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<document id="956021DA6EABA397A0E9560ED7ADF009" ID-CLB-Dataset="32334" ID-DOI="10.11646/zootaxa.4311.2.1" ID-GBIF-Dataset="fe39a11b-43d1-44ed-b047-19f2096eb6c0" ID-ISSN="1175-5326" ID-Zenodo-Dep="847417" ID-ZooBank="4E905C58-3E70-489B-8B14-190152523810" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1503490079451" checkinUser="plazi" docAuthor="Castelló, José" docDate="2017" docId="03D7E344FFE8382AFF3DF963FB5EB1DA" docLanguage="en" docName="zootaxa.4311.2.1.pdf" docOrigin="Zootaxa 4311 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atarbolana beirutensis Castelló 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="163" masterDocId="FFEE9B3CFFE03826FFAAFC72FFB1B41A" masterDocTitle="New and little-known species of isopods (Crustacea, Isopoda) from the eastern Mediterranean" masterLastPageNumber="182" masterPageNumber="151" pageNumber="159" updateTime="1698467061358" updateUser="ExternalLinkService">
<mods:mods id="4EDF7DDA6037219CCE94A88985E7F7D8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="65A3E09319166F19AC4EC17A69735A97">
<mods:title id="DD2BC3201458F8C7E4F0A293A83B58D7">New and little-known species of isopods (Crustacea, Isopoda) from the eastern Mediterranean</mods:title>
</mods:titleInfo>
<mods:name id="C49ADB109E5C6FA429D5428FEF3B30D6" type="personal">
<mods:role id="1B42BA98F5A17F2EF881DBC7C9733AEE">
<mods:roleTerm id="5D115F9B755A9F5D392A53DBCAA8D6D5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="78F24CCB645B97BE2981824C0E7B7A99">Castelló, José</mods:namePart>
</mods:name>
<mods:typeOfResource id="47AC276D10FCDA475F312F3E3ECF99A4">text</mods:typeOfResource>
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<mods:title id="4D1F482835F0A84E8B91A598DA478BDD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="41E7BA0DE1ADBB882D2416025B7D0295">
<mods:date id="32A133D23727ECFB679A71BDE0BB814C">2017</mods:date>
<mods:detail id="386462565DDEFBDF4A18578078FAA999" type="volume">
<mods:number id="0D0C6F4235F3E6F207A3ADDD146B94C1">4311</mods:number>
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<mods:detail id="FBC7355795C498C0866982F4CACEA0CB" type="issue">
<mods:number id="536CCE87ECA8D74E9A347107F04351E3">2</mods:number>
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<mods:classification id="8D84F5DF4284B912194082B1BFCE4434">journal article</mods:classification>
<mods:identifier id="864BD80C3ADD7FD554EC0F80A724FB62" type="CLB-Dataset">32334</mods:identifier>
<mods:identifier id="01F94801D5F5D489AF2C93701B14C865" type="DOI">10.11646/zootaxa.4311.2.1</mods:identifier>
<mods:identifier id="0F6257C5277E9708F2A0B62FA592E074" type="GBIF-Dataset">fe39a11b-43d1-44ed-b047-19f2096eb6c0</mods:identifier>
<mods:identifier id="EF47072811B7A49B09C3E132AAA4A9D7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="06A4604882CDDFBC6CF720E5BBA4D483" type="Zenodo-Dep">847417</mods:identifier>
<mods:identifier id="936D3286A93C0A7153AFF59814064689" type="ZooBank">4E905C58-3E70-489B-8B14-190152523810</mods:identifier>
</mods:mods>
<treatment id="03D7E344FFE8382AFF3DF963FB5EB1DA" ID-DOI="http://doi.org/10.5281/zenodo.6024220" ID-GBIF-Taxon="132797456" ID-Zenodo-Dep="6024220" LSID="urn:lsid:plazi:treatment:03D7E344FFE8382AFF3DF963FB5EB1DA" httpUri="http://treatment.plazi.org/id/03D7E344FFE8382AFF3DF963FB5EB1DA" lastPageId="12" lastPageNumber="163" pageId="8" pageNumber="159">
<subSubSection id="C36401D9FFE8382EFF3DF963FEBBB157" pageId="8" pageNumber="159" type="nomenclature">
<paragraph id="8BC15252FFE8382EFF3DF963FDBEB131" blockId="8.[151,527,1297,1357]" box="[151,527,1297,1323]" pageId="8" pageNumber="159">
<heading id="D089E53EFFE8382EFF3DF963FDBEB131" bold="true" box="[151,527,1297,1323]" fontSize="11" level="1" pageId="8" pageNumber="159" reason="1">
<emphasis id="B90A8E40FFE8382EFF3DF963FDBEB131" bold="true" box="[151,527,1297,1323]" pageId="8" pageNumber="159">
<taxonomicName id="4C7E29D1FFE8382EFF3DF963FE1BB131" ID-CoL="J8FG" authority="Castelló, 2017" authorityName="Castelló" authorityYear="2017" box="[151,426,1297,1323]" class="Malacostraca" family="Cirolanidae" genus="Atarbolana" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="beirutensis" status="sp. nov.">
<emphasis id="B90A8E40FFE8382EFF3DF963FE1BB131" bold="true" box="[151,426,1297,1323]" italics="true" pageId="8" pageNumber="159">Atarbolana beirutensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A239333BFFE8382EFE18F963FDBEB131" box="[434,527,1297,1323]" pageId="8" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC15252FFE8382EFF3DF946FEBBB157" blockId="8.[151,527,1297,1357]" box="[151,266,1332,1357]" pageId="8" pageNumber="159">
(
<figureCitation id="13454ED7FFE8382EFF35F946FEB2B157" box="[159,259,1332,1357]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="9.[151,250,1942,1964]" captionStartId-1="10.[151,250,1974,1996]" captionStartId-2="11.[151,250,1967,1989]" captionTargetBox-0="[207,1392,190,1928]" captionTargetBox-1="[194,1409,215,1946]" captionTargetBox-2="[179,1410,214,1927]" captionTargetId-0="figure@9.[185,1411,185,1933]" captionTargetId-1="figure@10.[164,1432,193,1965]" captionTargetId-2="figure@11.[151,1436,193,1946]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="FIGURE 6. Atarbolana beirutensis sp. nov. Holotype female [MZB 2016 - 3479]: A, body, dorsal View; B, frontal lamina; C, distal end of body, dorsolateral View; D, apex of pleotelson; E, uropod, dorsal View; F, uropod, lateral View; G, antenna 1; H, antenna 2. Scale bars: A = 1 mm; B, F – H = 0.5 mm; C = 1.5 mm; D – E = 0.25 mm." captionText-1="FIGURE 7. Atarbolana beirutensis sp. nov. Holotype female [MZB 2016 - 3479]: A, mandible; B, maxilla 1; C, maxilla 2; D, maxilliped; E, pereopod 1; F, pereopod 3; G, pereopod 5; H, pereopod 7. Scale bars: A – D = 0.1 mm; E – H = 0.25 mm." captionText-2="FIGURE 8. Atarbolana beirutensis sp. nov. Holotype female [MZB 2016 - 3479]: A – E, pleopods 1 – 5. Scale bar: A – E = 0.25 mm." httpUri-0="https://zenodo.org/record/847429/files/figure.png" httpUri-1="https://zenodo.org/record/847431/files/figure.png" httpUri-2="https://zenodo.org/record/847433/files/figure.png" pageId="8" pageNumber="159">Figs 6–8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36401D9FFE8382EFF3DF90EFCC0B23E" pageId="8" pageNumber="159" type="materials_examined">
<paragraph id="8BC15252FFE8382EFF3DF90EFA32B1A2" blockId="8.[151,1437,1403,2005]" pageId="8" pageNumber="159">
<materialsCitation id="3B16580FFFE8382EFF3DF90EFA32B1A2" ID-GBIF-Occurrence="1586888116" collectingDate="2002-09-17" collectionCode="MZB" collectorName="Bitar &amp; H. Zibrowius" country="Lebanon" elevation="5" latitude="33.888336" location="Material" longLatPrecision="19" longitude="35.466946" municipality="Raoucheh" pageId="8" pageNumber="159" specimenCode="MZB2016-3479, 20" specimenCount="1" stateProvince="Beirut" typeStatus="holotype">
<emphasis id="B90A8E40FFE8382EFF3DF90EFE4BB18F" bold="true" box="[151,506,1403,1429]" pageId="8" pageNumber="159">
<location id="8EA10489FFE8382EFF3DF90EFF4EB18F" LSID="urn:lsid:plazi:treatment:03D7E344FFE8382AFF3DF963FB5EB1DA:8EA10489FFE8382EFF3DF90EFF4EB18F" box="[151,255,1404,1429]" country="Lebanon" latitude="33.888336" longLatPrecision="19" longitude="35.466946" municipality="Raoucheh" name="Material" pageId="8" pageNumber="159" stateProvince="Beirut">Material</location>
examined.
<typeStatus id="54C5ECF0FFE8382EFE23F909FE40B18E" box="[393,497,1403,1428]" pageId="8" pageNumber="159" type="holotype">
<emphasis id="B90A8E40FFE8382EFE23F909FE40B18E" bold="true" box="[393,497,1403,1428]" italics="true" pageId="8" pageNumber="159">Holotype</emphasis>
</typeStatus>
:
</emphasis>
ovigerous female,
<quantity id="4C86FFB7FFE8382EFD7EF90EFC9CB18E" box="[724,813,1404,1429]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="159" unit="mm" value="4.5">4.5 mm</quantity>
, sample 5M15, station 11,
<collectingMunicipality id="6BA5C828FFE8382EFBF5F90EFB63B18F" box="[1119,1234,1404,1429]" pageId="8" pageNumber="159">Raoucheh</collectingMunicipality>
,
<collectingRegion id="49BA9CB0FFE8382EFB74F90EFA96B18F" box="[1246,1319,1404,1429]" country="Lebanon" name="Beyrouth" pageId="8" pageNumber="159">Beirut</collectingRegion>
,
<collectingCountry id="F36912C2FFE8382EFA99F90EFA29B18F" box="[1331,1432,1404,1429]" name="Lebanon" pageId="8" pageNumber="159">Lebanon</collectingCountry>
,
<geoCoordinate id="EE4A3495FFE8382EFF3DF9EDFEADB1A2" box="[151,284,1439,1464]" direction="north" orientation="latitude" pageId="8" pageNumber="159" precision="15" value="33.888336">33°53'18”N</geoCoordinate>
,
<geoCoordinate id="EE4A3495FFE8382EFE8DF9EDFE18B1A2" box="[295,425,1439,1464]" direction="east" orientation="longitude" pageId="8" pageNumber="159" precision="15" value="35.466946">35°28'01”E</geoCoordinate>
,
<elevation id="0053B561FFE8382EFE1FF9D2FE4CB1AD" box="[437,509,1440,1464]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="8" pageNumber="159" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">
<quantity id="4C86FFB7FFE8382EFE1FF9D2FE4CB1AD" box="[437,509,1440,1464]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="8" pageNumber="159" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">0–1 m</quantity>
</elevation>
, cave, G.
<collectorName id="268B3784FFE8382EFDC3F9EDFD10B1A2" box="[617,673,1439,1464]" pageId="8" pageNumber="159">Bitar</collectorName>
&amp;
<collectorName id="268B3784FFE8382EFD6FF9D2FCEAB1A2" box="[709,859,1439,1464]" pageId="8" pageNumber="159">H. Zibrowius</collectorName>
coll.,
<collectingDate id="EF848D7AFFE8382EFC0BF9D2FB8AB1A2" box="[929,1083,1440,1464]" pageId="8" pageNumber="159" value="2002-09-17">
<date id="FFC07492FFE8382EFC0BF9D2FB8AB1A2" box="[929,1083,1440,1464]" pageId="8" pageNumber="159" value="2002-09-17">17 Sept. 2002</date>
</collectingDate>
(
<specimenCode id="DBD8FA29FFE8382EFBE0F9EDFA9DB1A2" box="[1098,1324,1439,1464]" pageId="8" pageNumber="159">MZB2016-3479, 20</specimenCode>
slides).
</materialsCitation>
</paragraph>
<paragraph id="8BC15252FFE8382EFF6DF9B6FCC0B23E" blockId="8.[151,1437,1403,2005]" pageId="8" pageNumber="159">
<materialsCitation id="3B16580FFFE8382EFF6DF9B6FCDCB23E" ID-GBIF-Occurrence="1586888127" collectingDate="2002-09-17" collectionCode="MZB" collectorName="Bitar &amp; H. Zibrowius" country="Lebanon" elevation="3" latitude="33.888336" location="Material" longLatPrecision="19" longitude="35.466946" municipality="Raoucheh" pageId="8" pageNumber="159" specimenCode="MZB2016-3480, 15, MZB2016-3481" specimenCount="2" stateProvince="Beirut" typeStatus="paratype">
<emphasis id="B90A8E40FFE8382EFF6DF9B6FEF0B1C7" bold="true" box="[199,321,1476,1501]" pageId="8" pageNumber="159">
<typeStatus id="54C5ECF0FFE8382EFF6DF9B6FE88B1C1" box="[199,313,1476,1499]" pageId="8" pageNumber="159" type="paratype">
<emphasis id="B90A8E40FFE8382EFF6DF9B6FE88B1C1" bold="true" box="[199,313,1476,1499]" italics="true" pageId="8" pageNumber="159">Paratypes</emphasis>
</typeStatus>
:
</emphasis>
damaged female without posterior section of body, sample 5M15, same locality as holotype (
<specimenCode id="DBD8FA29FFE8382EFF34F995FE32B21A" box="[158,387,1511,1536]" pageId="8" pageNumber="159">MZB2016-3480, 15</specimenCode>
slides); juvenile,
<quantity id="4C86FFB7FFE8382EFDF9F99AFD28B1E5" box="[595,665,1512,1536]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="159" unit="mm" value="2.0">2 mm</quantity>
, sample 5M16, same locality as
<typeStatus id="54C5ECF0FFE8382EFBB2F995FBCBB21A" box="[1048,1146,1511,1536]" pageId="8" pageNumber="159" type="holotype">holotype</typeStatus>
(
<specimenCode id="DBD8FA29FFE8382EFB21F995FAF9B21A" box="[1163,1352,1511,1536]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="8" pageNumber="159">MZB2016-3481</specimenCode>
); depth between 0 and
<elevation id="0053B561FFE8382EFE95FA7EFEDCB23E" box="[319,365,1548,1572]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="159" unit="m" value="3.0">
<quantity id="4C86FFB7FFE8382EFE95FA7EFEDCB23E" box="[319,365,1548,1572]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="159" unit="m" value="3.0">3 m</quantity>
</elevation>
, G.
<collectorName id="268B3784FFE8382EFE3DFA7EFE7EB23F" box="[407,463,1548,1573]" pageId="8" pageNumber="159">Bitar</collectorName>
&amp;
<collectorName id="268B3784FFE8382EFE59FA7FFD39B23F" box="[499,648,1548,1573]" pageId="8" pageNumber="159">H. Zibrowius</collectorName>
coll.,
<collectingDate id="EF848D7AFFE8382EFD64FA7EFCDCB23E" box="[718,877,1548,1572]" pageId="8" pageNumber="159" value="2002-09-17">
<date id="FFC07492FFE8382EFD64FA7EFCDCB23E" box="[718,877,1548,1572]" pageId="8" pageNumber="159" value="2002-09-17">17 Sept. 2002</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36401D9FFE8382EFF6DFA5DFEE0B2AF" pageId="8" pageNumber="159" type="diagnosis">
<paragraph id="8BC15252FFE8382EFF6DFA5DFEE0B2AF" blockId="8.[151,1437,1403,2005]" pageId="8" pageNumber="159">
<emphasis id="B90A8E40FFE8382EFF6DFA5DFEF0B252" bold="true" box="[199,321,1583,1608]" pageId="8" pageNumber="159">Diagnosis.</emphasis>
<emphasis id="B90A8E40FFE8382EFEE3FA5DFE33B252" box="[329,386,1583,1608]" italics="true" pageId="8" pageNumber="159">Body</emphasis>
2.4 times longer than wide. Posterior margin of cephalon with groove on each side. Frontal lamina pentagonal, elongated, with triangulate anterior and concave lateral margins. Pereonites 4–7 with transverse grooves. Apex of pleotelson truncated, with 10 robust setae. Uropodal exopod like an elongated spoon, with concave section.
</paragraph>
</subSubSection>
<subSubSection id="C36401D9FFE8382AFF6DFACDFCFBB0BA" lastPageId="12" lastPageNumber="163" pageId="8" pageNumber="159" type="description">
<paragraph id="8BC15252FFE8382EFF6DFACDFA2CB3CE" blockId="8.[151,1437,1403,2005]" pageId="8" pageNumber="159">
<emphasis id="B90A8E40FFE8382EFF6DFACDFD0DB2C2" bold="true" box="[199,700,1727,1752]" pageId="8" pageNumber="159">
Description of
<typeStatus id="54C5ECF0FFE8382EFED0FACDFE50B2C2" box="[378,481,1727,1752]" pageId="8" pageNumber="159" type="holotype">holotype</typeStatus>
[MZB2016-3479].
</emphasis>
<emphasis id="B90A8E40FFE8382EFD6FFACDFD4EB2C2" box="[709,767,1727,1752]" italics="true" pageId="8" pageNumber="159">Body</emphasis>
2.4 times longer than wide. Cephalon with apex of rostral process overlapped by frontal lamina; posterior margin of cephalon with groove on each side. Frontal lamina pentagonal, with triangulate anterior and concave lateral margins. Pereonite 1 slightly longer than remaining pereonites, equal in length; pereonite 1 with lateral simple suture; coxae 2–3 with longitudinal suture almost parallel to the margin; coxae 3–7 with oblique suture. Few setae on margins of pereonites 3–7; surface of body smooth, without setae. Transverse dorsal groove on pereonites 4–7. Pleonites 2–5 visible from above, last three almost equal in length, without denticles. Pleotelson narrowed, concave in distal half, and truncated distally; apex with 10 robust setae and 4 long simple setae, two of them in the middle; lateral margins of pleotelson without setae.
</paragraph>
<caption id="DF0102DAFFE9382FFF3DFBE4FC51B3F0" httpUri="https://zenodo.org/record/847429/files/figure.png" pageId="9" pageNumber="160" targetBox="[207,1392,190,1928]" targetPageId="9">
<paragraph id="8BC15252FFE9382FFF3DFBE4FC51B3F0" blockId="9.[151,1435,1942,2026]" pageId="9" pageNumber="160">
<emphasis id="B90A8E40FFE9382FFF3DFBE4FEA5B3B6" bold="true" box="[151,276,1942,1964]" pageId="9" pageNumber="160">FIGURE 6.</emphasis>
<taxonomicName id="4C7E29D1FFE9382FFEB6FBE4FDB7B3B6" authority="Castelló, 2017" authorityName="Castelló" authorityYear="2017" box="[284,518,1942,1964]" class="Malacostraca" family="Cirolanidae" genus="Atarbolana" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="beirutensis" status="sp. nov.">
<emphasis id="B90A8E40FFE9382FFEB6FBE4FDB7B3B6" box="[284,518,1942,1964]" italics="true" pageId="9" pageNumber="160">Atarbolana beirutensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A239333BFFE9382FFDA4FBE5FDD0B3B6" box="[526,609,1943,1964]" pageId="9" pageNumber="160" rank="species">
<emphasis id="B90A8E40FFE9382FFDA4FBE5FDD0B3B6" bold="true" box="[526,609,1943,1964]" pageId="9" pageNumber="160">sp. nov.</emphasis>
</taxonomicNameLabel>
Holotype female [MZB2016-3479]: A, body, dorsal View; B, frontal lamina; C, distal end of body, dorsolateral View; D, apex of pleotelson; E, uropod, dorsal View; F, uropod, lateral View; G, antenna 1; H, antenna 2. Scale bars: A = 1 mm; B, F–H = 0.5 mm; C = 1.5 mm; D–E = 0.25 mm.
</paragraph>
</caption>
<caption id="DF0102DAFFEA382CFF3DFBC4FA8EB3F1" httpUri="https://zenodo.org/record/847431/files/figure.png" pageId="10" pageNumber="161" targetBox="[194,1409,215,1946]" targetPageId="10">
<paragraph id="8BC15252FFEA382CFF3DFBC4FA8EB3F1" blockId="10.[151,1435,1974,2027]" pageId="10" pageNumber="161">
<emphasis id="B90A8E40FFEA382CFF3DFBC4FEA2B3D6" bold="true" box="[151,275,1974,1996]" pageId="10" pageNumber="161">FIGURE 7.</emphasis>
<taxonomicName id="4C7E29D1FFEA382CFEB1FBC4FDB4B3D6" authority="Castelló, 2017" authorityName="Castelló" authorityYear="2017" box="[283,517,1974,1996]" class="Malacostraca" family="Cirolanidae" genus="Atarbolana" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="beirutensis" status="sp. nov.">
<emphasis id="B90A8E40FFEA382CFEB1FBC4FDB4B3D6" box="[283,517,1974,1996]" italics="true" pageId="10" pageNumber="161">Atarbolana beirutensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A239333BFFEA382CFDA6FBC5FDEFB3D6" box="[524,606,1975,1996]" pageId="10" pageNumber="161" rank="species">
<emphasis id="B90A8E40FFEA382CFDA6FBC5FDEFB3D6" bold="true" box="[524,606,1975,1996]" pageId="10" pageNumber="161">sp. nov.</emphasis>
</taxonomicNameLabel>
Holotype female [MZB2016-3479]: A, mandible; B, maxilla 1; C, maxilla 2; D, maxilliped; E, pereopod 1; F, pereopod 3; G, pereopod 5; H, pereopod 7. Scale bars: A–D = 0.1 mm; E–H = 0.25 mm.
</paragraph>
</caption>
<caption id="DF0102DAFFEB382DFF3DFBDDFF77B3FE" httpUri="https://zenodo.org/record/847433/files/figure.png" pageId="11" pageNumber="162" targetBox="[179,1410,214,1927]" targetPageId="11">
<paragraph id="8BC15252FFEB382DFF3DFBDDFF77B3FE" blockId="11.[151,1436,1967,2020]" pageId="11" pageNumber="162">
<emphasis id="B90A8E40FFEB382DFF3DFBDDFEA5B3DE" bold="true" box="[151,276,1967,1989]" pageId="11" pageNumber="162">FIGURE 8.</emphasis>
<taxonomicName id="4C7E29D1FFEB382DFEB6FBDDFDB7B3DF" authority="Castelló, 2017" authorityName="Castelló" authorityYear="2017" box="[284,518,1967,1989]" class="Malacostraca" family="Cirolanidae" genus="Atarbolana" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="beirutensis" status="sp. nov.">
<emphasis id="B90A8E40FFEB382DFEB6FBDDFDB7B3DF" box="[284,518,1967,1989]" italics="true" pageId="11" pageNumber="162">Atarbolana beirutensis</emphasis>
</taxonomicName>
<emphasis id="B90A8E40FFEB382DFDA4FBC2FDD0B3DF" bold="true" box="[526,609,1968,1989]" pageId="11" pageNumber="162">
<taxonomicNameLabel id="A239333BFFEB382DFDA4FBC2FDD0B3DF" box="[526,609,1968,1989]" pageId="11" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype female [MZB2016-3479]: A–E, pleopods 1–5. Scale bar: A–E = 0.25 mm.
</paragraph>
</caption>
<paragraph id="8BC15252FFEC382AFF6DFCEAFD95B507" blockId="12.[151,1437,151,1473]" pageId="12" pageNumber="163">
<emphasis id="B90A8E40FFEC382AFF6DFCEAFE8BB4AA" box="[199,314,152,176]" italics="true" pageId="12" pageNumber="163">Antenna 1</emphasis>
reaching pereonite 3, with 4 peduncular articles; article 3 longest and article 4 shortest; articulation of articles 3–4 oblique; flagellum of 13 articles (
<quantity id="4C86FFB7FFEC382AFD1AFCCEFD6AB4CF" box="[688,731,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="12" pageNumber="163" unit="in" value="9.0">9 in</quantity>
damaged female); articles 3–13 bearing aesthetascs.
<emphasis id="B90A8E40FFEC382AFA80FCCEFA2DB4CE" box="[1322,1436,188,212]" italics="true" pageId="12" pageNumber="163">Antenna 2</emphasis>
with peduncular article 4 longest; articles 1–4 each with group of long setae on anterodistal angle; flagellum with 16 articles (
<quantity id="4C86FFB7FFEC382AFEB0FD76FEE5B507" box="[282,340,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="12" pageNumber="163" unit="in" value="15.0">15 in</quantity>
damaged female).
</paragraph>
<paragraph id="8BC15252FFEC382AFF6DFD55FE64B602" blockId="12.[151,1437,151,1473]" pageId="12" pageNumber="163">
<emphasis id="B90A8E40FFEC382AFF6DFD55FEFCB55A" box="[199,333,295,320]" italics="true" pageId="12" pageNumber="163">Mandibular</emphasis>
palp articles 2–3 with 12 pectinated setae each (9 and 11 respectively in the damaged female); setae of article 2 arranged in two rows.
<emphasis id="B90A8E40FFEC382AFDF5FD3EFD7CB57E" box="[607,717,332,357]" italics="true" pageId="12" pageNumber="163">Maxilla 1</emphasis>
with 3 plumose robust setae on endopod and 12 curved robust setae and one slender seta on exopod.
<emphasis id="B90A8E40FFEC382AFDE3FD1DFD07B592" box="[585,694,367,392]" italics="true" pageId="12" pageNumber="163">Maxilla 2</emphasis>
inner lobe with 7, medial lobe with 4, and outer lobe with 5 long plumose setae. Maxillipedal palp article 1 with 3 long plumose setae placed distally; articles 2–5 with 4, 12, 9, and 8 setae on medial margin respectively, usually plumose (simple setae if placed some way from the margin); lateral margin with 3, 7, 1, and 2 long simple setae respectively; endite with 3 lateral and 2 subterminal stout plumose setae, and 2 coupling hooks.
</paragraph>
<paragraph id="8BC15252FFEC382AFF6DFE56FC4EB72D" blockId="12.[151,1437,151,1473]" pageId="12" pageNumber="163">
<emphasis id="B90A8E40FFEC382AFF6DFE56FEF6B626" box="[199,327,548,573]" italics="true" pageId="12" pageNumber="163">Pereopod 1</emphasis>
basis with 2 plumose setae on posterodistal angle; ischium with long seta on anterodistal angle and 3 long subterminal setae; merus with 3 long setae on anterodistal angle and subterminal seta, inner margin with 2 robust and 1 simple seta; carpus with 1 robust and 1 simple seta on inner margin, and 2 subterminal setae; propodus with 6 robust bifid setae interspersed with simple setae arranged in groups of 2–3; dactylus with distinct secondary unguis; row of 9 setae at base of unguis.
<emphasis id="B90A8E40FFEC382AFDCFFEC6FCBCB6D6" box="[613,781,692,717]" italics="true" pageId="12" pageNumber="163">Pereopods 2–3</emphasis>
shorter and more setose than pereopod 1; inner margin of merus-carpus-propodus with stout setae, especially in pereopod 2.
<emphasis id="B90A8E40FFEC382AFC37FEA5FB93B6EA" box="[925,1058,727,752]" italics="true" pageId="12" pageNumber="163">Pereopod 4</emphasis>
robust, with stout setae on both anterior and posterior margins.
<emphasis id="B90A8E40FFEC382AFE50FE8EFD12B70E" box="[506,675,764,789]" italics="true" pageId="12" pageNumber="163">Pereopods 5–7</emphasis>
similar, increasing in length posteriorly; numerous simple plumose setae and stout bifid setae on ischium-merus-carpus, especially in pereopod 7.
</paragraph>
<paragraph id="8BC15252FFEC382AFF6DFF36FE55B7F7" blockId="12.[151,1437,151,1473]" pageId="12" pageNumber="163">
<emphasis id="B90A8E40FFEC382AFF6DFF36FE8BB746" box="[199,314,836,861]" italics="true" pageId="12" pageNumber="163">Pleopod 1</emphasis>
endopod narrow; peduncle with 4 coupling hooks and plumose seta on inner margin; endopod and exopod with 13 and 26 plumose setae respectively.
<emphasis id="B90A8E40FFEC382AFD4BFF15FCE4B79A" box="[737,853,871,896]" italics="true" pageId="12" pageNumber="163">Pleopod 2</emphasis>
peduncle with 3 coupling hooks and plumose seta; endopod and exopod with 13 and 33 plumose setae.
<emphasis id="B90A8E40FFEC382AFC86FFFEFC7FB7BE" box="[812,974,908,933]" italics="true" pageId="12" pageNumber="163">Pleopods 2–5</emphasis>
peduncle increasingly short and rami progressively rounded. Pleopods 3–5 without setae on margin of endopod, and exopod with incomplete suture; endopod shorter than exopod.
</paragraph>
<paragraph id="8BC15252FFEC382AFF6DFF85FC51B067" blockId="12.[151,1437,151,1473]" pageId="12" pageNumber="163">
<emphasis id="B90A8E40FFEC382AFF6DFF85FE82B00A" box="[199,307,1015,1040]" italics="true" pageId="12" pageNumber="163">Uropodal</emphasis>
endopod rounded, flat, shorter than telsonic apex; 11 robust setae interspersed with stout setae on the margin of distal half. Uropodal exopod thin and concave surpassing telsonic apex; 5 robust setae on inner margin, apex with robust seta and a tuft of thinner setae, outer margin without robust setae. Tegument of exopod with simple setae and rounded scales with groups of short setae on their margin.
</paragraph>
<paragraph id="8BC15252FFEC382AFF6DF8F5FCFBB0BA" blockId="12.[151,1437,151,1473]" box="[199,842,1159,1184]" pageId="12" pageNumber="163">
<emphasis id="B90A8E40FFEC382AFF6DF8F5FEB8B0BA" box="[199,265,1159,1184]" italics="true" pageId="12" pageNumber="163">Color</emphasis>
in alcohol pale with dorsal brown chromatophores.
</paragraph>
</subSubSection>
<subSubSection id="C36401D9FFEC382AFF6DF8DEFCFAB0DF" box="[199,843,1196,1221]" pageId="12" pageNumber="163" type="etymology">
<paragraph id="8BC15252FFEC382AFF6DF8DEFCFAB0DF" blockId="12.[151,1437,151,1473]" box="[199,843,1196,1221]" pageId="12" pageNumber="163">
<emphasis id="B90A8E40FFEC382AFF6DF8DEFEFFB0DF" bold="true" box="[199,334,1196,1221]" pageId="12" pageNumber="163">Etymology.</emphasis>
Beirutensis, from
<collectingRegion id="49BA9CB0FFEC382AFDB7F8DEFDD7B0DF" box="[541,614,1196,1221]" country="Lebanon" name="Beyrouth" pageId="12" pageNumber="163">Beirut</collectingRegion>
, capital of
<collectingCountry id="F36912C2FFEC382AFD4BF8DEFCF6B0DF" box="[737,839,1196,1221]" name="Lebanon" pageId="12" pageNumber="163">Lebanon</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36401D9FFEC382AFF6DF8BDFB5EB1DA" pageId="12" pageNumber="163" type="discussion">
<paragraph id="8BC15252FFEC382AFF6DF8BDFB35B12A" blockId="12.[151,1437,151,1473]" pageId="12" pageNumber="163">
<emphasis id="B90A8E40FFEC382AFF6DF8BDFE8AB0FD" bold="true" box="[199,315,1231,1256]" pageId="12" pageNumber="163">Remarks.</emphasis>
<taxonomicName id="4C7E29D1FFEC382AFEEEF8BDFDF8B0F2" authority="Castelló, 2017" authorityName="Castelló" authorityYear="2017" box="[324,585,1231,1256]" class="Malacostraca" family="Cirolanidae" genus="Atarbolana" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="beirutensis" status="sp. nov.">
<emphasis id="B90A8E40FFEC382AFEEEF8BDFDF8B0F2" box="[324,585,1231,1256]" italics="true" pageId="12" pageNumber="163">Atarbolana beirutensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A239333BFFEC382AFDF8F8BDFD1FB0F2" box="[594,686,1231,1256]" pageId="12" pageNumber="163" rank="species">
<emphasis id="B90A8E40FFEC382AFDF8F8BDFD1FB0F2" bold="true" box="[594,686,1231,1256]" pageId="12" pageNumber="163">sp. nov.</emphasis>
</taxonomicNameLabel>
has a uropodal exopod with a concave section like an elongated spoon; this feature, a spoon-like uropodal exopod without the cylindrical section found in the other species, is an autapomorphic character, and may be added to the generic diagnosis as a variable feature.
</paragraph>
<paragraph id="8BC15252FFEC382AFF6DF94EFB5EB1DA" blockId="12.[151,1437,151,1473]" pageId="12" pageNumber="163">
The genus
<taxonomicName id="4C7E29D1FFEC382AFEE8F94EFD04B14E" authority="Bruce &amp; Javed, 1987" authorityName="Bruce &amp; Javed" authorityYear="1987" box="[322,693,1340,1365]" class="Malacostraca" family="Cirolanidae" genus="Atarbolana" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="B90A8E40FFEC382AFEE8F94EFE70B14F" box="[322,449,1340,1365]" italics="true" pageId="12" pageNumber="163">Atarbolana</emphasis>
<bibRefCitation id="EFEF2FA3FFEC382AFE62F94FFD04B14E" author="Bruce" box="[456,693,1340,1365]" pageId="12" pageNumber="163" refString="Bruce, N. L. &amp; JaVed, W. (1987) A new genus and species of cirolanid isopod Crustacea from the northern Indian Ocean. Journal of Natural History, 21, 1451 - 1460. https: // doi. org / 10.1080 / 00222938700770911" type="journal article" year="1987">Bruce &amp; Javed, 1987</bibRefCitation>
</taxonomicName>
consists of 4 species (
<taxonomicName id="4C7E29D1FFEC382AFC1BF94EFA9DB14E" authority="Bruce &amp; Javed, 1987" authorityName="Bruce &amp; Javed" authorityYear="1987" box="[945,1324,1340,1365]" class="Malacostraca" family="Cirolanidae" genus="Atarbolana" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="exoconta">
<emphasis id="B90A8E40FFEC382AFC1BF94EFB84B14E" box="[945,1077,1340,1364]" italics="true" pageId="12" pageNumber="163">A. exoconta</emphasis>
<bibRefCitation id="EFEF2FA3FFEC382AFB96F94FFA9DB14E" author="Bruce" box="[1084,1324,1340,1365]" pageId="12" pageNumber="163" refString="Bruce, N. L. &amp; JaVed, W. (1987) A new genus and species of cirolanid isopod Crustacea from the northern Indian Ocean. Journal of Natural History, 21, 1451 - 1460. https: // doi. org / 10.1080 / 00222938700770911" type="journal article" year="1987">Bruce &amp; Javed, 1987</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C7E29D1FFEC382AFA9DF94EFE0DB162" authority="Javed &amp; Yasmeen, 1989" authorityName="Javed &amp; Yasmeen" authorityYear="1989" class="Malacostraca" family="Cirolanidae" genus="Atarbolana" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="setosa">
<emphasis id="B90A8E40FFEC382AFA9DF94EFA2DB14E" box="[1335,1436,1340,1364]" italics="true" pageId="12" pageNumber="163">A. setosa</emphasis>
<bibRefCitation id="EFEF2FA3FFEC382AFF3DF92DFE0DB162" author="Javed" box="[151,444,1375,1400]" pageId="12" pageNumber="163" refString="JaVed, W. &amp; Yasmeen, R. (1989) Atarbolana setosa, a new cirolanid isopod from the northern Arabian Sea. Crustaceana, 56, 78 - 82. https: // doi. org / 10.1163 / 156854089 X 00815" type="journal article" year="1989">Javed &amp; Yasmeen, 1989</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C7E29D1FFEC382AFE64F912FC94B162" authority="Yasmeen, 2004" authorityName="Yasmeen" authorityYear="2004" box="[462,805,1375,1400]" class="Malacostraca" family="Cirolanidae" genus="Atarbolana" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="dasycolus">
<emphasis id="B90A8E40FFEC382AFE64F912FDD3B162" box="[462,610,1375,1400]" italics="true" pageId="12" pageNumber="163">A. dasycolus</emphasis>
<bibRefCitation id="EFEF2FA3FFEC382AFDDAF913FC94B162" author="Yasmeen" box="[624,805,1376,1400]" pageId="12" pageNumber="163" refString="Yasmeen, R. (2004) A new species of cirolanid isopod of the genus Atarbolana Bruce and JaVed, 1987 (Crustacea) from the Karachi coast of Pakistan. Pakistan Journal of Marine Sciences, 13, 21 - 26." type="journal article" year="2004">Yasmeen, 2004</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C7E29D1FFEC382AFCC6F912FEDAB187" authority="Khalaji-Pirbalouty, Naderloo &amp; Keikhosravi, 2015" authorityName="Khalaji-Pirbalouty, Naderloo &amp; Keikhosravi" authorityYear="2015" class="Malacostraca" family="Cirolanidae" genus="Atarbolana" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="makranensis">
<emphasis id="B90A8E40FFEC382AFCC6F912FB91B162" box="[876,1056,1375,1400]" italics="true" pageId="12" pageNumber="163">A. makranensis</emphasis>
Khalaji-Pirbalouty,
<bibRefCitation id="EFEF2FA3FFEC382AFABAF92DFEDAB187" author="Naderloo" pageId="12" pageNumber="163" refString="Khalaji-Pirbalouty, V., Naderloo, R. &amp; KeikhosraVi, A. (2015) Atarbolana makranensis, a new species of Cirolanidae (Crustacea, Isopoda) from Makran, Iranian coast of the Gulf of Oman. ZooKeys, 522, 129 - 140. https: // doi. org / 10.3897 / Zookeys. 522.6056" type="journal article" year="2015">Naderloo &amp; Keikhosravi, 2015</bibRefCitation>
</taxonomicName>
) distributed in the Pakistani coast and Gulf of
<collectingCountry id="F36912C2FFEC382AFC38F9F6FC66B186" box="[914,983,1412,1436]" name="Oman" pageId="12" pageNumber="163">Oman</collectingCountry>
(Iranian coast).
<taxonomicName id="4C7E29D1FFEC382AFB3DF9F6FA84B187" authority="Castelló, 2017" authorityName="Castelló" authorityYear="2017" box="[1175,1333,1412,1437]" class="Malacostraca" family="Cirolanidae" genus="Atarbolana" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="beirutensis" status="sp. nov.">
<emphasis id="B90A8E40FFEC382AFB3DF9F6FA84B187" box="[1175,1333,1412,1437]" italics="true" pageId="12" pageNumber="163">A. beirutensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A239333BFFEC382AFA95F9F6FA2DB187" box="[1343,1436,1412,1437]" pageId="12" pageNumber="163" rank="species">
<emphasis id="B90A8E40FFEC382AFA95F9F6FA2DB187" bold="true" box="[1343,1436,1412,1437]" pageId="12" pageNumber="163">sp. nov.</emphasis>
</taxonomicNameLabel>
expands the distribution of this genus, which is recorded for the first time in the Mediterranean sea.
</paragraph>
</subSubSection>
</treatment>
</document>
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