treatments-xml/data/03/D7/FF/03D7FF1CFF6DFF4BFF48F9473322FB86.xml
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<mods:title id="7AD1DC2EC50D592AA34A1F2BD6B48A8F">An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas</mods:title>
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<emphasis id="B90A9218FF6DFF4BFF48F94733A5FDB1" bold="true" box="[192,729,553,577]" pageId="166" pageNumber="165">
§
<taxonomicName id="4C7E3589FF6DFF4BFF5DF94733A5FDB1" authority="Arrow, 1911" authorityName="Arrow" authorityYear="1911" box="[213,729,553,577]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="166" pageNumber="165" phylum="Arthropoda" rank="species" species="carbonaria">
<emphasis id="B90A9218FF6DFF4BFF5DF9473352FDB1" bold="true" box="[213,558,553,577]" italics="true" pageId="166" pageNumber="165">Cyclocephala carbonaria</emphasis>
<bibRefCitation id="EFEF33FBFF6DFF4BFDBEF94733A5FDB1" author="Arrow, G. J." box="[566,729,553,577]" journalOrPublisher="Annals and Magazine of Natural History, including Zoology, Botany, and Geology 8" pageId="166" pageNumber="165" pagination="136 - 141" part="37" refId="ref191742" refString="Arrow, G. J. 1911 a. Notes on the lamellicorn beetles of the genus Golofa, with descriptions of three new species. Annals and Magazine of Natural History, including Zoology, Botany, and Geology 8 (7) (37): 136 - 141." title="Notes on the lamellicorn beetles of the genus Golofa, with descriptions of three new species" type="journal article" year="1911">Arrow, 1911</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<paragraph id="8BC14E0AFF6DFF4BFF60F93E32C4FD95" blockId="166.[232,952,592,614]" box="[232,952,592,614]" pageId="166" pageNumber="165">
<treatmentCitationGroup id="AB6E6924FF6DFF4BFF60F93E32C4FD95" box="[232,952,592,614]" pageId="166" pageNumber="165">
<treatmentCitation id="0ADF681BFF6DFF4BFF60F93E33BDFD95" author="Arrow, G. J." box="[232,705,592,614]" page="173" pageId="166" pageNumber="165" year="1911">
<taxonomicName id="4C7E3589FF6DFF4BFF60F93E33BDFD95" authority="Arrow 1911 b: 173" authorityName="Arrow" authorityPageNumber="173" authorityYear="1911" box="[232,705,592,614]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="166" pageNumber="165" phylum="Arthropoda" rank="species" species="carbonaria">
<emphasis id="B90A9218FF6DFF4BFF60F93E3085FD95" box="[232,505,592,613]" italics="true" pageId="166" pageNumber="165">Cyclocephala carbonaria</emphasis>
<bibRefCitation id="EFEF33FBFF6DFF4BFD88F93E33BDFD95" author="Arrow, G. J." box="[512,705,592,614]" journalOrPublisher="Annals and Magazine of Natural History, including Zoology, Botany, and Geology 8" pageId="166" pageNumber="165" pagination="151 - 176" part="43" refId="ref191794" refString="Arrow, G. J. 1911 b. Notes on the coleopterous subfamily Dynastinae, with descriptions of new genera and species. Annals and Magazine of Natural History, including Zoology, Botany, and Geology 8 (8) (43): 151 - 176." title="Notes on the coleopterous subfamily Dynastinae, with descriptions of new genera and species" type="journal article" year="1911">Arrow 1911b: 173</bibRefCitation>
</taxonomicName>
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(
<collectingCountry id="F3690E9AFF6DFF4BFD47F93E3234FD96" box="[719,840,592,614]" name="Nicaragua" pageId="166" pageNumber="165">Nicaragua</collectingCountry>
,
<collectingCountry id="F3690E9AFF6DFF4BFCDCF93E32CCFD95" box="[852,944,592,613]" name="Panama" pageId="166" pageNumber="165">Panama</collectingCountry>
)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3641D81FF6DFF4BFF48F9183322FB86" pageId="166" pageNumber="165" type="distribution">
<paragraph id="8BC14E0AFF6DFF4BFF48F9183356FCBE" blockId="166.[192,1440,630,846]" pageId="166" pageNumber="165">
<emphasis id="B90A9218FF6DFF4BFF48F9183331FD7E" bold="true" box="[192,589,630,654]" pageId="166" pageNumber="165">
Distribution.
<collectingCountry id="F3690E9AFF6DFF4BFEFDF918337DFD7E" box="[373,513,630,654]" name="Venezuela" pageId="166" pageNumber="165">Venezuela</collectingCountry>
(EO):
</emphasis>
<bibRefCitation id="EFEF33FBFF6DFF4BFDDAF9183380FD7E" author="Joly" box="[594,764,630,654]" firstAuthor="Joly" pageId="166" pageNumber="165" pagination="167 - 175" refId="ref209957" refString="Joly, L. J. 1995. Nuevos registros y redescripcion de dos especies negras de Cyclocephala (Coleoptera, Melolonthidae, Dynastinae) de Venezuela. Boletin de Entomologia Venezolana (N. S.) 10 (2): 167 - 175." type="journal article" year="1995">Joly 1995: 171</bibRefCitation>
(EO);
<bibRefCitation id="EFEF33FBFF6DFF4BFCCEF9193524FD7E" author="Moore" box="[838,1112,630,654]" etAl="et al." firstAuthor="Moore" pageId="166" pageNumber="165" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 156</bibRefCitation>
(EO).
<emphasis id="B90A9218FF6DFF4BFB2AF9183586FD7E" bold="true" box="[1186,1274,630,654]" pageId="166" pageNumber="165">Other:</emphasis>
<bibRefCitation id="EFEF33FBFF6DFF4BFB77F9193190FD5E" author="Arrow, G. J." journalOrPublisher="Annals and Magazine of Natural History, including Zoology, Botany, and Geology 8" pageId="166" pageNumber="165" pagination="151 - 176" part="43" refId="ref191794" refString="Arrow, G. J. 1911 b. Notes on the coleopterous subfamily Dynastinae, with descriptions of new genera and species. Annals and Magazine of Natural History, including Zoology, Botany, and Geology 8 (8) (43): 151 - 176." title="Notes on the coleopterous subfamily Dynastinae, with descriptions of new genera and species" type="journal article" year="1911">Arrow 1911b: 173</bibRefCitation>
(Nicaragua, Panama), 1937b: 9 (Ecuador, Nicaragua, Panama);
<bibRefCitation id="EFEF33FBFF6DFF4BFB81F9F8345DFD5E" author="Blackwelder, R. E." box="[1033,1313,662,686]" journalOrPublisher="United States National Museum Bulletin" pageId="166" pageNumber="165" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 251</bibRefCitation>
(Ecuador, Nicaragua, Panama);
<bibRefCitation id="EFEF33FBFF6DFF4BFE43F9D833C5FD3E" author="Endrodi, S." box="[459,697,694,718]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="166" pageNumber="165" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 164</bibRefCitation>
(Bolivia, Colombia, Costa Rica, Ecuador, Mexico, Nicaragua, Panama), 1985a: 40, 117 (Bolivia, Costa Rica, Ecuador, Mexico, Nicaragua, Panama);
<bibRefCitation id="EFEF33FBFF6DFF4BFB65F9B834E3FD1E" author="Joly" box="[1261,1439,726,750]" firstAuthor="Joly" pageId="166" pageNumber="165" pagination="167 - 175" refId="ref209957" refString="Joly, L. J. 1995. Nuevos registros y redescripcion de dos especies negras de Cyclocephala (Coleoptera, Melolonthidae, Dynastinae) de Venezuela. Boletin de Entomologia Venezolana (N. S.) 10 (2): 167 - 175." type="journal article" year="1995">Joly 1995: 171</bibRefCitation>
(Colombia, Costa Rica, Ecuador, Nicaragua, Panama);
<bibRefCitation id="EFEF33FBFF6DFF4BFCD8F998355DFCFE" author="Ratcliffe" box="[848,1057,758,782]" firstAuthor="Ratcliffe" pageId="166" pageNumber="165" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe 2003: 99</bibRefCitation>
(Costa Rica, Honduras, Mexico, Panama);
<bibRefCitation id="EFEF33FBFF6DFF4BFEB2F8793337FCDE" author="Moore" box="[314,587,790,814]" etAl="et al." firstAuthor="Moore" pageId="166" pageNumber="165" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 156</bibRefCitation>
(Belize, Bolivia, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama).
</paragraph>
<paragraph id="8BC14E0AFF6DFF4BFF48F8303322FB86" blockId="166.[192,1441,862,1142]" pageId="166" pageNumber="165">
<emphasis id="B90A9218FF6DFF4BFF48F8303074FC86" bold="true" box="[192,264,862,886]" pageId="166" pageNumber="165">Note.</emphasis>
<bibRefCitation id="EFEF33FBFF6DFF4BFE9AF830337FFC86" author="Dechambre, R. - P." box="[274,515,862,886]" journalOrPublisher="Coleopteres" pageId="166" pageNumber="165" pagination="13 - 27" part="3" refId="ref198483" refString="Dechambre, R. - P. 1997 a. Revision des Cyclocephala du groupe cribrata Burmeister (Coleoptera, Dynastidae). Coleopteres 3 (2): 13 - 27." title="Revision des Cyclocephala du groupe cribrata Burmeister (Coleoptera, Dynastidae)" type="journal article" year="1997">Dechambre (1997a)</bibRefCitation>
splits the species
<taxonomicName id="4C7E3589FF6DFF4BFD63F8303567FC86" authorityName="Arrow" authorityYear="1911" box="[747,1051,862,886]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="166" pageNumber="165" phylum="Arthropoda" rank="species" species="carbonaria">
<emphasis id="B90A9218FF6DFF4BFD63F8303567FC86" box="[747,1051,862,886]" italics="true" pageId="166" pageNumber="165">Cyclocephala carbonaria</emphasis>
</taxonomicName>
into a number of species, with
<taxonomicName id="4C7E3589FF6DFF4BFF48F8103014FC66" authorityName="Arrow" authorityYear="1911" box="[192,360,894,918]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="166" pageNumber="165" phylum="Arthropoda" rank="species" species="carbonaria">
<emphasis id="B90A9218FF6DFF4BFF48F8103014FC66" box="[192,360,894,918]" italics="true" pageId="166" pageNumber="165">C. carbonaria</emphasis>
</taxonomicName>
itself being restricted to Central America. According to this, South American specimens previously identified as
<taxonomicName id="4C7E3589FF6DFF4BFE6FF8F033F3FC46" authorityName="Arrow" authorityYear="1911" box="[487,655,926,950]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="166" pageNumber="165" phylum="Arthropoda" rank="species" species="carbonaria">
<emphasis id="B90A9218FF6DFF4BFE6FF8F033F3FC46" box="[487,655,926,950]" italics="true" pageId="166" pageNumber="165">C. carbonaria</emphasis>
</taxonomicName>
, such as those mentioned for
<collectingCountry id="F3690E9AFF6DFF4BFC77F8F03502FC46" box="[1023,1150,926,950]" name="Venezuela" pageId="166" pageNumber="165">Venezuela</collectingCountry>
in
<bibRefCitation id="EFEF33FBFF6DFF4BFB2FF8F03416FC46" author="Joly" box="[1191,1386,926,950]" firstAuthor="Joly" pageId="166" pageNumber="165" pagination="167 - 175" refId="ref209957" refString="Joly, L. J. 1995. Nuevos registros y redescripcion de dos especies negras de Cyclocephala (Coleoptera, Melolonthidae, Dynastinae) de Venezuela. Boletin de Entomologia Venezolana (N. S.) 10 (2): 167 - 175." type="journal article" year="1995">Joly (1995: 171)</bibRefCitation>
and
<bibRefCitation id="EFEF33FBFF6DFF4BFF48F8D13089FC26" author="Moore" box="[192,501,958,982]" etAl="et al." firstAuthor="Moore" pageId="166" pageNumber="165" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. (2018b: 156)</bibRefCitation>
(the latter who simply copied from older publications), must thus belong to other species.
<bibRefCitation id="EFEF33FBFF6DFF4BFEE3F8B0332CFC06" author="Ratcliffe" box="[363,592,990,1014]" firstAuthor="Ratcliffe" pageId="166" pageNumber="165" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe (2003: 99)</bibRefCitation>
suggests that more research needs to be done to establish if the South American specimens actually represent separate species or if they are merely intraspecific variations within
<taxonomicName id="4C7E3589FF6DFF4BFE92FF7030B8FBC6" authorityName="Arrow" authorityYear="1911" box="[282,452,1054,1078]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="166" pageNumber="165" phylum="Arthropoda" rank="species" species="carbonaria">
<emphasis id="B90A9218FF6DFF4BFE92FF7030B8FBC6" box="[282,452,1054,1078]" italics="true" pageId="166" pageNumber="165">C. carbonaria</emphasis>
</taxonomicName>
. The taxa which are described by
<bibRefCitation id="EFEF33FBFF6DFF4BFCFEFF70351AFBC6" author="Dechambre, R. - P." box="[886,1126,1054,1078]" journalOrPublisher="Coleopteres" pageId="166" pageNumber="165" pagination="13 - 27" part="3" refId="ref198483" refString="Dechambre, R. - P. 1997 a. Revision des Cyclocephala du groupe cribrata Burmeister (Coleoptera, Dynastidae). Coleopteres 3 (2): 13 - 27." title="Revision des Cyclocephala du groupe cribrata Burmeister (Coleoptera, Dynastidae)" type="journal article" year="1997">Dechambre (1997a)</bibRefCitation>
and which are known to occur in the research area are
<taxonomicName id="4C7E3589FF6DFF4BFDB6FF5033B7FBA6" authorityName="Dechambre" authorityYear="1997" box="[574,715,1086,1110]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="166" pageNumber="165" phylum="Arthropoda" rank="species" species="boucheri">
<emphasis id="B90A9218FF6DFF4BFDB6FF5033B7FBA6" box="[574,715,1086,1110]" italics="true" pageId="166" pageNumber="165">C. boucheri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7E3589FF6DFF4BFC83FF5032E6FBA6" authorityName="Dechambre" authorityYear="1997" box="[779,922,1086,1110]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="166" pageNumber="165" phylum="Arthropoda" rank="species" species="ergastuli">
<emphasis id="B90A9218FF6DFF4BFC83FF5032E6FBA6" box="[779,922,1086,1110]" italics="true" pageId="166" pageNumber="165">C. ergastuli</emphasis>
</taxonomicName>
. These are considered valid species by us and are included in this checklist.
</paragraph>
</subSubSection>
</treatment>
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