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<document id="25040BE5280AB214EC10D718F5A654D4" ID-DOI="10.11646/zootaxa.4755.2.12" ID-GBIF-Dataset="6b11c9bd-eae8-4973-bd53-5ebe9b658b1d" ID-ISSN="1175-5334" ID-Zenodo-Dep="3733721" ID-ZooBank="urn:lsid:zoobank.org:pub:375D7830-D90A-4D91-905F-786A1215DFED" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1585597547394" checkinUser="carolina" docAuthor="Liu, Chen &amp; Wang, Shuxia" docDate="2020" docId="0381830DFFB0FFA1FF0BFF5D018FB758" docLanguage="en" docName="zootaxa.4755.2.12.pdf.imf" docOrigin="Zootaxa 4755 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Promalactis tenuiclavifera Liu &amp; Wang 2020, sp. nov." docType="treatment" docVersion="9" lastPageNumber="392" masterDocId="FFB8FB75FFB2FFA3FF9CFFC4006BB223" masterDocTitle="Genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from Laos, with descriptions of five new species" masterLastPageNumber="398" masterPageNumber="390" pageNumber="392" updateTime="1698733430596" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="CDA2602815FA426738B5A2AB4678B9A0">Genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from Laos, with descriptions of five new species</mods:title>
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<mods:namePart id="49A02B6D4184C652D47FE39DD3F07439">Liu, Chen</mods:namePart>
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<mods:namePart id="9D0F5EB18C420F286D2A98B6CF28D2F6">Wang, Shuxia</mods:namePart>
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<mods:date id="86B366A257568802C570151C4AE13357">2020</mods:date>
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<treatment id="0381830DFFB0FFA1FF0BFF5D018FB758" ID-DOI="http://doi.org/10.5281/zenodo.3812468" ID-GBIF-Taxon="164401365" ID-Zenodo-Dep="3812468" LSID="urn:lsid:plazi:treatment:0381830DFFB0FFA1FF0BFF5D018FB758" httpUri="http://treatment.plazi.org/id/0381830DFFB0FFA1FF0BFF5D018FB758" lastPageNumber="392" pageId="2" pageNumber="392">
<subSubSection id="C3326190FFB0FFA1FF0BFF5D02FFB297" box="[151,660,153,180]" pageId="2" pageNumber="392" type="nomenclature">
<paragraph id="8B97321BFFB0FFA1FF0BFF5D02FFB297" blockId="2.[151,660,153,215]" box="[151,660,153,180]" pageId="2" pageNumber="392">
<heading id="D0DF8577FFB0FFA1FF0BFF5D02FFB297" bold="true" box="[151,660,153,180]" fontSize="10" level="2" pageId="2" pageNumber="392" reason="2">
<emphasis id="B95CEE09FFB0FFA1FF0BFF5D02FFB297" bold="true" box="[151,660,153,180]" pageId="2" pageNumber="392">
<taxonomicName id="4C284998FFB0FFA1FF0BFF5D0241B297" authority="Wang" authorityName="Liu &amp; Wang" authorityYear="2020" box="[151,554,153,180]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="392" phylum="Arthropoda" rank="species" species="tenuiclavifera" status="sp. nov.">
<emphasis id="B95CEE09FFB0FFA1FF0BFF5D01B3B297" bold="true" box="[151,472,153,180]" italics="true" pageId="2" pageNumber="392">Promalactis tenuiclavifera</emphasis>
Wang
</taxonomicName>
,
<taxonomicNameLabel id="A26F5372FFB0FFA1FDABFF5E02FFB297" box="[567,660,154,180]" pageId="2" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3326190FFB0FFA1FF0BFF79014AB2F4" box="[151,289,189,215]" pageId="2" pageNumber="392" type="description">
<paragraph id="8B97321BFFB0FFA1FF0BFF79014AB2F4" blockId="2.[151,660,153,215]" box="[151,289,189,215]" pageId="2" pageNumber="392">
(
<figureCitation id="13132E9EFFB0FFA1FF03FF790084B2F4" box="[159,239,189,215]" captionStart="FIGURES 1‒8" captionStartId="5.[151,264,1881,1906]" captionTargetBox="[179,1435,179,1854]" captionTargetId="figure@5.[151,1435,179,1854]" captionTargetPageId="5" captionText="FIGURES 1‒8. Adults of Promalactis spp. 1, P. unidentalis, holotype ♂; 2, P. tenuiclavifera, holotype ♂; 3, P. introflexa, holotype ♂; 4, P. bolikhamsaiana, holotype ♂; 5, P. serrulata, holotype ♂; 6, P. carinata Du, Li et Wang, 2011; 7, P. flavidia Kim, 2018; 8, P. apicuncata Wang, 2018. Scales=2 mm." figureDoi="http://doi.org/10.5281/zenodo.3733723" httpUri="https://zenodo.org/record/3733723/files/figure.png" pageId="2" pageNumber="392">Figs. 2</figureCitation>
,
<figureCitation id="13132E9EFFB0FFA1FF67FF790173B2F4" box="[251,280,189,215]" captionStart="FIGURES 9‒14" captionStartId="6.[151,264,1898,1923]" captionTargetBox="[248,1351,190,1853]" captionTargetId="figure@6.[226,1361,181,1872]" captionTargetPageId="6" captionText="FIGURES 9‒14. Genitalia of Promalactis spp. 9, P. unidentalis, holotype ♂, slide No. LC19247; 10, P. tenuiclavifera, holotype ♂, slide No. LC19241; 11, P. introflexa, holotype ♂, slide No. LC19242; 12a, P. introflexa, paratype ♀, slide No. LC19302, 12b, enlarged signum, slide No. LC19246; 13, P. bolikhamsaiana, holotype ♂, slide No. LC19262; 14, P. serrulata, holotype ♂, slide No. LC19250. Scales=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3733725" httpUri="https://zenodo.org/record/3733725/files/figure.png" pageId="2" pageNumber="392">10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3326190FFB0FFA1FF0BFEC2020CB344" pageId="2" pageNumber="392" type="materials_examined">
<paragraph id="8B97321BFFB0FFA1FF0BFEC204FEB360" blockId="2.[151,1437,261,1404]" pageId="2" pageNumber="392">
<emphasis id="B95CEE09FFB0FFA1FF0BFEC2012EB303" bold="true" box="[151,325,261,288]" pageId="2" pageNumber="392">Type material.</emphasis>
<materialsCitation id="3B403846FFB0FFA1FED0FEC104FEB360" ID-GBIF-Occurrence="2592567759" collectingDate="2019-07-29" collectorName="YS Bae &amp; DJ Lee &amp; No. LC" country="Laos" elevation="958" latitude="15.18" location="Dong Hua Sao National Biodiversity" longLatPrecision="772" longitude="106.13" municipality="Hua Sao National Biodiversity Conservation Area" pageId="2" pageNumber="392" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54938CB9FFB0FFA1FED0FEC101DFB33C" box="[332,436,261,287]" pageId="2" pageNumber="392" type="holotype">Holotype</typeStatus>
<specimenCount id="9D2EF992FFB0FFA1FE27FEC201B9B303" box="[443,466,262,288]" pageId="2" pageNumber="392" type="male">♂</specimenCount>
,
<location id="8EF764C0FFB0FFA1FE42FEC20310B33C" LSID="urn:lsid:plazi:treatment:0381830DFFB0FFA1FF0BFF5D018FB758:8EF764C0FFB0FFA1FE42FEC20310B33C" box="[478,891,261,288]" country="Laos" latitude="15.18" longLatPrecision="772" longitude="106.13" municipality="Hua Sao National Biodiversity Conservation Area" name="Dong Hua Sao National Biodiversity" pageId="2" pageNumber="392">Dong Hua Sao National Biodiversity</location>
Conservation Area (
<geoCoordinate id="EE1C54DCFFB0FFA1FBFEFEC104AFB303" box="[1122,1220,261,288]" degrees="15.18" direction="north" orientation="latitude" pageId="2" pageNumber="392" precision="555" value="15.18">15.18°N</geoCoordinate>
,
<geoCoordinate id="EE1C54DCFFB0FFA1FB53FEC10556B303" box="[1231,1341,261,288]" degrees="106.13" direction="east" orientation="longitude" pageId="2" pageNumber="392" precision="555" value="106.13">106.13°E</geoCoordinate>
), Cham- pasak,
<collectingCountry id="F33F728BFFB0FFA1FF79FEEE0175B367" box="[229,286,298,324]" name="Laos" pageId="2" pageNumber="392">Laos</collectingCountry>
,
<quantity id="4CD09FFEFFB0FFA1FEB5FEED0119B367" box="[297,370,297,324]" metricMagnitude="2" metricUnit="m" metricValue="9.58" pageId="2" pageNumber="392" unit="m" value="958.0">
<elevation id="0005D528FFB0FFA1FEB5FEED0119B367" box="[297,370,297,324]" metricMagnitude="2" metricUnit="m" metricValue="9.58" pageId="2" pageNumber="392" unit="m" value="958.0">958 m</elevation>
</quantity>
,
<date id="FF9614DBFFB0FFA1FEE2FEED0261B360" box="[382,522,297,323]" pageId="2" pageNumber="392" value="2019-07-29">
<collectingDate id="EFD2ED33FFB0FFA1FEE2FEED0261B360" box="[382,522,297,323]" pageId="2" pageNumber="392" value="2019-07-29">29.VII.2019</collectingDate>
</date>
, coll.
<collectorName id="26DD57CDFFB0FFA1FDD0FEED02CEB367" box="[588,677,297,324]" pageId="2" pageNumber="392">YS Bae</collectorName>
, MJ Qi &amp;
<collectorName id="26DD57CDFFB0FFA1FC87FEEE0300B367" box="[795,875,298,324]" pageId="2" pageNumber="392">DJ Lee</collectorName>
<emphasis id="B95CEE09FFB0FFA1FCEEFEEE03C1B360" box="[882,938,297,323]" italics="true" pageId="2" pageNumber="392">et al.</emphasis>
, slide
<collectorName id="26DD57CDFFB0FFA1FC68FEEE0421B360" box="[1012,1098,297,324]" pageId="2" pageNumber="392">No. LC</collectorName>
19241.
</materialsCitation>
</paragraph>
<paragraph id="8B97321BFFB0FFA1FF5BFE8A020CB344" blockId="2.[151,1437,261,1404]" box="[199,615,333,360]" pageId="2" pageNumber="392">
<materialsCitation id="3B403846FFB0FFA1FF5BFE8A020CB344" ID-GBIF-Occurrence="2592567760" box="[199,615,333,360]" collectingDate="2019-07-29" collectorName="YS Bae &amp; DJ Lee &amp; No. LC" country="Laos" elevation="958" latitude="15.18" location="Dong Hua Sao National Biodiversity" longLatPrecision="772" longitude="106.13" municipality="Hua Sao National Biodiversity Conservation Area" pageId="2" pageNumber="392" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54938CB9FFB0FFA1FF5BFE8A0146B34B" box="[199,301,334,360]" pageId="2" pageNumber="392" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D2EF992FFB0FFA1FEA4FE890137B344" box="[312,348,333,359]" pageId="2" pageNumber="392" type="male">1♂</specimenCount>
, same data as holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3326190FFB0FFA1FF5BFEB501A6B3D4" pageId="2" pageNumber="392" type="diagnosis">
<paragraph id="8B97321BFFB0FFA1FF5BFEB501A6B3D4" blockId="2.[151,1437,261,1404]" pageId="2" pageNumber="392">
<emphasis id="B95CEE09FFB0FFA1FF5BFEB50129B3AF" bold="true" box="[199,322,369,396]" pageId="2" pageNumber="392">Diagnosis.</emphasis>
This new species is similar to
<taxonomicName id="4C284998FFB0FFA1FD30FEB6045AB3A8" authority="Kim &amp; Park, 2014" authorityName="Kim &amp; Park" authorityYear="2014" box="[684,1073,369,395]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="392" phylum="Arthropoda" rank="species" species="caperatiella">
<emphasis id="B95CEE09FFB0FFA1FD30FEB6033AB3A8" box="[684,849,369,395]" italics="true" pageId="2" pageNumber="392">P. caperatiella</emphasis>
<bibRefCitation id="EFB94FEAFFB0FFA1FCC7FEB5045AB3A8" author="Kim &amp; Park" box="[859,1073,369,395]" firstAuthor="Kim" pageId="2" pageNumber="392" pagination="1327 - 1346" refId="ref4796" refString="Kim, S., Park, K. T., Heppner, J. B. &amp; Lee, S. (2014) Genus Promalactis Meyrick (Lepidoptera: Oecophoridae) in northern Vietnam, Part III: seven new species and four newly recorded species. Journal of Natural History, 48 (21 - 22), 1327 - 1346. https: // doi. org / 10.1080 / 00222933.2013.862576" type="journal article" year="2014">
Kim &amp; Park,
<quantity id="4CD09FFEFFB0FFA1FC65FEB5045AB3A8" box="[1017,1073,369,395]" metricMagnitude="1" metricUnit="m" metricValue="5.11556" pageId="2" pageNumber="392" unit="in" value="2014.0">2014</quantity>
</bibRefCitation>
</taxonomicName>
in the male genitalia. It can be distinguished from the latter by the valva produced to a rectangular process in distal 1/6 and the two equally sized cornuti; whereas in
<taxonomicName id="4C284998FFB0FFA1FEE7FE7E024BB3F0" authorityName="Kim &amp; Park" authorityYear="2014" box="[379,544,441,467]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="392" phylum="Arthropoda" rank="species" species="caperatiella">
<emphasis id="B95CEE09FFB0FFA1FEE7FE7E024BB3F0" box="[379,544,441,467]" italics="true" pageId="2" pageNumber="392">P. caperatiella</emphasis>
</taxonomicName>
, the distal 1/5 of the valva is produced to a sub-crescent process and the two cornuti are unequally sized.
</paragraph>
</subSubSection>
<subSubSection id="C3326190FFB0FFA1FF5BFDC501FAB72C" pageId="2" pageNumber="392" type="description">
<paragraph id="8B97321BFFB0FFA1FF5BFDC5034CB03F" blockId="2.[151,1437,261,1404]" box="[199,807,513,540]" pageId="2" pageNumber="392">
<emphasis id="B95CEE09FFB0FFA1FF5BFDC50132B03F" bold="true" box="[199,345,513,540]" pageId="2" pageNumber="392">Description.</emphasis>
Adult (
<figureCitation id="13132E9EFFB0FFA1FE31FDC50198B038" box="[429,499,513,539]" captionStart="FIGURES 1‒8" captionStartId="5.[151,264,1881,1906]" captionTargetBox="[179,1435,179,1854]" captionTargetId="figure@5.[151,1435,179,1854]" captionTargetPageId="5" captionText="FIGURES 1‒8. Adults of Promalactis spp. 1, P. unidentalis, holotype ♂; 2, P. tenuiclavifera, holotype ♂; 3, P. introflexa, holotype ♂; 4, P. bolikhamsaiana, holotype ♂; 5, P. serrulata, holotype ♂; 6, P. carinata Du, Li et Wang, 2011; 7, P. flavidia Kim, 2018; 8, P. apicuncata Wang, 2018. Scales=2 mm." figureDoi="http://doi.org/10.5281/zenodo.3733723" httpUri="https://zenodo.org/record/3733723/files/figure.png" pageId="2" pageNumber="392">Fig. 2</figureCitation>
). Wingspan 10.0‒11.0 mm.
</paragraph>
<paragraph id="8B97321BFFB0FFA1FF5BFDE102C9B088" blockId="2.[151,1437,261,1404]" pageId="2" pageNumber="392">
<emphasis id="B95CEE09FFB0FFA1FF5BFDE1016FB01C" box="[199,260,549,575]" italics="true" pageId="2" pageNumber="392">Head</emphasis>
with vertex and frons yellow, tinged with pale brownish yellow laterally. Labial palpus with second segment dim ochreous yellow on outer surface, orange yellow on inner surface; third segment with basal 1/3 whitish yellow, distal 2/3 dark ochreous brown mixed with black scales. Antenna with scape white, black at apex; flagellum black annulated with yellow on dorsal surface.
</paragraph>
<paragraph id="8B97321BFFB0FFA1FF5BFD7103CEB1E8" blockId="2.[151,1437,261,1404]" pageId="2" pageNumber="392">
<emphasis id="B95CEE09FFB0FFA1FF5BFD710173B0EC" box="[199,280,693,719]" italics="true" pageId="2" pageNumber="392">Thorax</emphasis>
and tegula pale rust brown. Forewing with basal 3/4 dark yellowish brown, distal 1/4 greyish yellow; costal spot large, rounded, cream white, with pale ochreous yellow scales on outer 1/3, with dense black scales on inner and posterior margins, extending from beyond distal 1/3 to above lower angle of cell; basal fascia slender, edged with black scales along inner margin; antemedian fascia from basal 1/4 of costal margin sinuate to distal 1/4 on dorsum, parallel with basal fascia, widened anteriorly, edged with black scales on inner margin, with orange yellow on outer margin; ill-defined blackish brown band from before end of fold to below costal spot, not diffused to tornus; apex with dense blackish brown scales; cilia orange yellow, grey along distal part of costal margin, mixed with greyish brown around tornus distally. Hindwing and cilia brown.
</paragraph>
<paragraph id="8B97321BFFB0FFA1FF5BFC1101EEB6C8" blockId="2.[151,1437,261,1404]" pageId="2" pageNumber="392">
<emphasis id="B95CEE09FFB0FFA1FF5BFC110100B1CC" box="[199,363,981,1007]" italics="true" pageId="2" pageNumber="392">Male genitalia</emphasis>
(
<figureCitation id="13132E9EFFB0FFA1FEE5FC1101A0B1CC" box="[377,459,981,1007]" captionStart="FIGURES 9‒14" captionStartId="6.[151,264,1898,1923]" captionTargetBox="[248,1351,190,1853]" captionTargetId="figure@6.[226,1361,181,1872]" captionTargetPageId="6" captionText="FIGURES 9‒14. Genitalia of Promalactis spp. 9, P. unidentalis, holotype ♂, slide No. LC19247; 10, P. tenuiclavifera, holotype ♂, slide No. LC19241; 11, P. introflexa, holotype ♂, slide No. LC19242; 12a, P. introflexa, paratype ♀, slide No. LC19302, 12b, enlarged signum, slide No. LC19246; 13, P. bolikhamsaiana, holotype ♂, slide No. LC19262; 14, P. serrulata, holotype ♂, slide No. LC19250. Scales=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3733725" httpUri="https://zenodo.org/record/3733725/files/figure.png" pageId="2" pageNumber="392">Fig. 10</figureCitation>
). Uncus wide at base, narrowed to basal 3/5, distinctly narrowed from distal 2/5 to apex. Gnathos almost as long as uncus; gnathos plate short, uniformly narrow, rounded at apex; lateral arm wide, bandshaped, longer than gnathos plate. Tegumen bifurcate from posterior 1/3; lateral arm narrowed anteriorly. Valva with basal 2/3 rectangular, then abruptly narrowed to distal 1/6, distal 1/6 produced to a rectangular process, serrate on edges, straight at apex; costa very narrow, reaching basal 2/3; sacculus slightly wide at base, concave at dorsal 2/5, distal 1/3 produced to a free clavate process, setose distally. Saccus clubbed, as long as valva, narrowly rounded apically. Juxta being a large irregular plate. Aedeagus strong, straight, longer than valva; two cornuti equally sized, strong, spine-shaped.
</paragraph>
<paragraph id="8B97321BFFB0FFA1FF5BFB3101FAB72C" blockId="2.[151,1437,261,1404]" box="[199,401,1269,1295]" pageId="2" pageNumber="392">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C3326190FFB0FFA1FF5BFADD0229B717" box="[199,578,1305,1332]" pageId="2" pageNumber="392" type="distribution">
<paragraph id="8B97321BFFB0FFA1FF5BFADD0229B717" blockId="2.[151,1437,261,1404]" box="[199,578,1305,1332]" pageId="2" pageNumber="392">
<emphasis id="B95CEE09FFB0FFA1FF5BFADD0132B717" bold="true" box="[199,345,1305,1332]" pageId="2" pageNumber="392">Distribution</emphasis>
.
<collectingCountry id="F33F728BFFB0FFA1FEFBFADE01F6B717" box="[359,413,1306,1332]" name="Laos" pageId="2" pageNumber="392">Laos</collectingCountry>
(
<collectingRegion id="49ECFCF9FFB0FFA1FE31FADD025CB717" box="[429,567,1305,1332]" country="Laos" name="Champasak" pageId="2" pageNumber="392">Champasak</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3326190FFB0FFA1FF5BFAFA018FB758" pageId="2" pageNumber="392" type="etymology">
<paragraph id="8B97321BFFB0FFA1FF5BFAFA018FB758" blockId="2.[151,1437,261,1404]" pageId="2" pageNumber="392">
<emphasis id="B95CEE09FFB0FFA1FF5BFAFA0125B77B" bold="true" box="[199,334,1342,1368]" pageId="2" pageNumber="392">Etymology.</emphasis>
The epithet of this new species is derived the Latin
<emphasis id="B95CEE09FFB0FFA1FC36FAFA0380B774" box="[938,1003,1342,1367]" italics="true" pageId="2" pageNumber="392">tenui-</emphasis>
and
<emphasis id="B95CEE09FFB0FFA1FBB9FAF904E2B774" box="[1061,1161,1341,1367]" italics="true" pageId="2" pageNumber="392">clavifera</emphasis>
, referring to the clavate distal process of the sacculus.
</paragraph>
</subSubSection>
</treatment>
</document>
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