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<document id="F7447CE15CD9DD573630434DF70D4475" ID-CLB-Dataset="55596" ID-DOI="10.1093/zoolinnean/zlab113" ID-GBIF-Dataset="35d20d55-5bc0-405e-a04b-53c6a06d4df5" ID-ISSN="0024-4082" ID-Zenodo-Dep="6530795" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652084785049" checkinUser="plazi" docAuthor="Rotatori, Filippo Maria, Moreno-Azanza, Miguel &amp; Mateus, Octávio" docDate="2022" docId="03FB87A0EB6E6411FF4FFCB6B4D2FDCF" docLanguage="en" docName="zlab113.pdf" docOrigin="Zoological Journal of the Linnean Society 195 (1)" docSource="https://academic.oup.com/zoolinnean/article/195/1/125/6535466" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Draconyx loureiroi Mateus &amp; Antunes 2001" docType="treatment" docVersion="6" lastPageNumber="143" masterDocId="FFC2FFD8EB6A6403FFE6FFFBB01CFFC2" masterDocTitle="Reappraisal and new material of the holotype of Draconyx loureiroi (Ornithischia: Iguanodontia) provide insights on the tempo and modo of evolution of thumb-spiked dinosaurs" masterLastPageNumber="156" masterPageNumber="125" pageNumber="129" updateTime="1699329958474" updateUser="ExternalLinkService">
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<mods:title id="DB4FB2FE85721A0AC0207FBCD6757E62">Reappraisal and new material of the holotype of Draconyx loureiroi (Ornithischia: Iguanodontia) provide insights on the tempo and modo of evolution of thumb-spiked dinosaurs</mods:title>
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<mods:namePart id="66C6A57F8060844AF87062B34588E460">Rotatori, Filippo Maria</mods:namePart>
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<mods:namePart id="9B3D993EDFFE0173D2A2A8CB20AAE5AC">Moreno-Azanza, Miguel</mods:namePart>
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<mods:namePart id="81DD280EF97362544E90DCDE0790CC80">Mateus, Octávio</mods:namePart>
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<treatment id="03FB87A0EB6E6411FF4FFCB6B4D2FDCF" ID-DOI="http://doi.org/10.5281/zenodo.6536743" ID-GBIF-Taxon="195298191" ID-Zenodo-Dep="6536743" LSID="urn:lsid:plazi:treatment:03FB87A0EB6E6411FF4FFCB6B4D2FDCF" httpUri="http://treatment.plazi.org/id/03FB87A0EB6E6411FF4FFCB6B4D2FDCF" lastPageId="18" lastPageNumber="143" pageId="4" pageNumber="129">
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<paragraph id="8BED36B6EB6E6407FF4FFCB6B2FCFCA4" blockId="4.[169,736,845,870]" box="[169,736,845,870]" pageId="4" pageNumber="129">
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<taxonomicName id="4C524D35EB6E6407FF4FFCB6B2FCFCA4" authority="MATEUS &amp; ANTUNES, 2001" authorityName="Mateus &amp; Antunes" authorityYear="2001" box="[169,736,845,870]" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="126" phylum="Chordata" rank="species" species="loureiroi">
<emphasis id="B926EAA4EB6E6407FF4FFCB6B1BAFCA6" box="[169,422,845,868]" italics="true" pageId="4" pageNumber="129">DRACONYX LOUREIROI</emphasis>
<bibRefCitation id="EFC34B47EB6E6407FE4BFCB6B2FCFCA4" author="Mateus O &amp; Antunes MT" box="[429,736,845,870]" pageId="4" pageNumber="129" pagination="61 - 73" refId="ref17993" refString="Mateus O, Antunes MT. 2001. Draconyx loureiroi, a new camptosauridae (Dinosauria, Ornithopoda) from the Late Jurassic of Lourinha, Portugal. Annales de Paleontologie 87: 61 - 73." type="journal article" year="2001">MATEUS &amp; ANTUNES, 2001</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C348653DEB6E6407FE6FFC88B1CCFB05" pageId="4" pageNumber="129" type="description">
<paragraph id="8BED36B6EB6E6407FE6FFC88B21CFC4E" blockId="4.[393,512,883,908]" box="[393,512,883,908]" pageId="4" pageNumber="129">
(
<figureCitation id="13692A33EB6E6407FE77FC88B1E4FC4E" box="[401,504,883,908]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStart-3="Figure 5" captionStart-4="Figure 6" captionStart-5="Figure 7" captionStartId-0="8.[146,225,1610,1632]" captionStartId-1="9.[163,242,1778,1800]" captionStartId-2="10.[145,223,1463,1485]" captionStartId-3="11.[163,244,1604,1626]" captionStartId-4="12.[145,224,1096,1118]" captionStartId-5="13.[163,242,1760,1782]" captionTargetBox-0="[353,1217,199,1567]" captionTargetBox-1="[227,1379,195,1738]" captionTargetBox-2="[145,1425,195,1423]" captionTargetBox-3="[166,1446,198,1561]" captionTargetBox-4="[145,1425,195,1056]" captionTargetBox-5="[168,1441,1049,1715]" captionTargetId-0="figure-134@8.[350,1220,195,1570]" captionTargetId-1="figure-27@9.[227,1379,195,1738]" captionTargetId-2="figure-26@10.[145,1425,195,1423]" captionTargetId-3="figure-26@11.[163,1449,195,1564]" captionTargetId-4="figure-432@12.[145,1425,195,1056]" captionTargetId-5="figure-480@13.[163,1449,1046,1721]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="12" captionTargetPageId-5="13" captionText-0="Figure 2. Two maxillary teeth of Draconyx loureiroi holotype ML 357 in labial and lingual views. Completely preserved maxillary in labial (A) and lingual (C) views; isolated maxillary crown in labial (B) and lingual (D) views.Abbreviations: ne: neck, os: occlusal surface, pr: primary ridge, tr: tertiary ridges." captionText-1="Figure 3. General overview of Draconyx loureiroi holotype ML 357 post-cranial skeleton. Caudal vertebrae (K-M) in left lateral view. Right femur (AE) in frontal (A), medial (B), caudal (C), lateral (D) and distal (E) views. Right tibia in medial (F) and lateral (G) views. Proximal end of the fibula in medial (I) and lateral (J) views. Right articulated pes in dorsal (N) and medial (Q) view. Pedal digit III (O) and pedal digit II (P) in dorsal view." captionText-2="Figure 4. Three dimensional models of axial skeleton of Draconyx loureiroi holotype ML 357 (AF):Caudal n.2 ML 357-10 in cranial view (A), left lateral view (B), caudal view (C), right lateral view (D), ventral view (E) and dorsal view (F). Caudal n.3 ML 357-11 (GL) in cranial view (G), left lateral view (H), caudal view (I), right lateral view (J), ventral view (K) and dorsal view (L). Caudal n.1 ML 357-9 (M-R): cranial view (M), left lateral view (N), caudal view (O), right lateral view (P), ventral view (Q) and dorsal view (R).Abbreviations: cf: chevron facet, k: keel, nc: neural canal, ns: neural spine, tp: transverse process." captionText-3="Figure 5. Manual elements of Draconyx loureiroi holotype ML 357 (ML 357 1-5, 20-26), including articulated partial carpus, manual phalanges and unguals (AL). Articulated partial carpus (ML 357 5) in extensor view (A), right phalanx n.1(ML 357 4) in dorsal view (B), lateral view (B1), proximal view (B2) and distal view (B3), left phalanx n.2 (ML 357 22) in dorsal view (C), lateral view (C1), proximal view (C2) and distal view (C3), left phalanx n.3 (ML 357 21) in dorsal view (D), lateral view (D1), proximal view (D2) and distal view (D3), left phalanx n.4 (ML 357 24) in dorsal view (E), lateral view (E1), proximal view (E2) and distal view (E3), isolated distal ends of tarsals (ML 357 25,26) in extensor view (F, G), left ungual phalanx n.1 in dorsal view (D), mediolateral view (D1) and proximal view (D2), left ungual phalanx n.1 (ML 357- 1) in dorsal view (H), mediolateral view (H1) and proximal view (H2), right ungual phalanx n.2 (ML 357 2) in dorsal view (I), mediolateral view (I1) and proximal view (I2), right ungual phalanx n.3 (ML 357 3) in dorsal view (J), mediolateral view (J1) and proximal view (J2), left ungual phalanx n.4 in dorsal view (K), mediolateral view (K1) and proximal view (K2), ungual phalanx n.5 (ML 357 20) in dorsal view (L), mediolateral view (L1) and proximal view (L2)." captionText-4="Figure 6. Three dimensional models of limb bones elements of Draconyx loureiroi holotype ML 357 (ML 357 6-8), femur (AE) and tibia and proximal end of fibula (FJ). Femur (ML 357 6) in cranial view (A), medial view (B), caudal view (C), lateral view (D) and distal view (E). Tibia (ML 357 7) and proximal end of fibula (ML 357 8) in cranial view (F), medial view (G), caudal view (H), lateral (I) and proximal (J) views." captionText-5="Figure 7. Comparison of distal sections of femoral shaft from selected iguanodontians. A, Draconyx loureiroi, B, Uteodon aphanoecetes, C, Cumnoria prestwichii, D, Camptosaurus dispar, E, Barilium dawsoni, F, Mantellisaurus atherfieldensis and G, Iguanodon galvensis. Abbreviations: cod: condylid, eg: extensor groove, fg: flexor groove, lc: lateral condyle, mc: medial condyle. Distal sections drawn from: Carpenter &amp; Wilson, 2008; Galton &amp; Powell, 1980; Norman, 2011; Norman, 1986; Verdù et al., 2018." figureDoi-0="http://doi.org/10.5281/zenodo.6530801" figureDoi-1="http://doi.org/10.5281/zenodo.6530803" figureDoi-2="http://doi.org/10.5281/zenodo.6530805" figureDoi-3="http://doi.org/10.5281/zenodo.6530807" figureDoi-4="http://doi.org/10.5281/zenodo.6530809" figureDoi-5="http://doi.org/10.5281/zenodo.6530811" httpUri-0="https://zenodo.org/record/6530801/files/figure.png" httpUri-1="https://zenodo.org/record/6530803/files/figure.png" httpUri-2="https://zenodo.org/record/6530805/files/figure.png" httpUri-3="https://zenodo.org/record/6530807/files/figure.png" httpUri-4="https://zenodo.org/record/6530809/files/figure.png" httpUri-5="https://zenodo.org/record/6530811/files/figure.png" pageId="4" pageNumber="129">FIGS 27</figureCitation>
)
</paragraph>
<paragraph id="8BED36B6EB6E6407FF77FC46B1CCFB05" blockId="4.[145,761,956,1223]" pageId="4" pageNumber="129">
<emphasis id="B926EAA4EB6E6407FF77FC46B12FFC13" box="[145,307,956,978]" italics="true" pageId="4" pageNumber="129">Type material:</emphasis>
<materialsCitation id="3B3A3CEBEB6E6407FED9FC47B210FC32" ID-GBIF-Occurrence="3765335301" collectionCode="ML" country="Portugal" county="Mya" location="Vale de Frades" municipality="lower Tithonian" pageId="4" pageNumber="129" specimenCode="ML 357" specimenCount="1" typeStatus="holotype">
The
<typeStatus id="54E98814EB6E6407FE96FC47B1CDFC10" box="[368,465,956,978]" pageId="4" pageNumber="129" type="holotype">holotype</typeStatus>
specimen (newly reported material marked with *),
<specimenCode id="DBF49ECDEB6E6407FE52FC20B210FC32" box="[436,524,986,1008]" collectionCode="ML" country="France" name="Musee de Lectoure" pageId="4" pageNumber="129">ML 357</specimenCode>
</materialsCitation>
(subnumbers from 1 to 31) includes two maxillary teeth, carpal bones, two metacarpal distal ends*, three left carpal phalanges, one right carpal phalanx*, two left unguals, three right unguals* and one right leg including proximal femoral epiphysis, proximal and distal epiphysis of the tibia, astragalus, calcaneum, four metatarsals, five phalanges and two unguals.
</paragraph>
</subSubSection>
<subSubSection id="C348653DEB6E6407FF77FB0BB21DFAA0" pageId="4" pageNumber="129" type="reference_group">
<paragraph id="8BED36B6EB6E6407FF77FB0BB21DFAA0" blockId="4.[145,761,1264,1378]" pageId="4" pageNumber="129">
<emphasis id="B926EAA4EB6E6407FF77FB0BB178FAC7" box="[145,356,1264,1285]" italics="true" pageId="4" pageNumber="129">Referred specimen:</emphasis>
An isolated left femur,
<specimenCode id="DBF49ECDEB6E6407FD8FFB0BB2A3FAC7" box="[617,703,1264,1285]" collectionCode="ML" country="France" name="Musee de Lectoure" pageId="4" pageNumber="129">ML 434</specimenCode>
from
<quantity id="4CAA9B53EB6E6407FF77FAF4B0D7FAE6" box="[145,203,1294,1316]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="129" unit="km" value="1.0">1 km</quantity>
south of the
<typeStatus id="54E98814EB6E6407FE83FAF4B18BFAE6" box="[357,407,1295,1316]" pageId="4" pageNumber="129">type</typeStatus>
locality previously referred to
<taxonomicName id="4C524D35EB6E6407FF77FAD5B174FA80" authorityName="Mateus &amp; Antunes" authorityYear="2001" box="[145,360,1325,1347]" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="129" phylum="Chordata" rank="species" species="loureiroi">
<emphasis id="B926EAA4EB6E6407FF77FAD5B174FA80" box="[145,360,1325,1347]" italics="true" pageId="4" pageNumber="129">Draconyx loureiroi</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC34B47EB6E6407FE74FAD5B2A1FA81" author="Mateus O &amp; Antunes MT" box="[402,701,1325,1347]" pageId="4" pageNumber="129" pagination="61 - 73" refId="ref17993" refString="Mateus O, Antunes MT. 2001. Draconyx loureiroi, a new camptosauridae (Dinosauria, Ornithopoda) from the Late Jurassic of Lourinha, Portugal. Annales de Paleontologie 87: 61 - 73." type="journal article" year="2001">Mateus &amp; Antunes (2001)</bibRefCitation>
, now referred to
<emphasis id="B926EAA4EB6E6407FEF7FAB7B1A5FAA3" box="[273,441,1356,1377]" italics="true" pageId="4" pageNumber="129">Ankylopollexia</emphasis>
indet.
</paragraph>
</subSubSection>
<subSubSection id="C348653DEB6E6407FF77FA70B2E6FA1F" pageId="4" pageNumber="129" type="materials_examined">
<paragraph id="8BED36B6EB6E6407FF77FA70B2E6FA1F" blockId="4.[145,762,1418,1502]" pageId="4" pageNumber="129">
<emphasis id="B926EAA4EB6E6407FF77FA70B206FA62" box="[145,538,1418,1440]" italics="true" pageId="4" pageNumber="129">
<typeStatus id="54E98814EB6E6407FF77FA70B0D6FA62" box="[145,202,1419,1440]" pageId="4" pageNumber="129">Type</typeStatus>
locality, horizon and age:
</emphasis>
<location id="8E8D606DEB6E6407FDD5FA71B2EEFA62" LSID="urn:lsid:plazi:treatment:03FB87A0EB6E6411FF4FFCB6B4D2FDCF:8E8D606DEB6E6407FDD5FA71B2EEFA62" box="[563,754,1418,1440]" country="Portugal" county="Mya" municipality="lower Tithonian" name="Vale de Frades" pageId="4" pageNumber="129">Vale de Frades</location>
,
<location id="8E8D606DEB6E6407FF77FA52B184FA7D" LSID="urn:lsid:plazi:treatment:03FB87A0EB6E6411FF4FFCB6B4D2FDCF:8E8D606DEB6E6407FF77FA52B184FA7D" box="[145,408,1449,1471]" country="Portugal" county="Mya" municipality="lower Tithonian" name="Lourinha Municipality" pageId="4" pageNumber="129">Lourinhã Municipality</location>
,
<collectingCountry id="F3457626EB6E6407FE42FA52B217FA7D" box="[420,523,1449,1471]" name="Portugal" pageId="4" pageNumber="129">Portugal</collectingCountry>
.
<location id="8E8D606DEB6E6407FDF0FA52B18FFA1C" LSID="urn:lsid:plazi:treatment:03FB87A0EB6E6411FF4FFCB6B4D2FDCF:8E8D606DEB6E6407FDF0FA52B18FFA1C" country="Portugal" county="Mya" municipality="lower Tithonian" name="Praia Azul Member of Lourinha Formation" pageId="4" pageNumber="129">Praia Azul Member of Lourinhã Formation</location>
,
<collectingMunicipality id="6B89ACCCEB6E6407FE7AFA33B24EFA1C" box="[412,594,1480,1502]" pageId="4" pageNumber="129">lower Tithonian</collectingMunicipality>
, 151152
<collectingCounty id="628C4E3AEB6E6407FD27FA33B2EBFA1F" box="[705,759,1480,1501]" pageId="4" pageNumber="129">Mya</collectingCounty>
.
</paragraph>
</subSubSection>
<subSubSection id="C348653DEB6E6407FF77F9FDB2CFF88D" pageId="4" pageNumber="129" type="diagnosis">
<paragraph id="8BED36B6EB6E6407FF77F9FDB2CFF88D" blockId="4.[145,761,1542,1871]" pageId="4" pageNumber="129">
<emphasis id="B926EAA4EB6E6407FF77F9FDB161F9D9" box="[145,381,1542,1563]" italics="true" pageId="4" pageNumber="129">Emended diagnosis:</emphasis>
Styracosternan iguanodontian distinguished from other basal iguanodontians by the following combination of characters: unfused and noninterlocked carpus; absence of a sharp crest running from the medial condyle of the femur towards of the lesser trochanter, fully open U-shaped extensor groove on distal epiphysis of the femur; fully open V-shaped flexor groove without overhangs on distal epiphysis of the femur; concave medial margin of proximal epiphysis of the tibia; caudally pointing fibular condyle of the tibia; and a splint-like metatarsal I.
</paragraph>
</subSubSection>
<subSubSection id="C348653DEB6E6407FCCFFF3DB598FCCB" pageId="4" pageNumber="129" type="description">
<paragraph id="8BED36B6EB6E6407FCCFFF3DB492FF1F" blockId="4.[809,1166,197,221]" box="[809,1166,197,221]" pageId="4" pageNumber="129">
<emphasis id="B926EAA4EB6E6407FCCFFF3DB492FF1F" box="[809,1166,197,221]" italics="true" pageId="4" pageNumber="129">Description and comparisons</emphasis>
</paragraph>
<paragraph id="8BED36B6EB6E6407FCCFFF10B598FCCB" blockId="4.[809,1426,235,778]" pageId="4" pageNumber="129">
<emphasis id="B926EAA4EB6E6407FCCFFF10B366FEDD" italics="true" pageId="4" pageNumber="129">
Cranial material: Maxillary teeth (ML 35731) (
<figureCitation id="13692A33EB6E6407FA83FF10B376FEDD" captionStart="Figure 2" captionStartId="8.[146,225,1610,1632]" captionTargetBox="[353,1217,199,1567]" captionTargetId="figure-134@8.[350,1220,195,1570]" captionTargetPageId="8" captionText="Figure 2. Two maxillary teeth of Draconyx loureiroi holotype ML 357 in labial and lingual views. Completely preserved maxillary in labial (A) and lingual (C) views; isolated maxillary crown in labial (B) and lingual (D) views.Abbreviations: ne: neck, os: occlusal surface, pr: primary ridge, tr: tertiary ridges." figureDoi="http://doi.org/10.5281/zenodo.6530801" httpUri="https://zenodo.org/record/6530801/files/figure.png" pageId="4" pageNumber="129">Fig. 2A, B</figureCitation>
):
</emphasis>
One of the maxillary teeth preserves part of the root and its crown is complete (
<figureCitation id="13692A33EB6E6407FB5DFED3B511FEFF" box="[1211,1293,296,318]" captionStart="Figure 2" captionStartId="8.[146,225,1610,1632]" captionTargetBox="[353,1217,199,1567]" captionTargetId="figure-134@8.[350,1220,195,1570]" captionTargetPageId="8" captionText="Figure 2. Two maxillary teeth of Draconyx loureiroi holotype ML 357 in labial and lingual views. Completely preserved maxillary in labial (A) and lingual (C) views; isolated maxillary crown in labial (B) and lingual (D) views.Abbreviations: ne: neck, os: occlusal surface, pr: primary ridge, tr: tertiary ridges." figureDoi="http://doi.org/10.5281/zenodo.6530801" httpUri="https://zenodo.org/record/6530801/files/figure.png" pageId="4" pageNumber="129">Fig. 2A</figureCitation>
). The other tooth is just an isolated crown (
<figureCitation id="13692A33EB6E6407FB61FEBCB4CBFE9E" box="[1159,1239,327,349]" captionStart="Figure 2" captionStartId="8.[146,225,1610,1632]" captionTargetBox="[353,1217,199,1567]" captionTargetId="figure-134@8.[350,1220,195,1570]" captionTargetPageId="8" captionText="Figure 2. Two maxillary teeth of Draconyx loureiroi holotype ML 357 in labial and lingual views. Completely preserved maxillary in labial (A) and lingual (C) views; isolated maxillary crown in labial (B) and lingual (D) views.Abbreviations: ne: neck, os: occlusal surface, pr: primary ridge, tr: tertiary ridges." figureDoi="http://doi.org/10.5281/zenodo.6530801" httpUri="https://zenodo.org/record/6530801/files/figure.png" pageId="4" pageNumber="129">Fig. 2B</figureCitation>
). The specimens appear to have suffered some erosion and post-mortem breakage. The root is slightly labiolingually curved and tapers smoothly into the crown (
<figureCitation id="13692A33EB6E6407FB07FE58B529FE7A" box="[1249,1333,419,441]" captionStart="Figure 2" captionStartId="8.[146,225,1610,1632]" captionTargetBox="[353,1217,199,1567]" captionTargetId="figure-134@8.[350,1220,195,1570]" captionTargetPageId="8" captionText="Figure 2. Two maxillary teeth of Draconyx loureiroi holotype ML 357 in labial and lingual views. Completely preserved maxillary in labial (A) and lingual (C) views; isolated maxillary crown in labial (B) and lingual (D) views.Abbreviations: ne: neck, os: occlusal surface, pr: primary ridge, tr: tertiary ridges." figureDoi="http://doi.org/10.5281/zenodo.6530801" httpUri="https://zenodo.org/record/6530801/files/figure.png" pageId="4" pageNumber="129">Fig. 2A</figureCitation>
). There is no cingulum at the junction between the root and the crown. Overall, the crown is leaf-shaped and the veneer of enamel is thicker on the labial side (
<figureCitation id="13692A33EB6E6407FAD4FE04B59EFDD6" box="[1330,1410,511,533]" captionStart="Figure 2" captionStartId="8.[146,225,1610,1632]" captionTargetBox="[353,1217,199,1567]" captionTargetId="figure-134@8.[350,1220,195,1570]" captionTargetPageId="8" captionText="Figure 2. Two maxillary teeth of Draconyx loureiroi holotype ML 357 in labial and lingual views. Completely preserved maxillary in labial (A) and lingual (C) views; isolated maxillary crown in labial (B) and lingual (D) views.Abbreviations: ne: neck, os: occlusal surface, pr: primary ridge, tr: tertiary ridges." figureDoi="http://doi.org/10.5281/zenodo.6530801" httpUri="https://zenodo.org/record/6530801/files/figure.png" pageId="4" pageNumber="129">Fig. 2A</figureCitation>
). Labially a thick primary ridge is distally offset and five accessory ridges are present on the mesial surface (
<figureCitation id="13692A33EB6E6407FCD7FDA0B3BEFDB2" box="[817,930,603,625]" captionStart="Figure 2" captionStartId="8.[146,225,1610,1632]" captionTargetBox="[353,1217,199,1567]" captionTargetId="figure-134@8.[350,1220,195,1570]" captionTargetPageId="8" captionText="Figure 2. Two maxillary teeth of Draconyx loureiroi holotype ML 357 in labial and lingual views. Completely preserved maxillary in labial (A) and lingual (C) views; isolated maxillary crown in labial (B) and lingual (D) views.Abbreviations: ne: neck, os: occlusal surface, pr: primary ridge, tr: tertiary ridges." figureDoi="http://doi.org/10.5281/zenodo.6530801" httpUri="https://zenodo.org/record/6530801/files/figure.png" pageId="4" pageNumber="129">Fig. 2A, B</figureCitation>
). Non-mammillated hook-like denticles are coarsely present on the mesial crown margin, while distally they appear to have been obliterated by either erosion or occlusal wear. An extensive occlusal surface develops on the apex of the crown and is inclined labiolingually approximately around 30° (
<figureCitation id="13692A33EB6E6407FAE3FD0FB568FCCB" box="[1285,1396,756,778]" captionStart="Figure 2" captionStartId="8.[146,225,1610,1632]" captionTargetBox="[353,1217,199,1567]" captionTargetId="figure-134@8.[350,1220,195,1570]" captionTargetPageId="8" captionText="Figure 2. Two maxillary teeth of Draconyx loureiroi holotype ML 357 in labial and lingual views. Completely preserved maxillary in labial (A) and lingual (C) views; isolated maxillary crown in labial (B) and lingual (D) views.Abbreviations: ne: neck, os: occlusal surface, pr: primary ridge, tr: tertiary ridges." figureDoi="http://doi.org/10.5281/zenodo.6530801" httpUri="https://zenodo.org/record/6530801/files/figure.png" pageId="4" pageNumber="129">Fig. 2A, B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C348653DEB6E6411FCCFFCCEB4D2FDCF" lastPageId="18" lastPageNumber="143" pageId="4" pageNumber="129" type="discussion">
<paragraph id="8BED36B6EB6E6407FCCFFCCEB528FB18" blockId="4.[809,1426,821,1242]" pageId="4" pageNumber="129">
<emphasis id="B926EAA4EB6E6407FCCFFCCEB38BFC88" box="[809,919,821,842]" italics="true" pageId="4" pageNumber="129">Remarks:</emphasis>
The tapering root, slightly labiolingually recurved and the leaf-shaped crown are common characteristics of Dryomorpha (
<bibRefCitation id="EFC34B47EB6E6407FB4EFC88B55EFC4A" author="Galton PM" box="[1192,1346,883,905]" pageId="4" pageNumber="129" pagination="207 - 243" refId="ref17077" refString="Galton PM. 1983. The cranial anatomy of Dryosaurus, a hypsilophodontid dinosaur from the Upper Jurassic of North America and East Africa, with a review of hypsilophodontids from the Upper Jurassic of North America. Geologica et Palaeontologica 17: 207 - 243." type="journal article" year="1983">Galton, 1983</bibRefCitation>
, 2006;
<bibRefCitation id="EFC34B47EB6E6407FCCFFC69B3D0FC65" author="Norman DB" box="[809,972,913,935]" pageId="4" pageNumber="129" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">Norman, 1986</bibRefCitation>
, 2004). The maxillary crowns possess a distally offset primary ridge resembling the condition of ankylopollexians but differing from dryosaurids in which the primary ridge is located towards the centre of the crown (
<bibRefCitation id="EFC34B47EB6E6407FC9FFBF7B418FBE0" author="Galton PM" box="[889,1028,1036,1058]" pageId="4" pageNumber="129" pagination="17 - 47" refId="ref17124" refString="Galton PM. 2006. Teeth of ornithischian dinosaurs (mostly Ornithopoda) from the Morrison Formation (Upper Jurassic) of western United States. Horns and beaks. Ceratopsian and Ornithopod dinosaurs. Bloomington: Indiana University Press, 17 - 47." type="book chapter" year="2006">Galton, 2006</bibRefCitation>
). In the original description,
<bibRefCitation id="EFC34B47EB6E6407FAD8FBF7B3FBFB83" author="Mateus O &amp; Antunes MT" pageId="4" pageNumber="129" pagination="61 - 73" refId="ref17993" refString="Mateus O, Antunes MT. 2001. Draconyx loureiroi, a new camptosauridae (Dinosauria, Ornithopoda) from the Late Jurassic of Lourinha, Portugal. Annales de Paleontologie 87: 61 - 73." type="journal article" year="2001">Mateus &amp; Antunes (2001)</bibRefCitation>
indicated the presence of five accessory ridges in the distal half of the crown as an autapomorphy of
<emphasis id="B926EAA4EB6E6407FCA5FB93B40AFBBF" box="[835,1046,1128,1149]" italics="true" pageId="4" pageNumber="129">
<taxonomicName id="4C524D35EB6E6407FCA5FB93B40EFBBF" authorityName="Mateus &amp; Antunes" authorityYear="2001" box="[835,1042,1128,1149]" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="129" phylum="Chordata" rank="species" species="loureiroi">Draconyx loureiroi</taxonomicName>
.
</emphasis>
However, the number of accessory ridges is variable throughout the dental series (
<bibRefCitation id="EFC34B47EB6E6407FAD8FB7DB37DFB79" author="Galton PM" pageId="4" pageNumber="129" pagination="17 - 47" refId="ref17124" refString="Galton PM. 2006. Teeth of ornithischian dinosaurs (mostly Ornithopoda) from the Morrison Formation (Upper Jurassic) of western United States. Horns and beaks. Ceratopsian and Ornithopod dinosaurs. Bloomington: Indiana University Press, 17 - 47." type="book chapter" year="2006">Galton, 2006</bibRefCitation>
). Therefore, we regard it as a non-informative character if the tooth position is not articulated.
</paragraph>
<paragraph id="8BED36B6EB6E6404FCCFFAEBB561FEF5" blockId="4.[809,1425,1296,1870]" lastBlockId="7.[827,1444,197,311]" lastPageId="7" lastPageNumber="132" pageId="4" pageNumber="129">
<emphasis id="B926EAA4EB6E6407FCCFFAEBB3D1FA86" italics="true" pageId="4" pageNumber="129">
Axial skeleton: Caudal vertebrae (ML 357911) (
<figureCitation id="13692A33EB6E6407FA86FAEBB372FA86" captionStart="Figure 3" captionStartId="9.[163,242,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@9.[227,1379,195,1738]" captionTargetPageId="9" captionText="Figure 3. General overview of Draconyx loureiroi holotype ML 357 post-cranial skeleton. Caudal vertebrae (K-M) in left lateral view. Right femur (AE) in frontal (A), medial (B), caudal (C), lateral (D) and distal (E) views. Right tibia in medial (F) and lateral (G) views. Proximal end of the fibula in medial (I) and lateral (J) views. Right articulated pes in dorsal (N) and medial (Q) view. Pedal digit III (O) and pedal digit II (P) in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6530803" httpUri="https://zenodo.org/record/6530803/files/figure.png" pageId="4" pageNumber="129">Figs 3KM</figureCitation>
,
<figureCitation id="13692A33EB6E6407FC98FAD4B3A0FA86" box="[894,956,1327,1348]" captionStart="Figure 4" captionStartId="10.[145,223,1463,1485]" captionTargetBox="[145,1425,195,1423]" captionTargetId="figure-26@10.[145,1425,195,1423]" captionTargetPageId="10" captionText="Figure 4. Three dimensional models of axial skeleton of Draconyx loureiroi holotype ML 357 (AF):Caudal n.2 ML 357-10 in cranial view (A), left lateral view (B), caudal view (C), right lateral view (D), ventral view (E) and dorsal view (F). Caudal n.3 ML 357-11 (GL) in cranial view (G), left lateral view (H), caudal view (I), right lateral view (J), ventral view (K) and dorsal view (L). Caudal n.1 ML 357-9 (M-R): cranial view (M), left lateral view (N), caudal view (O), right lateral view (P), ventral view (Q) and dorsal view (R).Abbreviations: cf: chevron facet, k: keel, nc: neural canal, ns: neural spine, tp: transverse process." figureDoi="http://doi.org/10.5281/zenodo.6530805" httpUri="https://zenodo.org/record/6530805/files/figure.png" pageId="4" pageNumber="129">4AF</figureCitation>
):
</emphasis>
Three proximal caudal vertebrae are preserved and, as for the rest of the skeleton, they have suffered breakage and erosion. The centra are stout and craniocaudally compressed. The largest centrum is ML 3579, while ML 35710 and ML 35711 are slightly smaller. The cranial and caudal surfaces are slightly amphicoelous, having a subelliptical, rounded shape (
<figureCitation id="13692A33EB6E6407FC67F9FEB49BF9D8" box="[897,1159,1541,1563]" captionStart="Figure 4" captionStartId="10.[145,223,1463,1485]" captionTargetBox="[145,1425,195,1423]" captionTargetId="figure-26@10.[145,1425,195,1423]" captionTargetPageId="10" captionText="Figure 4. Three dimensional models of axial skeleton of Draconyx loureiroi holotype ML 357 (AF):Caudal n.2 ML 357-10 in cranial view (A), left lateral view (B), caudal view (C), right lateral view (D), ventral view (E) and dorsal view (F). Caudal n.3 ML 357-11 (GL) in cranial view (G), left lateral view (H), caudal view (I), right lateral view (J), ventral view (K) and dorsal view (L). Caudal n.1 ML 357-9 (M-R): cranial view (M), left lateral view (N), caudal view (O), right lateral view (P), ventral view (Q) and dorsal view (R).Abbreviations: cf: chevron facet, k: keel, nc: neural canal, ns: neural spine, tp: transverse process." figureDoi="http://doi.org/10.5281/zenodo.6530805" httpUri="https://zenodo.org/record/6530805/files/figure.png" pageId="4" pageNumber="129">Fig. 4 A, C, G, I, M, O</figureCitation>
). Dorsally, the neural arch preserves the ventralmost portion of the neural spine but the neural suture is not visible. Immediately caudal to the spine, in the most complete specimens ML 357-9 and ML 357 - 10, (
<figureCitation id="13692A33EB6E6407FB6FF97BB51EF957" box="[1161,1282,1664,1686]" captionStart="Figure 4" captionStartId="10.[145,223,1463,1485]" captionTargetBox="[145,1425,195,1423]" captionTargetId="figure-26@10.[145,1425,195,1423]" captionTargetPageId="10" captionText="Figure 4. Three dimensional models of axial skeleton of Draconyx loureiroi holotype ML 357 (AF):Caudal n.2 ML 357-10 in cranial view (A), left lateral view (B), caudal view (C), right lateral view (D), ventral view (E) and dorsal view (F). Caudal n.3 ML 357-11 (GL) in cranial view (G), left lateral view (H), caudal view (I), right lateral view (J), ventral view (K) and dorsal view (L). Caudal n.1 ML 357-9 (M-R): cranial view (M), left lateral view (N), caudal view (O), right lateral view (P), ventral view (Q) and dorsal view (R).Abbreviations: cf: chevron facet, k: keel, nc: neural canal, ns: neural spine, tp: transverse process." figureDoi="http://doi.org/10.5281/zenodo.6530805" httpUri="https://zenodo.org/record/6530805/files/figure.png" pageId="4" pageNumber="129">Fig. 4C, O</figureCitation>
) two small, rounded and steeply inclined post-zygapophyses diverge laterally. On the lateral surface of the neural arch, the articulation for the transverse processes is overlain by a broken surface. The presence of this surface indicates that these vertebrae represent the cranialmost portion of the caudal series. In ventral view, the centra have an hourglass-shaped outline (
<figureCitation id="13692A33EB6D6404FA9EFF3EB387FF3B" captionStart="Figure 4" captionStartId="10.[145,223,1463,1485]" captionTargetBox="[145,1425,195,1423]" captionTargetId="figure-26@10.[145,1425,195,1423]" captionTargetPageId="10" captionText="Figure 4. Three dimensional models of axial skeleton of Draconyx loureiroi holotype ML 357 (AF):Caudal n.2 ML 357-10 in cranial view (A), left lateral view (B), caudal view (C), right lateral view (D), ventral view (E) and dorsal view (F). Caudal n.3 ML 357-11 (GL) in cranial view (G), left lateral view (H), caudal view (I), right lateral view (J), ventral view (K) and dorsal view (L). Caudal n.1 ML 357-9 (M-R): cranial view (M), left lateral view (N), caudal view (O), right lateral view (P), ventral view (Q) and dorsal view (R).Abbreviations: cf: chevron facet, k: keel, nc: neural canal, ns: neural spine, tp: transverse process." figureDoi="http://doi.org/10.5281/zenodo.6530805" httpUri="https://zenodo.org/record/6530805/files/figure.png" pageId="7" pageNumber="132">Fig. 4E, K, Q</figureCitation>
). The ventral surfaces of the centra possess a narrow keel and the margin is highly concave (
<figureCitation id="13692A33EB6D6404FAABFEF9B357FEF5" captionStart="Figure 4" captionStartId="10.[145,223,1463,1485]" captionTargetBox="[145,1425,195,1423]" captionTargetId="figure-26@10.[145,1425,195,1423]" captionTargetPageId="10" captionText="Figure 4. Three dimensional models of axial skeleton of Draconyx loureiroi holotype ML 357 (AF):Caudal n.2 ML 357-10 in cranial view (A), left lateral view (B), caudal view (C), right lateral view (D), ventral view (E) and dorsal view (F). Caudal n.3 ML 357-11 (GL) in cranial view (G), left lateral view (H), caudal view (I), right lateral view (J), ventral view (K) and dorsal view (L). Caudal n.1 ML 357-9 (M-R): cranial view (M), left lateral view (N), caudal view (O), right lateral view (P), ventral view (Q) and dorsal view (R).Abbreviations: cf: chevron facet, k: keel, nc: neural canal, ns: neural spine, tp: transverse process." figureDoi="http://doi.org/10.5281/zenodo.6530805" httpUri="https://zenodo.org/record/6530805/files/figure.png" pageId="7" pageNumber="132">Fig. 4B, P</figureCitation>
). Caudally, broad facets for chevrons are present.
</paragraph>
<caption id="DF2D663EEB6F6406FF6CFE2AB53CFE25" ID-Table-UUID="DF2D663EEB6F6406FF6CFE2AB53CFE25" box="[138,1312,465,487]" httpUri="http://table.plazi.org/id/DF2D663EEB6F6406FF6CFE2AB53CFE25" pageId="5" pageNumber="130" startId="5.[138,202,465,487]" targetBox="[138,1816,523,1645]" targetIsTable="true" targetPageId="5">
<paragraph id="8BED36B6EB6F6406FF6CFE2AB53CFE25" blockId="5.[138,1312,465,487]" box="[138,1312,465,487]" pageId="5" pageNumber="130">
<emphasis id="B926EAA4EB6F6406FF6CFE2AB0F8FE25" bold="true" box="[138,228,465,487]" pageId="5" pageNumber="130">Table 1.</emphasis>
Selected measurements of ML 357. All measurements are in mm (fragmented material marked with *)
</paragraph>
</caption>
<paragraph id="8BED36B6EB6F6406FE4BFDF0B0F0F9AF" pageId="5" pageNumber="130">
<table id="F952C416EB6F9BFCFF6CFDF0B704F9AF" box="[138,1816,523,1645]" gridcols="8" gridrows="38" pageId="5" pageNumber="130">
<tr id="356234F4EB6F9BFCFF6CFDF0B704FDE3" box="[138,1816,523,545]" gridrow="0" pageId="5" pageNumber="130" rowspan-0="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFE4BFDF0B228FDE3" box="[429,564,523,545]" gridcol="1" gridrow="0" pageId="5" pageNumber="130">Height</th>
<th id="76B35D88EB6F9BFCFD93FDF0B321FDE3" box="[629,829,523,545]" gridcol="2" gridrow="0" pageId="5" pageNumber="130">Mesiodistal width</th>
<th id="76B35D88EB6F9BFCFC87FDF0B434FDE3" box="[865,1064,523,545]" gridcol="3" gridrow="0" pageId="5" pageNumber="130">Labiolingual width</th>
</tr>
<tr id="356234F4EB6F9BFCFF6CFDD2B704FDFD" box="[138,1816,553,575]" gridrow="1" pageId="5" pageNumber="130" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFDD2B189FDFD" box="[138,405,553,575]" gridcol="0" gridrow="1" pageId="5" pageNumber="130">Complete maxillary tooth</th>
<td id="76B35D88EB6F9BFCFE4BFDD2B228FDFD" box="[429,564,553,575]" gridcol="1" gridrow="1" pageId="5" pageNumber="130">16</td>
<td id="76B35D88EB6F9BFCFD93FDD2B321FDFD" box="[629,829,553,575]" gridcol="2" gridrow="1" pageId="5" pageNumber="130">10</td>
<td id="76B35D88EB6F9BFCFC87FDD2B434FDFD" box="[865,1064,553,575]" gridcol="3" gridrow="1" pageId="5" pageNumber="130">6</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFDBCB704FD9F" box="[138,1816,583,605]" gridrow="2" pageId="5" pageNumber="130">
<th id="76B35D88EB6F9BFCFF6CFDBCB704FD9F" box="[138,1816,583,605]" colspan="8" colspanRight="7" gridcol="0" gridrow="2" pageId="5" pageNumber="130">ML 35731</th>
</tr>
<tr id="356234F4EB6F9BFCFF6CFD9EB704FDB9" box="[138,1816,613,635]" gridrow="3" pageId="5" pageNumber="130" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFD9EB189FDB9" box="[138,405,613,635]" gridcol="0" gridrow="3" pageId="5" pageNumber="130">Maxillary crown tooh</th>
<td id="76B35D88EB6F9BFCFE4BFD9EB228FDB9" box="[429,564,613,635]" gridcol="1" gridrow="3" pageId="5" pageNumber="130">11</td>
<td id="76B35D88EB6F9BFCFD93FD9EB321FDB9" box="[629,829,613,635]" gridcol="2" gridrow="3" pageId="5" pageNumber="130">9</td>
<td id="76B35D88EB6F9BFCFC87FD9EB434FDB9" box="[865,1064,613,635]" gridcol="3" gridrow="3" pageId="5" pageNumber="130">9</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFD79B704FD5A" box="[138,1816,642,664]" gridrow="4" pageId="5" pageNumber="130" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFD79B189FD5A" box="[138,405,642,664]" gridcol="0" gridrow="4" pageId="5" pageNumber="130">ML 35731</th>
</tr>
<tr id="356234F4EB6F9BFCFF6CFD5BB704FD74" box="[138,1816,672,694]" gridrow="5" pageId="5" pageNumber="130" rowspan-0="1">
<td id="76B35D88EB6F9BFCFE4BFD5BB228FD74" box="[429,564,672,694]" gridcol="1" gridrow="5" pageId="5" pageNumber="130">Total Length</td>
<td id="76B35D88EB6F9BFCFD93FD5BB321FD74" box="[629,829,672,694]" gridcol="2" gridrow="5" pageId="5" pageNumber="130">
<collectingRegion id="4996F854EB6F6406FD93FD5BB2CFFD74" box="[629,723,672,694]" country="France" name="Centre" pageId="5" pageNumber="130">Centrum</collectingRegion>
cranial
</td>
<td id="76B35D88EB6F9BFCFC87FD5BB434FD74" box="[865,1064,672,694]" gridcol="3" gridrow="5" pageId="5" pageNumber="130">
<collectingRegion id="4996F854EB6F6406FC84FD5BB3DCFD74" box="[866,960,672,694]" country="France" name="Centre" pageId="5" pageNumber="130">Centrum</collectingRegion>
cranial
</td>
<td id="76B35D88EB6F9BFCFBA2FD5BB4F2FD74" box="[1092,1262,672,694]" gridcol="4" gridrow="5" pageId="5" pageNumber="130">
<collectingRegion id="4996F854EB6F6406FBA2FD5BB4BEFD74" box="[1092,1186,672,694]" country="France" name="Centre" pageId="5" pageNumber="130">Centrum</collectingRegion>
caudal
</td>
<td id="76B35D88EB6F9BFCFAE2FD5BB5DEFD74" box="[1284,1474,672,694]" gridcol="5" gridrow="5" pageId="5" pageNumber="130">
<collectingRegion id="4996F854EB6F6406FAE2FD5BB57EFD74" box="[1284,1378,672,694]" country="France" name="Centre" pageId="5" pageNumber="130">Centrum</collectingRegion>
caudal
</td>
<td id="76B35D88EB6F9BFCFA32FD5BB640FD74" box="[1492,1628,672,694]" gridcol="6" gridrow="5" pageId="5" pageNumber="130">Neural spine</td>
<td id="76B35D88EB6F9BFCF99BFD5BB704FD74" box="[1661,1816,672,694]" gridcol="7" gridrow="5" pageId="5" pageNumber="130">Neural arch</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFD46B704FD11" box="[138,1816,701,723]" gridrow="6" pageId="5" pageNumber="130" rowspan-0="1" rowspan-1="1">
<td id="76B35D88EB6F9BFCFD93FD46B321FD11" box="[629,829,701,723]" gridcol="2" gridrow="6" pageId="5" pageNumber="130">mediolateral</td>
<td id="76B35D88EB6F9BFCFC87FD46B434FD11" box="[865,1064,701,723]" gridcol="3" gridrow="6" pageId="5" pageNumber="130">dorsoventral</td>
<td id="76B35D88EB6F9BFCFBA2FD46B4F2FD11" box="[1092,1262,701,723]" gridcol="4" gridrow="6" pageId="5" pageNumber="130">mediolateral</td>
<td id="76B35D88EB6F9BFCFAE2FD46B5DEFD11" box="[1284,1474,701,723]" gridcol="5" gridrow="6" pageId="5" pageNumber="130">dorsoventral</td>
<td id="76B35D88EB6F9BFCFA32FD46B640FD11" box="[1492,1628,701,723]" gridcol="6" gridrow="6" pageId="5" pageNumber="130">height</td>
<td id="76B35D88EB6F9BFCF99BFD46B704FD11" box="[1661,1816,701,723]" gridcol="7" gridrow="6" pageId="5" pageNumber="130">mediolateral</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFD20B704FD33" box="[138,1816,731,753]" gridrow="7" pageId="5" pageNumber="130" rowspan-0="1" rowspan-1="1" rowspan-6="1">
<td id="76B35D88EB6F9BFCFD93FD20B321FD33" box="[629,829,731,753]" gridcol="2" gridrow="7" pageId="5" pageNumber="130">width</td>
<td id="76B35D88EB6F9BFCFC87FD20B434FD33" box="[865,1064,731,753]" gridcol="3" gridrow="7" pageId="5" pageNumber="130">height</td>
<td id="76B35D88EB6F9BFCFBA2FD20B4F2FD33" box="[1092,1262,731,753]" gridcol="4" gridrow="7" pageId="5" pageNumber="130">width</td>
<td id="76B35D88EB6F9BFCFAE2FD20B5DEFD33" box="[1284,1474,731,753]" gridcol="5" gridrow="7" pageId="5" pageNumber="130">height</td>
<td id="76B35D88EB6F9BFCF99BFD20B704FD33" box="[1661,1816,731,753]" gridcol="7" gridrow="7" pageId="5" pageNumber="130">width</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFD02B704FCCD" box="[138,1816,761,783]" gridrow="8" pageId="5" pageNumber="130">
<th id="76B35D88EB6F9BFCFF6CFD02B189FCCD" box="[138,405,761,783]" gridcol="0" gridrow="8" pageId="5" pageNumber="130">Caudal vertebra n.1</th>
<td id="76B35D88EB6F9BFCFE4BFD02B228FCCD" box="[429,564,761,783]" gridcol="1" gridrow="8" pageId="5" pageNumber="130">59</td>
<td id="76B35D88EB6F9BFCFD93FD02B321FCCD" box="[629,829,761,783]" gridcol="2" gridrow="8" pageId="5" pageNumber="130">58</td>
<td id="76B35D88EB6F9BFCFC87FD02B434FCCD" box="[865,1064,761,783]" gridcol="3" gridrow="8" pageId="5" pageNumber="130">52</td>
<td id="76B35D88EB6F9BFCFBA2FD02B4F2FCCD" box="[1092,1262,761,783]" gridcol="4" gridrow="8" pageId="5" pageNumber="130">55</td>
<td id="76B35D88EB6F9BFCFAE2FD02B5DEFCCD" box="[1284,1474,761,783]" gridcol="5" gridrow="8" pageId="5" pageNumber="130">56</td>
<td id="76B35D88EB6F9BFCFA32FD02B640FCCD" box="[1492,1628,761,783]" gridcol="6" gridrow="8" pageId="5" pageNumber="130">56</td>
<td id="76B35D88EB6F9BFCF99BFD02B704FCCD" box="[1661,1816,761,783]" gridcol="7" gridrow="8" pageId="5" pageNumber="130">80</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFCEDB704FCEE" box="[138,1816,790,812]" gridrow="9" pageId="5" pageNumber="130" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFCEDB189FCEE" box="[138,405,790,812]" gridcol="0" gridrow="9" pageId="5" pageNumber="130">ML 3579</th>
</tr>
<tr id="356234F4EB6F9BFCFF6CFCCFB704FC88" box="[138,1816,820,842]" gridrow="10" pageId="5" pageNumber="130">
<th id="76B35D88EB6F9BFCFF6CFCCFB189FC88" box="[138,405,820,842]" gridcol="0" gridrow="10" pageId="5" pageNumber="130">Caudal vertebra n.2</th>
<td id="76B35D88EB6F9BFCFE4BFCCFB228FC88" box="[429,564,820,842]" gridcol="1" gridrow="10" pageId="5" pageNumber="130">47</td>
<td id="76B35D88EB6F9BFCFD93FCCFB321FC88" box="[629,829,820,842]" gridcol="2" gridrow="10" pageId="5" pageNumber="130">48</td>
<td id="76B35D88EB6F9BFCFC87FCCFB434FC88" box="[865,1064,820,842]" gridcol="3" gridrow="10" pageId="5" pageNumber="130">53</td>
<td id="76B35D88EB6F9BFCFBA2FCCFB4F2FC88" box="[1092,1262,820,842]" gridcol="4" gridrow="10" pageId="5" pageNumber="130">47</td>
<td id="76B35D88EB6F9BFCFAE2FCCFB5DEFC88" box="[1284,1474,820,842]" gridcol="5" gridrow="10" pageId="5" pageNumber="130">52</td>
<td id="76B35D88EB6F9BFCFA32FCCFB640FC88" box="[1492,1628,820,842]" gridcol="6" gridrow="10" pageId="5" pageNumber="130">45</td>
<td id="76B35D88EB6F9BFCF99BFCCFB704FC88" box="[1661,1816,820,842]" gridcol="7" gridrow="10" pageId="5" pageNumber="130">60</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFCA9B704FCAA" box="[138,1816,850,872]" gridrow="11" pageId="5" pageNumber="130">
<th id="76B35D88EB6F9BFCFF6CFCA9B704FCAA" box="[138,1816,850,872]" colspan="8" colspanRight="7" gridcol="0" gridrow="11" pageId="5" pageNumber="130">ML 35710</th>
</tr>
<tr id="356234F4EB6F9BFCFF6CFC8BB704FC44" box="[138,1816,880,902]" gridrow="12" pageId="5" pageNumber="130">
<th id="76B35D88EB6F9BFCFF6CFC8BB189FC44" box="[138,405,880,902]" gridcol="0" gridrow="12" pageId="5" pageNumber="130">Caudal vertebra n.3</th>
<td id="76B35D88EB6F9BFCFE4BFC8BB228FC44" box="[429,564,880,902]" gridcol="1" gridrow="12" pageId="5" pageNumber="130">47</td>
<td id="76B35D88EB6F9BFCFD93FC8BB321FC44" box="[629,829,880,902]" gridcol="2" gridrow="12" pageId="5" pageNumber="130">44</td>
<td id="76B35D88EB6F9BFCFC87FC8BB434FC44" box="[865,1064,880,902]" gridcol="3" gridrow="12" pageId="5" pageNumber="130">48</td>
<td id="76B35D88EB6F9BFCFBA2FC8BB4F2FC44" box="[1092,1262,880,902]" gridcol="4" gridrow="12" pageId="5" pageNumber="130">40</td>
<td id="76B35D88EB6F9BFCFAE2FC8BB5DEFC44" box="[1284,1474,880,902]" gridcol="5" gridrow="12" pageId="5" pageNumber="130">50</td>
<td id="76B35D88EB6F9BFCFA32FC8BB640FC44" box="[1492,1628,880,902]" gridcol="6" gridrow="12" pageId="5" pageNumber="130">45</td>
<td id="76B35D88EB6F9BFCF99BFC8BB704FC44" box="[1661,1816,880,902]" gridcol="7" gridrow="12" pageId="5" pageNumber="130">60</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFC76B704FC61" box="[138,1816,909,931]" gridrow="13" pageId="5" pageNumber="130" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFC76B189FC61" box="[138,405,909,931]" gridcol="0" gridrow="13" pageId="5" pageNumber="130">ML 35711</th>
</tr>
<tr id="356234F4EB6F9BFCFF6CFC50B704FC03" box="[138,1816,939,961]" gridrow="14" pageId="5" pageNumber="130" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFC50B189FC03" box="[138,405,939,961]" gridcol="0" gridrow="14" pageId="5" pageNumber="130">Articulated carpus</th>
<td id="76B35D88EB6F9BFCFE4BFC50B228FC03" box="[429,564,939,961]" gridcol="1" gridrow="14" pageId="5" pageNumber="130">Total Length</td>
<td id="76B35D88EB6F9BFCFD93FC50B321FC03" box="[629,829,939,961]" gridcol="2" gridrow="14" pageId="5" pageNumber="130">Mediolateral width</td>
<td id="76B35D88EB6F9BFCFC87FC50B4F2FC03" box="[865,1262,939,961]" colspan="2" colspanRight="1" gridcol="3" gridrow="14" pageId="5" pageNumber="130">Craniocaudal height</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFC32B704FC1D" box="[138,1816,969,991]" gridrow="15" pageId="5" pageNumber="130" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFC32B189FC1D" box="[138,405,969,991]" gridcol="0" gridrow="15" pageId="5" pageNumber="130">Metacarpal (fragment)</th>
<td id="76B35D88EB6F9BFCFE4BFC32B228FC1D" box="[429,564,969,991]" gridcol="1" gridrow="15" pageId="5" pageNumber="130">32</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFC1DB704FC3E" box="[138,1816,998,1020]" gridrow="16" pageId="5" pageNumber="130" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFC1DB189FC3E" box="[138,405,998,1020]" gridcol="0" gridrow="16" pageId="5" pageNumber="130">ML 3575</th>
</tr>
<tr id="356234F4EB6F9BFCFF6CFBFFB704FBD8" box="[138,1816,1028,1050]" gridrow="17" pageId="5" pageNumber="130" rowspan-1="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFBFFB189FBD8" box="[138,405,1028,1050]" gridcol="0" gridrow="17" pageId="5" pageNumber="130">
Carpal n.1
<specimenCode id="DBF49ECDEB6F6406FF19FBFFB175FBD8" box="[255,361,1028,1050]" collectionCode="ML" country="France" name="Musee de Lectoure" pageId="5" pageNumber="130">ML 3575</specimenCode>
</th>
<td id="76B35D88EB6F9BFCFD93FBFFB321FBD8" box="[629,829,1028,1050]" gridcol="2" gridrow="17" pageId="5" pageNumber="130">20</td>
<td id="76B35D88EB6F9BFCFC87FBFFB434FBD8" box="[865,1064,1028,1050]" gridcol="3" gridrow="17" pageId="5" pageNumber="130">22</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFBD9B704FBFA" box="[138,1816,1058,1080]" gridrow="18" pageId="5" pageNumber="130" rowspan-1="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFBD9B189FBFA" box="[138,405,1058,1080]" gridcol="0" gridrow="18" pageId="5" pageNumber="130">Carpal n.2 ML 3575</th>
<td id="76B35D88EB6F9BFCFD93FBD9B321FBFA" box="[629,829,1058,1080]" gridcol="2" gridrow="18" pageId="5" pageNumber="130">13</td>
<td id="76B35D88EB6F9BFCFC87FBD9B434FBFA" box="[865,1064,1058,1080]" gridcol="3" gridrow="18" pageId="5" pageNumber="130">13</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFBBBB704FB94" box="[138,1816,1088,1110]" gridrow="19" pageId="5" pageNumber="130" rowspan-1="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFBBBB189FB94" box="[138,405,1088,1110]" gridcol="0" gridrow="19" pageId="5" pageNumber="130">Carpal n.3 ML 3575</th>
<td id="76B35D88EB6F9BFCFD93FBBBB321FB94" box="[629,829,1088,1110]" gridcol="2" gridrow="19" pageId="5" pageNumber="130">14</td>
<td id="76B35D88EB6F9BFCFC87FBBBB434FB94" box="[865,1064,1088,1110]" gridcol="3" gridrow="19" pageId="5" pageNumber="130">18</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFBA5B704FBB6" box="[138,1816,1118,1140]" gridrow="20" pageId="5" pageNumber="130" rowspan-0="1" rowspan-1="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<td id="76B35D88EB6F9BFCFD93FBA5B321FBB6" box="[629,829,1118,1140]" gridcol="2" gridrow="20" pageId="5" pageNumber="130">Mediolateral distal</td>
<td id="76B35D88EB6F9BFCFC87FBA5B434FBB6" box="[865,1064,1118,1140]" gridcol="3" gridrow="20" pageId="5" pageNumber="130">Extensor-palmar</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFB87B704FB50" box="[138,1816,1148,1170]" gridrow="21" pageId="5" pageNumber="130" rowspan-0="1" rowspan-1="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<td id="76B35D88EB6F9BFCFD93FB87B321FB50" box="[629,829,1148,1170]" gridcol="2" gridrow="21" pageId="5" pageNumber="130">width</td>
<td id="76B35D88EB6F9BFCFC87FB87B434FB50" box="[865,1064,1148,1170]" gridcol="3" gridrow="21" pageId="5" pageNumber="130">distal height</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFB61B704FB72" box="[138,1816,1178,1200]" gridrow="22" pageId="5" pageNumber="130" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFB61B189FB72" box="[138,405,1178,1200]" gridcol="0" gridrow="22" pageId="5" pageNumber="130">Distal metacarpal 1 ML</th>
<td id="76B35D88EB6F9BFCFE4BFB61B228FB72" box="[429,564,1178,1200]" gridcol="1" gridrow="22" pageId="5" pageNumber="130">23</td>
<td id="76B35D88EB6F9BFCFD93FB61B321FB72" box="[629,829,1178,1200]" gridcol="2" gridrow="22" pageId="5" pageNumber="130">18</td>
<td id="76B35D88EB6F9BFCFC87FB61B434FB72" box="[865,1064,1178,1200]" gridcol="3" gridrow="22" pageId="5" pageNumber="130">6</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFB4CB704FB0F" box="[138,1816,1207,1229]" gridrow="23" pageId="5" pageNumber="130" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFB4CB189FB0F" box="[138,405,1207,1229]" gridcol="0" gridrow="23" pageId="5" pageNumber="130">35725</th>
</tr>
<tr id="356234F4EB6F9BFCFF6CFB2EB704FB29" box="[138,1816,1237,1259]" gridrow="24" pageId="5" pageNumber="130" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFB2EB189FB29" box="[138,405,1237,1259]" gridcol="0" gridrow="24" pageId="5" pageNumber="130">Distal metacarpal 2 ML</th>
<td id="76B35D88EB6F9BFCFE4BFB2EB228FB29" box="[429,564,1237,1259]" gridcol="1" gridrow="24" pageId="5" pageNumber="130">18</td>
<td id="76B35D88EB6F9BFCFD93FB2EB321FB29" box="[629,829,1237,1259]" gridcol="2" gridrow="24" pageId="5" pageNumber="130">18</td>
<td id="76B35D88EB6F9BFCFC87FB2EB434FB29" box="[865,1064,1237,1259]" gridcol="3" gridrow="24" pageId="5" pageNumber="130">16</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFB09B704FACA" box="[138,1816,1266,1288]" gridrow="25" pageId="5" pageNumber="130" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFB09B189FACA" box="[138,405,1266,1288]" gridcol="0" gridrow="25" pageId="5" pageNumber="130">35726</th>
</tr>
<tr id="356234F4EB6F9BFCFF6CFAEAB704FAE5" box="[138,1816,1297,1319]" gridrow="26" pageId="5" pageNumber="130" rowspan-1="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFAEAB189FAE5" box="[138,405,1297,1319]" gridcol="0" gridrow="26" pageId="5" pageNumber="130">Left manual elements</th>
<td id="76B35D88EB6F9BFCFD93FAEAB321FAE5" box="[629,829,1297,1319]" gridcol="2" gridrow="26" pageId="5" pageNumber="130">Mediolateral distal</td>
<td id="76B35D88EB6F9BFCFC87FAEAB434FAE5" box="[865,1064,1297,1319]" gridcol="3" gridrow="26" pageId="5" pageNumber="130">Mediolateral</td>
<td id="76B35D88EB6F9BFCFBA2FAEAB4F2FAE5" box="[1092,1262,1297,1319]" gridcol="4" gridrow="26" pageId="5" pageNumber="130">Dorsovental</td>
<td id="76B35D88EB6F9BFCFAE2FAEAB5DEFAE5" box="[1284,1474,1297,1319]" gridcol="5" gridrow="26" pageId="5" pageNumber="130">Dorsovental</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFAD5B704FA86" box="[138,1816,1326,1348]" gridrow="27" pageId="5" pageNumber="130" rowspan-0="1" rowspan-1="1" rowspan-6="1" rowspan-7="1">
<td id="76B35D88EB6F9BFCFD93FAD5B321FA86" box="[629,829,1326,1348]" gridcol="2" gridrow="27" pageId="5" pageNumber="130">width</td>
<td id="76B35D88EB6F9BFCFC87FAD5B434FA86" box="[865,1064,1326,1348]" gridcol="3" gridrow="27" pageId="5" pageNumber="130">proximal width</td>
<td id="76B35D88EB6F9BFCFBA2FAD5B4F2FA86" box="[1092,1262,1326,1348]" gridcol="4" gridrow="27" pageId="5" pageNumber="130">distal height</td>
<td id="76B35D88EB6F9BFCFAE2FAD5B5DEFA86" box="[1284,1474,1326,1348]" gridcol="5" gridrow="27" pageId="5" pageNumber="130">proximal height</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFAB7B704FAA0" box="[138,1816,1356,1378]" gridrow="28" pageId="5" pageNumber="130" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFAB7B189FAA0" box="[138,405,1356,1378]" gridcol="0" gridrow="28" pageId="5" pageNumber="130">Manual phalanx n.2 ML</th>
<td id="76B35D88EB6F9BFCFE4BFAB7B228FAA0" box="[429,564,1356,1378]" gridcol="1" gridrow="28" pageId="5" pageNumber="130">8</td>
<td id="76B35D88EB6F9BFCFD93FAB7B321FAA0" box="[629,829,1356,1378]" gridcol="2" gridrow="28" pageId="5" pageNumber="130">15</td>
<td id="76B35D88EB6F9BFCFC87FAB7B434FAA0" box="[865,1064,1356,1378]" gridcol="3" gridrow="28" pageId="5" pageNumber="130">18</td>
<td id="76B35D88EB6F9BFCFBA2FAB7B4F2FAA0" box="[1092,1262,1356,1378]" gridcol="4" gridrow="28" pageId="5" pageNumber="130">10</td>
<td id="76B35D88EB6F9BFCFAE2FAB7B5DEFAA0" box="[1284,1474,1356,1378]" gridcol="5" gridrow="28" pageId="5" pageNumber="130">10</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFA92B704FABD" box="[138,1816,1385,1407]" gridrow="29" pageId="5" pageNumber="130" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFA92B189FABD" box="[138,405,1385,1407]" gridcol="0" gridrow="29" pageId="5" pageNumber="130">35722</th>
</tr>
<tr id="356234F4EB6F9BFCFF6CFA7CB704FA5F" box="[138,1816,1415,1437]" gridrow="30" pageId="5" pageNumber="130" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFA7CB189FA5F" box="[138,405,1415,1437]" gridcol="0" gridrow="30" pageId="5" pageNumber="130">Manual phalanx n.3 ML</th>
<td id="76B35D88EB6F9BFCFE4BFA7CB228FA5F" box="[429,564,1415,1437]" gridcol="1" gridrow="30" pageId="5" pageNumber="130">17</td>
<td id="76B35D88EB6F9BFCFD93FA7CB321FA5F" box="[629,829,1415,1437]" gridcol="2" gridrow="30" pageId="5" pageNumber="130">16</td>
<td id="76B35D88EB6F9BFCFC87FA7CB434FA5F" box="[865,1064,1415,1437]" gridcol="3" gridrow="30" pageId="5" pageNumber="130">19</td>
<td id="76B35D88EB6F9BFCFBA2FA7CB4F2FA5F" box="[1092,1262,1415,1437]" gridcol="4" gridrow="30" pageId="5" pageNumber="130">10</td>
<td id="76B35D88EB6F9BFCFAE2FA7CB5DEFA5F" box="[1284,1474,1415,1437]" gridcol="5" gridrow="30" pageId="5" pageNumber="130">13</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFA5EB704FA79" box="[138,1816,1445,1467]" gridrow="31" pageId="5" pageNumber="130">
<th id="76B35D88EB6F9BFCFF6CFA5EB704FA79" box="[138,1816,1445,1467]" colspan="8" colspanRight="7" gridcol="0" gridrow="31" pageId="5" pageNumber="130">35721</th>
</tr>
<tr id="356234F4EB6F9BFCFF6CFA38B704FA1B" box="[138,1816,1475,1497]" gridrow="32" pageId="5" pageNumber="130" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFA38B189FA1B" box="[138,405,1475,1497]" gridcol="0" gridrow="32" pageId="5" pageNumber="130">Manual phalanx n.4 ML</th>
<td id="76B35D88EB6F9BFCFE4BFA38B228FA1B" box="[429,564,1475,1497]" gridcol="1" gridrow="32" pageId="5" pageNumber="130">19</td>
<td id="76B35D88EB6F9BFCFD93FA38B321FA1B" box="[629,829,1475,1497]" gridcol="2" gridrow="32" pageId="5" pageNumber="130">18</td>
<td id="76B35D88EB6F9BFCFC87FA38B434FA1B" box="[865,1064,1475,1497]" gridcol="3" gridrow="32" pageId="5" pageNumber="130">20</td>
<td id="76B35D88EB6F9BFCFBA2FA38B4F2FA1B" box="[1092,1262,1475,1497]" gridcol="4" gridrow="32" pageId="5" pageNumber="130">9</td>
<td id="76B35D88EB6F9BFCFAE2FA38B5DEFA1B" box="[1284,1474,1475,1497]" gridcol="5" gridrow="32" pageId="5" pageNumber="130">15</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CFA1BB704FA34" box="[138,1816,1504,1526]" gridrow="33" pageId="5" pageNumber="130" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFA1BB189FA34" box="[138,405,1504,1526]" gridcol="0" gridrow="33" pageId="5" pageNumber="130">35724</th>
</tr>
<tr id="356234F4EB6F9BFCFF6CFA05B704F9D6" box="[138,1816,1534,1556]" gridrow="34" pageId="5" pageNumber="130" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CFA05B189F9D6" box="[138,405,1534,1556]" gridcol="0" gridrow="34" pageId="5" pageNumber="130">Manual ungual n.1 ML</th>
<td id="76B35D88EB6F9BFCFE4BFA05B228F9D6" box="[429,564,1534,1556]" gridcol="1" gridrow="34" pageId="5" pageNumber="130">35</td>
<td id="76B35D88EB6F9BFCFC87FA05B434F9D6" box="[865,1064,1534,1556]" gridcol="3" gridrow="34" pageId="5" pageNumber="130">22</td>
<td id="76B35D88EB6F9BFCFAE2FA05B5DEF9D6" box="[1284,1474,1534,1556]" gridcol="5" gridrow="34" pageId="5" pageNumber="130">15</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CF9E0B704F9F3" box="[138,1816,1563,1585]" gridrow="35" pageId="5" pageNumber="130" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CF9E0B189F9F3" box="[138,405,1563,1585]" gridcol="0" gridrow="35" pageId="5" pageNumber="130">3571</th>
</tr>
<tr id="356234F4EB6F9BFCFF6CF9C2B704F98D" box="[138,1816,1593,1615]" gridrow="36" pageId="5" pageNumber="130" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-7="1">
<th id="76B35D88EB6F9BFCFF6CF9C2B189F98D" box="[138,405,1593,1615]" gridcol="0" gridrow="36" pageId="5" pageNumber="130">Manual ungual n.4 ML</th>
<td id="76B35D88EB6F9BFCFE4BF9C2B228F98D" box="[429,564,1593,1615]" gridcol="1" gridrow="36" pageId="5" pageNumber="130">38</td>
<td id="76B35D88EB6F9BFCFC87F9C2B434F98D" box="[865,1064,1593,1615]" gridcol="3" gridrow="36" pageId="5" pageNumber="130">15</td>
<td id="76B35D88EB6F9BFCFAE2F9C2B5DEF98D" box="[1284,1474,1593,1615]" gridcol="5" gridrow="36" pageId="5" pageNumber="130">15</td>
</tr>
<tr id="356234F4EB6F9BFCFF6CF9ACB704F9AF" box="[138,1816,1623,1645]" gridrow="37" pageId="5" pageNumber="130">
<th id="76B35D88EB6F9BFCFF6CF9ACB704F9AF" box="[138,1816,1623,1645]" colspan="8" colspanRight="7" gridcol="0" gridrow="37" pageId="5" pageNumber="130">35723</th>
</tr>
</table>
</paragraph>
<caption id="DF2D663EEB6C6405FF6CFDD9B141FDFA" box="[138,349,546,568]" isContinuationCaption="true" pageId="6" pageNumber="131" startId="6.[138,202,546,568]" targetBox="[138,1660,596,1720]" targetIsTable="true" targetPageId="6">
<paragraph id="8BED36B6EB6C6405FF6CFDD9B141FDFA" blockId="6.[138,349,546,568]" box="[138,349,546,568]" pageId="6" pageNumber="131">
<emphasis id="B926EAA4EB6C6405FF6CFDD9B0F8FDFA" bold="true" box="[138,228,546,568]" pageId="6" pageNumber="131">Table 1.</emphasis>
Continued
</paragraph>
</caption>
<paragraph id="8BED36B6EB6C6405FF6CFDAFB0A6F97A" pageId="6" pageNumber="131">
<table id="F952C416EB6C9BFCFF6CFDAFB660F97A" box="[138,1660,596,1720]" gridcols="7" gridrows="38" pageId="6" pageNumber="131">
<tr id="356234F4EB6C9BFCFF6CFDAFB660FDA8" box="[138,1660,596,618]" gridrow="0" pageId="6" pageNumber="131">
<th id="76B35D88EB6C9BFCFF6CFDAFB660FDA8" box="[138,1660,596,618]" colspan="7" colspanRight="6" gridcol="0" gridrow="0" pageId="6" pageNumber="131">Right manual elements</th>
</tr>
<tr id="356234F4EB6C9BFCFF6CFD89B660FD4A" box="[138,1660,626,648]" gridrow="1" pageId="6" pageNumber="131" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CFD89B181FD4A" box="[138,413,626,648]" gridcol="0" gridrow="1" pageId="6" pageNumber="131">Manual phalanx n.1</th>
<td id="76B35D88EB6C9BFCFE4BFD89B226FD4A" box="[429,570,626,648]" gridcol="1" gridrow="1" pageId="6" pageNumber="131">17</td>
<td id="76B35D88EB6C9BFCFD93FD89B357FD4A" box="[629,843,626,648]" gridcol="2" gridrow="1" pageId="6" pageNumber="131">17</td>
<td id="76B35D88EB6C9BFCFC84FD89B436FD4A" box="[866,1066,626,648]" gridcol="3" gridrow="1" pageId="6" pageNumber="131">22</td>
<td id="76B35D88EB6C9BFCFBA2FD89B4FBFD4A" box="[1092,1255,626,648]" gridcol="4" gridrow="1" pageId="6" pageNumber="131">10</td>
<td id="76B35D88EB6C9BFCFAE2FD89B5DEFD4A" box="[1284,1474,626,648]" gridcol="5" gridrow="1" pageId="6" pageNumber="131">17</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFD6BB660FD64" box="[138,1660,656,678]" gridrow="2" pageId="6" pageNumber="131">
<th id="76B35D88EB6C9BFCFF6CFD6BB660FD64" box="[138,1660,656,678]" colspan="7" colspanRight="6" gridcol="0" gridrow="2" pageId="6" pageNumber="131">ML 3574</th>
</tr>
<tr id="356234F4EB6C9BFCFF6CFD55B660FD06" box="[138,1660,686,708]" gridrow="3" pageId="6" pageNumber="131" rowspan-2="1" rowspan-4="1" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CFD55B181FD06" box="[138,413,686,708]" gridcol="0" gridrow="3" pageId="6" pageNumber="131">Manual ungual n.2 zML</th>
<td id="76B35D88EB6C9BFCFE4BFD55B226FD06" box="[429,570,686,708]" gridcol="1" gridrow="3" pageId="6" pageNumber="131">30</td>
<td id="76B35D88EB6C9BFCFC84FD55B436FD06" box="[866,1066,686,708]" gridcol="3" gridrow="3" pageId="6" pageNumber="131">17</td>
<td id="76B35D88EB6C9BFCFAE2FD55B5DEFD06" box="[1284,1474,686,708]" gridcol="5" gridrow="3" pageId="6" pageNumber="131">15</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFD30B660FD23" box="[138,1660,715,737]" gridrow="4" pageId="6" pageNumber="131" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CFD30B181FD23" box="[138,413,715,737]" gridcol="0" gridrow="4" pageId="6" pageNumber="131">3572</th>
</tr>
<tr id="356234F4EB6C9BFCFF6CFD12B660FD3D" box="[138,1660,745,767]" gridrow="5" pageId="6" pageNumber="131" rowspan-2="1" rowspan-4="1" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CFD12B181FD3D" box="[138,413,745,767]" gridcol="0" gridrow="5" pageId="6" pageNumber="131">Manual ungual n.3</th>
<td id="76B35D88EB6C9BFCFE4BFD12B226FD3D" box="[429,570,745,767]" gridcol="1" gridrow="5" pageId="6" pageNumber="131">33</td>
<td id="76B35D88EB6C9BFCFC84FD12B436FD3D" box="[866,1066,745,767]" gridcol="3" gridrow="5" pageId="6" pageNumber="131">12</td>
<td id="76B35D88EB6C9BFCFAE2FD12B5DEFD3D" box="[1284,1474,745,767]" gridcol="5" gridrow="5" pageId="6" pageNumber="131">14</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFCFDB660FCDE" box="[138,1660,774,796]" gridrow="6" pageId="6" pageNumber="131" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CFCFDB181FCDE" box="[138,413,774,796]" gridcol="0" gridrow="6" pageId="6" pageNumber="131">ML 3573</th>
</tr>
<tr id="356234F4EB6C9BFCFF6CFCDFB660FCF8" box="[138,1660,804,826]" gridrow="7" pageId="6" pageNumber="131" rowspan-2="1" rowspan-4="1" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CFCDFB181FCF8" box="[138,413,804,826]" gridcol="0" gridrow="7" pageId="6" pageNumber="131">Manual ungual n.5</th>
<td id="76B35D88EB6C9BFCFE4BFCDFB226FCF8" box="[429,570,804,826]" gridcol="1" gridrow="7" pageId="6" pageNumber="131">33</td>
<td id="76B35D88EB6C9BFCFC84FCDFB436FCF8" box="[866,1066,804,826]" gridcol="3" gridrow="7" pageId="6" pageNumber="131">18</td>
<td id="76B35D88EB6C9BFCFAE2FCDFB5DEFCF8" box="[1284,1474,804,826]" gridcol="5" gridrow="7" pageId="6" pageNumber="131">13</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFCB9B660FC9A" box="[138,1660,834,856]" gridrow="8" pageId="6" pageNumber="131">
<th id="76B35D88EB6C9BFCFF6CFCB9B660FC9A" box="[138,1660,834,856]" colspan="7" colspanRight="6" gridcol="0" gridrow="8" pageId="6" pageNumber="131">ML 35720</th>
</tr>
<tr id="356234F4EB6C9BFCFF6CFC9BB660FCB4" box="[138,1660,864,886]" gridrow="9" pageId="6" pageNumber="131" rowspan-0="1">
<td id="76B35D88EB6C9BFCFE4BFC9BB226FCB4" box="[429,570,864,886]" gridcol="1" gridrow="9" pageId="6" pageNumber="131">Total Length</td>
<td id="76B35D88EB6C9BFCFD93FC9BB357FCB4" box="[629,843,864,886]" gridcol="2" gridrow="9" pageId="6" pageNumber="131">Mediolateral distal</td>
<td id="76B35D88EB6C9BFCFC84FC9BB436FCB4" box="[866,1066,864,886]" gridcol="3" gridrow="9" pageId="6" pageNumber="131">Medial condyle</td>
<td id="76B35D88EB6C9BFCFBA2FC9BB4FBFCB4" box="[1092,1255,864,886]" gridcol="4" gridrow="9" pageId="6" pageNumber="131">Lateral condyle</td>
<td id="76B35D88EB6C9BFCFAE2FC9BB5DEFCB4" box="[1284,1474,864,886]" gridcol="5" gridrow="9" pageId="6" pageNumber="131">Flexor groove</td>
<td id="76B35D88EB6C9BFCFA32FC9BB660FCB4" box="[1492,1660,864,886]" gridcol="6" gridrow="9" pageId="6" pageNumber="131">Extensor groove</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFC86B660FC51" box="[138,1660,893,915]" gridrow="10" pageId="6" pageNumber="131" rowspan-0="1" rowspan-1="1">
<td id="76B35D88EB6C9BFCFD93FC86B357FC51" box="[629,843,893,915]" gridcol="2" gridrow="10" pageId="6" pageNumber="131">width</td>
<td id="76B35D88EB6C9BFCFC84FC86B436FC51" box="[866,1066,893,915]" gridcol="3" gridrow="10" pageId="6" pageNumber="131">length</td>
<td id="76B35D88EB6C9BFCFBA2FC86B4FBFC51" box="[1092,1255,893,915]" gridcol="4" gridrow="10" pageId="6" pageNumber="131">length</td>
<td id="76B35D88EB6C9BFCFAE2FC86B5DEFC51" box="[1284,1474,893,915]" gridcol="5" gridrow="10" pageId="6" pageNumber="131">width</td>
<td id="76B35D88EB6C9BFCFA32FC86B660FC51" box="[1492,1660,893,915]" gridcol="6" gridrow="10" pageId="6" pageNumber="131">width</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFC60B660FC73" box="[138,1660,923,945]" gridrow="11" pageId="6" pageNumber="131">
<th id="76B35D88EB6C9BFCFF6CFC60B181FC73" box="[138,413,923,945]" gridcol="0" gridrow="11" pageId="6" pageNumber="131">Femur ML 3576</th>
<td id="76B35D88EB6C9BFCFE4BFC60B226FC73" box="[429,570,923,945]" gridcol="1" gridrow="11" pageId="6" pageNumber="131">280</td>
<td id="76B35D88EB6C9BFCFD93FC60B357FC73" box="[629,843,923,945]" gridcol="2" gridrow="11" pageId="6" pageNumber="131">120</td>
<td id="76B35D88EB6C9BFCFC84FC60B436FC73" box="[866,1066,923,945]" gridcol="3" gridrow="11" pageId="6" pageNumber="131">120</td>
<td id="76B35D88EB6C9BFCFBA2FC60B4FBFC73" box="[1092,1255,923,945]" gridcol="4" gridrow="11" pageId="6" pageNumber="131">110</td>
<td id="76B35D88EB6C9BFCFAE2FC60B5DEFC73" box="[1284,1474,923,945]" gridcol="5" gridrow="11" pageId="6" pageNumber="131">50</td>
<td id="76B35D88EB6C9BFCFA32FC60B660FC73" box="[1492,1660,923,945]" gridcol="6" gridrow="11" pageId="6" pageNumber="131">30</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFC42B660FC0D" box="[138,1660,953,975]" gridrow="12" pageId="6" pageNumber="131" rowspan-0="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td id="76B35D88EB6C9BFCFE4BFC42B226FC0D" box="[429,570,953,975]" gridcol="1" gridrow="12" pageId="6" pageNumber="131">Total Length</td>
<td id="76B35D88EB6C9BFCFD93FC42B357FC0D" box="[629,843,953,975]" gridcol="2" gridrow="12" pageId="6" pageNumber="131">Craniocaudal distal</td>
<td id="76B35D88EB6C9BFCFC84FC42B436FC0D" box="[866,1066,953,975]" gridcol="3" gridrow="12" pageId="6" pageNumber="131">Mediolateral width</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFC2DB660FC2E" box="[138,1660,982,1004]" gridrow="13" pageId="6" pageNumber="131" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td id="76B35D88EB6C9BFCFD93FC2DB357FC2E" box="[629,843,982,1004]" gridcol="2" gridrow="13" pageId="6" pageNumber="131">width</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFC0FB660FBC8" box="[138,1660,1012,1034]" gridrow="14" pageId="6" pageNumber="131" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CFC0FB181FBC8" box="[138,413,1012,1034]" gridcol="0" gridrow="14" pageId="6" pageNumber="131">Tibia proximal epiphysis</th>
<td id="76B35D88EB6C9BFCFE4BFC0FB226FBC8" box="[429,570,1012,1034]" gridcol="1" gridrow="14" pageId="6" pageNumber="131">210</td>
<td id="76B35D88EB6C9BFCFD93FC0FB357FBC8" box="[629,843,1012,1034]" gridcol="2" gridrow="14" pageId="6" pageNumber="131">140</td>
<td id="76B35D88EB6C9BFCFC84FC0FB436FBC8" box="[866,1066,1012,1034]" gridcol="3" gridrow="14" pageId="6" pageNumber="131">135</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFBE9B660FBEA" box="[138,1660,1042,1064]" gridrow="15" pageId="6" pageNumber="131">
<th id="76B35D88EB6C9BFCFF6CFBE9B660FBEA" box="[138,1660,1042,1064]" colspan="7" colspanRight="6" gridcol="0" gridrow="15" pageId="6" pageNumber="131">ML 3577</th>
</tr>
<tr id="356234F4EB6C9BFCFF6CFBCBB660FB84" box="[138,1660,1072,1094]" gridrow="16" pageId="6" pageNumber="131" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td id="76B35D88EB6C9BFCFE4BFBCBB226FB84" box="[429,570,1072,1094]" gridcol="1" gridrow="16" pageId="6" pageNumber="131">Total Length</td>
<td id="76B35D88EB6C9BFCFD93FBCBB357FB84" box="[629,843,1072,1094]" gridcol="2" gridrow="16" pageId="6" pageNumber="131">Mediolateral distal</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFBB6B660FBA1" box="[138,1660,1101,1123]" gridrow="17" pageId="6" pageNumber="131" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td id="76B35D88EB6C9BFCFD93FBB6B357FBA1" box="[629,843,1101,1123]" gridcol="2" gridrow="17" pageId="6" pageNumber="131">width</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFB90B660FB43" box="[138,1660,1131,1153]" gridrow="18" pageId="6" pageNumber="131" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CFB90B181FB43" box="[138,413,1131,1153]" gridcol="0" gridrow="18" pageId="6" pageNumber="131">Tibia distal epiphysis ML</th>
<td id="76B35D88EB6C9BFCFE4BFB90B226FB43" box="[429,570,1131,1153]" gridcol="1" gridrow="18" pageId="6" pageNumber="131">210</td>
<td id="76B35D88EB6C9BFCFD93FB90B357FB43" box="[629,843,1131,1153]" gridcol="2" gridrow="18" pageId="6" pageNumber="131">150</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFB73B660FB5C" box="[138,1660,1160,1182]" gridrow="19" pageId="6" pageNumber="131" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CFB73B181FB5C" box="[138,413,1160,1182]" gridcol="0" gridrow="19" pageId="6" pageNumber="131">35712</th>
</tr>
<tr id="356234F4EB6C9BFCFF6CFB5CB660FB7F" box="[138,1660,1191,1213]" gridrow="20" pageId="6" pageNumber="131" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td id="76B35D88EB6C9BFCFE4BFB5CB226FB7F" box="[429,570,1191,1213]" gridcol="1" gridrow="20" pageId="6" pageNumber="131">Craniocaudal</td>
<td id="76B35D88EB6C9BFCFD93FB5CB357FB7F" box="[629,843,1191,1213]" gridcol="2" gridrow="20" pageId="6" pageNumber="131">Craniocaudal length</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFB3FB660FB18" box="[138,1660,1220,1242]" gridrow="21" pageId="6" pageNumber="131" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td id="76B35D88EB6C9BFCFE4BFB3FB226FB18" box="[429,570,1220,1242]" gridcol="1" gridrow="21" pageId="6" pageNumber="131">length</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFB19B660FB3A" box="[138,1660,1250,1272]" gridrow="22" pageId="6" pageNumber="131" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CFB19B181FB3A" box="[138,413,1250,1272]" gridcol="0" gridrow="22" pageId="6" pageNumber="131">Fibula (proximal) ML</th>
<td id="76B35D88EB6C9BFCFE4BFB19B226FB3A" box="[429,570,1250,1272]" gridcol="1" gridrow="22" pageId="6" pageNumber="131">75</td>
<td id="76B35D88EB6C9BFCFD93FB19B357FB3A" box="[629,843,1250,1272]" gridcol="2" gridrow="22" pageId="6" pageNumber="131">90</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFB04B660FAD7" box="[138,1660,1279,1301]" gridrow="23" pageId="6" pageNumber="131" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CFB04B181FAD7" box="[138,413,1279,1301]" gridcol="0" gridrow="23" pageId="6" pageNumber="131">3578</th>
</tr>
<tr id="356234F4EB6C9BFCFF6CFAE6B660FAF1" box="[138,1660,1309,1331]" gridrow="24" pageId="6" pageNumber="131">
<th id="76B35D88EB6C9BFCFF6CFAE6B660FAF1" box="[138,1660,1309,1331]" colspan="7" colspanRight="6" gridcol="0" gridrow="24" pageId="6" pageNumber="131">Pes ML 35712</th>
</tr>
<tr id="356234F4EB6C9BFCFF6CFAC0B660FA93" box="[138,1660,1339,1361]" gridrow="25" pageId="6" pageNumber="131" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td id="76B35D88EB6C9BFCFE4BFAC0B226FA93" box="[429,570,1339,1361]" gridcol="1" gridrow="25" pageId="6" pageNumber="131">Mediolateral</td>
<td id="76B35D88EB6C9BFCFD93FAC0B357FA93" box="[629,843,1339,1361]" gridcol="2" gridrow="25" pageId="6" pageNumber="131">Dorsoventral height</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFAA2B660FAAD" box="[138,1660,1369,1391]" gridrow="26" pageId="6" pageNumber="131">
<th id="76B35D88EB6C9BFCFF6CFAA2B660FAAD" box="[138,1660,1369,1391]" colspan="7" colspanRight="6" gridcol="0" gridrow="26" pageId="6" pageNumber="131">width</th>
</tr>
<tr id="356234F4EB6C9BFCFF6CFA8CB660FA4F" box="[138,1660,1399,1421]" gridrow="27" pageId="6" pageNumber="131" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CFA8CB181FA4F" box="[138,413,1399,1421]" gridcol="0" gridrow="27" pageId="6" pageNumber="131">Tarsal 3 ML 35712</th>
<td id="76B35D88EB6C9BFCFE4BFA8CB226FA4F" box="[429,570,1399,1421]" gridcol="1" gridrow="27" pageId="6" pageNumber="131">50</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFA6EB660FA69" box="[138,1660,1429,1451]" gridrow="28" pageId="6" pageNumber="131" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CFA6EB181FA69" box="[138,413,1429,1451]" gridcol="0" gridrow="28" pageId="6" pageNumber="131">Tarsal 4 ML 35712</th>
<td id="76B35D88EB6C9BFCFE4BFA6EB226FA69" box="[429,570,1429,1451]" gridcol="1" gridrow="28" pageId="6" pageNumber="131">70</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFA48B660FA0B" box="[138,1660,1459,1481]" gridrow="29" pageId="6" pageNumber="131" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CFA48B181FA0B" box="[138,413,1459,1481]" gridcol="0" gridrow="29" pageId="6" pageNumber="131">Astragalus ML 35712</th>
<td id="76B35D88EB6C9BFCFE4BFA48B226FA0B" box="[429,570,1459,1481]" gridcol="1" gridrow="29" pageId="6" pageNumber="131">90</td>
<td id="76B35D88EB6C9BFCFD93FA48B357FA0B" box="[629,843,1459,1481]" gridcol="2" gridrow="29" pageId="6" pageNumber="131">50</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFA2AB660FA25" box="[138,1660,1489,1511]" gridrow="30" pageId="6" pageNumber="131" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CFA2AB181FA25" box="[138,413,1489,1511]" gridcol="0" gridrow="30" pageId="6" pageNumber="131">Calcaneum ML 35712</th>
<td id="76B35D88EB6C9BFCFE4BFA2AB226FA25" box="[429,570,1489,1511]" gridcol="1" gridrow="30" pageId="6" pageNumber="131">30</td>
<td id="76B35D88EB6C9BFCFD93FA2AB357FA25" box="[629,843,1489,1511]" gridcol="2" gridrow="30" pageId="6" pageNumber="131">70</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CFA14B660F9C7" box="[138,1660,1519,1541]" gridrow="31" pageId="6" pageNumber="131" rowspan-0="1" rowspan-6="1">
<td id="76B35D88EB6C9BFCFE4BFA14B226F9C7" box="[429,570,1519,1541]" gridcol="1" gridrow="31" pageId="6" pageNumber="131">Total Length</td>
<td id="76B35D88EB6C9BFCFD93FA14B357F9C7" box="[629,843,1519,1541]" gridcol="2" gridrow="31" pageId="6" pageNumber="131">Mediolateral distal</td>
<td id="76B35D88EB6C9BFCFC84FA14B436F9C7" box="[866,1066,1519,1541]" gridcol="3" gridrow="31" pageId="6" pageNumber="131">Mediolateral</td>
<td id="76B35D88EB6C9BFCFBA2FA14B4FBF9C7" box="[1092,1255,1519,1541]" gridcol="4" gridrow="31" pageId="6" pageNumber="131">Dorsoplantar</td>
<td id="76B35D88EB6C9BFCFAE2FA14B5DEF9C7" box="[1284,1474,1519,1541]" gridcol="5" gridrow="31" pageId="6" pageNumber="131">Dorsplantar</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CF9F7B660F9E0" box="[138,1660,1548,1570]" gridrow="32" pageId="6" pageNumber="131" rowspan-0="1" rowspan-1="1" rowspan-6="1">
<td id="76B35D88EB6C9BFCFD93F9F7B357F9E0" box="[629,843,1548,1570]" gridcol="2" gridrow="32" pageId="6" pageNumber="131">width</td>
<td id="76B35D88EB6C9BFCFC84F9F7B436F9E0" box="[866,1066,1548,1570]" gridcol="3" gridrow="32" pageId="6" pageNumber="131">proximal width</td>
<td id="76B35D88EB6C9BFCFBA2F9F7B4FBF9E0" box="[1092,1255,1548,1570]" gridcol="4" gridrow="32" pageId="6" pageNumber="131">distal height</td>
<td id="76B35D88EB6C9BFCFAE2F9F7B5DEF9E0" box="[1284,1474,1548,1570]" gridcol="5" gridrow="32" pageId="6" pageNumber="131">proximal height</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CF9D1B660F982" box="[138,1660,1578,1600]" gridrow="33" pageId="6" pageNumber="131" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CF9D1B181F982" box="[138,413,1578,1600]" gridcol="0" gridrow="33" pageId="6" pageNumber="131">Metatarsal I ML 35712</th>
<td id="76B35D88EB6C9BFCFE4BF9D1B226F982" box="[429,570,1578,1600]" gridcol="1" gridrow="33" pageId="6" pageNumber="131">30*</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CF9B3B660F99C" box="[138,1660,1608,1630]" gridrow="34" pageId="6" pageNumber="131" rowspan-5="1" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CF9B3B181F99C" box="[138,413,1608,1630]" gridcol="0" gridrow="34" pageId="6" pageNumber="131">Metatarsal II ML 35712</th>
<td id="76B35D88EB6C9BFCFE4BF9B3B226F99C" box="[429,570,1608,1630]" gridcol="1" gridrow="34" pageId="6" pageNumber="131">140</td>
<td id="76B35D88EB6C9BFCFD93F9B3B357F99C" box="[629,843,1608,1630]" gridcol="2" gridrow="34" pageId="6" pageNumber="131">50</td>
<td id="76B35D88EB6C9BFCFC84F9B3B436F99C" box="[866,1066,1608,1630]" gridcol="3" gridrow="34" pageId="6" pageNumber="131">25</td>
<td id="76B35D88EB6C9BFCFBA2F9B3B4FBF99C" box="[1092,1255,1608,1630]" gridcol="4" gridrow="34" pageId="6" pageNumber="131">45</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CF99DB660F9BE" box="[138,1660,1638,1660]" gridrow="35" pageId="6" pageNumber="131" rowspan-5="1" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CF99DB181F9BE" box="[138,413,1638,1660]" gridcol="0" gridrow="35" pageId="6" pageNumber="131">Metatarsal III ML 35712</th>
<td id="76B35D88EB6C9BFCFE4BF99DB226F9BE" box="[429,570,1638,1660]" gridcol="1" gridrow="35" pageId="6" pageNumber="131">175</td>
<td id="76B35D88EB6C9BFCFD93F99DB357F9BE" box="[629,843,1638,1660]" gridcol="2" gridrow="35" pageId="6" pageNumber="131">70</td>
<td id="76B35D88EB6C9BFCFC84F99DB436F9BE" box="[866,1066,1638,1660]" gridcol="3" gridrow="35" pageId="6" pageNumber="131">40</td>
<td id="76B35D88EB6C9BFCFBA2F99DB4FBF9BE" box="[1092,1255,1638,1660]" gridcol="4" gridrow="35" pageId="6" pageNumber="131">40</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CF97FB660F958" box="[138,1660,1668,1690]" gridrow="36" pageId="6" pageNumber="131" rowspan-5="1" rowspan-6="1">
<th id="76B35D88EB6C9BFCFF6CF97FB226F958" box="[138,570,1668,1690]" colspan="2" colspanRight="1" gridcol="0" gridrow="36" pageId="6" pageNumber="131">Metatarsal IV ML 35712, 60*</th>
<td id="76B35D88EB6C9BFCFD93F97FB357F958" box="[629,843,1668,1690]" gridcol="2" gridrow="36" pageId="6" pageNumber="131">35</td>
<td id="76B35D88EB6C9BFCFC84F97FB436F958" box="[866,1066,1668,1690]" gridcol="3" gridrow="36" pageId="6" pageNumber="131">70</td>
<td id="76B35D88EB6C9BFCFBA2F97FB4FBF958" box="[1092,1255,1668,1690]" gridcol="4" gridrow="36" pageId="6" pageNumber="131">45</td>
</tr>
<tr id="356234F4EB6C9BFCFF6CF959B660F97A" box="[138,1660,1698,1720]" gridrow="37" pageId="6" pageNumber="131">
<th id="76B35D88EB6C9BFCFF6CF959B660F97A" box="[138,1660,1698,1720]" colspan="7" colspanRight="6" gridcol="0" gridrow="37" pageId="6" pageNumber="131">18</th>
</tr>
</table>
</paragraph>
<paragraph id="8BED36B6EB6D6404FCDDFEA1B3D3FCDF" blockId="7.[827,1450,346,798]" pageId="7" pageNumber="132">
<emphasis id="B926EAA4EB6D6404FCDDFEA1B3B4FEAD" box="[827,936,346,367]" italics="true" pageId="7" pageNumber="132">Remarks:</emphasis>
The three caudal vertebrae are stout and cylindrical, similar in overall shape and proportions to those of other basal iguanodontians, such as:
<taxonomicName id="4C524D35EB6D6404FCDDFE4CB4E4FE0E" authority=", Cumnoria" authorityName="Cumnoria" box="[827,1272,438,460]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="132" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B926EAA4EB6D6404FCDDFE4CB441FE0E" box="[827,1117,438,460]" italics="true" pageId="7" pageNumber="132">Camptosaurus dispar</emphasis>
,
<emphasis id="B926EAA4EB6D6404FB93FE4CB4E4FE0E" box="[1141,1272,439,460]" italics="true" pageId="7" pageNumber="132">Cumnoria</emphasis>
</taxonomicName>
<taxonomicName id="4C524D35EB6D6404FAE1FE4CB586FE0E" authorityName="Hulke" authorityYear="1880" box="[1287,1434,438,460]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="132" phylum="Chordata" rank="species" species="prestwichii">
<emphasis id="B926EAA4EB6D6404FAE1FE4CB586FE0E" box="[1287,1434,438,460]" italics="true" pageId="7" pageNumber="132">prestwichii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C524D35EB6D6404FCDDFE2DB40DFE28" authorityName="Marsh" authorityYear="1878" box="[827,1041,469,491]" class="Reptilia" family="Dryosauridae" genus="Dryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="132" phylum="Chordata" rank="species" species="altus">
<emphasis id="B926EAA4EB6D6404FCDDFE2DB40DFE28" box="[827,1041,469,491]" italics="true" pageId="7" pageNumber="132">Dryosaurus altus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C524D35EB6D6404FBC5FE2EB587FE28" authorityName="Virchow" authorityYear="1919" box="[1059,1435,469,490]" class="Reptilia" family="Dryosauridae" genus="Dysalotosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="132" phylum="Chordata" rank="species" species="lettowvorbecki">
<emphasis id="B926EAA4EB6D6404FBC5FE2EB587FE28" box="[1059,1435,469,490]" italics="true" pageId="7" pageNumber="132">Dysalotosaurus lettowvorbecki</emphasis>
</taxonomicName>
,
<taxonomicName id="4C524D35EB6D6404FCDDFE0FB577FDC8" authority="Mantell, 1825" authorityName="Mantell" authorityYear="1825" box="[827,1387,500,522]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="132" phylum="Chordata" rank="species" species="atherfieldensis">
<emphasis id="B926EAA4EB6D6404FCDDFE0FB4A8FDCB" box="[827,1204,500,521]" italics="true" pageId="7" pageNumber="132">Mantellisaurus atherfieldensis</emphasis>
<bibRefCitation id="EFC34B47EB6D6404FB26FE0FB577FDC8" author="Mantell GA" box="[1216,1387,500,522]" pageId="7" pageNumber="132" pagination="179 - 186" refId="ref17874" refString="Mantell GA. 1825. Notice on the Iguanodon, a newly discovered fossil reptile, from the sandstone of Tilgate Forest, in Sussex. Philosophical Transactions of the Royal Society of London 115: 179 - 186." type="journal article" year="1825">Mantell, 1825</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C524D35EB6D6404FCDDFDE9B428FDE5" baseAuthorityName="Carpenter &amp; Wilson" baseAuthorityYear="2008" box="[827,1076,530,551]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="132" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="B926EAA4EB6D6404FCDDFDE9B428FDE5" box="[827,1076,530,551]" italics="true" pageId="7" pageNumber="132">Uteodon aphanoecetes</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC34B47EB6D6404FBA2FDE9B55EFDEA" author="Galton PM &amp; Powell HP" box="[1092,1346,530,552]" pageId="7" pageNumber="132" pagination="411 - 443" refId="ref17215" refString="Galton PM, Powell HP. 1980. The ornithischian dinosaur Camptosaurus prestwichii from the Upper Jurassic of England. Palaeontology 23: 411 - 443." type="journal article" year="1980">Galton &amp; Powell, 1980</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB6D6404FAA9FDE9B36AFD84" author="Galton PM" pageId="7" pageNumber="132" pagination="271 - 312" refId="ref17047" refString="Galton PM. 1981. Dryosaurus, a hypsilophodontid dinosaur from the Upper Jurassic of North America and Africa postcranial skeleton. Palaontologische Zeitschrift 55: 271 - 312." type="journal article" year="1981">Galton, 1981</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB6D6404FC6FFDC9B427FD85" author="Norman DB" box="[905,1083,561,583]" pageId="7" pageNumber="132" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">Norman, 1986</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB6D6404FBA8FDCAB580FD84" author="Carpenter K &amp; Wilson Y" box="[1102,1436,561,583]" pageId="7" pageNumber="132" pagination="227 - 264" refId="ref16499" refString="Carpenter K, Wilson Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter &amp; Wilson, 2008</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB6D6404FCDDFDABB478FDA7" author="Carpenter K &amp; Galton PM" box="[827,1124,592,614]" pageId="7" pageNumber="132" pagination="167 - 207" refId="ref16420" refString="Carpenter K, Galton PM. 2018. A photo documentation of bipedal ornithischian dinosaurs from the Upper Jurassic Morrison Formation, USA. Geology of the Intermountain West 5: 167 - 207." type="journal article" year="2018">Carpenter &amp; Galton, 2018</bibRefCitation>
). They do differ remarkably from those of
<taxonomicName id="4C524D35EB6D6404FC30FD95B36EFD60" authority="Van Beneden, 1881" authorityName="Van Beneden" authorityYear="1881" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="132" phylum="Chordata" rank="species" species="bernissartensis">
<emphasis id="B926EAA4EB6D6404FC30FD95B51DFD41" box="[982,1281,622,643]" italics="true" pageId="7" pageNumber="132">Iguanodon bernissartensis</emphasis>
<bibRefCitation id="EFC34B47EB6D6404FAE1FD94B36EFD60" author="Van Beneden PJ" pageId="7" pageNumber="132" pagination="600 - 608" refId="ref19216" refString="Van Beneden PJ. 1881. Sur l'arc pelvien chez les dinosauriens de Bernissart (review of G. A. Boulenger). Bulletin de l'Academie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique (3 rd ser.) 1: 600 - 608." type="journal article" year="1881">Van Beneden, 1881</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C524D35EB6D6404FC4CFD76B5BEFD61" authority="McDonald et al., 2010 b" authorityName="McDonald" authorityYear="2010" box="[938,1442,653,675]" class="Reptilia" genus="Iguanacolossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="132" phylum="Chordata" rank="species" species="fortis">
<emphasis id="B926EAA4EB6D6404FC4CFD76B484FD60" box="[938,1176,653,674]" italics="true" pageId="7" pageNumber="132">Iguanacolossus fortis</emphasis>
<bibRefCitation id="EFC34B47EB6D6404FB79FD76B5BEFD61" author="McDonald AT &amp; Kirkland JI &amp; DeBlieux DD &amp; Madsen SK &amp; Cavin J &amp; Milner AR &amp; Panzarin L" box="[1183,1442,653,675]" pageId="7" pageNumber="132" pagination="14075" refId="ref18126" refString="McDonald AT, Kirkland JI, DeBlieux DD, Madsen SK, Cavin J, Milner AR, Panzarin L. 2010 b. New basal iguanodonts from the Cedar Mountain Formation of Utah and the evolution of thumb-spiked dinosaurs. PLoS One 5: e 14075." type="journal article" year="2010">
McDonald
<emphasis id="B926EAA4EB6D6404FAFCFD75B556FD60" box="[1306,1354,653,675]" italics="true" pageId="7" pageNumber="132">et al</emphasis>
., 2010b
</bibRefCitation>
</taxonomicName>
in being less discoidal in shape, and from
<taxonomicName id="4C524D35EB6D6404FAD1FD57B477FD22" authority="Lydekker, 1888 a" authorityName="Lydekker" authorityYear="1888" class="Reptilia" family="Iguanodontidae" genus="Barilium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="132" phylum="Chordata" rank="species" species="dawsoni">
<emphasis id="B926EAA4EB6D6404FAD1FD57B382FD1D" italics="true" pageId="7" pageNumber="132">Barilium dawsoni</emphasis>
<bibRefCitation id="EFC34B47EB6D6404FC41FD31B477FD22" author="Lydekker R" box="[935,1131,714,736]" pageId="7" pageNumber="132" pagination="46 - 61" refId="ref17744" refString="Lydekker R. 1888 a. Note on a new Wealden iguanodont and other dinosaurs. Quarterly Journal of the Geological Society of London 44: 46 - 61." type="journal article" year="1888">Lydekker, 1888a</bibRefCitation>
</taxonomicName>
by being less compressed dorsoventrally (
<bibRefCitation id="EFC34B47EB6D6404FBECFD11B4A3FD3C" author="Norman DB" box="[1034,1215,745,767]" pageId="7" pageNumber="132" pagination="1 - 105" refId="ref18270" refString="Norman DB. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman, 1980</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB6D6404FB32FD12B513FD3C" author="Norman DB" box="[1236,1295,745,766]" pageId="7" pageNumber="132" pagination="165 - 194" refId="ref18373" refString="Norman DB. 2011. On the osteology of the lower Wealden (Valanginian) ornithopod Barilium dawsoni (Iguanodontia: Styracosterna). Special Papers in Palaeontology 86: 165 - 194." type="journal article" year="2011">2011</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB6D6404FAC2FD12B3DDFCDC" author="McDonald AT &amp; Kirkland JI &amp; DeBlieux DD &amp; Madsen SK &amp; Cavin J &amp; Milner AR &amp; Panzarin L" pageId="7" pageNumber="132" pagination="14075" refId="ref18126" refString="McDonald AT, Kirkland JI, DeBlieux DD, Madsen SK, Cavin J, Milner AR, Panzarin L. 2010 b. New basal iguanodonts from the Cedar Mountain Formation of Utah and the evolution of thumb-spiked dinosaurs. PLoS One 5: e 14075." type="journal article" year="2010">
<collectingCountry id="F3457626EB6D6404FAC2FD12B5BEFD3D" box="[1316,1442,745,767]" name="Heard Island and McDonald Islands" pageId="7" pageNumber="132">McDonald</collectingCountry>
<emphasis id="B926EAA4EB6D6404FCDDFCF3B376FCDF" box="[827,874,776,797]" italics="true" pageId="7" pageNumber="132">et al</emphasis>
., 2010b
</bibRefCitation>
).
</paragraph>
<paragraph id="8BED36B6EB6D6404FCDDFCA7B54CF9B2" blockId="7.[827,1443,860,1893]" pageId="7" pageNumber="132">
<emphasis id="B926EAA4EB6D6404FCDDFCA7B43DFCB3" box="[827,1057,860,881]" italics="true" pageId="7" pageNumber="132">Manus (ML 35715</emphasis>
,
<emphasis id="B926EAA4EB6D6404FBCBFCA7B4CEFCB3" box="[1069,1234,860,881]" italics="true" pageId="7" pageNumber="132">
2026) (
<figureCitation id="13692A33EB6D6404FB60FCA7B4D8FCB3" box="[1158,1220,860,881]" captionStart="Figure 5" captionStartId="11.[163,244,1604,1626]" captionTargetBox="[166,1446,198,1561]" captionTargetId="figure-26@11.[163,1449,195,1564]" captionTargetPageId="11" captionText="Figure 5. Manual elements of Draconyx loureiroi holotype ML 357 (ML 357 1-5, 20-26), including articulated partial carpus, manual phalanges and unguals (AL). Articulated partial carpus (ML 357 5) in extensor view (A), right phalanx n.1(ML 357 4) in dorsal view (B), lateral view (B1), proximal view (B2) and distal view (B3), left phalanx n.2 (ML 357 22) in dorsal view (C), lateral view (C1), proximal view (C2) and distal view (C3), left phalanx n.3 (ML 357 21) in dorsal view (D), lateral view (D1), proximal view (D2) and distal view (D3), left phalanx n.4 (ML 357 24) in dorsal view (E), lateral view (E1), proximal view (E2) and distal view (E3), isolated distal ends of tarsals (ML 357 25,26) in extensor view (F, G), left ungual phalanx n.1 in dorsal view (D), mediolateral view (D1) and proximal view (D2), left ungual phalanx n.1 (ML 357- 1) in dorsal view (H), mediolateral view (H1) and proximal view (H2), right ungual phalanx n.2 (ML 357 2) in dorsal view (I), mediolateral view (I1) and proximal view (I2), right ungual phalanx n.3 (ML 357 3) in dorsal view (J), mediolateral view (J1) and proximal view (J2), left ungual phalanx n.4 in dorsal view (K), mediolateral view (K1) and proximal view (K2), ungual phalanx n.5 (ML 357 20) in dorsal view (L), mediolateral view (L1) and proximal view (L2)." figureDoi="http://doi.org/10.5281/zenodo.6530807" httpUri="https://zenodo.org/record/6530807/files/figure.png" pageId="7" pageNumber="132">Fig. 5</figureCitation>
):
</emphasis>
The partial carpus (ML 3575) is composed of the proximal part of a metacarpal, two distal carpals and a proximal carpal (
<figureCitation id="13692A33EB6D6404FCA5FC43B388FC0C" box="[835,916,952,974]" captionStart="Figure 5" captionStartId="11.[163,244,1604,1626]" captionTargetBox="[166,1446,198,1561]" captionTargetId="figure-26@11.[163,1449,195,1564]" captionTargetPageId="11" captionText="Figure 5. Manual elements of Draconyx loureiroi holotype ML 357 (ML 357 1-5, 20-26), including articulated partial carpus, manual phalanges and unguals (AL). Articulated partial carpus (ML 357 5) in extensor view (A), right phalanx n.1(ML 357 4) in dorsal view (B), lateral view (B1), proximal view (B2) and distal view (B3), left phalanx n.2 (ML 357 22) in dorsal view (C), lateral view (C1), proximal view (C2) and distal view (C3), left phalanx n.3 (ML 357 21) in dorsal view (D), lateral view (D1), proximal view (D2) and distal view (D3), left phalanx n.4 (ML 357 24) in dorsal view (E), lateral view (E1), proximal view (E2) and distal view (E3), isolated distal ends of tarsals (ML 357 25,26) in extensor view (F, G), left ungual phalanx n.1 in dorsal view (D), mediolateral view (D1) and proximal view (D2), left ungual phalanx n.1 (ML 357- 1) in dorsal view (H), mediolateral view (H1) and proximal view (H2), right ungual phalanx n.2 (ML 357 2) in dorsal view (I), mediolateral view (I1) and proximal view (I2), right ungual phalanx n.3 (ML 357 3) in dorsal view (J), mediolateral view (J1) and proximal view (J2), left ungual phalanx n.4 in dorsal view (K), mediolateral view (K1) and proximal view (K2), ungual phalanx n.5 (ML 357 20) in dorsal view (L), mediolateral view (L1) and proximal view (L2)." figureDoi="http://doi.org/10.5281/zenodo.6530807" httpUri="https://zenodo.org/record/6530807/files/figure.png" pageId="7" pageNumber="132">Fig. 5A</figureCitation>
). Two isolated distal epiphyses of metacarpals (ML 35725, 26) are associated with the semiarticulated carpus, being both consistent in size and their state of preservation (
<figureCitation id="13692A33EB6D6404FB67FBEFB4EFFBEB" box="[1153,1267,1044,1066]" captionStart="Figure 5" captionStartId="11.[163,244,1604,1626]" captionTargetBox="[166,1446,198,1561]" captionTargetId="figure-26@11.[163,1449,195,1564]" captionTargetPageId="11" captionText="Figure 5. Manual elements of Draconyx loureiroi holotype ML 357 (ML 357 1-5, 20-26), including articulated partial carpus, manual phalanges and unguals (AL). Articulated partial carpus (ML 357 5) in extensor view (A), right phalanx n.1(ML 357 4) in dorsal view (B), lateral view (B1), proximal view (B2) and distal view (B3), left phalanx n.2 (ML 357 22) in dorsal view (C), lateral view (C1), proximal view (C2) and distal view (C3), left phalanx n.3 (ML 357 21) in dorsal view (D), lateral view (D1), proximal view (D2) and distal view (D3), left phalanx n.4 (ML 357 24) in dorsal view (E), lateral view (E1), proximal view (E2) and distal view (E3), isolated distal ends of tarsals (ML 357 25,26) in extensor view (F, G), left ungual phalanx n.1 in dorsal view (D), mediolateral view (D1) and proximal view (D2), left ungual phalanx n.1 (ML 357- 1) in dorsal view (H), mediolateral view (H1) and proximal view (H2), right ungual phalanx n.2 (ML 357 2) in dorsal view (I), mediolateral view (I1) and proximal view (I2), right ungual phalanx n.3 (ML 357 3) in dorsal view (J), mediolateral view (J1) and proximal view (J2), left ungual phalanx n.4 in dorsal view (K), mediolateral view (K1) and proximal view (K2), ungual phalanx n.5 (ML 357 20) in dorsal view (L), mediolateral view (L1) and proximal view (L2)." figureDoi="http://doi.org/10.5281/zenodo.6530807" httpUri="https://zenodo.org/record/6530807/files/figure.png" pageId="7" pageNumber="132">Fig. 5F, G</figureCitation>
). Four manual phalanges and five ungual phalanges are associated with the carpus (
<figureCitation id="13692A33EB6D6404FC18FBAAB473FBA5" box="[1022,1135,1105,1127]" captionStart="Figure 5" captionStartId="11.[163,244,1604,1626]" captionTargetBox="[166,1446,198,1561]" captionTargetId="figure-26@11.[163,1449,195,1564]" captionTargetPageId="11" captionText="Figure 5. Manual elements of Draconyx loureiroi holotype ML 357 (ML 357 1-5, 20-26), including articulated partial carpus, manual phalanges and unguals (AL). Articulated partial carpus (ML 357 5) in extensor view (A), right phalanx n.1(ML 357 4) in dorsal view (B), lateral view (B1), proximal view (B2) and distal view (B3), left phalanx n.2 (ML 357 22) in dorsal view (C), lateral view (C1), proximal view (C2) and distal view (C3), left phalanx n.3 (ML 357 21) in dorsal view (D), lateral view (D1), proximal view (D2) and distal view (D3), left phalanx n.4 (ML 357 24) in dorsal view (E), lateral view (E1), proximal view (E2) and distal view (E3), isolated distal ends of tarsals (ML 357 25,26) in extensor view (F, G), left ungual phalanx n.1 in dorsal view (D), mediolateral view (D1) and proximal view (D2), left ungual phalanx n.1 (ML 357- 1) in dorsal view (H), mediolateral view (H1) and proximal view (H2), right ungual phalanx n.2 (ML 357 2) in dorsal view (I), mediolateral view (I1) and proximal view (I2), right ungual phalanx n.3 (ML 357 3) in dorsal view (J), mediolateral view (J1) and proximal view (J2), left ungual phalanx n.4 in dorsal view (K), mediolateral view (K1) and proximal view (K2), ungual phalanx n.5 (ML 357 20) in dorsal view (L), mediolateral view (L1) and proximal view (L2)." figureDoi="http://doi.org/10.5281/zenodo.6530807" httpUri="https://zenodo.org/record/6530807/files/figure.png" pageId="7" pageNumber="132">Fig. 5BE</figureCitation>
). Three manual phalanges (phalanx n.2 ML 35722, phalanx n.3 ML 35721 and phalanx n.4 ML 35724) and two unguals (ungual n.1 ML 3571 and ungual n.4 ML 35723) are likely to belong to the left manus (
<figureCitation id="13692A33EB6D6404FB85FB37B4BFFB23" box="[1123,1187,1228,1250]" captionStart="Figure 4" captionStartId="10.[145,223,1463,1485]" captionTargetBox="[145,1425,195,1423]" captionTargetId="figure-26@10.[145,1425,195,1423]" captionTargetPageId="10" captionText="Figure 4. Three dimensional models of axial skeleton of Draconyx loureiroi holotype ML 357 (AF):Caudal n.2 ML 357-10 in cranial view (A), left lateral view (B), caudal view (C), right lateral view (D), ventral view (E) and dorsal view (F). Caudal n.3 ML 357-11 (GL) in cranial view (G), left lateral view (H), caudal view (I), right lateral view (J), ventral view (K) and dorsal view (L). Caudal n.1 ML 357-9 (M-R): cranial view (M), left lateral view (N), caudal view (O), right lateral view (P), ventral view (Q) and dorsal view (R).Abbreviations: cf: chevron facet, k: keel, nc: neural canal, ns: neural spine, tp: transverse process." figureDoi="http://doi.org/10.5281/zenodo.6530805" httpUri="https://zenodo.org/record/6530805/files/figure.png" pageId="7" pageNumber="132">Fig. 4</figureCitation>
), while the remaining three unguals (ungual n.2 ML 3572, ungual n.3 ML 3573 and ungual n.5 ML 35720) and manual phalanx n.1 ML 3574 are attributed to the right manus. Given the fragmentary condition and the weathering of the specimen, it is not possible to identify the bones of each individual carpal. The two distal carpals are cuboid in shape and are stout elements, while the proximal carpal is more lightly built and slightly arched (
<figureCitation id="13692A33EB6D6404FAF7FA3AB57FFA15" box="[1297,1379,1473,1495]" captionStart="Figure 5" captionStartId="11.[163,244,1604,1626]" captionTargetBox="[166,1446,198,1561]" captionTargetId="figure-26@11.[163,1449,195,1564]" captionTargetPageId="11" captionText="Figure 5. Manual elements of Draconyx loureiroi holotype ML 357 (ML 357 1-5, 20-26), including articulated partial carpus, manual phalanges and unguals (AL). Articulated partial carpus (ML 357 5) in extensor view (A), right phalanx n.1(ML 357 4) in dorsal view (B), lateral view (B1), proximal view (B2) and distal view (B3), left phalanx n.2 (ML 357 22) in dorsal view (C), lateral view (C1), proximal view (C2) and distal view (C3), left phalanx n.3 (ML 357 21) in dorsal view (D), lateral view (D1), proximal view (D2) and distal view (D3), left phalanx n.4 (ML 357 24) in dorsal view (E), lateral view (E1), proximal view (E2) and distal view (E3), isolated distal ends of tarsals (ML 357 25,26) in extensor view (F, G), left ungual phalanx n.1 in dorsal view (D), mediolateral view (D1) and proximal view (D2), left ungual phalanx n.1 (ML 357- 1) in dorsal view (H), mediolateral view (H1) and proximal view (H2), right ungual phalanx n.2 (ML 357 2) in dorsal view (I), mediolateral view (I1) and proximal view (I2), right ungual phalanx n.3 (ML 357 3) in dorsal view (J), mediolateral view (J1) and proximal view (J2), left ungual phalanx n.4 in dorsal view (K), mediolateral view (K1) and proximal view (K2), ungual phalanx n.5 (ML 357 20) in dorsal view (L), mediolateral view (L1) and proximal view (L2)." figureDoi="http://doi.org/10.5281/zenodo.6530807" httpUri="https://zenodo.org/record/6530807/files/figure.png" pageId="7" pageNumber="132">Fig. 5A</figureCitation>
). The proximal end of the metacarpal preserves a concave proximal margin. Despite the poor preservation, it is possible to determine that the carpus does not include fused elements, as all of the contacts between all the identifiable carpals are clearly visible (
<figureCitation id="13692A33EB6D6404FB16F9A1B55DF9B2" box="[1264,1345,1626,1648]" captionStart="Figure 5" captionStartId="11.[163,244,1604,1626]" captionTargetBox="[166,1446,198,1561]" captionTargetId="figure-26@11.[163,1449,195,1564]" captionTargetPageId="11" captionText="Figure 5. Manual elements of Draconyx loureiroi holotype ML 357 (ML 357 1-5, 20-26), including articulated partial carpus, manual phalanges and unguals (AL). Articulated partial carpus (ML 357 5) in extensor view (A), right phalanx n.1(ML 357 4) in dorsal view (B), lateral view (B1), proximal view (B2) and distal view (B3), left phalanx n.2 (ML 357 22) in dorsal view (C), lateral view (C1), proximal view (C2) and distal view (C3), left phalanx n.3 (ML 357 21) in dorsal view (D), lateral view (D1), proximal view (D2) and distal view (D3), left phalanx n.4 (ML 357 24) in dorsal view (E), lateral view (E1), proximal view (E2) and distal view (E3), isolated distal ends of tarsals (ML 357 25,26) in extensor view (F, G), left ungual phalanx n.1 in dorsal view (D), mediolateral view (D1) and proximal view (D2), left ungual phalanx n.1 (ML 357- 1) in dorsal view (H), mediolateral view (H1) and proximal view (H2), right ungual phalanx n.2 (ML 357 2) in dorsal view (I), mediolateral view (I1) and proximal view (I2), right ungual phalanx n.3 (ML 357 3) in dorsal view (J), mediolateral view (J1) and proximal view (J2), left ungual phalanx n.4 in dorsal view (K), mediolateral view (K1) and proximal view (K2), ungual phalanx n.5 (ML 357 20) in dorsal view (L), mediolateral view (L1) and proximal view (L2)." figureDoi="http://doi.org/10.5281/zenodo.6530807" httpUri="https://zenodo.org/record/6530807/files/figure.png" pageId="7" pageNumber="132">Fig. 5A</figureCitation>
).
</paragraph>
<paragraph id="8BED36B6EB6D6409FCB5F982B133F91A" blockId="7.[827,1443,860,1893]" lastBlockId="10.[145,761,1638,1752]" lastPageId="10" lastPageNumber="135" pageId="7" pageNumber="132">
The two isolated distal metacarpals fragments are similar in shape: they are compressed along their extensorpalmar axis, broadening mediolaterally towards their distal ends. The medial and lateral ginglymi are subequal in size and preserved on their collateral ligament pits. The manual phalanges, ML 35722 and ML 35721, articulate with each other. Furthermore, phalanx ML 35722 articulates with the ungual ML 35723. Therefore, these latter manual elements compose a complete finger (
<figureCitation id="13692A33EB62640BFDDCF924B2D6F937" box="[570,714,1759,1781]" captionStart="Figure 5" captionStartId="11.[163,244,1604,1626]" captionTargetBox="[166,1446,198,1561]" captionTargetId="figure-26@11.[163,1449,195,1564]" captionTargetPageId="11" captionText="Figure 5. Manual elements of Draconyx loureiroi holotype ML 357 (ML 357 1-5, 20-26), including articulated partial carpus, manual phalanges and unguals (AL). Articulated partial carpus (ML 357 5) in extensor view (A), right phalanx n.1(ML 357 4) in dorsal view (B), lateral view (B1), proximal view (B2) and distal view (B3), left phalanx n.2 (ML 357 22) in dorsal view (C), lateral view (C1), proximal view (C2) and distal view (C3), left phalanx n.3 (ML 357 21) in dorsal view (D), lateral view (D1), proximal view (D2) and distal view (D3), left phalanx n.4 (ML 357 24) in dorsal view (E), lateral view (E1), proximal view (E2) and distal view (E3), isolated distal ends of tarsals (ML 357 25,26) in extensor view (F, G), left ungual phalanx n.1 in dorsal view (D), mediolateral view (D1) and proximal view (D2), left ungual phalanx n.1 (ML 357- 1) in dorsal view (H), mediolateral view (H1) and proximal view (H2), right ungual phalanx n.2 (ML 357 2) in dorsal view (I), mediolateral view (I1) and proximal view (I2), right ungual phalanx n.3 (ML 357 3) in dorsal view (J), mediolateral view (J1) and proximal view (J2), left ungual phalanx n.4 in dorsal view (K), mediolateral view (K1) and proximal view (K2), ungual phalanx n.5 (ML 357 20) in dorsal view (L), mediolateral view (L1) and proximal view (L2)." figureDoi="http://doi.org/10.5281/zenodo.6530807" httpUri="https://zenodo.org/record/6530807/files/figure.png" pageId="8" pageNumber="133">Fig. 5C, D, K</figureCitation>
). In general, all of the phalanges have a distal triangular section, moderately arched ventral surface and strongly inclined and robust ginglymi. The lateral ginglymus is smaller with respect to the medial one, with the exception of phalanx n.2 where they are subequal (
<figureCitation id="13692A33EB62640BFA80F93AB374F937" captionStart="Figure 5" captionStartId="11.[163,244,1604,1626]" captionTargetBox="[166,1446,198,1561]" captionTargetId="figure-26@11.[163,1449,195,1564]" captionTargetPageId="11" captionText="Figure 5. Manual elements of Draconyx loureiroi holotype ML 357 (ML 357 1-5, 20-26), including articulated partial carpus, manual phalanges and unguals (AL). Articulated partial carpus (ML 357 5) in extensor view (A), right phalanx n.1(ML 357 4) in dorsal view (B), lateral view (B1), proximal view (B2) and distal view (B3), left phalanx n.2 (ML 357 22) in dorsal view (C), lateral view (C1), proximal view (C2) and distal view (C3), left phalanx n.3 (ML 357 21) in dorsal view (D), lateral view (D1), proximal view (D2) and distal view (D3), left phalanx n.4 (ML 357 24) in dorsal view (E), lateral view (E1), proximal view (E2) and distal view (E3), isolated distal ends of tarsals (ML 357 25,26) in extensor view (F, G), left ungual phalanx n.1 in dorsal view (D), mediolateral view (D1) and proximal view (D2), left ungual phalanx n.1 (ML 357- 1) in dorsal view (H), mediolateral view (H1) and proximal view (H2), right ungual phalanx n.2 (ML 357 2) in dorsal view (I), mediolateral view (I1) and proximal view (I2), right ungual phalanx n.3 (ML 357 3) in dorsal view (J), mediolateral view (J1) and proximal view (J2), left ungual phalanx n.4 in dorsal view (K), mediolateral view (K1) and proximal view (K2), ungual phalanx n.5 (ML 357 20) in dorsal view (L), mediolateral view (L1) and proximal view (L2)." figureDoi="http://doi.org/10.5281/zenodo.6530807" httpUri="https://zenodo.org/record/6530807/files/figure.png" pageId="8" pageNumber="133">Fig. 5BE</figureCitation>
). The unguals are generally elongated and clawlike, slightly arched along the extensorpalmar axis and have a subtriangular articular facet (
<figureCitation id="13692A33EB62640BFAE8F8E6B59EF8F0" box="[1294,1410,1820,1843]" captionStart="Figure 5" captionStartId="11.[163,244,1604,1626]" captionTargetBox="[166,1446,198,1561]" captionTargetId="figure-26@11.[163,1449,195,1564]" captionTargetPageId="11" captionText="Figure 5. Manual elements of Draconyx loureiroi holotype ML 357 (ML 357 1-5, 20-26), including articulated partial carpus, manual phalanges and unguals (AL). Articulated partial carpus (ML 357 5) in extensor view (A), right phalanx n.1(ML 357 4) in dorsal view (B), lateral view (B1), proximal view (B2) and distal view (B3), left phalanx n.2 (ML 357 22) in dorsal view (C), lateral view (C1), proximal view (C2) and distal view (C3), left phalanx n.3 (ML 357 21) in dorsal view (D), lateral view (D1), proximal view (D2) and distal view (D3), left phalanx n.4 (ML 357 24) in dorsal view (E), lateral view (E1), proximal view (E2) and distal view (E3), isolated distal ends of tarsals (ML 357 25,26) in extensor view (F, G), left ungual phalanx n.1 in dorsal view (D), mediolateral view (D1) and proximal view (D2), left ungual phalanx n.1 (ML 357- 1) in dorsal view (H), mediolateral view (H1) and proximal view (H2), right ungual phalanx n.2 (ML 357 2) in dorsal view (I), mediolateral view (I1) and proximal view (I2), right ungual phalanx n.3 (ML 357 3) in dorsal view (J), mediolateral view (J1) and proximal view (J2), left ungual phalanx n.4 in dorsal view (K), mediolateral view (K1) and proximal view (K2), ungual phalanx n.5 (ML 357 20) in dorsal view (L), mediolateral view (L1) and proximal view (L2)." figureDoi="http://doi.org/10.5281/zenodo.6530807" httpUri="https://zenodo.org/record/6530807/files/figure.png" pageId="8" pageNumber="133">Fig. 5HL</figureCitation>
). The only exception, ungual n.5 (
<figureCitation id="13692A33EB62640BFB58F8C0B509F893" box="[1214,1301,1851,1873]" captionStart="Figure 5" captionStartId="11.[163,244,1604,1626]" captionTargetBox="[166,1446,198,1561]" captionTargetId="figure-26@11.[163,1449,195,1564]" captionTargetPageId="11" captionText="Figure 5. Manual elements of Draconyx loureiroi holotype ML 357 (ML 357 1-5, 20-26), including articulated partial carpus, manual phalanges and unguals (AL). Articulated partial carpus (ML 357 5) in extensor view (A), right phalanx n.1(ML 357 4) in dorsal view (B), lateral view (B1), proximal view (B2) and distal view (B3), left phalanx n.2 (ML 357 22) in dorsal view (C), lateral view (C1), proximal view (C2) and distal view (C3), left phalanx n.3 (ML 357 21) in dorsal view (D), lateral view (D1), proximal view (D2) and distal view (D3), left phalanx n.4 (ML 357 24) in dorsal view (E), lateral view (E1), proximal view (E2) and distal view (E3), isolated distal ends of tarsals (ML 357 25,26) in extensor view (F, G), left ungual phalanx n.1 in dorsal view (D), mediolateral view (D1) and proximal view (D2), left ungual phalanx n.1 (ML 357- 1) in dorsal view (H), mediolateral view (H1) and proximal view (H2), right ungual phalanx n.2 (ML 357 2) in dorsal view (I), mediolateral view (I1) and proximal view (I2), right ungual phalanx n.3 (ML 357 3) in dorsal view (J), mediolateral view (J1) and proximal view (J2), left ungual phalanx n.4 in dorsal view (K), mediolateral view (K1) and proximal view (K2), ungual phalanx n.5 (ML 357 20) in dorsal view (L), mediolateral view (L1) and proximal view (L2)." figureDoi="http://doi.org/10.5281/zenodo.6530807" httpUri="https://zenodo.org/record/6530807/files/figure.png" pageId="8" pageNumber="133">Fig. 5L</figureCitation>
), appears instead to be compressed along its extensorpalmar axis. A fracture develops mediolaterally and is slightly inclined craniocaudally. Both parts are separated by this fracture and have suffered mediolateral displacement.
</paragraph>
<caption id="DF2D663EEB6D6404FF45F8DFB0A5F8C2" box="[163,185,1682,1892]" isContinuationCaption="true" pageId="7" pageNumber="132" startId="7.[163,185,1828,1892]" targetBox="[207,555,721,1892]" targetIsTable="true" targetPageId="7">
<paragraph id="8BED36B6EB6D6404FF45F8DFB0A5F8C2" blockId="7.[163,185,1682,1892]" box="[163,185,1682,1892]" pageId="7" pageNumber="132">
<emphasis id="B926EAA4EB6D6404FF45F8DFB0A5F8DF" bold="true" box="[163,185,1808,1892]" pageId="7" pageNumber="132">Table 1</emphasis>
. Continued
</paragraph>
</caption>
<paragraph id="8BED36B6EB6D6404FF29F8D3B212F983" box="[207,526,721,1892]" pageId="7" pageNumber="132">
<table id="F952C416EB6D9BFCFF29FD2AB237F8A6" box="[207,555,721,1892]" gridcols="6" gridrows="12" pageId="7" pageNumber="132" textDirection="bu">
<tr id="356234F4EB6D9BFCFF29FD2AB0F9F8A6" box="[207,229,721,1892]" gridrow="0" pageId="7" pageNumber="132" rowspan-2="1" rowspan-4="1" textDirection="bu">
<th id="76B35D88EB6D9BFCFF29F997B0F9F8A6" box="[207,229,1644,1892]" gridcol="0" gridrow="0" pageId="7" pageNumber="132" textDirection="bu">Pedal Phalanx II- 1 ML</th>
<th id="76B35D88EB6D9BFCFF29F9E6B0F9F983" box="[207,229,1565,1601]" gridcol="1" gridrow="0" pageId="7" pageNumber="132" textDirection="bu">50 *</th>
<th id="76B35D88EB6D9BFCFF29FB88B0F9FB4E" box="[207,229,1139,1164]" gridcol="3" gridrow="0" pageId="7" pageNumber="132" textDirection="bu">48</th>
<th id="76B35D88EB6D9BFCFF29FD2AB0F9FD28" box="[207,229,721,746]" gridcol="5" gridrow="0" pageId="7" pageNumber="132" textDirection="bu">55</th>
</tr>
<tr id="356234F4EB6D9BFCFF0AFD2AB11EF8A6" box="[236,258,721,1892]" gridrow="1" pageId="7" pageNumber="132" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" textDirection="bu">
<th id="76B35D88EB6D9BFCFF0AF997B11EF8A6" box="[236,258,1644,1892]" gridcol="0" gridrow="1" pageId="7" pageNumber="132" textDirection="bu">357 17</th>
</tr>
<tr id="356234F4EB6D9BFCFEECFD2AB13CF8A6" box="[266,288,721,1892]" gridrow="2" pageId="7" pageNumber="132" textDirection="bu">
<th id="76B35D88EB6D9BFCFEECF997B13CF8A6" box="[266,288,1644,1892]" gridcol="0" gridrow="2" pageId="7" pageNumber="132" textDirection="bu">Pedal Phalanx II-2 ML</th>
<td id="76B35D88EB6D9BFCFEECF9E6B13CF983" box="[266,288,1565,1601]" gridcol="1" gridrow="2" pageId="7" pageNumber="132" textDirection="bu">40</td>
<td id="76B35D88EB6D9BFCFEECFA9BB13CFABB" box="[266,288,1376,1401]" gridcol="2" gridrow="2" pageId="7" pageNumber="132" textDirection="bu">30</td>
<td id="76B35D88EB6D9BFCFEECFB88B13CFB4E" box="[266,288,1139,1164]" gridcol="3" gridrow="2" pageId="7" pageNumber="132" textDirection="bu">45</td>
<td id="76B35D88EB6D9BFCFEECFC6AB13CFC68" box="[266,288,913,938]" gridcol="4" gridrow="2" pageId="7" pageNumber="132" textDirection="bu">20</td>
<td id="76B35D88EB6D9BFCFEECFD2AB13CFD28" box="[266,288,721,746]" gridcol="5" gridrow="2" pageId="7" pageNumber="132" textDirection="bu">40</td>
</tr>
<tr id="356234F4EB6D9BFCFECEFD2AB122F8A6" box="[296,318,721,1892]" gridrow="3" pageId="7" pageNumber="132" textDirection="bu">
<th id="76B35D88EB6D9BFCFECEFD2AB122F8A6" box="[296,318,721,1892]" colspan="6" colspanRight="5" gridcol="0" gridrow="3" pageId="7" pageNumber="132" textDirection="bu">357 14</th>
</tr>
<tr id="356234F4EB6D9BFCFEA0FD2AB140F8A6" box="[326,348,721,1892]" gridrow="4" pageId="7" pageNumber="132" textDirection="bu">
<th id="76B35D88EB6D9BFCFEA0F997B140F8A6" box="[326,348,1644,1892]" gridcol="0" gridrow="4" pageId="7" pageNumber="132" textDirection="bu">Pedal Phalanx III- 1 ML</th>
<td id="76B35D88EB6D9BFCFEA0F9E6B140F983" box="[326,348,1565,1601]" gridcol="1" gridrow="4" pageId="7" pageNumber="132" textDirection="bu">63</td>
<td id="76B35D88EB6D9BFCFEA0FA9BB140FABB" box="[326,348,1376,1401]" gridcol="2" gridrow="4" pageId="7" pageNumber="132" textDirection="bu">45</td>
<td id="76B35D88EB6D9BFCFEA0FB88B140FB4E" box="[326,348,1139,1164]" gridcol="3" gridrow="4" pageId="7" pageNumber="132" textDirection="bu">69</td>
<td id="76B35D88EB6D9BFCFEA0FC6AB140FC68" box="[326,348,913,938]" gridcol="4" gridrow="4" pageId="7" pageNumber="132" textDirection="bu">25</td>
<td id="76B35D88EB6D9BFCFEA0FD2AB140FD28" box="[326,348,721,746]" gridcol="5" gridrow="4" pageId="7" pageNumber="132" textDirection="bu">45</td>
</tr>
<tr id="356234F4EB6D9BFCFE85FD2AB165F8A6" box="[355,377,721,1892]" gridrow="5" pageId="7" pageNumber="132" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" textDirection="bu">
<th id="76B35D88EB6D9BFCFE85F997B165F8A6" box="[355,377,1644,1892]" gridcol="0" gridrow="5" pageId="7" pageNumber="132" textDirection="bu">357 13</th>
</tr>
<tr id="356234F4EB6D9BFCFE67FD2AB18BF8A6" box="[385,407,721,1892]" gridrow="6" pageId="7" pageNumber="132" rowspan-5="1" textDirection="bu">
<th id="76B35D88EB6D9BFCFE67F997B18BF8A6" box="[385,407,1644,1892]" gridcol="0" gridrow="6" pageId="7" pageNumber="132" textDirection="bu">Pedal Phalanx III- 3 ML</th>
<td id="76B35D88EB6D9BFCFE67F9E6B18BF983" box="[385,407,1565,1601]" gridcol="1" gridrow="6" pageId="7" pageNumber="132" textDirection="bu">35</td>
<td id="76B35D88EB6D9BFCFE67FA9BB18BFABB" box="[385,407,1376,1401]" gridcol="2" gridrow="6" pageId="7" pageNumber="132" textDirection="bu">35</td>
<td id="76B35D88EB6D9BFCFE67FB88B18BFB4E" box="[385,407,1139,1164]" gridcol="3" gridrow="6" pageId="7" pageNumber="132" textDirection="bu">28</td>
<td id="76B35D88EB6D9BFCFE67FC6AB18BFC68" box="[385,407,913,938]" gridcol="4" gridrow="6" pageId="7" pageNumber="132" textDirection="bu">20</td>
</tr>
<tr id="356234F4EB6D9BFCFE78FD2AB1A8F8A6" box="[414,436,721,1892]" gridrow="7" pageId="7" pageNumber="132" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" textDirection="bu">
<th id="76B35D88EB6D9BFCFE78F997B1A8F8A6" box="[414,436,1644,1892]" gridcol="0" gridrow="7" pageId="7" pageNumber="132" textDirection="bu">357 15</th>
</tr>
<tr id="356234F4EB6D9BFCFE5AFD2AB1CEF8A6" box="[444,466,721,1892]" gridrow="8" pageId="7" pageNumber="132" rowspan-3="1" rowspan-5="1" textDirection="bu">
<th id="76B35D88EB6D9BFCFE5AF997B1CEF8A6" box="[444,466,1644,1892]" gridcol="0" gridrow="8" pageId="7" pageNumber="132" textDirection="bu">Pedal Ungual II ML</th>
<td id="76B35D88EB6D9BFCFE5AF9E6B1CEF983" box="[444,466,1565,1601]" gridcol="1" gridrow="8" pageId="7" pageNumber="132" textDirection="bu">60</td>
<td id="76B35D88EB6D9BFCFE5AFA9BB1CEFABB" box="[444,466,1376,1401]" gridcol="2" gridrow="8" pageId="7" pageNumber="132" textDirection="bu">25</td>
<td id="76B35D88EB6D9BFCFE5AFC6AB1CEFC68" box="[444,466,913,938]" gridcol="4" gridrow="8" pageId="7" pageNumber="132" textDirection="bu">30</td>
</tr>
<tr id="356234F4EB6D9BFCFE3CFD2AB1ECF8A6" box="[474,496,721,1892]" gridrow="9" pageId="7" pageNumber="132" textDirection="bu">
<th id="76B35D88EB6D9BFCFE3CFD2AB1ECF8A6" box="[474,496,721,1892]" colspan="6" colspanRight="5" gridcol="0" gridrow="9" pageId="7" pageNumber="132" textDirection="bu">357 16</th>
</tr>
<tr id="356234F4EB6D9BFCFE1EFD2AB212F8A6" box="[504,526,721,1892]" gridrow="10" pageId="7" pageNumber="132" rowspan-3="1" rowspan-5="1" textDirection="bu">
<th id="76B35D88EB6D9BFCFE1EF997B212F8A6" box="[504,526,1644,1892]" gridcol="0" gridrow="10" pageId="7" pageNumber="132" textDirection="bu">Pedal Ungual III ML</th>
<td id="76B35D88EB6D9BFCFE1EF9E6B212F983" box="[504,526,1565,1601]" gridcol="1" gridrow="10" pageId="7" pageNumber="132" textDirection="bu">60</td>
<paragraph id="8BED36B6EB6D6404FE1EFA9BB237F88C" box="[504,555,913,1870]" pageId="7" pageNumber="132">
<td id="76B35D88EB6D9BFCFE1EFA9BB212FABB" box="[504,526,1376,1401]" gridcol="2" gridrow="10" pageId="7" pageNumber="132" textDirection="bu">30</td>
<td id="76B35D88EB6D9BFCFE1EFC6AB212FC68" box="[504,526,913,938]" gridcol="4" gridrow="10" pageId="7" pageNumber="132" textDirection="bu">25</td>
<tr id="356234F4EB6D9BFCFDF3FD2AB237F8A6" box="[533,555,721,1892]" gridrow="11" pageId="7" pageNumber="132" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" textDirection="bu">
<th id="76B35D88EB6D9BFCFDF3F997B237F8A6" box="[533,555,1644,1892]" gridcol="0" gridrow="11" pageId="7" pageNumber="132" textDirection="bu">
357
<paragraph id="8BED36B6EB6D6404FDF3F8E1B237F8DB" box="[533,555,1794,1832]" pageId="7" pageNumber="132"> 19</paragraph>
</th>
</tr>
</paragraph>
</tr>
</table>
</paragraph>
<caption id="DF2D663EEB62640BFF74F9B1B307F958" ID-DOI="http://doi.org/10.5281/zenodo.6530801" ID-Zenodo-Dep="6530801" httpUri="https://zenodo.org/record/6530801/files/figure.png" pageId="8" pageNumber="133" startId="8.[146,225,1610,1632]" targetBox="[353,1217,199,1567]" targetPageId="8">
<paragraph id="8BED36B6EB62640BFF74F9B1B307F958" blockId="8.[145,1425,1610,1690]" pageId="8" pageNumber="133">
<emphasis id="B926EAA4EB62640BFF74F9B1B0E7F99D" bold="true" box="[146,251,1610,1632]" pageId="8" pageNumber="133">Figure 2.</emphasis>
Two maxillary teeth of
<taxonomicName id="4C524D35EB62640BFE18F9B1B2DEF9A2" authorityName="Mateus &amp; Antunes" authorityYear="2001" box="[510,706,1610,1632]" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="133" phylum="Chordata" rank="species" species="loureiroi">
<emphasis id="B926EAA4EB62640BFE18F9B1B2DEF9A2" box="[510,706,1610,1632]" italics="true" pageId="8" pageNumber="133">Draconyx loureiroi</emphasis>
</taxonomicName>
holotype ML 357 in labial and lingual views. Completely preserved maxillary in labial (A) and lingual (C) views; isolated maxillary crown in labial (B) and lingual (D) views. Abbreviations: ne: neck, os: occlusal surface, pr: primary ridge, tr: tertiary ridges.
</paragraph>
</caption>
<caption id="DF2D663EEB63640AFF45F909B3ABF8A2" ID-DOI="http://doi.org/10.5281/zenodo.6530803" ID-Zenodo-Dep="6530803" httpUri="https://zenodo.org/record/6530803/files/figure.png" pageId="9" pageNumber="134" startId="9.[163,242,1778,1800]" targetBox="[227,1379,195,1738]" targetPageId="9">
<paragraph id="8BED36B6EB63640AFF45F909B3ABF8A2" blockId="9.[163,1443,1778,1888]" pageId="9" pageNumber="134">
<emphasis id="B926EAA4EB63640AFF45F909B110F8C5" bold="true" box="[163,268,1778,1800]" pageId="9" pageNumber="134">Figure 3.</emphasis>
General overview of
<taxonomicName id="4C524D35EB63640AFE14F909B2AAF8CA" authorityName="Mateus &amp; Antunes" authorityYear="2001" box="[498,694,1778,1800]" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="134" phylum="Chordata" rank="species" species="loureiroi">
<emphasis id="B926EAA4EB63640AFE14F909B2AAF8CA" box="[498,694,1778,1800]" italics="true" pageId="9" pageNumber="134">Draconyx loureiroi</emphasis>
</taxonomicName>
holotype ML 357 post-cranial skeleton. Caudal vertebrae (K-M) in left lateral view. Right femur (AE) in frontal (A), medial (B), caudal (C), lateral (D) and distal (E) views. Right tibia in medial (F) and lateral (G) views. Proximal end of the fibula in medial (I) and lateral (J) views. Right articulated pes in dorsal (N) and medial (Q) view. Pedal digit III (O) and pedal digit II (P) in dorsal view.
</paragraph>
</caption>
<caption id="DF2D663EEB606409FF77FA4CB58FF980" ID-DOI="http://doi.org/10.5281/zenodo.6530805" ID-Zenodo-Dep="6530805" httpUri="https://zenodo.org/record/6530805/files/figure.png" pageId="10" pageNumber="135" startId="10.[145,223,1463,1485]" targetBox="[145,1425,195,1423]" targetPageId="10">
<paragraph id="8BED36B6EB606409FF77FA4CB58FF980" blockId="10.[145,1427,1463,1602]" pageId="10" pageNumber="135">
<emphasis id="B926EAA4EB606409FF77FA4CB0EBFA0E" bold="true" box="[145,247,1463,1485]" pageId="10" pageNumber="135">Figure 4.</emphasis>
Three dimensional models of axial skeleton of
<taxonomicName id="4C524D35EB606409FD3FFA4CB38BFA0F" authorityName="Mateus &amp; Antunes" authorityYear="2001" box="[729,919,1463,1485]" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="10" pageNumber="135" phylum="Chordata" rank="species" species="loureiroi">
<emphasis id="B926EAA4EB606409FD3FFA4CB38BFA0F" box="[729,919,1463,1485]" italics="true" pageId="10" pageNumber="135">Draconyx loureiroi</emphasis>
</taxonomicName>
holotype ML 357 (AF): Caudal n.2 ML 357-10 in cranial view (A), left lateral view (B), caudal view (C), right lateral view (D), ventral view (E) and dorsal view (F). Caudal n.3 ML 357-11 (GL) in cranial view (G), left lateral view (H), caudal view (I), right lateral view (J), ventral view (K) and dorsal view (L). Caudal n.1 ML 357-9 (M-R): cranial view (M), left lateral view (N), caudal view (O), right lateral view (P), ventral view (Q) and dorsal view (R). Abbreviations: cf: chevron facet, k: keel, nc: neural canal, ns: neural spine, tp: transverse process.
</paragraph>
</caption>
<paragraph id="8BED36B6EB60640FFF77F907B162F91A" blockId="10.[145,760,1788,1902]" lastBlockId="12.[145,762,1301,1752]" lastPageId="12" lastPageNumber="137" pageId="10" pageNumber="135">
<emphasis id="B926EAA4EB606409FF77F907B112F8D3" box="[145,270,1788,1809]" italics="true" pageId="10" pageNumber="135">R e m a r k s:</emphasis>
D e s p i t e t h e p o o r p r e s e r v a t i o n o f the carpus of ML 3 5 7, certain features allow comparisons with other taxa. The small metacarpals are not significantly different from those of other iguanodontians (Norman, 2004). However, the carpus is constituted by isolated blocky elements, resembling the condition in dryosaurids and
<taxonomicName id="4C524D35EB606409FB56F95FB58DF97B" box="[1200,1425,1700,1721]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" kingdom="Animalia" order="Ornithischia" pageId="10" pageNumber="135" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B926EAA4EB606409FB56F95FB547F97B" box="[1200,1371,1700,1721]" italics="true" pageId="10" pageNumber="135">Tenontosaurus</emphasis>
spp.
</taxonomicName>
(
<bibRefCitation id="EFC34B47EB606409FCD4F939B3CDF915" author="Dodson P" box="[818,977,1730,1752]" pageId="10" pageNumber="135" pagination="81 - 85" refId="ref16752" refString="Dodson P. 1980. Comparative osteology of the American ornithopods Camptosaurus and Tenontosaurus. Memoirs of the Society of Geology, France 139: 81 - 85." type="journal article" year="1980">Dodson, 1980</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB606409FC06F939B465F915" author="Galton PM" box="[992,1145,1730,1752]" pageId="10" pageNumber="135" pagination="271 - 312" refId="ref17047" refString="Galton PM. 1981. Dryosaurus, a hypsilophodontid dinosaur from the Upper Jurassic of North America and Africa postcranial skeleton. Palaontologische Zeitschrift 55: 271 - 312." type="journal article" year="1981">Galton, 1981</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB606409FB61F938B53FF915" author="Forster CA" box="[1159,1315,1730,1752]" pageId="10" pageNumber="135" pagination="273 - 294" refId="ref16896" refString="Forster CA. 1990. The postcranial skeleton of the ornithopod dinosaur Tenontosaurus tilletti. Journal of Vertebrate Paleontology 10: 273 - 294." type="journal article" year="1990">Forster, 1990</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB606409FAD7F939B3B4F934" author="Winkler DA &amp; Murry PA &amp; Jacobs LL" pageId="10" pageNumber="135" pagination="330 - 348" refId="ref19543" refString="Winkler DA, Murry PA, Jacobs LL. 1997. A new species of Tenontosaurus (Dinosauria: Ornithopoda) from the Early Cretaceous of Texas. Journal of Vertebrate Paleontology 17: 330 - 348." type="journal article" year="1997">
Winkler
<emphasis id="B926EAA4EB606409FCCFF919B37EF934" box="[809,866,1761,1783]" italics="true" pageId="10" pageNumber="135">et al.</emphasis>
, 1997
</bibRefCitation>
). The distal carpals of ML 357 differ from the ones of
<taxonomicName id="4C524D35EB606409FC4CF8FBB4B8F8D6" box="[938,1188,1791,1813]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="10" pageNumber="135" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B926EAA4EB606409FC4CF8FBB4B8F8D6" box="[938,1188,1791,1813]" italics="true" pageId="10" pageNumber="135">Camptosaurus dispar</emphasis>
</taxonomicName>
, which are arranged in two co-ossified and highly interlocked blocks. Furthermore, the carpus of ML 357 does not exhibit the total ossified condition present in
<taxonomicName id="4C524D35EB60640FFAEBF8A0B215FAE9" authority=", Magnamanus" authorityName="Magnamanus" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="12" lastPageNumber="137" order="Ornithischia" pageId="10" pageNumber="135" phylum="Chordata" rank="species" species="bernissartensis">
<emphasis id="B926EAA4EB60640FFAEBF8A0B152FAE8" italics="true" lastPageId="12" lastPageNumber="137" pageId="10" pageNumber="135">Iguanodon bernissartensis</emphasis>
,
<emphasis id="B926EAA4EB66640FFE86FAEDB215FAE9" box="[352,521,1302,1323]" italics="true" pageId="12" pageNumber="137">Magnamanus</emphasis>
</taxonomicName>
<emphasis id="B926EAA4EB66640FFDF2FAEDB28DFAE9" box="[532,657,1302,1323]" italics="true" pageId="12" pageNumber="137">soriaensis</emphasis>
<bibRefCitation id="EFC34B47EB66640FFD7DFAEEB10CFA88" author="Vidarte CF &amp; Calvo MM &amp; Fuentes FM &amp; Fuentes MM" pageId="12" pageNumber="137" pagination="407 - 446" refId="ref19357" refString="Vidarte CF, Calvo MM, Fuentes FM, Fuentes MM. 2016. Un nuevo dinosaurio estiracosterno (Ornithopoda: Ankylopollexia) del Cretacico Inferior de Espana. Spanish Journal of Palaeontology 31: 407 - 446." type="journal article" year="2016">
Vidarte
<emphasis id="B926EAA4EB66640FFF77FACEB0D6FA8B" box="[145,202,1332,1354]" italics="true" pageId="12" pageNumber="137">et al.</emphasis>
, 2016
</bibRefCitation>
,
<taxonomicName id="4C524D35EB66640FFEFBFACFB29CFA8B" authorityName="Mantell" authorityYear="1825" box="[285,640,1332,1353]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="137" phylum="Chordata" rank="species" species="atherfieldensis">
<emphasis id="B926EAA4EB66640FFEFBFACFB29CFA8B" box="[285,640,1332,1353]" italics="true" pageId="12" pageNumber="137">Mantellisaurus atherfieldensis</emphasis>
</taxonomicName>
and other styracosternans (
<figureCitation id="13692A33EB66640FFEBFFAA9B1C9FAAA" box="[345,469,1362,1384]" captionStart="Figure 12" captionStartId="22.[145,223,1787,1809]" captionTargetBox="[210,1361,198,1744]" captionTargetId="figure-26@22.[206,1364,195,1747]" captionTargetPageId="22" captionText="Figure 12. Time calibrated strict consensus tree of the analysis including Draconyx loureiroi in the dataset of Xu et al. (2018) (A). Bootstrap value in blue, Bremer support in red. Red colour is associated with an increasing Homoplasy Concentration index. XY graphs of Average Evolutionary Rates (B) and Average Homoplasy Rates (C) through time. Note the coinciding peaks of both AER and AHR. Silhouettes from phylopic.org. Credit: Michael Keesey, Matthew Dempsey and Pete Buchholz." figureDoi="http://doi.org/10.5281/zenodo.6530821" httpUri="https://zenodo.org/record/6530821/files/figure.png" pageId="12" pageNumber="137">Fig. 12AJ</figureCitation>
) (
<bibRefCitation id="EFC34B47EB66640FFE0AFAA9B29BFAAA" author="Dodson P" box="[492,647,1362,1384]" pageId="12" pageNumber="137" pagination="81 - 85" refId="ref16752" refString="Dodson P. 1980. Comparative osteology of the American ornithopods Camptosaurus and Tenontosaurus. Memoirs of the Society of Geology, France 139: 81 - 85." type="journal article" year="1980">Dodson, 1980</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB66640FFD75FAA8B0D6FA44" author="Norman DB" pageId="12" pageNumber="137" pagination="1 - 105" refId="ref18270" refString="Norman DB. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman, 1980</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB66640FFF3CFA8AB10FFA45" author="Norman DB" box="[218,275,1393,1415]" pageId="12" pageNumber="137" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
, 2004, 2011, 2015,
<bibRefCitation id="EFC34B47EB66640FFE19FA8AB2F5FA45" author="Vidarte CF &amp; Calvo MM &amp; Fuentes FM &amp; Fuentes MM" box="[511,745,1393,1415]" pageId="12" pageNumber="137" pagination="407 - 446" refId="ref19357" refString="Vidarte CF, Calvo MM, Fuentes FM, Fuentes MM. 2016. Un nuevo dinosaurio estiracosterno (Ornithopoda: Ankylopollexia) del Cretacico Inferior de Espana. Spanish Journal of Palaeontology 31: 407 - 446." type="journal article" year="2016">
Vidarte
<emphasis id="B926EAA4EB66640FFD85FA89B2BCFA44" box="[611,672,1393,1415]" italics="true" pageId="12" pageNumber="137">et al.</emphasis>
, 2016
</bibRefCitation>
). The carpals and the manual unguals are similar to the ones present in
<taxonomicName id="4C524D35EB66640FFE9EFA54B2E4FA06" authority=", Cumnoria" authorityName="Cumnoria" box="[376,760,1454,1476]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="137" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B926EAA4EB66640FFE9EFA54B269FA01" box="[376,629,1454,1476]" italics="true" pageId="12" pageNumber="137">Camptosaurus dispar</emphasis>
,
<emphasis id="B926EAA4EB66640FFD65FA54B2E4FA06" box="[643,760,1455,1476]" italics="true" pageId="12" pageNumber="137">Cumnoria</emphasis>
</taxonomicName>
<taxonomicName id="4C524D35EB66640FFF77FA36B100FA20" authorityName="Hulke" authorityYear="1880" box="[145,284,1485,1506]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="137" phylum="Chordata" rank="species" species="prestwichii">
<emphasis id="B926EAA4EB66640FFF77FA36B100FA20" box="[145,284,1485,1506]" italics="true" pageId="12" pageNumber="137">prestwichii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D35EB66640FFE87FA36B264FA20" baseAuthorityName="Carpenter &amp; Wilson" baseAuthorityYear="2008" box="[353,632,1485,1506]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="137" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="B926EAA4EB66640FFE87FA36B264FA20" box="[353,632,1485,1506]" italics="true" pageId="12" pageNumber="137">Uteodon aphanoecetes</emphasis>
</taxonomicName>
, differing from the ones of more derived styracosternans (i.e.
<taxonomicName id="4C524D35EB66640FFF77F9F1B26AF9E2" authority=", Magnamanus" authorityName="Magnamanus" box="[145,630,1546,1568]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="137" phylum="Chordata" rank="species" species="bernissartensis">
<emphasis id="B926EAA4EB66640FFF77F9F1B1DAF9DD" box="[145,454,1546,1567]" italics="true" pageId="12" pageNumber="137">Iguanodon bernissartensis</emphasis>
,
<emphasis id="B926EAA4EB66640FFE33F9F0B26AF9E2" box="[469,630,1547,1568]" italics="true" pageId="12" pageNumber="137">Magnamanus</emphasis>
</taxonomicName>
<emphasis id="B926EAA4EB66640FFD98F9F0B2EEF9E2" box="[638,754,1547,1568]" italics="true" pageId="12" pageNumber="137">soriaensis</emphasis>
,
<taxonomicName id="4C524D35EB66640FFF77F9D1B228F9FD" authorityName="Mantell" authorityYear="1825" box="[145,564,1577,1599]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="137" phylum="Chordata" rank="species" species="atherfieldensis">
<emphasis id="B926EAA4EB66640FFF77F9D1B228F9FD" box="[145,564,1577,1599]" italics="true" pageId="12" pageNumber="137">Mantellisaurus atherfieldensis</emphasis>
</taxonomicName>
and all other hadrosauriformes) in being, respectively, more arched and more claw-like instead of extremely compressed and hoof-like (
<figureCitation id="13692A33EB66640FFEA4F97EB1D9F958" box="[322,453,1669,1691]" captionStart="Figure 12" captionStartId="22.[145,223,1787,1809]" captionTargetBox="[210,1361,198,1744]" captionTargetId="figure-26@22.[206,1364,195,1747]" captionTargetPageId="22" captionText="Figure 12. Time calibrated strict consensus tree of the analysis including Draconyx loureiroi in the dataset of Xu et al. (2018) (A). Bootstrap value in blue, Bremer support in red. Red colour is associated with an increasing Homoplasy Concentration index. XY graphs of Average Evolutionary Rates (B) and Average Homoplasy Rates (C) through time. Note the coinciding peaks of both AER and AHR. Silhouettes from phylopic.org. Credit: Michael Keesey, Matthew Dempsey and Pete Buchholz." figureDoi="http://doi.org/10.5281/zenodo.6530821" httpUri="https://zenodo.org/record/6530821/files/figure.png" pageId="12" pageNumber="137">Fig. 12AJ</figureCitation>
) (
<bibRefCitation id="EFC34B47EB66640FFE06F97EB2EDF958" author="Galton PM &amp; Powell HP" box="[480,753,1669,1691]" pageId="12" pageNumber="137" pagination="411 - 443" refId="ref17215" refString="Galton PM, Powell HP. 1980. The ornithischian dinosaur Camptosaurus prestwichii from the Upper Jurassic of England. Palaeontology 23: 411 - 443." type="journal article" year="1980">Galton &amp; Powell, 1980</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB66640FFF77F95FB129F97B" author="Norman DB" box="[145,309,1700,1721]" pageId="12" pageNumber="137" pagination="1 - 105" refId="ref18270" refString="Norman DB. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman, 1980</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB66640FFEA7F95FB165F978" author="Norman DB" box="[321,377,1700,1722]" pageId="12" pageNumber="137" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
, 2004;
<bibRefCitation id="EFC34B47EB66640FFE2FF95FB2EFF97B" author="Carpenter K &amp; Wilson Y" box="[457,755,1700,1722]" pageId="12" pageNumber="137" pagination="227 - 264" refId="ref16499" refString="Carpenter K, Wilson Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter &amp; Wilson, 2008</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB66640FFF77F939B172F91A" author="Vidarte CF &amp; Calvo MM &amp; Fuentes FM &amp; Fuentes MM" box="[145,366,1730,1752]" pageId="12" pageNumber="137" pagination="407 - 446" refId="ref19357" refString="Vidarte CF, Calvo MM, Fuentes FM, Fuentes MM. 2016. Un nuevo dinosaurio estiracosterno (Ornithopoda: Ankylopollexia) del Cretacico Inferior de Espana. Spanish Journal of Palaeontology 31: 407 - 446." type="journal article" year="2016">
Vidarte
<emphasis id="B926EAA4EB66640FFF16F938B135F915" box="[240,297,1730,1752]" italics="true" pageId="12" pageNumber="137">et al.</emphasis>
, 2016
</bibRefCitation>
).
</paragraph>
<caption id="DF2D663EEB616408FF45F9BFB4A1F8BD" ID-DOI="http://doi.org/10.5281/zenodo.6530807" ID-Zenodo-Dep="6530807" httpUri="https://zenodo.org/record/6530807/files/figure.png" pageId="11" pageNumber="136" startId="11.[163,244,1604,1626]" targetBox="[166,1446,198,1561]" targetPageId="11">
<paragraph id="8BED36B6EB616408FF45F9BFB4A1F8BD" blockId="11.[163,1443,1604,1919]" pageId="11" pageNumber="136">
<emphasis id="B926EAA4EB616408FF45F9BFB113F99B" bold="true" box="[163,271,1604,1626]" pageId="11" pageNumber="136">Figure 5.</emphasis>
Manual elements of
<taxonomicName id="4C524D35EB616408FE1CF9BEB2DFF998" authorityName="Mateus &amp; Antunes" authorityYear="2001" box="[506,707,1604,1626]" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="11" pageNumber="136" phylum="Chordata" rank="species" species="loureiroi">
<emphasis id="B926EAA4EB616408FE1CF9BEB2DFF998" box="[506,707,1604,1626]" italics="true" pageId="11" pageNumber="136">Draconyx loureiroi</emphasis>
</taxonomicName>
holotype ML 357 (ML 357 1-5, 20-26), including articulated partial carpus, manual phalanges and unguals (AL). Articulated partial carpus (ML 357 5) in extensor view (A), right phalanx n.1(ML 357 4) in dorsal view (B), lateral view (B1), proximal view (B2) and distal view (B3), left phalanx n.2 (ML 357 22) in dorsal view (C), lateral view (C1), proximal view (C2) and distal view (C3), left phalanx n.3 (ML 357 21) in dorsal view (D), lateral view (D1), proximal view (D2) and distal view (D3), left phalanx n.4 (ML 357 24) in dorsal view (E), lateral view (E1), proximal view (E2) and distal view (E3), isolated distal ends of tarsals (ML 357 25,26) in extensor view (F, G), left ungual phalanx n.1 in dorsal view (D), mediolateral view (D1) and proximal view (D2), left ungual phalanx n.1 (ML 357- 1) in dorsal view (H), mediolateral view (H1) and proximal view (H2), right ungual phalanx n.2 (ML 357 2) in dorsal view (I), mediolateral view (I1) and proximal view (I2), right ungual phalanx n.3 (ML 357 3) in dorsal view (J), mediolateral view (J1) and proximal view (J2), left ungual phalanx n.4 in dorsal view (K), mediolateral view (K1) and proximal view (K2), ungual phalanx n.5 (ML 357 20) in dorsal view (L), mediolateral view (L1) and proximal view (L2).
</paragraph>
</caption>
<caption id="DF2D663EEB66640FFF77FBB3B31FFB74" ID-DOI="http://doi.org/10.5281/zenodo.6530809" ID-Zenodo-Dep="6530809" httpUri="https://zenodo.org/record/6530809/files/figure.png" pageId="12" pageNumber="137" startId="12.[145,224,1096,1118]" targetBox="[145,1425,195,1056]" targetPageId="12">
<paragraph id="8BED36B6EB66640FFF77FBB3B31FFB74" blockId="12.[145,1425,1096,1206]" pageId="12" pageNumber="137">
<emphasis id="B926EAA4EB66640FFF77FBB3B0E7FB9F" bold="true" box="[145,251,1096,1118]" pageId="12" pageNumber="137">Figure 6.</emphasis>
Three dimensional models of limb bones elements of
<taxonomicName id="4C524D35EB66640FFCDEFBB2B3E0FB9C" authorityName="Mateus &amp; Antunes" authorityYear="2001" box="[824,1020,1096,1118]" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="137" phylum="Chordata" rank="species" species="loureiroi">
<emphasis id="B926EAA4EB66640FFCDEFBB2B3E0FB9C" box="[824,1020,1096,1118]" italics="true" pageId="12" pageNumber="137">Draconyx loureiroi</emphasis>
</taxonomicName>
holotype ML 357 (ML 357 6-8), femur (AE) and tibia and proximal end of fibula (FJ). Femur (ML 357 6) in cranial view (A), medial view (B), caudal view (C), lateral view (D) and distal view (E). Tibia (ML 357 7) and proximal end of fibula (ML 357 8) in cranial view (F), medial view (G), caudal view (H), lateral (I) and proximal (J) views.
</paragraph>
</caption>
<paragraph id="8BED36B6EB66640EFF77F8F9B200FE94" blockId="12.[145,761,1793,1876]" lastBlockId="13.[163,778,197,342]" lastPageId="13" lastPageNumber="138" pageId="12" pageNumber="137">
<emphasis id="B926EAA4EB66640FFF77F8F9B25BF8D4" box="[145,583,1793,1815]" italics="true" pageId="12" pageNumber="137">
Femur ML 3576 (
<figureCitation id="13692A33EB66640FFE88F8F9B1F4F8D4" box="[366,488,1793,1815]" captionStart="Figure 3" captionStartId="9.[163,242,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@9.[227,1379,195,1738]" captionTargetPageId="9" captionText="Figure 3. General overview of Draconyx loureiroi holotype ML 357 post-cranial skeleton. Caudal vertebrae (K-M) in left lateral view. Right femur (AE) in frontal (A), medial (B), caudal (C), lateral (D) and distal (E) views. Right tibia in medial (F) and lateral (G) views. Proximal end of the fibula in medial (I) and lateral (J) views. Right articulated pes in dorsal (N) and medial (Q) view. Pedal digit III (O) and pedal digit II (P) in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6530803" httpUri="https://zenodo.org/record/6530803/files/figure.png" pageId="12" pageNumber="137">Figs 3AE</figureCitation>
,
<figureCitation id="13692A33EB66640FFE11F8FAB22BF8D4" box="[503,567,1793,1814]" captionStart="Figure 6" captionStartId="12.[145,224,1096,1118]" captionTargetBox="[145,1425,195,1056]" captionTargetId="figure-432@12.[145,1425,195,1056]" captionTargetPageId="12" captionText="Figure 6. Three dimensional models of limb bones elements of Draconyx loureiroi holotype ML 357 (ML 357 6-8), femur (AE) and tibia and proximal end of fibula (FJ). Femur (ML 357 6) in cranial view (A), medial view (B), caudal view (C), lateral view (D) and distal view (E). Tibia (ML 357 7) and proximal end of fibula (ML 357 8) in cranial view (F), medial view (G), caudal view (H), lateral (I) and proximal (J) views." figureDoi="http://doi.org/10.5281/zenodo.6530809" httpUri="https://zenodo.org/record/6530809/files/figure.png" pageId="12" pageNumber="137">6AE</figureCitation>
):
</emphasis>
The preserved right femur consists of the heavily eroded distalmost part of the shaft and the distal epiphysis. The proximalmost part of the preserved shaft is mediolaterally crushed and compressed, due to taphonomic processes. The femur is strongly bowed craniocaudally; the section of the shaft was subcircular but is heavily distorted proximally by the compression and damaged distally (
<figureCitation id="13692A33EB66640FFAEFFA55B59DFA06" box="[1289,1409,1454,1476]" captionStart="Figure 6" captionStartId="12.[145,224,1096,1118]" captionTargetBox="[145,1425,195,1056]" captionTargetId="figure-432@12.[145,1425,195,1056]" captionTargetPageId="12" captionText="Figure 6. Three dimensional models of limb bones elements of Draconyx loureiroi holotype ML 357 (ML 357 6-8), femur (AE) and tibia and proximal end of fibula (FJ). Femur (ML 357 6) in cranial view (A), medial view (B), caudal view (C), lateral view (D) and distal view (E). Tibia (ML 357 7) and proximal end of fibula (ML 357 8) in cranial view (F), medial view (G), caudal view (H), lateral (I) and proximal (J) views." figureDoi="http://doi.org/10.5281/zenodo.6530809" httpUri="https://zenodo.org/record/6530809/files/figure.png" pageId="12" pageNumber="137">Fig. 6B, D</figureCitation>
). The distal epiphysis is subrectangular in shape in distal view, extending slightly more mediolaterally than craniocaudally. The two distal condyles are preserved, appearing subequal in size with the lateral condyle slightly larger than the medial one (
<figureCitation id="13692A33EB66640FFCD7F99DB398F9BE" box="[817,900,1638,1660]" captionStart="Figure 6" captionStartId="12.[145,224,1096,1118]" captionTargetBox="[145,1425,195,1056]" captionTargetId="figure-432@12.[145,1425,195,1056]" captionTargetPageId="12" captionText="Figure 6. Three dimensional models of limb bones elements of Draconyx loureiroi holotype ML 357 (ML 357 6-8), femur (AE) and tibia and proximal end of fibula (FJ). Femur (ML 357 6) in cranial view (A), medial view (B), caudal view (C), lateral view (D) and distal view (E). Tibia (ML 357 7) and proximal end of fibula (ML 357 8) in cranial view (F), medial view (G), caudal view (H), lateral (I) and proximal (J) views." figureDoi="http://doi.org/10.5281/zenodo.6530809" httpUri="https://zenodo.org/record/6530809/files/figure.png" pageId="12" pageNumber="137">Fig. 6E</figureCitation>
). A deep, extensive, fully open and U-shaped extensor groove separates the two condyles cranially (
<figureCitation id="13692A33EB66640FFCD4F95FB39BF978" box="[818,903,1700,1722]" captionStart="Figure 6" captionStartId="12.[145,224,1096,1118]" captionTargetBox="[145,1425,195,1056]" captionTargetId="figure-432@12.[145,1425,195,1056]" captionTargetPageId="12" captionText="Figure 6. Three dimensional models of limb bones elements of Draconyx loureiroi holotype ML 357 (ML 357 6-8), femur (AE) and tibia and proximal end of fibula (FJ). Femur (ML 357 6) in cranial view (A), medial view (B), caudal view (C), lateral view (D) and distal view (E). Tibia (ML 357 7) and proximal end of fibula (ML 357 8) in cranial view (F), medial view (G), caudal view (H), lateral (I) and proximal (J) views." figureDoi="http://doi.org/10.5281/zenodo.6530809" httpUri="https://zenodo.org/record/6530809/files/figure.png" pageId="12" pageNumber="137">Fig. 6E</figureCitation>
). The cranial process of the lateral condyle is rounded and deflected caudally. The caudal fingerlike process of the lateral condyle (condylid, according to
<bibRefCitation id="EFC34B47EB66640FFCA3F8FBB437F8D7" author="Bertozzo F &amp; Dalla Vecchia FM &amp; Fabbri M" box="[837,1067,1792,1813]" pageId="12" pageNumber="137" pagination="3403" refId="ref16369" refString="Bertozzo F, Dalla Vecchia FM, Fabbri M. 2017. The Venice specimen of Ouranosaurus nigeriensis (Dinosauria, Ornithopoda). PeerJ 5: e 3403." type="journal article" year="2017">
Bertozzo
<emphasis id="B926EAA4EB66640FFC56F8FBB3FBF8D7" box="[944,999,1792,1813]" italics="true" pageId="12" pageNumber="137">et al.</emphasis>
, 2017
</bibRefCitation>
) is not preserved, being broken at its base, but the crest for the muscular insertion is distinguishable. The medial condyle appears stout and rectangular, although its cranial and caudal processs have been eroded. The flexor groove is fully open and its margin, consisting of the caudal process of the lateral condyle and the condylid of the medial condyle, are V-shaped in outline.
</paragraph>
<paragraph id="8BED36B6EB67640DFF45FE85B2E6FC9D" blockId="13.[163,780,382,956]" lastBlockId="14.[145,762,197,863]" lastPageId="14" lastPageNumber="139" pageId="13" pageNumber="138">
<emphasis id="B926EAA4EB67640EFF45FE85B10FFE51" box="[163,275,382,403]" italics="true" pageId="13" pageNumber="138">Remarks:</emphasis>
The preserved femoral shaft is strongly bowed craniocaudally, as in dryosaurids, most elasmarians,
<taxonomicName id="4C524D35EB67640EFEA0FE47B250FE13" box="[326,588,444,465]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B926EAA4EB67640EFEA0FE47B250FE13" box="[326,588,444,465]" italics="true" pageId="13" pageNumber="138">Camptosaurus dispar</emphasis>
</taxonomicName>
and differently from
<taxonomicName id="4C524D35EB67640EFF02FE20B1DCFE32" box="[228,448,475,496]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B926EAA4EB67640EFF02FE20B189FE32" box="[228,405,475,496]" italics="true" pageId="13" pageNumber="138">Tenontosaurus</emphasis>
sp.
</taxonomicName>
and other styracosternans (
<bibRefCitation id="EFC34B47EB67640EFF4AFE01B141FDCC" author="Norman DB" box="[172,349,505,527]" pageId="13" pageNumber="138" pagination="1 - 105" refId="ref18270" refString="Norman DB. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman, 1980</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB67640EFE89FE02B1B6FDCD" author="Norman DB" box="[367,426,505,527]" pageId="13" pageNumber="138" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
, 2004;
<bibRefCitation id="EFC34B47EB67640EFDEEFE02B0FCFDEF" author="Carpenter K &amp; Wilson Y" pageId="13" pageNumber="138" pagination="227 - 264" refId="ref16499" refString="Carpenter K, Wilson Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter &amp; Wilson, 2008</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB67640EFF12FDE3B253FDEF" author="Carpenter K &amp; Galton PM" box="[244,591,536,558]" pageId="13" pageNumber="138" pagination="167 - 207" refId="ref16420" refString="Carpenter K, Galton PM. 2018. A photo documentation of bipedal ornithischian dinosaurs from the Upper Jurassic Morrison Formation, USA. Geology of the Intermountain West 5: 167 - 207." type="journal article" year="2018">Carpenter &amp; Galton, 2018</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB67640EFD82FDE3B0C1FD8E" author="Herne MC &amp; Nair JP &amp; Evans AR &amp; Tait AM" pageId="13" pageNumber="138" pagination="1 - 42" refId="ref17444" refString="Herne MC, Nair JP, Evans AR, Tait AM. 2019. New smallbodied ornithopods (Dinosauria, Neornithischia) from the Early Cretaceous Wonthaggi Formation (Strzelecki Group) of the Australian - Antarctic rift system, with revision of Qantassaurus intrepidus Rich and Vickers-Rich, 1999. Journal of Paleontology 93: 1 - 42." type="journal article" year="2019">
Herne
<emphasis id="B926EAA4EB67640EFD59FDE3B31FFDEF" box="[703,771,536,557]" italics="true" pageId="13" pageNumber="138">et al.</emphasis>
, 2019
</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB67640EFF0BFDCDB1EEFD8E" author="Rozadilla S &amp; Agnolin FL &amp; Novas FE" box="[237,498,566,588]" pageId="13" pageNumber="138" pagination="2043 - 2089" refId="ref18763" refString="Rozadilla S, Agnolin FL, Novas FE. 2019. Osteology of the Patagonian ornithopod Talenkauen santacrucensis (Dinosauria, Ornithischia). Journal of Systematic Palaeontology 17: 2043 - 2089." type="journal article" year="2019">
Rozadilla
<emphasis id="B926EAA4EB67640EFE8CFDCCB183FD89" box="[362,415,566,588]" italics="true" pageId="13" pageNumber="138">et al</emphasis>
., 2019
</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB67640EFDE5FDCCB221FD8E" author="Rozadilla S &amp; Cruzado-Caballero P &amp; Calvo JO" box="[515,573,567,588]" pageId="13" pageNumber="138" pagination="104311" refId="ref18797" refString="Rozadilla S, Cruzado-Caballero P, Calvo JO. 2020. Osteology of Ornithopod Macrogryphosaurus gondwanicus (Dinosauria, Ornithischia) from the Upper Cretaceous of Patagonia, Argentina. Cretaceous Research 108: 104311." type="journal article" year="2020">2020</bibRefCitation>
). On the cranial surface there is no crest developing from the medial condyle, extending proximally towards the lesser trochanter, as seen in other ankylopollexians, such as
<taxonomicName id="4C524D35EB67640EFEF5FD4AB22DFD04" box="[275,561,689,710]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B926EAA4EB67640EFEF5FD4AB22DFD04" box="[275,561,689,710]" italics="true" pageId="13" pageNumber="138">Camptosaurus dispar</emphasis>
</taxonomicName>
,
<taxonomicName id="4C524D35EB67640EFDA0FD4AB31FFD05" box="[582,771,689,711]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B926EAA4EB67640EFDA0FD4AB2CDFD04" box="[582,721,689,710]" italics="true" pageId="13" pageNumber="138">Iguanodon</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="4C524D35EB67640EFF45FD2BB251FD27" authorityName="Mantell" authorityYear="1825" box="[163,589,720,741]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="atherfieldensis">
<emphasis id="B926EAA4EB67640EFF45FD2BB251FD27" box="[163,589,720,741]" italics="true" pageId="13" pageNumber="138">Mantellisaurus atherfieldensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D35EB67640EFD7BFD2BB15DFCC1" baseAuthorityName="Carpenter &amp; Wilson" baseAuthorityYear="2008" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="B926EAA4EB67640EFD7BFD2BB15DFCC1" italics="true" pageId="13" pageNumber="138">Uteodon aphanoecetes</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC34B47EB67640EFEB5FD15B1E2FCC6" author="Gilmore CW" box="[339,510,750,772]" pageId="13" pageNumber="138" pagination="197 - 332" refId="ref17242" refString="Gilmore CW. 1909. Osteology of the Jurassic reptile Camptosaurus: with a revision of the species of the genus, and description of two new species. Proceedings of the United States National Museum 36: 197 - 332." type="journal article" year="1909">Gilmore, 1909</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB67640EFDE8FD14B2A7FCC6" author="Norman DB" box="[526,699,751,772]" pageId="13" pageNumber="138" pagination="1 - 105" refId="ref18270" refString="Norman DB. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman, 1980</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB67640EFD2CFD15B318FCC6" author="Norman DB" box="[714,772,750,772]" pageId="13" pageNumber="138" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
, 2004;
<bibRefCitation id="EFC34B47EB67640EFF01FCF6B20DFCE0" author="Carpenter K &amp; Wilson Y" box="[231,529,781,803]" pageId="13" pageNumber="138" pagination="227 - 264" refId="ref16499" refString="Carpenter K, Wilson Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter &amp; Wilson, 2008</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB67640EFDFBFCF6B0C0FC83" author="Carpenter K &amp; Galton PM" pageId="13" pageNumber="138" pagination="167 - 207" refId="ref16420" refString="Carpenter K, Galton PM. 2018. A photo documentation of bipedal ornithischian dinosaurs from the Upper Jurassic Morrison Formation, USA. Geology of the Intermountain West 5: 167 - 207." type="journal article" year="2018">Carpenter &amp; Galton, 2018</bibRefCitation>
). The lateral condyle of the distal epiphysis of the femur in
<taxonomicName id="4C524D35EB67640EFEDDFCB0B20AFC9D" authorityName="Mateus &amp; Antunes" authorityYear="2001" box="[315,534,842,864]" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="loureiroi">
<emphasis id="B926EAA4EB67640EFEDDFCB0B20AFC9D" box="[315,534,842,864]" italics="true" pageId="13" pageNumber="138">Draconyx loureiroi</emphasis>
</taxonomicName>
is concave in outline and it extends more craniocaudally than the ones of
<taxonomicName id="4C524D35EB67640EFF23FC73B26CFC5F" authority=", Cumnoria" authorityName="Cumnoria" box="[197,624,904,925]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B926EAA4EB67640EFF23FC73B1C0FC5F" box="[197,476,904,925]" italics="true" pageId="13" pageNumber="138">Camptosaurus dispar</emphasis>
,
<emphasis id="B926EAA4EB67640EFE17FC73B26CFC5F" box="[497,624,904,925]" italics="true" pageId="13" pageNumber="138">Cumnoria</emphasis>
</taxonomicName>
<taxonomicName id="4C524D35EB67640EFD9AFC73B316FC5F" authorityName="Hulke" authorityYear="1880" box="[636,778,904,925]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="prestwichii">
<emphasis id="B926EAA4EB67640EFD9AFC73B316FC5F" box="[636,778,904,925]" italics="true" pageId="13" pageNumber="138">prestwichii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D35EB67640EFF31FC5DB1C1FC79" baseAuthorityName="Carpenter &amp; Wilson" baseAuthorityYear="2008" box="[215,477,934,955]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="B926EAA4EB67640EFF31FC5DB1C1FC79" box="[215,477,934,955]" italics="true" pageId="13" pageNumber="138">Uteodon aphanoecetes</emphasis>
</taxonomicName>
(
<figureCitation id="13692A33EB67640EFE09FC5DB22DFC7E" box="[495,561,934,956]" captionStart="Figure 7" captionStartId="13.[163,242,1760,1782]" captionTargetBox="[168,1441,1049,1715]" captionTargetId="figure-480@13.[163,1449,1046,1721]" captionTargetPageId="13" captionText="Figure 7. Comparison of distal sections of femoral shaft from selected iguanodontians. A, Draconyx loureiroi, B, Uteodon aphanoecetes, C, Cumnoria prestwichii, D, Camptosaurus dispar, E, Barilium dawsoni, F, Mantellisaurus atherfieldensis and G, Iguanodon galvensis. Abbreviations: cod: condylid, eg: extensor groove, fg: flexor groove, lc: lateral condyle, mc: medial condyle. Distal sections drawn from: Carpenter &amp; Wilson, 2008; Galton &amp; Powell, 1980; Norman, 2011; Norman, 1986; Verdù et al., 2018." figureDoi="http://doi.org/10.5281/zenodo.6530811" httpUri="https://zenodo.org/record/6530811/files/figure.png" pageId="13" pageNumber="138">Fig. 7</figureCitation>
;
<bibRefCitation id="EFC34B47EB67640EFDA6FC5DB36AFF18" author="Galton PM &amp; Powell HP" box="[576,886,197,956]" pageId="13" pageNumber="138" pagination="411 - 443" refId="ref17215" refString="Galton PM, Powell HP. 1980. The ornithischian dinosaur Camptosaurus prestwichii from the Upper Jurassic of England. Palaeontology 23: 411 - 443." type="journal article" year="1980">Galton &amp; Powell, 1980</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB67640EFC6EFF3EB4C9FF18" author="Carpenter K &amp; Wilson Y" box="[904,1237,197,219]" pageId="13" pageNumber="138" pagination="227 - 264" refId="ref16499" refString="Carpenter K, Wilson Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter &amp; Wilson, 2008</bibRefCitation>
). The inflection point of the curvature is located more cranially in
<taxonomicName id="4C524D35EB67640EFCDDFEF8B438FED5" authorityName="Mateus &amp; Antunes" authorityYear="2001" box="[827,1060,258,280]" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="loureiroi">
<emphasis id="B926EAA4EB67640EFCDDFEF8B438FED5" box="[827,1060,258,280]" italics="true" pageId="13" pageNumber="138">Draconyx loureiroi</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C524D35EB67640EFB73FEF8B5BFFED5" box="[1173,1443,258,280]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B926EAA4EB67640EFB73FEF8B5BFFED5" box="[1173,1443,258,280]" italics="true" pageId="13" pageNumber="138">Camptosaurus dispar</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D35EB67640EFC95FEDAB494FEF4" baseAuthorityName="Carpenter &amp; Wilson" baseAuthorityYear="2008" box="[883,1160,289,310]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="B926EAA4EB67640EFC95FEDAB494FEF4" box="[883,1160,289,310]" italics="true" pageId="13" pageNumber="138">Uteodon aphanoecetes</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC34B47EB67640EFB7BFEDAB369FE97" author="Carpenter K &amp; Wilson Y" pageId="13" pageNumber="138" pagination="227 - 264" refId="ref16499" refString="Carpenter K, Wilson Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter &amp; Wilson, 2008</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB67640EFC63FEBBB4D8FE97" author="Carpenter K &amp; Galton PM" box="[901,1220,320,342]" pageId="13" pageNumber="138" pagination="167 - 207" refId="ref16420" refString="Carpenter K, Galton PM. 2018. A photo documentation of bipedal ornithischian dinosaurs from the Upper Jurassic Morrison Formation, USA. Geology of the Intermountain West 5: 167 - 207." type="journal article" year="2018">Carpenter &amp; Galton, 2018</bibRefCitation>
), while
<emphasis id="B926EAA4EB67640EFACEFEBBB3DBFEB1" italics="true" pageId="13" pageNumber="138">
Cumnoria
<taxonomicName id="4C524D35EB67640EFCDDFEA5B3DBFEB1" authorityName="Hulke" authorityYear="1880" box="[827,967,350,371]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
</emphasis>
exhibits a smoother outline without abrupt changes in curvature. The lateral condyle of Early Cretaceous species, such as
<taxonomicName id="4C524D35EB67640EFB2BFE67B5BFFE73" authorityName="Lydekker" authorityYear="1888" box="[1229,1443,412,433]" class="Reptilia" family="Iguanodontidae" genus="Barilium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="dawsoni">
<emphasis id="B926EAA4EB67640EFB2BFE67B5BFFE73" box="[1229,1443,412,433]" italics="true" pageId="13" pageNumber="138">Barilium dawsoni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D35EB67640EFC94FE41B4EFFE0D" authorityName="Mantell" authorityYear="1825" box="[882,1267,442,463]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="atherfieldensis">
<emphasis id="B926EAA4EB67640EFC94FE41B4EFFE0D" box="[882,1267,442,463]" italics="true" pageId="13" pageNumber="138">Mantellisaurus atherfieldensis</emphasis>
</taxonomicName>
, are larger in proportions with respect to the total size of the epiphysis and extend more caudally than the ones of the above-mentioned taxa, including
<taxonomicName id="4C524D35EB67640EFAFCFDECB5BFFDE9" authorityName="Mateus &amp; Antunes" authorityYear="2001" box="[1306,1443,534,556]" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="loureiroi">
<emphasis id="B926EAA4EB67640EFAFCFDECB5BFFDE9" box="[1306,1443,534,556]" italics="true" pageId="13" pageNumber="138">D. loureiroi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC34B47EB67640EFCA3FDCEB41EFD89" author="Norman DB" box="[837,1026,565,587]" pageId="13" pageNumber="138" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">Norman, 1986</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB67640EFBFEFDCEB44AFD88" author="Norman DB" box="[1048,1110,565,586]" pageId="13" pageNumber="138" pagination="165 - 194" refId="ref18373" refString="Norman DB. 2011. On the osteology of the lower Wealden (Valanginian) ornithopod Barilium dawsoni (Iguanodontia: Styracosterna). Special Papers in Palaeontology 86: 165 - 194." type="journal article" year="2011">2011</bibRefCitation>
). The medial condyle of
<taxonomicName id="4C524D35EB67640EFCDDFDAFB40BFDAB" authorityName="Mateus &amp; Antunes" authorityYear="2001" box="[827,1047,596,617]" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="loureiroi">
<emphasis id="B926EAA4EB67640EFCDDFDAFB40BFDAB" box="[827,1047,596,617]" italics="true" pageId="13" pageNumber="138">Draconyx loureiroi</emphasis>
</taxonomicName>
is subrectangular and its medial margin is straight, as is the one of
<taxonomicName id="4C524D35EB67640EFB0CFD89B3EAFD64" authorityName="Mantell" authorityYear="1825" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="atherfieldensis">
<emphasis id="B926EAA4EB67640EFB0CFD89B3EAFD64" italics="true" pageId="13" pageNumber="138">Mantellisaurus atherfieldensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D35EB67640EFBDDFD6AB53DFD64" authorityName="Lydekker" authorityYear="1888" box="[1083,1313,657,678]" class="Reptilia" family="Iguanodontidae" genus="Barilium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="dawsoni">
<emphasis id="B926EAA4EB67640EFBDDFD6AB53DFD64" box="[1083,1313,657,678]" italics="true" pageId="13" pageNumber="138">Barilium dawsoni</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC34B47EB67640EFAD0FD6AB36BFD04" author="Norman DB" pageId="13" pageNumber="138" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">Norman, 1986</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB67640EFC6CFD4BB3DAFD07" author="Norman DB" box="[906,966,688,709]" pageId="13" pageNumber="138" pagination="165 - 194" refId="ref18373" refString="Norman DB. 2011. On the osteology of the lower Wealden (Valanginian) ornithopod Barilium dawsoni (Iguanodontia: Styracosterna). Special Papers in Palaeontology 86: 165 - 194." type="journal article" year="2011">2011</bibRefCitation>
). However, in Jurassic taxa such as
<taxonomicName id="4C524D35EB67640EFCDDFD35B450FD21" baseAuthorityName="Carpenter &amp; Wilson" baseAuthorityYear="2008" box="[827,1100,718,739]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="B926EAA4EB67640EFCDDFD35B450FD21" box="[827,1100,718,739]" italics="true" pageId="13" pageNumber="138">Uteodon aphanoecetes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D35EB67640EFB68FD34B587FD21" box="[1166,1435,718,740]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B926EAA4EB67640EFB68FD34B587FD21" box="[1166,1435,718,740]" italics="true" pageId="13" pageNumber="138">Camptosaurus dispar</emphasis>
</taxonomicName>
, the medial margin of the medial condyle is rounded (
<bibRefCitation id="EFC34B47EB67640EFCA2FCF7B491FCE3" author="Carpenter K &amp; Wilson Y" box="[836,1165,780,802]" pageId="13" pageNumber="138" pagination="227 - 264" refId="ref16499" refString="Carpenter K, Wilson Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter &amp; Wilson, 2008</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB67640EFB79FCF7B36AFCFD" author="Carpenter K &amp; Galton PM" pageId="13" pageNumber="138" pagination="167 - 207" refId="ref16420" refString="Carpenter K, Galton PM. 2018. A photo documentation of bipedal ornithischian dinosaurs from the Upper Jurassic Morrison Formation, USA. Geology of the Intermountain West 5: 167 - 207." type="journal article" year="2018">Carpenter &amp; Galton, 2018</bibRefCitation>
). The flexor and extensor grooves are fully open as in many basal and cursorial iguanodontians, contrasting with more derived forms (Norman, 2004). Extensor grooves of the Jurassic taxa
<taxonomicName id="4C524D35EB67640EFB11FC7CB397FC78" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B926EAA4EB67640EFB11FC7CB397FC78" italics="true" pageId="13" pageNumber="138">Camptosaurus dispar</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D35EB67640EFC28FC5EB4FEFC78" baseAuthorityName="Carpenter &amp; Wilson" baseAuthorityYear="2008" box="[974,1250,933,954]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="B926EAA4EB67640EFC28FC5EB4FEFC78" box="[974,1250,933,954]" italics="true" pageId="13" pageNumber="138">Uteodon aphanoecetes</emphasis>
</taxonomicName>
are fully open, but shallower compared to the ones of
<taxonomicName id="4C524D35EB64640DFD60FF3EB166FF3B" authority=", Cumnoria" authorityName="Cumnoria" class="Reptilia" family="Iguanodontidae" genus="Barilium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="dawsoni">
<emphasis id="B926EAA4EB64640DFD60FF3EB0E8FF3B" italics="true" pageId="14" pageNumber="139">Barilium dawsoni</emphasis>
,
<emphasis id="B926EAA4EB64640DFEE4FF1FB166FF3B" box="[258,378,228,249]" italics="true" pageId="14" pageNumber="139">Cumnoria</emphasis>
</taxonomicName>
<taxonomicName id="4C524D35EB64640DFE64FF1FB219FF3B" authorityName="Hulke" authorityYear="1880" box="[386,517,228,249]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="prestwichii">
<emphasis id="B926EAA4EB64640DFE64FF1FB219FF3B" box="[386,517,228,249]" italics="true" pageId="14" pageNumber="139">prestwichii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C524D35EB64640DFDF2FF1EB2EEFF3B" authorityName="Mateus &amp; Antunes" authorityYear="2001" box="[532,754,228,250]" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="loureiroi">
<emphasis id="B926EAA4EB64640DFDF2FF1EB2EEFF3B" box="[532,754,228,250]" italics="true" pageId="14" pageNumber="139">Draconyx loureiroi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C524D35EB64640DFF77FEF9B122FEDA" box="[145,318,258,280]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B926EAA4EB64640DFF77FEF9B108FED5" box="[145,276,258,279]" italics="true" pageId="14" pageNumber="139">Iguanodon</emphasis>
sp.
</taxonomicName>
and
<taxonomicName id="4C524D35EB64640DFE99FEF9B2E5FED5" authorityName="Mantell" authorityYear="1825" box="[383,761,258,279]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="atherfieldensis">
<emphasis id="B926EAA4EB64640DFE99FEF9B2E5FED5" box="[383,761,258,279]" italics="true" pageId="14" pageNumber="139">Mantellisaurus atherfieldensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC34B47EB64640DFF7DFED9B14DFEF4" author="Norman DB" box="[155,337,289,311]" pageId="14" pageNumber="139" pagination="1 - 105" refId="ref18270" refString="Norman DB. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman, 1980</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB64640DFE83FEDAB1BDFEF5" author="Norman DB" box="[357,417,289,311]" pageId="14" pageNumber="139" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
, 2004, 2011,
<bibRefCitation id="EFC34B47EB64640DFDB0FEDAB13AFE97" author="Carpenter K &amp; Wilson Y" pageId="14" pageNumber="139" pagination="227 - 264" refId="ref16499" refString="Carpenter K, Wilson Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter &amp; Wilson, 2008</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB64640DFED5FEBBB27DFE97" author="Carpenter K &amp; Galton PM" box="[307,609,320,342]" pageId="14" pageNumber="139" pagination="167 - 207" refId="ref16420" refString="Carpenter K, Galton PM. 2018. A photo documentation of bipedal ornithischian dinosaurs from the Upper Jurassic Morrison Formation, USA. Geology of the Intermountain West 5: 167 - 207." type="journal article" year="2018">Carpenter &amp; Galton, 2018</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB64640DFD8BFEBBB0D5FEB6" author="Verdu FJ &amp; Royo-Torres R &amp; Cobos A &amp; Alcala L" pageId="14" pageNumber="139" pagination="437 - 474" refId="ref19266" refString="Verdu FJ, Royo-Torres R, Cobos A, Alcala L. 2018. New systematic and phylogenetic data about the early Barremian Iguanodon galvensis (Ornithopoda: Iguanodontoidea) from Spain. Historical Biology 30: 437 - 474." type="journal article" year="2018">
Verdú
<emphasis id="B926EAA4EB64640DFD5CFEBBB2EEFE97" box="[698,754,320,341]" italics="true" pageId="14" pageNumber="139">et al.</emphasis>
, 2018
</bibRefCitation>
). Moreover, the flexor groove walls of
<emphasis id="B926EAA4EB64640DFD64FEA4B10BFE50" italics="true" pageId="14" pageNumber="139">
Cumnoria
<taxonomicName id="4C524D35EB64640DFF77FE86B10BFE50" authorityName="Hulke" authorityYear="1880" box="[145,279,381,402]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
</emphasis>
and
<taxonomicName id="4C524D35EB64640DFEB1FE86B27EFE50" baseAuthorityName="Carpenter &amp; Wilson" baseAuthorityYear="2008" box="[343,610,381,402]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="B926EAA4EB64640DFEB1FE86B27EFE50" box="[343,610,381,402]" italics="true" pageId="14" pageNumber="139">Uteodon aphanoecetes</emphasis>
</taxonomicName>
are slightly divergent from one another (
<figureCitation id="13692A33EB64640DFE1FFE67B223FE73" box="[505,575,412,434]" captionStart="Figure 7" captionStartId="13.[163,242,1760,1782]" captionTargetBox="[168,1441,1049,1715]" captionTargetId="figure-480@13.[163,1449,1046,1721]" captionTargetPageId="13" captionText="Figure 7. Comparison of distal sections of femoral shaft from selected iguanodontians. A, Draconyx loureiroi, B, Uteodon aphanoecetes, C, Cumnoria prestwichii, D, Camptosaurus dispar, E, Barilium dawsoni, F, Mantellisaurus atherfieldensis and G, Iguanodon galvensis. Abbreviations: cod: condylid, eg: extensor groove, fg: flexor groove, lc: lateral condyle, mc: medial condyle. Distal sections drawn from: Carpenter &amp; Wilson, 2008; Galton &amp; Powell, 1980; Norman, 2011; Norman, 1986; Verdù et al., 2018." figureDoi="http://doi.org/10.5281/zenodo.6530811" httpUri="https://zenodo.org/record/6530811/files/figure.png" pageId="14" pageNumber="139">Fig. 7</figureCitation>
). The extensor groove of the Early Cretaceous iguanodontians
<taxonomicName id="4C524D35EB64640DFF77FE22B173FE2C" box="[145,367,473,494]" class="Reptilia" family="Iguanodontidae" genus="Barilium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="dawsoni">
<emphasis id="B926EAA4EB64640DFF77FE22B173FE2C" box="[145,367,473,494]" italics="true" pageId="14" pageNumber="139">Barilium dawsoni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C524D35EB64640DFE66FE22B2E4FE2C" authorityName="Mantell" authorityYear="1825" box="[384,760,473,494]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="atherfieldensis">
<emphasis id="B926EAA4EB64640DFE66FE22B2E4FE2C" box="[384,760,473,494]" italics="true" pageId="14" pageNumber="139">Mantellisaurus atherfieldensis</emphasis>
</taxonomicName>
and other styracosternans are partially enclosed by overhangs of medial and lateral condyles.
<taxonomicName id="4C524D35EB64640DFD6FFDECB0EAFD88" authorityName="Mateus &amp; Antunes" authorityYear="2001" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="loureiroi">
<emphasis id="B926EAA4EB64640DFD6FFDECB0EAFD88" italics="true" pageId="14" pageNumber="139">Draconyx loureiroi</emphasis>
</taxonomicName>
does not exhibit an overhang of the medial condyle on the flexor groove, unlike the condition present in
<taxonomicName id="4C524D35EB64640DFED6FD88B2EAFD4A" authority=", Cumnoria" authorityName="Cumnoria" box="[304,758,626,648]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B926EAA4EB64640DFED6FD88B246FD4A" box="[304,602,626,648]" italics="true" pageId="14" pageNumber="139">Camptosaurus dispar</emphasis>
,
<emphasis id="B926EAA4EB64640DFD94FD88B2EAFD4A" box="[626,758,627,648]" italics="true" pageId="14" pageNumber="139">Cumnoria</emphasis>
</taxonomicName>
<taxonomicName id="4C524D35EB64640DFF77FD69B135FD65" authorityName="Hulke" authorityYear="1880" box="[145,297,657,679]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="prestwichii">
<emphasis id="B926EAA4EB64640DFF77FD69B135FD65" box="[145,297,657,679]" italics="true" pageId="14" pageNumber="139">prestwichii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C524D35EB64640DFEA4FD69B2ECFD65" authorityName="Mantell" authorityYear="1825" box="[322,752,657,679]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="atherfieldensis">
<emphasis id="B926EAA4EB64640DFEA4FD69B2ECFD65" box="[322,752,657,679]" italics="true" pageId="14" pageNumber="139">Mantellisaurus atherfieldensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C524D35EB64640DFF77FD4BB241FD04" authority="Taquet, 1976" authorityName="Taquet" authorityYear="1976" box="[145,605,688,710]" class="Reptilia" family="Iguanodontidae" genus="Ouranosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="nigeriensis">
<emphasis id="B926EAA4EB64640DFF77FD4BB1DDFD07" box="[145,449,688,709]" italics="true" pageId="14" pageNumber="139">Ouranosaurus nigeriensis</emphasis>
<bibRefCitation id="EFC34B47EB64640DFE20FD4BB241FD04" author="Taquet P" box="[454,605,688,710]" pageId="14" pageNumber="139" refId="ref19143" refString="Taquet P. 1976. Geologie et paleontologie du Gisement de Gadoufaoua (Aptien du Niger). Paris: Editions du Centre National de la Recherque Scientifique." type="book" year="1976">Taquet, 1976</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C524D35EB64640DFD71FD4BB124FD21" baseAuthorityName="Carpenter &amp; Wilson" baseAuthorityYear="2008" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="B926EAA4EB64640DFD71FD4BB124FD21" italics="true" pageId="14" pageNumber="139">Uteodon aphanoecetes</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC34B47EB64640DFEABFD34B21FFD26" author="Norman DB" box="[333,515,719,740]" pageId="14" pageNumber="139" pagination="1 - 105" refId="ref18270" refString="Norman DB. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman, 1980</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB64640DFDF0FD35B24EFD26" author="Norman DB" box="[534,594,718,740]" pageId="14" pageNumber="139" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
, 2004, 2011;
<bibRefCitation id="EFC34B47EB64640DFF77FD16B1C7FCC0" author="Carpenter K &amp; Wilson Y" box="[145,475,749,771]" pageId="14" pageNumber="139" pagination="227 - 264" refId="ref16499" refString="Carpenter K, Wilson Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter &amp; Wilson, 2008</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB64640DFE0BFD15B2EEFCC0" author="Bertozzo F &amp; Dalla Vecchia FM &amp; Fabbri M" box="[493,754,749,771]" pageId="14" pageNumber="139" pagination="3403" refId="ref16369" refString="Bertozzo F, Dalla Vecchia FM, Fabbri M. 2017. The Venice specimen of Ouranosaurus nigeriensis (Dinosauria, Ornithopoda). PeerJ 5: e 3403." type="journal article" year="2017">
Bertozzo
<emphasis id="B926EAA4EB64640DFD83FD15B2B8FCC0" box="[613,676,749,771]" italics="true" pageId="14" pageNumber="139">et al.</emphasis>
, 2017
</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB64640DFF77FCF7B1D6FCE3" author="Carpenter K &amp; Galton PM" box="[145,458,780,802]" pageId="14" pageNumber="139" pagination="167 - 207" refId="ref16420" refString="Carpenter K, Galton PM. 2018. A photo documentation of bipedal ornithischian dinosaurs from the Upper Jurassic Morrison Formation, USA. Geology of the Intermountain West 5: 167 - 207." type="journal article" year="2018">Carpenter &amp; Galton, 2018</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB64640DFE3FFCF7B2B1FCE3" author="Verdu FJ &amp; Royo-Torres R &amp; Cobos A &amp; Alcala L" box="[473,685,780,802]" pageId="14" pageNumber="139" pagination="437 - 474" refId="ref19266" refString="Verdu FJ, Royo-Torres R, Cobos A, Alcala L. 2018. New systematic and phylogenetic data about the early Barremian Iguanodon galvensis (Ornithopoda: Iguanodontoidea) from Spain. Historical Biology 30: 437 - 474." type="journal article" year="2018">
Verdú
<emphasis id="B926EAA4EB64640DFDCFFCF7B278FCE3" box="[553,612,780,801]" italics="true" pageId="14" pageNumber="139">et al.</emphasis>
, 2018
</bibRefCitation>
). This condition more closely resembles the plesiomorphic condition within Ornithopoda (
<bibRefCitation id="EFC34B47EB64640DFDE6FCB1B2F5FC9C" author="Norman DB &amp; Sues HD &amp; Witmer LM &amp; Coria RA" box="[512,745,841,863]" pageId="14" pageNumber="139" pagination="393 - 412" refId="ref18449" refString="Norman DB, Sues HD, Witmer LM, Coria RA. 2004. Basal ornithopoda. In: Weishampel DB, Dodson P, Osmolska H, eds. The Dinosauria. Berkeley: University of California, 393 - 412." type="book chapter" year="2004">
Norman
<emphasis id="B926EAA4EB64640DFD8FFCB1B2BFFC9C" box="[617,675,841,863]" italics="true" pageId="14" pageNumber="139">et al.</emphasis>
, 2004
</bibRefCitation>
).
</paragraph>
<caption id="DF2D663EEB67640EFF45F91BB106F8AE" ID-DOI="http://doi.org/10.5281/zenodo.6530811" ID-Zenodo-Dep="6530811" httpUri="https://zenodo.org/record/6530811/files/figure.png" pageId="13" pageNumber="138" startId="13.[163,242,1760,1782]" targetBox="[168,1441,1049,1715]" targetPageId="13">
<paragraph id="8BED36B6EB67640EFF45F91BB106F8AE" blockId="13.[163,1443,1760,1900]" pageId="13" pageNumber="138">
<emphasis id="B926EAA4EB67640EFF45F91BB110F934" bold="true" box="[163,268,1760,1782]" pageId="13" pageNumber="138">Figure 7.</emphasis>
Comparison of distal sections of femoral shaft from selected iguanodontians. A,
<taxonomicName id="4C524D35EB67640EFBB9F91AB53FF935" authorityName="Mateus &amp; Antunes" authorityYear="2001" box="[1119,1315,1761,1783]" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="loureiroi">
<emphasis id="B926EAA4EB67640EFBB9F91AB53FF935" box="[1119,1315,1761,1783]" italics="true" pageId="13" pageNumber="138">Draconyx loureiroi</emphasis>
</taxonomicName>
, B,
<taxonomicName id="4C524D35EB67640EFAADF91AB131F8D6" baseAuthorityName="Carpenter &amp; Wilson" baseAuthorityYear="2008" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="B926EAA4EB67640EFAADF91AB131F8D6" italics="true" pageId="13" pageNumber="138">Uteodon aphanoecetes</emphasis>
</taxonomicName>
, C,
<emphasis id="B926EAA4EB67640EFEB0F905B221F8D6" box="[342,573,1790,1812]" italics="true" pageId="13" pageNumber="138">
Cumnoria
<taxonomicName id="4C524D35EB67640EFE2EF905B221F8D6" authorityName="Hulke" authorityYear="1880" box="[456,573,1790,1812]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
</emphasis>
, D,
<taxonomicName id="4C524D35EB67640EFD81F905B351F8D6" box="[615,845,1790,1812]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B926EAA4EB67640EFD81F905B351F8D6" box="[615,845,1790,1812]" italics="true" pageId="13" pageNumber="138">Camptosaurus dispar</emphasis>
</taxonomicName>
, E,
<taxonomicName id="4C524D35EB67640EFC91F905B42BF8D6" box="[887,1079,1790,1812]" class="Reptilia" family="Iguanodontidae" genus="Barilium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="dawsoni">
<emphasis id="B926EAA4EB67640EFC91F905B42BF8D6" box="[887,1079,1790,1812]" italics="true" pageId="13" pageNumber="138">Barilium dawsoni</emphasis>
</taxonomicName>
, F,
<taxonomicName id="4C524D35EB67640EFBBBF905B5BEF8D6" authorityName="Mantell" authorityYear="1825" box="[1117,1442,1790,1812]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="atherfieldensis">
<emphasis id="B926EAA4EB67640EFBBBF905B5BEF8D6" box="[1117,1442,1790,1812]" italics="true" pageId="13" pageNumber="138">Mantellisaurus atherfieldensis</emphasis>
</taxonomicName>
and G,
<taxonomicName id="4C524D35EB67640EFF0DF8E0B1DCF8F3" box="[235,448,1819,1841]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="138" phylum="Chordata" rank="species" species="galvensis">
<emphasis id="B926EAA4EB67640EFF0DF8E0B1DCF8F3" box="[235,448,1819,1841]" italics="true" pageId="13" pageNumber="138">Iguanodon galvensis</emphasis>
</taxonomicName>
. Abbreviations: cod: condylid, eg: extensor groove, fg: flexor groove, lc: lateral condyle, mc: medial condyle. Distal sections drawn from:
<bibRefCitation id="EFC34B47EB67640EFDF8F8C2B337F88D" author="Carpenter K &amp; Wilson Y" box="[542,811,1849,1871]" pageId="13" pageNumber="138" pagination="227 - 264" refId="ref16499" refString="Carpenter K, Wilson Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter &amp; Wilson, 2008</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB67640EFCD0F8C2B400F88D" author="Galton PM &amp; Powell HP" box="[822,1052,1849,1871]" pageId="13" pageNumber="138" pagination="411 - 443" refId="ref17215" refString="Galton PM, Powell HP. 1980. The ornithischian dinosaur Camptosaurus prestwichii from the Upper Jurassic of England. Palaeontology 23: 411 - 443." type="journal article" year="1980">Galton &amp; Powell, 1980</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB67640EFBCEF8C2B4A7F88D" author="Norman DB" box="[1064,1211,1849,1871]" pageId="13" pageNumber="138" pagination="165 - 194" refId="ref18373" refString="Norman DB. 2011. On the osteology of the lower Wealden (Valanginian) ornithopod Barilium dawsoni (Iguanodontia: Styracosterna). Special Papers in Palaeontology 86: 165 - 194." type="journal article" year="2011">Norman, 2011</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB67640EFB20F8C2B545F88D" author="Norman DB" box="[1222,1369,1849,1871]" pageId="13" pageNumber="138" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">Norman, 1986</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB67640EFA82F8C2B108F8AE" author="Verdu FJ &amp; Royo-Torres R &amp; Cobos A &amp; Alcala L" pageId="13" pageNumber="138" pagination="437 - 474" refId="ref19266" refString="Verdu FJ, Royo-Torres R, Cobos A, Alcala L. 2018. New systematic and phylogenetic data about the early Barremian Iguanodon galvensis (Ornithopoda: Iguanodontoidea) from Spain. Historical Biology 30: 437 - 474." type="journal article" year="2018">
Verdù
<emphasis id="B926EAA4EB67640EFF45F8ACB0CCF8AE" box="[163,208,1878,1900]" italics="true" pageId="13" pageNumber="138">et al</emphasis>
., 2018
</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="8BED36B6EB64640DFF77FC73B3FFFEDA" blockId="14.[145,762,904,1907]" lastBlockId="14.[809,1425,197,280]" pageId="14" pageNumber="139">
<emphasis id="B926EAA4EB64640DFF77FC73B176FC5F" box="[145,362,904,925]" italics="true" pageId="14" pageNumber="139">Tibia (ML 3577</emphasis>
,
<emphasis id="B926EAA4EB64640DFE9BFC73B2E5FC5F" box="[381,761,904,925]" italics="true" pageId="14" pageNumber="139">
12) (
<figureCitation id="13692A33EB64640DFE5DFC73B25CFC5F" box="[443,576,904,925]" captionStart="Figure 3" captionStartId="9.[163,242,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@9.[227,1379,195,1738]" captionTargetPageId="9" captionText="Figure 3. General overview of Draconyx loureiroi holotype ML 357 post-cranial skeleton. Caudal vertebrae (K-M) in left lateral view. Right femur (AE) in frontal (A), medial (B), caudal (C), lateral (D) and distal (E) views. Right tibia in medial (F) and lateral (G) views. Proximal end of the fibula in medial (I) and lateral (J) views. Right articulated pes in dorsal (N) and medial (Q) view. Pedal digit III (O) and pedal digit II (P) in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6530803" httpUri="https://zenodo.org/record/6530803/files/figure.png" pageId="14" pageNumber="139">Figs 3F, G</figureCitation>
,
<figureCitation id="13692A33EB64640DFDB5FC73B28CFC5F" box="[595,656,904,925]" captionStart="Figure 6" captionStartId="12.[145,224,1096,1118]" captionTargetBox="[145,1425,195,1056]" captionTargetId="figure-432@12.[145,1425,195,1056]" captionTargetPageId="12" captionText="Figure 6. Three dimensional models of limb bones elements of Draconyx loureiroi holotype ML 357 (ML 357 6-8), femur (AE) and tibia and proximal end of fibula (FJ). Femur (ML 357 6) in cranial view (A), medial view (B), caudal view (C), lateral view (D) and distal view (E). Tibia (ML 357 7) and proximal end of fibula (ML 357 8) in cranial view (F), medial view (G), caudal view (H), lateral (I) and proximal (J) views." figureDoi="http://doi.org/10.5281/zenodo.6530809" httpUri="https://zenodo.org/record/6530809/files/figure.png" pageId="14" pageNumber="139">6FJ</figureCitation>
,
<figureCitation id="13692A33EB64640DFD45FC73B2FBFC5F" box="[675,743,904,925]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="14" pageNumber="139">8AD</figureCitation>
):
</emphasis>
The proximal (
<figureCitation id="13692A33EB64640DFEA9FC5DB1DBFC7E" box="[335,455,934,956]" captionStart="Figure 6" captionStartId="12.[145,224,1096,1118]" captionTargetBox="[145,1425,195,1056]" captionTargetId="figure-432@12.[145,1425,195,1056]" captionTargetPageId="12" captionText="Figure 6. Three dimensional models of limb bones elements of Draconyx loureiroi holotype ML 357 (ML 357 6-8), femur (AE) and tibia and proximal end of fibula (FJ). Femur (ML 357 6) in cranial view (A), medial view (B), caudal view (C), lateral view (D) and distal view (E). Tibia (ML 357 7) and proximal end of fibula (ML 357 8) in cranial view (F), medial view (G), caudal view (H), lateral (I) and proximal (J) views." figureDoi="http://doi.org/10.5281/zenodo.6530809" httpUri="https://zenodo.org/record/6530809/files/figure.png" pageId="14" pageNumber="139">Fig. 6FJ</figureCitation>
) and distal (
<figureCitation id="13692A33EB64640DFD94FC5DB2F3FC7E" box="[626,751,934,956]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="14" pageNumber="139">Fig. 8AE</figureCitation>
) epiphyses of the tibia are preserved, but the tibial shaft is missing. Both extremities are heavily eroded and covered by matrix and adhesives, but relevant characters are still distinguishable. The proximal epiphysis preserves a conspicuous cnemial crest, a robust fibular condyle and the internal condyle. The cnemial crest tapers dorsally forming a smooth edge; in dorsal view its margins are laterally concave and medially convex (
<figureCitation id="13692A33EB64640DFEBFFB41B1DFFB12" box="[345,451,1210,1232]" captionStart="Figure 6" captionStartId="12.[145,224,1096,1118]" captionTargetBox="[145,1425,195,1056]" captionTargetId="figure-432@12.[145,1425,195,1056]" captionTargetPageId="12" captionText="Figure 6. Three dimensional models of limb bones elements of Draconyx loureiroi holotype ML 357 (ML 357 6-8), femur (AE) and tibia and proximal end of fibula (FJ). Femur (ML 357 6) in cranial view (A), medial view (B), caudal view (C), lateral view (D) and distal view (E). Tibia (ML 357 7) and proximal end of fibula (ML 357 8) in cranial view (F), medial view (G), caudal view (H), lateral (I) and proximal (J) views." figureDoi="http://doi.org/10.5281/zenodo.6530809" httpUri="https://zenodo.org/record/6530809/files/figure.png" pageId="14" pageNumber="139">Fig. 6G, I</figureCitation>
). Laterally, it is divided by the fibular condyle to form a deep and extensive scar (sulcus tibialis). The fibular condyle is a stout process that deflects strongly caudally, its articular facet is rounded and it blends smoothly with the anterior and posterior surfaces of the condyle. The internal condyle is a blunt eminence directed caudally and projecting gently laterally. Like the cnemial crest, its lateral margin is concave, while the medial margin is strongly convex (
<figureCitation id="13692A33EB64640DFEB5FA35B1B8FA26" box="[339,420,1486,1508]" captionStart="Figure 6" captionStartId="12.[145,224,1096,1118]" captionTargetBox="[145,1425,195,1056]" captionTargetId="figure-432@12.[145,1425,195,1056]" captionTargetPageId="12" captionText="Figure 6. Three dimensional models of limb bones elements of Draconyx loureiroi holotype ML 357 (ML 357 6-8), femur (AE) and tibia and proximal end of fibula (FJ). Femur (ML 357 6) in cranial view (A), medial view (B), caudal view (C), lateral view (D) and distal view (E). Tibia (ML 357 7) and proximal end of fibula (ML 357 8) in cranial view (F), medial view (G), caudal view (H), lateral (I) and proximal (J) views." figureDoi="http://doi.org/10.5281/zenodo.6530809" httpUri="https://zenodo.org/record/6530809/files/figure.png" pageId="14" pageNumber="139">Fig. 6E</figureCitation>
). This convexity forms a deep sulcus that separates the internal condyle from the fibular condyle. Immediately ventral to the proximal epiphysis, the proximal part of the diaphyseal shaft is preserved and it is teardrop-shaped in cross-section. The distal part of the tibia is articulated with the astragalus, calcaneum and the metatarsals (
<figureCitation id="13692A33EB64640DFD2CF97DB0CFF978" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="14" pageNumber="139">Fig. 8AE</figureCitation>
). The distal epiphysis flares mediolaterally, the medial malleolus is rounded and slightly more expanded mediolaterally than the lateral one, which is narrow and elongate, expanding proximodistally. They articulate between each other forming an angle of approximately 45°. The intermuscular line is located on the caudal surface of the distal epiphysis, separating the surface of the two malleoli and reaching the astragalus, forming with its apex a continuous concave surface.
</paragraph>
<paragraph id="8BED36B6EB64640DFCCFFEBAB3FEFB97" blockId="14.[809,1426,321,1109]" pageId="14" pageNumber="139">
<emphasis id="B926EAA4EB64640DFCCFFEBAB3BDFE94" box="[809,929,321,342]" italics="true" pageId="14" pageNumber="139">Remarks:</emphasis>
The medial margin of the proximal epiphysis of the tibia is convex, as is typical of many iguandontians (Norman, 2004), except for
<taxonomicName id="4C524D35EB64640DFCCFFE66B546FE70" authority="Novas et al., 2004" authorityName="Novas" authorityYear="2004" box="[809,1370,413,435]" class="Reptilia" family="Hadrosauridae" genus="Talenkauen" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="santacrucensis">
<emphasis id="B926EAA4EB64640DFCCFFE66B46FFE70" box="[809,1139,413,434]" italics="true" pageId="14" pageNumber="139">Talenkauen santacrucensis</emphasis>
<bibRefCitation id="EFC34B47EB64640DFB9BFE66B546FE70" author="Novas F &amp; Cambiaso A &amp; Ambrosio A" box="[1149,1370,413,435]" pageId="14" pageNumber="139" pagination="75 - 82" refId="ref18492" refString="Novas F, Cambiaso A, Ambrosio A. 2004. A new basal iguanodontian (Dinosauria, Ornithischia) from the Upper Cretaceous of Patagonia. Ameghiniana 41: 75 - 82." type="journal article" year="2004">
Novas
<emphasis id="B926EAA4EB64640DFB37FE65B513FE70" box="[1233,1295,413,435]" italics="true" pageId="14" pageNumber="139">et al.</emphasis>
, 2004
</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C524D35EB64640DFCCFFE40B58CFE13" authority="Escaso et al., 2014" authorityName="Escaso" authorityYear="2014" box="[809,1424,443,465]" class="Reptilia" family="Dryosauridae" genus="Eousdryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="nanohallucis">
<emphasis id="B926EAA4EB64640DFCCFFE40B487FE12" box="[809,1179,443,464]" italics="true" pageId="14" pageNumber="139">Eousdryosaurus nanohallucis</emphasis>
<bibRefCitation id="EFC34B47EB64640DFB40FE47B58CFE13" author="Escaso F &amp; Ortega F &amp; Dantas P &amp; Malafaia E &amp; Silva BC &amp; Gasulla JM &amp; Mocho P &amp; Narvaez I &amp; Sanz JL" box="[1190,1424,443,465]" pageId="14" pageNumber="139" pagination="1102 - 1112" refId="ref16813" refString="Escaso F, Ortega F, Dantas P, Malafaia E, Silva BC, Gasulla JM, Mocho P, Narvaez I, Sanz JL. 2014. A new dryosaurid ornithopod (Dinosauria, Ornithischia) from the Late Jurassic of Portugal. Journal of Vertebrate Paleontology 34: 1102 - 1112." type="journal article" year="2014">
Escaso
<emphasis id="B926EAA4EB64640DFAE3FE47B558FE12" box="[1285,1348,443,465]" italics="true" pageId="14" pageNumber="139">et al.</emphasis>
, 2014
</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFC34B47EB64640DFCD7FE20B419FE2D" author="Escaso F &amp; Ortega F &amp; Dantas P &amp; Malafaia E &amp; Silva BC &amp; Gasulla JM &amp; Mocho P &amp; Narvaez I &amp; Sanz JL" box="[817,1029,474,496]" pageId="14" pageNumber="139" pagination="1102 - 1112" refId="ref16813" refString="Escaso F, Ortega F, Dantas P, Malafaia E, Silva BC, Gasulla JM, Mocho P, Narvaez I, Sanz JL. 2014. A new dryosaurid ornithopod (Dinosauria, Ornithischia) from the Late Jurassic of Portugal. Journal of Vertebrate Paleontology 34: 1102 - 1112." type="journal article" year="2014">
Escaso
<emphasis id="B926EAA4EB64640DFC61FE20B3DDFE2D" box="[903,961,474,496]" italics="true" pageId="14" pageNumber="139">et al.</emphasis>
, 2014
</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB64640DFBF5FE21B519FE2D" author="Rozadilla S &amp; Agnolin FL &amp; Novas FE" box="[1043,1285,474,496]" pageId="14" pageNumber="139" pagination="2043 - 2089" refId="ref18763" refString="Rozadilla S, Agnolin FL, Novas FE. 2019. Osteology of the Patagonian ornithopod Talenkauen santacrucensis (Dinosauria, Ornithischia). Journal of Systematic Palaeontology 17: 2043 - 2089." type="journal article" year="2019">
Rozadilla
<emphasis id="B926EAA4EB64640DFB6EFE20B4DDFE2D" box="[1160,1217,474,496]" italics="true" pageId="14" pageNumber="139">et al.</emphasis>
, 2019
</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB64640DFAF2FE21B3B4FDCC" author="Dieudonne PE &amp; Cruzado-Caballero P &amp; Godefroit P &amp; Tortosa T" pageId="14" pageNumber="139" pagination="2335 - 2355" refId="ref16723" refString="Dieudonne PE, Cruzado-Caballero P, Godefroit P, Tortosa T. 2021. A new phylogeny of cerapodan dinosaurs. Historical Biology 33: 2335 - 2355." type="journal article" year="2021">
Dieudonné
<emphasis id="B926EAA4EB64640DFCCFFE01B37EFDCC" box="[809,866,505,527]" italics="true" pageId="14" pageNumber="139">et al.</emphasis>
, 2021
</bibRefCitation>
). As noted by
<bibRefCitation id="EFC34B47EB64640DFBA9FE02B541FDCD" author="Dieudonne PE &amp; Cruzado-Caballero P &amp; Godefroit P &amp; Tortosa T" box="[1103,1373,505,527]" pageId="14" pageNumber="139" pagination="2335 - 2355" refId="ref16723" refString="Dieudonne PE, Cruzado-Caballero P, Godefroit P, Tortosa T. 2021. A new phylogeny of cerapodan dinosaurs. Historical Biology 33: 2335 - 2355." type="journal article" year="2021">
Dieudonné
<emphasis id="B926EAA4EB64640DFB32FE01B51BFDCC" box="[1236,1287,505,527]" italics="true" pageId="14" pageNumber="139">et al</emphasis>
. (2021)
</bibRefCitation>
, the cnemial crest apex is directed strongly craniolaterally, a characteristic common to Laurasian dryomorphans and
<taxonomicName id="4C524D35EB64640DFCBFFDAEB4DCFDA8" authority="Pompeckj, 1920" authorityName="Pompeckj" authorityYear="1920" box="[857,1216,597,619]" class="Reptilia" family="Dryosauridae" genus="Dysalotosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="genus">
<emphasis id="B926EAA4EB64640DFCBFFDAEB414FDA8" box="[857,1032,597,618]" italics="true" pageId="14" pageNumber="139">Dysalotosaurus</emphasis>
Pompeckj, 1920
</taxonomicName>
, differing from the condition exhibited by Elasmaria and the Portuguese ornithopod
<taxonomicName id="4C524D35EB64640DFC5FFD69B3ACFD04" authority="(Escaso et al., 2014)" baseAuthorityName="Escaso" baseAuthorityYear="2014" class="Reptilia" family="Dryosauridae" genus="Eousdryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="nanohallucis">
<emphasis id="B926EAA4EB64640DFC5FFD69B535FD65" box="[953,1321,658,679]" italics="true" pageId="14" pageNumber="139">Eousdryosaurus nanohallucis</emphasis>
(
<bibRefCitation id="EFC34B47EB64640DFAD8FD68B3BAFD04" author="Escaso F &amp; Ortega F &amp; Dantas P &amp; Malafaia E &amp; Silva BC &amp; Gasulla JM &amp; Mocho P &amp; Narvaez I &amp; Sanz JL" pageId="14" pageNumber="139" pagination="1102 - 1112" refId="ref16813" refString="Escaso F, Ortega F, Dantas P, Malafaia E, Silva BC, Gasulla JM, Mocho P, Narvaez I, Sanz JL. 2014. A new dryosaurid ornithopod (Dinosauria, Ornithischia) from the Late Jurassic of Portugal. Journal of Vertebrate Paleontology 34: 1102 - 1112." type="journal article" year="2014">
Escaso
<emphasis id="B926EAA4EB64640DFCCFFD4AB37EFD04" box="[809,866,689,710]" italics="true" pageId="14" pageNumber="139">et al.</emphasis>
, 2014
</bibRefCitation>
)
</taxonomicName>
. As in many styracosternans, the cnemial crest is well developed (Norman, 2004). The fibular condyle differs from dryosaurids (with the exception of
<taxonomicName id="4C524D35EB64640DFCA8FCF6B4AFFCE0" authorityName="Galton" authorityYear="1975" box="[846,1203,781,802]" class="Reptilia" family="Dryosauridae" genus="Valdosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="canaliculatus">
<emphasis id="B926EAA4EB64640DFCA8FCF6B4AFFCE0" box="[846,1203,781,802]" italics="true" pageId="14" pageNumber="139">Valdosaurus canaliculatus</emphasis>
</taxonomicName>
),
<taxonomicName id="4C524D35EB64640DFB35FCF6B369FC82" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B926EAA4EB64640DFB35FCF6B369FC82" italics="true" pageId="14" pageNumber="139">Camptosaurus dispar</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D35EB64640DFC55FCD0B4ABFC82" baseAuthorityName="Carpenter &amp; Wilson" baseAuthorityYear="2008" box="[947,1207,811,832]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="B926EAA4EB64640DFC55FCD0B4ABFC82" box="[947,1207,811,832]" italics="true" pageId="14" pageNumber="139">Uteodon aphanoecetes</emphasis>
</taxonomicName>
in being partially posterolaterally deflected, similar to the condition seen in
<emphasis id="B926EAA4EB64640DFCAEFC92B45BFCBC" box="[840,1095,873,894]" italics="true" pageId="14" pageNumber="139">
Cumnoria
<taxonomicName id="4C524D35EB64640DFC20FC92B45BFCBC" authorityName="Hulke" authorityYear="1880" box="[966,1095,873,894]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
</emphasis>
,
<taxonomicName id="4C524D35EB64640DFBB3FC92B58DFCBC" box="[1109,1425,873,894]" class="Reptilia" family="Hadrosauridae" genus="Talenkauen" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="santacrucensis">
<emphasis id="B926EAA4EB64640DFBB3FC92B58DFCBC" box="[1109,1425,873,894]" italics="true" pageId="14" pageNumber="139">Talenkauen santacrucensis</emphasis>
</taxonomicName>
and more derived styracosternanans (
<bibRefCitation id="EFC34B47EB64640DFAFDFC7CB3D8FC79" author="Galton PM &amp; Powell HP" pageId="14" pageNumber="139" pagination="411 - 443" refId="ref17215" refString="Galton PM, Powell HP. 1980. The ornithischian dinosaur Camptosaurus prestwichii from the Upper Jurassic of England. Palaeontology 23: 411 - 443." type="journal article" year="1980">Galton &amp; Powell, 1980</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB64640DFC30FC5CB499FC79" author="Norman DB" box="[982,1157,934,956]" pageId="14" pageNumber="139" pagination="1 - 105" refId="ref18270" refString="Norman DB. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman, 1980</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB64640DFB70FC5DB4CCFC7E" author="Norman DB" box="[1174,1232,934,956]" pageId="14" pageNumber="139" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB64640DFB04FC5DB500FC79" author="Norman DB" box="[1250,1308,934,955]" pageId="14" pageNumber="139" pagination="165 - 194" refId="ref18373" refString="Norman DB. 2011. On the osteology of the lower Wealden (Valanginian) ornithopod Barilium dawsoni (Iguanodontia: Styracosterna). Special Papers in Palaeontology 86: 165 - 194." type="journal article" year="2011">2011</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB64640DFACBFC5CB3B4FC18" author="Norman DB &amp; Sues HD &amp; Witmer LM &amp; Coria RA" pageId="14" pageNumber="139" pagination="393 - 412" refId="ref18449" refString="Norman DB, Sues HD, Witmer LM, Coria RA. 2004. Basal ornithopoda. In: Weishampel DB, Dodson P, Osmolska H, eds. The Dinosauria. Berkeley: University of California, 393 - 412." type="book chapter" year="2004">
Norman
<emphasis id="B926EAA4EB64640DFCCFFC3EB37EFC18" box="[809,866,965,986]" italics="true" pageId="14" pageNumber="139">et al.</emphasis>
, 2004
</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB64640DFC50FC3EB4F9FC18" author="Carpenter K &amp; Wilson Y" box="[950,1253,965,987]" pageId="14" pageNumber="139" pagination="227 - 264" refId="ref16499" refString="Carpenter K, Wilson Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter &amp; Wilson, 2008</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB64640DFB15FC3EB37FFC3B" author="Barrett PM &amp; Butler RJ &amp; Twitchett RJ &amp; Hutt S" pageId="14" pageNumber="139" pagination="131 - 163" refId="ref16131" refString="Barrett PM, Butler RJ, Twitchett RJ, Hutt S. 2011. New material of Valdosaurus canaliculatus (Ornithischia: Ornithopoda) from the Lower Cretaceous of southern England. Special Papers in Palaeontology 86: 131 - 163." type="journal article" year="2011">
Barrett
<emphasis id="B926EAA4EB64640DFAB7FC3EB597FC18" box="[1361,1419,965,986]" italics="true" pageId="14" pageNumber="139">et al.</emphasis>
, 2011
</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB64640DFC95FC18B464FC3B" author="Rozadilla S &amp; Agnolin FL &amp; Novas FE" box="[883,1144,995,1017]" pageId="14" pageNumber="139" pagination="2043 - 2089" refId="ref18763" refString="Rozadilla S, Agnolin FL, Novas FE. 2019. Osteology of the Patagonian ornithopod Talenkauen santacrucensis (Dinosauria, Ornithischia). Journal of Systematic Palaeontology 17: 2043 - 2089." type="journal article" year="2019">
Rozadilla
<emphasis id="B926EAA4EB64640DFC16FC1FB431FC3A" box="[1008,1069,995,1017]" italics="true" pageId="14" pageNumber="139">et al.</emphasis>
, 2019
</bibRefCitation>
). The distal epiphysis, found in articulation with the rest of the pes, does not differ significantly from that of other iguanodontians (Norman, 2004).
</paragraph>
<paragraph id="8BED36B6EB64640DFCCFFB85B461FA6A" blockId="14.[809,1426,1150,1540]" pageId="14" pageNumber="139">
<emphasis id="B926EAA4EB64640DFCCFFB85B3E3FB51" box="[809,1023,1150,1171]" italics="true" pageId="14" pageNumber="139">Fibula (ML 3578</emphasis>
,
<emphasis id="B926EAA4EB64640DFBE8FB85B540FB51" box="[1038,1372,1150,1172]" italics="true" pageId="14" pageNumber="139">
12) (
<figureCitation id="13692A33EB64640DFBA3FB84B4A8FB56" box="[1093,1204,1150,1172]" captionStart="Figure 3" captionStartId="9.[163,242,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@9.[227,1379,195,1738]" captionTargetPageId="9" captionText="Figure 3. General overview of Draconyx loureiroi holotype ML 357 post-cranial skeleton. Caudal vertebrae (K-M) in left lateral view. Right femur (AE) in frontal (A), medial (B), caudal (C), lateral (D) and distal (E) views. Right tibia in medial (F) and lateral (G) views. Proximal end of the fibula in medial (I) and lateral (J) views. Right articulated pes in dorsal (N) and medial (Q) view. Pedal digit III (O) and pedal digit II (P) in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6530803" httpUri="https://zenodo.org/record/6530803/files/figure.png" pageId="14" pageNumber="139">Figs 3I, J</figureCitation>
,
<figureCitation id="13692A33EB64640DFB25FB85B4E0FB51" box="[1219,1276,1150,1171]" captionStart="Figure 6" captionStartId="12.[145,224,1096,1118]" captionTargetBox="[145,1425,195,1056]" captionTargetId="figure-432@12.[145,1425,195,1056]" captionTargetPageId="12" captionText="Figure 6. Three dimensional models of limb bones elements of Draconyx loureiroi holotype ML 357 (ML 357 6-8), femur (AE) and tibia and proximal end of fibula (FJ). Femur (ML 357 6) in cranial view (A), medial view (B), caudal view (C), lateral view (D) and distal view (E). Tibia (ML 357 7) and proximal end of fibula (ML 357 8) in cranial view (F), medial view (G), caudal view (H), lateral (I) and proximal (J) views." figureDoi="http://doi.org/10.5281/zenodo.6530809" httpUri="https://zenodo.org/record/6530809/files/figure.png" pageId="14" pageNumber="139">6FJ</figureCitation>
,
<figureCitation id="13692A33EB64640DFAEDFB85B550FB51" box="[1291,1356,1150,1171]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="14" pageNumber="139">8AD</figureCitation>
):
</emphasis>
The distal and proximal epiphyses of the fibula are preserved, whereas the diaphysis is completely absent. The proximal end is a flattened subtriangular element. Dorsally, it appears to be slightly medially convex. The cranial margin deflects abruptly dorsally into the cranial process, which is slightly eroded (
<figureCitation id="13692A33EB64640DFAD1FACDB59EFA8E" box="[1335,1410,1334,1356]" captionStart="Figure 6" captionStartId="12.[145,224,1096,1118]" captionTargetBox="[145,1425,195,1056]" captionTargetId="figure-432@12.[145,1425,195,1056]" captionTargetPageId="12" captionText="Figure 6. Three dimensional models of limb bones elements of Draconyx loureiroi holotype ML 357 (ML 357 6-8), femur (AE) and tibia and proximal end of fibula (FJ). Femur (ML 357 6) in cranial view (A), medial view (B), caudal view (C), lateral view (D) and distal view (E). Tibia (ML 357 7) and proximal end of fibula (ML 357 8) in cranial view (F), medial view (G), caudal view (H), lateral (I) and proximal (J) views." figureDoi="http://doi.org/10.5281/zenodo.6530809" httpUri="https://zenodo.org/record/6530809/files/figure.png" pageId="14" pageNumber="139">Fig. 6I</figureCitation>
). In contrast, the caudal margin deflects less abruptly dorsally. A deep fossa is present close to the caudal margin on the lateral surface.
</paragraph>
<paragraph id="8BED36B6EB64640DFCA7FA4AB4E9F9C1" blockId="14.[809,1426,1150,1540]" pageId="14" pageNumber="139">
The distal epiphysis is in articulation with the rest of the pes, located on the lateral surface of the tibia and contacts the calcaneum (
<figureCitation id="13692A33EB64640DFB95FA15B4F9F9C1" box="[1139,1253,1518,1540]" captionStart="Figure 7" captionStartId="13.[163,242,1760,1782]" captionTargetBox="[168,1441,1049,1715]" captionTargetId="figure-480@13.[163,1449,1046,1721]" captionTargetPageId="13" captionText="Figure 7. Comparison of distal sections of femoral shaft from selected iguanodontians. A, Draconyx loureiroi, B, Uteodon aphanoecetes, C, Cumnoria prestwichii, D, Camptosaurus dispar, E, Barilium dawsoni, F, Mantellisaurus atherfieldensis and G, Iguanodon galvensis. Abbreviations: cod: condylid, eg: extensor groove, fg: flexor groove, lc: lateral condyle, mc: medial condyle. Distal sections drawn from: Carpenter &amp; Wilson, 2008; Galton &amp; Powell, 1980; Norman, 2011; Norman, 1986; Verdù et al., 2018." figureDoi="http://doi.org/10.5281/zenodo.6530811" httpUri="https://zenodo.org/record/6530811/files/figure.png" pageId="14" pageNumber="139">Fig. 7AD</figureCitation>
).
</paragraph>
<paragraph id="8BED36B6EB646413FCCFF9D6B2D2FF3B" blockId="14.[809,1426,1581,1909]" lastBlockId="16.[145,760,197,250]" lastPageId="16" lastPageNumber="141" pageId="14" pageNumber="139">
<emphasis id="B926EAA4EB64640DFCCFF9D6B385F980" box="[809,921,1581,1602]" italics="true" pageId="14" pageNumber="139">Remarks:</emphasis>
The proximal epiphysis has cranial and caudal margins that diverge smoothly, as in
<taxonomicName id="4C524D35EB64640DFAEBF9B7B37EF9BD" authorityName="Marsh" authorityYear="1878" class="Reptilia" family="Dryosauridae" genus="Dryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="altus">
<emphasis id="B926EAA4EB64640DFAEBF9B7B37EF9BD" italics="true" pageId="14" pageNumber="139">Dryosaurus altus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C524D35EB64640DFC96F991B4D7F9BD" authorityName="Virchow" authorityYear="1919" box="[880,1227,1642,1663]" class="Reptilia" family="Dryosauridae" genus="Dysalotosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="lettowvorbecki">
<emphasis id="B926EAA4EB64640DFC96F991B4D7F9BD" box="[880,1227,1642,1663]" italics="true" pageId="14" pageNumber="139">Dysalotosaurus lettowvorbecki</emphasis>
</taxonomicName>
,
<taxonomicName id="4C524D35EB64640DFB31F991B3C8F95C" class="Reptilia" family="Dryosauridae" genus="Eousdryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="nanohallucis">
<emphasis id="B926EAA4EB64640DFB31F991B3C8F95C" italics="true" pageId="14" pageNumber="139">Eousdryosaurus nanohallucis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C524D35EB64640DFC0CF972B54FF95C" box="[1002,1363,1673,1694]" class="Reptilia" family="Hadrosauridae" genus="Talenkauen" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="santacrucensis">
<emphasis id="B926EAA4EB64640DFC0CF972B54FF95C" box="[1002,1363,1673,1694]" italics="true" pageId="14" pageNumber="139">Talenkauen santacrucensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D35EB64640DFCCFF95CB44AF97E" authorityName="Galton" authorityYear="1975" box="[809,1110,1703,1724]" class="Reptilia" family="Dryosauridae" genus="Valdosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="canaliculatus">
<emphasis id="B926EAA4EB64640DFCCFF95CB44AF97E" box="[809,1110,1703,1724]" italics="true" pageId="14" pageNumber="139">Valdosaurus canaliculatus</emphasis>
</taxonomicName>
, but differs from
<taxonomicName id="4C524D35EB64640DFAF0F95CB3C1F919" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="bernissartensis">
<emphasis id="B926EAA4EB64640DFAF0F95CB3C1F919" italics="true" pageId="14" pageNumber="139">Iguanodon bernissartensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C524D35EB64640DFC0AF93DB540F919" authorityName="Mantell" authorityYear="1825" box="[1004,1372,1734,1755]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="atherfieldensis">
<emphasis id="B926EAA4EB64640DFC0AF93DB540F919" box="[1004,1372,1734,1755]" italics="true" pageId="14" pageNumber="139">Mantellisaurus atherfieldensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D35EB64640DFCCFF91EB46DF938" authorityName="Taquet" authorityYear="1976" box="[809,1137,1765,1786]" class="Reptilia" family="Iguanodontidae" genus="Ouranosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="nigeriensis">
<emphasis id="B926EAA4EB64640DFCCFF91EB46DF938" box="[809,1137,1765,1786]" italics="true" pageId="14" pageNumber="139">Ouranosaurus nigeriensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC34B47EB64640DFB61F91EB527F938" author="Norman DB" box="[1159,1339,1765,1786]" pageId="14" pageNumber="139" pagination="1 - 105" refId="ref18270" refString="Norman DB. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman, 1980</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB64640DFAA8F91EB595F939" author="Norman DB" box="[1358,1417,1765,1787]" pageId="14" pageNumber="139" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB64640DFCCFF8F8B3A1F8DA" author="Galton PM" box="[809,957,1795,1817]" pageId="14" pageNumber="139" pagination="271 - 312" refId="ref17047" refString="Galton PM. 1981. Dryosaurus, a hypsilophodontid dinosaur from the Upper Jurassic of North America and Africa postcranial skeleton. Palaontologische Zeitschrift 55: 271 - 312." type="journal article" year="1981">Galton, 1981</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB64640DFC2DF8FFB4B4F8DA" author="Barrett PM &amp; Butler RJ &amp; Twitchett RJ &amp; Hutt S" box="[971,1192,1795,1817]" pageId="14" pageNumber="139" pagination="131 - 163" refId="ref16131" refString="Barrett PM, Butler RJ, Twitchett RJ, Hutt S. 2011. New material of Valdosaurus canaliculatus (Ornithischia: Ornithopoda) from the Lower Cretaceous of southern England. Special Papers in Palaeontology 86: 131 - 163." type="journal article" year="2011">
Barrett
<emphasis id="B926EAA4EB64640DFBCFF8FFB47EF8DA" box="[1065,1122,1795,1817]" italics="true" pageId="14" pageNumber="139">et al.</emphasis>
, 2011
</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB64640DFB53F8FFB597F8DA" author="Escaso F &amp; Ortega F &amp; Dantas P &amp; Malafaia E &amp; Silva BC &amp; Gasulla JM &amp; Mocho P &amp; Narvaez I &amp; Sanz JL" box="[1205,1419,1795,1817]" pageId="14" pageNumber="139" pagination="1102 - 1112" refId="ref16813" refString="Escaso F, Ortega F, Dantas P, Malafaia E, Silva BC, Gasulla JM, Mocho P, Narvaez I, Sanz JL. 2014. A new dryosaurid ornithopod (Dinosauria, Ornithischia) from the Late Jurassic of Portugal. Journal of Vertebrate Paleontology 34: 1102 - 1112." type="journal article" year="2014">
Escaso
<emphasis id="B926EAA4EB64640DFAEAF8FFB559F8DA" box="[1292,1349,1795,1817]" italics="true" pageId="14" pageNumber="139">et al.</emphasis>
, 2014
</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB64640DFCCFF8D9B3D4F8FA" author="Barrett PM" box="[809,968,1826,1848]" pageId="14" pageNumber="139" pagination="29 - 48" refId="ref16093" refString="Barrett PM. 2016. A new specimen of Valdosaurus canaliculatus (Ornithopoda: Dryosauridae) from the Lower Cretaceous of the Isle of Wight, England. Memoirs of Museum Victoria 74: 29 - 48." type="journal article" year="2016">Barrett, 2016</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB64640DFC31F8D9B4D4F8F5" author="Bertozzo F &amp; Dalla Vecchia FM &amp; Fabbri M" box="[983,1224,1826,1848]" pageId="14" pageNumber="139" pagination="3403" refId="ref16369" refString="Bertozzo F, Dalla Vecchia FM, Fabbri M. 2017. The Venice specimen of Ouranosaurus nigeriensis (Dinosauria, Ornithopoda). PeerJ 5: e 3403." type="journal article" year="2017">
Bertozzo
<emphasis id="B926EAA4EB64640DFBA0F8D8B49DF8F5" box="[1094,1153,1826,1848]" italics="true" pageId="14" pageNumber="139">et al.</emphasis>
, 2017
</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB64640DFB3EF8D9B37CF894" author="Rozadilla S &amp; Agnolin FL &amp; Novas FE" pageId="14" pageNumber="139" pagination="2043 - 2089" refId="ref18763" refString="Rozadilla S, Agnolin FL, Novas FE. 2019. Osteology of the Patagonian ornithopod Talenkauen santacrucensis (Dinosauria, Ornithischia). Journal of Systematic Palaeontology 17: 2043 - 2089." type="journal article" year="2019">
Rozadilla
<emphasis id="B926EAA4EB64640DFAA9F8D8B596F8F5" box="[1359,1418,1826,1848]" italics="true" pageId="14" pageNumber="139">et al.</emphasis>
, 2019
</bibRefCitation>
). As in
<emphasis id="B926EAA4EB64640DFC51F8BAB4E4F894" box="[951,1272,1857,1878]" italics="true" pageId="14" pageNumber="139">Sektensaurus sanjuanboscoi</emphasis>
<bibRefCitation id="EFC34B47EB64640DFB19F8BAB378F8B6" author="Ibiricu LM &amp; Casal GA &amp; Martinez RD &amp; Luna M &amp; Canale JI &amp; Alvarez BN &amp; Gonzalez Riga B" pageId="14" pageNumber="139" pagination="276 - 291" refId="ref17601" refString="Ibiricu LM, Casal GA, Martinez RD, Luna M, Canale JI, Alvarez BN, Gonzalez Riga B. 2019. A new ornithopod dinosaur (Dinosauria: Ornithischia) from the Late Cretaceous of central Patagonia. Cretaceous Research 98: 276 - 291." type="journal article" year="2019">
Ibiricu
<emphasis id="B926EAA4EB64640DFAB5F8BAB597F894" box="[1363,1419,1857,1878]" italics="true" pageId="14" pageNumber="139">et al.</emphasis>
, 2019
</bibRefCitation>
and
<taxonomicName id="4C524D35EB64640DFC4EF8A4B4E5F8B7" box="[936,1273,1887,1909]" class="Reptilia" family="Hadrosauridae" genus="Talenkauen" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="139" phylum="Chordata" rank="species" species="santacrucensis">
<emphasis id="B926EAA4EB64640DFC4EF8A4B4E5F8B7" box="[936,1273,1887,1909]" italics="true" pageId="14" pageNumber="139">Talenkauen santacrucensis</emphasis>
</taxonomicName>
, the caudal margin of the proximal part of the fibula is almost vertical (
<bibRefCitation id="EFC34B47EB7A6413FF10FF1FB1D9FF3B" author="Ibiricu LM &amp; Casal GA &amp; Martinez RD &amp; Luna M &amp; Canale JI &amp; Alvarez BN &amp; Gonzalez Riga B" box="[246,453,228,250]" pageId="16" pageNumber="141" pagination="276 - 291" refId="ref17601" refString="Ibiricu LM, Casal GA, Martinez RD, Luna M, Canale JI, Alvarez BN, Gonzalez Riga B. 2019. A new ornithopod dinosaur (Dinosauria: Ornithischia) from the Late Cretaceous of central Patagonia. Cretaceous Research 98: 276 - 291." type="journal article" year="2019">
Ibiricu
<emphasis id="B926EAA4EB7A6413FEAFFF1EB19DFF3B" box="[329,385,228,250]" italics="true" pageId="16" pageNumber="141">et al.</emphasis>
, 2019
</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB7A6413FE37FF1FB2A3FF3B" author="Rozadilla S &amp; Agnolin FL &amp; Novas FE" box="[465,703,228,250]" pageId="16" pageNumber="141" pagination="2043 - 2089" refId="ref18763" refString="Rozadilla S, Agnolin FL, Novas FE. 2019. Osteology of the Patagonian ornithopod Talenkauen santacrucensis (Dinosauria, Ornithischia). Journal of Systematic Palaeontology 17: 2043 - 2089." type="journal article" year="2019">
Rozadilla
<emphasis id="B926EAA4EB7A6413FDA5FF1EB267FF3B" box="[579,635,228,250]" italics="true" pageId="16" pageNumber="141">et al.</emphasis>
, 2019
</bibRefCitation>
).
</paragraph>
<caption id="DF2D663EEB65640CFF45F909B415F8BF" ID-DOI="http://doi.org/10.5281/zenodo.6530813" ID-Zenodo-Dep="6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="15" pageNumber="140" startId="15.[163,241,1778,1800]" targetBox="[227,1379,195,1738]" targetPageId="15">
<paragraph id="8BED36B6EB65640CFF45F909B415F8BF" blockId="15.[163,1443,1778,1917]" pageId="15" pageNumber="140">
<emphasis id="B926EAA4EB65640CFF45F909B117F8C5" bold="true" box="[163,267,1778,1800]" pageId="15" pageNumber="140">Figure 8.</emphasis>
Three dimensional models of the pes of
<taxonomicName id="4C524D35EB65640CFD53F908B36BF8CA" authorityName="Mateus &amp; Antunes" authorityYear="2001" box="[693,887,1778,1800]" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="15" pageNumber="140" phylum="Chordata" rank="species" species="loureiroi">
<emphasis id="B926EAA4EB65640CFD53F908B36BF8CA" box="[693,887,1778,1800]" italics="true" pageId="15" pageNumber="140">Draconyx loureiroi</emphasis>
</taxonomicName>
holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I. Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III.
</paragraph>
</caption>
<paragraph id="8BED36B6EB7A6413FF77FED8B274FD48" blockId="16.[145,761,290,650]" pageId="16" pageNumber="141">
<emphasis id="B926EAA4EB7A6413FF77FED8B2DFFEFA" box="[145,707,291,312]" italics="true" pageId="16" pageNumber="141">
Pes (ML 3571219) (
<figureCitation id="13692A33EB7A6413FE73FED8B209FEFA" box="[405,533,291,312]" captionStart="Figure 3" captionStartId="9.[163,242,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@9.[227,1379,195,1738]" captionTargetPageId="9" captionText="Figure 3. General overview of Draconyx loureiroi holotype ML 357 post-cranial skeleton. Caudal vertebrae (K-M) in left lateral view. Right femur (AE) in frontal (A), medial (B), caudal (C), lateral (D) and distal (E) views. Right tibia in medial (F) and lateral (G) views. Proximal end of the fibula in medial (I) and lateral (J) views. Right articulated pes in dorsal (N) and medial (Q) view. Pedal digit III (O) and pedal digit II (P) in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6530803" httpUri="https://zenodo.org/record/6530803/files/figure.png" pageId="16" pageNumber="141">Figs 3N, Q</figureCitation>
,
<figureCitation id="13692A33EB7A6413FDC3FED8B27DFEFA" box="[549,609,291,312]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="16" pageNumber="141">8AF</figureCitation>
,
<figureCitation id="13692A33EB7A6413FD96FED8B2AEFEFA" box="[624,690,291,312]" captionStart="Figure 9" captionStartId="16.[146,224,1848,1870]" captionTargetBox="[305,1265,920,1809]" captionTargetId="figure-367@16.[305,1265,920,1809]" captionTargetPageId="16" captionText="Figure 9. Three dimensional models of the articulated leg (AB) and pes (CD) of Draconyx loureiroi holotype ML 357. Leg in medial view (A) and lateral view (B). Pes in medial (C) and lateral (D) views." figureDoi="http://doi.org/10.5281/zenodo.6530815" httpUri="https://zenodo.org/record/6530815/files/figure.png" pageId="16" pageNumber="141">9C, D</figureCitation>
):
</emphasis>
The pes is articulated with the distal tibia, distal fibula, astragalus and calcaneum and proximal metatarsals. The diagenetic permineralization between the tibia and the rest of the pes prevented a reliable segmentation by CT techniques. This also hampered views of the internal anatomy of the astragalus and calcaneum. The articulated elements share the same subnumber (ML 35712), while the isolated elements have their own numbers. Therefore, the same element may have two subnumbers, depending on whether part of it belongs to the articulated block.
</paragraph>
<paragraph id="8BED36B6EB7A6413FF77FD39B100FD18" blockId="16.[145,284,706,730]" box="[145,284,706,730]" pageId="16" pageNumber="141">
<heading id="D0A581DAEB7A6413FF77FD39B100FD18" box="[145,284,706,730]" centered="true" fontSize="9" level="2" pageId="16" pageNumber="141" reason="2">
<emphasis id="B926EAA4EB7A6413FF77FD39B100FD18" box="[145,284,706,730]" italics="true" pageId="16" pageNumber="141">Calcaneum</emphasis>
</heading>
</paragraph>
<paragraph id="8BED36B6EB7A6413FF77FD11B413FDCC" blockId="16.[145,761,746,860]" lastBlockId="16.[809,1426,197,526]" pageId="16" pageNumber="141">
The calcaneum is preserved in its entirety and does not show signs of significant breakage, erosion or distortion. In its general appearance, the calcaneum i s a c o m p a c t a n d n a r r o w e l e m e n t, e x p a n d i n g craniocaudally more than mediolaterally (
<figureCitation id="13692A33EB7A6413FAF7FF3EB59EFF19" box="[1297,1410,197,219]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="16" pageNumber="141">Fig. 8B, E</figureCitation>
). Distally, its surface is subrectangular in outline and extends dorsally, appearing as a small rhomboidal element in caudal view (
<figureCitation id="13692A33EB7A6413FBBCFEDAB4C8FEF5" box="[1114,1236,289,311]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="16" pageNumber="141">Fig. 8B, E</figureCitation>
). Caudally, the medial margin of the calcaneum forms a smooth concavity that accommodates the lateralmost margin of the astragalus. In lateral view, the calcaneum appears triangular with a caudal apex and is slightly deflected proximally. The proximal and distal margins progressively flare cranially, with the latter forming a scoop-like concavity.
</paragraph>
<paragraph id="8BED36B6EB7A6413FCCFFDBDB3B3FD9C" blockId="16.[809,943,582,606]" box="[809,943,582,606]" pageId="16" pageNumber="141">
<heading id="D0A581DAEB7A6413FCCFFDBDB3B3FD9C" box="[809,943,582,606]" centered="true" fontSize="9" level="2" pageId="16" pageNumber="141" reason="2">
<emphasis id="B926EAA4EB7A6413FCCFFDBDB3B3FD9C" box="[809,943,582,606]" italics="true" pageId="16" pageNumber="141">Astragalus</emphasis>
</heading>
</paragraph>
<paragraph id="8BED36B6EB7A6412FCCFFD95B1CFFEDA" blockId="16.[809,1425,622,859]" lastBlockId="17.[163,779,197,280]" lastPageId="17" lastPageNumber="142" pageId="16" pageNumber="141">
Like the calcaneum, the astragalus is complete and not significantly distorted or eroded. In distal view, the bone has a subtrapezoidal shape (
<figureCitation id="13692A33EB7A6413FB42FD57B50DFD03" box="[1188,1297,684,706]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="16" pageNumber="141">Fig. 8A, E</figureCitation>
). Laterally, it contacts the medial malleolus of the tibia via a deep concavity, while medially the astragalus/calcaneum contact is straight (
<figureCitation id="13692A33EB7A6413FBFFFCF3B48EFCDF" box="[1049,1170,776,798]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="16" pageNumber="141">Fig. 8A, E</figureCitation>
). In caudal view, the astragalus is triangular, with a high ascending process representing the apex, which protrudes extensively dorsally. Immediately ventral to the ascending process, a stout steeply inclined caudomedial process forms a well-distinguishable relief.
</paragraph>
<caption id="DF2D663EEB7A6413FF74F8C3B3D1F8AE" ID-DOI="http://doi.org/10.5281/zenodo.6530815" ID-Zenodo-Dep="6530815" httpUri="https://zenodo.org/record/6530815/files/figure.png" pageId="16" pageNumber="141" startId="16.[146,224,1848,1870]" targetBox="[305,1265,920,1809]" targetPageId="16">
<paragraph id="8BED36B6EB7A6413FF74F8C3B3D1F8AE" blockId="16.[145,1425,1848,1900]" pageId="16" pageNumber="141">
<emphasis id="B926EAA4EB7A6413FF74F8C3B0E5F88F" bold="true" box="[146,249,1848,1870]" pageId="16" pageNumber="141">Figure 9.</emphasis>
Three dimensional models of the articulated leg (AB) and pes (CD) of
<taxonomicName id="4C524D35EB7A6413FC0CF8C2B4B7F88D" authorityName="Mateus &amp; Antunes" authorityYear="2001" box="[1002,1195,1849,1871]" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="16" pageNumber="141" phylum="Chordata" rank="species" species="loureiroi">
<emphasis id="B926EAA4EB7A6413FC0CF8C2B4B7F88D" box="[1002,1195,1849,1871]" italics="true" pageId="16" pageNumber="141">Draconyx loureiroi</emphasis>
</taxonomicName>
holotype ML 357. Leg in medial view (A) and lateral view (B). Pes in medial (C) and lateral (D) views.
</paragraph>
</caption>
<paragraph id="8BED36B6EB7B6412FF45FEBAB23FFC7E" blockId="17.[163,780,321,956]" pageId="17" pageNumber="142">
<emphasis id="B926EAA4EB7B6412FF45FEBAB113FE94" box="[163,271,321,342]" italics="true" pageId="17" pageNumber="142">Remarks:</emphasis>
The astragalus of ML 357 is subtriangular and its caudal surface is moderately high, as in dryosaurids,
<taxonomicName id="4C524D35EB7B6412FEB2FE84B315FE56" authority=", Cumnoria" authorityName="Cumnoria" box="[340,777,382,404]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="142" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B926EAA4EB7B6412FEB2FE84B26FFE51" box="[340,627,382,404]" italics="true" pageId="17" pageNumber="142">Camptosaurus dispar</emphasis>
,
<emphasis id="B926EAA4EB7B6412FD6FFE84B315FE56" box="[649,777,383,404]" italics="true" pageId="17" pageNumber="142">Cumnoria</emphasis>
</taxonomicName>
<taxonomicName id="4C524D35EB7B6412FF45FE66B13CFE70" authorityName="Hulke" authorityYear="1880" box="[163,288,413,434]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="142" phylum="Chordata" rank="species" species="prestwichii">
<emphasis id="B926EAA4EB7B6412FF45FE66B13CFE70" box="[163,288,413,434]" italics="true" pageId="17" pageNumber="142">prestwichii</emphasis>
</taxonomicName>
,
<emphasis id="B926EAA4EB7B6412FECAFE66B14AFE13" italics="true" pageId="17" pageNumber="142">
<taxonomicName id="4C524D35EB7B6412FECAFE66B24BFE70" box="[300,599,413,434]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="142" phylum="Chordata" rank="species" species="bernissartensis">Iguanodon bernissartensis</taxonomicName>
<taxonomicName id="4C524D35EB7B6412FDBBFE66B14AFE13" authorityName="Mantell" authorityYear="1825" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="142" phylum="Chordata" rank="species" species="atherfieldensis">Mantellisaurus atherfieldensis</taxonomicName>
</emphasis>
and
<taxonomicName id="4C524D35EB7B6412FE71FE47B2B9FE13" box="[407,677,444,465]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="142" phylum="Chordata" rank="species" species="aphanaecetes">
<emphasis id="B926EAA4EB7B6412FE71FE47B2B9FE13" box="[407,677,444,465]" italics="true" pageId="17" pageNumber="142">Uteodon aphanaecetes</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC34B47EB7B6412FD5FFE47B14BFE32" author="Galton PM &amp; Powell HP" pageId="17" pageNumber="142" pagination="411 - 443" refId="ref17215" refString="Galton PM, Powell HP. 1980. The ornithischian dinosaur Camptosaurus prestwichii from the Upper Jurassic of England. Palaeontology 23: 411 - 443." type="journal article" year="1980">Galton &amp; Powell, 1980</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB7B6412FE81FE20B20EFE32" author="Norman DB" box="[359,530,475,496]" pageId="17" pageNumber="142" pagination="1 - 105" refId="ref18270" refString="Norman DB. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman, 1980</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB7B6412FDC7FE21B246FE32" author="Norman DB" box="[545,602,474,496]" pageId="17" pageNumber="142" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB7B6412FD8CFE21B318FE32" author="Galton PM" box="[618,772,474,496]" pageId="17" pageNumber="142" pagination="271 - 312" refId="ref17047" refString="Galton PM. 1981. Dryosaurus, a hypsilophodontid dinosaur from the Upper Jurassic of North America and Africa postcranial skeleton. Palaontologische Zeitschrift 55: 271 - 312." type="journal article" year="1981">Galton, 1981</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB7B6412FF45FE02B1CBFDCC" author="Carpenter K &amp; Wilson Y" box="[163,471,505,527]" pageId="17" pageNumber="142" pagination="227 - 264" refId="ref16499" refString="Carpenter K, Wilson Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter &amp; Wilson, 2008</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB7B6412FE00FE01B2DBFDCC" author="Barrett PM &amp; Butler RJ &amp; Twitchett RJ &amp; Hutt S" box="[486,711,505,527]" pageId="17" pageNumber="142" pagination="131 - 163" refId="ref16131" refString="Barrett PM, Butler RJ, Twitchett RJ, Hutt S. 2011. New material of Valdosaurus canaliculatus (Ornithischia: Ornithopoda) from the Lower Cretaceous of southern England. Special Papers in Palaeontology 86: 131 - 163." type="journal article" year="2011">
Barrett
<emphasis id="B926EAA4EB7B6412FDA0FE01B29CFDCC" box="[582,640,505,527]" italics="true" pageId="17" pageNumber="142">et al.</emphasis>
, 2011
</bibRefCitation>
). The ascending process is not as developed as in
<taxonomicName id="4C524D35EB7B6412FD46FDE3B11EFD89" class="Reptilia" family="Iguanodontidae" genus="Barilium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="142" phylum="Chordata" rank="species" species="dawsoni">
<emphasis id="B926EAA4EB7B6412FD46FDE3B11EFD89" italics="true" pageId="17" pageNumber="142">Barilium dawsoni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C524D35EB7B6412FEE8FDCDB225FD89" box="[270,569,566,587]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="142" phylum="Chordata" rank="species" species="bernissartensis">
<emphasis id="B926EAA4EB7B6412FEE8FDCDB225FD89" box="[270,569,566,587]" italics="true" pageId="17" pageNumber="142">Iguanodon bernissartensis</emphasis>
</taxonomicName>
or
<taxonomicName id="4C524D35EB7B6412FDBBFDCDB158FDA8" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="142" phylum="Chordata" rank="species" species="athefieldensis">
<emphasis id="B926EAA4EB7B6412FDBBFDCDB158FDA8" italics="true" pageId="17" pageNumber="142">Mantellisaurus athefieldensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC34B47EB7B6412FEB3FDADB1E7FDA8" author="Norman DB" box="[341,507,597,619]" pageId="17" pageNumber="142" pagination="1 - 105" refId="ref18270" refString="Norman DB. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman, 1980</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB7B6412FDECFDAEB25EFDA9" author="Norman DB" box="[522,578,597,619]" pageId="17" pageNumber="142" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB7B6412FDB7FDAEB295FDA8" author="Norman DB" box="[593,649,597,618]" pageId="17" pageNumber="142" pagination="165 - 194" refId="ref18373" refString="Norman DB. 2011. On the osteology of the lower Wealden (Valanginian) ornithopod Barilium dawsoni (Iguanodontia: Styracosterna). Special Papers in Palaeontology 86: 165 - 194." type="journal article" year="2011">2011</bibRefCitation>
), but more closely resembles the condition found in
<taxonomicName id="4C524D35EB7B6412FD82FD8FB1D8FD6A" authority="(NMNH 2210)" baseAuthorityName="NMNH" baseAuthorityYear="2210" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="142" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B926EAA4EB7B6412FD82FD8FB0E4FD6A" italics="true" pageId="17" pageNumber="142">Camptosaurus dispar</emphasis>
(NMNH 2210)
</taxonomicName>
,
<emphasis id="B926EAA4EB7B6412FE3DFD68B31EFD6A" box="[475,770,658,680]" italics="true" pageId="17" pageNumber="142">
Cumnoria
<taxonomicName id="4C524D35EB7B6412FD88FD68B31EFD6A" authorityName="Hulke" authorityYear="1880" box="[622,770,658,680]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="142" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
</emphasis>
,
<taxonomicName id="4C524D35EB7B6412FF45FD4AB1CCFD05" authorityName="Taquet" authorityYear="1976" box="[163,464,689,711]" class="Reptilia" family="Iguanodontidae" genus="Ouranosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="142" phylum="Chordata" rank="species" species="nigeriensis">
<emphasis id="B926EAA4EB7B6412FF45FD4AB1CCFD05" box="[163,464,689,711]" italics="true" pageId="17" pageNumber="142">Ouranosaurus nigeriensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D35EB7B6412FDEDFD4AB317FD04" baseAuthorityName="Carpenter &amp; Wilson" baseAuthorityYear="2008" box="[523,779,689,710]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="142" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="B926EAA4EB7B6412FDEDFD4AB317FD04" box="[523,779,689,710]" italics="true" pageId="17" pageNumber="142">Uteodon aphanoecetes</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC34B47EB7B6412FF4BFD2BB1C2FD27" author="Galton PM &amp; Powell HP" box="[173,478,720,742]" pageId="17" pageNumber="142" pagination="411 - 443" refId="ref17215" refString="Galton PM, Powell HP. 1980. The ornithischian dinosaur Camptosaurus prestwichii from the Upper Jurassic of England. Palaeontology 23: 411 - 443." type="journal article" year="1980">Galton &amp; Powell, 1980</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB7B6412FE13FD2BB0C0FCC6" author="Carpenter K &amp; Wilson Y" pageId="17" pageNumber="142" pagination="227 - 264" refId="ref16499" refString="Carpenter K, Wilson Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter &amp; Wilson, 2008</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB7B6412FF0AFD14B1FFFCC6" author="Bertozzo F &amp; Dalla Vecchia FM &amp; Fabbri M" box="[236,483,750,772]" pageId="17" pageNumber="142" pagination="3403" refId="ref16369" refString="Bertozzo F, Dalla Vecchia FM, Fabbri M. 2017. The Venice specimen of Ouranosaurus nigeriensis (Dinosauria, Ornithopoda). PeerJ 5: e 3403." type="journal article" year="2017">
Bertozzo
<emphasis id="B926EAA4EB7B6412FEBBFD14B186FCC1" box="[349,410,750,772]" italics="true" pageId="17" pageNumber="142">et al.</emphasis>
, 2017
</bibRefCitation>
). As in the other known Portuguese ornithopod,
<taxonomicName id="4C524D35EB7B6412FE54FCF6B318FCE0" box="[434,772,781,802]" class="Reptilia" family="Dryosauridae" genus="Eousdryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="142" phylum="Chordata" rank="species" species="nanohallucis">
<emphasis id="B926EAA4EB7B6412FE54FCF6B318FCE0" box="[434,772,781,802]" italics="true" pageId="17" pageNumber="142">Eousdryosaurus nanohallucis</emphasis>
</taxonomicName>
, it shows a caudomedial process, which casts doubt on the diagnostic status of this character for this latter taxon (
<bibRefCitation id="EFC34B47EB7B6412FEDDFC91B212FCBC" author="Escaso F &amp; Ortega F &amp; Dantas P &amp; Malafaia E &amp; Silva BC &amp; Gasulla JM &amp; Mocho P &amp; Narvaez I &amp; Sanz JL" box="[315,526,873,895]" pageId="17" pageNumber="142" pagination="1102 - 1112" refId="ref16813" refString="Escaso F, Ortega F, Dantas P, Malafaia E, Silva BC, Gasulla JM, Mocho P, Narvaez I, Sanz JL. 2014. A new dryosaurid ornithopod (Dinosauria, Ornithischia) from the Late Jurassic of Portugal. Journal of Vertebrate Paleontology 34: 1102 - 1112." type="journal article" year="2014">
Escaso
<emphasis id="B926EAA4EB7B6412FE76FC91B1D5FCBC" box="[400,457,873,895]" italics="true" pageId="17" pageNumber="142">et al.</emphasis>
, 2014
</bibRefCitation>
). The calcaneum does not possess any remarkable differences from other ankylopollexians (Norman, 2004).
</paragraph>
<paragraph id="8BED36B6EB7B6412FF45FC0EB155FBCF" blockId="17.[163,329,1013,1037]" box="[163,329,1013,1037]" pageId="17" pageNumber="142">
<heading id="D0A581DAEB7B6412FF45FC0EB155FBCF" box="[163,329,1013,1037]" centered="true" fontSize="9" level="2" pageId="17" pageNumber="142" reason="2">
<emphasis id="B926EAA4EB7B6412FF45FC0EB155FBCF" box="[163,329,1013,1037]" italics="true" pageId="17" pageNumber="142">Distal tarsals</emphasis>
</heading>
</paragraph>
<paragraph id="8BED36B6EB7B6412FF45FBE5B31AFB29" blockId="17.[163,780,1053,1259]" pageId="17" pageNumber="142">
Two distal tarsals are preserved, which are identified as tarsal 3 and tarsal 4 (
<figureCitation id="13692A33EB7B6412FE2EFBC7B201FB93" box="[456,541,1084,1106]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="17" pageNumber="142">Fig. 8D</figureCitation>
). Tarsal 3 is a large element that is mediolaterally elongated and contacts metatarsal III along a straight and linear surface (
<figureCitation id="13692A33EB7B6412FD06FB82B0D8FB6F" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="17" pageNumber="142">Fig. 8D</figureCitation>
). Tarsal 4 is a stout, elongated element and contacts the metatarsal IV along a concave surface, while its medialmost tip overlaps tarsal 3 proximally (
<figureCitation id="13692A33EB7B6412FD42FB2EB2EBFB28" box="[676,759,1237,1259]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="17" pageNumber="142">Fig. 8D</figureCitation>
).
</paragraph>
<paragraph id="8BED36B6EB7B6412FF45FAEFB201F9C2" blockId="17.[163,779,1300,1536]" pageId="17" pageNumber="142">
<emphasis id="B926EAA4EB7B6412FF45FAEFB112FAEB" box="[163,270,1300,1321]" italics="true" pageId="17" pageNumber="142">Remarks:</emphasis>
Distal tarsals 3 and 4, found in articulation with metatarsals III and IV, resemble the cushionlike condition described by
<bibRefCitation id="EFC34B47EB7B6412FDF9FAAAB2F9FAA5" author="Gilmore CW" box="[543,741,1361,1383]" pageId="17" pageNumber="142" pagination="197 - 332" refId="ref17242" refString="Gilmore CW. 1909. Osteology of the Jurassic reptile Camptosaurus: with a revision of the species of the genus, and description of two new species. Proceedings of the United States National Museum 36: 197 - 332." type="journal article" year="1909">Gilmore (1909)</bibRefCitation>
in
<taxonomicName id="4C524D35EB7B6412FF45FA8BB1B4FA47" baseAuthorityName="NMNH" baseAuthorityYear="2210" box="[163,424,1392,1413]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="142" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B926EAA4EB7B6412FF45FA8BB1B4FA47" box="[163,424,1392,1413]" italics="true" pageId="17" pageNumber="142">Camptosaurus dispar</emphasis>
</taxonomicName>
, and differing from the thinlike and subrounded condition observed in
<taxonomicName id="4C524D35EB7B6412FD76FA75B144FA00" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="142" phylum="Chordata" rank="species" species="bernissartensis">
<emphasis id="B926EAA4EB7B6412FD76FA75B144FA00" italics="true" pageId="17" pageNumber="142">Iguanodon bernissartensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D35EB7B6412FE7EFA56B316FA00" authorityName="Mantell" authorityYear="1825" box="[408,778,1453,1474]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="142" phylum="Chordata" rank="species" species="atherfieldensis">
<emphasis id="B926EAA4EB7B6412FE7EFA56B316FA00" box="[408,778,1453,1474]" italics="true" pageId="17" pageNumber="142">Mantellisaurus atherfieldensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC34B47EB7B6412FF4AFA37B146FA23" author="Norman DB" box="[172,346,1484,1505]" pageId="17" pageNumber="142" pagination="1 - 105" refId="ref18270" refString="Norman DB. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman, 1980</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB7B6412FE8AFA37B1BAFA20" author="Norman DB" box="[364,422,1484,1506]" pageId="17" pageNumber="142" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
) or the one present in basal ornithopods (
<bibRefCitation id="EFC34B47EB7B6412FEDFFA11B1D6F9C2" author="Galton PM" box="[313,458,1514,1536]" pageId="17" pageNumber="142" refId="ref16969" refString="Galton PM. 1974. The ornithischian dinosaur Hypsilophodon from the Wealden of the Isle of Wight. London: British Museum (Natural History)." type="book" year="1974">Galton, 1974</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB7B6412FE31FA10B212F9C2" author="Galton PM" box="[471,526,1515,1536]" pageId="17" pageNumber="142" pagination="271 - 312" refId="ref17047" refString="Galton PM. 1981. Dryosaurus, a hypsilophodontid dinosaur from the Upper Jurassic of North America and Africa postcranial skeleton. Palaontologische Zeitschrift 55: 271 - 312." type="journal article" year="1981">1981</bibRefCitation>
).
</paragraph>
<paragraph id="8BED36B6EB7B6412FF45F9C2B131F992" blockId="17.[163,301,1593,1616]" box="[163,301,1593,1616]" pageId="17" pageNumber="142">
<heading id="D0A581DAEB7B6412FF45F9C2B131F992" box="[163,301,1593,1616]" centered="true" fontSize="9" level="2" pageId="17" pageNumber="142" reason="2">
<emphasis id="B926EAA4EB7B6412FF45F9C2B131F992" box="[163,301,1593,1616]" italics="true" pageId="17" pageNumber="142">Metatarsus</emphasis>
</heading>
</paragraph>
<paragraph id="8BED36B6EB7B6412FF45F99AB3E1FE2D" blockId="17.[163,779,1633,1900]" lastBlockId="17.[827,1444,197,495]" pageId="17" pageNumber="142">
Although they are fragmented, metatarsals IIIV are long and slender (
<figureCitation id="13692A33EB7B6412FE43F97BB1F1F957" box="[421,493,1664,1686]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="17" pageNumber="142">Figs 8</figureCitation>
,
<figureCitation id="13692A33EB7B6412FE1DF97BB227F957" box="[507,571,1664,1685]" captionStart="Figure 9" captionStartId="16.[146,224,1848,1870]" captionTargetBox="[305,1265,920,1809]" captionTargetId="figure-367@16.[305,1265,920,1809]" captionTargetPageId="16" captionText="Figure 9. Three dimensional models of the articulated leg (AB) and pes (CD) of Draconyx loureiroi holotype ML 357. Leg in medial view (A) and lateral view (B). Pes in medial (C) and lateral (D) views." figureDoi="http://doi.org/10.5281/zenodo.6530815" httpUri="https://zenodo.org/record/6530815/files/figure.png" pageId="17" pageNumber="142">9C, D</figureCitation>
). A thick splinter of bone next to metatarsal II was interpreted as the proximalmost part of metatarsal I in the original description of
<bibRefCitation id="EFC34B47EB7B6412FED9F927B243F930" author="Mateus O &amp; Antunes MT" box="[319,607,1756,1778]" pageId="17" pageNumber="142" pagination="61 - 73" refId="ref17993" refString="Mateus O, Antunes MT. 2001. Draconyx loureiroi, a new camptosauridae (Dinosauria, Ornithopoda) from the Late Jurassic of Lourinha, Portugal. Annales de Paleontologie 87: 61 - 73." type="journal article" year="2001">Mateus &amp; Antunes (2001)</bibRefCitation>
and a vestigial digit I has been proposed as a possible autapomorphy for
<taxonomicName id="4C524D35EB7B6412FF2DF8E1B1B8F8EC" authorityName="Mateus &amp; Antunes" authorityYear="2001" box="[203,420,1817,1839]" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="142" phylum="Chordata" rank="species" species="loureiroi">
<emphasis id="B926EAA4EB7B6412FF2DF8E1B1B8F8EC" box="[203,420,1817,1839]" italics="true" pageId="17" pageNumber="142">Draconyx loureiroi</emphasis>
</taxonomicName>
. Despite being heavily eroded, the bone is preserved in anatomical position and the rod-like structure supports the interpretation of this bone as the first metatarsal (
<figureCitation id="13692A33EB7B6412FB4BFF3EB518FF18" box="[1197,1284,197,219]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="17" pageNumber="142">Fig. 8F</figureCitation>
). The status of this character as an autapomorphy is discussed below. A deep groove in continuity with this fragment and impressed on metatarsal II probably represents the entire articulation surface with metatarsal I (
<figureCitation id="13692A33EB7B6412FA9EFEBBB346FEB6" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="17" pageNumber="142">Fig. 8F</figureCitation>
). Metatarsals II and III are complete and do not appear to have suffered extensive breakage or fracture (
<figureCitation id="13692A33EB7B6412FCA5FE67B3AEFE73" box="[835,946,412,434]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="17" pageNumber="142">Fig. 8C, D</figureCitation>
). Metatarsal IV in contrast is broken at the midshaft with the distal epiphysis preserved as an isolated element.
</paragraph>
<paragraph id="8BED36B6EB7B6412FCDDFDDCB3CFFDFD" blockId="17.[827,979,551,575]" box="[827,979,551,575]" pageId="17" pageNumber="142">
<heading id="D0A581DAEB7B6412FCDDFDDCB3CFFDFD" box="[827,979,551,575]" centered="true" fontSize="9" level="2" pageId="17" pageNumber="142" reason="2">
<emphasis id="B926EAA4EB7B6412FCDDFDDCB3CFFDFD" box="[827,979,551,575]" italics="true" pageId="17" pageNumber="142">Pedal digit I</emphasis>
</heading>
</paragraph>
<paragraph id="8BED36B6EB7B6412FCDDFDABB42AFD46" blockId="17.[827,1442,592,644]" pageId="17" pageNumber="142">
The first pedal digit consists only of the splint-like metatarsal I (
<figureCitation id="13692A33EB7B6412FC31FD95B43BFD46" box="[983,1063,622,644]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="17" pageNumber="142">Fig. 8F</figureCitation>
).
</paragraph>
<paragraph id="8BED36B6EB7B6412FCDDFD46B3C2FD16" blockId="17.[827,990,701,725]" box="[827,990,701,725]" pageId="17" pageNumber="142">
<heading id="D0A581DAEB7B6412FCDDFD46B3C2FD16" box="[827,990,701,725]" centered="true" fontSize="9" level="2" pageId="17" pageNumber="142" reason="2">
<emphasis id="B926EAA4EB7B6412FCDDFD46B3C2FD16" box="[827,990,701,725]" italics="true" pageId="17" pageNumber="142">Pedal digit II</emphasis>
</heading>
</paragraph>
<paragraph id="8BED36B6EB7B6412FCDDFD1EB3FDFC56" blockId="17.[827,1444,741,1714]" pageId="17" pageNumber="142">
Pedal digit II is represented by metatarsal II, the pedal phalanx II-1 (ML 35717) and the ungual phalanx II-2 (35716), which articulates with the phalanx II-2 (357 14). Pedal phalanx II-1 is broken at the midshaft, but the proximal and distalmost epiphyses are preserved (
<figureCitation id="13692A33EB7B6412FCA5FC85B3CDFC56" box="[835,977,894,916]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="17" pageNumber="142">Fig. 8A, C, D</figureCitation>
).
</paragraph>
<paragraph id="8BED36B6EB7B6412FCB5FC66B513F973" blockId="17.[827,1444,741,1714]" pageId="17" pageNumber="142">
Metatarsal II is positioned more proximally with respect to Mt IIIIV. In mediolateral view, metatarsal II has a keyhole shape, as described by
<bibRefCitation id="EFC34B47EB7B6412FAF1FC20B39BFBCD" author="Herne MC &amp; Tait AM &amp; Weisbecker V &amp; Hall M &amp; Nair JP &amp; Cleeland M &amp; Salisbury SW" pageId="17" pageNumber="142" pagination="4113" refId="ref17392" refString="Herne MC, Tait AM, Weisbecker V, Hall M, Nair JP, Cleeland M, Salisbury SW. 2018. A new small-bodied ornithopod (Dinosauria, Ornithischia) from a deep, highenergy Early Cretaceous river of the Australian-Antarctic rift system. PeerJ 5: e 4113." type="journal article" year="2018">
Herne
<emphasis id="B926EAA4EB7B6412FA8EFC20B580FC2D" box="[1384,1436,986,1008]" italics="true" pageId="17" pageNumber="142">et al</emphasis>
. (2018)
</bibRefCitation>
for the Australian ornithopod
<taxonomicName id="4C524D35EB7B6412FAECFC02B495FBEF" authority="Herne et al., 2018" authorityName="Herne" authorityYear="2018" class="Reptilia" genus="Diluvicursor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="142" phylum="Chordata" rank="species" species="pickeringi">
<emphasis id="B926EAA4EB7B6412FAECFC02B3B3FBEF" italics="true" pageId="17" pageNumber="142">Diluvicursor pickeringi</emphasis>
<bibRefCitation id="EFC34B47EB7B6412FC51FBE3B495FBEF" author="Herne MC &amp; Tait AM &amp; Weisbecker V &amp; Hall M &amp; Nair JP &amp; Cleeland M &amp; Salisbury SW" box="[951,1161,1048,1069]" pageId="17" pageNumber="142" pagination="4113" refId="ref17392" refString="Herne MC, Tait AM, Weisbecker V, Hall M, Nair JP, Cleeland M, Salisbury SW. 2018. A new small-bodied ornithopod (Dinosauria, Ornithischia) from a deep, highenergy Early Cretaceous river of the Australian-Antarctic rift system. PeerJ 5: e 4113." type="journal article" year="2018">
Herne
<emphasis id="B926EAA4EB7B6412FBEEFBE3B427FBEF" box="[1032,1083,1048,1069]" italics="true" pageId="17" pageNumber="142">et al</emphasis>
., 2018
</bibRefCitation>
</taxonomicName>
, with the proximalmost part dorsoplantarily higher than the distal epiphysis (
<figureCitation id="13692A33EB7B6412FCA2FBAEB3DFFBA9" box="[836,963,1109,1131]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="17" pageNumber="142">Fig. 8A, F</figureCitation>
). The distal epiphysis develops more dorsoplantarily than mediolaterally, and the articular surface is convex. The shaft of Mt II is piriform (teardrop) in cross-section, with the plantar surface bearing a keel. The distalmost part of the epiphysis expands dorsoplantarily, giving the condyle a subtrapezoidal outline (
<figureCitation id="13692A33EB7B6412FBADFAF6B4A0FAE1" box="[1099,1212,1293,1315]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="17" pageNumber="142">Fig. 8A, D</figureCitation>
). The condyle of the metatarsal articulates perfectly with pedal phalanx II-1, which accommodates the condyle in a gently convex triangular facet. The distal epiphysis of the phalanx is in articulation with pedal phalanx II-2, which is a thick, subrectangular element in dorsal view (
<figureCitation id="13692A33EB7B6412FCA5FA3EB3B2FA18" box="[835,942,1477,1499]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="17" pageNumber="142">Fig. 8C, D</figureCitation>
). Proximally, the articular facet is not visible, but distally the two condyles are well distinguishable, the lateral slightly inclined with respect to the medial one. On both lateral surfaces, deep extensor grooves are present for tendon insertion. The ungual (phalanx II-2) of the pedal digit II is preserved, being a pointed claw-like element, which is dorsoplantarly arched and proximodistally elongated (
<figureCitation id="13692A33EB7B6412FB97F967B51CF973" box="[1137,1280,1692,1714]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="17" pageNumber="142">Fig. 8A, C, D</figureCitation>
).
</paragraph>
<paragraph id="8BED36B6EB7B6412FCDDF909B3F5F8C8" blockId="17.[827,1001,1778,1802]" box="[827,1001,1778,1802]" pageId="17" pageNumber="142">
<heading id="D0A581DAEB7B6412FCDDF909B3F5F8C8" box="[827,1001,1778,1802]" centered="true" fontSize="9" level="2" pageId="17" pageNumber="142" reason="2">
<emphasis id="B926EAA4EB7B6412FCDDF909B3F5F8C8" box="[827,1001,1778,1802]" italics="true" pageId="17" pageNumber="142">Pedal digit III</emphasis>
</heading>
</paragraph>
<paragraph id="8BED36B6EB7B6411FCDDF8E1B1C3FC1B" blockId="17.[827,1442,1818,1902]" lastBlockId="18.[145,761,197,986]" lastPageId="18" lastPageNumber="143" pageId="17" pageNumber="142">
Pedal digit III is composed of metatarsal III, the largest pedal phalanx preserved III-1 (ML 35713), the smallest pedal phalanx is interpreted to be III-3 (ML 35715) and the largest ungual, pedal phalanx III-4 (ML 35719). The pedal phalanx III-2 is missing. Metatarsal III is the longest element of the metatarsus, being a slender, yet compact element. Proximally, the Mt III articulates with the tarsal 3 and intervenes between Mt II and Mt IV (
<figureCitation id="13692A33EB786411FDA2FEA5B2ABFEB6" box="[580,695,350,372]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="18" pageNumber="143">Fig. 8BD</figureCitation>
). The subrectangular shaft ends distally in a mediolaterally expanded condyle, the only metatarsal exhibiting this condition. Pedal phalanx III-1 is a stout, robust element, which is moderately dorsoplantarly arched. The proximal articular facet is concave and subellipsoidal in outline, being the long axis oriented mediolaterally (
<figureCitation id="13692A33EB786411FE57FDCEB238FD88" box="[433,548,565,587]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="18" pageNumber="143">Fig. 8BD</figureCitation>
). Distally, the two ginglymi are subequal in size, being marked by deep collateral ligament pits. Pedal phalanx III-3 is a small proximodistally compressed element, which possesses a dorsoplantarly deep articular facet (
<figureCitation id="13692A33EB786411FDA3FD4BB2A4FD07" box="[581,696,688,710]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="18" pageNumber="143">Fig. 8BD</figureCitation>
). The ventral margin of pedal phalanx III-3 is not as arched as the other phalanges and the collateral ligament pits occupy almost the entire lateral and medial surfaces. The ungual of the digit III (pedal phalanx III-3) is a stout, pointed element, which is more robust than the ungual preserved on pedal digit II. The ungual of digit III differs also from the other preserved ungual in having a less arched and more dorsoplantarly flattened outline (
<figureCitation id="13692A33EB786411FEBBFC3FB1D3FC1B" box="[349,463,964,986]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="18" pageNumber="143">Fig. 8BD</figureCitation>
).
</paragraph>
<paragraph id="8BED36B6EB786411FF77FBE9B120FBE8" blockId="18.[145,316,1042,1066]" box="[145,316,1042,1066]" pageId="18" pageNumber="143">
<heading id="D0A581DAEB786411FF77FBE9B120FBE8" box="[145,316,1042,1066]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="143" reason="2">
<emphasis id="B926EAA4EB786411FF77FBE9B120FBE8" box="[145,316,1042,1066]" italics="true" pageId="18" pageNumber="143">Pedal digit IV</emphasis>
</heading>
</paragraph>
<paragraph id="8BED36B6EB786411FF77FBC1B101FA87" blockId="18.[145,760,1082,1350]" pageId="18" pageNumber="143">
Metatarsal IV (ML 35712, 18) is the only element of pedal digit IV available. Proximally it is the most mediolaterally expanded and dorsoplantarly compressed of the metatarsals and the proximal epiphysis is fan-shaped in dorsal and plantar view (
<figureCitation id="13692A33EB786411FF3DFB2FB149FB2B" box="[219,341,1236,1258]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="18" pageNumber="143">Fig. 8BD</figureCitation>
). Distally, the section of the shaft becomes progressively subrectangular, culminating in a moderately dorsoplantarly deep distal condyle (
<figureCitation id="13692A33EB786411FF7FFACBB111FA87" box="[153,269,1328,1350]" captionStart="Figure 8" captionStartId="15.[163,241,1778,1800]" captionTargetBox="[227,1379,195,1738]" captionTargetId="figure-27@15.[227,1379,195,1738]" captionTargetPageId="15" captionText="Figure 8. Three dimensional models of the pes of Draconyx loureiroi holotype ML 357. Articulated pes (ML 357-12) (AF) in medial (A), lateral (B), dorsal (C), plantar (D) and caudal (E) views, F, detail of metatarsal I.Abbreviations: as: astragalus, ca, calcalneum, fi: fibula, lm: lateral malleolus, mm: medial malleolus, mt-I: metatarsal I, mt-II: metatarsal II, mt-III: metatarsal III, mt-IV: metatarsal IV, p II-1: pedal phalanx II-1, p II-1: pedal phalanx II-2, p III-1: pedal phalanx III-1, p III-3: pedal phalanx III-3, ta-3: tarsal 3, ta-4: tarsal 4, un-II: ungual II, un-III: ungual III." figureDoi="http://doi.org/10.5281/zenodo.6530813" httpUri="https://zenodo.org/record/6530813/files/figure.png" pageId="18" pageNumber="143">Fig. 8BD</figureCitation>
).
</paragraph>
<paragraph id="8BED36B6EB786411FF77FA95B4D2FDCF" blockId="18.[145,762,1390,1903]" lastBlockId="18.[809,1426,197,526]" pageId="18" pageNumber="143">
<emphasis id="B926EAA4EB786411FF77FA95B0E2FA41" box="[145,254,1390,1411]" italics="true" pageId="18" pageNumber="143">Remarks:</emphasis>
The metatarsus of ML 357 differs strongly from those of
<taxonomicName id="4C524D35EB786411FEAFFA76B242FA60" baseAuthorityName="NMNH" baseAuthorityYear="2210" box="[329,606,1421,1442]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="18" pageNumber="143" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B926EAA4EB786411FEAFFA76B242FA60" box="[329,606,1421,1442]" italics="true" pageId="18" pageNumber="143">Camptosaurus dispar</emphasis>
</taxonomicName>
,
<taxonomicName id="4C524D35EB786411FD94FA76B158FA03" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="18" pageNumber="143" phylum="Chordata" rank="species" species="bernissartensis">
<emphasis id="B926EAA4EB786411FD94FA76B158FA03" italics="true" pageId="18" pageNumber="143">Iguanodon bernissartensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C524D35EB786411FEB5FA57B294FA03" authorityName="Taquet" authorityYear="1976" box="[339,648,1452,1473]" class="Reptilia" family="Iguanodontidae" genus="Ouranosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="18" pageNumber="143" phylum="Chordata" rank="species" species="nigeriensis">
<emphasis id="B926EAA4EB786411FEB5FA57B294FA03" box="[339,648,1452,1473]" italics="true" pageId="18" pageNumber="143">Ouranosaurus nigeriensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C524D35EB786411FD71FA57B136FA1D" baseAuthorityName="Carpenter &amp; Wilson" baseAuthorityYear="2008" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="18" pageNumber="143" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="B926EAA4EB786411FD71FA57B136FA1D" italics="true" pageId="18" pageNumber="143">Uteodon aphanoecetes</emphasis>
</taxonomicName>
and other large-sized hadrosauriforms in general proportions, while exhibiting the slender and gracile proportions seen in
<emphasis id="B926EAA4EB786411FE18F9F3B2E5F9DF" box="[510,761,1544,1565]" italics="true" pageId="18" pageNumber="143">
Cumnoria
<taxonomicName id="4C524D35EB786411FD9CF9F3B2E5F9DF" authorityName="Hulke" authorityYear="1880" box="[634,761,1544,1565]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="18" pageNumber="143" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
</emphasis>
and
<taxonomicName id="4C524D35EB786411FF22F9DDB23BF9F9" authorityName="Mantell" authorityYear="1825" box="[196,551,1574,1595]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="18" pageNumber="143" phylum="Chordata" rank="species" species="atherfieldensis">
<emphasis id="B926EAA4EB786411FF22F9DDB23BF9F9" box="[196,551,1574,1595]" italics="true" pageId="18" pageNumber="143">Mantellisaurus atherfieldensis</emphasis>
</taxonomicName>
instead (
<bibRefCitation id="EFC34B47EB786411FD73F9DDB0D6F998" author="Gilmore CW" pageId="18" pageNumber="143" pagination="197 - 332" refId="ref17242" refString="Gilmore CW. 1909. Osteology of the Jurassic reptile Camptosaurus: with a revision of the species of the genus, and description of two new species. Proceedings of the United States National Museum 36: 197 - 332." type="journal article" year="1909">Gilmore, 1909</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB786411FF3CF9BEB1F0F998" author="Galton PM &amp; Powell HP" box="[218,492,1605,1627]" pageId="18" pageNumber="143" pagination="411 - 443" refId="ref17215" refString="Galton PM, Powell HP. 1980. The ornithischian dinosaur Camptosaurus prestwichii from the Upper Jurassic of England. Palaeontology 23: 411 - 443." type="journal article" year="1980">Galton &amp; Powell, 1980</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB786411FE1BF9BDB2B4F998" author="Norman DB" box="[509,680,1605,1627]" pageId="18" pageNumber="143" pagination="1 - 105" refId="ref18270" refString="Norman DB. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman, 1980</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB786411FD5FF9BEB2EEF999" author="Norman DB" box="[697,754,1605,1627]" pageId="18" pageNumber="143" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
, 2004;
<bibRefCitation id="EFC34B47EB786411FF3FF99FB1D2F9BB" author="Bertozzo F &amp; Dalla Vecchia FM &amp; Fabbri M" box="[217,462,1636,1657]" pageId="18" pageNumber="143" pagination="3403" refId="ref16369" refString="Bertozzo F, Dalla Vecchia FM, Fabbri M. 2017. The Venice specimen of Ouranosaurus nigeriensis (Dinosauria, Ornithopoda). PeerJ 5: e 3403." type="journal article" year="2017">
Bertozzo
<emphasis id="B926EAA4EB786411FEAFF99FB199F9BB" box="[329,389,1636,1657]" italics="true" pageId="18" pageNumber="143">et al.</emphasis>
, 2017
</bibRefCitation>
). Furthermore, as in the latter taxa, ML 357 possesses a metatarsal III that is sensibly more elongated with respect to metatarsals II and IV (
<bibRefCitation id="EFC34B47EB786411FF14F93BB1E5F917" author="Galton PM &amp; Powell HP" box="[242,505,1728,1750]" pageId="18" pageNumber="143" pagination="411 - 443" refId="ref17215" refString="Galton PM, Powell HP. 1980. The ornithischian dinosaur Camptosaurus prestwichii from the Upper Jurassic of England. Palaeontology 23: 411 - 443." type="journal article" year="1980">Galton &amp; Powell, 1980</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB786411FDEEF93BB2B3F914" author="Norman DB" box="[520,687,1728,1750]" pageId="18" pageNumber="143" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">Norman, 1986</bibRefCitation>
). This is in stark contrast to elasmarians
<taxonomicName id="4C524D35EB786411FDAFF924B17DF8D0" authority=", Cumnoria" authorityName="Cumnoria" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="18" pageNumber="143" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B926EAA4EB786411FDAFF924B0C1F8D0" italics="true" pageId="18" pageNumber="143">Camptosaurus dispar</emphasis>
,
<emphasis id="B926EAA4EB786411FF0DF906B17DF8D0" box="[235,353,1789,1810]" italics="true" pageId="18" pageNumber="143">Cumnoria</emphasis>
</taxonomicName>
<taxonomicName id="4C524D35EB786411FE8CF906B1F7F8D0" authorityName="Hulke" authorityYear="1880" box="[362,491,1789,1810]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="18" pageNumber="143" phylum="Chordata" rank="species" species="prestwichii">
<emphasis id="B926EAA4EB786411FE8CF906B1F7F8D0" box="[362,491,1789,1810]" italics="true" pageId="18" pageNumber="143">prestwichii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C524D35EB786411FE1CF905B2D9F8D0" authorityName="Marsh" authorityYear="1878" box="[506,709,1789,1811]" class="Reptilia" family="Dryosauridae" genus="Dryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="18" pageNumber="143" phylum="Chordata" rank="species" species="altus">
<emphasis id="B926EAA4EB786411FE1CF905B2D9F8D0" box="[506,709,1789,1811]" italics="true" pageId="18" pageNumber="143">Dryosaurus altus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D35EB786411FF77F8E7B1F3F8F3" authorityName="Virchow" authorityYear="1919" box="[145,495,1820,1841]" class="Reptilia" family="Dryosauridae" genus="Dysalotosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="18" pageNumber="143" phylum="Chordata" rank="species" species="lettowvorbecki">
<emphasis id="B926EAA4EB786411FF77F8E7B1F3F8F3" box="[145,495,1820,1841]" italics="true" pageId="18" pageNumber="143">Dysalotosaurus lettowvorbecki</emphasis>
</taxonomicName>
, whose metatarsal I is extremely reduced, losing any functional capability (
<bibRefCitation id="EFC34B47EB786411FF7FF8A2B121F8AC" author="Gilmore CW" box="[153,317,1881,1903]" pageId="18" pageNumber="143" pagination="197 - 332" refId="ref17242" refString="Gilmore CW. 1909. Osteology of the Jurassic reptile Camptosaurus: with a revision of the species of the genus, and description of two new species. Proceedings of the United States National Museum 36: 197 - 332." type="journal article" year="1909">Gilmore, 1909</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB786411FEADF8A2B24CF8AC" author="Galton PM &amp; Powell HP" box="[331,592,1881,1903]" pageId="18" pageNumber="143" pagination="411 - 443" refId="ref17215" refString="Galton PM, Powell HP. 1980. The ornithischian dinosaur Camptosaurus prestwichii from the Upper Jurassic of England. Palaeontology 23: 411 - 443." type="journal article" year="1980">Galton &amp; Powell, 1980</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB786411FDB8F8A2B2EFF8AC" author="Galton PM" box="[606,755,1881,1903]" pageId="18" pageNumber="143" pagination="271 - 312" refId="ref17047" refString="Galton PM. 1981. Dryosaurus, a hypsilophodontid dinosaur from the Upper Jurassic of North America and Africa postcranial skeleton. Palaontologische Zeitschrift 55: 271 - 312." type="journal article" year="1981">Galton, 1981</bibRefCitation>
;
<bibRefCitation id="EFC34B47EB786411FCCFFF3EB443FF18" author="Carpenter K &amp; Galton PM" box="[809,1119,197,219]" pageId="18" pageNumber="143" pagination="167 - 207" refId="ref16420" refString="Carpenter K, Galton PM. 2018. A photo documentation of bipedal ornithischian dinosaurs from the Upper Jurassic Morrison Formation, USA. Geology of the Intermountain West 5: 167 - 207." type="journal article" year="2018">Carpenter &amp; Galton, 2018</bibRefCitation>
). However, this condition is preserved in some specimens of
<taxonomicName id="4C524D35EB786411FAE1FF1FB3C2FED5" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="18" pageNumber="143" phylum="Chordata" rank="species" species="bernissartensis">
<emphasis id="B926EAA4EB786411FAE1FF1FB3C2FED5" italics="true" pageId="18" pageNumber="143">Iguanodon bernissartensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D35EB786411FBF8FEF9B58CFED5" authorityName="Mantell" authorityYear="1825" box="[1054,1424,258,279]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="18" pageNumber="143" phylum="Chordata" rank="species" species="atherfieldensis">
<emphasis id="B926EAA4EB786411FBF8FEF9B58CFED5" box="[1054,1424,258,279]" italics="true" pageId="18" pageNumber="143">Mantellisaurus atherfieldensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC34B47EB786411FCD7FEDAB3CEFEF4" author="Norman DB" box="[817,978,289,310]" pageId="18" pageNumber="143" pagination="1 - 105" refId="ref18270" refString="Norman DB. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman 1980</bibRefCitation>
,
<bibRefCitation id="EFC34B47EB786411FC06FEDAB404FEF5" author="Norman DB" box="[992,1048,289,311]" pageId="18" pageNumber="143" pagination="281 - 372" refId="ref18307" refString="Norman DB. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
). Assessing the presence of this character in other taxa is problematic, because of the extreme fragility of the reduced metatarsal I, whose absence/presence is highly subject to preservation bias. The alleged dryosaurid,
<taxonomicName id="4C524D35EB786411FBDFFE67B596FE73" box="[1081,1418,412,433]" class="Reptilia" family="Dryosauridae" genus="Eousdryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="18" pageNumber="143" phylum="Chordata" rank="species" species="nanohallucis">
<emphasis id="B926EAA4EB786411FBDFFE67B596FE73" box="[1081,1418,412,433]" italics="true" pageId="18" pageNumber="143">Eousdryosaurus nanohallucis</emphasis>
</taxonomicName>
, from the same formation as ML 357, has been shown to possess a reduced metatarsal I in articulation with a pedal phalanx (
<bibRefCitation id="EFC34B47EB786411FC08FE03B4A2FDCF" author="Escaso F &amp; Ortega F &amp; Dantas P &amp; Malafaia E &amp; Silva BC &amp; Gasulla JM &amp; Mocho P &amp; Narvaez I &amp; Sanz JL" box="[1006,1214,504,525]" pageId="18" pageNumber="143" pagination="1102 - 1112" refId="ref16813" refString="Escaso F, Ortega F, Dantas P, Malafaia E, Silva BC, Gasulla JM, Mocho P, Narvaez I, Sanz JL. 2014. A new dryosaurid ornithopod (Dinosauria, Ornithischia) from the Late Jurassic of Portugal. Journal of Vertebrate Paleontology 34: 1102 - 1112." type="journal article" year="2014">
Escaso
<emphasis id="B926EAA4EB786411FBA5FE03B467FDCF" box="[1091,1147,504,525]" italics="true" pageId="18" pageNumber="143">et al.</emphasis>
, 2014
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>