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<document id="122E97BC840809C55EA52DC6F463E2BA" ID-CLB-Dataset="21232" ID-DOI="10.11646/zootaxa.4608.1.3" ID-GBIF-Dataset="c4d067fa-4c56-4248-8084-caf4dd1eea21" ID-ISSN="1175-5326" ID-Zenodo-Dep="3993611" ID-ZooBank="3540218C-34FA-4D91-925A-3E89C6C3A7C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1597950731833" checkinUser="felipe" docAuthor="Venugopal, Akhil S. &amp; Thomas, Sabu K." docDate="2019" docId="03FB8782FFFB775D4AFEFF343669FD20" docLanguage="en" docName="zootaxa.4608.1.3.pdf" docOrigin="Zootaxa 4608 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pheropsophus andrewesi Jedlicka 1964" docType="treatment" docVersion="8" lastPageNumber="68" masterDocId="FFC2FFFAFFF8775E4A69FFAD3561FE32" masterDocTitle="Bombardier beetles of the genus Pheropsophus Solier 1833 (Carabidae: Brachininae: Brachinini) from Indian subcontinent" masterLastPageNumber="89" masterPageNumber="65" pageNumber="68" updateTime="1698845024281" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FCCE60E40921059EF613B9CBC3DE30CD">Bombardier beetles of the genus Pheropsophus Solier 1833 (Carabidae: Brachininae: Brachinini) from Indian subcontinent</mods:title>
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<mods:namePart id="3F0DC8BB3FA9845DCA9E913D356021E3">Venugopal, Akhil S.</mods:namePart>
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<treatment id="03FB8782FFFB775D4AFEFF343669FD20" ID-DOI="http://doi.org/10.5281/zenodo.4335061" ID-GBIF-Taxon="167284773" ID-Zenodo-Dep="4335061" LSID="urn:lsid:plazi:treatment:03FB8782FFFB775D4AFEFF343669FD20" httpUri="http://treatment.plazi.org/id/03FB8782FFFB775D4AFEFF343669FD20" lastPageNumber="68" pageId="3" pageNumber="68">
<subSubSection id="C348651FFFFB775D4AFEFF3437CFFE81" box="[151,686,152,180]" pageId="3" pageNumber="68" type="nomenclature">
<paragraph id="8BED3694FFFB775D4AFEFF3437CFFE81" blockId="3.[151,686,152,216]" box="[151,686,152,180]" pageId="3" pageNumber="68">
<heading id="D0A581F8FFFB775D4AFEFF3437CFFE81" bold="true" box="[151,686,152,180]" fontSize="11" level="1" pageId="3" pageNumber="68" reason="1">
<taxonomicName id="4C524D17FFFB775D4AFEFF3437CFFE81" ID-CoL="4G385" authority="JEDLICKA, 1964" authorityName="Jedlicka" authorityYear="1964" box="[151,686,152,180]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B926EA86FFFB775D4AFEFF3437CFFE81" bold="true" box="[151,686,152,180]" pageId="3" pageNumber="68">
<emphasis id="B926EA86FFFB775D4AFEFF3434A4FE86" bold="true" box="[151,453,153,180]" italics="true" pageId="3" pageNumber="68">Pheropsophus andrewesi</emphasis>
<bibRefCitation id="EFC34B65FFFB775D4BA4FF3537CFFE81" author="Jedlicka, A." box="[461,686,152,180]" pageId="3" pageNumber="68" pagination="1 - 6" refId="ref19878" refString="Jedlicka, A. (1964) Neue Carabiden aus der Palaarctischen region (Coleoptera). Annotationes Zoologicae et Botanicae, 3, 1 - 6." type="journal article" year="1964">JEDLIČKA, 1964</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C348651FFFFB775D4AFEFF103464FEEA" box="[151,261,189,216]" pageId="3" pageNumber="68" type="description">
<paragraph id="8BED3694FFFB775D4AFEFF103464FEEA" blockId="3.[151,686,152,216]" box="[151,261,189,216]" pageId="3" pageNumber="68">
(
<figureCitation id="13692A11FFFB775D4AC9FF10359CFEEA" box="[160,253,189,216]" captionStart="FIGURE 1" captionStartId="4.[151,250,1869,1894]" captionTargetBox="[205,1381,207,1825]" captionTargetId="figure@4.[196,1392,196,1829]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of Pheropsophus from India A) Pheropsophus andrewesi (Holotype, India: Assam, ZMHB), B) P. aptinoides (Holotype, East Indies, MNHN), C) P. bimaculatus (Holotype, Ind.Or., LSUK), D) P. cardoni (Syntype, Chota Nagpore, Nowatoli, ZMHB), E) P. catoirei (Holotype, Bengalia, MNHN), F) P. chaudoiri (Holotype, Nepal, BMNH), G) P. consularis (Syntype, Burma, NMPC), H) P. javanus (Holotype, Java, MNHN), I) P. lineifrons (Holotype, Ind.Or., ZMHB)" figureDoi="http://doi.org/10.5281/zenodo.3993613" httpUri="https://zenodo.org/record/3993613/files/figure.png" pageId="3" pageNumber="68">Fig. 1 A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C348651FFFFB775D4AFEFEAF3776FFB5" pageId="3" pageNumber="68" type="reference_group">
<paragraph id="8BED3694FFFB775D4AFEFEAF3624FF28" blockId="3.[151,837,257,282]" box="[151,837,257,282]" pageId="3" pageNumber="68">
<taxonomicName id="4C524D17FFFB775D4AFEFEAF3722FF28" ID-CoL="4G385" authority="Jedlicka, 1964: 5" authorityName="Jedlicka" authorityPageNumber="5" authorityYear="1964" box="[151,579,257,282]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B926EA86FFFB775D4AFEFEAF34F4FF28" box="[151,405,258,282]" italics="true" pageId="3" pageNumber="68">Pheropsophus andrewesi</emphasis>
<bibRefCitation id="EFC34B65FFFB775D4BF2FEAC3722FF28" author="Jedlicka, A." box="[411,579,257,282]" pageId="3" pageNumber="68" pagination="1 - 6" refId="ref19878" refString="Jedlicka, A. (1964) Neue Carabiden aus der Palaarctischen region (Coleoptera). Annotationes Zoologicae et Botanicae, 3, 1 - 6." type="journal article" year="1964">Jedlička, 1964:5</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EFC34B65FFFB775D4826FEAC3620FF28" author="Kirschenhofer, E." box="[591,833,257,282]" pageId="3" pageNumber="68" pagination="25 - 64" refId="ref19906" refString="Kirschenhofer, E. (2010) New and Little known species of Carabidae from Middle East and Southeast Asia (Coleoptera: Carabidae: Lebiini, Brachinini). Annales Historico-Naturales Musei Nationalis Hungarici, 102, 25 - 64." type="journal article" year="2010">Kirschenhofer, 2010:55</bibRefCitation>
.
</paragraph>
<paragraph id="8BED3694FFFB775D4AFEFEEA3776FFB5" blockId="3.[151,1437,327,787]" pageId="3" pageNumber="68">
<emphasis id="B926EA86FFFB775D4AFEFEEA34B5FF50" bold="true" box="[151,468,327,354]" pageId="3" pageNumber="68">Specimens examined (n=1)</emphasis>
:
<materialsCitation id="3B3A3CC9FFFB775D4B8BFEE53776FFB5" ID-GBIF-Occurrence="2846550334" collectionCode="ZMHB" country="India" determinerName="Ing. Jedlicka" location="Assam" pageId="3" pageNumber="68" specimenCount="2" stateProvince="Assam" typeStatus="holotype">
<typeStatus id="54E98836FFFB775D4B8BFEE5372FFF50" box="[482,590,328,354]" pageId="3" pageNumber="68" type="holotype">Holotype</typeStatus>
; labelled “
<collectingCountry id="F3457604FFFB775D48A8FEE53661FF50" box="[705,768,328,354]" name="India" pageId="3" pageNumber="68">India</collectingCountry>
,
<collectingRegion id="4996F876FFFB775D4961FEE53636FF50" box="[776,855,328,354]" country="India" name="Assam" pageId="3" pageNumber="68">Assam</collectingRegion>
,
<typeStatus id="54E98836FFFB775D4908FEE536B8FF50" box="[865,985,328,354]" pageId="3" pageNumber="68" type="holotype">Holotypus</typeStatus>
,
<taxonomicName id="4C524D17FFFB775D498AFEE530F4FF50" ID-CoL="4G385" authorityName="Jedlicka" authorityYear="1964" box="[995,1429,328,354]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="andrewesi" subGenus="Stenaptinus">
<emphasis id="B926EA86FFFB775D498AFEE530F4FF50" box="[995,1429,328,354]" italics="true" pageId="3" pageNumber="68">Pheropsophus (Stenaptinus) andrewesi</emphasis>
</taxonomicName>
, det.
<determinerName id="7E31DA50FFFB775D4AAEFEC0343CFFB4" box="[199,349,364,391]" pageId="3" pageNumber="68">Ing. Jedlička</determinerName>
”; type in
<collectionCode id="ED43AE51FFFB775D4BAFFEC03776FFB5" box="[454,535,365,391]" pageId="3" pageNumber="68">ZMHB</collectionCode>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C348651FFFFB775D4AAEFE223015FCF8" pageId="3" pageNumber="68" type="description">
<paragraph id="8BED3694FFFB775D4AAEFE223707FF98" blockId="3.[151,1437,327,787]" box="[199,614,399,427]" pageId="3" pageNumber="68">
<emphasis id="B926EA86FFFB775D4AAEFE223438FF98" bold="true" box="[199,345,399,426]" pageId="3" pageNumber="68">Description.</emphasis>
Length: 11.015.0 mm.
</paragraph>
<paragraph id="8BED3694FFFB775D4AAEFE1E315AFFC1" blockId="3.[151,1437,327,787]" pageId="3" pageNumber="68">
<emphasis id="B926EA86FFFB775D4AAEFE1E347AFFFC" bold="true" box="[199,283,435,462]" pageId="3" pageNumber="68">Colour</emphasis>
: Head, pronotum and elytra black; palps, legs, clypeus, labrum and two spots next to the antennae yellow-red. Segments 14 of antennae (yellowish red) lighter than 511 (dark brown).
</paragraph>
<paragraph id="8BED3694FFFB775D4AAEFE513014FC24" blockId="3.[151,1437,327,787]" box="[199,1397,508,535]" pageId="3" pageNumber="68">
<emphasis id="B926EA86FFFB775D4AAEFE513471FC24" bold="true" box="[199,272,508,534]" pageId="3" pageNumber="68">Head:</emphasis>
With fairly protruding eyes, flat frontal furrows, more or less wrinkled and punctured towards neck.
</paragraph>
<paragraph id="8BED3694FFFB775D4AAEFD8D31B7FC6C" blockId="3.[151,1437,327,787]" pageId="3" pageNumber="68">
<emphasis id="B926EA86FFFB775D4AAEFD8D3428FC08" bold="true" box="[199,329,544,570]" pageId="3" pageNumber="68">Pronotum:</emphasis>
Wider than the head, longer than broad, apex and base straight, equally wide, sides convex, narrowed to the base, hind-quarter of the disc rectangular, basal fovea missing, median furrow deep.
</paragraph>
<paragraph id="8BED3694FFFB775D4AAEFDC534FFFC94" blockId="3.[151,1437,327,787]" pageId="3" pageNumber="68">
<emphasis id="B926EA86FFFB775D4AAEFDC5347CFCB0" bold="true" box="[199,285,616,642]" pageId="3" pageNumber="68">Elytra:</emphasis>
Humerus rounded; distinctly broadened towards the apex, strongly striated, intervals broad, flattened, apex straight truncated.
</paragraph>
<paragraph id="8BED3694FFFB775D4AAEFD023015FCF8" blockId="3.[151,1437,327,787]" box="[199,1396,687,715]" pageId="3" pageNumber="68">
<emphasis id="B926EA86FFFB775D4AAEFD023764FCF8" bold="true" box="[199,517,687,714]" pageId="3" pageNumber="68">Geographical distribution.</emphasis>
<collectingCountry id="F3457604FFFB775D4865FD1D3701FCF8" box="[524,608,688,714]" name="India" pageId="3" pageNumber="68">INDIA</collectingCountry>
:
<collectingRegion id="4996F876FFFB775D4801FD1D37D5FCF8" box="[616,692,688,714]" country="India" name="Assam" pageId="3" pageNumber="68">Assam</collectingRegion>
(
<bibRefCitation id="EFC34B65FFFB775D48AAFD1D3602FCF9" author="Jedlicka, A." box="[707,867,688,715]" pageId="3" pageNumber="68" pagination="1 - 6" refId="ref19878" refString="Jedlicka, A. (1964) Neue Carabiden aus der Palaarctischen region (Coleoptera). Annotationes Zoologicae et Botanicae, 3, 1 - 6." type="journal article" year="1964">Jedlička 1964</bibRefCitation>
);
<collectingRegion id="4996F876FFFB775D491EFD1D3699FCF8" box="[887,1016,688,714]" country="India" name="Meghalaya" pageId="3" pageNumber="68">Meghalaya</collectingRegion>
: Khasi hills (
<bibRefCitation id="EFC34B65FFFB775D4EE3FD1D300CFCF8" author="Kirschenhofer, E." box="[1162,1389,688,714]" pageId="3" pageNumber="68" pagination="25 - 64" refId="ref19906" refString="Kirschenhofer, E. (2010) New and Little known species of Carabidae from Middle East and Southeast Asia (Coleoptera: Carabidae: Lebiini, Brachinini). Annales Historico-Naturales Musei Nationalis Hungarici, 102, 25 - 64." type="journal article" year="2010">Kirschenhofer 2010</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C348651FFFFB775D4AAEFD793669FD20" pageId="3" pageNumber="68" type="discussion">
<paragraph id="8BED3694FFFB775D4AAEFD793669FD20" blockId="3.[151,1437,327,787]" pageId="3" pageNumber="68">
<emphasis id="B926EA86FFFB775D4AAEFD79345AFCDC" bold="true" box="[199,315,724,750]" pageId="3" pageNumber="68">Remarks.</emphasis>
Morphologically similar to
<taxonomicName id="4C524D17FFFB775D4810FD78366BFCDC" authorityName="Chaudoir" authorityYear="1876" box="[633,778,724,750]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="aptinoides">
<emphasis id="B926EA86FFFB775D4810FD78366BFCDC" box="[633,778,724,750]" italics="true" pageId="3" pageNumber="68">P. aptinoides</emphasis>
</taxonomicName>
, but differs with head, pronotum and elytra entirely black. It is endemic to the Indo-Burma hotspot of biodiversity.
</paragraph>
</subSubSection>
</treatment>
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