161 lines
20 KiB
XML
161 lines
20 KiB
XML
<document id="B4C636DC1028C1A2990B3A800F751142" ID-CLB-Dataset="8837" ID-DOI="10.11646/zootaxa.4908.2.3" ID-GBIF-Dataset="65624457-9eba-46fa-93dc-dfadc60cb618" ID-ISSN="1175-5326" ID-Zenodo-Dep="4438304" ID-ZooBank="BEE08FC0-68C2-469D-BDEA-4F1D9843012E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610615188102" checkinUser="plazi" docAuthor="Shear, William A." docDate="2021" docId="7670A27AFF88FFC7FF69EFB5D9DB4FE9" docLanguage="en" docName="zootaxa.4908.2.3.pdf" docOrigin="Zootaxa 4908 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amplaria mendocino Shear 2021, new species" docType="treatment" docVersion="8" lastPageNumber="221" masterDocId="8A49DA02FF87FFD7FFFEE902DB294D40" masterDocTitle="The millipede family Striariidae Bollman, 1893. II. New records and species of the genus Amplaria Chamberlin, 1941 (Diplopoda, Chordeumatida, Striarioidea)" masterLastPageNumber="224" masterPageNumber="205" pageNumber="220" updateTime="1698924432405" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="BAE95908E0782A8F7E2C7DBB175961CB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1930355B7613FC0EECEAD1391D18DC5D">
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<mods:title id="F69CBED896271D3ECBADBE1BBACB9115">The millipede family Striariidae Bollman, 1893. II. New records and species of the genus Amplaria Chamberlin, 1941 (Diplopoda, Chordeumatida, Striarioidea)</mods:title>
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</mods:titleInfo>
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<mods:name id="C35BC06597D8017FD6D87B06498518A8" type="personal">
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<mods:role id="405AF166EA6297530B9C6EFA710BFE56">
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<mods:roleTerm id="04C20C8E962F22E159C08A1FC3570638">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E492FC935C9983E935C02F5108B60910">Shear, William A.</mods:namePart>
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<mods:affiliation id="CDA82B3095851C796B40C131B7825E2B">Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943 USA; current address: 1950 Price Drive, Farmville VA 23901 USA.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="4FBC4D4ABB9A2603E2E2D783B79AA7F8">text</mods:typeOfResource>
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<mods:titleInfo id="312BD256667C40412E11D0110BA0CD69">
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<mods:title id="D8FE0BA79D41753BE80006EA961CAD92">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="298950E7E25ED2DAA8D4B22629CA024F">
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<mods:date id="C23AC46589590EBC91C8B8C32D6101DA">2021</mods:date>
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<mods:detail id="5A0A1DBBA20188D5075873140847E0A1" type="pubDate">
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<mods:number id="C4370BF7943D25D9D8CA102A080FE957">2021-01-14</mods:number>
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<mods:number id="8A761591BA707CD184A8952ECE322743">4908</mods:number>
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<mods:detail id="D334A2398DE6C373662A4C7CD6C0D9F5" type="issue">
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<mods:number id="976DB22C54D7B625912A80644371E277">2</mods:number>
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<mods:start id="4F8E07CE071428247DA1E8588E4EDD12">205</mods:start>
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<mods:identifier id="68D822FDBF29C4926783E029CCD5152D" type="CLB-Dataset">8837</mods:identifier>
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<mods:identifier id="D6B12C15E0B58704C3AC4402D46C0843" type="DOI">10.11646/zootaxa.4908.2.3</mods:identifier>
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<mods:identifier id="D545BF8B97AA1D2C0E7D8CDF12EE913E" type="GBIF-Dataset">65624457-9eba-46fa-93dc-dfadc60cb618</mods:identifier>
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<mods:identifier id="FEBC675F80D8361D902BC144F7F05066" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="7670A27AFF88FFC7FF69EFB5D9DB4FE9" ID-DOI="http://doi.org/10.5281/zenodo.4448045" ID-GBIF-Taxon="177006410" ID-Zenodo-Dep="4448045" LSID="urn:lsid:plazi:treatment:7670A27AFF88FFC7FF69EFB5D9DB4FE9" httpUri="http://treatment.plazi.org/id/7670A27AFF88FFC7FF69EFB5D9DB4FE9" lastPageId="16" lastPageNumber="221" pageId="15" pageNumber="220">
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<subSubSection id="B6C340E7FF88FFD8FF69EFB5D91C4B92" box="[151,565,1719,1746]" pageId="15" pageNumber="220" type="nomenclature">
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<paragraph id="FE66136CFF88FFD8FF69EFB5D91C4B92" blockId="15.[151,565,1719,1783]" box="[151,565,1719,1746]" pageId="15" pageNumber="220">
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<heading id="A52EA400FF88FFD8FF69EFB5D91C4B92" bold="true" box="[151,565,1719,1746]" fontSize="11" level="1" pageId="15" pageNumber="220" reason="1">
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<emphasis id="CCADCF7EFF88FFD8FF69EFB5D91C4B92" bold="true" box="[151,565,1719,1746]" pageId="15" pageNumber="220">
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<emphasis id="CCADCF7EFF88FFD8FF69EFB5DAB44B92" bold="true" box="[151,413,1719,1746]" italics="true" pageId="15" pageNumber="220">
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<taxonomicName id="39D968EFFF88FFD8FF69EFB5DAB14B92" authority="Shear, 2021" authorityName="Shear" authorityYear="2021" box="[151,408,1719,1746]" class="Diplopoda" family="Striariidae" genus="Amplaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="mendocino" status="sp. nov.">Amplaria mendocino</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="D79E7205FF88FFD8FE5BEFBAD91C4B92" box="[421,565,1720,1746]" pageId="15" pageNumber="220" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B6C340E7FF88FFD8FF69EFDEDA9D4AC6" pageId="15" pageNumber="220" type="description">
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<paragraph id="FE66136CFF88FFD8FF69EFDEDA324BB7" blockId="15.[151,565,1719,1783]" box="[151,283,1756,1783]" pageId="15" pageNumber="220">
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<figureCitation id="66E20FE9FF88FFD8FF69EFDEDBDA4BB6" box="[151,243,1756,1782]" captionStart="FIGS" captionStartId="16.[151,210,1820,1844]" captionTargetBox="[264,1323,707,1795]" captionTargetId="figure-219@16.[264,1323,705,1797]" captionTargetPageId="16" captionText="FIGS. 41–44. Amplaria menocino, A. flucticulus. Fig. 41. Amplaria mendocino, legpairs 2–4 of male, ventral view. FIGS. 42–44. A. flucticulus. Fig. 42. Head, collum and rings 2–6, dorsolateral view. Fig. 43. Legpairs 2 and 3 of male, lateral view. Fig. 44. Right gonopod, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4438329" httpUri="https://zenodo.org/record/4438329/files/figure.png" pageId="15" pageNumber="220">Figs. 41</figureCitation>
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,
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<figureCitation id="66E20FE9FF88FFD8FF01EFDEDA324BB7" box="[255,283,1756,1783]" captionStart="FIGS" captionStartId="18.[151,210,1748,1772]" captionTargetBox="[300,1283,786,1715]" captionTargetId="figure-251@18.[294,1293,777,1725]" captionTargetPageId="18" captionText="FIGS. 55–59. Drawings of gonopods of Amplaria species. Fig. 55. Right gonopod of A. umatilla, mesal view. Fig. 56. Right gonopod of A. cervus, mesal view. Fig. 57. Right gonopod of A. mendocino, mesal view. Fig. 58. Right gonopod of A. flucticulus, lateral view. Fig. 59. Right gonopod of A. flucticulus, mesal view." figureDoi="http://doi.org/10.5281/zenodo.4438335" httpUri="https://zenodo.org/record/4438335/files/figure.png" pageId="15" pageNumber="220">57</figureCitation>
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</paragraph>
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<paragraph id="FE66136CFF88FFD8FF69EE27DA9D4AC6" blockId="15.[151,1437,1828,2034]" pageId="15" pageNumber="220">
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<emphasis id="CCADCF7EFF88FFD8FF69EE27DBFF4A7E" box="[151,214,1829,1854]" italics="true" pageId="15" pageNumber="220">Types</emphasis>
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:
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<materialsCitation id="4EB11931FF88FFD8FF19EE26DA994AC6" ID-GBIF-Occurrence="3019425315" collectingDate="1990-09-20" collectorName="D. Ubick. Parts" country="United States of America" county="Mendocino Co." location="Usal Road" municipality="Male" pageId="15" pageNumber="220" specimenCount="4" stateProvince="California" typeStatus="holotype">
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<collectingMunicipality id="1E028916FF88FFD8FF19EE26DA084A7E" box="[231,289,1828,1854]" pageId="15" pageNumber="220">Male</collectingMunicipality>
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<typeStatus id="2162ADCEFF88FFD8FED7EE26DAA24A7E" box="[297,395,1828,1854]" pageId="15" pageNumber="220" type="holotype">holotype</typeStatus>
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and male
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<typeStatus id="2162ADCEFF88FFD8FDFDEE26D94A4A7E" box="[515,611,1828,1854]" pageId="15" pageNumber="220" type="paratype">paratype</typeStatus>
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from
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<location id="FB0645B7FF88FFD8FD55EE26D80C4A7E" LSID="urn:lsid:plazi:treatment:7670A27AFF88FFC7FF69EFB5D9DB4FE9:FB0645B7FF88FFD8FD55EE26D80C4A7E" box="[683,805,1828,1854]" country="United States of America" county="Mendocino Co." municipality="Male" name="Usal Road" pageId="15" pageNumber="220" stateProvince="California">Usal Road</location>
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,
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<quantity id="3921BE89FF88FFD8FCCFEE26D84F4A7E" box="[817,870,1828,1854]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="15" pageNumber="220" unit="mi" value="1.0">1 mi</quantity>
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N of
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<collectingCountry id="86CE53FCFF88FFD8FC55EE26D8E64A7E" box="[939,975,1828,1854]" name="United States of America" pageId="15" pageNumber="220">US</collectingCountry>
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1, 39.7910°, -123.8290°, 700’ asl,
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<collectingCounty id="17076BE0FF88FFD8FA9EEE26DA3C4A22" pageId="15" pageNumber="220">Mendocino Co.</collectingCounty>
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,
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<collectingRegion id="3C1DDD8EFF88FFD8FEDFEE4ADABC4A22" box="[289,405,1864,1890]" country="United States of America" name="California" pageId="15" pageNumber="220">California</collectingRegion>
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, collected
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<date id="8A6735ACFF88FFD8FDF3EE4AD9C14A22" box="[525,744,1864,1890]" pageId="15" pageNumber="220" value="1990-09-20">
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<collectingDate id="9A23CC44FF88FFD8FDF3EE4AD9C14A22" box="[525,744,1864,1890]" pageId="15" pageNumber="220" value="1990-09-20">20 September 1990</collectingDate>
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</date>
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by
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<collectorName id="532C76BAFF88FFD8FCEDEE4AD8E94A22" box="[787,960,1864,1890]" pageId="15" pageNumber="220">D. Ubick. Parts</collectorName>
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of the
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<typeStatus id="2162ADCEFF88FFD8FBEEEE4ADF5B4A22" box="[1040,1138,1864,1890]" pageId="15" pageNumber="220" type="holotype">holotype</typeStatus>
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and
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<typeStatus id="2162ADCEFF88FFD8FB57EE4ADE204A22" box="[1193,1289,1864,1890]" pageId="15" pageNumber="220" type="paratype">paratype</typeStatus>
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are mounted on SEM stub WAS34-16
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B6C340E7FF88FFD8FF39EE92DA214AB2" pageId="15" pageNumber="220" type="diagnosis">
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<paragraph id="FE66136CFF88FFD8FF39EE92DA214AB2" blockId="15.[151,1437,1828,2034]" pageId="15" pageNumber="220">
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<emphasis id="CCADCF7EFF88FFD8FF39EE92DA134AEA" bold="true" box="[199,314,1936,1962]" pageId="15" pageNumber="220">Diagnosis</emphasis>
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: Very similar in gonopod anatomy to the foregoing species,
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<emphasis id="CCADCF7EFF88FFD8FC25EE92DF624AEA" box="[987,1099,1936,1962]" italics="true" pageId="15" pageNumber="220">
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<taxonomicName id="39D968EFFF88FFD8FC25EE92DF6E4AEA" authorityName="Shear" authorityYear="2021" box="[987,1095,1936,1962]" class="Diplopoda" family="Striariidae" genus="Amplaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="cervus">A. cervus</taxonomicName>
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.
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</emphasis>
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Distinct in the ventral depressions on male coxae 4 and 5, which also bear fine, dense setae (
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<figureCitation id="66E20FE9FF88FFD8FCA6EEB6D8824A8E" box="[856,939,1972,1998]" captionStart="FIGS" captionStartId="16.[151,210,1820,1844]" captionTargetBox="[264,1323,707,1795]" captionTargetId="figure-219@16.[264,1323,705,1797]" captionTargetPageId="16" captionText="FIGS. 41–44. Amplaria menocino, A. flucticulus. Fig. 41. Amplaria mendocino, legpairs 2–4 of male, ventral view. FIGS. 42–44. A. flucticulus. Fig. 42. Head, collum and rings 2–6, dorsolateral view. Fig. 43. Legpairs 2 and 3 of male, lateral view. Fig. 44. Right gonopod, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4438329" httpUri="https://zenodo.org/record/4438329/files/figure.png" pageId="15" pageNumber="220">Fig. 41</figureCitation>
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). Legpair 1 is not enlarged and lacks needlelike setae.
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</paragraph>
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</subSubSection>
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<subSubSection id="B6C340E7FF97FFC7FF39E995DBE84D95" pageId="16" pageNumber="221" type="etymology">
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<paragraph id="FE66136CFF97FFC7FF39E995DBE84D95" blockId="16.[151,1437,151,681]" pageId="16" pageNumber="221">
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<emphasis id="CCADCF7EFF97FFC7FF39E995DA614DF1" bold="true" box="[199,328,151,177]" pageId="16" pageNumber="221">Etymology</emphasis>
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: The species name, a noun in apposition, recognizes the
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<typeStatus id="2162ADCEFF97FFC7FC2DE995DF2A4DF1" box="[979,1027,151,177]" pageId="16" pageNumber="221">type</typeStatus>
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locality in Mendocino Co.,
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<collectingRegion id="3C1DDD8EFF97FFC7FABAE995DB944D95" country="United States of America" name="California" pageId="16" pageNumber="221">California</collectingRegion>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B6C340E7FF97FFC7FF39E9DCDAEA4FC5" pageId="16" pageNumber="221" type="description">
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<paragraph id="FE66136CFF97FFC7FF39E9DCD9A04CB5" blockId="16.[151,1437,151,681]" pageId="16" pageNumber="221">
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<emphasis id="CCADCF7EFF97FFC7FF39E9DCDA784DB8" bold="true" box="[199,337,222,248]" pageId="16" pageNumber="221">Description</emphasis>
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:
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<emphasis id="CCADCF7EFF97FFC7FE9DE9DDD9264DB9" box="[355,527,223,249]" italics="true" pageId="16" pageNumber="221">
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Male
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<typeStatus id="2162ADCEFF97FFC7FE58E9DDD9234DB9" box="[422,522,223,249]" pageId="16" pageNumber="221" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Length, about
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<quantity id="3921BE89FF97FFC7FD3EE9DDD83A4DB9" box="[704,787,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="16" pageNumber="221" unit="mm" value="11.0">11 mm</quantity>
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, width 1.0 mm. Fourteen ommatidia in triangular patch. Labrum with long, slightly sinuous hooks at each angle. Mandibles not modified. Legpairs 1 and 2 similar in size; legpair 1 lacks long needle-like setae. Legpairs 2–7 telopodites of similar size, with flattened podomeres. Second trochanters with relatively small posteriorventral processes, bearing specialized setae. Legs 3 with coxal flasks of moderate length, divergent, with dense setae anterioapically, telopodites not reduced (
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<figureCitation id="66E20FE9FF97FFC7FBA6E86DDF824CC9" box="[1112,1195,367,393]" captionStart="FIGS" captionStartId="16.[151,210,1820,1844]" captionTargetBox="[264,1323,707,1795]" captionTargetId="figure-219@16.[264,1323,705,1797]" captionTargetPageId="16" captionText="FIGS. 41–44. Amplaria menocino, A. flucticulus. Fig. 41. Amplaria mendocino, legpairs 2–4 of male, ventral view. FIGS. 42–44. A. flucticulus. Fig. 42. Head, collum and rings 2–6, dorsolateral view. Fig. 43. Legpairs 2 and 3 of male, lateral view. Fig. 44. Right gonopod, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4438329" httpUri="https://zenodo.org/record/4438329/files/figure.png" pageId="16" pageNumber="221">Fig. 41</figureCitation>
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). Pleurotergal bars of ring 3 narrow, just touching in midline. Coxae of legpairs 4, 5, distally depressed, with fine, dense setae (
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<figureCitation id="66E20FE9FF97FFC7FAC5E891DEA64CED" box="[1339,1423,403,429]" captionStart="FIGS" captionStartId="16.[151,210,1820,1844]" captionTargetBox="[264,1323,707,1795]" captionTargetId="figure-219@16.[264,1323,705,1797]" captionTargetPageId="16" captionText="FIGS. 41–44. Amplaria menocino, A. flucticulus. Fig. 41. Amplaria mendocino, legpairs 2–4 of male, ventral view. FIGS. 42–44. A. flucticulus. Fig. 42. Head, collum and rings 2–6, dorsolateral view. Fig. 43. Legpairs 2 and 3 of male, lateral view. Fig. 44. Right gonopod, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4438329" httpUri="https://zenodo.org/record/4438329/files/figure.png" pageId="16" pageNumber="221">Fig. 41</figureCitation>
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). Crests moderate. Pygidium short. Color pale tan, each metazonite with dark purplish dots on posterior ends of crests 3 and 4, metazonites shaded darker laterally.
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</paragraph>
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<paragraph id="FE66136CFF97FFC7FF39E8FDDBE44F21" blockId="16.[151,1437,151,681]" pageId="16" pageNumber="221">
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Gonopods (
|
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<figureCitation id="66E20FE9FF97FFC7FEB4E8FDDAB54F59" box="[330,412,510,537]" captionStart="FIGS" captionStartId="18.[151,210,1748,1772]" captionTargetBox="[300,1283,786,1715]" captionTargetId="figure-251@18.[294,1293,777,1725]" captionTargetPageId="18" captionText="FIGS. 55–59. Drawings of gonopods of Amplaria species. Fig. 55. Right gonopod of A. umatilla, mesal view. Fig. 56. Right gonopod of A. cervus, mesal view. Fig. 57. Right gonopod of A. mendocino, mesal view. Fig. 58. Right gonopod of A. flucticulus, lateral view. Fig. 59. Right gonopod of A. flucticulus, mesal view." figureDoi="http://doi.org/10.5281/zenodo.4438335" httpUri="https://zenodo.org/record/4438335/files/figure.png" pageId="16" pageNumber="221">Fig. 57</figureCitation>
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) almost identical to those of
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<emphasis id="CCADCF7EFF97FFC7FD1BE8FDD87D4F58" box="[741,852,511,537]" italics="true" pageId="16" pageNumber="221">
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||
<taxonomicName id="39D968EFFF97FFC7FD1BE8FDD8794F58" authorityName="Shear" authorityYear="2021" box="[741,848,511,537]" class="Diplopoda" family="Striariidae" genus="Amplaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="16" pageNumber="221" phylum="Arthropoda" rank="species" species="cervus">A. cervus</taxonomicName>
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,
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</emphasis>
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but anterior angiocoxites differing slightly in shape of distal end (compare
|
||
<figureCitation id="66E20FE9FF97FFC7FE65EB21DAC24F7D" box="[411,491,546,573]" captionStart="FIGS" captionStartId="18.[151,210,1748,1772]" captionTargetBox="[300,1283,786,1715]" captionTargetId="figure-251@18.[294,1293,777,1725]" captionTargetPageId="18" captionText="FIGS. 55–59. Drawings of gonopods of Amplaria species. Fig. 55. Right gonopod of A. umatilla, mesal view. Fig. 56. Right gonopod of A. cervus, mesal view. Fig. 57. Right gonopod of A. mendocino, mesal view. Fig. 58. Right gonopod of A. flucticulus, lateral view. Fig. 59. Right gonopod of A. flucticulus, mesal view." figureDoi="http://doi.org/10.5281/zenodo.4438335" httpUri="https://zenodo.org/record/4438335/files/figure.png" pageId="16" pageNumber="221">Fig. 56</figureCitation>
|
||
and
|
||
<figureCitation id="66E20FE9FF97FFC7FDDDEB21D95E4F7D" box="[547,631,546,573]" captionStart="FIGS" captionStartId="18.[151,210,1748,1772]" captionTargetBox="[300,1283,786,1715]" captionTargetId="figure-251@18.[294,1293,777,1725]" captionTargetPageId="18" captionText="FIGS. 55–59. Drawings of gonopods of Amplaria species. Fig. 55. Right gonopod of A. umatilla, mesal view. Fig. 56. Right gonopod of A. cervus, mesal view. Fig. 57. Right gonopod of A. mendocino, mesal view. Fig. 58. Right gonopod of A. flucticulus, lateral view. Fig. 59. Right gonopod of A. flucticulus, mesal view." figureDoi="http://doi.org/10.5281/zenodo.4438335" httpUri="https://zenodo.org/record/4438335/files/figure.png" pageId="16" pageNumber="221">Fig. 57</figureCitation>
|
||
). Ninth legpair with flattened coxae, very small, acute mesal coxal process.
|
||
</paragraph>
|
||
<paragraph id="FE66136CFF97FFC7FF39EB69DAEA4FC5" blockId="16.[151,1437,151,681]" box="[199,451,619,645]" pageId="16" pageNumber="221">Females not collected.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B6C340E7FF97FFC7FF39EB8CD9DB4FE9" box="[199,754,654,681]" pageId="16" pageNumber="221" type="distribution">
|
||
<paragraph id="FE66136CFF97FFC7FF39EB8CD9DB4FE9" blockId="16.[151,1437,151,681]" box="[199,754,654,681]" pageId="16" pageNumber="221">
|
||
<emphasis id="CCADCF7EFF97FFC7FF39EB8CDA704FE8" bold="true" box="[199,345,654,680]" pageId="16" pageNumber="221">Distribution</emphasis>
|
||
: Known only from the
|
||
<typeStatus id="2162ADCEFF97FFC7FD9FEB8DD9B84FE9" box="[609,657,655,681]" pageId="16" pageNumber="221">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |