231 lines
31 KiB
XML
231 lines
31 KiB
XML
<document id="6BEEF6F97025CEA39C3B3D1C3A780FCA" ID-CLB-Dataset="26356" ID-DOI="10.11646/zootaxa.4624.1.8" ID-GBIF-Dataset="4d45525c-5a3e-4e37-a64a-bbf25ba47313" ID-ISSN="1175-5326" ID-Zenodo-Dep="3259349" ID-ZooBank="7DBFCA0A-C648-4222-837C-D47DADE2232D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1561702388238" checkinUser="plazi" docAuthor="Brooks, Scott E., Cumming, Jeffrey M. & Grimaldi, David A." docDate="2019" docId="8A5E87C987172547FF0D24B7FAECF97D" docLanguage="en" docName="zootaxa.4624.1.8.pdf.imf" docOrigin="Zootaxa 4624 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Schistostoma foliatum Brooks, Cumming & Grimaldi, 2019, sp. nov." docType="treatment" docVersion="7" lastPageNumber="126" masterDocId="7667FFB187142542FF9A223DFFF1FF9E" masterDocTitle="Remarkable new fossil species of Schistostoma Becker (Diptera: Dolichopodidae Microphorinae) from mid-Cretaceous Burmese amber" masterLastPageNumber="131" masterPageNumber="121" pageNumber="124" updateTime="1698737691181" updateUser="ExternalLinkService">
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<mods:mods id="B0E014AAAB014BF293777D60F6DE7E78" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0745919016D606483993E87102840C7E">
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<mods:title id="B2E07B70BF6B46CB0A56A9440BAC3E42">Remarkable new fossil species of Schistostoma Becker (Diptera: Dolichopodidae Microphorinae) from mid-Cretaceous Burmese amber</mods:title>
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<mods:name id="A7B15355F73FF867C648BBE4F3E62ECC" type="personal">
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<mods:namePart id="F6690C4B0578AC0A0DFEA776A6B48B2E">Brooks, Scott E.</mods:namePart>
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<mods:roleTerm id="B6F889170064771BD889B730DD0026D4">Author</mods:roleTerm>
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<mods:namePart id="E6F49C6750D40126AD6524B7BC6BC693">Cumming, Jeffrey M.</mods:namePart>
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<mods:roleTerm id="366D9053497E3C30BEF465A2BE7F034B">Author</mods:roleTerm>
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<mods:namePart id="E908E9BDEBFA25D72B966656107FC17F">Grimaldi, David A.</mods:namePart>
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<mods:typeOfResource id="D733A35F52565BF68B6AB3F6845C5F74">text</mods:typeOfResource>
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<mods:title id="9D1A9CA58954236A4B461482BD352427">Zootaxa</mods:title>
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<mods:part id="19454716354A75A00FB9A27746BB1652">
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<mods:date id="69235DD5DF2D4856F71CC670BE13C0B3">2019</mods:date>
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<mods:number id="20F85DEF3BB794B1FCD7508DF98722F4">2019-06-27</mods:number>
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<mods:number id="6A464FBCFA921B2D05F4FD3213FFC540">4624</mods:number>
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<mods:classification id="7587C535C2DA2C00EAD559C54401D933">journal article</mods:classification>
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</mods:mods>
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<treatment id="8A5E87C987172547FF0D24B7FAECF97D" ID-GBIF-Taxon="158414990" LSID="urn:lsid:plazi:treatment:8A5E87C987172547FF0D24B7FAECF97D" httpUri="http://treatment.plazi.org/id/8A5E87C987172547FF0D24B7FAECF97D" lastPageId="5" lastPageNumber="126" pageId="3" pageNumber="124">
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<subSubSection id="4AED655487172541FF0D24B7FDFBF93A" box="[151,522,1674,1701]" pageId="3" pageNumber="124" type="nomenclature">
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<paragraph id="024836DF87172541FF0D24B7FDFBF93A" blockId="3.[151,522,1674,1736]" box="[151,522,1674,1701]" pageId="3" pageNumber="124">
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<heading id="590081B387172541FF0D24B7FDFBF93A" bold="true" box="[151,522,1674,1701]" fontSize="11" level="1" pageId="3" pageNumber="124" reason="1">
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<emphasis id="3083EACD87172541FF0D24B7FDFBF93A" bold="true" box="[151,522,1674,1701]" pageId="3" pageNumber="124">
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<taxonomicName id="C5F74D5C87172541FF0D24B7FE54F93B" box="[151,421,1674,1701]" class="Insecta" family="Dolichopodidae" genus="Schistostoma" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="foliatum" status="sp. nov.">
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<emphasis id="3083EACD87172541FF0D24B7FE54F93B" bold="true" box="[151,421,1674,1701]" italics="true" pageId="3" pageNumber="124">Schistostoma foliatum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="2BB057B687172541FE3624B7FDFBF93A" box="[428,522,1674,1700]" pageId="3" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="4AED655487172541FF0D2493FEE8F956" box="[151,281,1710,1736]" pageId="3" pageNumber="124" type="description">
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<paragraph id="024836DF87172541FF0D2493FEE8F956" blockId="3.[151,522,1674,1736]" box="[151,281,1710,1736]" pageId="3" pageNumber="124">
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(
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<figureCitation id="9ACC2A5A87172541FF052493FEE0F956" box="[159,273,1710,1736]" captionStart-0="FIGURES 5–8" captionStart-1="FIGURES 11–12" captionStartId-0="6.[151,264,1933,1958]" captionStartId-1="8.[151,264,1740,1765]" captionTargetBox-0="[224,1362,502,1899]" captionTargetBox-1="[151,1436,181,1712]" captionTargetId-0="figure@6.[212,1376,489,1914]" captionTargetId-1="figure@8.[151,1436,179,1713]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionText-0="FIGURES 5–8. Schistostoma burmanicum sp. nov. and Schistostoma foliatum sp. nov. 5. Right antenna of S. burmanicum, left medial view. 6. Left foreleg of S. burmanicum, left lateral view. 7. Hypopygium of S. burmanicum, ventral view. 8. Left foreleg of S. foliatum, left lateral view. Abbreviations: epand—epandrium; hypd—hypandrium; hypd proc—hypandrial process." captionText-1="FIGURES 11–12. Schistostoma foliatum sp. nov. male holotype (AMNH Bu-175). 11. Wing. 12. Legs, right lateral view. Abbreviations: C—costal vein; CuA—anterior branch of cubital vein; cua—anterior cubital cell; dm—discal medial cell; dmm—discal medial crossvein; h–humeral crossvein; M 1, M 2, M 4 —medial veins; R 1, R 2+3, R 4+5 —radial veins; r-m—radial-medial crossvein; Sc—subcostal vein." httpUri-0="https://zenodo.org/record/3259355/files/figure.png" httpUri-1="https://zenodo.org/record/3259357/files/figure.png" pageId="3" pageNumber="124">Figs 8–12</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="4AED655487172541FF0D24CBFB62F8E2" pageId="3" pageNumber="124" type="materials_examined">
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<paragraph id="024836DF87172541FF0D24CBFB62F8E2" blockId="3.[151,1436,1781,1988]" pageId="3" pageNumber="124">
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<emphasis id="3083EACD87172541FF0D24CBFEB2F88E" bold="true" box="[151,323,1781,1808]" pageId="3" pageNumber="124">Type material.</emphasis>
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<materialsCitation id="B29F3C8287172541FED224C8FB62F8E2" ID-GBIF-Occurrence="2270855924" collectionCode="AMNH, AMBER, MYAN-, MAR" collectorName="Leeward Capitol Corp." country="Myanmar" county="Upper Cretaceous" location="Ledo Rd." municipality="Tanai Village" pageId="3" pageNumber="124" specimenCount="1" specimenCount-male="1" stateProvince="Kachin" typeStatus="holotype">
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<emphasis id="3083EACD87172541FED224C8FE16F88E" bold="true" box="[328,487,1781,1808]" pageId="3" pageNumber="124">
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<typeStatus id="DD4C887D87172541FED224C8FE16F88E" box="[328,487,1781,1808]" pageId="3" pageNumber="124" type="holotype">HOLOTYPE</typeStatus>
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</emphasis>
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<specimenCount id="14F1FD5687172541FE7624CBFDF2F88E" box="[492,515,1782,1808]" pageId="3" pageNumber="124" type="male">♂</specimenCount>
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,
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<collectionCode id="64E6AE1A87172541FD9624CBFD93F88E" box="[524,610,1782,1808]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="124">AMNH</collectionCode>
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Bu-175, in Burmese amber with accompanying labels: “
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<collectionCode id="64E6AE1A87172541FB5724CBFAC9F88E" box="[1229,1336,1782,1808]" pageId="3" pageNumber="124">AMBER</collectionCode>
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:
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<collectionCode id="64E6AE1A87172541FADA24CBFA6DF88E" box="[1344,1436,1782,1808]" pageId="3" pageNumber="124">MYAN-</collectionCode>
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<collectionCode id="64E6AE1A87172541FF0D2527FF26F8AA" box="[151,215,1818,1844]" country="Zimbabwe" httpUri="http://grbio.org/cool/p96j-ip08" name="Grasslands Rhizobium Collection" pageId="3" pageNumber="124">MAR</collectionCode>
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(
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<collectingCountry id="7AE0764F87172541FF7D2527FEA1F8AA" box="[231,336,1818,1844]" name="Myanmar" pageId="3" pageNumber="124">BURMA</collectingCountry>
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)/
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<collectingCounty id="EB294E5387172541FEF22527FDC3F8AA" box="[360,562,1818,1844]" pageId="3" pageNumber="124">Upper Cretaceous</collectingCounty>
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/
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<collectingRegion id="C033F83D87172541FDD82527FD66F8AA" box="[578,663,1818,1844]" country="Myanmar" name="Kachin" pageId="3" pageNumber="124">Kachin</collectingRegion>
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:
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<collectingMunicipality id="E22CACA587172541FD392527FCC8F8AA" box="[675,825,1818,1844]" pageId="3" pageNumber="124">Tanai Village</collectingMunicipality>
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(on
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<location id="0728600487172541FCF42527FC26F8AA" LSID="urn:lsid:plazi:treatment:8A5E87C987172547FF0D24B7FAECF97D:0728600487172541FCF42527FC26F8AA" box="[878,983,1818,1844]" country="Myanmar" county="Upper Cretaceous" municipality="Tanai Village" name="Ledo Rd." pageId="3" pageNumber="124" stateProvince="Kachin">Ledo Rd.</location>
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/
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<locationDeviation id="13FA2FFD87172541FC7D2524FB18F8AB" box="[999,1257,1817,1845]" location="Myitkyna" pageId="3" pageNumber="124">
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<quantity id="C50F9B3A87172541FC7D2524FBCCF8AA" box="[999,1085,1817,1844]" metricMagnitude="5" metricUnit="m" metricValue="1.05" pageId="3" pageNumber="124" unit="km" value="105.0">105 km</quantity>
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NW Myitkyna
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</locationDeviation>
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)/ coll.
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<collectorName id="AF02530987172541FAA02527FEC4F8C6" pageId="3" pageNumber="124">Leeward Capitol Corp.</collectorName>
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, 2000/
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<collectionCode id="64E6AE1A87172541FE102503FE11F8C6" box="[394,480,1854,1880]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="124">AMNH</collectionCode>
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Bu-175”; “Burmese Amber/
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<taxonomicName id="C5F74D5C87172541FCA82503FC5DF8C6" box="[818,940,1854,1880]" kingdom="Plantae" order="Diptera" pageId="3" pageNumber="124" rank="order">DIPTERA</taxonomicName>
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:/
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<taxonomicName id="C5F74D5C87172541FC5B2503FB96F8C6" authorityName="Collin" authorityYear="1960" box="[961,1127,1854,1880]" class="Insecta" family="Microphoridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="family">Microphoridae</taxonomicName>
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(raptorial male)*/ 1 ACA-
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<taxonomicName id="C5F74D5C87172541FF0D255FFF10F8E2" authorityName="Poletaeva" authorityYear="1964" box="[151,225,1890,1916]" class="Trilobita" family="Solenopleuridae" genus="Rina" higherTaxonomySource="CoL" kingdom="Animalia" order="Agnostida" pageId="3" pageNumber="124" phylum="Arthropoda" rank="genus">RINA</taxonomicName>
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”; “
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<typeStatus id="DD4C887D87172541FE9A255FFE67F8E2" box="[256,406,1890,1916]" pageId="3" pageNumber="124" type="holotype">HOLOTYPE</typeStatus>
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/
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<taxonomicName id="C5F74D5C87172541FE3C255FFD6FF8E2" box="[422,670,1890,1916]" class="Insecta" family="Dolichopodidae" genus="Schistostoma" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="foliatum">
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<emphasis id="3083EACD87172541FE3C255FFD6FF8E2" box="[422,670,1890,1916]" italics="true" pageId="3" pageNumber="124">Schistostoma foliatum</emphasis>
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</taxonomicName>
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/ Cumming, Brooks & Grimaldi [red label]”.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="4AED655487172541FF5D25B8FC24F85A" pageId="3" pageNumber="124" type="diagnosis">
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<paragraph id="024836DF87172541FF5D25B8FC24F85A" blockId="3.[151,1436,1781,1988]" pageId="3" pageNumber="124">
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<emphasis id="3083EACD87172541FF5D25B8FEB3F83E" bold="true" box="[199,322,1925,1952]" pageId="3" pageNumber="124">Diagnosis.</emphasis>
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This species is characterized by its enlarged lamellate apically acute fore tibia (
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<figureCitation id="9ACC2A5A87172541FB2725BBFB0EF83E" box="[1213,1279,1926,1952]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1933,1958]" captionTargetBox="[224,1362,502,1899]" captionTargetId="figure@6.[212,1376,489,1914]" captionTargetPageId="6" captionText="FIGURES 5–8. Schistostoma burmanicum sp. nov. and Schistostoma foliatum sp. nov. 5. Right antenna of S. burmanicum, left medial view. 6. Left foreleg of S. burmanicum, left lateral view. 7. Hypopygium of S. burmanicum, ventral view. 8. Left foreleg of S. foliatum, left lateral view. Abbreviations: epand—epandrium; hypd—hypandrium; hypd proc—hypandrial process." httpUri="https://zenodo.org/record/3259355/files/figure.png" pageId="3" pageNumber="124">Fig. 8</figureCitation>
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) with distinctive colouration pattern (lacking ‘eyespot’) and fringe-like marginal setae.
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</paragraph>
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</subSubSection>
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<caption id="5688665787102546FF0D2411FD9FF9F7" httpUri="https://zenodo.org/record/3259353/files/figure.png" pageId="4" pageNumber="125" startId="4.[151,264,1580,1605]" targetBox="[151,1436,181,1560]" targetPageId="4">
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<paragraph id="024836DF87102546FF0D2411FD9FF9F7" blockId="4.[151,1437,1580,1641]" pageId="4" pageNumber="125">
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<emphasis id="3083EACD87102546FF0D2411FECBF9DB" bold="true" box="[151,314,1580,1605]" pageId="4" pageNumber="125">FIGURES 3–4.</emphasis>
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<taxonomicName id="C5F74D5C87102546FEA52411FDBBF9DA" box="[319,586,1580,1604]" class="Insecta" family="Dolichopodidae" genus="Schistostoma" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="burmanicum" status="sp. nov.">
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<emphasis id="3083EACD87102546FEA52411FDBBF9DA" box="[319,586,1580,1604]" italics="true" pageId="4" pageNumber="125">Schistostoma burmanicum</emphasis>
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</taxonomicName>
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<emphasis id="3083EACD87102546FDD52411FD6EF9DA" bold="true" box="[591,671,1580,1604]" pageId="4" pageNumber="125">
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<taxonomicNameLabel id="2BB057B687102546FDD52411FD6EF9DA" box="[591,671,1580,1604]" pageId="4" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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male holotype (AMNH Bu-0029).
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<emphasis id="3083EACD87102546FB9B2411FBE5F9DB" bold="true" box="[1025,1044,1580,1605]" pageId="4" pageNumber="125">3.</emphasis>
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Habitus photograph, right lateral view.
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<emphasis id="3083EACD87102546FF0D246DFF5BF9F7" bold="true" box="[151,170,1616,1641]" pageId="4" pageNumber="125">4.</emphasis>
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Right foreleg and midleg, right lateral view.
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</paragraph>
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</caption>
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<subSubSection id="4AED655487102547FF5D24ABFB95F921" lastPageId="5" lastPageNumber="126" pageId="4" pageNumber="125" type="description">
|
||
<paragraph id="024836DF87102547FF5D24ABFB95F921" blockId="4.[151,1437,1686,2037]" lastBlockId="5.[151,1437,153,1763]" lastPageId="5" lastPageNumber="126" pageId="4" pageNumber="125">
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<emphasis id="3083EACD87102546FF5D24ABFE55F92F" bold="true" box="[199,420,1686,1713]" pageId="4" pageNumber="125">Description. Male.</emphasis>
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Body length
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<quantity id="C50F9B3A87102546FDA624ABFD64F92F" box="[572,661,1686,1713]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="125" unit="mm" value="1.6">1.6 mm</quantity>
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. Wing length
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<quantity id="C50F9B3A87102546FCA824ABFC7AF92F" box="[818,907,1686,1713]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="125" unit="mm" value="1.5">1.5 mm</quantity>
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||
.
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<emphasis id="3083EACD87102546FC0C24AAFC2EF92F" bold="true" box="[918,991,1687,1713]" pageId="4" pageNumber="125">Head:</emphasis>
|
||
Elliptical in lateral view. Neck inserted near middle of head, with several short setae adjacent to occipital foramen. Ocellar triangle distinctly protruded with a pair of strong ocellar bristles and a pair of weaker postocellar setae. Holoptic (Fig. 10) with compound eyes contiguous for entire length of frons; compound eye ovoid in lateral view, occupying most of head, bare, dorsal ommatidia distinctly larger than ventral ommatidia, medial edge of eye with emargination slightly below level of antenna giving rise to demarcation line between larger dorsal and smaller ventral ommatidia. Frons entirely obliterated by eyes. Face and clypeus recessed in oral cavity. Face broad, subrectangular. Clypeus weakly bulging, mostly separated from frons, only connected medially by narrow sclerotized bridge. Occiput with series of 8–9 visible postocular setae per side. Gena narrow. Postgena broad. Antenna (Fig. 10) inserted near middle of head in profile; scape short about half length of pedicel; pedicel about
|
||
<date id="7649101F87102546FD6325E6FCEAF86B" box="[761,795,2011,2037]" pageId="4" pageNumber="125">2X</date>
|
||
longer than wide than wide, with several setae; postpedi- cel strongly narrowing beyond basal third, with dorsal margin concave, about
|
||
<date id="7649101F87112547FB9F22A4FBD6FF2D" box="[1029,1063,153,179]" pageId="5" pageNumber="126">3X</date>
|
||
longer than wide, clothed in fine hairs (longer ventrally); stylus arista-like, terminal, about 2.5X longer than postpedicel, 2-articled, basal article short. Palpus digitiform, with short setae. Proboscis narrow, directed anteroventrally, labellum with marginal setae.
|
||
<emphasis id="3083EACD87112547FF0D2338FF08FE81" bold="true" box="[151,249,261,287]" pageId="5" pageNumber="126">Thorax:</emphasis>
|
||
Slightly longer than wide in dorsal view. Mesoscutum strongly arched (Fig. 10), prescutellar depression present. Proepisternum bare. Postpronotal lobe with 1 strong seta, with adjacent short seta. Acrostichal setae present, apparently biserial; other bristles of thorax well-differentiated, each side with: apparently 10–11 dorsocentrals, anterior dorsocentrals short and closely spaced, 2 posteriormost dorsocentrals strong and widely separated; several apparent strong supra-alar setae; 2 strong notopleural setae; 1 strong postalar seta. Scutellum with posterior margin rounded, with 2 pairs of setae, outer pair weak, inner pair strong. Mesopleuron apparently bare.
|
||
<emphasis id="3083EACD87112547FB4A2384FAE0FE4D" bold="true" box="[1232,1297,441,467]" pageId="5" pageNumber="126">Legs:</emphasis>
|
||
Fairly long; foreleg and midleg with specialized morphology and chaetotaxy; tarsal claws, pulvilli and empodium normally developed on all legs.
|
||
<emphasis id="3083EACD87112547FEEF203CFE3CFD85" box="[373,461,513,539]" italics="true" pageId="5" pageNumber="126">Foreleg</emphasis>
|
||
(
|
||
<figureCitation id="9ACC2A5A87112547FE41203CFDEFFD85" box="[475,542,513,539]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1933,1958]" captionTargetBox="[224,1362,502,1899]" captionTargetId="figure@6.[212,1376,489,1914]" captionTargetPageId="6" captionText="FIGURES 5–8. Schistostoma burmanicum sp. nov. and Schistostoma foliatum sp. nov. 5. Right antenna of S. burmanicum, left medial view. 6. Left foreleg of S. burmanicum, left lateral view. 7. Hypopygium of S. burmanicum, ventral view. 8. Left foreleg of S. foliatum, left lateral view. Abbreviations: epand—epandrium; hypd—hypandrium; hypd proc—hypandrial process." httpUri="https://zenodo.org/record/3259355/files/figure.png" pageId="5" pageNumber="126">Fig. 8</figureCitation>
|
||
): Coxa with a few short subapical setae visible; femur with several strong dorsal and dorsolateral setae, with large strongly projecting basiventral tubercle bearing 5 apical setae, ventral surface immediately distal to basiventral tubercle with series of about 6 short setae (not visibly borne on tubercles); tibia greatly enlarged and lamellate, leaf-shaped with apical margin acute, flattened laterally and very slightly concave medially, distinctly patterned (
|
||
<figureCitation id="9ACC2A5A87112547FE7720ACFDC4FD35" box="[493,565,657,683]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1933,1958]" captionTargetBox="[224,1362,502,1899]" captionTargetId="figure@6.[212,1376,489,1914]" captionTargetPageId="6" captionText="FIGURES 5–8. Schistostoma burmanicum sp. nov. and Schistostoma foliatum sp. nov. 5. Right antenna of S. burmanicum, left medial view. 6. Left foreleg of S. burmanicum, left lateral view. 7. Hypopygium of S. burmanicum, ventral view. 8. Left foreleg of S. foliatum, left lateral view. Abbreviations: epand—epandrium; hypd—hypandrium; hypd proc—hypandrial process." httpUri="https://zenodo.org/record/3259355/files/figure.png" pageId="5" pageNumber="126">Figs 8</figureCitation>
|
||
,
|
||
<figureCitation id="9ACC2A5A87112547FDD820ACFDAEFD35" box="[578,607,657,683]" captionStart="FIGURES 11–12" captionStartId="8.[151,264,1740,1765]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure@8.[151,1436,179,1713]" captionTargetPageId="8" captionText="FIGURES 11–12. Schistostoma foliatum sp. nov. male holotype (AMNH Bu-175). 11. Wing. 12. Legs, right lateral view. Abbreviations: C—costal vein; CuA—anterior branch of cubital vein; cua—anterior cubital cell; dm—discal medial cell; dmm—discal medial crossvein; h–humeral crossvein; M 1, M 2, M 4 —medial veins; R 1, R 2+3, R 4+5 —radial veins; r-m—radial-medial crossvein; Sc—subcostal vein." httpUri="https://zenodo.org/record/3259357/files/figure.png" pageId="5" pageNumber="126">12</figureCitation>
|
||
) but lacking discrete ‘eye spot’, lateral surface with longitudinal series of 5 short setae along middle and 6 longer setae basiventrally, dorsal and ventral margins with fringe of setae, medial surface bare; tarsus with short setae, tarsomere 1 as long as combined length of tarsomeres 2–3.
|
||
<emphasis id="3083EACD87112547FB4020E4FAD8FD6D" box="[1242,1321,729,755]" italics="true" pageId="5" pageNumber="126">Midleg</emphasis>
|
||
(
|
||
<figureCitation id="9ACC2A5A87112547FAA020E4FA7EFD6D" box="[1338,1423,729,755]" captionStart="FIGURES 11–12" captionStartId="8.[151,264,1740,1765]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure@8.[151,1436,179,1713]" captionTargetPageId="8" captionText="FIGURES 11–12. Schistostoma foliatum sp. nov. male holotype (AMNH Bu-175). 11. Wing. 12. Legs, right lateral view. Abbreviations: C—costal vein; CuA—anterior branch of cubital vein; cua—anterior cubital cell; dm—discal medial cell; dmm—discal medial crossvein; h–humeral crossvein; M 1, M 2, M 4 —medial veins; R 1, R 2+3, R 4+5 —radial veins; r-m—radial-medial crossvein; Sc—subcostal vein." httpUri="https://zenodo.org/record/3259357/files/figure.png" pageId="5" pageNumber="126">Fig. 12</figureCitation>
|
||
): Coxa with strong elongate anterior seta at extreme base and series of 4 gradually shorter setae towards apex; trochanter with several weak setae; femur with strong elongate basidorsal seta, series of 3 strong anterodorsal setae along basal half (about half-length basidorsal seta), series of short dorsal setae towards apex, prominent basiventral tubercle bearing thick curved seta, 2 ventral setae near middle (of similar size) and several apparent anterior setae; tibia with flange on basal half bearing close-set series of short modified setae (some hook-shaped), 1 long erect anterior seta near mid-length, several hook-shaped setae on distal half; tarsomere 1 with setae somewhat longer than on tarsomeres 2–5.
|
||
<emphasis id="3083EACD87112547FEEF21E8FE3EFC71" box="[373,463,981,1007]" italics="true" pageId="5" pageNumber="126">Hindleg</emphasis>
|
||
: Coxal setae obscured from view; femur with series of strong dorsal and ventral setae of similar length, dorsal setae gradually shorter apically, ventral setae gradually shorter basally; tibia with series of strong dorsal setae, gradually stronger distally, series of similar ventral setae, gradually weaker distally; tarsomere 1 slightly shorter than combined length of tarsomeres 2–5, with prominent basiventral seta, otherwise with short setae similar to other tarsomeres.
|
||
<emphasis id="3083EACD87112547FD892659FDACFBE1" bold="true" box="[531,605,1124,1151]" pageId="5" pageNumber="126">Wing:</emphasis>
|
||
Pterostigma absent, membrane entirely covered with minute microtrichia (
|
||
<figureCitation id="9ACC2A5A87112547FF3A26B4FF00FB3D" box="[160,241,1161,1187]" captionStart="FIGURES 11–12" captionStartId="8.[151,264,1740,1765]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure@8.[151,1436,179,1713]" captionTargetPageId="8" captionText="FIGURES 11–12. Schistostoma foliatum sp. nov. male holotype (AMNH Bu-175). 11. Wing. 12. Legs, right lateral view. Abbreviations: C—costal vein; CuA—anterior branch of cubital vein; cua—anterior cubital cell; dm—discal medial cell; dmm—discal medial crossvein; h–humeral crossvein; M 1, M 2, M 4 —medial veins; R 1, R 2+3, R 4+5 —radial veins; r-m—radial-medial crossvein; Sc—subcostal vein." httpUri="https://zenodo.org/record/3259357/files/figure.png" pageId="5" pageNumber="126">Fig. 11</figureCitation>
|
||
). Anal lobe well-developed and right-angled, alula absent. Costa circumambient. Extreme anterior base of costa with 2 strong setae. Anterior costal section bearing row of short spine-like setae intermixed with fine setae from humeral crossvein to R
|
||
<subScript id="9E73349A87112547FE4826DFFE1DFB6E" attach="left" box="[466,492,1250,1264]" fontSize="6" pageId="5" pageNumber="126">2+3</subScript>
|
||
, posterior part of costa beyond R
|
||
<subScript id="9E73349A87112547FCC026DFFC82FB6E" attach="left" box="[858,883,1250,1264]" fontSize="6" pageId="5" pageNumber="126">2+3</subScript>
|
||
with only fine setae. Longitudinal veins complete reaching wing margin (CuA+CuP obscured from view). Sc faint apically. R
|
||
<subScript id="9E73349A87112547FC41273BFC12FA8A" attach="left" box="[987,995,1286,1300]" fontSize="6" pageId="5" pageNumber="126">1</subScript>
|
||
weakly sinuous, reaching costa beyond middle of wing (or beyond base of M
|
||
<subScript id="9E73349A87112547FDAF2717FDCCFAA6" attach="left" box="[565,573,1322,1336]" fontSize="6" pageId="5" pageNumber="126">2</subScript>
|
||
). Base of Rs originating opposite humeral crossvein. R
|
||
<subScript id="9E73349A87112547FB392717FB4DFAA6" attach="left" box="[1187,1212,1322,1336]" fontSize="6" pageId="5" pageNumber="126">2+3</subScript>
|
||
diverging from R
|
||
<subScript id="9E73349A87112547FA192717FA6CFAA6" attach="left" box="[1411,1437,1322,1336]" fontSize="6" pageId="5" pageNumber="126">4+5</subScript>
|
||
apically. R
|
||
<subScript id="9E73349A87112547FE8A2773FEDBFAC2" attach="left" box="[272,298,1358,1372]" fontSize="6" pageId="5" pageNumber="126">4+5</subScript>
|
||
weakly sinuous. M
|
||
<subScript id="9E73349A87112547FD932773FDE0FAC2" attach="left" box="[521,529,1358,1372]" fontSize="6" pageId="5" pageNumber="126">1</subScript>
|
||
diverging from R
|
||
<subScript id="9E73349A87112547FD442773FD09FAC2" attach="left" box="[734,760,1358,1372]" fontSize="6" pageId="5" pageNumber="126">4+5</subScript>
|
||
beyond cell dm. M
|
||
<subScript id="9E73349A87112547FC4D2773FC2EFAC2" attach="left" box="[983,991,1358,1372]" fontSize="6" pageId="5" pageNumber="126">1</subScript>
|
||
and M
|
||
<subScript id="9E73349A87112547FBAB2773FBC8FAC2" attach="left" box="[1073,1081,1358,1372]" fontSize="6" pageId="5" pageNumber="126">2</subScript>
|
||
strongly diverging beyond cell dm. M
|
||
<subScript id="9E73349A87112547FF7E274FFF1DFA1E" attach="left" box="[228,236,1394,1408]" fontSize="6" pageId="5" pageNumber="126">2</subScript>
|
||
and M
|
||
<subScript id="9E73349A87112547FEA4274FFEB7FA1E" attach="left" box="[318,326,1394,1408]" fontSize="6" pageId="5" pageNumber="126">4</subScript>
|
||
weakly diverging beyond cell dm. Short r-m crossvein present (but nearly indiscernible) in basal portion of wing, distal to base of R
|
||
<subScript id="9E73349A87112547FD8727ABFDC6FA3A" attach="left" box="[541,567,1430,1444]" fontSize="6" pageId="5" pageNumber="126">4+5</subScript>
|
||
. Crossvein bm-m complete. Cell dm present, closed by base of M
|
||
<subScript id="9E73349A87112547FA8127ABFAD2FA3A" attach="left" box="[1307,1315,1430,1444]" fontSize="6" pageId="5" pageNumber="126">2</subScript>
|
||
and crossvein dm-m, cell extended to middle of wing. Cells br, bm and cua in basal fourth of wing. Cells bm and cua broader than br. Cell cua closed, rounded apically with CuA curved. Vein CuA+CuP obscured from view. Calypter obscured from view.
|
||
<emphasis id="3083EACD87112547FE8127CDFE68F995" bold="true" box="[283,409,1520,1547]" pageId="5" pageNumber="126">Abdomen:</emphasis>
|
||
Tergites and sternites 1–6 with relatively long setae. Segment 7 tubular, surface obscured from view. Sternite 8 ovoid, forming dome-like cap over dorsal region of hypopygium, with several strong marginal setae.
|
||
<emphasis id="3083EACD87112547FF442407FE9CF9CD" box="[222,365,1594,1619]" italics="true" pageId="5" pageNumber="126">Hypopygium</emphasis>
|
||
(
|
||
<figureCitation id="9ACC2A5A87112547FEE72404FE4CF9CD" box="[381,445,1593,1619]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1933,1958]" captionTargetBox="[224,1362,502,1899]" captionTargetId="figure@6.[212,1376,489,1914]" captionTargetPageId="6" captionText="FIGURES 5–8. Schistostoma burmanicum sp. nov. and Schistostoma foliatum sp. nov. 5. Right antenna of S. burmanicum, left medial view. 6. Left foreleg of S. burmanicum, left lateral view. 7. Hypopygium of S. burmanicum, ventral view. 8. Left foreleg of S. foliatum, left lateral view. Abbreviations: epand—epandrium; hypd—hypandrium; hypd proc—hypandrial process." httpUri="https://zenodo.org/record/3259355/files/figure.png" pageId="5" pageNumber="126">Fig 7</figureCitation>
|
||
): Lateroflexed to right; inverted with posterior end directed anteriorly; small, about 1/5 length of abdomen, apparently symmetrical. Right and left epandrial lamellae rounded basally and oblong in ventral view, apices obscured from view. Hypandrium rounded basally with narrow elongate medial hypandrial process flanked by pair of strong setae. Cercus short, subtriangular, setose.
|
||
<emphasis id="3083EACD87112547FC1D2498FC16F921" bold="true" box="[903,999,1701,1727]" pageId="5" pageNumber="126">Female:</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4AED655487112547FF5D24F4FAECF97D" box="[199,1309,1737,1763]" pageId="5" pageNumber="126" type="etymology">
|
||
<paragraph id="024836DF87112547FF5D24F4FAECF97D" blockId="5.[151,1437,153,1763]" box="[199,1309,1737,1763]" pageId="5" pageNumber="126">
|
||
<emphasis id="3083EACD87112547FF5D24F4FEBFF97D" bold="true" box="[199,334,1737,1763]" pageId="5" pageNumber="126">Etymology.</emphasis>
|
||
The specific epithet refers to the greatly enlarged leaf-shaped tibia of the male foreleg.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |