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<document id="4B9E145F2D862C6F005AEFD7EDBF47B6" ID-DOI="10.5281/zenodo.10396362" ID-ISSN="1942-1354" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1707168658757" checkinUser="felipe" docAuthor="Zhang, Jing, Cong, Qian &amp; Grishin, Nick V." docDate="2023" docId="03810139FFEFBB60C0CAFE20E775B7BD" docLanguage="en" docName="InsectaMundi.2023.1026.1-115.pdf" docOrigin="Insecta Mundi 2023 (26)" docSource="http://dx.doi.org/10.5281/zenodo.10396362" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Alychna ayonis Zhang &amp; Cong &amp; Grishin 2023, new species" docType="treatment" docVersion="1" lastPageNumber="47" masterDocId="FFB87941FFDFBB50C00AFF56E355B262" masterDocTitle="Supplementary Materials and Appendix" masterLastPageNumber="115" masterPageNumber="1" pageNumber="47" updateTime="1707169288913" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="7F66E3D2A6137DCDC8E6C373F1B809D3">
<mods:title id="5CE5AAC6F4306BC8804574CECE42A5F1">Supplementary Materials and Appendix</mods:title>
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<mods:name id="36BB123D8531BB9DCAE2C89E6EBF128E" type="personal">
<mods:role id="C549A46F13322B82574C37B8D35E71F7">
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<mods:namePart id="767A4C67FAA8584764FBBE7ABFB538F3">Zhang, Jing</mods:namePart>
<mods:affiliation id="9C227331F47B3021D86553181A3DBA02">McDermott Center for Human Growth and Development and Department of Biophysics University of Texas Southwestern Medical Center 5323 Harry Hines Blvd., Dallas, TX, 75390 - 8816 USA</mods:affiliation>
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<mods:namePart id="2DACEF5A647BC9FB8ED97C836C276CEC">Cong, Qian</mods:namePart>
<mods:affiliation id="069D5E603C824D3D4318A7F23BD054AC">McDermott Center for Human Growth and Development and Department of Biophysics University of Texas Southwestern Medical Center 5323 Harry Hines Blvd., Dallas, TX, 75390 - 8816 USA</mods:affiliation>
</mods:name>
<mods:name id="009E1260B9E5B6044F31B767CF6EF4BD" type="personal">
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</mods:role>
<mods:namePart id="9C03C52AC179398D42F3E9C1B1A3DBDF">Grishin, Nick V.</mods:namePart>
<mods:affiliation id="213B602D013F99F7FDF784DBC4098B61">Departments of Biophysics and Biochemistry University of Texas Southwestern Medical Center 5323 Harry Hines Blvd., Dallas, TX, 75390 - 9050 USA</mods:affiliation>
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<mods:title id="63D9B75DEE75DD5CCA665031CE441DA7">Insecta Mundi</mods:title>
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<mods:date id="C5B2ECC8BC8D21EDAC114946FB08CEC9">2023</mods:date>
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<mods:identifier id="8E7A269E9B76CD2D3B2F7DDAB1DF9AC4" type="DOI">10.5281/zenodo.10396362</mods:identifier>
<mods:identifier id="65E29217CBAE95FD0CA381D16C92684E" type="ISSN">1942-1354</mods:identifier>
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<treatment id="03810139FFEFBB60C0CAFE20E775B7BD" LSID="urn:lsid:plazi:treatment:03810139FFEFBB60C0CAFE20E775B7BD" httpUri="http://treatment.plazi.org/id/03810139FFEFBB60C0CAFE20E775B7BD" lastPageNumber="47" pageId="48" pageNumber="47">
<subSubSection id="C332E3A4FFEFBB60C0CAFE20E0F2B3D4" pageId="48" pageNumber="47" type="nomenclature">
<paragraph id="8B97B02FFFEFBB60C0CAFE20E1D9B3F1" blockId="48.[192,935,374,472]" box="[192,652,374,403]" pageId="48" pageNumber="47">
<heading id="D0DF0743FFEFBB60C0CAFE20E1D9B3F1" box="[192,652,374,403]" fontSize="12" level="2" pageId="48" pageNumber="47" reason="2">
<emphasis id="B95C6C3DFFEFBB60C0CAFE20E1D9B3F1" bold="true" box="[192,652,374,403]" pageId="48" pageNumber="47">
<taxonomicName id="4C28CBACFFEFBB60C0CAFE20E2BEB3F1" authority="Grishin" authorityName="Zhang &amp; Cong &amp; Grishin" authorityYear="2023" box="[192,491,374,403]" class="Insecta" family="Hesperiidae" genus="Alychna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="ayonis" status="sp. nov.">
<emphasis id="B95C6C3DFFEFBB60C0CAFE20E2D5B3F1" bold="true" box="[192,384,374,403]" italics="true" pageId="48" pageNumber="47">Alychna ayonis</emphasis>
Grishin
</taxonomicName>
,
<taxonomicNameLabel id="A26FD146FFEFBB60C1FDFE21E1D9B3F1" box="[503,652,374,403]" pageId="48" pageNumber="47" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<paragraph id="8B97B02FFFEFBB60C0CAFECDE0F2B3D4" blockId="48.[192,935,374,472]" box="[192,935,411,438]" pageId="48" pageNumber="47">
https://zoobank.org/
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</paragraph>
</subSubSection>
<subSubSection id="C332E3A4FFEFBB60C0CAFEE8E148B3BA" box="[192,541,445,472]" pageId="48" pageNumber="47" type="description">
<paragraph id="8B97B02FFFEFBB60C0CAFEE8E148B3BA" blockId="48.[192,935,374,472]" box="[192,541,445,472]" pageId="48" pageNumber="47">
(
<figureCitation id="1313ACAAFFEFBB60C0C2FEE8E252B3BA" box="[200,263,445,472]" captionStart="Figure 5" captionStartId="49.[192,261,1225,1249]" captionTargetBox="[193,1437,230,1172]" captionTargetId="figure-175@49.[192,1440,229,1175]" captionTargetPageId="49" captionText="Figure 5. Phylogenetic trees of Moncina (part 1) inferred from protein-coding regions in a) the Z chromosome and b) the mitochondrial genome (in segments). See Fig. 1 legend for other notations." pageId="48" pageNumber="47">Fig. 5</figureCitation>
part, 107108, 334335)
</paragraph>
</subSubSection>
<subSubSection id="C332E3A4FFEFBB60C0CAFEBEE7D8B601" pageId="48" pageNumber="47" type="diagnosis">
<paragraph id="8B97B02FFFEFBB60C0CAFEBEE70EB11C" blockId="48.[192,1441,488,895]" pageId="48" pageNumber="47">
<emphasis id="B95C6C3DFFEFBB60C0CAFEBEE2B2B060" bold="true" box="[192,487,488,514]" pageId="48" pageNumber="47">Definition and diagnosis.</emphasis>
Phylogenetic trees reveal that specimens from
<collectingCountry id="F33FF0BFFFEFBB60C3E7FEBEE71EB060" box="[1005,1099,488,514]" name="Ecuador" pageId="48" pageNumber="47">Ecuador</collectingCountry>
and
<collectingCountry id="F33FF0BFFFEFBB60C48AFEBFE7E1B061" box="[1152,1204,489,515]" name="Peru" pageId="48" pageNumber="47">Peru</collectingCountry>
identified as
<taxonomicName id="4C28CBACFFEFBB60C54CFEBEE152B047" authority="(Mabille, 1898)" baseAuthorityName="Mabille" baseAuthorityYear="1898" class="Insecta" family="Hesperiidae" genus="Alychna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="exclamationis">
<emphasis id="B95C6C3DFFEFBB60C54CFEBEE203B047" italics="true" pageId="48" pageNumber="47">Alychna exclamationis</emphasis>
(Mabille, 1898)
</taxonomicName>
(type locality in
<collectingCountry id="F33FF0BFFFEFBB60C2C8FD5DE046B047" box="[706,787,523,549]" name="Bolivia" pageId="48" pageNumber="47">Bolivia</collectingCountry>
,
<typeStatus id="54930E8DFFEFBB60C317FD5DE0D6B047" box="[797,899,523,549]" pageId="48" pageNumber="47" type="lectotype">lectotype</typeStatus>
sequenced as NVG-18042G08) show prominent genetic differentiation from it while being its sister (
<figureCitation id="1313ACAAFFEFBB60C303FD7BE01FB02A" box="[777,842,557,584]" captionStart="Figure 5" captionStartId="49.[192,261,1225,1249]" captionTargetBox="[193,1437,230,1172]" captionTargetId="figure-175@49.[192,1440,229,1175]" captionTargetPageId="49" captionText="Figure 5. Phylogenetic trees of Moncina (part 1) inferred from protein-coding regions in a) the Z chromosome and b) the mitochondrial genome (in segments). See Fig. 1 legend for other notations." pageId="48" pageNumber="47">Fig. 5</figureCitation>
): e.g., their COI barcodes differ by 1.5% (10 bp) and, therefore, represent a new species, more distantly related to
<taxonomicName id="4C28CBACFFEFBB60C354FD06E7F4B008" authority="(E. Bell, 1942)" baseAuthorityName="E. Bell" baseAuthorityYear="1942" box="[862,1185,592,618]" class="Insecta" family="Hesperiidae" genus="Alychna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="zenus">
<emphasis id="B95C6C3DFFEFBB60C354FD06E0A8B008" box="[862,1021,592,618]" italics="true" pageId="48" pageNumber="47">Alychna zenus</emphasis>
(E. Bell, 1942)
</taxonomicName>
(type locality in
<collectingCountry id="F33FF0BFFFEFBB60C555FD06E3BCB0EE" name="Ecuador" pageId="48" pageNumber="47">Ecuador</collectingCountry>
) (COI barcodes differ by 3.5%, 23 bp). This new species keys to “
<taxonomicName id="4C28CBACFFEFBB60C3D7FD24E785B0EE" baseAuthorityName="Mabille" baseAuthorityYear="1897" box="[989,1232,626,652]" class="Insecta" family="Hesperiidae" genus="Psoralis" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="exclamationis">
<emphasis id="B95C6C3DFFEFBB60C3D7FD24E785B0EE" box="[989,1232,626,652]" italics="true" pageId="48" pageNumber="47">Psoralis exclamationis</emphasis>
</taxonomicName>
” (J.43.3) in
<bibRefCitation id="EFB9CDDEFFEFBB60C554FD25E25CB0D2" author="Evans WH" pageId="48" pageNumber="47" pagination="54 - 88" refId="ref77169" refString="Evans WH. 1955. A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. The Trustees of the British Museum (Natural History); London. v + 499 p., pl. 54 - 88." type="book chapter" year="1955">Evans (1955)</bibRefCitation>
but differs from it by narrower stigma, narrower white spots and streaks on forewing, a semi-circle of discal pale dots on ventral hindwing (
<figureCitation id="1313ACAAFFEFBB60C26CFDE1E1A9B0B0" box="[614,764,695,722]" captionStart="Figures 105122" captionStartId="100.[192,271,1841,1865]" captionTargetBox="[195,1432,334,1727]" captionTargetId="figure-77@100.[192,1440,327,1735]" captionTargetPageId="100" captionText="Figures 105122. Holotypes, data in text.105106) Psoralis (Saniba) magnamacus sp. n.107108) Alychna ayonis sp. n.109 110) Wahydra banios sp. n. 111112) Wahydra cuzcona sp. n. 113114) Cynea (Cynea) aureofimbra sp. n. 115116) Cynea (Nycea) quada sp. n. 117118) Cynea (Quinta) achirae sp. n. 119120) Eutus amazonicus sp. n. 121122) Eutus incus sp. n." pageId="48" pageNumber="47">Fig. 107108</figureCitation>
), undivided uncus, harpe strongly upcurved and protruding beyond ampulla dorsad, terminally rounded in lateral view and plate-like in dorsal view, costa-ampulla with a bump closer to ampulla, concave on both sides of the bump (
<figureCitation id="1313ACAAFFEFBB60C388FDABE74FB175" box="[898,1050,764,791]" captionStart="Figures 334360" captionStartId="112.[192,271,1750,1774]" captionTargetBox="[208,1419,269,1711]" captionTargetId="figure-128@112.[192,1440,247,1720]" captionTargetPageId="112" captionText="Figures 334360. Genitalia of holotypes (unless indicated), data in text. 334335) Alychna ayonis sp. n. 336338) Wahydra banios sp. n. 339341) Wahydra cuzcona sp. n. 342344) Cynea (Cynea) aureofimbra sp. n. 345347) Cynea (Nycea) quada sp. n. 348349) Cynea (Quinta) achirae sp. n. paratype ♂ NVG-22035B12.350352) Eutus amazonicus sp. n. 353354) Eutus incus sp. n. 355356) Eutus septemaculatus sp. n. 357358) Godmia viridicapita sp. n. 359360) Rhomba pulla sp. n. Views: dorsal (334, 337, 340, 342, 346, 348, 350, 353, 355, 358, 359), left lateral (335, 338, 341, 344, 347, 349, 352, 354, 356, 358, 360), right lateral of left valva (343, 345, 351), dorsoanterior (336, 339)." pageId="48" pageNumber="47">Fig. 334335</figureCitation>
). Due to the cryptic nature of this species, most reliable identification is achieved by DNA and a combination of the following base pairs is diagnostic in the nuclear genome: aly420.62.5:G73T, aly420.62.5:G84A, aly
<date id="FF9696EFFFEFBB60C3E0FC14E71BB13F" box="[1002,1102,834,861]" pageId="48" pageNumber="47" value="2041-08-02">2041.8.2</date>
:A195C, aly2012.47.1:G1269C, aly25.10.1:G48C, and COI barcode: A85T, T163A, T169A, T205A, A553C, A631G.
</paragraph>
<paragraph id="8B97B02FFFEFBB60C0CAFCD9E7D8B601" blockId="48.[192,1440,911,1123]" pageId="48" pageNumber="47">
<emphasis id="B95C6C3DFFEFBB60C0CAFCD9E11BB1CB" bold="true" box="[192,590,911,937]" pageId="48" pageNumber="47">
Barcode sequence of the
<typeStatus id="54930E8DFFEFBB60C1EAFCD9E11CB1CB" box="[480,585,911,937]" pageId="48" pageNumber="47" type="holotype">holotype</typeStatus>
.
</emphasis>
Sample NVG-22035H07, GenBank OR837672, 658 base pairs: AACTTTATATTTTATTTTTGGTATTTGAGCAGGTATACTAGGAACTTCATTAAGTTTATTAATTCGTACAGAATTAGGAAATCCTGGATCTTTAATT GGAGATGATCAAATTTATAATACTATTGTAACAGCTCATGCTTTTATTATAATTTTTTTTATAGTAATACCAATTATAATTGGAGGATTTGGAAATT GATTAGTTCCATTAATATTAGGAGCTCCTGATATAGCTTTCCCACGAATAAATAATATAAGATTTTGAATATTACCCCCTTCCTTAATATTATTAAT TTCAAGAAGAATTGTTGAAAATGGTGCAGGTACTGGATGAACTGTTTATCCCCCCCTTTCTTCTAATATCGCACACCAAGGATCATCTGTTGATTTA GCAATTTTTTCTTTACATTTAGCTGGGATCTCTTCTATTTTAGGAGCTATTAATTTTATTACTACAATTATTAATATACGAATTAGAAATATATCCT TTGATCAAATACCTTTATTTGTATGATCTGTAGGAATTACAGCTTTATTATTACTTTTATCTTTACCCGTATTAGCTGGAGCTATTACAATACTTTT AACTGATCGAAACTTAAATACTTCTTTTTTTGATCCAGCTGGAGGAGGGGACCCTATCTTATATCAACATTTATTT
</paragraph>
</subSubSection>
<subSubSection id="C332E3A4FFEFBB60C0CAFB20E091B727" pageId="48" pageNumber="47" type="materials_examined">
<paragraph id="8B97B02FFFEFBB60C0CAFB20E670B778" blockId="48.[192,1440,1142,1307]" pageId="48" pageNumber="47">
<emphasis id="B95C6C3DFFEFBB60C0CAFB20E239B6F2" bold="true" box="[192,364,1142,1168]" pageId="48" pageNumber="47">Type material.</emphasis>
<materialsCitation id="3B40BA72FFEFBB60C17FFB20E76BB69A" collectionCode="DC, USNM, V" collectorName="Sept. &amp; S. S. Nicolay &amp; Nick &amp; Grishin" country="Ecuador" elevation="2800" location="Smithsonian Institution" municipality="National Museum of Natural History" pageId="48" pageNumber="47" specimenCount="2" specimenCount-male="2" stateProvince="Napo" typeStatus="holotype">
<emphasis id="B95C6C3DFFEFBB60C17FFB20E2BCB6F2" bold="true" box="[373,489,1142,1168]" pageId="48" pageNumber="47">
<typeStatus id="54930E8DFFEFBB60C17FFB20E2B1B6F2" box="[373,484,1142,1168]" pageId="48" pageNumber="47" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D2E7BA6FFEFBB60C1F9FB21E15DB6F1" box="[499,520,1143,1171]" count="1" pageId="48" pageNumber="47" type="male"></specimenCount>
deposited in the
<collectingMunicipality id="6BF32A55FFEFBB60C2DFFB20E72AB6F2" box="[725,1151,1142,1168]" pageId="48" pageNumber="47">National Museum of Natural History</collectingMunicipality>
,
<location id="8EF7E6F4FFEFBB60C486FB20E6C9B6F2" LSID="urn:lsid:plazi:treatment:03810139FFEFBB60C0CAFE20E775B7BD:8EF7E6F4FFEFBB60C486FB20E6C9B6F2" box="[1164,1436,1142,1168]" country="Ecuador" municipality="National Museum of Natural History" name="Smithsonian Institution" pageId="48" pageNumber="47" stateProvince="Napo">Smithsonian Institution</location>
, Washington,
<collectionCode id="ED3928EAFFEFBB60C15DFBCFE2D4B6D1" box="[343,385,1177,1203]" pageId="48" pageNumber="47">DC</collectionCode>
, USA (
<collectionCode id="ED3928EAFFEFBB60C1DEFBCFE17CB6D1" box="[468,553,1177,1203]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="48" pageNumber="47" type="Museum">USNM</collectionCode>
), illustrated in
<figureCitation id="1313ACAAFFEFBB60C2D2FBCEE039B6D1" box="[728,876,1176,1203]" captionStart="Figures 105122" captionStartId="100.[192,271,1841,1865]" captionTargetBox="[195,1432,334,1727]" captionTargetId="figure-77@100.[192,1440,327,1735]" captionTargetPageId="100" captionText="Figures 105122. Holotypes, data in text.105106) Psoralis (Saniba) magnamacus sp. n.107108) Alychna ayonis sp. n.109 110) Wahydra banios sp. n. 111112) Wahydra cuzcona sp. n. 113114) Cynea (Cynea) aureofimbra sp. n. 115116) Cynea (Nycea) quada sp. n. 117118) Cynea (Quinta) achirae sp. n. 119120) Eutus amazonicus sp. n. 121122) Eutus incus sp. n." pageId="48" pageNumber="47">Fig. 107108</figureCitation>
, bears the following three rectangular labels, two white: [
<collectingCountry id="F33FF0BFFFEFBB60C110FBEDE2C8B6B7" box="[282,413,1211,1237]" name="Ecuador" pageId="48" pageNumber="47">ECUADOR</collectingCountry>
<collectingRegion id="49EC7ECDFFEFBB60C1AEFBEAE2B4B6B4" box="[420,481,1212,1238]" country="Ecuador" name="Napo" pageId="48" pageNumber="47">Napo</collectingRegion>
| Papallacta
<quantity id="4CD01DCAFFEFBB60C264FBEDE1EEB6B7" box="[622,699,1211,1237]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="48" pageNumber="47" unit="m" value="2800.0">
<elevation id="0005571CFFEFBB60C264FBEDE1EEB6B7" box="[622,699,1211,1237]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="48" pageNumber="47" unit="m" value="2800.0">2800m</elevation>
</quantity>
| 23
<collectorName id="26DDD5F9FFEFBB60C2E5FBEDE072B6B7" box="[751,807,1211,1237]" pageId="48" pageNumber="47">Sept.</collectorName>
87 |
<collectorName id="26DDD5F9FFEFBB60C368FBEDE0BEB6B7" box="[866,1003,1211,1237]" pageId="48" pageNumber="47">S. S. Nicolay</collectorName>
], [DNA sample ID: | NVG-22035H07 | c/o
<collectorName id="26DDD5F9FFEFBB60C0FDFB8BE279B695" box="[247,300,1245,1271]" pageId="48" pageNumber="47">Nick</collectorName>
<collectionCode id="ED3928EAFFEFBB60C138FB88E210B69A" box="[306,325,1246,1272]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="48" pageNumber="47" type="Museum">V</collectionCode>
.
<collectorName id="26DDD5F9FFEFBB60C144FB8BE2F1B695" box="[334,420,1245,1271]" pageId="48" pageNumber="47">Grishin</collectorName>
], and one red [
<typeStatus id="54930E8DFFEFBB60C250FB88E1BEB69A" box="[602,747,1246,1272]" pageId="48" pageNumber="47" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D2E7BA6FFEFBB60C2FBFB89E053B699" box="[753,774,1247,1275]" count="1" pageId="48" pageNumber="47" type="male"></specimenCount>
|
<taxonomicName id="4C28CBACFFEFBB60C310FB8BE022B695" authorityName="Grishin" authorityYear="2019" box="[794,887,1245,1271]" class="Insecta" family="Hesperiidae" genus="Alychna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="47" phylum="Arthropoda" rank="genus">Alychna</taxonomicName>
|
<taxonomicName id="4C28CBACFFEFBB60C380FB8BE77BB695" authority="Grishin" authorityName="Zhang &amp; Cong &amp; Grishin" authorityYear="2023" box="[906,1070,1245,1271]" class="Insecta" family="Hesperiidae" genus="Alychna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="ayonis">ayonis Grishin</taxonomicName>
]
</materialsCitation>
.
<emphasis id="B95C6C3DFFEFBB60C440FB88E7ECB69A" bold="true" box="[1098,1209,1246,1272]" pageId="48" pageNumber="47">
<typeStatus id="54930E8DFFEFBB60C440FB88E7E1B69A" box="[1098,1204,1246,1272]" pageId="48" pageNumber="47" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D2E7BA6FFEFBB60C4B5FB88E7B4B699" box="[1215,1249,1246,1275]" count="1" pageId="48" pageNumber="47" type="male">1♂</specimenCount>
NVG-19021H01 the same data as the
<typeStatus id="54930E8DFFEFBB60C1ACFA56E15DB778" box="[422,520,1280,1306]" pageId="48" pageNumber="47" type="holotype">holotype</typeStatus>
but collected on
<date id="FF9696EFFFEFBB60C2CDFA56E003B778" box="[711,854,1280,1306]" pageId="48" pageNumber="47" value="1987-09-17">17-Sep-1987</date>
, genitalia H962 by S. S. Nicolay [
<collectionCode id="ED3928EAFFEFBB60C4C8FA56E64FB778" box="[1218,1306,1280,1306]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="48" pageNumber="47" type="Museum">USNM</collectionCode>
].
</paragraph>
<paragraph id="8B97B02FFFEFBB60C0CAFA7DE091B727" blockId="48.[192,964,1322,1349]" box="[192,964,1322,1349]" pageId="48" pageNumber="47">
<emphasis id="B95C6C3DFFEFBB60C0CAFA7DE20EB726" bold="true" box="[192,347,1322,1349]" pageId="48" pageNumber="47">
<typeStatus id="54930E8DFFEFBB60C0CAFA7DE3ACB727" box="[192,249,1323,1349]" pageId="48" pageNumber="47">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F33FF0BFFFEFBB60C16BFA7DE296B727" box="[353,451,1323,1349]" name="Ecuador" pageId="48" pageNumber="47">Ecuador</collectingCountry>
:
<collectingRegion id="49EC7ECDFFEFBB60C1C7FA7DE123B727" box="[461,630,1323,1349]" country="Ecuador" name="Napo" pageId="48" pageNumber="47">Napo Province</collectingRegion>
, Papallacta, elevation
<quantity id="4CD01DCAFFEFBB60C360FA7DE095B727" box="[874,960,1323,1349]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="48" pageNumber="47" unit="m" value="2800.0">2800 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C332E3A4FFEFBB60C0CAFA03E74CB7D6" pageId="48" pageNumber="47" type="etymology">
<paragraph id="8B97B02FFFEFBB60C0CAFA03E74CB7D6" blockId="48.[192,1440,1365,1461]" pageId="48" pageNumber="47">
<emphasis id="B95C6C3DFFEFBB60C0CAFA03E211B70D" bold="true" box="[192,324,1365,1391]" pageId="48" pageNumber="47">Etymology.</emphasis>
The name is given for the “i” on the forewing. A close relative of this species is named exclamati-
<emphasis id="B95C6C3DFFEFBB60C567FA00E6CCB70D" box="[1389,1433,1366,1391]" italics="true" pageId="48" pageNumber="47">onis</emphasis>
, probably for the exclamation mark pattern on the forewing. In the new species, the exclamation mark is thinner and resembles the letter “i” [
<emphasis id="B95C6C3DFFEFBB60C1F6FACDE140B7D6" box="[508,533,1435,1460]" italics="true" pageId="48" pageNumber="47">ay</emphasis>
]. The name is treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C332E3A4FFEFBB60C0CAFA93E775B7BD" box="[192,1056,1477,1503]" pageId="48" pageNumber="47" type="distribution">
<paragraph id="8B97B02FFFEFBB60C0CAFA93E775B7BD" blockId="48.[192,1056,1477,1503]" box="[192,1056,1477,1503]" pageId="48" pageNumber="47">
<emphasis id="B95C6C3DFFEFBB60C0CAFA93E20EB7BD" bold="true" box="[192,347,1477,1503]" pageId="48" pageNumber="47">Distribution.</emphasis>
Currently known only from the
<typeStatus id="54930E8DFFEFBB60C2CDFA93E1A2B7BD" box="[711,759,1477,1503]" pageId="48" pageNumber="47">type</typeStatus>
locality in
<collectingRegion id="49EC7ECDFFEFBB60C379FA93E0E4B7BD" box="[883,945,1477,1503]" country="Ecuador" name="Napo" pageId="48" pageNumber="47">Napo</collectingRegion>
,
<collectingCountry id="F33FF0BFFFEFBB60C3B6FA93E749B7BD" box="[956,1052,1477,1503]" name="Ecuador" pageId="48" pageNumber="47">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>