232 lines
25 KiB
XML
232 lines
25 KiB
XML
<document id="86B0067FF92F33F7FE5E5C6BBD75DBF9" ID-DOI="10.5281/zenodo.10396362" ID-ISSN="1942-1354" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1707168658757" checkinUser="felipe" docAuthor="Zhang, Jing, Cong, Qian & Grishin, Nick V." docDate="2023" docId="03810139FFFCBB74C0CAFB07E192B028" docLanguage="en" docName="InsectaMundi.2023.1026.1-115.pdf" docOrigin="Insecta Mundi 2023 (26)" docSource="http://dx.doi.org/10.5281/zenodo.10396362" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Staphylus (Vulga) vulga vulga Grishin 2023, new species" docType="treatment" docVersion="1" lastPageNumber="35" masterDocId="FFB87941FFDFBB50C00AFF56E355B262" masterDocTitle="Supplementary Materials and Appendix" masterLastPageNumber="115" masterPageNumber="1" pageNumber="34" updateTime="1707169288913" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="9D598A81E956FB1DABAFCEDD99CD05ED">
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<mods:title id="4BCF7140876B3E18FA666CAEC846F884">Supplementary Materials and Appendix</mods:title>
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</mods:titleInfo>
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<mods:name id="B09791F6417C4F1966118CEFBF57999B" type="personal">
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<mods:roleTerm id="991B6D69BD68F945A832D29D873B3B19">Author</mods:roleTerm>
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<mods:namePart id="A429D8E2ED117EAE5AAF4F0A2CA6F67A">Zhang, Jing</mods:namePart>
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<mods:affiliation id="966B95DAE1B7F4A25A8EAC33F317F490">McDermott Center for Human Growth and Development and Department of Biophysics University of Texas Southwestern Medical Center 5323 Harry Hines Blvd., Dallas, TX, 75390 - 8816 USA</mods:affiliation>
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<mods:roleTerm id="2C411E384804AB58E14C008695BBAE63">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D6BECC9B00AD8A40A8E3EB130E230BF4">Cong, Qian</mods:namePart>
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<mods:affiliation id="6F20D7964DB32AA22C8249B742D4D61B">McDermott Center for Human Growth and Development and Department of Biophysics University of Texas Southwestern Medical Center 5323 Harry Hines Blvd., Dallas, TX, 75390 - 8816 USA</mods:affiliation>
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<mods:roleTerm id="222FE83939131C9D34852EC19E28DC13">Author</mods:roleTerm>
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<mods:namePart id="A3344DAD74442BBF9C5FFE3D6BBB974A">Grishin, Nick V.</mods:namePart>
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<mods:affiliation id="58ECBF50F7963071EA1317A95897182F">Departments of Biophysics and Biochemistry University of Texas Southwestern Medical Center 5323 Harry Hines Blvd., Dallas, TX, 75390 - 9050 USA</mods:affiliation>
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<mods:typeOfResource id="1A3AECC54E2CCCF6B59D9531464ADFCD">text</mods:typeOfResource>
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<mods:title id="D250F4B4CB368E87D32E46AD02613461">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="0D560D3BBE091F42C37036E09AD74979">
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<mods:date id="FC1E8C54E152D7E5E9510B434A906652">2023</mods:date>
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<mods:number id="DB805D75C08C9EB51E559823A580DBCC">2023-12-29</mods:number>
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<mods:number id="8903D4D90CCCAEC7987E64093754CF70">2023</mods:number>
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<mods:number id="37018FEF0E01D78BEDC3ADBB878391B9">26</mods:number>
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<mods:start id="197BFF1E673B594622E43796F856057B">1</mods:start>
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<mods:url id="B06AB593B98B2F97D6D8610D999DD4DF">http://dx.doi.org/10.5281/zenodo.10396362</mods:url>
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<mods:identifier id="D097D7CCCA01BB3EF1826877C98286E0" type="DOI">10.5281/zenodo.10396362</mods:identifier>
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<mods:identifier id="71420896323EF29A71D4F3DCD84B8CB9" type="ISSN">1942-1354</mods:identifier>
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<treatment id="03810139FFFCBB74C0CAFB07E192B028" LSID="urn:lsid:plazi:treatment:03810139FFFCBB74C0CAFB07E192B028" httpUri="http://treatment.plazi.org/id/03810139FFFCBB74C0CAFB07E192B028" lastPageId="36" lastPageNumber="35" pageId="35" pageNumber="34">
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<subSubSection id="C332E3A4FFFCBB73C0CAFB07E0FFB6F3" pageId="35" pageNumber="34" type="nomenclature">
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<paragraph id="8B97B02FFFFCBB73C0CAFB07E1ADB60C" blockId="35.[192,938,1105,1203]" box="[192,760,1105,1134]" pageId="35" pageNumber="34">
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<heading id="D0DF0743FFFCBB73C0CAFB07E1ADB60C" box="[192,760,1105,1134]" fontSize="12" level="2" pageId="35" pageNumber="34" reason="2">
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<emphasis id="B95C6C3DFFFCBB73C0CAFB07E1ADB60C" bold="true" box="[192,760,1105,1134]" pageId="35" pageNumber="34">
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<taxonomicName id="4C28CBACFFFCBB73C0CAFB07E102B60C" authority="Grishin" authorityName="vulga Grishin" authorityYear="2023" box="[192,599,1105,1134]" class="Insecta" family="Hesperiidae" genus="Staphylus" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="34" phylum="Arthropoda" rank="species" species="vulga" status="sp. nov." subGenus="Vulga">
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<emphasis id="B95C6C3DFFFCBB73C0CAFB07E26EB60C" bold="true" box="[192,315,1105,1134]" italics="true" pageId="35" pageNumber="34">Staphylus</emphasis>
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(
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<emphasis id="B95C6C3DFFFCBB73C141FB07E2C1B60C" bold="true" box="[331,404,1105,1134]" italics="true" pageId="35" pageNumber="34">Vulga</emphasis>
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)
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<emphasis id="B95C6C3DFFFCBB73C1ACFB07E2BEB60C" bold="true" box="[422,491,1105,1134]" italics="true" pageId="35" pageNumber="34">vulga</emphasis>
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Grishin
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26FD146FFFCBB73C269FB04E1ADB60C" box="[611,760,1105,1134]" pageId="35" pageNumber="34" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8B97B02FFFFCBB73C0CAFB20E0FFB6F3" blockId="35.[192,938,1105,1203]" box="[192,938,1142,1169]" pageId="35" pageNumber="34">
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||
https://zoobank.org/
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||
<uuid id="FF8E8AFAFFFCBB73C1A9FB20E0FFB6F3" box="[419,938,1142,1169]" pageId="35" pageNumber="34">1ECE61F4-A057-4903-8706-D6CD468A8B87</uuid>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C332E3A4FFFCBB73C0CAFBCEE157B6D1" box="[192,514,1176,1203]" pageId="35" pageNumber="34" type="description">
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<paragraph id="8B97B02FFFFCBB73C0CAFBCEE157B6D1" blockId="35.[192,938,1105,1203]" box="[192,514,1176,1203]" pageId="35" pageNumber="34">
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(
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<figureCitation id="1313ACAAFFFCBB73C0C2FBCEE252B6D0" box="[200,263,1176,1202]" captionStart="Figure 3" captionStartId="33.[192,261,1387,1411]" captionTargetBox="[193,1438,244,1339]" captionTargetId="figure-11@33.[192,1440,242,1340]" captionTargetPageId="33" captionText="Figure 3. Phylogenetic trees of remaining Pyrginae (excluding Achlyodini) inferred from protein-coding regions in a) the Z chromosome and b) the mitochondrial genome (in segments). See Fig. 1 legend for other notations." pageId="35" pageNumber="34">Fig. 3</figureCitation>
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part, 77–78, 295–297)
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</paragraph>
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</subSubSection>
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<subSubSection id="C332E3A4FFFCBB73C0CAFB94E7D8B555" pageId="35" pageNumber="34" type="diagnosis">
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<paragraph id="8B97B02FFFFCBB73C0CAFB94E7E7B437" blockId="35.[192,1441,1218,1622]" pageId="35" pageNumber="34">
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<emphasis id="B95C6C3DFFFCBB73C0CAFB94E2B8B6BE" bold="true" box="[192,493,1218,1244]" pageId="35" pageNumber="34">Definition and diagnosis.</emphasis>
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Phylogenetic analysis that included
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<typeStatus id="54930E8DFFFCBB73C387FB95E0BBB6BF" box="[909,1006,1219,1245]" pageId="35" pageNumber="34" type="syntype">syntypes</typeStatus>
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of
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<taxonomicName id="4C28CBACFFFCBB73C41DFB94E6F5B6BE" authority="(Moschler, 1879)" baseAuthorityName="Moschler" baseAuthorityYear="1879" box="[1047,1440,1218,1244]" class="Insecta" family="Hesperiidae" genus="Staphylus" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="34" phylum="Arthropoda" rank="species" species="vulgata">
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<emphasis id="B95C6C3DFFFCBB73C41DFB94E78FB6BE" box="[1047,1242,1218,1244]" italics="true" pageId="35" pageNumber="34">Staphylus vulgata</emphasis>
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(Möschler, 1879)
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</taxonomicName>
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(type locality in
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<collectingCountry id="F33FF0BFFFFCBB73C171FBB2E2B9B69C" box="[379,492,1252,1278]" name="Colombia" pageId="35" pageNumber="34">Colombia</collectingCountry>
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) reveals that Panamanian specimens identified as this species are strongly differentiated genetically from and are not even monophyletic with it (
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<figureCitation id="1313ACAAFFFCBB73C365FA51E0E4B743" box="[879,945,1287,1313]" captionStart="Figure 3" captionStartId="33.[192,261,1387,1411]" captionTargetBox="[193,1438,244,1339]" captionTargetId="figure-11@33.[192,1440,242,1340]" captionTargetPageId="33" captionText="Figure 3. Phylogenetic trees of remaining Pyrginae (excluding Achlyodini) inferred from protein-coding regions in a) the Z chromosome and b) the mitochondrial genome (in segments). See Fig. 1 legend for other notations." pageId="35" pageNumber="34">Fig. 3</figureCitation>
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), e.g., their COI barcodes differ by 5.9% (39 bp). These specimens are sister to
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<taxonomicName id="4C28CBACFFFCBB73C24CFA7FE1B3B721" authority="Zhang & Cong & Grishin, 2023" authorityName="Zhang & Cong & Grishin" authorityYear="2023" box="[582,742,1321,1347]" class="Insecta" family="Hesperiidae" genus="Staphylus" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="34" phylum="Arthropoda" rank="species" species="vula" status="sp. nov.">
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<emphasis id="B95C6C3DFFFCBB73C24CFA7FE1B3B721" box="[582,742,1321,1347]" italics="true" pageId="35" pageNumber="34">Staphylus vula</emphasis>
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</taxonomicName>
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<emphasis id="B95C6C3DFFFCBB73C2E7FA7FE023B721" bold="true" box="[749,886,1321,1347]" pageId="35" pageNumber="34">
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||
<taxonomicNameLabel id="A26FD146FFFCBB73C2E7FA7FE023B721" box="[749,886,1321,1347]" pageId="35" pageNumber="34" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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but differ genetically from it (e.g., COI barcodes differ by 1.5%, 10 bp), and, therefore, represent a new species. This new species keys to
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<taxonomicName id="4C28CBACFFFCBB73C49DFA1AE656B707" baseAuthorityName="Moschler" baseAuthorityYear="1879" box="[1175,1283,1355,1382]" class="Insecta" family="Hesperiidae" genus="Staphylus" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="34" phylum="Arthropoda" rank="species" species="vulgata">
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<emphasis id="B95C6C3DFFFCBB73C49DFA1AE656B707" box="[1175,1283,1355,1382]" italics="true" pageId="35" pageNumber="34">S. vulgata</emphasis>
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</taxonomicName>
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(E.32.6(a)) in
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<bibRefCitation id="EFB9CDDEFFFCBB73C0CBFA38E200B7EA" author="Evans WH" box="[193,341,1389,1416]" pageId="35" pageNumber="34" pagination="26 - 53" refId="ref77110" refString="Evans WH. 1953. A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part III. Pyrginae. Section 2. The Trustees of the British Museum (Natural History); London. v + 246 p., pl. 26 - 53." type="book chapter" year="1953">Evans (1953)</bibRefCitation>
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and differs from its relatives by the following combination of characters: costa of valva strongly convex, nearly angled at the dorsal-most point, valva broader in lateral view, expansion of ampulla directed distad and slightly ventrad towards its end, and reaches about the middle of harpe, its terminal spines longer, harpe semi-triangular, distally rounded, with concave near its base dorsoposterior margin (
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<figureCitation id="1313ACAAFFFCBB73C477FA82E641B78D" box="[1149,1300,1492,1519]" captionStart="Figures 285–311" captionStartId="110.[192,271,1719,1743]" captionTargetBox="[205,1420,253,1687]" captionTargetId="figure-150@110.[192,1440,229,1702]" captionTargetPageId="110" captionText="Figures 285–311. Genitalia of holotypes (unless indicated), data in text.285–287) Polyctor (Fenops) lamperus sp. n.288–290) Nisoniades (Nisoniades) lutum sp. n. 291–292) Bolla (Stolla) vena sp. n. 293–294) Staphylus (Vulga) vula sp. n. 295–297) Staphylus (Vulga) vulga sp. n. 298–299) Staphylus (Staphylus) rotundalus sp. n. 300–301) Staphylus (Staphylus) yucatanus sp. n. paratype ♂ NVG-20063A08. 302–303) Heliopetes (Heliopetes) lana sp. n. paratype ♂ NVG-20062H06. 304–306) Canesia ella sp. n. 307–309) Paches (Paches) loxeca sp. n. 310–311) Clito congruens sp. n. Views: dorsal (285, 288, 291, 293, 295, 298, 300, 302, 305, 307, 310), left lateral (287, 290, 292, 294, 297, 299, 301, 303, 304 aedeagus, 306, 308, 309 aedeagus, 311), right lateral (286, 289, 296)." pageId="35" pageNumber="34">Fig. 295–297</figureCitation>
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). Due to the cryptic nature of this species, most reliable identification is achieved by DNA and a combination of the following base pairs is diagnostic in the nuclear genome: aly361.16.7:C229T, aly887.42.4:A63G, aly116.34.5:G392A, aly727.17.5:G94T, aly728.49.1:T1077C, and COI barcode: A40G, T50T, 85A, A202A, 400T.
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</paragraph>
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<paragraph id="8B97B02FFFFCBB73C0CAF933E7D8B555" blockId="35.[192,1440,1637,1847]" pageId="35" pageNumber="34">
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<emphasis id="B95C6C3DFFFCBB73C0CAF933E11BB41D" bold="true" box="[192,590,1637,1663]" pageId="35" pageNumber="34">
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Barcode sequence of the
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<typeStatus id="54930E8DFFFCBB73C1EAF933E11CB41D" box="[480,585,1637,1663]" pageId="35" pageNumber="34" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Sample NVG-18058C11, GenBank OR837657, 658 base pairs: AACTTTATATTTTATTTTTGGTATTTGAGCAGGAATAGTGGGAACTTCTTTAAGTATTCTTATTCGATCAGAATTAGGAACCCCAGGATCTTTAATT GGAGATGATCAAATTTATAATACTATTGTAACTGCTCATGCTTTTATTATAATTTTTTTTATAGTTATACCTATTATAATTGGAGGATTTGGAAATT GACTTGTACCTCTTATATTAGGAGCTCCTGATATAGCTTTTCCTCGAATAAATAATATAAGTTTTTGATTATTACCCCCATCCTTAATACTTTTAAT TTCAAGTAGCATTGTAGAAAATGGAGCAGGAACTGGATGAACTGTATATCCCCCACTTTCAGCCAATATTGCCCATCAAGGTTCATCTGTAGATTTA GCTATTTTTTCTCTTCATTTAGCTGGAATTTCTTCCATTTTAGGAGCAATTAACTTCATCACAACTATTATTAATATACGAATTAATAATTTATCAT TTGATCAAATACCTTTATTTGTATGAGCTGTAGGAATTACAGCATTACTTTTACTCCTATCTTTACCAGTATTAGCTGGAGCTATTACTATACTTTT AACTGATCGAAATCTTAATACATCATTTTTTGATCCCGCTGGAGGAGGTGATCCTATTTTATATCAACATTTATTT
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C332E3A4FFFCBB74C0CAF81CE0DFB3D0" lastPageId="36" lastPageNumber="35" pageId="35" pageNumber="34" type="materials_examined">
|
||
<paragraph id="8B97B02FFFFCBB73C0CAF81CE6F4B5E4" blockId="35.[192,1441,1866,1926]" pageId="35" pageNumber="34">
|
||
<emphasis id="B95C6C3DFFFCBB73C0CAF81CE239B506" bold="true" box="[192,364,1866,1892]" pageId="35" pageNumber="34">Type material.</emphasis>
|
||
<materialsCitation id="3B40BA72FFFCBB73C17FF81CE6F4B5E4" collectionCode="DC, USNM" country="United States of America" location="National Museum of Natural History" municipality="Smithsonian Institution" pageId="35" pageNumber="34" specimenCount="1" specimenCount-male="1" stateProvince="Washington" typeStatus="holotype">
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<emphasis id="B95C6C3DFFFCBB73C17FF81CE2BCB506" bold="true" box="[373,489,1866,1892]" pageId="35" pageNumber="34">
|
||
<typeStatus id="54930E8DFFFCBB73C17FF81CE2B1B506" box="[373,484,1866,1892]" pageId="35" pageNumber="34" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
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||
<specimenCount id="9D2E7BA6FFFCBB73C1F9F81DE15DB505" box="[499,520,1867,1895]" count="1" pageId="35" pageNumber="34" type="male">♂</specimenCount>
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||
deposited in the
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||
<location id="8EF7E6F4FFFCBB73C2DFF81CE72AB506" LSID="urn:lsid:plazi:treatment:03810139FFFCBB74C0CAFB07E192B028:8EF7E6F4FFFCBB73C2DFF81CE72AB506" box="[725,1151,1866,1892]" country="United States of America" municipality="Smithsonian Institution" name="National Museum of Natural History" pageId="35" pageNumber="34" stateProvince="Washington">National Museum of Natural History</location>
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||
,
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||
<collectingMunicipality id="6BF32A55FFFCBB73C486F81CE6C9B506" box="[1164,1436,1866,1892]" pageId="35" pageNumber="34">Smithsonian Institution</collectingMunicipality>
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,
|
||
<collectingRegion id="49EC7ECDFFFCBB73C0CAF83AE21CB5E4" box="[192,329,1900,1926]" country="United States of America" name="Washington" pageId="35" pageNumber="34">Washington</collectingRegion>
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||
,
|
||
<collectionCode id="ED3928EAFFFCBB73C15EF83AE22BB5E4" box="[340,382,1900,1926]" pageId="35" pageNumber="34">DC</collectionCode>
|
||
,
|
||
<collectingCountry id="F33FF0BFFFFCBB73C183F83AE2EBB5E4" box="[393,446,1900,1926]" name="United States of America" pageId="35" pageNumber="34">USA</collectingCountry>
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(
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||
<collectionCode id="ED3928EAFFFCBB73C1C6F83AE174B5E4" box="[460,545,1900,1926]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="35" pageNumber="34" type="Museum">USNM</collectionCode>
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), illustrated in
|
||
<figureCitation id="1313ACAAFFFCBB73C2C3F83AE015B5E4" box="[713,832,1900,1926]" captionStart="Figures 63–84" captionStartId="98.[192,271,1810,1834]" captionTargetBox="[196,1434,233,1782]" captionTargetId="figure-99@98.[192,1440,229,1789]" captionTargetPageId="98" captionText="Figures 63–84. Holotypes, data in text. 63–64) Myrinia aragua sp. n. 65–66) Myrinia maculosa sp. n. 67–68) Myrinia manchada sp. n. 69–70) Polyctor (Fenops) lamperus sp. n. 71–72) Nisoniades (Nisoniades) lutum sp. n. 73–74) Bolla (Stolla) vena sp. n. 75–76) Staphylus (Vulga) vula sp. n. 77–78) Staphylus (Vulga) vulga sp. n. 79–80) Staphylus (Staphylus) rotundalus sp. n. 81–82) Staphylus (Staphylus) yucatanus sp. n. 83–84) Heliopetes (Heliopetes) lana sp. n." pageId="35" pageNumber="34">Fig. 77–78</figureCitation>
|
||
, bears the following five rectangular labels, four white:
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B97B02FFFFBBB74C0CAFFB0E143B3EA" blockId="36.[191,1440,230,392]" pageId="36" pageNumber="35">
|
||
[
|
||
<collectingCountry id="F33FF0BFFFFBBB74C0DBFFB0E219B363" box="[209,332,230,257]" name="Panama" pageId="36" pageNumber="35">PANAMA</collectingCountry>
|
||
:
|
||
<specimenCount id="9D2E7BA6FFFBBB74C145FFB0E2C7B363" box="[335,402,230,257]" count="500" pageId="36" pageNumber="35" type="generic">
|
||
<quantity id="4CD01DCAFFFBBB74C145FFB0E2DBB363" box="[335,398,230,257]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="35" unit="m" value="500.0">500m</quantity>
|
||
.
|
||
</specimenCount>
|
||
|
|
||
<collectingRegion id="49EC7ECDFFFBBB74C1ADFFB0E2A2B362" box="[423,503,230,256]" country="Panama" name="Darien" pageId="36" pageNumber="35">Darien</collectingRegion>
|
||
, Cana |
|
||
<date id="FF9696EFFFFBBB74C259FFB0E19AB362" box="[595,719,230,256]" pageId="36" pageNumber="35" value="1984-01-03">3 Jan. 1984</date>
|
||
| Gordon Small], [
|
||
<taxonomicName id="4C28CBACFFFBBB74C3B8FFB0E74BB362" authorityName="Godman and Salvin" authorityYear="1896" box="[946,1054,230,256]" class="Insecta" family="Hesperiidae" genus="Staphylus" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="35" phylum="Arthropoda" rank="genus">Staphylus</taxonomicName>
|
||
<specimenCount id="9D2E7BA6FFFBBB74C42FFFB1E76FB361" box="[1061,1082,231,259]" count="1" pageId="36" pageNumber="35" type="male">♂</specimenCount>
|
||
| caribbea (Williams | and Bell) | det. H. A. Freeman], [DNA sample ID: | NVG-18058C11 | c/o Nick
|
||
<collectionCode id="ED3928EAFFFBBB74C3E8FE5EE0A0B340" box="[994,1013,264,290]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="36" pageNumber="35" type="Museum">V</collectionCode>
|
||
. Grishin], [USNMENT | {QR Code} | 01466697], and one red [
|
||
<typeStatus id="54930E8DFFFBBB74C1F7FE7CE1DBB326" box="[509,654,298,324]" pageId="36" pageNumber="35" type="holotype">HOLOTYPE</typeStatus>
|
||
<specimenCount id="9D2E7BA6FFFBBB74C29CFE7DE1FEB325" box="[662,683,299,327]" count="1" pageId="36" pageNumber="35" type="male">♂</specimenCount>
|
||
|
|
||
<taxonomicName id="4C28CBACFFFBBB74C2CAFE7CE07EB326" authorityName="Godman and Salvin" authorityYear="1896" box="[704,811,298,324]" class="Insecta" family="Hesperiidae" genus="Staphylus" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="35" phylum="Arthropoda" rank="genus">Staphylus</taxonomicName>
|
||
|
|
||
<taxonomicName id="4C28CBACFFFBBB74C349FE7CE088B326" authority="Grishin" authorityName="vulga Grishin" authorityYear="2023" box="[835,989,298,324]" class="Insecta" family="Hesperiidae" genus="Staphylus" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="35" phylum="Arthropoda" rank="species" species="vulga">vulga Grishin</taxonomicName>
|
||
].
|
||
<emphasis id="B95C6C3DFFFBBB74C3F4FE7CE738B326" bold="true" box="[1022,1133,298,324]" pageId="36" pageNumber="35">
|
||
<typeStatus id="54930E8DFFFBBB74C3F4FE7CE73DB326" box="[1022,1128,298,324]" pageId="36" pageNumber="35" type="paratype">Paratype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D2E7BA6FFFBBB74C479FE7CE7C3B325" box="[1139,1174,298,327]" count="1" pageId="36" pageNumber="35" type="male">1♂</specimenCount>
|
||
NVG-18058B12, USNMENT_01466686
|
||
<collectingRegion id="49EC7ECDFFFBBB74C184FE1AE2BEB304" box="[398,491,332,358]" country="Panama" name="Panama" pageId="36" pageNumber="35">Panama</collectingRegion>
|
||
:
|
||
<taxonomicName id="4C28CBACFFFBBB74C1FEFE1AE1F8B304" authority=", Bayano" authorityName="Bayano" box="[500,685,332,358]" class="Insecta" family="Hesperiidae" genus="Aguna" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="35" phylum="Arthropoda" rank="species" species="panama">Panama, Bayano</taxonomicName>
|
||
, “8°[likely 9°]03′N
|
||
<geoCoordinate id="EE1CD6E8FFFBBB74C38AFE1AE0B4B304" box="[896,993,332,360]" degrees="78" direction="west" minutes="40" orientation="longitude" pageId="36" pageNumber="35" precision="925" value="-78.666664">78°40′W</geoCoordinate>
|
||
”,
|
||
<date id="FF9696EFFFFBBB74C3FEFE1AE7D0B304" box="[1012,1157,332,358]" pageId="36" pageNumber="35" value="1974-10-26">26-Oct-1974</date>
|
||
, G. B. Small leg., genitalia vial H-2198 by H. A. Freeman.
|
||
</paragraph>
|
||
<paragraph id="8B97B02FFFFBBB74C0CAFECEE0DFB3D0" blockId="36.[192,906,407,434]" box="[192,906,407,434]" pageId="36" pageNumber="35">
|
||
<emphasis id="B95C6C3DFFFBBB74C0CAFECEE20EB3D3" bold="true" box="[192,347,407,434]" pageId="36" pageNumber="35">
|
||
<typeStatus id="54930E8DFFFBBB74C0CAFECEE3ACB3D0" box="[192,249,408,434]" pageId="36" pageNumber="35">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="F33FF0BFFFFBBB74C16BFECEE2EBB3D0" box="[353,446,408,434]" name="Panama" pageId="36" pageNumber="35">Panama</collectingCountry>
|
||
:
|
||
<collectingRegion id="49EC7ECDFFFBBB74C1C2FECEE1D4B3D0" box="[456,641,408,434]" country="Panama" name="Darien" pageId="36" pageNumber="35">Darien Province</collectingRegion>
|
||
, Cana, elevation
|
||
<specimenCount id="9D2E7BA6FFFBBB74C334FEC1E0DFB3D0" box="[830,906,407,434]" count="500" pageId="36" pageNumber="35" type="generic">
|
||
<quantity id="4CD01DCAFFFBBB74C334FEC1E0D2B3D0" box="[830,903,407,434]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="35" unit="m" value="500.0">500 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C332E3A4FFFBBB74C0CAFE94E158B042" pageId="36" pageNumber="35" type="etymology">
|
||
<paragraph id="8B97B02FFFFBBB74C0CAFE94E158B042" blockId="36.[192,1440,450,544]" pageId="36" pageNumber="35">
|
||
<emphasis id="B95C6C3DFFFBBB74C0CAFE94E211B3BE" bold="true" box="[192,324,450,476]" pageId="36" pageNumber="35">Etymology.</emphasis>
|
||
The name is formed from the species epithet of
|
||
<taxonomicName id="4C28CBACFFFBBB74C37CFE94E0B4B3BE" baseAuthorityName="Moschler" baseAuthorityYear="1879" box="[886,993,450,476]" class="Insecta" family="Hesperiidae" genus="Staphylus" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="35" phylum="Arthropoda" rank="species" species="vulgata">
|
||
<emphasis id="B95C6C3DFFFBBB74C37CFE94E0B4B3BE" box="[886,993,450,476]" italics="true" pageId="36" pageNumber="35">S. vulgata</emphasis>
|
||
</taxonomicName>
|
||
(in Latin,
|
||
<emphasis id="B95C6C3DFFFBBB74C455FE94E7FBB3BE" box="[1119,1198,450,476]" italics="true" pageId="36" pageNumber="35">vulgate</emphasis>
|
||
means common). By slightly shortening the name, it refers to this species in the middle of the range of the
|
||
<taxonomicName id="4C28CBACFFFBBB74C49FFEB2E657B39C" baseAuthorityName="Moschler" baseAuthorityYear="1879" box="[1173,1282,484,510]" class="Insecta" family="Hesperiidae" genus="Staphylus" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="35" phylum="Arthropoda" rank="species" species="vulgata">
|
||
<emphasis id="B95C6C3DFFFBBB74C49FFEB2E657B39C" box="[1173,1282,484,510]" italics="true" pageId="36" pageNumber="35">S. vulgata</emphasis>
|
||
</taxonomicName>
|
||
complex. The name is a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C332E3A4FFFBBB74C0CAFD79E192B028" box="[192,711,559,586]" pageId="36" pageNumber="35" type="distribution">
|
||
<paragraph id="8B97B02FFFFBBB74C0CAFD79E192B028" blockId="36.[192,711,559,586]" box="[192,711,559,586]" pageId="36" pageNumber="35">
|
||
<emphasis id="B95C6C3DFFFBBB74C0CAFD79E20EB02B" bold="true" box="[192,347,559,585]" pageId="36" pageNumber="35">Distribution.</emphasis>
|
||
Recorded from eastern
|
||
<collectingCountry id="F33FF0BFFFFBBB74C26DFD66E191B028" box="[615,708,560,586]" name="Panama" pageId="36" pageNumber="35">Panama</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |