348 lines
37 KiB
XML
348 lines
37 KiB
XML
<document id="D7145DDC8067E2DAC311635675095AC4" ID-CLB-Dataset="21232" ID-DOI="10.11646/zootaxa.4608.1.3" ID-GBIF-Dataset="c4d067fa-4c56-4248-8084-caf4dd1eea21" ID-ISSN="1175-5326" ID-Zenodo-Dep="3993611" ID-ZooBank="3540218C-34FA-4D91-925A-3E89C6C3A7C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1597950731833" checkinUser="felipe" docAuthor="Venugopal, Akhil S. & Thomas, Sabu K." docDate="2019" docId="03FB8782FFEA774B4AFEF880304BFFE1" docLanguage="en" docName="zootaxa.4608.1.3.pdf" docOrigin="Zootaxa 4608 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pheropsophus devagiriensis Venugopal & Thomas 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="86" masterDocId="FFC2FFFAFFF8775E4A69FFAD3561FE32" masterDocTitle="Bombardier beetles of the genus Pheropsophus Solier 1833 (Carabidae: Brachininae: Brachinini) from Indian subcontinent" masterLastPageNumber="89" masterPageNumber="65" pageNumber="83" updateTime="1698845024281" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="4376F99E8D3DA85D721811CDA5B141CE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D9C3C698E203F93D57F07D7BA8EA81C4">
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<mods:title id="BEA762CB49D1BE331CD0DB4EBF3A018D">Bombardier beetles of the genus Pheropsophus Solier 1833 (Carabidae: Brachininae: Brachinini) from Indian subcontinent</mods:title>
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</mods:titleInfo>
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<mods:name id="A1409DDFA4F28E6970863E741739D5BF" type="personal">
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<mods:roleTerm id="6EC22BC94BA5F151045AFCBADCC3F565">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6C562C1502DEC0C0DFB11D80E1B8C08A">Venugopal, Akhil S.</mods:namePart>
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<mods:name id="94733B5C6A1AC9C45AF310D34C048EFC" type="personal">
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<mods:roleTerm id="FBF767503BBE3D720EBF7042D3DD8188">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9FE826FA3CF6809C7DC1B19C4839D796">Thomas, Sabu K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A42B5FF8A594ACBFC6622FE543DFCDEB">text</mods:typeOfResource>
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<mods:titleInfo id="4B1ABB4C61BF8053297C8EABFB2C6119">
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<mods:title id="54106D61723A5C487F65C8E952571A04">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B91D201B5153C070D5EB69BBCEEB0040">
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<mods:date id="3A7F0F2ACEE246F0A7258F780437A761">2019</mods:date>
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<mods:detail id="623DDD5F31856F0F5A723AF8B54EE001" type="pubDate">
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<mods:number id="6DA87AEF1245AEDA70778A5AF8CE7C9D">2019-05-17</mods:number>
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</mods:detail>
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<mods:detail id="52BC3EB43125DDBC7BE202E78230AFA6" type="volume">
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<mods:number id="53DFF34CA11C9B96B801B4F57ED40D81">4608</mods:number>
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</mods:detail>
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<mods:detail id="9BD72EF52375783C49B0BEFF4245215A" type="issue">
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<mods:number id="0AF66B87EB8887529A3477393274B00A">1</mods:number>
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<mods:extent id="B5C3F9FA4D4CBCEFC6FE4D5A01DA3870" unit="page">
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<mods:start id="94B61B420A41E787D47148694AC1FD09">65</mods:start>
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<mods:classification id="3FFAA2D049A170E7FA24C8BF5E34DC4C">journal article</mods:classification>
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<mods:identifier id="27C2612DC219B64A0C86584BF06CC14D" type="CLB-Dataset">21232</mods:identifier>
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<mods:identifier id="E38D3133BF796F8762CDD2F78B422542" type="DOI">10.11646/zootaxa.4608.1.3</mods:identifier>
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<mods:identifier id="317A11BFF9A6351AD12E44F2D90CD7DD" type="GBIF-Dataset">c4d067fa-4c56-4248-8084-caf4dd1eea21</mods:identifier>
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<mods:identifier id="BFB55CB448BCCC5DF60546BE3208FD2E" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="03FB8782FFEA774B4AFEF880304BFFE1" ID-DOI="http://doi.org/10.5281/zenodo.4335135" ID-GBIF-Taxon="167284790" ID-Zenodo-Dep="4335135" LSID="urn:lsid:plazi:treatment:03FB8782FFEA774B4AFEF880304BFFE1" httpUri="http://treatment.plazi.org/id/03FB8782FFEA774B4AFEF880304BFFE1" lastPageId="21" lastPageNumber="86" pageId="18" pageNumber="83">
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<subSubSection id="C348651FFFEA774C4AFEF880372FF975" box="[151,590,1837,1864]" pageId="18" pageNumber="83" type="nomenclature">
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<paragraph id="8BED3694FFEA774C4AFEF880372FF975" blockId="18.[151,590,1837,1899]" box="[151,590,1837,1864]" pageId="18" pageNumber="83">
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<heading id="D0A581F8FFEA774C4AFEF880372FF975" bold="true" box="[151,590,1837,1864]" fontSize="11" level="1" pageId="18" pageNumber="83" reason="1">
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||
<emphasis id="B926EA86FFEA774C4AFEF880372FF975" bold="true" box="[151,590,1837,1864]" pageId="18" pageNumber="83">
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||
<taxonomicName id="4C524D17FFEA774C4AFEF8803488F97A" ID-CoL="8MFW3" authority="Venugopal & Thomas, 2019" authorityName="Venugopal & Thomas" authorityYear="2019" box="[151,489,1837,1864]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="83" phylum="Arthropoda" rank="species" species="devagiriensis" status="sp. nov.">
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<emphasis id="B926EA86FFEA774C4AFEF8803488F97A" bold="true" box="[151,489,1837,1864]" italics="true" pageId="18" pageNumber="83">Pheropsophus devagiriensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21557FDFFEA774C4B98F880372FF975" box="[497,590,1837,1863]" pageId="18" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C348651FFFEA774D4AFEF8FC3465F984" lastPageId="19" lastPageNumber="84" pageId="18" pageNumber="83" type="description">
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<paragraph id="8BED3694FFEA774C4AFEF8FC345AF959" blockId="18.[151,590,1837,1899]" box="[151,315,1872,1899]" pageId="18" pageNumber="83">
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||
(
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<figureCitation id="13692A11FFEA774C4AC9F8FC359FF959" box="[160,254,1873,1899]" captionStart="FIGURE 4" captionStartId="19.[151,250,1696,1721]" captionTargetBox="[159,1426,1001,1656]" captionTargetId="figure@19.[153,1437,990,1664]" captionTargetPageId="19" captionText="FIGURE 4. Dorsal habitus of– A) Pheropsophus devagiriensis sp. nov. (Male, ‘India: Tamil Nadu: Courtallam’, Holotype), B) P. indicus sp. nov. (Male, ‘India: Kerala: Palakkad’, Holotype)" figureDoi="http://doi.org/10.5281/zenodo.3993619" httpUri="https://zenodo.org/record/3993619/files/figure.png" pageId="18" pageNumber="83">Fig. 4 A</figureCitation>
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,
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<figureCitation id="13692A11FFEA774C4B63F8FD3452F959" box="[266,307,1872,1899]" captionStart="FIGURE 5" captionStartId="20.[151,250,1882,1907]" captionTargetBox="[174,1420,189,1789]" captionTargetId="figure@20.[153,1438,176,1822]" captionTargetPageId="20" captionText="FIGURE 5. Pheropsophus male genitalia, Right lateral aspect–A) Pheropsophus devagiriensis sp. nov. (Holotype, ‘India: Tamilnadu: Courtallam’), B) P. indicus sp. nov. (Holotype, ‘India: Kerala: Palakkad’), C) P. bimaculatus (‘India: Kerala: Nilambur’), D) P. lissoderus (‘India: Kerala: Kalpetta’), E) P. occipitalis (‘India: Kerala: Kalpetta’), F) P. picicollis (‘India: Kerala: Chelari’), G) P. sobrinus (‘India: Kerala: Palakkad’), H) P. tripustulatus (‘India: Kerala: Nilambur’). Scale Bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3993621" httpUri="https://zenodo.org/record/3993621/files/figure.png" pageId="18" pageNumber="83">5 A</figureCitation>
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)
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</paragraph>
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<paragraph id="8BED3694FFEA774C4AFEF8353678F9C9" blockId="18.[151,1436,1944,2043]" pageId="18" pageNumber="83">
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<emphasis id="B926EA86FFEA774C4AFEF83534BDF981" bold="true" box="[151,476,1944,1971]" pageId="18" pageNumber="83">Specimens examined (n=6):</emphasis>
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<materialsCitation id="3B3A3CC9FFEA774C4B89F8343602F9E5" ID-GBIF-Occurrence="2846550359" collectingDate="2015-10-14" collectorName="S. V. Akhil" country="India" latitude="8.930855" location="Courtallam" longLatPrecision="1" longitude="77.268845" municipality="Male" pageId="18" pageNumber="83" specimenCount="1" stateProvince="Tamil Nadu" typeStatus="holotype">
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<typeStatus id="54E98836FFEA774C4B89F834372AF981" box="[480,587,1945,1971]" pageId="18" pageNumber="83" type="holotype">Holotype</typeStatus>
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, labelled “
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<collectingMunicipality id="6B89ACEEFFEA774C48D5F834379BF981" box="[700,762,1945,1971]" pageId="18" pageNumber="83">Male</collectingMunicipality>
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||
,
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<collectingCountry id="F3457604FFEA774C496DF8343623F981" box="[772,834,1945,1971]" name="India" pageId="18" pageNumber="83">India</collectingCountry>
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||
:
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<collectingRegion id="4996F876FFEA774C4920F83436B1F981" box="[841,976,1945,1971]" country="India" name="Tamil Nadu" pageId="18" pageNumber="83">Tamil Nadu</collectingRegion>
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:
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<location id="8E8D604FFFEA774C49B1F8343137F981" LSID="urn:lsid:plazi:treatment:03FB8782FFEA774B4AFEF880304BFFE1:8E8D604FFFEA774C49B1F8343137F981" box="[984,1110,1945,1971]" country="India" latitude="8.930855" longLatPrecision="1" longitude="77.268845" municipality="Male" name="Courtallam" pageId="18" pageNumber="83" stateProvince="Tamil Nadu">Courtallam</location>
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||
(
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||
<geoCoordinate id="EE665053FFEA774C4E0DF8353191F981" box="[1124,1264,1944,1971]" degrees="8.930855" direction="north" orientation="latitude" pageId="18" pageNumber="83" precision="1" value="8.930855">8.930855°N</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE665053FFEA774C4E90F83430EEF981" box="[1273,1423,1945,1971]" degrees="77.268843" direction="east" orientation="longitude" pageId="18" pageNumber="83" precision="1" value="77.268845">77.268843°E</geoCoordinate>
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), Hand Picking, from under stones,
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<date id="FFEC1054FFEA774C4877F81137F6F9E5" box="[542,663,1980,2007]" pageId="18" pageNumber="83" value="2015-10-14">
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||
<collectingDate id="EFA8E9BCFFEA774C4877F81137F6F9E5" box="[542,663,1980,2007]" pageId="18" pageNumber="83" value="2015-10-14">14.X.2015</collectingDate>
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</date>
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||
, coll.
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||
<collectorName id="26A75342FFEA774C48B7F8103638F9E5" box="[734,857,1981,2007]" pageId="18" pageNumber="83">S.V. Akhil</collectorName>
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||
”
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B3A3CC9FFEA774C4906F8103678F9C9" ID-GBIF-Occurrence="2846550371" collectingDate="2015-10-14" collectorName="S. V. Akhil." country="India" latitude="8.930855" location="Courtallam" longLatPrecision="1" longitude="77.268845" pageId="18" pageNumber="83" specimenCount="5" stateProvince="Tamil Nadu" typeStatus="paratype">
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||
<typeStatus id="54E98836FFEA774C4906F81036BEF9E5" box="[879,991,1981,2007]" pageId="18" pageNumber="83" type="paratype">Paratypes</typeStatus>
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||
,
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<specimenCount id="9D54FD1DFFEA774C4985F8113150F9E5" box="[1004,1073,1980,2007]" pageId="18" pageNumber="83" type="generic">5 exs.</specimenCount>
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||
,
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||
<collectingCountry id="F3457604FFEA774C4E57F810311DF9E5" box="[1086,1148,1981,2007]" name="India" pageId="18" pageNumber="83">India</collectingCountry>
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||
:
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||
<collectingRegion id="4996F876FFEA774C4EE1F8103073F9E5" box="[1160,1298,1981,2007]" country="India" name="Tamil Nadu" pageId="18" pageNumber="83">Tamil Nadu</collectingRegion>
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||
:
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<location id="8E8D604FFFEA774C4F77F81030FDF9E5" LSID="urn:lsid:plazi:treatment:03FB8782FFEA774B4AFEF880304BFFE1:8E8D604FFFEA774C4F77F81030FDF9E5" box="[1310,1436,1981,2007]" country="India" latitude="8.930855" longLatPrecision="1" longitude="77.268845" name="Courtallam" pageId="18" pageNumber="83" stateProvince="Tamil Nadu">Courtallam</location>
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||
(
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||
<geoCoordinate id="EE665053FFEA774C4AC8F84D344DF9C9" box="[161,300,2016,2043]" degrees="8.930855" direction="north" orientation="latitude" pageId="18" pageNumber="83" precision="1" value="8.930855">8.930855°N</geoCoordinate>
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,
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||
<geoCoordinate id="EE665053FFEA774C4B5EF84C34ACF9C9" box="[311,461,2017,2043]" degrees="77.268843" direction="east" orientation="longitude" pageId="18" pageNumber="83" precision="1" value="77.268845">77.268843°E</geoCoordinate>
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),
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||
<date id="FFEC1054FFEA774C4B8BF84D373BF9C9" box="[482,602,2016,2043]" pageId="18" pageNumber="83" value="2015-10-14">
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<collectingDate id="EFA8E9BCFFEA774C4B8BF84D373BF9C9" box="[482,602,2016,2043]" pageId="18" pageNumber="83" value="2015-10-14">14.X.2015</collectingDate>
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||
</date>
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, coll.
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||
<collectorName id="26A75342FFEA774C48F7F84C3678F9C9" box="[670,793,2017,2043]" pageId="18" pageNumber="83">S.V. Akhil.</collectorName>
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</materialsCitation>
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</paragraph>
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<paragraph id="8BED3694FFEB774D4AAEFF3537C4FE81" blockId="19.[151,1437,152,971]" box="[199,677,152,179]" pageId="19" pageNumber="84">
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<emphasis id="B926EA86FFEB774D4AAEFF353438FE81" bold="true" box="[199,345,152,179]" pageId="19" pageNumber="84">Description.</emphasis>
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TLA: 20.0 mm; TW:
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<quantity id="4CAA9B71FFEB774D4826FF3437C4FE81" box="[591,677,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="19" pageNumber="84" unit="mm" value="8.1">8.1 mm</quantity>
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</paragraph>
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<paragraph id="8BED3694FFEB774D4AA7FF113704FC0C" blockId="19.[151,1437,152,971]" pageId="19" pageNumber="84">
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<emphasis id="B926EA86FFEB774D4AA7FF11344AFEE5" bold="true" box="[206,299,188,215]" pageId="19" pageNumber="84">Colour:</emphasis>
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Head completely yellow without spots; clypeus (lateral edge brownish black) and labrum yellow with anterior brownish yellow border; fronto-clypeal suture light reddish brown; antennal segments 1–4 yellowish brown and 5–11 dark reddish brown; pronotum black with two large yellow spots on both sides of a narrow but prominent black median furrow, the two spots not reaching the lateral edge, lateral border being black; anterior and posterior margins of pronotum black; elytra with yellow patches including two small, yellow, rounded humeral spots, wide median transverse bands interrupted at suture and extreme apex; apical yellow band of elytra very narrow, just a small, fine yellow line; scutellum black; legs with coxa, trochanter, femur and tibia pale reddish yellow, tarsal end of tibia reddish brown, first tarsi with middle portion yellow and rest reddish brown, tarsi 2–5 reddish brown; claws dark brown; underside of head completely yellow; lateral side of the thorax with pale reddish yellow spot in the form of “T”;
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<emphasis id="B926EA86FFEB774D4B45FDAC34DCFC28" box="[300,445,513,538]" italics="true" pageId="19" pageNumber="84">metasternum</emphasis>
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reddish yellow posteriorly and reddish brown anteriorly;
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<emphasis id="B926EA86FFEB774D4E2AFDAC3189FC28" box="[1091,1256,513,538]" italics="true" pageId="19" pageNumber="84">metepisternum</emphasis>
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reddish yellow; rest of the ventral region brownish black.
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</paragraph>
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<paragraph id="8BED3694FFEB774D4AAEFDE4371DFCFC" blockId="19.[151,1437,152,971]" pageId="19" pageNumber="84">
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<emphasis id="B926EA86FFEB774D4AAEFDE43471FC51" bold="true" box="[199,272,585,611]" pageId="19" pageNumber="84">Head:</emphasis>
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<emphasis id="B926EA86FFEB774D4B70FDE4342FFC50" box="[281,334,585,610]" italics="true" pageId="19" pageNumber="84">Eyes</emphasis>
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prominent and protruding;
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<emphasis id="B926EA86FFEB774D48E2FDE537BDFC50" box="[651,732,584,610]" italics="true" pageId="19" pageNumber="84">labrum</emphasis>
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convex anteriorly;
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<emphasis id="B926EA86FFEB774D49D5FDE43161FC50" box="[956,1024,585,610]" italics="true" pageId="19" pageNumber="84">vertex</emphasis>
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with long supra orbital setae. Third antennal segment longer than segments 1 and 2 combined.
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<emphasis id="B926EA86FFEB774D494BFDC136F9FCB4" box="[802,920,620,646]" italics="true" pageId="19" pageNumber="84">Mandibles</emphasis>
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wide but short, protruding beyond the labrum; mandibular scrobe unisetose.
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<emphasis id="B926EA86FFEB774D4B8DFD3D3740FC98" box="[484,545,656,682]" italics="true" pageId="19" pageNumber="84">Palpi</emphasis>
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pubescent with terminal segment dilated.
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<emphasis id="B926EA86FFEB774D4992FD3D311FFC98" box="[1019,1150,656,682]" italics="true" pageId="19" pageNumber="84">Submentum</emphasis>
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setose with 4–5 long and 10–12 short setae.
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<emphasis id="B926EA86FFEB774D4B00FD1934CDFCFC" box="[361,428,692,718]" italics="true" pageId="19" pageNumber="84">Gular</emphasis>
|
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sutures diverging.
|
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</paragraph>
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<paragraph id="8BED3694FFEB774D4AAEFD74311CFD09" blockId="19.[151,1437,152,971]" pageId="19" pageNumber="84">
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<emphasis id="B926EA86FFEB774D4AAEFD743428FCC1" bold="true" box="[199,329,729,755]" pageId="19" pageNumber="84">Pronotum:</emphasis>
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Narrowed posteriorly; sides rounded in the anterior half and parallel behind; beaded dark lateral margin; anterior and posterior margin straight; disc finely punctate but glabrous except for very few brown setae along lateral edges; anterior and posterior margins each with a fringe of short dense setae
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</paragraph>
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<paragraph id="8BED3694FFEB774D4AAEFCE837A6FD95" blockId="19.[151,1437,152,971]" pageId="19" pageNumber="84">
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<emphasis id="B926EA86FFEB774D4AAEFCE8347CFD6D" bold="true" box="[199,285,837,863]" pageId="19" pageNumber="84">Elytra:</emphasis>
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convex on sides; with almost erased humerus; base narrower than apex, broadest just before apex. All nine striations (counting from the suture) well marked. Elytra very sparsely punctate with 1–3 setae on each intervals and a row of short yet sparse setae near apex.
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</paragraph>
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<paragraph id="8BED3694FFEB774D4AAEFC1D376DFDF8" blockId="19.[151,1437,152,971]" box="[199,524,944,971]" pageId="19" pageNumber="84">
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<emphasis id="B926EA86FFEB774D4AAEFC1D3424FDF9" bold="true" box="[199,325,944,971]" pageId="19" pageNumber="84">Abdomen:</emphasis>
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finely pubescent.
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</paragraph>
|
||
<caption id="DF2D661CFFEB774D4AFEF90D3675F8EF" ID-DOI="http://doi.org/10.5281/zenodo.3993619" ID-Zenodo-Dep="3993619" httpUri="https://zenodo.org/record/3993619/files/figure.png" pageId="19" pageNumber="84" startId="19.[151,250,1696,1721]" targetBox="[159,1426,1001,1656]" targetPageId="19">
|
||
<paragraph id="8BED3694FFEB774D4AFEF90D3675F8EF" blockId="19.[151,1436,1696,1757]" pageId="19" pageNumber="84">
|
||
<emphasis id="B926EA86FFEB774D4AFEF90D3472F88B" bold="true" box="[151,275,1696,1721]" pageId="19" pageNumber="84">FIGURE 4.</emphasis>
|
||
Dorsal habitus of– A)
|
||
<taxonomicName id="4C524D17FFEB774D4B90F90C3677F88B" authority="Venugopal & Thomas, 2019" authorityName="Venugopal & Thomas" authorityYear="2019" box="[505,790,1697,1721]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="84" phylum="Arthropoda" rank="species" species="devagiriensis" status="sp. nov.">
|
||
<emphasis id="B926EA86FFEB774D4B90F90C3677F88B" box="[505,790,1697,1721]" italics="true" pageId="19" pageNumber="84">Pheropsophus devagiriensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B926EA86FFEB774D4975F90C360DF88B" bold="true" box="[796,876,1697,1721]" pageId="19" pageNumber="84">
|
||
<taxonomicNameLabel id="A21557FDFFEB774D4975F90C360DF88B" box="[796,876,1697,1721]" pageId="19" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Male, ‘India: Tamil Nadu: Courtallam’, Holotype), B)
|
||
<taxonomicName id="4C524D17FFEB774D4AFEF96B3598F8EF" authority="Venugopal & Thomas, 2019" authorityName="Venugopal & Thomas" authorityYear="2019" box="[151,249,1733,1757]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="84" phylum="Arthropoda" rank="species" species="indicus" status="sp. nov.">
|
||
<emphasis id="B926EA86FFEB774D4AFEF96B3598F8EF" box="[151,249,1733,1757]" italics="true" pageId="19" pageNumber="84">P. indicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B926EA86FFEB774D4B69F9683431F8EF" bold="true" box="[256,336,1733,1757]" pageId="19" pageNumber="84">
|
||
<taxonomicNameLabel id="A21557FDFFEB774D4B69F9683431F8EF" box="[256,336,1733,1757]" pageId="19" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Male, ‘India: Kerala: Palakkad’, Holotype)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BED3694FFEB774D4AAEF8A13460F97B" blockId="19.[151,1437,1802,1974]" pageId="19" pageNumber="84">
|
||
<emphasis id="B926EA86FFEB774D4AAEF8A13419F917" bold="true" box="[199,376,1802,1830]" pageId="19" pageNumber="84">Male Genitalia</emphasis>
|
||
: Aedeagus with median lobe stout, low degree of curvature (only slightly curved); apex round- ed, blunt.
|
||
</paragraph>
|
||
<paragraph id="8BED3694FFEB774D4AAEF8F93465F984" blockId="19.[151,1437,1802,1974]" pageId="19" pageNumber="84">
|
||
<emphasis id="B926EA86FFEB774D4AAEF8F93417F95C" bold="true" box="[199,374,1876,1902]" pageId="19" pageNumber="84">Measurements</emphasis>
|
||
:
|
||
<emphasis id="B926EA86FFEB774D4BEFF8F93756F95C" bold="true" box="[390,567,1876,1902]" pageId="19" pageNumber="84">
|
||
<typeStatus id="54E98836FFEB774D4BEFF8F93492F95C" box="[390,499,1876,1902]" pageId="19" pageNumber="84" type="holotype">Holotype</typeStatus>
|
||
Male
|
||
</emphasis>
|
||
, TLA=
|
||
<quantity id="4CAA9B71FFEB774D48E5F8FE37BFF95C" box="[652,734,1875,1902]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="84" unit="mm" value="20.0">20 mm</quantity>
|
||
; TLB=
|
||
<quantity id="4CAA9B71FFEB774D495FF8FE36FCF95C" box="[822,925,1875,1902]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" pageId="19" pageNumber="84" unit="mm" value="17.7">17.7 mm</quantity>
|
||
; TW=
|
||
<quantity id="4CAA9B71FFEB774D4985F8FE3124F95C" box="[1004,1093,1875,1902]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="19" pageNumber="84" unit="mm" value="8.1">8.1 mm</quantity>
|
||
; PL =
|
||
<quantity id="4CAA9B71FFEB774D4EF9F8FE318BF95C" box="[1168,1258,1875,1902]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="19" pageNumber="84" unit="mm" value="3.6">3.6 mm</quantity>
|
||
; PW =
|
||
<quantity id="4CAA9B71FFEB774D4F57F8FE30F9F95C" box="[1342,1432,1875,1902]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="19" pageNumber="84" unit="mm" value="3.7">3.7 mm</quantity>
|
||
; EL =
|
||
<quantity id="4CAA9B71FFEB774D4ABFF8DA345DF9A0" box="[214,316,1911,1938]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="19" pageNumber="84" unit="mm" value="10.2">10.2 mm</quantity>
|
||
.
|
||
<emphasis id="B926EA86FFEB774D4B2FF8D53770F9A0" bold="true" box="[326,529,1912,1938]" pageId="19" pageNumber="84">
|
||
<typeStatus id="54E98836FFEB774D4B2FF8D534D2F9A0" box="[326,435,1912,1938]" pageId="19" pageNumber="84" type="paratype">Paratype</typeStatus>
|
||
Female
|
||
</emphasis>
|
||
, TLA=
|
||
<quantity id="4CAA9B71FFEB774D4802F8DA37DCF9A0" box="[619,701,1911,1938]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="19" pageNumber="84" unit="mm" value="21.0">21 mm</quantity>
|
||
; TLB =
|
||
<quantity id="4CAA9B71FFEB774D4970F8DA36E1F9A0" box="[793,896,1911,1938]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="19" pageNumber="84" unit="mm" value="16.8">16.8 mm</quantity>
|
||
; TW=
|
||
<quantity id="4CAA9B71FFEB774D49A4F8DA3144F9A0" box="[973,1061,1911,1938]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="19" pageNumber="84" unit="mm" value="7.2">7.2 mm</quantity>
|
||
; PL =
|
||
<quantity id="4CAA9B71FFEB774D4E04F8DA31A7F9A0" box="[1133,1222,1911,1938]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="19" pageNumber="84" unit="mm" value="3.6">3.6 mm</quantity>
|
||
; PW =
|
||
<quantity id="4CAA9B71FFEB774D4F7EF8DB3011F9A0" box="[1303,1392,1910,1938]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="19" pageNumber="84" unit="mm" value="3.5">3.5 mm</quantity>
|
||
; EL =
|
||
<quantity id="4CAA9B71FFEB774D4AC7F8363465F984" box="[174,260,1947,1974]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="19" pageNumber="84" unit="mm" value="9.6">9.6 mm</quantity>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF2D661CFFEC774A4AFEF8F73020F9EC" ID-DOI="http://doi.org/10.5281/zenodo.3993621" ID-Zenodo-Dep="3993621" httpUri="https://zenodo.org/record/3993621/files/figure.png" pageId="20" pageNumber="85" startId="20.[151,250,1882,1907]" targetBox="[174,1420,189,1789]" targetPageId="20">
|
||
<paragraph id="8BED3694FFEC774A4AFEF8F73020F9EC" blockId="20.[151,1437,1882,2015]" pageId="20" pageNumber="85">
|
||
<emphasis id="B926EA86FFEC774A4AFEF8F73477F941" bold="true" box="[151,278,1882,1907]" pageId="20" pageNumber="85">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C524D17FFEC774A4B41F8F734D9F940" authorityName="Solier" authorityYear="1833" box="[296,440,1882,1906]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="85" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B926EA86FFEC774A4B41F8F734D9F940" box="[296,440,1882,1906]" italics="true" pageId="20" pageNumber="85">Pheropsophus</emphasis>
|
||
</taxonomicName>
|
||
male genitalia, Right lateral aspect–A)
|
||
<taxonomicName id="4C524D17FFEC774A4935F8F7311DF940" authority="Venugopal & Thomas, 2019" authorityName="Venugopal & Thomas" authorityYear="2019" box="[860,1148,1882,1906]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="85" phylum="Arthropoda" rank="species" species="devagiriensis" status="sp. nov.">
|
||
<emphasis id="B926EA86FFEC774A4935F8F7311DF940" box="[860,1148,1882,1906]" italics="true" pageId="20" pageNumber="85">Pheropsophus devagiriensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B926EA86FFEC774A4EEFF8F631B8F941" bold="true" box="[1158,1241,1883,1907]" pageId="20" pageNumber="85">
|
||
<taxonomicNameLabel id="A21557FDFFEC774A4EEFF8F631B8F941" box="[1158,1241,1883,1907]" pageId="20" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Holotype, ‘India: Tamilnadu: Courtallam’), B)
|
||
<taxonomicName id="4C524D17FFEC774A4BABF8D23744F9A4" authority="Venugopal & Thomas, 2019" authorityName="Venugopal & Thomas" authorityYear="2019" box="[450,549,1918,1942]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="85" phylum="Arthropoda" rank="species" species="indicus" status="sp. nov.">
|
||
<emphasis id="B926EA86FFEC774A4BABF8D23744F9A4" box="[450,549,1918,1942]" italics="true" pageId="20" pageNumber="85">P. indicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B926EA86FFEC774A4845F8D2371FF9A5" bold="true" box="[556,638,1919,1943]" pageId="20" pageNumber="85">
|
||
<taxonomicNameLabel id="A21557FDFFEC774A4845F8D2371FF9A5" box="[556,638,1919,1943]" pageId="20" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Holotype, ‘India: Kerala: Palakkad’), C)
|
||
<taxonomicName id="4C524D17FFEC774A4E5BF8D231A9F9A4" baseAuthorityName="Linnaeus" baseAuthorityYear="1771" box="[1074,1224,1918,1942]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="85" phylum="Arthropoda" rank="species" species="bimaculatus">
|
||
<emphasis id="B926EA86FFEC774A4E5BF8D231A9F9A4" box="[1074,1224,1918,1942]" italics="true" pageId="20" pageNumber="85">P. bimaculatus</emphasis>
|
||
</taxonomicName>
|
||
(‘India: Kerala: Nilambur’), D)
|
||
<taxonomicName id="4C524D17FFEC774A4B7FF80E34F4F988" authorityName="Chaudoir" authorityYear="1850" box="[278,405,1954,1978]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="85" phylum="Arthropoda" rank="species" species="lissoderus">
|
||
<emphasis id="B926EA86FFEC774A4B7FF80E34F4F988" box="[278,405,1954,1978]" italics="true" pageId="20" pageNumber="85">P. lissoderus</emphasis>
|
||
</taxonomicName>
|
||
(‘India: Kerala: Kalpetta’), E)
|
||
<taxonomicName id="4C524D17FFEC774A48A4F80E362EF988" authorityName="Macleay" authorityYear="1825" box="[717,847,1954,1978]" class="Insecta" family="Carabidae" genus="Aptinus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="85" phylum="Arthropoda" rank="species" species="occipitalis">
|
||
<emphasis id="B926EA86FFEC774A48A4F80E362EF988" box="[717,847,1954,1978]" italics="true" pageId="20" pageNumber="85">P. occipitalis</emphasis>
|
||
</taxonomicName>
|
||
(‘India: Kerala: Kalpetta’), F)
|
||
<taxonomicName id="4C524D17FFEC774A4EECF80E319BF988" authorityName="Chaudoir" authorityYear="1876" box="[1157,1274,1954,1978]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="85" phylum="Arthropoda" rank="species" species="picicollis">
|
||
<emphasis id="B926EA86FFEC774A4EECF80E319BF988" box="[1157,1274,1954,1978]" italics="true" pageId="20" pageNumber="85">P. picicollis</emphasis>
|
||
</taxonomicName>
|
||
(‘India: Kerala: Chelari’), G)
|
||
<taxonomicName id="4C524D17FFEC774A4B49F86A34F1F9EC" baseAuthorityName="Dejean" baseAuthorityYear="1826" box="[288,400,1990,2014]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="85" phylum="Arthropoda" rank="species" species="sobrinus">
|
||
<emphasis id="B926EA86FFEC774A4B49F86A34F1F9EC" box="[288,400,1990,2014]" italics="true" pageId="20" pageNumber="85">P. sobrinus</emphasis>
|
||
</taxonomicName>
|
||
(‘India: Kerala: Palakkad’), H)
|
||
<taxonomicName id="4C524D17FFEC774A48BCF86A360FF9EC" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[725,878,1990,2014]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="85" phylum="Arthropoda" rank="species" species="tripustulatus">
|
||
<emphasis id="B926EA86FFEC774A48BCF86A360FF9EC" box="[725,878,1990,2014]" italics="true" pageId="20" pageNumber="85">P. tripustulatus</emphasis>
|
||
</taxonomicName>
|
||
(‘India: Kerala: Nilambur’). Scale Bar 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C348651FFFED774B4AAEFF343024FF71" pageId="21" pageNumber="86" type="materials_examined">
|
||
<paragraph id="8BED3694FFED774B4AAEFF343651FEC9" blockId="21.[151,1437,152,467]" pageId="21" pageNumber="86">
|
||
<emphasis id="B926EA86FFED774B4AAEFF34340CFE81" bold="true" box="[199,365,152,179]" pageId="21" pageNumber="86">Type material</emphasis>
|
||
:
|
||
<materialsCitation id="3B3A3CC9FFED774B4B15FF34365CFEE5" ID-GBIF-Occurrence="2846550339" collectingDate="2015-10-14" collectorName="S. V. Akhil" country="India" latitude="8.930855" location="Courtallam" longLatPrecision="1" longitude="77.268845" pageId="21" pageNumber="86" specimenCount="1" stateProvince="Tamil Nadu" typeStatus="holotype">
|
||
<typeStatus id="54E98836FFED774B4B15FF343485FE81" box="[380,484,153,179]" pageId="21" pageNumber="86" type="holotype">Holotype</typeStatus>
|
||
male:
|
||
<collectingCountry id="F3457604FFED774B4859FF34370FFE81" box="[560,622,153,179]" name="India" pageId="21" pageNumber="86">India</collectingCountry>
|
||
:
|
||
<collectingRegion id="4996F876FFED774B481EFF343661FE81" box="[631,768,153,179]" country="India" name="Tamil Nadu" pageId="21" pageNumber="86">Tamil Nadu</collectingRegion>
|
||
:
|
||
<location id="8E8D604FFFED774B4963FF3436E9FE81" LSID="urn:lsid:plazi:treatment:03FB8782FFEA774B4AFEF880304BFFE1:8E8D604FFFED774B4963FF3436E9FE81" box="[778,904,153,179]" country="India" latitude="8.930855" longLatPrecision="1" longitude="77.268845" name="Courtallam" pageId="21" pageNumber="86" stateProvince="Tamil Nadu">Courtallam</location>
|
||
(
|
||
<geoCoordinate id="EE665053FFED774B49FEFF353142FE81" box="[919,1059,152,179]" degrees="8.930855" direction="north" orientation="latitude" pageId="21" pageNumber="86" precision="1" value="8.930855">8.930855°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE665053FFED774B4E47FF3431A7FE86" box="[1070,1222,153,180]" degrees="77.268843" direction="east" orientation="longitude" pageId="21" pageNumber="86" precision="1" value="77.268845">77.268843°E</geoCoordinate>
|
||
),
|
||
<date id="FFEC1054FFED774B4EB0FF353030FE81" box="[1241,1361,152,179]" pageId="21" pageNumber="86" value="2015-10-14">
|
||
<collectingDate id="EFA8E9BCFFED774B4EB0FF353030FE81" box="[1241,1361,152,179]" pageId="21" pageNumber="86" value="2015-10-14">14.X.2015</collectingDate>
|
||
</date>
|
||
, handpicked from under stones, coll.
|
||
<collectorName id="26A75342FFED774B4B9AFF103709FEE5" box="[499,616,189,215]" pageId="21" pageNumber="86">S.V. Akhil</collectorName>
|
||
, deposited at ZSIC
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B3A3CC9FFED774B492FFF103651FEC9" ID-GBIF-Occurrence="2846550355" collectingDate="2015-10-14" country="India" latitude="8.930855" location="Courtallam" longLatPrecision="1" longitude="77.268845" pageId="21" pageNumber="86" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Tamil Nadu" typeStatus="paratype">
|
||
<typeStatus id="54E98836FFED774B492FFF1036D6FEE5" box="[838,951,189,215]" pageId="21" pageNumber="86" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D54FD1DFFED774B49A9FF10316CFEE5" box="[960,1037,189,215]" pageId="21" pageNumber="86" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D54FD1DFFED774B4E7EFF1031E4FEE5" box="[1047,1157,189,215]" pageId="21" pageNumber="86" type="female">4 females</specimenCount>
|
||
,
|
||
<collectingRegion id="4996F876FFED774B4EE7FF103075FEE5" box="[1166,1300,189,215]" country="India" name="Tamil Nadu" pageId="21" pageNumber="86">Tamil Nadu</collectingRegion>
|
||
,
|
||
<location id="8E8D604FFFED774B4F77FF1030FDFEE5" LSID="urn:lsid:plazi:treatment:03FB8782FFEA774B4AFEF880304BFFE1:8E8D604FFFED774B4F77FF1030FDFEE5" box="[1310,1436,189,215]" country="India" latitude="8.930855" longLatPrecision="1" longitude="77.268845" name="Courtallam" pageId="21" pageNumber="86" stateProvince="Tamil Nadu">Courtallam</location>
|
||
(
|
||
<geoCoordinate id="EE665053FFED774B4AF6FF4D344AFEC9" box="[159,299,224,251]" degrees="8.930855" direction="north" orientation="latitude" pageId="21" pageNumber="86" precision="1" value="8.930855">8.930855°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE665053FFED774B4B5EFF4C34AEFECE" box="[311,463,225,252]" degrees="77.268843" direction="east" orientation="longitude" pageId="21" pageNumber="86" precision="1" value="77.268845">77.268843°E</geoCoordinate>
|
||
),
|
||
<date id="FFEC1054FFED774B4B8BFF4D373BFEC9" box="[482,602,224,251]" pageId="21" pageNumber="86" value="2015-10-14">
|
||
<collectingDate id="EFA8E9BCFFED774B4B8BFF4D373BFEC9" box="[482,602,224,251]" pageId="21" pageNumber="86" value="2015-10-14">14.X.2015</collectingDate>
|
||
</date>
|
||
, from under stones
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BED3694FFED774B4AAEFEA9304AFF12" blockId="21.[151,1437,152,467]" box="[199,1323,260,288]" pageId="21" pageNumber="86">
|
||
<emphasis id="B926EA86FFED774B4AAEFEA93764FF2D" bold="true" box="[199,517,260,287]" pageId="21" pageNumber="86">Geographical distribution.</emphasis>
|
||
<collectingCountry id="F3457604FFED774B4865FEA8372BFF2D" box="[524,586,261,287]" name="India" pageId="21" pageNumber="86">India</collectingCountry>
|
||
:
|
||
<collectingRegion id="4996F876FFED774B483AFEA837BDFF2D" box="[595,732,261,287]" country="India" name="Tamil Nadu" pageId="21" pageNumber="86">Tamil Nadu</collectingRegion>
|
||
: Courtallam (Moist south Western Ghats eco-region)
|
||
</paragraph>
|
||
<paragraph id="8BED3694FFED774B4AAEFE853024FF71" blockId="21.[151,1437,152,467]" box="[199,1349,296,323]" pageId="21" pageNumber="86">
|
||
<emphasis id="B926EA86FFED774B4AAEFE853490FF71" bold="true" box="[199,497,296,323]" pageId="21" pageNumber="86">Collecting circumstances</emphasis>
|
||
: All the six specimens were handpicked from under stones near to a stream.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C348651FFFED774B4AAEFEE036A8FF55" box="[199,969,333,359]" pageId="21" pageNumber="86" type="etymology">
|
||
<paragraph id="8BED3694FFED774B4AAEFEE036A8FF55" blockId="21.[151,1437,152,467]" box="[199,969,333,359]" pageId="21" pageNumber="86">
|
||
<emphasis id="B926EA86FFED774B4AAEFEE03429FF55" bold="true" box="[199,328,333,359]" pageId="21" pageNumber="86">Etymology</emphasis>
|
||
: This species is named after the host research institution.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C348651FFFED774B4AAEFEDC304BFFE1" pageId="21" pageNumber="86" type="discussion">
|
||
<paragraph id="8BED3694FFED774B4AAEFEDC304BFFE1" blockId="21.[151,1437,152,467]" pageId="21" pageNumber="86">
|
||
<emphasis id="B926EA86FFED774B4AAEFEDC345AFFB9" bold="true" box="[199,315,369,395]" pageId="21" pageNumber="86">Remarks.</emphasis>
|
||
<taxonomicName id="4C524D17FFED774B4B2AFEDC3710FFB9" authority="Venugopal & Thomas, 2019" authorityName="Venugopal & Thomas" authorityYear="2019" box="[323,625,369,395]" class="Insecta" family="Carabidae" genus="Pherosophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="86" phylum="Arthropoda" rank="species" species="devagiriensis" status="sp. nov.">
|
||
<emphasis id="B926EA86FFED774B4B2AFEDC3710FFB9" box="[323,625,369,395]" italics="true" pageId="21" pageNumber="86">Pherosophus devagiriensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B926EA86FFED774B4813FEDC37B4FFB9" bold="true" box="[634,725,369,395]" pageId="21" pageNumber="86">
|
||
<taxonomicNameLabel id="A21557FDFFED774B4813FEDC37B4FFB9" box="[634,725,369,395]" pageId="21" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is very similar to
|
||
<taxonomicName id="4C524D17FFED774B49CCFEDF316AFFB9" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[933,1035,369,395]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="86" phylum="Arthropoda" rank="species" species="hilaris">
|
||
<emphasis id="B926EA86FFED774B49CCFEDF316AFFB9" box="[933,1035,369,395]" italics="true" pageId="21" pageNumber="86">P. hilaris</emphasis>
|
||
</taxonomicName>
|
||
but differs from it by head entirely yellow, pronotum with bright yellow spot not reaching the lateral margin, lateral margin black, pronontal median furrow deep, disc glabrous and shiny, elytral humeral spot very small, elytral yellow apical band narrow.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |