309 lines
36 KiB
XML
309 lines
36 KiB
XML
<document id="A376E6E1B551ADFAA351253216F02EC4" ID-DOI="10.5281/zenodo.10396362" ID-ISSN="1942-1354" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1707168658757" checkinUser="felipe" docAuthor="Zhang, Jing, Cong, Qian & Grishin, Nick V." docDate="2023" docId="03810139FFCDBB42C0CAFD97E1DAB5E4" docLanguage="en" docName="InsectaMundi.2023.1026.1-115.pdf" docOrigin="Insecta Mundi 2023 (26)" docSource="http://dx.doi.org/10.5281/zenodo.10396362" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Ridens angulinea Zhang & Cong & Grishin 2023, new species" docType="treatment" docVersion="1" lastPageNumber="17" masterDocId="FFB87941FFDFBB50C00AFF56E355B262" masterDocTitle="Supplementary Materials and Appendix" masterLastPageNumber="115" masterPageNumber="1" pageNumber="17" updateTime="1707169288913" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="F1768BB0E3B46F9C8F124D9196D5B18B">
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<mods:title id="3DBD92BB91AC30A1973A4A4434C6AE8B">Supplementary Materials and Appendix</mods:title>
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<mods:name id="7EE81CBB3CE95A9BDD926003240085AD" type="personal">
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<mods:roleTerm id="0F8973D32B85B7001115F38618177CE3">Author</mods:roleTerm>
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<mods:namePart id="D2D4775130B638D5971FE24F1C99DB51">Zhang, Jing</mods:namePart>
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<mods:affiliation id="D2C74271601DF4B14B89A14576FEBD3B">McDermott Center for Human Growth and Development and Department of Biophysics University of Texas Southwestern Medical Center 5323 Harry Hines Blvd., Dallas, TX, 75390 - 8816 USA</mods:affiliation>
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<mods:roleTerm id="578894E935C5E9B42EEF6F2E78C28F2A">Author</mods:roleTerm>
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<mods:namePart id="1B421BDE86F07BD8C480281471B1EFFD">Cong, Qian</mods:namePart>
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<mods:affiliation id="37080012F57740A5F7355C2AEB02124D">McDermott Center for Human Growth and Development and Department of Biophysics University of Texas Southwestern Medical Center 5323 Harry Hines Blvd., Dallas, TX, 75390 - 8816 USA</mods:affiliation>
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<mods:roleTerm id="0AF54785F505CFEED69C31368BB26BF7">Author</mods:roleTerm>
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<mods:namePart id="8F28FA0666F56F73EF554FF6A6173A2D">Grishin, Nick V.</mods:namePart>
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<mods:affiliation id="C7F76DCBDDA2DBBB53AD077EF78E8650">Departments of Biophysics and Biochemistry University of Texas Southwestern Medical Center 5323 Harry Hines Blvd., Dallas, TX, 75390 - 9050 USA</mods:affiliation>
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<mods:typeOfResource id="F43FE15CF74C81FF6BA9C1ED43CC64D6">text</mods:typeOfResource>
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<mods:title id="8ECB2804F26C1C63C09EDE7C593CD19D">Insecta Mundi</mods:title>
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<mods:part id="2BBA25411CD787BCB5556D84257FD6F9">
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<mods:date id="1FD08BE8D0D324F9614CDCA282F357F5">2023</mods:date>
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<mods:number id="8A44C6B9D2743E2F999F55CCF311FA38">2023-12-29</mods:number>
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<mods:number id="DDD3DEFF015480730436EA6512EA6D61">2023</mods:number>
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<mods:number id="14C3FD31A2F3782636504D60ED5A2C46">26</mods:number>
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<treatment id="03810139FFCDBB42C0CAFD97E1DAB5E4" LSID="urn:lsid:plazi:treatment:03810139FFCDBB42C0CAFD97E1DAB5E4" httpUri="http://treatment.plazi.org/id/03810139FFCDBB42C0CAFD97E1DAB5E4" lastPageNumber="17" pageId="18" pageNumber="17">
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<subSubSection id="C332E3A4FFCDBB42C0CAFD97E0EFB162" pageId="18" pageNumber="17" type="nomenclature">
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<paragraph id="8B97B02FFFCDBB42C0CAFD97E1F2B0BC" blockId="18.[192,954,705,803]" box="[192,679,705,734]" pageId="18" pageNumber="17">
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<heading id="D0DF0743FFCDBB42C0CAFD97E1F2B0BC" box="[192,679,705,734]" fontSize="12" level="2" pageId="18" pageNumber="17" reason="2">
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<emphasis id="B95C6C3DFFCDBB42C0CAFD97E1F2B0BC" bold="true" box="[192,679,705,734]" pageId="18" pageNumber="17">
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<taxonomicName id="4C28CBACFFCDBB42C0CAFD97E153B0BC" authority="Grishin" authorityName="Zhang & Cong & Grishin" authorityYear="2023" box="[192,518,705,734]" class="Insecta" family="Hesperiidae" genus="Ridens" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="angulinea" status="sp. nov.">
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<emphasis id="B95C6C3DFFCDBB42C0CAFD97E2CEB0BC" bold="true" box="[192,411,705,734]" italics="true" pageId="18" pageNumber="17">Ridens angulinea</emphasis>
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Grishin
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26FD146FFCDBB42C218FD97E1F2B0BC" box="[530,679,705,734]" pageId="18" pageNumber="17" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B97B02FFFCDBB42C0CAFDB0E0EFB162" blockId="18.[192,954,705,803]" box="[192,954,742,768]" pageId="18" pageNumber="17">
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https://zoobank.org/
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<uuid id="FF8E8AFAFFCDBB42C1AAFDB0E0EFB162" box="[416,954,742,768]" pageId="18" pageNumber="17">F16FA82A-8646-4B46-A2F2-7C7A8DDECC6B</uuid>
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</paragraph>
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</subSubSection>
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<subSubSection id="C332E3A4FFCDBB42C0CAFC5EE157B140" box="[192,514,776,803]" pageId="18" pageNumber="17" type="description">
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<paragraph id="8B97B02FFFCDBB42C0CAFC5EE157B140" blockId="18.[192,954,705,803]" box="[192,514,776,803]" pageId="18" pageNumber="17">
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(
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<figureCitation id="1313ACAAFFCDBB42C0C2FC5EE252B141" box="[200,263,776,803]" captionStart="Figure 1" captionStartId="8.[192,260,1506,1530]" captionTargetBox="[193,1439,222,1459]" captionTargetId="figure-13@8.[192,1440,219,1461]" captionTargetPageId="8" captionText="Figure 1. Phylogenetic trees of Eudaminae inferred from protein-coding regions in a) the Z chromosome; gaps in branches indicate places where vertical slices of the tree were removed to reduce its horizontal dimension (to allow an increase of the font size), i.e., branches with gaps, are longer than shown; and b) the mitochondrial genome (in segments): clades are cut from the tree and rearranged to match better the order of taxa in the Z chromosome tree and to save space. Statistical support values are shown by nodes.For each specimen, the name adopted in this work is given first, and a previously used name (or misidentification for the new species) is listed in square brackets (if different), supplemented with the DNA sample number, type status (see Materials and Methods for abbreviations), general locality, and year of collection. See Table S1 in the Supplemental file (Zhang et al. 2023a) or NCBI database entries for additional data about these specimens. Synonyms are given in parentheses preceded by “=”. The type status refers to this synonym if the synonym name is provided. The same notations are used throughout this work in other figures showing phylogenetic trees.Clades of new species proposed here are shown in red color, and their species epithets are highlighted in red. In other phylogenetic tree figures (e.g., Fig. 2), yellow highlight indicates taxonomic changes, e.g., transfer to a different genus (genus name highlighted) or change in species, subspecies, or synonym status (species epithet highlighted) and clades with taxonomic changes are shown in different colors. Green arrows point from the clade of the genus/ species where a species/synonym was previously placed to the clade of its placement proposed in this work." pageId="18" pageNumber="17">Fig. 1</figureCitation>
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part, 33–34, 246–247)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C332E3A4FFCDBB42C0CAFC60E7D8B7F7" pageId="18" pageNumber="17" type="diagnosis">
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<paragraph id="8B97B02FFFCDBB42C0CAFC60E240B6CE" blockId="18.[192,1440,822,1196]" pageId="18" pageNumber="17">
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<emphasis id="B95C6C3DFFCDBB42C0CAFC60E2B9B132" bold="true" box="[192,492,822,848]" pageId="18" pageNumber="17">Definition and diagnosis.</emphasis>
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Phylogenetic trees reveal that specimens from
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<collectingCountry id="F33FF0BFFFCDBB42C408FC61E762B133" box="[1026,1079,823,849]" name="Peru" pageId="18" pageNumber="17">Peru</collectingCountry>
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and
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<collectingCountry id="F33FF0BFFFCDBB42C465FC60E7EBB132" box="[1135,1214,822,848]" name="Bolivia" pageId="18" pageNumber="17">Bolivia</collectingCountry>
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identified as
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<taxonomicName id="4C28CBACFFCDBB42C55CFC60E11BB111" authority="(Herrich-Schaffer, 1869)" baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1869" class="Insecta" family="Hesperiidae" genus="Ridens" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="fulminans">
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<emphasis id="B95C6C3DFFCDBB42C55CFC60E27AB111" italics="true" pageId="18" pageNumber="17">Ridens fulminans</emphasis>
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(Herrich-Schäffer, 1869)
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</taxonomicName>
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(type locality not specified) are not monophyletic with it and are instead sister to several species of
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<taxonomicName id="4C28CBACFFCDBB42C1E7FC2DE191B1F4" authority="Evans, 1952" authorityName="Evans" authorityYear="1952" box="[493,708,891,918]" class="Insecta" family="Hesperiidae" genus="Ridens" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B95C6C3DFFCDBB42C1E7FC2DE162B1F7" box="[493,567,891,917]" italics="true" pageId="18" pageNumber="17">Ridens</emphasis>
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||
<bibRefCitation id="EFB9CDDEFFCDBB42C237FC2AE191B1F4" author="Evans WH" box="[573,708,891,918]" pageId="18" pageNumber="17" pagination="10 - 25" refId="ref77051" refString="Evans WH. 1952. A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part II. Pyrginae. Section I. The Trustees of the British Museum (Natural History); London. v + 178 p., pl. 10 - 25." type="book chapter" year="1952">Evans, 1952</bibRefCitation>
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||
</taxonomicName>
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||
(
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||
<figureCitation id="1313ACAAFFCDBB42C2D9FC2DE046B1F4" box="[723,787,891,918]" captionStart="Figure 1" captionStartId="8.[192,260,1506,1530]" captionTargetBox="[193,1439,222,1459]" captionTargetId="figure-13@8.[192,1440,219,1461]" captionTargetPageId="8" captionText="Figure 1. Phylogenetic trees of Eudaminae inferred from protein-coding regions in a) the Z chromosome; gaps in branches indicate places where vertical slices of the tree were removed to reduce its horizontal dimension (to allow an increase of the font size), i.e., branches with gaps, are longer than shown; and b) the mitochondrial genome (in segments): clades are cut from the tree and rearranged to match better the order of taxa in the Z chromosome tree and to save space. Statistical support values are shown by nodes.For each specimen, the name adopted in this work is given first, and a previously used name (or misidentification for the new species) is listed in square brackets (if different), supplemented with the DNA sample number, type status (see Materials and Methods for abbreviations), general locality, and year of collection. See Table S1 in the Supplemental file (Zhang et al. 2023a) or NCBI database entries for additional data about these specimens. Synonyms are given in parentheses preceded by “=”. The type status refers to this synonym if the synonym name is provided. The same notations are used throughout this work in other figures showing phylogenetic trees.Clades of new species proposed here are shown in red color, and their species epithets are highlighted in red. In other phylogenetic tree figures (e.g., Fig. 2), yellow highlight indicates taxonomic changes, e.g., transfer to a different genus (genus name highlighted) or change in species, subspecies, or synonym status (species epithet highlighted) and clades with taxonomic changes are shown in different colors. Green arrows point from the clade of the genus/ species where a species/synonym was previously placed to the clade of its placement proposed in this work." pageId="18" pageNumber="17">Fig. 1</figureCitation>
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), and therefore represent a new species. COI barcodes of this new species and a
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<typeStatus id="54930E8DFFCDBB42C1C1FCC9E177B1DB" box="[459,546,927,953]" pageId="18" pageNumber="17" type="syntype">syntype</typeStatus>
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of
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<taxonomicName id="4C28CBACFFCDBB42C241FCC9E18EB1DA" baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1869" box="[587,731,926,952]" class="Insecta" family="Hesperiidae" genus="Ridens" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="fulminans">
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||
<emphasis id="B95C6C3DFFCDBB42C241FCC9E18EB1DA" box="[587,731,926,952]" italics="true" pageId="18" pageNumber="17">R. fulminans</emphasis>
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</taxonomicName>
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differ by 6.8% (45 bp). The new species keys to
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<taxonomicName id="4C28CBACFFCDBB42C501FCC8E3A0B1B9" baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1869" class="Insecta" family="Hesperiidae" genus="Ridens" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="fulminans">
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||
<emphasis id="B95C6C3DFFCDBB42C501FCC8E3A0B1B9" italics="true" pageId="18" pageNumber="17">Ridens fulminans</emphasis>
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||
</taxonomicName>
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(C.12.7) in
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<bibRefCitation id="EFB9CDDEFFCDBB42C189FC94E14DB1BE" author="Evans WH" box="[387,536,961,988]" pageId="18" pageNumber="17" pagination="10 - 25" refId="ref77051" refString="Evans WH. 1952. A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part II. Pyrginae. Section I. The Trustees of the British Museum (Natural History); London. v + 178 p., pl. 10 - 25." type="book chapter" year="1952">Evans (1952)</bibRefCitation>
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and differs from
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<taxonomicName id="4C28CBACFFCDBB42C2ECFC94E023B1B9" baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1869" box="[742,886,961,987]" class="Insecta" family="Hesperiidae" genus="Ridens" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="fulminans">
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||
<emphasis id="B95C6C3DFFCDBB42C2ECFC94E023B1B9" box="[742,886,961,987]" italics="true" pageId="18" pageNumber="17">R. fulminans</emphasis>
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||
</taxonomicName>
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and
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<taxonomicName id="4C28CBACFFCDBB42C3B9FC97E7B5B1BE" authority="Evans, 1952" authorityName="Evans" authorityYear="1952" box="[947,1248,961,988]" class="Insecta" family="Hesperiidae" genus="Ridens" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="fulima">
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||
<emphasis id="B95C6C3DFFCDBB42C3B9FC97E71AB1B9" box="[947,1103,961,987]" italics="true" pageId="18" pageNumber="17">Ridens fulima</emphasis>
|
||
<bibRefCitation id="EFB9CDDEFFCDBB42C452FC94E7B5B1BE" author="Evans WH" box="[1112,1248,961,988]" pageId="18" pageNumber="17" pagination="10 - 25" refId="ref77051" refString="Evans WH. 1952. A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part II. Pyrginae. Section I. The Trustees of the British Museum (Natural History); London. v + 178 p., pl. 10 - 25." type="book chapter" year="1952">Evans, 1952</bibRefCitation>
|
||
</taxonomicName>
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||
(type locality in
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||
<collectingCountry id="F33FF0BFFFCDBB42C0CAFCB2E250B19C" box="[192,261,996,1022]" name="Brazil" pageId="18" pageNumber="17">Brazil</collectingCountry>
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||
:
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<collectingRegion id="49EC7ECDFFCDBB42C119FCB2E2EDB19C" box="[275,440,996,1022]" country="Brazil" name="Espirito Santo" pageId="18" pageNumber="17">Espírito Santo</collectingRegion>
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) in typically having narrower forewing hyaline spots (although some specimens have broader spots as well) and less prominent, mostly vestigial, pale discal band on the ventral hindwing (
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||
<figureCitation id="1313ACAAFFCDBB42C57EFB51E25CB626" captionStart="Figures 33–48" captionStartId="96.[192,271,1841,1865]" captionTargetBox="[198,1434,372,1745]" captionTargetId="figure-67@96.[192,1440,361,1751]" captionTargetPageId="96" captionText="Figures 33–48. Holotypes, data in text. 33–34) Ridens angulinea sp. n. 35–36) Pythonides lera sp. n. 37–38) Pythonides latemarginatus sp. n. 39–40) Gindanes variegatus sp. n. 41–42) Milanion (Milanion) virga sp. n. 43–44) Milanion (Milanion) furvus sp. n. 45–46) Milanion (Milanion) laricus sp. n. 47–48) Charidia ronda sp. n." pageId="18" pageNumber="17">Fig. 33–34</figureCitation>
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||
). Due to the cryptic nature of this species, most reliable identification is achieved by DNA and a combination of the following base pairs is diagnostic in the nuclear genome: aly116.28.11:A377G, aly116.28.11:A405G, aly2284.34.16:A61C, aly2284.34.16:A90G, aly
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||
<date id="FF9696EFFFCDBB42C2C9FB39E07DB6EB" box="[707,808,1135,1161]" pageId="18" pageNumber="17" value="2311-02-08">2311.2.8</date>
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||
:G123A, and COI barcode: T34C, T70C, A100C, T145C, T263C.
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||
</paragraph>
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||
<paragraph id="8B97B02FFFCDBB42C0CAFBE9E7D8B7F7" blockId="18.[192,1440,1215,1429]" pageId="18" pageNumber="17">
|
||
<emphasis id="B95C6C3DFFCDBB42C0CAFBE9E11BB6BB" bold="true" box="[192,590,1215,1241]" pageId="18" pageNumber="17">
|
||
Barcode sequence of the
|
||
<typeStatus id="54930E8DFFCDBB42C1EAFBE9E11CB6BB" box="[480,585,1215,1241]" pageId="18" pageNumber="17" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Sample NVG-14104E04, GenBank OR837636, 658 base pairs: AACTTTATATTTTATTTTTGGAATTTGAGCTGGCTTAATTGGAACTTCCTTAAGATTACTTATTCGTACCGAATTAGGAATTCCCGGTTCTTTAATT GGCGATGACCAAATTTATAATACTATTGTAACAGCACATGCCTTTATCATAATCTTTTTTATAGTAATACCAATTATAATTGGTGGATTTGGAAATT GATTAGTACCATTAATATTAGGAGCCCCTGATATAGCATTCCCCCGAATAAATAATATAAGATTTTGACTATTACCCCCATCTCTTACTCTTTTAAT TTCTAGAAGAATTGTAGAAAATGGTGCTGGAACAGGTTGAACAGTTTATCCCCCTCTTTCAACTAATATTGCCCATCAAGGGGCATCTGTCGACTTA GCCATTTTTTCTCTTCATTTAGCTGGAATTTCCTCAATTTTAGGAGCAATTAATTTTATTACAACAATTATTAATATGCGAATTAATAATTTATCAT TTGATCAAATACCATTATTTATTTGAGCCGTAGGAATTACAGCATTATTATTATTACTTTCTTTACCTGTATTGGCAGGAGCTATTACCATATTACT TACCGATCGAAACTTAAATACTTCATTTTTTGATCCTGCAGGAGGTGGAGATCCAATTTTATATCAACACTTATTT
|
||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C332E3A4FFCDBB42C0CAFAFDE022B56B" pageId="18" pageNumber="17" type="materials_examined">
|
||
<paragraph id="8B97B02FFFCDBB42C0CAFAFDE049B4B9" blockId="18.[191,1440,1450,1755]" pageId="18" pageNumber="17">
|
||
<emphasis id="B95C6C3DFFCDBB42C0CAFAFDE23DB7A6" bold="true" box="[192,360,1450,1477]" pageId="18" pageNumber="17">Type material.</emphasis>
|
||
<materialsCitation id="3B40BA72FFCDBB42C164FAFDE64EB44C" collectingDate="2013-01-29" collectionCode="DC, USNM, V" country="Peru" elevation="1194" location="Smithsonian Institution" municipality="National Museum of Natural History" pageId="18" pageNumber="17" specimenCount="2" specimenCount-male="2" stateProvince="Cusco" typeStatus="holotype">
|
||
<emphasis id="B95C6C3DFFCDBB42C164FAFDE2B7B7A7" bold="true" box="[366,482,1451,1477]" pageId="18" pageNumber="17">
|
||
<typeStatus id="54930E8DFFCDBB42C164FAFDE288B7A7" box="[366,477,1451,1477]" pageId="18" pageNumber="17" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D2E7BA6FFCDBB42C1E2FAFAE2A8B7AA" box="[488,509,1452,1480]" count="1" pageId="18" pageNumber="17" type="male">♂</specimenCount>
|
||
currently deposited in the
|
||
<collectingMunicipality id="6BF32A55FFCDBB42C323FAFDE793B7A7" box="[809,1222,1451,1477]" pageId="18" pageNumber="17">National Museum of Natural History</collectingMunicipality>
|
||
,
|
||
<location id="8EF7E6F4FFCDBB42C4DAFAFDE253B785" LSID="urn:lsid:plazi:treatment:03810139FFCDBB42C0CAFD97E1DAB5E4:8EF7E6F4FFCDBB42C4DAFAFDE253B785" country="Peru" municipality="National Museum of Natural History" name="Smithsonian Institution" pageId="18" pageNumber="17" stateProvince="Cusco">Smithsonian Institution</location>
|
||
, Washington,
|
||
<collectionCode id="ED3928EAFFCDBB42C1A9FA98E298B78A" box="[419,461,1486,1512]" pageId="18" pageNumber="17">DC</collectionCode>
|
||
, USA (
|
||
<collectionCode id="ED3928EAFFCDBB42C212FA98E138B78A" box="[536,621,1486,1512]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="17" type="Museum">USNM</collectionCode>
|
||
), illustrated in
|
||
<figureCitation id="1313ACAAFFCDBB42C31BFA9BE0D3B78A" box="[785,902,1485,1512]" captionStart="Figures 33–48" captionStartId="96.[192,271,1841,1865]" captionTargetBox="[198,1434,372,1745]" captionTargetId="figure-67@96.[192,1440,361,1751]" captionTargetPageId="96" captionText="Figures 33–48. Holotypes, data in text. 33–34) Ridens angulinea sp. n. 35–36) Pythonides lera sp. n. 37–38) Pythonides latemarginatus sp. n. 39–40) Gindanes variegatus sp. n. 41–42) Milanion (Milanion) virga sp. n. 43–44) Milanion (Milanion) furvus sp. n. 45–46) Milanion (Milanion) laricus sp. n. 47–48) Charidia ronda sp. n." pageId="18" pageNumber="17">Fig. 33–34</figureCitation>
|
||
, bears the following three rectangular labels, two white: [
|
||
<collectingCountry id="F33FF0BFFFCDBB42C110FAA6E236B468" box="[282,355,1520,1546]" name="Peru" pageId="18" pageNumber="17">PERU</collectingCountry>
|
||
:
|
||
<collectingRegion id="49EC7ECDFFCDBB42C16CFAA6E2F9B468" box="[358,428,1520,1546]" country="Peru" name="Cusco" pageId="18" pageNumber="17">Cuzco</collectingRegion>
|
||
<quantity id="4CD01DCAFFCDBB42C1B9FAA6E153B468" box="[435,518,1520,1546]" metricMagnitude="3" metricUnit="m" metricValue="1.194" pageId="18" pageNumber="17" unit="m" value="1194.0">
|
||
<elevation id="0005571CFFCDBB42C1B9FAA6E153B468" box="[435,518,1520,1546]" metricMagnitude="3" metricUnit="m" metricValue="1.194" pageId="18" pageNumber="17" unit="m" value="1194.0">1,194m</elevation>
|
||
</quantity>
|
||
| Bridge@
|
||
<quantity id="4CD01DCAFFCDBB42C270FAA6E198B469" box="[634,717,1520,1547]" metricMagnitude="3" metricUnit="m" metricValue="1.194" pageId="18" pageNumber="17" unit="m" value="1194.0">
|
||
<elevation id="0005571CFFCDBB42C270FAA6E198B469" box="[634,717,1520,1547]" metricMagnitude="3" metricUnit="m" metricValue="1.194" pageId="18" pageNumber="17" unit="m" value="1194.0">1194 m</elevation>
|
||
</quantity>
|
||
| Cosnipata Road 3167 |
|
||
<date id="FF9696EFFFCDBB42C3EDFAA6E718B468" box="[999,1101,1520,1546]" pageId="18" pageNumber="17" value="2013-01-29">
|
||
<collectingDate id="EFD26F07FFCDBB42C3EDFAA6E718B468" box="[999,1101,1520,1546]" pageId="18" pageNumber="17" value="2013-01-29">29.I.2013</collectingDate>
|
||
</date>
|
||
Kinyon], [DNA sample ID: | NVG-14104E04 | c/o Nick
|
||
<collectionCode id="ED3928EAFFCDBB42C1E5F942E157B44C" box="[495,514,1556,1582]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="18" pageNumber="17" type="Museum">V</collectionCode>
|
||
. Grishin], and one red [
|
||
<typeStatus id="54930E8DFFCDBB42C311F945E0F9B44F" box="[795,940,1555,1581]" pageId="18" pageNumber="17" type="holotype">HOLOTYPE</typeStatus>
|
||
<specimenCount id="9D2E7BA6FFCDBB42C3B9F942E09DB452" box="[947,968,1556,1584]" count="1" pageId="18" pageNumber="17" type="male">♂</specimenCount>
|
||
|
|
||
<taxonomicName id="4C28CBACFFCDBB42C3D7F945E7CBB44F" authorityName="Zhang & Cong & Grishin" authorityYear="2023" box="[989,1182,1555,1581]" class="Insecta" family="Hesperiidae" genus="Ridens" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="angulinea">Ridens angulinea</taxonomicName>
|
||
| Grishin]
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B40BA72FFCDBB42C52CF945E29FB411" collectingDate="2016-10-24" collectionCode="USNM" collectorName="S. Kinyon" country="Peru" location="Quebrada" municipality="Cosnipata Valley" pageId="18" pageNumber="17" specimenCount="6" specimenCount-male="6" stateProvince="Cusco" typeStatus="paratype">
|
||
<emphasis id="B95C6C3DFFCDBB42C52CF945E6F5B44F" bold="true" box="[1318,1440,1555,1581]" pageId="18" pageNumber="17">
|
||
<typeStatus id="54930E8DFFCDBB42C52CF945E6CEB44F" box="[1318,1435,1555,1581]" pageId="18" pageNumber="17" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D2E7BA6FFCDBB42C0CAF960E3A2B431" box="[192,247,1590,1619]" count="5" pageId="18" pageNumber="17" type="male">5♂♂</specimenCount>
|
||
:
|
||
<specimenCount id="9D2E7BA6FFCDBB42C109F960E270B431" box="[259,293,1590,1619]" count="1" pageId="18" pageNumber="17" type="male">1♂</specimenCount>
|
||
NVG-17103B12
|
||
<collectionCode id="ED3928EAFFCDBB42C1EEF960E167B432" box="[484,562,1590,1616]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="17" type="Museum">USNM</collectionCode>
|
||
_00913749
|
||
<collectingCountry id="F33FF0BFFFCDBB42C2B8F960E1BCB432" box="[690,745,1590,1616]" name="Peru" pageId="18" pageNumber="17">Peru</collectingCountry>
|
||
:
|
||
<collectingRegion id="49EC7ECDFFCDBB42C2F8F960E06FB432" box="[754,826,1590,1616]" country="Peru" name="Cusco" pageId="18" pageNumber="17">Cuzco</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BF32A55FFCDBB42C349F960E0AAB432" box="[835,1023,1590,1616]" pageId="18" pageNumber="17">Cosnipata Valley</collectingMunicipality>
|
||
,
|
||
<location id="8EF7E6F4FFCDBB42C402F960E722B432" LSID="urn:lsid:plazi:treatment:03810139FFCDBB42C0CAFD97E1DAB5E4:8EF7E6F4FFCDBB42C402F960E722B432" box="[1032,1143,1590,1616]" country="Peru" municipality="Cosnipata Valley" name="Quebrada" pageId="18" pageNumber="17" stateProvince="Cusco">Quebrada</location>
|
||
<taxonomicName id="4C28CBACFFCDBB42C477F960E657B432" authority="Isabel" authorityName="Isabel" box="[1149,1282,1590,1616]" class="Insecta" family="Hesperiidae" genus="Nisoniades" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="santa">Santa Isabel</taxonomicName>
|
||
,
|
||
<date id="FF9696EFFFCDBB42C501F960E6C9B432" box="[1291,1436,1590,1616]" pageId="18" pageNumber="17" value="2016-10-24">
|
||
<collectingDate id="EFD26F07FFCDBB42C501F960E6C9B432" box="[1291,1436,1590,1616]" pageId="18" pageNumber="17" value="2016-10-24">24-Oct-2016</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26DDD5F9FFCDBB42C0B5F90FE27BB411" box="[191,302,1625,1651]" pageId="18" pageNumber="17">S. Kinyon</collectorName>
|
||
leg. [
|
||
<collectionCode id="ED3928EAFFCDBB42C161F90FE296B411" box="[363,451,1625,1651]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="17" type="Museum">USNM</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B40BA72FFCDBB42C1DCF90FE067B4F7" collectingDate="1988-11" collectorName="Carlos Tello" country="Bolivia" elevation="950" location="Caranavi Province" municipality="Yungas Region" pageId="18" pageNumber="17" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="9D2E7BA6FFCDBB42C1DCF90FE2AFB414" box="[470,506,1625,1654]" count="1" pageId="18" pageNumber="17" type="male">1♂</specimenCount>
|
||
NVG-18089A11
|
||
<collectingCountry id="F33FF0BFFFCDBB42C2CBF90FE046B411" box="[705,787,1625,1651]" name="Bolivia" pageId="18" pageNumber="17">Bolivia</collectingCountry>
|
||
:
|
||
<collectingMunicipality id="6BF32A55FFCDBB42C314F90FE09CB411" box="[798,969,1625,1651]" pageId="18" pageNumber="17">Yungas Region</collectingMunicipality>
|
||
,
|
||
<location id="8EF7E6F4FFCDBB42C3DFF90FE7F2B411" LSID="urn:lsid:plazi:treatment:03810139FFCDBB42C0CAFD97E1DAB5E4:8EF7E6F4FFCDBB42C3DFF90FE7F2B411" box="[981,1191,1625,1651]" country="Bolivia" municipality="Yungas Region" name="Caranavi Province" pageId="18" pageNumber="17">Caranavi Province</location>
|
||
,
|
||
<locationDeviation id="9A25A90DFFCDBB42C4BEF90EE6C9B411" box="[1204,1436,1624,1651]" location="Coroico" pageId="18" pageNumber="17">
|
||
<quantity id="4CD01DCAFFCDBB42C4BEF90EE7A9B411" box="[1204,1276,1624,1651]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="17" unit="km" value="50.0">50 km</quantity>
|
||
N of Coroico
|
||
</locationDeviation>
|
||
,
|
||
<quantity id="4CD01DCAFFCDBB42C0C8F92AE21CB4F4" box="[194,329,1660,1686]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="12.0" metricValueMin="7.0" pageId="18" pageNumber="17" unit="m" value="950.0" valueMax="1200.0" valueMin="700.0">
|
||
<elevation id="0005571CFFCDBB42C0C8F92AE21CB4F4" box="[194,329,1660,1686]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="12.0" metricValueMin="7.0" pageId="18" pageNumber="17" unit="m" value="950.0" valueMax="1200.0" valueMin="700.0">700-1200 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FF9696EFFFCDBB42C15EF92AE290B4F4" box="[340,453,1660,1686]" pageId="18" pageNumber="17" value="1988-11">
|
||
<collectingDate id="EFD26F07FFCDBB42C15EF92AE290B4F4" box="[340,453,1660,1686]" pageId="18" pageNumber="17" value="1988-11">Nov-1988</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26DDD5F9FFCDBB42C1C5F92DE103B4F7" box="[463,598,1659,1685]" pageId="18" pageNumber="17">Carlos Tello</collectorName>
|
||
leg. [EBrockmann]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B40BA72FFCDBB42C337F92AE037B4DA" collectionCode="AMNH" collectorName="Fassl" country="Bolivia" elevation="750" location="Rio Songo" pageId="18" pageNumber="17" specimenCode="IZC 00337748" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
|
||
<specimenCount id="9D2E7BA6FFCDBB42C337F92AE027B4FB" box="[829,882,1660,1689]" count="3" pageId="18" pageNumber="17" type="male">3♂♂</specimenCount>
|
||
<collectingCountry id="F33FF0BFFFCDBB42C373F92DE09EB4F7" box="[889,971,1659,1685]" name="Bolivia" pageId="18" pageNumber="17">Bolivia</collectingCountry>
|
||
:
|
||
<location id="8EF7E6F4FFCDBB42C3DEF92DE71FB4F4" LSID="urn:lsid:plazi:treatment:03810139FFCDBB42C0CAFD97E1DAB5E4:8EF7E6F4FFCDBB42C3DEF92DE71FB4F4" box="[980,1098,1659,1686]" country="Bolivia" name="Rio Songo" pageId="18" pageNumber="17">Rio Songo</location>
|
||
: NVG-19041B10
|
||
<collectionCode id="ED3928EAFFCDBB42C504F92AE631B4F4" box="[1294,1380,1660,1686]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="18" pageNumber="17">AMNH</collectionCode>
|
||
_
|
||
<specimenCode id="DB8E1854FFCDBB42C579F92AE279B4DA" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="18" pageNumber="17">IZC 00337748</specimenCode>
|
||
<quantity id="4CD01DCAFFCDBB42C13FF9C8E2D5B4DB" box="[309,384,1694,1721]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="18" pageNumber="17" unit="m" value="750.0">
|
||
<elevation id="0005571CFFCDBB42C13FF9C8E2D5B4DB" box="[309,384,1694,1721]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="18" pageNumber="17" unit="m" value="750.0">750 m</elevation>
|
||
</quantity>
|
||
, coll.
|
||
<collectorName id="26DDD5F9FFCDBB42C1CFF9C8E2A8B4DA" box="[453,509,1694,1720]" pageId="18" pageNumber="17">Fassl</collectorName>
|
||
, genitalia slide G770 [
|
||
<collectionCode id="ED3928EAFFCDBB42C2F6F9C8E00EB4DA" box="[764,859,1694,1720]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="18" pageNumber="17">AMNH</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B40BA72FFCDBB42C365F9C8E042B4B9" collectionCode="R, ANSP" collectorName="C. Williams & Jr." country="Bolivia" location="No." pageId="18" pageNumber="17" specimenCount="1" typeStatus="paratype">
|
||
and
|
||
<collectionCode id="ED3928EAFFCDBB42C3A9F9C9E0EDB4DB" box="[931,952,1695,1721]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18" pageNumber="17">R</collectionCode>
|
||
.
|
||
<collectorName id="26DDD5F9FFCDBB42C3CEF9C8E718B4DA" box="[964,1101,1694,1720]" pageId="18" pageNumber="17">C. Williams</collectorName>
|
||
,
|
||
<collectorName id="26DDD5F9FFCDBB42C451F9C9E726B4DB" box="[1115,1139,1695,1721]" pageId="18" pageNumber="17">Jr.</collectorName>
|
||
collection [
|
||
<collectionCode id="ED3928EAFFCDBB42C4F3F9C8E612B4DA" box="[1273,1351,1694,1720]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="18" pageNumber="17">ANSP</collectionCode>
|
||
]: NVG-22042E10 and NVG-22042E11 genitalia slide
|
||
<location id="8EF7E6F4FFCDBB42C2B4F997E1B2B4B9" LSID="urn:lsid:plazi:treatment:03810139FFCDBB42C0CAFD97E1DAB5E4:8EF7E6F4FFCDBB42C2B4F997E1B2B4B9" box="[702,743,1729,1755]" country="Bolivia" name="No." pageId="18" pageNumber="17">No.</location>
|
||
316
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B97B02FFFCDBB42C0CAF9B9E022B56B" blockId="18.[192,887,1774,1801]" box="[192,887,1774,1801]" pageId="18" pageNumber="17">
|
||
<emphasis id="B95C6C3DFFCDBB42C0CAF9B9E20EB56A" bold="true" box="[192,347,1774,1801]" pageId="18" pageNumber="17">
|
||
<typeStatus id="54930E8DFFCDBB42C0CAF9B9E3ACB56B" box="[192,249,1775,1801]" pageId="18" pageNumber="17">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="F33FF0BFFFCDBB42C16BF9B9E2CDB56B" box="[353,408,1775,1801]" name="Peru" pageId="18" pageNumber="17">Peru</collectingCountry>
|
||
:
|
||
<collectingRegion id="49EC7ECDFFCDBB42C1A8F9B9E2BFB56B" box="[418,490,1775,1801]" country="Peru" name="Cusco" pageId="18" pageNumber="17">Cuzco</collectingRegion>
|
||
, Cosñipata Road, bridge at
|
||
<quantity id="4CD01DCAFFCDBB42C317F9B9E026B56B" box="[797,883,1775,1801]" metricMagnitude="3" metricUnit="m" metricValue="1.194" pageId="18" pageNumber="17" unit="m" value="1194.0">1194 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C332E3A4FFCDBB42C0CAF84AE799B53B" pageId="18" pageNumber="17" type="etymology">
|
||
<paragraph id="8B97B02FFFCDBB42C0CAF84AE799B53B" blockId="18.[192,1440,1820,1881]" pageId="18" pageNumber="17">
|
||
<emphasis id="B95C6C3DFFCDBB42C0CAF84AE211B554" bold="true" box="[192,324,1820,1846]" pageId="18" pageNumber="17">Etymology.</emphasis>
|
||
The name reflects narrower white bands and spots in many specimens of this species. It is derived from the Latin phrase “narrow lines”:
|
||
<emphasis id="B95C6C3DFFCDBB42C26CF869E1C8B53A" box="[614,669,1855,1880]" italics="true" pageId="18" pageNumber="17">angu</emphasis>
|
||
[stis]
|
||
<emphasis id="B95C6C3DFFCDBB42C2D1F869E056B53B" box="[731,771,1855,1881]" italics="true" pageId="18" pageNumber="17">line</emphasis>
|
||
[is]+a. The name is a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C332E3A4FFCDBB42C0CAF83AE1DAB5E4" box="[192,655,1900,1927]" pageId="18" pageNumber="17" type="distribution">
|
||
<paragraph id="8B97B02FFFCDBB42C0CAF83AE1DAB5E4" blockId="18.[192,655,1900,1927]" box="[192,655,1900,1927]" pageId="18" pageNumber="17">
|
||
<emphasis id="B95C6C3DFFCDBB42C0CAF83AE20EB5E4" bold="true" box="[192,347,1900,1926]" pageId="18" pageNumber="17">Distribution.</emphasis>
|
||
Southern
|
||
<collectingCountry id="F33FF0BFFFCDBB42C1C5F83BE156B5E5" box="[463,515,1901,1927]" name="Peru" pageId="18" pageNumber="17">Peru</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33FF0BFFFCDBB42C230F83AE1D9B5E4" box="[570,652,1900,1926]" name="Bolivia" pageId="18" pageNumber="17">Bolivia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |