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<document id="61F0F358DD9A3C124FA82D853765BF07" ID-DOI="10.5281/zenodo.3678492" ID-GBIF-Dataset="8e18452b-1d9d-462e-bac8-8c86eeb4ddc3" ID-ISSN="1942-1354" ID-Zenodo-Dep="3678492" ID-ZooBank="930EAB11-37FA-41B0-980A-1A4736527842" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1582295086673" checkinUser="felipe" docAuthor="Hielkema, Auke J. &amp; Hielkema, Meindert A." docDate="2019" docId="03D7FF1CFF54FF72FF48F8AD331DF9C8" docLanguage="en" docName="InsectaMundi.0732.1-306.pdf.imf" docOrigin="Insecta Mundi 732 (732)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Ancognatha vulgaris Arrow 1911" docType="treatment" docVersion="13" lastPageNumber="158" masterDocId="FFEE8764FFCBFFEDFF88FB6E317CFFF0" masterDocTitle="An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas" masterLastPageNumber="306" masterPageNumber="1" pageNumber="158" updateTime="1698721431862" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="E2CDFBCAD7EAB2042A152037D210B097">An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas</mods:title>
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<mods:namePart id="059487B117987659AC93CD3039DB0C21">Hielkema, Auke J.</mods:namePart>
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<treatment id="03D7FF1CFF54FF72FF48F8AD331DF9C8" ID-DOI="http://doi.org/10.5281/zenodo.3682983" ID-GBIF-Taxon="162246789" ID-Zenodo-Dep="3682983" LSID="urn:lsid:plazi:treatment:03D7FF1CFF54FF72FF48F8AD331DF9C8" httpUri="http://treatment.plazi.org/id/03D7FF1CFF54FF72FF48F8AD331DF9C8" lastPageNumber="158" pageId="159" pageNumber="158">
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<heading id="D089F966FF54FF72FF48F8AD33E3FC2B" bold="true" box="[192,671,962,987]" fontSize="10" level="2" pageId="159" pageNumber="158" reason="3">
<emphasis id="B90A9218FF54FF72FF48F8AD33E3FC2B" bold="true" box="[192,671,962,987]" pageId="159" pageNumber="158">
§
<taxonomicName id="4C7E3589FF54FF72FF5DF8AC33E3FC2B" authority="Arrow, 1911" authorityName="Arrow" authorityYear="1911" box="[213,671,962,987]" class="Insecta" family="Dynastidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" pageId="159" pageNumber="158" phylum="Arthropoda" rank="species" species="vulgaris">
<emphasis id="B90A9218FF54FF72FF5DF8AC3089FC2A" bold="true" box="[213,501,962,986]" italics="true" pageId="159" pageNumber="158">Ancognatha vulgaris</emphasis>
<bibRefCitation id="EFEF33FBFF54FF72FE74F8AD33E3FC2B" author="Arrow, G. J." box="[508,671,963,987]" journalOrPublisher="Annals and Magazine of Natural History, including Zoology, Botany, and Geology 8" pageId="159" pageNumber="158" pagination="136 - 141" part="37" refId="ref191742" refString="Arrow, G. J. 1911 a. Notes on the lamellicorn beetles of the genus Golofa, with descriptions of three new species. Annals and Magazine of Natural History, including Zoology, Botany, and Geology 8 (7) (37): 136 - 141." title="Notes on the lamellicorn beetles of the genus Golofa, with descriptions of three new species" type="journal article" year="1911">Arrow, 1911</bibRefCitation>
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<paragraph id="8BC14E0AFF54FF72FF60F887322DFBF0" blockId="159.[232,849,1001,1024]" box="[232,849,1001,1024]" pageId="159" pageNumber="158">
<treatmentCitationGroup id="AB6E6924FF54FF72FF60F887322DFBF0" box="[232,849,1001,1024]" pageId="159" pageNumber="158">
<treatmentCitation id="0ADF681BFF54FF72FF60F887322DFBF0" author="Arrow, G. J." box="[232,849,1001,1024]" page="169" pageId="159" pageNumber="158" year="1911">
<taxonomicName id="4C7E3589FF54FF72FF60F887322DFBF0" authority="Arrow 1911 b: 169 (Panama-Peru)" authorityName="Arrow" authorityYear="1911" box="[232,849,1001,1024]" class="Insecta" family="Dynastidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" pageId="159" pageNumber="158" phylum="Arthropoda" rank="species" species="vulgaris">
<emphasis id="B90A9218FF54FF72FF60F88730ADFC0E" box="[232,465,1001,1022]" italics="true" pageId="159" pageNumber="158">Ancognatha vulgaris</emphasis>
<bibRefCitation id="EFEF33FBFF54FF72FE5FF88433E4FC0F" author="Arrow, G. J." box="[471,664,1001,1023]" journalOrPublisher="Annals and Magazine of Natural History, including Zoology, Botany, and Geology 8" pageId="159" pageNumber="158" pagination="151 - 176" part="43" refId="ref191794" refString="Arrow, G. J. 1911 b. Notes on the coleopterous subfamily Dynastinae, with descriptions of new genera and species. Annals and Magazine of Natural History, including Zoology, Botany, and Geology 8 (8) (43): 151 - 176." title="Notes on the coleopterous subfamily Dynastinae, with descriptions of new genera and species" type="journal article" year="1911">Arrow 1911b: 169</bibRefCitation>
(Panama–Peru)
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="C3641D81FF54FF72FF48FF61331DF9C8" pageId="159" pageNumber="158" type="distribution">
<paragraph id="8BC14E0AFF54FF72FF48FF61357EFB18" blockId="159.[192,1440,1039,1256]" pageId="159" pageNumber="158">
<emphasis id="B90A9218FF54FF72FF48FF613333FBD7" bold="true" box="[192,591,1039,1064]" pageId="159" pageNumber="158">
Distribution.
<collectingCountry id="F3690E9AFF54FF72FEF1FF7E3335FBD7" box="[377,585,1039,1064]" name="French Guiana" pageId="159" pageNumber="158">French Guiana</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF54FF72FDDCFF7E3244FBD8" author="Duranton, M." box="[596,824,1040,1064]" journalOrPublisher="Coleopteres, numero special" pageId="159" pageNumber="158" pagination="1 - 31" part="1" refId="ref200092" refString="Duranton, M. 2011. Chorologie et ecologie des dynastides de Guyane francaise. Coleopteres, numero special 1: 1 - 31." title="Chorologie et ecologie des dynastides de Guyane francaise" type="journal article" year="2011">Duranton 2011: 17</bibRefCitation>
(?).
<emphasis id="B90A9218FF54FF72FCE1FF6132B9FBD7" bold="true" box="[873,965,1039,1063]" pageId="159" pageNumber="158">
<collectingCountry id="F3690E9AFF54FF72FCE1FF6132C3FBD7" box="[873,959,1039,1063]" name="Brazil" pageId="159" pageNumber="158">Brazil</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF54FF72FC43FF7E35C6FBD8" author="Endrodi, S." bookContentInfo="800 p." box="[971,1210,1040,1064]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="159" pageNumber="158" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">Endrödi 1985a: 161</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF54FF72FB4DFF7E34E7FBD8" author="Ratcliffe" box="[1221,1435,1040,1064]" firstAuthor="Ratcliffe" pageId="159" pageNumber="158" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe 2003: 45</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF54FF72FF48FF5E30A7FBB8" author="Moore" box="[192,475,1072,1096]" etAl="et al." firstAuthor="Moore" pageId="159" pageNumber="158" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 119</bibRefCitation>
.
<emphasis id="B90A9218FF54FF72FE6DFF5E3307FBB8" bold="true" box="[485,635,1072,1096]" pageId="159" pageNumber="158">
<collectingCountry id="F3690E9AFF54FF72FE6DFF5E3309FBB8" box="[485,629,1072,1096]" name="Venezuela" pageId="159" pageNumber="158">Venezuela</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF54FF72FD08FF5E3213FBB8" author="Endrodi, S." box="[640,879,1072,1096]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="159" pageNumber="158" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 382</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF54FF72FCF2FF5E35E9FBB8" author="Moore" box="[890,1173,1072,1096]" etAl="et al." firstAuthor="Moore" pageId="159" pageNumber="158" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 119</bibRefCitation>
.
<emphasis id="B90A9218FF54FF72FB16FF41358BFBB7" bold="true" box="[1182,1271,1071,1095]" pageId="159" pageNumber="158">Other:</emphasis>
<bibRefCitation id="EFEF33FBFF54FF72FB74FF5E3190FB98" author="Arrow, G. J." journalOrPublisher="Annals and Magazine of Natural History, including Zoology, Botany, and Geology 8" pageId="159" pageNumber="158" pagination="151 - 176" part="43" refId="ref191794" refString="Arrow, G. J. 1911 b. Notes on the coleopterous subfamily Dynastinae, with descriptions of new genera and species. Annals and Magazine of Natural History, including Zoology, Botany, and Geology 8 (8) (43): 151 - 176." title="Notes on the coleopterous subfamily Dynastinae, with descriptions of new genera and species" type="journal article" year="1911">Arrow 1911b: 169</bibRefCitation>
(Panama–Peru), 1937b: 6 (Colombia, Panama, Peru);
<bibRefCitation id="EFEF33FBFF54FF72FC00FF3E35E3FB98" author="Blackwelder, R. E." box="[904,1183,1104,1128]" journalOrPublisher="United States National Museum Bulletin" pageId="159" pageNumber="158" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 250</bibRefCitation>
(Colombia, Panama, Peru);
<bibRefCitation id="EFEF33FBFF54FF72FE99FF1E3081FB78" author="Endrodi, S." box="[273,509,1136,1160]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="159" pageNumber="158" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 382</bibRefCitation>
(Brazil
<collectingRegion id="49BA80E8FF54FF72FDD5FF1E33A0FB78" box="[605,732,1136,1160]" country="Venezuela" name="Amazonas" pageId="159" pageNumber="158">Amazonas</collectingRegion>
(Nauta) [&gt; Peru!], Bolivia, Colombia, Costa Rica, Ecuador, Panama, Peru), 1985a: 161 (Bolivia, Colombia, Costa Rica, Ecuador, Panama, Peru);
<bibRefCitation id="EFEF33FBFF54FF72FB43FFFE34DCFB58" author="Ratcliffe" box="[1227,1440,1168,1192]" firstAuthor="Ratcliffe" pageId="159" pageNumber="158" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe 2003: 45</bibRefCitation>
(Brazil/Bolivia–Costa Rica);
<bibRefCitation id="EFEF33FBFF54FF72FD91FFDE33C7FB38" author="Ponchel" box="[537,699,1200,1224]" firstAuthor="Ponchel" pageId="159" pageNumber="158" pagination="60 - 61" refId="ref220051" refString="Ponchel, Y. 2011. Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 4: 60 - 61. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." type="journal article" year="2011">Ponchel 2011</bibRefCitation>
(not mentioned);
<bibRefCitation id="EFEF33FBFF54FF72FC19FFDE3546FB38" author="Dupuis, F." box="[913,1082,1200,1224]" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="159" pageNumber="158" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis 2016b</bibRefCitation>
(not mentioned);
<bibRefCitation id="EFEF33FBFF54FF72FA98FFDE3038FB18" author="Moore" etAl="et al." firstAuthor="Moore" pageId="159" pageNumber="158" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 119</bibRefCitation>
(Bolivia, Colombia, Costa Rica, Ecuador, Panama, Peru).
</paragraph>
<paragraph id="8BC14E0AFF54FF72FF48FF9634DCFA7F" blockId="159.[192,1440,1272,1424]" pageId="159" pageNumber="158">
<emphasis id="B90A9218FF54FF72FF48FF963061FAE0" bold="true" box="[192,285,1272,1296]" pageId="159" pageNumber="158">Note 1.</emphasis>
<bibRefCitation id="EFEF33FBFF54FF72FEAAFF96337AFAE0" author="Arrow, G. J." box="[290,518,1272,1296]" journalOrPublisher="Annals and Magazine of Natural History, including Zoology, Botany, and Geology 8" pageId="159" pageNumber="158" pagination="151 - 176" part="43" refId="ref191794" refString="Arrow, G. J. 1911 b. Notes on the coleopterous subfamily Dynastinae, with descriptions of new genera and species. Annals and Magazine of Natural History, including Zoology, Botany, and Geology 8 (8) (43): 151 - 176." title="Notes on the coleopterous subfamily Dynastinae, with descriptions of new genera and species" type="journal article" year="1911">Arrow (1911b: 169)</bibRefCitation>
states in his description of
<taxonomicName id="4C7E3589FF54FF72FCC2FF963539FAE0" authorityName="Arrow" authorityYear="1911" box="[842,1093,1272,1296]" class="Insecta" family="Dynastidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" pageId="159" pageNumber="158" phylum="Arthropoda" rank="species" species="vulgaris">
<emphasis id="B90A9218FF54FF72FCC2FF963539FAE0" box="[842,1093,1272,1296]" italics="true" pageId="159" pageNumber="158">Ancognatha vulgaris</emphasis>
</taxonomicName>
that “it ranges from
<collectingCountry id="F3690E9AFF54FF72FAB2FF9634E3FAFF" box="[1338,1439,1272,1295]" name="Panama" pageId="159" pageNumber="158">Panama</collectingCountry>
to
<collectingCountry id="F3690E9AFF54FF72FF56FE763065FADF" box="[222,281,1304,1327]" name="Peru" pageId="159" pageNumber="158">Peru</collectingCountry>
and the Upper Amazons (Nauta).” Based on this statement,
<bibRefCitation id="EFEF33FBFF54FF72FC70FE76358BFAC0" author="Endrodi, S." box="[1016,1271,1304,1328]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="159" pageNumber="158" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi (1966a: 382)</bibRefCitation>
subsequently places Nauta in the Brazilian state of Amazonas. In fact, Nauta is a town situated on the Upper Amazon in the Peruvian region of
<collectingRegion id="49BA80E8FF54FF72FDA6FE3633FCFA9F" box="[558,640,1368,1391]" country="Peru" name="Loreto" pageId="159" pageNumber="158">Loreto</collectingRegion>
, well away from the border with
<collectingCountry id="F3690E9AFF54FF72FB90FE36351BFA80" box="[1048,1127,1368,1392]" name="Brazil" pageId="159" pageNumber="158">Brazil</collectingCountry>
. The records for
<collectingCountry id="F3690E9AFF54FF72FABCFE363403FA80" box="[1332,1407,1368,1392]" name="Brazil" pageId="159" pageNumber="158">Brazil</collectingCountry>
in
<bibRefCitation id="EFEF33FBFF54FF72FF48FE1630BDFA60" author="Endrodi, S." bookContentInfo="800 p." box="[192,449,1400,1424]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="159" pageNumber="158" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">Endrödi (1985a: 161)</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF54FF72FE45FE1633CEFA60" author="Ratcliffe" box="[461,690,1400,1424]" firstAuthor="Ratcliffe" pageId="159" pageNumber="158" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe (2003: 45)</bibRefCitation>
and
<bibRefCitation id="EFEF33FBFF54FF72FD65FE163564FA60" author="Moore" box="[749,1048,1400,1424]" etAl="et al." firstAuthor="Moore" pageId="159" pageNumber="158" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. (2018b: 119)</bibRefCitation>
may well be based on this error.
</paragraph>
<paragraph id="8BC14E0AFF54FF72FF48FECE331DF9C8" blockId="159.[192,1440,1440,1592]" pageId="159" pageNumber="158">
<emphasis id="B90A9218FF54FF72FF48FECE305CFA48" bold="true" box="[192,288,1440,1464]" pageId="159" pageNumber="158">Note 2.</emphasis>
<bibRefCitation id="EFEF33FBFF54FF72FEA0FECE335DFA48" author="Duranton, M." box="[296,545,1440,1464]" journalOrPublisher="Coleopteres, numero special" pageId="159" pageNumber="158" pagination="1 - 31" part="1" refId="ref200092" refString="Duranton, M. 2011. Chorologie et ecologie des dynastides de Guyane francaise. Coleopteres, numero special 1: 1 - 31." title="Chorologie et ecologie des dynastides de Guyane francaise" type="journal article" year="2011">Duranton (2011: 17)</bibRefCitation>
notes he saw no definite specimens of
<taxonomicName id="4C7E3589FF54FF72FB88FECE347CFA48" authorityName="Arrow" authorityYear="1911" box="[1024,1280,1440,1464]" class="Insecta" family="Dynastidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" pageId="159" pageNumber="158" phylum="Arthropoda" rank="species" species="vulgaris">
<emphasis id="B90A9218FF54FF72FB88FECE347CFA48" box="[1024,1280,1440,1464]" italics="true" pageId="159" pageNumber="158">Ancognatha vulgaris</emphasis>
</taxonomicName>
from
<collectingCountry id="F3690E9AFF54FF72FAC1FECE3062FA28" name="French Guiana" pageId="159" pageNumber="158">French Guiana</collectingCountry>
, and the species is not mentioned in the checklists for
<collectingCountry id="F3690E9AFF54FF72FC41FEAE35F8FA28" box="[969,1156,1472,1496]" name="French Guiana" pageId="159" pageNumber="158">French Guiana</collectingCountry>
by
<bibRefCitation id="EFEF33FBFF54FF72FB3BFEAE3415FA28" author="Ponchel" box="[1203,1385,1472,1496]" firstAuthor="Ponchel" pageId="159" pageNumber="158" pagination="60 - 61" refId="ref220051" refString="Ponchel, Y. 2011. Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 4: 60 - 61. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." type="journal article" year="2011">Ponchel (2011)</bibRefCitation>
and
<bibRefCitation id="EFEF33FBFF54FF72FF48FE8E3002FA08" author="Dupuis, F." box="[192,382,1504,1528]" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="159" pageNumber="158" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis (2016b)</bibRefCitation>
. We have not seen any specimens from
<collectingCountry id="F3690E9AFF54FF72FCD3FE8E32A8FA08" box="[859,980,1504,1528]" name="Suriname" pageId="159" pageNumber="158">Suriname</collectingCountry>
and are unaware of any other records from the research area. This species appears to be restricted to northwestern South America and the southern part of Central America.
</paragraph>
</subSubSection>
</treatment>
</document>
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