id="4C524D28FF87C30DFF75F45FFEA4BE68" authority="Nadein &amp; Leschen, 2017" authorityName="Nadein &amp; Leschen" authorityYear="2017" box="[151,342,296,322]" class="Insecta" family="Chrysomelidae" genus="Tyrannomolpus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B926EAB9FF87C30DFF75F45FFEA4BE68" bold="true" box="[151,342,296,322]" italics="true" pageId="1" pageNumber="272">Tyrannomolpus</emphasis>
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<taxonomicNameLabel id="A21557C2FF87C30DFEBFF45FFE39BE68" box="[349,459,296,322]" pageId="1" pageNumber="272" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BED36ABFF87C30DFF75F43DFEFFBE49" blockId="1.[151,459,296,355]" box="[151,269,330,355]" pageId="1" pageNumber="272">
<figureCitation id="13692A2EFF87C30DFF75F43DFEFBBE49" box="[151,265,330,355]" captionStart-0="FIGURES 1, 2" captionStart-1="FIGURES 3 – 12" captionStart-2="FIGURES 13 – 25" captionStart-3="FIGURES 26 – 31" captionStartId-0="1.[151,264,1938,1961]" captionStartId-1="2.[151,264,1926,1949]" captionStartId-2="3.[151,264,1702,1725]" captionStartId-3="5.[151,264,1468,1491]" captionTargetBox-0="[252,1353,1059,1872]" captionTargetBox-1="[204,1404,193,1905]" captionTargetBox-2="[151,1359,194,1682]" captionTargetBox-3="[151,1431,381,1444]" captionTargetId-0="figure@1.[211,1376,1036,1891]" captionTargetId-1="figure@2.[178,1409,193,1906]" captionTargetId-2="figure@3.[151,1436,193,1682]" captionTargetId-3="figure@5.[151,1436,373,1448]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="5" captionText-0="FIGURES 1, 2. Tyrannomolpus rex, gen. nov., spec. nov. general view; 1: dorsal; 2: ventral." captionText-1="FIGURES 3 – 12. Tyrannomolpus rex, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view." captionText-2="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." captionText-3="FIGURES 26 – 31. Tyrannomolpus rex, morphological details; 26: ovipositor; 27: gonocoxites; 28: spermatheca; 29: aedeagus ventral; 30: aedeagus lateral; 31: main body of penis ventral." httpUri-0="https://zenodo.org/record/831890/files/figure.png" httpUri-1="https://zenodo.org/record/831892/files/figure.png" httpUri-2="https://zenodo.org/record/831894/files/figure.png" httpUri-3="https://zenodo.org/record/831896/files/figure.png" pageId="1" pageNumber="272">Figs 1–31</figureCitation>
.
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<paragraph id="8BED36ABFF87C30DFF75F4E4FDA4BE82" blockId="1.[151,598,402,424]" box="[151,598,402,424]" pageId="1" pageNumber="272">
Type species: 
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<emphasis id="B926EAB9FF87C30DFEC4F4E5FE15BE82" box="[294,487,402,424]" italics="true" pageId="1" pageNumber="272">Tyrannomolpus rex</emphasis>
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<emphasis id="B926EAB9FF87C30DFE0DF4E4FDA4BE82" bold="true" box="[495,598,403,424]" pageId="1" pageNumber="272">spec. nov.</emphasis>
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<subSubSection id="C3486520FF87C30DFF75F4A2FE5CBC50" pageId="1" pageNumber="272" type="diagnosis">
<paragraph id="8BED36ABFF87C30DFF75F4A2FE5CBC50" blockId="1.[151,1437,469,962]" pageId="1" pageNumber="272">
<emphasis id="B926EAB9FF87C30DFF75F4A2FEE3BEC4" bold="true" box="[151,273,469,494]" pageId="1" pageNumber="272">Diagnosis.</emphasis>
Colour dark bronzy-greenish with metallic luster (
<figureCitation id="13692A2EFF87C30DFCAAF4A2FC5FBEC4" box="[840,941,469,494]" captionStart="FIGURES 1, 2" captionStartId="1.[151,264,1938,1961]" captionTargetBox="[252,1353,1059,1872]" captionTargetId="figure@1.[211,1376,1036,1891]" captionTargetPageId="1" captionText="FIGURES 1, 2. Tyrannomolpus rex, gen. nov., spec. nov. general view; 1: dorsal; 2: ventral." httpUri="https://zenodo.org/record/831890/files/figure.png" pageId="1" pageNumber="272">Figs 1, 2</figureCitation>
). Vertex of head and dorsum setose (
<figureCitation id="13692A2EFF87C30DFAA8F4A2FA7CBEC4" box="[1354,1422,469,494]" captionStart="FIGURES 3 – 12" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[204,1404,193,1905]" captionTargetId="figure@2.[178,1409,193,1906]" captionTargetPageId="2" captionText="FIGURES 3 – 12. Tyrannomolpus rex, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view." httpUri="https://zenodo.org/record/831892/files/figure.png" pageId="1" pageNumber="272">Fig. 3</figureCitation>
). Head with midcranial suture (
<figureCitation id="13692A2EFF87C30DFE12F48EFDC5BD3B" box="[496,567,505,530]" captionStart="FIGURES 3 – 12" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[204,1404,193,1905]" captionTargetId="figure@2.[178,1409,193,1906]" captionTargetPageId="2" captionText="FIGURES 3 – 12. Tyrannomolpus rex, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view." httpUri="https://zenodo.org/record/831892/files/figure.png" pageId="1" pageNumber="272">Fig. 3</figureCitation>
). Pronotum (
<figureCitation id="13692A2EFF87C30DFD33F48EFCE5BD3B" box="[721,791,505,530]" captionStart="FIGURES 3 – 12" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[204,1404,193,1905]" captionTargetId="figure@2.[178,1409,193,1906]" captionTargetPageId="2" captionText="FIGURES 3 – 12. Tyrannomolpus rex, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view." httpUri="https://zenodo.org/record/831892/files/figure.png" pageId="1" pageNumber="272">Fig. 9</figureCitation>
) widest at middle, converging anteriorly, lateral margin distinctly undulate; base without a row of dense punctures. Hypomera weakly convex medially. Prosternum anteriorly deflexed with leading edge hidden in ventral view; medially concave with distinct carinae confluent with sides of prosternal process, sexually dimorphic with male having a transverse anterior fovea, prosternal process with well-developed lateral coxal braces, apex expanded behind coxae with sides parallel-sided and depressed apically (
<figureCitation id="13692A2EFF87C30DFF1DF7DAFE73BDEC" box="[255,385,685,710]" captionStart="FIGURES 3 – 12" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[204,1404,193,1905]" captionTargetId="figure@2.[178,1409,193,1906]" captionTargetPageId="2" captionText="FIGURES 3 – 12. Tyrannomolpus rex, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view." httpUri="https://zenodo.org/record/831892/files/figure.png" pageId="1" pageNumber="272">Figs 10, 12</figureCitation>
). Procoxal cavities externally closed (
<figureCitation id="13692A2EFF87C30DFCCEF7DAFC87BDEC" box="[812,885,685,710]" captionStart="FIGURES 1, 2" captionStartId="1.[151,264,1938,1961]" captionTargetBox="[252,1353,1059,1872]" captionTargetId="figure@1.[211,1376,1036,1891]" captionTargetPageId="1" captionText="FIGURES 1, 2. Tyrannomolpus rex, gen. nov., spec. nov. general view; 1: dorsal; 2: ventral." httpUri="https://zenodo.org/record/831890/files/figure.png" pageId="1" pageNumber="272">Figs 2</figureCitation>
, 
<figureCitation id="13692A2EFF87C30DFC63F7D9FC52BDEC" box="[897,928,686,710]" captionStart="FIGURES 3 – 12" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[204,1404,193,1905]" captionTargetId="figure@2.[178,1409,193,1906]" captionTargetPageId="2" captionText="FIGURES 3 – 12. Tyrannomolpus rex, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view." httpUri="https://zenodo.org/record/831892/files/figure.png" pageId="1" pageNumber="272">10</figureCitation>
). Elytra (
<figureCitation id="13692A2EFF87C30DFBEAF7DAFBBBBDEC" box="[1032,1097,685,710]" captionStart="FIGURES 1, 2" captionStartId="1.[151,264,1938,1961]" captionTargetBox="[252,1353,1059,1872]" captionTargetId="figure@1.[211,1376,1036,1891]" captionTargetPageId="1" captionText="FIGURES 1, 2. Tyrannomolpus rex, gen. nov., spec. nov. general view; 1: dorsal; 2: ventral." httpUri="https://zenodo.org/record/831890/files/figure.png" pageId="1" pageNumber="272">Fig. 1</figureCitation>
) moderately convex, humeral calli present, elytral punctures arranged in 10 partially confused rows with weak carinae. Hind wings fully developed (
<figureCitation id="13692A2EFF87C30DFEF8F782FE9DBC24" box="[282,367,757,782]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="1" pageNumber="272">Fig. 17</figureCitation>
). Dorsal shaft of meso- and metatibiae (
<figureCitation id="13692A2EFF87C30DFCD4F782FC7BBC24" box="[822,905,757,782]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="1" pageNumber="272">Fig. 22</figureCitation>
) without groove along entire flank, excavation confined to apex and lacking dorsal carinae, tarsal claws appendiculate (
<figureCitation id="13692A2EFF87C30DFC5CF66EFBE3BC18" box="[958,1041,793,818]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="1" pageNumber="272">Fig. 21</figureCitation>
). Pygidium (
<figureCitation id="13692A2EFF87C30DFB40F66EFB00BC18" box="[1186,1266,793,818]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="1" pageNumber="272">Fig. 18</figureCitation>
) medially with deep groove. Aedeagus (
<figureCitation id="13692A2EFF87C30DFE4EF64AFDDFBC7C" box="[428,557,829,854]" captionStart="FIGURES 26 – 31" captionStartId="5.[151,264,1468,1491]" captionTargetBox="[151,1431,381,1444]" captionTargetId="figure@5.[151,1436,373,1448]" captionTargetPageId="5" captionText="FIGURES 26 – 31. Tyrannomolpus rex, morphological details; 26: ovipositor; 27: gonocoxites; 28: spermatheca; 29: aedeagus ventral; 30: aedeagus lateral; 31: main body of penis ventral." httpUri="https://zenodo.org/record/831896/files/figure.png" pageId="1" pageNumber="272">Figs 29–31</figureCitation>
) with tegmenal strut subequal in length to penis, basal hood and main body of penis subequal in length.
</paragraph>
</subSubSection>
<subSubSection id="C3486520FF87C309FF25F6F2FDF1BFFF" lastPageId="5" lastPageNumber="276" pageId="1" pageNumber="272" type="description">
<paragraph id="8BED36ABFF87C30DFF25F6F2FE43BCE8" blockId="1.[151,1437,469,962]" pageId="1" pageNumber="272">
<emphasis id="B926EAB9FF87C30DFF25F6F2FE53BCB4" bold="true" box="[199,417,901,926]" pageId="1" pageNumber="272">Description. Body</emphasis>
(
<figureCitation id="13692A2EFF87C30DFE50F6F2FDE9BCB4" box="[434,539,901,926]" captionStart="FIGURES 1, 2" captionStartId="1.[151,264,1938,1961]" captionTargetBox="[252,1353,1059,1872]" captionTargetId="figure@1.[211,1376,1036,1891]" captionTargetPageId="1" captionText="FIGURES 1, 2. Tyrannomolpus rex, gen. nov., spec. nov. general view; 1: dorsal; 2: ventral." httpUri="https://zenodo.org/record/831890/files/figure.png" pageId="1" pageNumber="272">Figs 1, 2</figureCitation>
) elongate robust and broad, moderately convex; cuticle glabrous and dorsally setose (
<figureCitation id="13692A2EFF87C30DFF08F6DEFEC3BCE8" box="[234,305,937,962]" captionStart="FIGURES 1, 2" captionStartId="1.[151,264,1938,1961]" captionTargetBox="[252,1353,1059,1872]" captionTargetId="figure@1.[211,1376,1036,1891]" captionTargetPageId="1" captionText="FIGURES 1, 2. Tyrannomolpus rex, gen. nov., spec. nov. general view; 1: dorsal; 2: ventral." httpUri="https://zenodo.org/record/831890/files/figure.png" pageId="1" pageNumber="272">Figs 1</figureCitation>
, 
<figureCitation id="13692A2EFF87C30DFEDDF6DDFEBDBCE8" box="[319,335,938,962]" captionStart="FIGURES 3 – 12" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[204,1404,193,1905]" captionTargetId="figure@2.[178,1409,193,1906]" captionTargetPageId="2" captionText="FIGURES 3 – 12. Tyrannomolpus rex, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view." httpUri="https://zenodo.org/record/831892/files/figure.png" pageId="1" pageNumber="272">9</figureCitation>
, 
<figureCitation id="13692A2EFF87C30DFEB9F6DEFE56BCE8" box="[347,420,937,962]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="1" pageNumber="272">15, 16</figureCitation>
).
</paragraph>
<caption id="DF2D6623FF87C30DFF75F2E5FBB7B882" box="[151,1093,1938,1961]" httpUri="https://zenodo.org/record/831890/files/figure.png" pageId="1" pageNumber="272" targetBox="[252,1353,1059,1872]" targetPageId="1">
<paragraph id="8BED36ABFF87C30DFF75F2E5FBB7B882" blockId="1.[151,1093,1938,1961]" box="[151,1093,1938,1961]" pageId="1" pageNumber="272">
<emphasis id="B926EAB9FF87C30DFF75F2E5FEC8B88D" bold="true" box="[151,314,1938,1961]" pageId="1" pageNumber="272">FIGURES 1, 2.</emphasis>
<taxonomicName id="4C524D28FF87C30DFEA2F2E5FDF1B882" authority="Nadein &amp; Leschen, 2017" authorityName="Nadein &amp; Leschen" authorityYear="2017" box="[320,515,1938,1960]" class="Insecta" family="Chrysomelidae" genus="Tyrannomolpus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="rex" status="gen. nov.">
<emphasis id="B926EAB9FF87C30DFEA2F2E5FDF1B882" box="[320,515,1938,1960]" italics="true" pageId="1" pageNumber="272">Tyrannomolpus rex</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21557C2FF87C30DFDF2F2E4FD10B882" box="[528,738,1939,1960]" pageId="1" pageNumber="272" rank="species">
<emphasis id="B926EAB9FF87C30DFDF2F2E4FD9CB882" bold="true" box="[528,622,1939,1960]" pageId="1" pageNumber="272">gen. nov.</emphasis>
, 
<emphasis id="B926EAB9FF87C30DFD99F2E4FD10B882" bold="true" box="[635,738,1939,1960]" pageId="1" pageNumber="272">spec. nov.</emphasis>
</taxonomicNameLabel>
general view; 1: dorsal; 2: ventral.
</paragraph>
</caption>
<caption id="DF2D6623FF84C30EFF75F2F1FBA0B8F0" httpUri="https://zenodo.org/record/831892/files/figure.png" pageId="2" pageNumber="273" targetBox="[204,1404,193,1905]" targetPageId="2">
<paragraph id="8BED36ABFF84C30EFF75F2F1FBA0B8F0" blockId="2.[151,1436,1926,2010]" pageId="2" pageNumber="273">
<emphasis id="B926EAB9FF84C30EFF75F2F1FEB5B8B6" bold="true" box="[151,327,1926,1949]" pageId="2" pageNumber="273">FIGURES 3–12.</emphasis>
<taxonomicName id="4C524D28FF84C30EFEAFF2F0FDE2B8B7" authorityName="Nadein &amp; Leschen" authorityYear="2017" box="[333,528,1927,1949]" class="Insecta" family="Chrysomelidae" genus="Tyrannomolpus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="species" species="rex">
<emphasis id="B926EAB9FF84C30EFEAFF2F0FDE2B8B7" box="[333,528,1927,1949]" italics="true" pageId="2" pageNumber="273">Tyrannomolpus rex</emphasis>
</taxonomicName>
, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view.
</paragraph>
</caption>
<caption id="DF2D6623FF85C30FFF75F3D1FBD3B9D0" httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="3" pageNumber="274" targetBox="[151,1359,194,1682]" targetPageId="3">
<paragraph id="8BED36ABFF85C30FFF75F3D1FBD3B9D0" blockId="3.[151,1436,1702,1786]" pageId="3" pageNumber="274">
<emphasis id="B926EAB9FF85C30FFF75F3D1FEABB996" bold="true" box="[151,345,1702,1725]" pageId="3" pageNumber="274">FIGURES 13–25.</emphasis>
<taxonomicName id="4C524D28FF85C30FFE86F3D0FDDEB997" authorityName="Nadein &amp; Leschen" authorityYear="2017" box="[356,556,1703,1725]" class="Insecta" family="Chrysomelidae" genus="Tyrannomolpus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="species" species="rex">
<emphasis id="B926EAB9FF85C30FFE86F3D0FDDEB997" box="[356,556,1703,1725]" italics="true" pageId="3" pageNumber="274">Tyrannomolpus rex</emphasis>
</taxonomicName>
, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female.
</paragraph>
</caption>
<paragraph id="8BED36ABFF85C30FFF25F255FA6EB8C4" blockId="3.[151,1436,1826,2030]" pageId="3" pageNumber="274">
<emphasis id="B926EAB9FF85C30FFF25F255FEF5B811" bold="true" box="[199,263,1826,1851]" pageId="3" pageNumber="274">Head</emphasis>
(
<figureCitation id="13692A2EFF85C30FFEF5F255FEA9B810" box="[279,347,1826,1851]" captionStart="FIGURES 3 – 12" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[204,1404,193,1905]" captionTargetId="figure@2.[178,1409,193,1906]" captionTargetPageId="2" captionText="FIGURES 3 – 12. Tyrannomolpus rex, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view." httpUri="https://zenodo.org/record/831892/files/figure.png" pageId="3" pageNumber="274">Fig. 3</figureCitation>
) with vertex wide, flat, with midcranial suture, covered with long and dense setae. Frons long, trapezoidal, flat, limited by thin grooves, anterior margin of frons emarginate with pair of short and acute processes, small portions of frons above antennal insertions convex; genae short, not longer than diameter of antennal insertion. Eyes (
<figureCitation id="13692A2EFF85C30FFE5DF2FAFDF5B88C" box="[447,519,1933,1958]" captionStart="FIGURES 3 – 12" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[204,1404,193,1905]" captionTargetId="figure@2.[178,1409,193,1906]" captionTargetPageId="2" captionText="FIGURES 3 – 12. Tyrannomolpus rex, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view." httpUri="https://zenodo.org/record/831892/files/figure.png" pageId="3" pageNumber="274">Fig. 4</figureCitation>
) large, elongate-elliptic, convex. Exposed part of labrum distinctly transverse, entire labrum weakly longer than wide (
<figureCitation id="13692A2EFF85C30FFD8FF2C5FD46B8E1" box="[621,692,1970,1995]" captionStart="FIGURES 3 – 12" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[204,1404,193,1905]" captionTargetId="figure@2.[178,1409,193,1906]" captionTargetPageId="2" captionText="FIGURES 3 – 12. Tyrannomolpus rex, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view." httpUri="https://zenodo.org/record/831892/files/figure.png" pageId="3" pageNumber="274">Fig. 5</figureCitation>
), anterior margin concave, anterolaterally with three long setae, labral plate with pair of pores; tormae shorter than labral plate. Mandibles large, strongly projecting. Maxillae (
<figureCitation id="13692A2EFF85C30FFA8DF2A2FA6EB8C4" box="[1391,1436,2005,2030]" captionStart="FIGURES 3 – 12" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[204,1404,193,1905]" captionTargetId="figure@2.[178,1409,193,1906]" captionTargetPageId="2" captionText="FIGURES 3 – 12. Tyrannomolpus rex, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view." httpUri="https://zenodo.org/record/831892/files/figure.png" pageId="3" pageNumber="274">Fig.</figureCitation>
</paragraph>
<paragraph id="8BED36ABFF82C308FF75F5E0FB21BEDF" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="275">
6) large, lacinia broad, shorter than galea, the latter about twice longer than wide, apically with a bunch of dense setae, maxillary palp long, 4-segmented, palpomere 1 much shorter than following, palpomere 2 longer than following, about 3 times longer than wide, palpomere 3 and 4 equal in length, about twice longer than wide, last palpomere somewhat fusiform with truncated apex. Labium (
<figureCitation id="13692A2EFF82C308FC9AF473FC31BE36" box="[888,963,260,285]" captionStart="FIGURES 3 – 12" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[204,1404,193,1905]" captionTargetId="figure@2.[178,1409,193,1906]" captionTargetPageId="2" captionText="FIGURES 3 – 12. Tyrannomolpus rex, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view." httpUri="https://zenodo.org/record/831892/files/figure.png" pageId="4" pageNumber="275">Fig. 7</figureCitation>
) small, submentum and mentum short, transverse, labial palpi long, 3-segmented, palpomere 1 shortest, about as long as wide, palpomere 2 the longest, about 3 times longer than wide, palpomere 3 about 2.5 times longer than wide, conical with obtuse apex. Antennal insertions placed at the distance of diameter of insertions from margin of eye and anterior part of frons. Antennae (
<figureCitation id="13692A2EFF82C308FF7DF4E3FF13BE86" box="[159,225,404,429]" captionStart="FIGURES 3 – 12" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[204,1404,193,1905]" captionTargetId="figure@2.[178,1409,193,1906]" captionTargetPageId="2" captionText="FIGURES 3 – 12. Tyrannomolpus rex, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view." httpUri="https://zenodo.org/record/831892/files/figure.png" pageId="4" pageNumber="275">Fig. 8</figureCitation>
) long, filiform, 11-segmented, extending to middle of the body; scapus thickened and slightly curved, about twice longer than wide, antennomere 2 small, globose, antennomeres 3–10 long and thin, at least three times longer than wide or longer, last antennomere shorter than previous, hardly three times longer than wide.
</paragraph>
<paragraph id="8BED36ABFF82C308FF25F488FABBBC8F" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="275">
<emphasis id="B926EAB9FF82C308FF25F488FEB0BD32" bold="true" box="[199,322,511,536]" pageId="4" pageNumber="275">Prothorax</emphasis>
(
<figureCitation id="13692A2EFF82C308FEB1F488FE6BBD32" box="[339,409,511,536]" captionStart="FIGURES 3 – 12" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[204,1404,193,1905]" captionTargetId="figure@2.[178,1409,193,1906]" captionTargetPageId="2" captionText="FIGURES 3 – 12. Tyrannomolpus rex, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view." httpUri="https://zenodo.org/record/831892/files/figure.png" pageId="4" pageNumber="275">Fig. 9</figureCitation>
) with pronotum transverse (1.20–1.42 times wider than long), convex, widest at middle, converging anteriorly; anterior margin straight in middle, lateral margins with two short angular projections, undulate, posterior margin convex, rounded; hind margin with a distinct bead, anterior angles protruding, with obtuse apex, posterior angles weakly protruding, acute, each angle bear setiferous pore; pronotal surface densely punctate, punctures small, bearing long setae, densest at sides, sparser at disc; base without a row of dense punctures. Prosternum (
<figureCitation id="13692A2EFF82C308FE43F7C3FE18BDE6" box="[417,490,692,717]" captionStart="FIGURES 1, 2" captionStartId="1.[151,264,1938,1961]" captionTargetBox="[252,1353,1059,1872]" captionTargetId="figure@1.[211,1376,1036,1891]" captionTargetPageId="1" captionText="FIGURES 1, 2. Tyrannomolpus rex, gen. nov., spec. nov. general view; 1: dorsal; 2: ventral." httpUri="https://zenodo.org/record/831890/files/figure.png" pageId="4" pageNumber="275">Figs 2</figureCitation>
, 
<figureCitation id="13692A2EFF82C308FE14F7C3FDCFBDE6" box="[502,573,692,716]" captionStart="FIGURES 3 – 12" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[204,1404,193,1905]" captionTargetId="figure@2.[178,1409,193,1906]" captionTargetPageId="2" captionText="FIGURES 3 – 12. Tyrannomolpus rex, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view." httpUri="https://zenodo.org/record/831892/files/figure.png" pageId="4" pageNumber="275">10, 12</figureCitation>
) short in front of procoxae, much shorter than wide; anteriorly deflexed with leading edge hidden in ventral view; medially concave with distinct carinae confluent with the sides of the prosternal process, sexually dimorphic with male having a transverse anterior fovea, prosternal process with welldeveloped lateral coxal braces, apex expanded behind coxae with sides parallel-sided and depressed apically (
<figureCitation id="13692A2EFF82C308FA89F668FF13BC76" captionStart="FIGURES 3 – 12" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[204,1404,193,1905]" captionTargetId="figure@2.[178,1409,193,1906]" captionTargetPageId="2" captionText="FIGURES 3 – 12. Tyrannomolpus rex, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view." httpUri="https://zenodo.org/record/831892/files/figure.png" pageId="4" pageNumber="275">Figs 10, 12</figureCitation>
) hypomera weakly convex medially, hypomeral suture distinct; prosternal process comparatively narrow, between coxae about twice narrower than diameter of coxa, strongly expanded behind coxae laterally and contacting the hypomeral process; procoxal cavities externally closed (
<figureCitation id="13692A2EFF82C308FC52F6FBFC0BBC8E" box="[944,1017,908,933]" captionStart="FIGURES 1, 2" captionStartId="1.[151,264,1938,1961]" captionTargetBox="[252,1353,1059,1872]" captionTargetId="figure@1.[211,1376,1036,1891]" captionTargetPageId="1" captionText="FIGURES 1, 2. Tyrannomolpus rex, gen. nov., spec. nov. general view; 1: dorsal; 2: ventral." httpUri="https://zenodo.org/record/831890/files/figure.png" pageId="4" pageNumber="275">Figs 2</figureCitation>
, 
<figureCitation id="13692A2EFF82C308FBE7F6FBFBD6BC8E" box="[1029,1060,908,932]" captionStart="FIGURES 3 – 12" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[204,1404,193,1905]" captionTargetId="figure@2.[178,1409,193,1906]" captionTargetPageId="2" captionText="FIGURES 3 – 12. Tyrannomolpus rex, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view." httpUri="https://zenodo.org/record/831892/files/figure.png" pageId="4" pageNumber="275">10</figureCitation>
); anterior margin straight.
</paragraph>
<paragraph id="8BED36ABFF82C308FF25F6D8FD73BB8A" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="275">
<emphasis id="B926EAB9FF82C308FF25F6D8FEB3BCE2" bold="true" box="[199,321,943,968]" pageId="4" pageNumber="275">Scutellary</emphasis>
shield (
<figureCitation id="13692A2EFF82C308FE7CF6D8FE03BCE2" box="[414,497,943,968]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="4" pageNumber="275">Fig. 14</figureCitation>
) transverse and obtuse. Mesoventrite partly hidden by prosternum, T-shaped, flat, intercoxal process (
<figureCitation id="13692A2EFF82C308FE9CF6A3FE27BCC6" box="[382,469,980,1005]" captionStart="FIGURES 3 – 12" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[204,1404,193,1905]" captionTargetId="figure@2.[178,1409,193,1906]" captionTargetPageId="2" captionText="FIGURES 3 – 12. Tyrannomolpus rex, morphological details; 3: head, midcranial suture marked with white arrow; 4: head, left lateral; 5: labrum; 6: maxilla; 7: labium and labial palpi; 8: antenna; 9: pronotum dorsal; 10: prosternum; 11: mesoventrite, mesanepisternum, mesepimeron, metaventrite, metanepimeron; 12: prosternum, anterior view." httpUri="https://zenodo.org/record/831892/files/figure.png" pageId="4" pageNumber="275">Fig. 11</figureCitation>
) long and relatively narrow, longer than wide, apical half of process expanded, rectangular and elevated, apex straight; mesanepisternum triangular, nearly medially transversely impressed; mesepimeron narrow and long. Metaventrite long, as long as abdominal ventrite 1, convex, intercoxal metaventral process rather short, wide with nearly straight apex, posterior margin medially angularly and deeply emarginate and impressed; metanepimeron narrow, flat. Metendosternite (
<figureCitation id="13692A2EFF82C308FC8FF113FC31BB56" box="[877,963,1124,1149]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="4" pageNumber="275">Fig. 13</figureCitation>
) with short and broad stalk, arms long, proximal half about twice wider than distal.
</paragraph>
<paragraph id="8BED36ABFF82C308FF25F1DBFD73BAB6" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="275">
<emphasis id="B926EAB9FF82C308FF25F1DBFEE6BBEF" bold="true" box="[199,276,1196,1221]" pageId="4" pageNumber="275">Elytra</emphasis>
(
<figureCitation id="13692A2EFF82C308FEC6F1DBFE97BBEE" box="[292,357,1196,1221]" captionStart="FIGURES 1, 2" captionStartId="1.[151,264,1938,1961]" captionTargetBox="[252,1353,1059,1872]" captionTargetId="figure@1.[211,1376,1036,1891]" captionTargetPageId="1" captionText="FIGURES 1, 2. Tyrannomolpus rex, gen. nov., spec. nov. general view; 1: dorsal; 2: ventral." httpUri="https://zenodo.org/record/831890/files/figure.png" pageId="4" pageNumber="275">Fig. 1</figureCitation>
) elongate (3.29–3.76 x longer than pronotal length) and broad (1.52–1.62 x the greatest elytral width), moderately convex, base 1.45 times wider than the combined based of the pronotum, lateral sides vertical; humeral calli large and projecting; elytral surface punctate (
<figureCitation id="13692A2EFF82C308FCB2F183FC54BA27" box="[848,934,1268,1293]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="4" pageNumber="275">Fig. 15</figureCitation>
), punctures larger than those on pronotum, arranged in 10 long rows, punctures in rows somewhat confused and irregular, sutural row short, not reaching middle of elytra, of multiply, dense, confused punctures; intervals between rows convex, finely punctate, surface covered with dense setae (
<figureCitation id="13692A2EFF82C308FE59F028FDFCBA52" box="[443,526,1375,1400]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="4" pageNumber="275">Fig. 16</figureCitation>
). Epipleura rather narrow and feebly concave, widest at base and visible in lateral view. Hind wings fully developed (
<figureCitation id="13692A2EFF82C308FDC3F0F3FD86BAB6" box="[545,628,1412,1437]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="4" pageNumber="275">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="8BED36ABFF82C308FF25F0D0FD40B97A" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="275">
<emphasis id="B926EAB9FF82C308FF25F0D0FEBFBAEA" bold="true" box="[199,333,1447,1472]" pageId="4" pageNumber="275">Abdominal</emphasis>
(
<figureCitation id="13692A2EFF82C308FEBDF0D0FE41BAEA" box="[351,435,1447,1472]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="4" pageNumber="275">Fig. 19</figureCitation>
) ventrite I moderately convex, intercoxal process long and wide, its apex triangularlyrounded; ventrites II–IV short, equal in length, much shorter than ventrite I, last ventrite nearly as long as previous or slightly longer, its margin rounded with straight apex, in male (
<figureCitation id="13692A2EFF82C308FC92F098FC33B922" box="[880,961,1519,1544]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="4" pageNumber="275">Fig. 20</figureCitation>
) and evenly arcuate in female, subapically with a medial deep spherical setose fovea; pygidium (tergum VII) wide and relatively short, medially with wide and deep groove (
<figureCitation id="13692A2EFF82C308FEBDF340FE40B97A" box="[351,434,1591,1616]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="4" pageNumber="275">Fig. 18</figureCitation>
), not reaching to apex.
</paragraph>
<paragraph id="8BED36ABFF82C308FF25F32BFAC1B867" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="275">
<emphasis id="B926EAB9FF82C308FF25F32BFF0DB95F" bold="true" box="[199,255,1628,1653]" pageId="4" pageNumber="275">Legs</emphasis>
(
<figureCitation id="13692A2EFF82C308FEF2F32BFE89B95E" box="[272,379,1628,1653]" captionStart="FIGURES 1, 2" captionStartId="1.[151,264,1938,1961]" captionTargetBox="[252,1353,1059,1872]" captionTargetId="figure@1.[211,1376,1036,1891]" captionTargetPageId="1" captionText="FIGURES 1, 2. Tyrannomolpus rex, gen. nov., spec. nov. general view; 1: dorsal; 2: ventral." httpUri="https://zenodo.org/record/831890/files/figure.png" pageId="4" pageNumber="275">Figs 1, 2</figureCitation>
, 
<figureCitation id="13692A2EFF82C308FE68F32BFE26B95E" box="[394,468,1628,1652]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="4" pageNumber="275">22, 23</figureCitation>
) large and somewhat thin, femora lacking teeth, tibiae (
<figureCitation id="13692A2EFF82C308FBBCF32BFB16B95E" box="[1118,1252,1628,1653]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="4" pageNumber="275">Figs 22, 23</figureCitation>
) thin, gradually widened towards apex, dorsal shaft of tibiae with two thin longitudinal ridges, without groove along entire flank, tibial spur indistinct, apex of middle and hind tibiae weakly expanded in two short and thin blades, with excavation confined to apex and lacking dorsal carinae; tarsomere (
<figureCitation id="13692A2EFF82C308FCD7F3B0FC1DB9CA" box="[821,1007,1735,1760]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="4" pageNumber="275">Figs 22, 24, 25</figureCitation>
) I somewhat longer than following, triangular, its apex hardly narrower than apex of following, tarsomere II triangular with wide apex, tarsomere III widest, long, deeply bilobed, last tarsomere as long as first, weakly projecting from third, tarsal claws appendiculate (
<figureCitation id="13692A2EFF82C308FEA0F243FE67B866" box="[322,405,1844,1869]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="4" pageNumber="275">Fig. 21</figureCitation>
); tarsi of males (
<figureCitation id="13692A2EFF82C308FDB2F243FD50B866" box="[592,674,1844,1869]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="4" pageNumber="275">Fig. 24</figureCitation>
) slightly wider and broader than those of female (
<figureCitation id="13692A2EFF82C308FB30F243FAD7B867" box="[1234,1317,1844,1869]" captionStart="FIGURES 13 – 25" captionStartId="3.[151,264,1702,1725]" captionTargetBox="[151,1359,194,1682]" captionTargetId="figure@3.[151,1436,193,1682]" captionTargetPageId="3" captionText="FIGURES 13 – 25. Tyrannomolpus rex, morphological details; 13: metendosternite; 14: mesonotum and scutellary shield dorsal; 15: elytral disc; 16: elytral apex; 17: hind wing right; 18: pygidium (tergum VII); 19: abdomen; 20: abdominal apex; 21: tarsal claws; 22: left hind leg; 23: right protibia; 24: protarsus male; 25: protarsus female." httpUri="https://zenodo.org/record/831894/files/figure.png" pageId="4" pageNumber="275">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="8BED36ABFF82C308FF25F220FDC0B893" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="275">
<emphasis id="B926EAB9FF82C308FF25F220FEC9B85A" bold="true" box="[199,315,1879,1904]" pageId="4" pageNumber="275">Aedeagus</emphasis>
(
<figureCitation id="13692A2EFF82C308FEAAF220FE3AB85A" box="[328,456,1879,1904]" captionStart="FIGURES 26 – 31" captionStartId="5.[151,264,1468,1491]" captionTargetBox="[151,1431,381,1444]" captionTargetId="figure@5.[151,1436,373,1448]" captionTargetPageId="5" captionText="FIGURES 26 – 31. Tyrannomolpus rex, morphological details; 26: ovipositor; 27: gonocoxites; 28: spermatheca; 29: aedeagus ventral; 30: aedeagus lateral; 31: main body of penis ventral." httpUri="https://zenodo.org/record/831896/files/figure.png" pageId="4" pageNumber="275">Figs 29–31</figureCitation>
) with tegmenal strut (tegmen) subequal in length to penis, basal hood and main body of penis subequal in length; overall shape long and broad, weakly depressed dorsoventrally, slightly curved in lateral view, basal hood weakly sclerotized.
</paragraph>
<paragraph id="8BED36ABFF82C309FF25F2B3FDF1BFFF" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1437,151,320]" lastPageId="5" lastPageNumber="276" pageId="4" pageNumber="275">
<emphasis id="B926EAB9FF82C308FF25F2B3FEB5B8F7" bold="true" box="[199,327,1988,2013]" pageId="4" pageNumber="275">Ovipositor</emphasis>
(
<figureCitation id="13692A2EFF82C308FEB7F2B3FE26B8F6" box="[341,468,1988,2013]" captionStart="FIGURES 26 – 31" captionStartId="5.[151,264,1468,1491]" captionTargetBox="[151,1431,381,1444]" captionTargetId="figure@5.[151,1436,373,1448]" captionTargetPageId="5" captionText="FIGURES 26 – 31. Tyrannomolpus rex, morphological details; 26: ovipositor; 27: gonocoxites; 28: spermatheca; 29: aedeagus ventral; 30: aedeagus lateral; 31: main body of penis ventral." httpUri="https://zenodo.org/record/831896/files/figure.png" pageId="4" pageNumber="275">Figs 26, 27</figureCitation>
) long; gonocoxites laterally compressed and somewhat crescent shaped, about 2 times longer than wide and gradually expanded apically with short subapical gonostyli about the length of coxite. Spermatheca (
<figureCitation id="13692A2EFF83C309FED5F5CBFE7ABFFE" box="[311,392,188,213]" captionStart="FIGURES 26 – 31" captionStartId="5.[151,264,1468,1491]" captionTargetBox="[151,1431,381,1444]" captionTargetId="figure@5.[151,1436,373,1448]" captionTargetPageId="5" captionText="FIGURES 26 – 31. Tyrannomolpus rex, morphological details; 26: ovipositor; 27: gonocoxites; 28: spermatheca; 29: aedeagus ventral; 30: aedeagus lateral; 31: main body of penis ventral." httpUri="https://zenodo.org/record/831896/files/figure.png" pageId="5" pageNumber="276">Fig. 28</figureCitation>
) c-shaped.
</paragraph>
</subSubSection>
<subSubSection id="C3486520FF83C309FF25F5A8FEE2BE6A" pageId="5" pageNumber="276" type="etymology">
<paragraph id="8BED36ABFF83C309FF25F5A8FEE2BE6A" blockId="5.[151,1437,151,320]" pageId="5" pageNumber="276">
<emphasis id="B926EAB9FF83C309FF25F5A8FEBCBFD2" bold="true" box="[199,334,223,248]" pageId="5" pageNumber="276">Etymology.</emphasis>
The genus name combines the dinosaur genus name 
<taxonomicName id="4C524D28FF83C309FC29F596FB81BFD2" authorityName="Osborn" authorityYear="1905" box="[971,1139,225,248]" class="Reptilia" family="Tyrannosauridae" genus="Tyrannosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Avetheropoda" pageId="5" pageNumber="276" phylum="Chordata" rank="genus">
<emphasis id="B926EAB9FF83C309FC29F596FB81BFD2" box="[971,1139,225,248]" italics="true" pageId="5" pageNumber="276">Tyrannosaurus</emphasis>
</taxonomicName>
with the suffix “molpus, referring to the relatively large size of the new genus to other chrysomelids endemic to 
<collectingCountry id="F345763BFF83C309FB7DF472FACEBE37" box="[1183,1340,260,285]" name="New Zealand" pageId="5" pageNumber="276">New Zealand</collectingCountry>
. Gender masculine.
</paragraph>
</subSubSection>
<caption id="DF2D6623FF83C309FF75F0CBFD0CBAD8" httpUri="https://zenodo.org/record/831896/files/figure.png" pageId="5" pageNumber="276" targetBox="[151,1431,381,1444]" targetPageId="5">
<paragraph id="8BED36ABFF83C309FF75F0CBFD0CBAD8" blockId="5.[151,1436,1468,1522]" pageId="5" pageNumber="276">
<emphasis id="B926EAB9FF83C309FF75F0CBFEA7BAF8" bold="true" box="[151,341,1468,1491]" pageId="5" pageNumber="276">FIGURES 26–31.</emphasis>
<taxonomicName id="4C524D28FF83C309FEB9F0CAFDECBAF9" authorityName="Nadein &amp; Leschen" authorityYear="2017" box="[347,542,1469,1491]" class="Insecta" family="Chrysomelidae" genus="Tyrannomolpus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="species" species="rex">
<emphasis id="B926EAB9FF83C309FEB9F0CAFDECBAF9" box="[347,542,1469,1491]" italics="true" pageId="5" pageNumber="276">Tyrannomolpus rex</emphasis>
</taxonomicName>
, morphological details; 26: ovipositor; 27: gonocoxites; 28: spermatheca; 29: aedeagus ventral; 30: aedeagus lateral; 31: main body of penis ventral.
</paragraph>
</caption>
<subSubSection id="C3486520FF83C30BFF25F369FCABBD17" lastPageId="7" lastPageNumber="278" pageId="5" pageNumber="276" type="discussion">
<paragraph id="8BED36ABFF83C309FF25F369FB78B8CC" blockId="5.[151,1437,1566,2023]" pageId="5" pageNumber="276">
<emphasis id="B926EAB9FF83C309FF25F369FEBDB91D" bold="true" box="[199,335,1566,1591]" pageId="5" pageNumber="276">Comments.</emphasis>
The attribution of the new genus to the tribe 
<taxonomicName id="4C524D28FF83C309FCB2F369FC34B91D" authorityName="Hope" authorityYear="1840" box="[848,966,1566,1591]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="tribe" tribe="Eumolpini">Eumolpini</taxonomicName>
is based on the following combination of characters: simple apices of meso- and metatibiae, carinate and irregularly curved sides of prothorax, appendiculate tarsal claws, pygidium with a longitudinal medial groove, aedeagus with a constriction between a median lobe and a basal hood (after 
<bibRefCitation id="EFC34B5AFF83C309FE8EF3FDFE1BB989" author="Chen" box="[364,489,1674,1699]" pageId="5" pageNumber="276" refString="Chen, S. H. (1940) Notes on Chinese Eumolpidae. Sinensia, 11, 483 - 528." type="journal article" year="1940">Chen 1940</bibRefCitation>
; 
<bibRefCitation id="EFC34B5AFF83C309FE16F3FDFD7BB989" author="Selman" box="[500,649,1674,1699]" pageId="5" pageNumber="276" refString="Selman, B. J. (1965) A revision of Nodini and a key to the genera of Eumolpidae of Africa (Coleoptera: Eumolpidae). Bulletin of the British Museum of Natural History, Entomology, 16, 141 - 174." type="journal article" year="1965">Selman 1965</bibRefCitation>
; 
<bibRefCitation id="EFC34B5AFF83C309FD77F3FDFC2BB989" author="Moseyko" box="[661,985,1674,1699]" pageId="5" pageNumber="276" refString="Moseyko, A. G. &amp; Kirejtshuk, A. G. (2013) On the taxonomic position of new taxa of the subfamily Eumolpinae (Coleoptera, Chrysomelidae) from Baltic amber. Terrestrial Arthropod Reviews, 6, 163 - 172." type="journal article" year="2013">Moseyko &amp; Kirejtshuk 2013</bibRefCitation>
). The eumolpines are in need in revision from most regions in the Pacific and Australia (
<bibRefCitation id="EFC34B5AFF83C309FD59F3D9FCC5B9ED" author="Reid" box="[699,823,1710,1735]" pageId="5" pageNumber="276" refString="Reid, C. A. M. (2017) Australopapuan leaf beetle diversity: the contributions of hosts plants and geography. Austral Entomology, 56 (2), 123 - 137" type="journal article" year="2017">Reid 2017</bibRefCitation>
), and 
<taxonomicName id="4C524D28FF83C309FC9FF3D9FBD9B9ED" authority="Nadein &amp; Leschen, 2017" authorityName="Nadein &amp; Leschen" authorityYear="2017" box="[893,1067,1710,1735]" class="Insecta" family="Chrysomelidae" genus="Tyrannomolpus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B926EAB9FF83C309FC9FF3D9FBD9B9ED" box="[893,1067,1710,1735]" italics="true" pageId="5" pageNumber="276">Tyrannomolpus</emphasis>
</taxonomicName>
can be distinguished from most Australian (
<bibRefCitation id="EFC34B5AFF83C309FEF8F3A5FE4BB9C1" author="Chapuis" box="[282,441,1746,1771]" pageId="5" pageNumber="276" refString="Chapuis, F. (1874) Histoire Naturelle des Insectes. Genera des Coleopteres. X. Phytophages. Libraire Encyclopedique de Roret, Paris, iv + 455 pp." type="book" year="1874">Chapuis 1874</bibRefCitation>
), New Caledonian (e.g., Gómez-Zurita &amp; Cardoso 2014; 
<bibRefCitation id="EFC34B5AFF83C309FBD5F3A5FAF3B9C1" author="Jolivet" box="[1079,1281,1746,1771]" pageId="5" pageNumber="276" refString="Jolivet, P., Verma, K. &amp; Mille, C. (2007) New species of Eumolpinae from the genera Dematochroma Baly, 1864 and Taophila Heller, 1916 from New Caledonia (Coleoptera: Eumolpinae). Revue Francaise d'Entomologie, New Series, 29 (1), 33 - 47." type="journal article" year="2007">
Jolivet 
<emphasis id="B926EAB9FF83C309FB6AF3A4FB4BB9C1" box="[1160,1209,1746,1771]" italics="true" pageId="5" pageNumber="276">et al</emphasis>
. 2007
</bibRefCitation>
, 
<bibRefCitation id="EFC34B5AFF83C309FAE9F3A4FAB4B9C1" author="Jolivet" box="[1291,1350,1747,1771]" pageId="5" pageNumber="276" refString="Jolivet, P., Verma, K. &amp; Mille, C. (2009) Eumolpinae recently collected in New Caledonia and Vanuatu (Coleoptera, Chrysomelidae). Nouvelle Revue d'Entomologie, New Series, 26 (1), 4 - 17." type="journal article" year="2009">2009</bibRefCitation>
; 
<bibRefCitation id="EFC34B5AFF83C309FAB3F3A5FEBBB825" author="Jolivet" pageId="5" pageNumber="276" refString="Jolivet, P. &amp; Verma, K. K. (2010) Biogeography and Biology of the New Caledonian Chrysomelidae (Coleoptera). In: Jolivet, P., Santiago-Blay, J. &amp; Schmitt, M. (Eds.), Research on Chrysomelidae. Uol. 2. Brill Publishers, Leiden, pp. 211 - 223." type="journal article" year="2010">Jolivet &amp; Verma 2010</bibRefCitation>
; 
<bibRefCitation id="EFC34B5AFF83C309FEB8F381FDE9B825" author="Samuelson" box="[346,539,1782,1807]" pageId="5" pageNumber="276" refString="Samuelson, G. A. (2010) Review of Taophila, a genus endemic to New Caledonia (Coleoptera: Chrysomelidae: Eumolpinae). Zootaxa, 2621, 45 - 62." type="journal article" year="2010">Samuelson 2010</bibRefCitation>
) and New Zealand eumolpine genera (
<figureCitation id="13692A2EFF83C309FC09F381FB82B825" box="[1003,1136,1782,1807]" captionStart="FIGURES 32 – 35" captionStartId="6.[151,264,1703,1726]" captionTargetBox="[277,1310,219,1652]" captionTargetId="figure@6.[267,1319,213,1667]" captionTargetPageId="6" captionText="FIGURES 32 – 35. New Zealand Eumolpinae; 32: Pilacolaspis sp., body length 5.1 mm; 33: Atrichatus aeneicollis Broun, body length 5.5. mm; 34: Peniticus suffusus Sharp, body length 4.6 mm; 35: Eucolaspis nr. brunnea (Fabricius), body length 4.1 mm." httpUri="https://zenodo.org/record/831898/files/figure.png" pageId="5" pageNumber="276">Figs 32–35</figureCitation>
) by the prothorax that is narrower than the elytral base. This character is also present in at least one species of 
<taxonomicName id="4C524D28FF83C309FB8CF26DFAD2B819" authority="Heller" authorityName="Heller" authorityYear="1916" box="[1134,1312,1818,1843]" class="Insecta" family="Chrysomelidae" genus="Taophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB9FF83C309FB8CF26DFB23B819" box="[1134,1233,1818,1843]" italics="true" pageId="5" pageNumber="276">Taophila</emphasis>
Heller
</taxonomicName>
from New Caledonia, but lacks the setose vestiture present in 
<taxonomicName id="4C524D28FF83C309FD39F249FC7BB87D" authority="Nadein &amp; Leschen, 2017" authorityName="Nadein &amp; Leschen" authorityYear="2017" box="[731,905,1854,1879]" class="Insecta" family="Chrysomelidae" genus="Tyrannomolpus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B926EAB9FF83C309FD39F249FC7BB87D" box="[731,905,1854,1879]" italics="true" pageId="5" pageNumber="276">Tyrannomolpus</emphasis>
</taxonomicName>
. Morphologically, 
<taxonomicName id="4C524D28FF83C309FB83F249FAFDB87D" authority="Nadein &amp; Leschen, 2017" authorityName="Nadein &amp; Leschen" authorityYear="2017" box="[1121,1295,1854,1879]" class="Insecta" family="Chrysomelidae" genus="Tyrannomolpus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B926EAB9FF83C309FB83F249FAFDB87D" box="[1121,1295,1854,1879]" italics="true" pageId="5" pageNumber="276">Tyrannomolpus</emphasis>
</taxonomicName>
is closest to 
<taxonomicName id="4C524D28FF83C309FF75F215FED0B851" authorityName="Sharp. A" authorityYear="1886" box="[151,290,1890,1915]" class="Insecta" family="Chrysomelidae" genus="Pilacolaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB9FF83C309FF75F215FED0B851" box="[151,290,1890,1915]" italics="true" pageId="5" pageNumber="276">Pilacolaspis</emphasis>
</taxonomicName>
(
<figureCitation id="13692A2EFF83C309FED2F215FE73B851" box="[304,385,1890,1915]" captionStart="FIGURES 32 – 35" captionStartId="6.[151,264,1703,1726]" captionTargetBox="[277,1310,219,1652]" captionTargetId="figure@6.[267,1319,213,1667]" captionTargetPageId="6" captionText="FIGURES 32 – 35. New Zealand Eumolpinae; 32: Pilacolaspis sp., body length 5.1 mm; 33: Atrichatus aeneicollis Broun, body length 5.5. mm; 34: Peniticus suffusus Sharp, body length 4.6 mm; 35: Eucolaspis nr. brunnea (Fabricius), body length 4.1 mm." httpUri="https://zenodo.org/record/831898/files/figure.png" pageId="5" pageNumber="276">Fig. 32</figureCitation>
) from the mainland of New Zealand from which it differs by the much larger size (
<quantity id="4CAA9B4EFF83C309FAF8F214FA8DB850" box="[1306,1407,1891,1915]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="5" pageNumber="276" unit="mm" value="10.3">10.3 mm</quantity>
or more versus up to about 
<quantity id="4CAA9B4EFF83C309FE4CF2F1FDF7B8B4" box="[430,517,1926,1951]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="5" pageNumber="276" unit="mm" value="5.9">5.9 mm</quantity>
in 
<taxonomicName id="4C524D28FF83C309FDC8F2F1FD46B8B5" authorityName="Sharp. A" authorityYear="1886" box="[554,692,1926,1951]" class="Insecta" family="Chrysomelidae" genus="Pilacolaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB9FF83C309FDC8F2F1FD46B8B5" box="[554,692,1926,1951]" italics="true" pageId="5" pageNumber="276">Pilacolaspis</emphasis>
</taxonomicName>
) and undulate outline of the pronotum (simple in 
<taxonomicName id="4C524D28FF83C309FB0BF2F1FA81B8B5" authorityName="Sharp. A" authorityYear="1886" box="[1257,1395,1926,1951]" class="Insecta" family="Chrysomelidae" genus="Pilacolaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB9FF83C309FB0BF2F1FA81B8B5" box="[1257,1395,1926,1951]" italics="true" pageId="5" pageNumber="276">Pilacolaspis</emphasis>
</taxonomicName>
) as well as the longer eyes (length/width ratio is about 2, in 
<taxonomicName id="4C524D28FF83C309FCDBF2DDFC31B8E9" authorityName="Sharp. A" authorityYear="1886" box="[825,963,1962,1987]" class="Insecta" family="Chrysomelidae" genus="Pilacolaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB9FF83C309FCDBF2DDFC31B8E9" box="[825,963,1962,1987]" italics="true" pageId="5" pageNumber="276">Pilacolaspis</emphasis>
</taxonomicName>
1.2), metaventrite as long as abdominal ventrite I (shorter in 
<taxonomicName id="4C524D28FF83C309FE9CF2B9FDFBB8CD" authorityName="Sharp. A" authorityYear="1886" box="[382,521,1998,2023]" class="Insecta" family="Chrysomelidae" genus="Pilacolaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB9FF83C309FE9CF2B9FDFBB8CD" box="[382,521,1998,2023]" italics="true" pageId="5" pageNumber="276">Pilacolaspis</emphasis>
</taxonomicName>
) and the vertex of head with a distinct midcranial suture.
</paragraph>
<caption id="DF2D6623FF80C30AFF75F3D0FA6EB9F6" httpUri="https://zenodo.org/record/831898/files/figure.png" pageId="6" pageNumber="277" targetBox="[277,1310,219,1652]" targetPageId="6">
<paragraph id="8BED36ABFF80C30AFF75F3D0FA6EB9F6" blockId="6.[151,1436,1703,1756]" pageId="6" pageNumber="277">
<emphasis id="B926EAB9FF80C30AFF75F3D0FEA1B996" bold="true" box="[151,339,1703,1726]" pageId="6" pageNumber="277">FIGURES 32–35.</emphasis>
New Zealand Eumolpinae; 32: 
<taxonomicName id="4C524D28FF80C30AFD70F3D0FCFDB997" authorityName="Sharp. A" authorityYear="1886" box="[658,783,1703,1725]" class="Insecta" family="Chrysomelidae" genus="Pilacolaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB9FF80C30AFD70F3D0FCFDB997" box="[658,783,1703,1725]" italics="true" pageId="6" pageNumber="277">Pilacolaspis</emphasis>
</taxonomicName>
sp., body length 5.1 mm; 33: 
<taxonomicName id="4C524D28FF80C30AFBD9F3D0FA92B997" authority="Broun" authorityName="Broun" box="[1083,1376,1703,1725]" class="Insecta" family="Chrysomelidae" genus="Atrichatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="277" phylum="Arthropoda" rank="species" species="aeneicollis">
<emphasis id="B926EAB9FF80C30AFBD9F3D0FAEBB997" box="[1083,1305,1703,1725]" italics="true" pageId="6" pageNumber="277">Atrichatus aeneicollis</emphasis>
Broun
</taxonomicName>
, body length 5.5. mm; 34: 
<taxonomicName id="4C524D28FF80C30AFE80F3B0FDA8B9F6" authority="Sharp" authorityName="Sharp" box="[354,602,1734,1756]" class="Insecta" family="Chrysomelidae" genus="Peniticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="277" phylum="Arthropoda" rank="species" species="suffusus">
<emphasis id="B926EAB9FF80C30AFE80F3B0FE32B9F6" box="[354,448,1735,1756]" italics="true" pageId="6" pageNumber="277">Peniticus</emphasis>
<emphasis id="B926EAB9FF80C30AFE24F3B1FDEAB9F6" box="[454,536,1734,1756]" italics="true" pageId="6" pageNumber="277">suffusus</emphasis>
Sharp
</taxonomicName>
, body length 4.6 mm; 35: 
<taxonomicName id="4C524D28FF80C30AFC83F3B1FC22B9F6" authorityName="Sharp" authorityYear="1886" box="[865,976,1734,1756]" class="Insecta" family="Chrysomelidae" genus="Eucolaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB9FF80C30AFC83F3B1FC22B9F6" box="[865,976,1734,1756]" italics="true" pageId="6" pageNumber="277">Eucolaspis</emphasis>
</taxonomicName>
nr. 
<emphasis id="B926EAB9FF80C30AFC14F3B1FBB8B9F6" box="[1014,1098,1734,1756]" italics="true" pageId="6" pageNumber="277">brunnea</emphasis>
(Fabricius), body length 4.1 mm.
</paragraph>
</caption>
<paragraph id="8BED36ABFF80C30BFF25F27EFBB0BE4F" blockId="6.[151,1436,1801,2006]" lastBlockId="7.[151,1436,151,573]" lastPageId="7" lastPageNumber="278" pageId="6" pageNumber="277">
<materialsCitation id="3B3A3CF6FF80C30BFF25F27EFBB0BE4F" ID-GBIF-Occurrence="1576159165" country="New Zealand" lastPageId="7" lastPageNumber="278" location="Loyalty Islands" pageId="6" pageNumber="277" specimenCount="1" stateProvince="Loyalty Islands">
The most notable feature of 
<taxonomicName id="4C524D28FF80C30AFDFEF27EFD38B808" authority="Nadein &amp; Leschen, 2017" authorityName="Nadein &amp; Leschen" authorityYear="2017" box="[540,714,1801,1826]" class="Insecta" family="Chrysomelidae" genus="Tyrannomolpus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B926EAB9FF80C30AFDFEF27EFD38B808" box="[540,714,1801,1826]" italics="true" pageId="6" pageNumber="277">Tyrannomolpus</emphasis>
</taxonomicName>
is its large body size, which might be an example of island gigantism well known to vertebrates (e.g., 
<bibRefCitation id="EFC34B5AFF80C30AFD78F25BFC93B86F" author="Jaffe" box="[666,865,1836,1861]" pageId="6" pageNumber="277" refString="Jaffe, A. L., Slater, G. J. &amp; Alfaro, M. E. (2011) The evolution of island gigantism and body size variation in tortoises and turtles. Biology letters, 7 (4), 558 - 561." type="journal article" year="2011">
Jaffe 
<emphasis id="B926EAB9FF80C30AFD3FF259FCE1B86F" box="[733,787,1836,1861]" italics="true" pageId="6" pageNumber="277">et al</emphasis>
. 2011
</bibRefCitation>
) and occurs among 
<collectingCountry id="F345763BFF80C30AFBB9F259FB05B86F" box="[1115,1271,1836,1861]" name="New Zealand" pageId="6" pageNumber="277">New Zealand</collectingCountry>
invertebrates, including weevils (
<bibRefCitation id="EFC34B5AFF80C30AFE93F226FDABB840" author="Buckley" box="[369,601,1873,1898]" pageId="6" pageNumber="277" refString="Buckley, T. B., Krosch, M. &amp; Leschen, R. A. B. (2015) Evolution of New Zealand insects: summary and prospectus for future research. Austral Entomology, 54, 1 - 27." type="journal article" year="2015">
Buckley 
<emphasis id="B926EAB9FF80C30AFE38F225FDFCB840" box="[474,526,1873,1898]" italics="true" pageId="6" pageNumber="277">et al</emphasis>
. 2015
</bibRefCitation>
; 
<bibRefCitation id="EFC34B5AFF80C30AFD8AF226FC77B840" author="Kuschel" box="[616,901,1873,1898]" pageId="6" pageNumber="277" refString="Kuschel, G. &amp; worthy, T. H. (1996) Past distribution of large weevils (Coleoptera: Curculionidae) in the South Island, New Zealand, based on Holocene fossil remains title. New Zealand Entomologist, 19, 15 - 19. https: // doi. org / 10.1080 / 00779962.1996.9722016" type="journal article" year="1996">Kuschel &amp; Worthy 1996</bibRefCitation>
; 
<bibRefCitation id="EFC34B5AFF80C30AFC71F226FB5BB843" author="Leschen" box="[915,1193,1873,1898]" pageId="6" pageNumber="277" refString="Leschen, R. A. B. &amp; Rhode, B. E. (2002) A new genus and species of large extinct Ulodidae (Coleoptera) from New Zealand. New Zealand Entomologist, 25, 57 - 64." type="journal article" year="2002">Leschen &amp; Rhode 2002</bibRefCitation>
). But the exact sister relationships of 
<taxonomicName id="4C524D28FF80C30AFEB7F202FDF0B8A4" authority="Nadein &amp; Leschen, 2017" authorityName="Nadein &amp; Leschen" authorityYear="2017" box="[341,514,1909,1934]" class="Insecta" family="Chrysomelidae" genus="Tyrannomolpus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B926EAB9FF80C30AFEB7F202FDF0B8A4" box="[341,514,1909,1934]" italics="true" pageId="6" pageNumber="277">Tyrannomolpus</emphasis>
</taxonomicName>
among Melanesian Rift eumolpines is unknown, though it is most similar to 
<taxonomicName id="4C524D28FF80C30AFF75F2EEFED0B898" authorityName="Sharp. A" authorityYear="1886" box="[151,290,1945,1970]" class="Insecta" family="Chrysomelidae" genus="Pilacolaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB9FF80C30AFF75F2EEFED0B898" box="[151,290,1945,1970]" italics="true" pageId="6" pageNumber="277">Pilacolaspis</emphasis>
</taxonomicName>
based on similar elytral punctuation and pubescence (especially in the apical half). In a biogeographic analysis of the region, 
<bibRefCitation id="EFC34B5AFF80C30AFE7FF2C9FD91B8FF" box="[413,611,1981,2006]" pageId="6" pageNumber="277" refString="Ung, V., Michaux, B. &amp; Leschen, R. A. B. (2017) A comprehensive vicariant model for Southwest Pacific biotas. Systematic Botany, 29 (60), 424 - 439." type="journal article">
Ung 
<emphasis id="B926EAB9FF80C30AFE34F2C9FDFBB8FC" box="[470,521,1981,2006]" italics="true" pageId="6" pageNumber="277">et al</emphasis>
. (2017)
</bibRefCitation>
propose a sister relationship of the Three Kings with 
<collectingCountry id="F345763BFF80C30AFB32F2C9FA99B8FC" box="[1232,1387,1981,2006]" name="New Zealand" pageId="6" pageNumber="277">New Zealand</collectingCountry>
, but also that 
<collectingCountry id="F345763BFF81C30BFF1FF5E0FE96BF9A" box="[253,356,151,176]" name="Australia" pageId="7" pageNumber="278">Australia</collectingCountry>
is the sister to the Melanesian Rift with 
<collectingCountry id="F345763BFF81C30BFCCFF5EFFC2EBF9A" box="[813,988,151,176]" name="New Caledonia" pageId="7" pageNumber="278">New Caledonia</collectingCountry>
and the 
<collectingRegion id="4996F849FF81C30BFBDEF5E0FB18BF9A" box="[1084,1258,151,176]" country="New Caledonia" name="Province des Iles" pageId="7" pageNumber="278">Loyalty Islands</collectingRegion>
at the base. We may predict that 
<taxonomicName id="4C524D28FF81C30BFE8EF5CBFDEBBFFF" authority="Nadein &amp; Leschen, 2017" authorityName="Nadein &amp; Leschen" authorityYear="2017" box="[364,537,188,213]" class="Insecta" family="Chrysomelidae" genus="Tyrannomolpus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B926EAB9FF81C30BFE8EF5CBFDEBBFFF" box="[364,537,188,213]" italics="true" pageId="7" pageNumber="278">Tyrannomolpus</emphasis>
</taxonomicName>
may indeed be related to the 
<collectingCountry id="F345763BFF81C30BFC7DF5CAFBCFBFFF" box="[927,1085,188,213]" name="New Zealand" pageId="7" pageNumber="278">New Zealand</collectingCountry>
<taxonomicName id="4C524D28FF81C30BFBAEF5CBFB24BFFF" authorityName="Sharp. A" authorityYear="1886" box="[1100,1238,188,213]" class="Insecta" family="Chrysomelidae" genus="Pilacolaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB9FF81C30BFBAEF5CBFB24BFFF" box="[1100,1238,188,213]" italics="true" pageId="7" pageNumber="278">Pilacolaspis</emphasis>
</taxonomicName>
, but at present, phylogenetic relationships among eumolpines are incomplete (e.g., 
<bibRefCitation id="EFC34B5AFF81C30BFC6AF5A8FB5ABFD2" author="Papadopoulou" box="[904,1192,223,248]" pageId="7" pageNumber="278" refString="Papadopoulou, A., Cardoso, A. &amp; Gomez? Zurita, J. (2013) Diversity and diversification of Eumolpinae (Coleoptera: Chrysomelidae) in New Caledonia. Zoological Journal of the Linnean Society, 168 (3), 473 - 495. https: // doi. org / 10.1111 / zoj. 12039" type="journal article" year="2013">
Papadopoulou 
<emphasis id="B926EAB9FF81C30BFBCCF596FBACBFD2" box="[1070,1118,223,248]" italics="true" pageId="7" pageNumber="278">et al</emphasis>
. 2013
</bibRefCitation>
), though strides in the taxonomic and phylogenetic relationships of the 
<collectingCountry id="F345763BFF81C30BFD2DF472FC72BE37" box="[719,896,260,285]" name="New Caledonia" pageId="7" pageNumber="278">New Caledonia</collectingCountry>
fauna have been made recently (e.g., Gómez- 
<bibRefCitation id="EFC34B5AFF81C30BFF75F450FEE8BE6A" author="Zurita" box="[151,282,295,320]" pageId="7" pageNumber="278" refString="Gomez-Zurita, J. (2011) Revision of New Caledonian species of Eumolpinae described by K. M. Heller (Coleoptera: Chrysomelidae). Zootaxa, 3060, 31 - 46." type="journal article" year="2011">Zurita 2011</bibRefCitation>
and references above) while discussions about its origin are mostly verbal (
<bibRefCitation id="EFC34B5AFF81C30BFB96F450FAA3BE6A" author="Jolivet" box="[1140,1361,295,320]" pageId="7" pageNumber="278" refString="Jolivet, P. &amp; Verma, K. K. (2008 a) Eumolpinae-a widely distributed and much diversified subfamily of leaf beetles (Coleoptera, Chrysomelidae). Terrestrial Arthropod Reviews, 1, 3 - 37." type="journal article" year="2008" yearSuffix="a">
Jolivet 
<emphasis id="B926EAB9FF81C30BFB2AF45EFB0BBE6A" box="[1224,1273,295,320]" italics="true" pageId="7" pageNumber="278">et al</emphasis>
. 2008a
</bibRefCitation>
, 
<bibRefCitation id="EFC34B5AFF81C30BFAB7F450FA91BE6A" author="Jolivet" box="[1365,1379,295,320]" pageId="7" pageNumber="278" refString="Jolivet, P. &amp; Verma, K. K. (2008 b) On the origin of the chrysomelid fauna of New Caledonia. In: Jolivet, P., Santiago-Blay, J. &amp; Schmitt, M. (Eds.), Research on Chrysomelidae. Uol. 1. Brill Publishers, Leiden, pp. 30 8 - 319." type="journal article" year="2008" yearSuffix="b">b</bibRefCitation>
) and require a complete phylogenetic backbone to understand the evolution of the group.
</materialsCitation>
</paragraph>
<paragraph id="8BED36ABFF81C30BFF25F418FCABBD17" blockId="7.[151,1436,151,573]" pageId="7" pageNumber="278">
<taxonomicName id="4C524D28FF81C30BFF25F418FE87BEA2" authority="Nadein &amp; Leschen, 2017" authorityName="Nadein &amp; Leschen" authorityYear="2017" box="[199,373,367,392]" class="Insecta" family="Chrysomelidae" genus="Tyrannomolpus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B926EAB9FF81C30BFF25F418FE87BEA2" box="[199,373,367,392]" italics="true" pageId="7" pageNumber="278">Tyrannomolpus</emphasis>
</taxonomicName>
is known by only by a few specimens, and the single specimen with plant host data indicate it was collected at night on 
<taxonomicName id="4C524D28FF81C30BFE5EF4E2FD88BE87" authorityName="Seem" authorityYear="1862" box="[444,634,404,429]" class="Magnoliopsida" family="Araliaceae" genus="Meryta" kingdom="Plantae" order="Apiales" pageId="7" pageNumber="278" phylum="Tracheophyta" rank="species" species="sinclairii">
<emphasis id="B926EAB9FF81C30BFE5EF4E2FD88BE87" box="[444,634,404,429]" italics="true" pageId="7" pageNumber="278">Meryta sinclairii</emphasis>
</taxonomicName>
, referred to as Puka, a tree that is endemic to the Three Kings Islands and now planted commonly in gardens on the mainland. Despite further collecting to the islands, including a short visit by the second author in 2008 to Great Island with the intention to re-collect this and other rare endemic species (see 
<bibRefCitation id="EFC34B5AFF81C30BFF29F488FE1FBD32" author="Buckley" box="[203,493,511,536]" pageId="7" pageNumber="278" refString="Buckley, T. B. &amp; Leschen, R. A. B. (2013) Comparative phylogenetic analysis reveals long-term isolation of lineages on the Three Kings Islands, New Zealand. Biological Journal of the Linnaean Society, 106, 361 - 377. https: // doi. org / 10.1111 / j. 1095 - 8312.2012.02009. x" type="journal article" year="2013">Buckley &amp; Leschen 2013</bibRefCitation>
), there are no other specimens we know in existence. Additional surveys are needed to determine whether the species is still thriving on the islands.
</paragraph>
</subSubSection>
</treatment>
</document>