175 lines
21 KiB
XML
175 lines
21 KiB
XML
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<mods:title id="D98B41DBC83019C506CCF46EC61C7E6A">Taxonomy of Neotropical genera Compsodactylus and Dasyus and notes on claw movement in Macrodactylini (Coleoptera: Scarabaeidae: Melolonthinae)</mods:title>
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<mods:namePart id="ADFE880D9AA4545A0452B772A7A41426">Fuhrmann, Juares</mods:namePart>
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<mods:title id="2ADB21C9DE9A8705E2184DCFC234BE00">Zootaxa</mods:title>
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<mods:part id="1FDFBE3DA3EAA0B8734E2A0C1AB03C51">
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<mods:date id="E70183EEFAED8209428A612FF1258AFA">2019</mods:date>
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<mods:number id="0F65E24837958A04C16FD23B002CAECA">4679</mods:number>
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<treatment id="6902F1027936046AFF484CA314DFFD27" ID-DOI="http://doi.org/10.5281/zenodo.5874092" ID-GBIF-Taxon="159609687" ID-Zenodo-Dep="5874092" LSID="urn:lsid:plazi:treatment:6902F1027936046AFF484CA314DFFD27" httpUri="http://treatment.plazi.org/id/6902F1027936046AFF484CA314DFFD27" lastPageNumber="141" pageId="2" pageNumber="141">
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<subSubSection id="A9B1139F7936046AFF484CA312CEFEDA" pageId="2" pageNumber="141" type="nomenclature">
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<paragraph id="E11440147936046AFF484CA31351FF49" blockId="2.[151,531,153,180]" box="[151,531,153,180]" pageId="2" pageNumber="141">
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<heading id="BA5CF7787936046AFF484CA31351FF49" bold="true" box="[151,531,153,180]" fontSize="11" level="1" pageId="2" pageNumber="141" reason="1">
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<emphasis id="D3DF9C067936046AFF484CA31351FF49" bold="true" box="[151,531,153,180]" pageId="2" pageNumber="141">
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Key to
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<taxonomicName id="26AB3B977936046AFF2E4CA010F4FF4E" authorityName="Fuhrmann" authorityYear="2012" box="[241,438,153,180]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="genus">
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<emphasis id="D3DF9C067936046AFF2E4CA010F4FF4E" bold="true" box="[241,438,153,180]" italics="true" pageId="2" pageNumber="141">Compsodactylus</emphasis>
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</taxonomicName>
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species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="E11440147936046AFF484CD812CEFEDA" blockId="2.[151,1437,225,288]" pageId="2" pageNumber="141">
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Modified from
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<bibRefCitation id="853A3DE57936046AFE984CD810B9FF01" author="Fuhrmann, J." box="[327,507,225,251]" pageId="2" pageNumber="141" pagination="43 - 57" refId="ref12727" refString="Fuhrmann, J. (2012) Compsodactylus, a new South American genus with one new species and two new combinations (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 3577 (1), 43 - 57. https: // doi. org / 10.11646 / zootaxa. 3577.1.2" type="journal article" year="2012">Fuhrmann 2012</bibRefCitation>
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and using data of
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<bibRefCitation id="853A3DE57936046AFD0D4CD81502FF01" author="Figueroa, L. & Neita-Moreno, J. C." box="[722,1088,225,251]" pageId="2" pageNumber="141" pagination="587 - 591" refId="ref12528" refString="Figueroa, L. & Neita-Moreno, J. C. (2019) A new Compsodactylus Fuhrmann (Coleoptera: Scarabaeidae: Melolonthinae) from Peru. Zootaxa, 4560 (3), 587 - 591. https: // doi. org / 10.11646 / zootaxa. 4560.3.12" type="journal article" year="2019">Figueroa & Neita-Moreno 2019</bibRefCitation>
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. Males do not have metatibiae spurs while females have two metatibial spurs (see remarks above).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A9B1139F7936046AFF484D7314DFFD27" pageId="2" pageNumber="141" type="key">
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<keyStep id="5A5F58B17936046AFF484D7314DEFE4D" pageId="2" pageNumber="141">
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<paragraph id="E11440147936046AFF484D7314DEFE86" blockId="2.[151,1437,330,733]" pageId="2" pageNumber="141">
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<keyLead id="5A5AE3217936046AFF484D7314DEFE86" pageId="2" pageNumber="141">
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<emphasis id="D3DF9C067936046AFF484D7311E1FEA5" bold="true" box="[151,163,330,351]" pageId="2" pageNumber="141">1</emphasis>
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. Metafemur of males greatly widened and about three times wider than width of medial area of metatibia; females with gonostylus...............................................................................................
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<emphasis id="D3DF9C067936046AFA4F4D5E14DEFE86" bold="true" box="[1424,1436,359,380]" pageId="2" pageNumber="141">2</emphasis>
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</keyLead>
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</paragraph>
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<paragraph id="E11440147936046AFF484DBC14DEFE4D" blockId="2.[151,1437,330,733]" pageId="2" pageNumber="141">
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<keyLead id="5A5AE3217936046AFF484DBC14DEFE4D" pageId="2" pageNumber="141">
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- Metafemur of males slightly widened and less than two times the width of the medial area of the metatibia (
|
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<figureCitation id="79905C917936046AFB264DBC142CFE60" box="[1273,1390,388,411]" captionStart="FIGURES 9–20" captionStartId="6.[151,264,1633,1658]" captionTargetBox="[155,1436,246,1595]" captionTargetId="figure@6.[151,1436,229,1613]" captionTargetPageId="6" captionText="FIGURES 9–20. Male appendages. 9, Dasyus collaris metacoxa; 10, Compsodactylus argentinus metacoxa; 11, C. argentinus apex of metatibia; 12, C. argentinus metafemur and tibia; 13, C. argentinus tibia, posterior view; 14, C. parvulus apex of metatibia; 15, C. parvulus metafemur and tibia; 16, C. parvulus tibia, posterior view; 17–18, C. argentinus aedeagus (lateral and parameres in dorsal view); 19–20, C. parvulus aedeagus (lateral and parameres in dorsal view). cxk, coxa-katepisternum articulation; cxp, coxopleural articulation; cxt, coxa-trochanter articulation; mbd, marginal bead; mip, metacoxal inner process (limit in dotted line). Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3468199" httpUri="https://zenodo.org/record/3468199/files/figure.png" pageId="2" pageNumber="141">Figs. 12, 15</figureCitation>
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); females without gonostylus...............................................................................
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<emphasis id="D3DF9C067936046AFA4F4D9B14DEFE4D" bold="true" box="[1424,1436,418,439]" pageId="2" pageNumber="141">3</emphasis>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="5A5F58B17936046AFF484D8614DEFD9D" pageId="2" pageNumber="141">
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<paragraph id="E11440147936046AFF484D8614DFFDEA" blockId="2.[151,1437,330,733]" pageId="2" pageNumber="141">
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<keyLead id="5A5AE3217936046AFF484D8614DFFDEA" pageId="2" pageNumber="141">
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<emphasis id="D3DF9C067936046AFF484D8611E1FE2E" bold="true" box="[151,163,447,468]" pageId="2" pageNumber="141">2</emphasis>
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. Pronotum and elytra with metallic green reflection; apex of protarsomere I of males slightly wider than apex of II, metatarsus of males somewhat moniliform; pygidium of males with scale-like setae; females with distal gonocoxite somewhat trapezoid or oval;
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<collectingCountry id="99BC00847936046AFEED4DC3101CFDF5" box="[306,350,506,527]" name="Peru" pageId="2" pageNumber="141">Peru</collectingCountry>
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(
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<collectingRegion id="236F8EF67936046AFEB54DC3108DFDEA" box="[362,463,506,528]" country="Peru" name="Apurimac" pageId="2" pageNumber="141">Apurímac</collectingRegion>
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;
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<figureCitation id="79905C917936046AFE074DC3135FFDEA" box="[472,541,506,528]" captionStart="FIGURES 98–99" captionStartId="22.[151,264,1517,1542]" captionTargetBox="[214,1363,182,1469]" captionTargetId="figure@22.[151,1436,174,1498]" captionTargetPageId="22" captionText="FIGURES 98–99. South America distributional records. 98, Compsodactylus; 99, Dasyus. Scale = 500 km." figureDoi="http://doi.org/10.5281/zenodo.3468229" httpUri="https://zenodo.org/record/3468229/files/figure.png" pageId="2" pageNumber="141">Fig. 98</figureCitation>
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)....................................................
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<taxonomicName id="26AB3B977936046AFB7C4DC314DFFDEA" authority="(Frey, 1972)" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[1187,1437,506,528]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="martinezi">
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||
<emphasis id="D3DF9C067936046AFB7C4DC314DFFDEA" bold="true" box="[1187,1437,506,528]" pageId="2" pageNumber="141">
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||
<emphasis id="D3DF9C067936046AFB7C4DC3145EFDF5" bold="true" box="[1187,1308,506,527]" italics="true" pageId="2" pageNumber="141">C. martinezi</emphasis>
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(
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<bibRefCitation id="853A3DE57936046AFAF64DC314D7FDEA" author="Frey, G." box="[1321,1429,506,528]" pageId="2" pageNumber="141" pagination="67 - 85" refId="ref12652" refString="Frey, G. (1972) Bestimmungstabelle der Gattung Dicrania Serv. (Coleoptera - Melolonthinae - Macrodactylini). Entomologische Arbeiten aus dem Museum G. Frey, 23, 67 - 85." type="journal article" year="1972">Frey, 1972</bibRefCitation>
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)
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||
</emphasis>
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||
</taxonomicName>
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||
</keyLead>
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||
</paragraph>
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||
<paragraph id="E11440147936046AFF484E2114DEFD9D" blockId="2.[151,1437,330,733]" pageId="2" pageNumber="141">
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||
<keyLead id="5A5AE3217936046AFF484E2114DEFD9D" pageId="2" pageNumber="141">
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||
- Dorsal surface without metallic green reflection; apex of protarsomere I of males twice wider than apex of II or almost so, metatarsus of males somewhat filiform; pygidium of males with hair-like setae; females with distal gonocoxite coma-like;
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||
<collectingCountry id="99BC00847936046AFAAF4E0C14DEFDB0" box="[1392,1436,565,586]" name="Peru" pageId="2" pageNumber="141">Peru</collectingCountry>
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(
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||
<collectingRegion id="236F8EF67936046AFF374E6B101FFD92" box="[232,349,594,616]" country="Peru" name="La Libertad" pageId="2" pageNumber="141">La Libertad</collectingRegion>
|
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;
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||
<figureCitation id="79905C917936046AFEB84E6B10EEFD92" box="[359,428,594,616]" captionStart="FIGURES 98–99" captionStartId="22.[151,264,1517,1542]" captionTargetBox="[214,1363,182,1469]" captionTargetId="figure@22.[151,1436,174,1498]" captionTargetPageId="22" captionText="FIGURES 98–99. South America distributional records. 98, Compsodactylus; 99, Dasyus. Scale = 500 km." figureDoi="http://doi.org/10.5281/zenodo.3468229" httpUri="https://zenodo.org/record/3468229/files/figure.png" pageId="2" pageNumber="141">Fig. 98</figureCitation>
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)...............................................
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<taxonomicName id="26AB3B977936046AFC344E6B14DEFD9D" authority="Figueroa & Neita-Moreno, 2019" authorityName="Figueroa & Neita-Moreno" authorityYear="2019" box="[1003,1436,594,616]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="vallejoi">
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<emphasis id="D3DF9C067936046AFC344E6B14DEFD9D" bold="true" box="[1003,1436,594,616]" pageId="2" pageNumber="141">
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<emphasis id="D3DF9C067936046AFC344E6B12B9FD9D" bold="true" box="[1003,1019,594,615]" italics="true" pageId="2" pageNumber="141">C</emphasis>
|
||
.
|
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<emphasis id="D3DF9C067936046AFBD84E6B150DFD92" bold="true" box="[1031,1103,594,616]" italics="true" pageId="2" pageNumber="141">vallejoi</emphasis>
|
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<bibRefCitation id="853A3DE57936046AFB8A4E6B14DEFD9D" author="Figueroa, L. & Neita-Moreno, J. C." box="[1109,1436,594,616]" pageId="2" pageNumber="141" pagination="587 - 591" refId="ref12528" refString="Figueroa, L. & Neita-Moreno, J. C. (2019) A new Compsodactylus Fuhrmann (Coleoptera: Scarabaeidae: Melolonthinae) from Peru. Zootaxa, 4560 (3), 587 - 591. https: // doi. org / 10.11646 / zootaxa. 4560.3.12" type="journal article" year="2019">Figueroa & Neita-Moreno, 2019</bibRefCitation>
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</emphasis>
|
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</taxonomicName>
|
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</keyLead>
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</paragraph>
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</keyStep>
|
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<keyStep id="5A5F58B17936046AFF484E5614DFFD27" pageId="2" pageNumber="141">
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<paragraph id="E11440147936046AFF484E5614DFFD58" blockId="2.[151,1437,330,733]" pageId="2" pageNumber="141">
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<keyLead id="5A5AE3217936046AFF484E5614DFFD58" pageId="2" pageNumber="141">
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<emphasis id="D3DF9C067936046AFF484E5611E1FD7E" bold="true" box="[151,163,623,644]" pageId="2" pageNumber="141">3</emphasis>
|
||
. Elytral striae strongly and regularly punctate; metatibia of males gradually enlarged (
|
||
<figureCitation id="79905C917936046AFBCF4E561515FD7F" box="[1040,1111,623,645]" captionStart="FIGURES 9–20" captionStartId="6.[151,264,1633,1658]" captionTargetBox="[155,1436,246,1595]" captionTargetId="figure@6.[151,1436,229,1613]" captionTargetPageId="6" captionText="FIGURES 9–20. Male appendages. 9, Dasyus collaris metacoxa; 10, Compsodactylus argentinus metacoxa; 11, C. argentinus apex of metatibia; 12, C. argentinus metafemur and tibia; 13, C. argentinus tibia, posterior view; 14, C. parvulus apex of metatibia; 15, C. parvulus metafemur and tibia; 16, C. parvulus tibia, posterior view; 17–18, C. argentinus aedeagus (lateral and parameres in dorsal view); 19–20, C. parvulus aedeagus (lateral and parameres in dorsal view). cxk, coxa-katepisternum articulation; cxp, coxopleural articulation; cxt, coxa-trochanter articulation; mbd, marginal bead; mip, metacoxal inner process (limit in dotted line). Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3468199" httpUri="https://zenodo.org/record/3468199/files/figure.png" pageId="2" pageNumber="141">Fig. 12</figureCitation>
|
||
); females with inner distal area of gonocoxite rounded;
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||
<collectingCountry id="99BC00847936046AFE734EB4134EFD59" box="[428,524,653,675]" name="Argentina" pageId="2" pageNumber="141">Argentina</collectingCountry>
|
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(
|
||
<collectingRegion id="236F8EF67936046AFDC74EB413C1FD58" box="[536,643,653,674]" country="Argentina" name="Catamarca" pageId="2" pageNumber="141">Catamarca</collectingRegion>
|
||
,
|
||
<collectingRegion id="236F8EF67936046AFD524EB413FDFD59" box="[653,703,653,675]" country="Argentina" name="Salta" pageId="2" pageNumber="141">Salta</collectingRegion>
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||
,
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||
<collectingRegion id="236F8EF67936046AFD174EB41267FD59" box="[712,805,653,675]" country="Argentina" name="Tucuman" pageId="2" pageNumber="141">Tucumán</collectingRegion>
|
||
;
|
||
<figureCitation id="79905C917936046AFCF04EB41236FD59" box="[815,884,653,675]" captionStart="FIGURES 98–99" captionStartId="22.[151,264,1517,1542]" captionTargetBox="[214,1363,182,1469]" captionTargetId="figure@22.[151,1436,174,1498]" captionTargetPageId="22" captionText="FIGURES 98–99. South America distributional records. 98, Compsodactylus; 99, Dasyus. Scale = 500 km." figureDoi="http://doi.org/10.5281/zenodo.3468229" httpUri="https://zenodo.org/record/3468229/files/figure.png" pageId="2" pageNumber="141">Fig. 98</figureCitation>
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||
)......................
|
||
<taxonomicName id="26AB3B977936046AFB5B4EB414DFFD58" authority="(Moser, 1919)" baseAuthorityName="Moser" baseAuthorityYear="1919" box="[1156,1437,652,674]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="argentinus">
|
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<emphasis id="D3DF9C067936046AFB5B4EB414DFFD58" bold="true" box="[1156,1437,652,674]" pageId="2" pageNumber="141">
|
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<emphasis id="D3DF9C067936046AFB5B4EB415D6FD58" bold="true" box="[1156,1172,653,674]" italics="true" pageId="2" pageNumber="141">C</emphasis>
|
||
.
|
||
<emphasis id="D3DF9C067936046AFB7F4EB41449FD58" bold="true" box="[1184,1291,653,674]" italics="true" pageId="2" pageNumber="141">argentinus</emphasis>
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||
(
|
||
<bibRefCitation id="853A3DE57936046AFAC74EB514D7FD58" author="Moser, J." box="[1304,1429,652,674]" pageId="2" pageNumber="141" pagination="3 - 64" refId="ref13786" refString="Moser, J. (1919) Beitrag zur Kenntnis der Melolonthiden (Col.). (IX). Stettiner Entomologische Zeitung, 80, 3 - 64." type="journal article" year="1919">Moser, 1919</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="E11440147936046AFF484E9314DFFD27" blockId="2.[151,1437,330,733]" pageId="2" pageNumber="141">
|
||
<keyLead id="5A5AE3217936046AFF484E9314DFFD27" pageId="2" pageNumber="141">
|
||
- Elytral striae shallowly and irregularly punctate; metatibia of males abruptly enlarged in proximal third (
|
||
<figureCitation id="79905C917936046AFB174E93144DFD3A" box="[1224,1295,682,704]" captionStart="FIGURES 9–20" captionStartId="6.[151,264,1633,1658]" captionTargetBox="[155,1436,246,1595]" captionTargetId="figure@6.[151,1436,229,1613]" captionTargetPageId="6" captionText="FIGURES 9–20. Male appendages. 9, Dasyus collaris metacoxa; 10, Compsodactylus argentinus metacoxa; 11, C. argentinus apex of metatibia; 12, C. argentinus metafemur and tibia; 13, C. argentinus tibia, posterior view; 14, C. parvulus apex of metatibia; 15, C. parvulus metafemur and tibia; 16, C. parvulus tibia, posterior view; 17–18, C. argentinus aedeagus (lateral and parameres in dorsal view); 19–20, C. parvulus aedeagus (lateral and parameres in dorsal view). cxk, coxa-katepisternum articulation; cxp, coxopleural articulation; cxt, coxa-trochanter articulation; mbd, marginal bead; mip, metacoxal inner process (limit in dotted line). Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3468199" httpUri="https://zenodo.org/record/3468199/files/figure.png" pageId="2" pageNumber="141">Fig. 15</figureCitation>
|
||
); females with inner distal area of gonocoxite straight;
|
||
<collectingCountry id="99BC00847936046AFDBF4EFE13E5FD27" box="[608,679,711,733]" name="Bolivia" pageId="2" pageNumber="141">Bolivia</collectingCountry>
|
||
(
|
||
<collectingRegion id="236F8EF67936046AFD6B4EFE1277FD27" box="[692,821,711,733]" country="Bolivia" name="Cochabamba" pageId="2" pageNumber="141">Cochabamba</collectingRegion>
|
||
,
|
||
<collectingRegion id="236F8EF67936046AFCE14EFE12EEFD26" box="[830,940,711,732]" country="Bolivia" name="Santa Cruz" pageId="2" pageNumber="141">Santa Cruz</collectingRegion>
|
||
;
|
||
<figureCitation id="79905C917936046AFC6A4EFE12B8FD27" box="[949,1018,711,733]" captionStart="FIGURES 98–99" captionStartId="22.[151,264,1517,1542]" captionTargetBox="[214,1363,182,1469]" captionTargetId="figure@22.[151,1436,174,1498]" captionTargetPageId="22" captionText="FIGURES 98–99. South America distributional records. 98, Compsodactylus; 99, Dasyus. Scale = 500 km." figureDoi="http://doi.org/10.5281/zenodo.3468229" httpUri="https://zenodo.org/record/3468229/files/figure.png" pageId="2" pageNumber="141">Fig. 98</figureCitation>
|
||
)..............
|
||
<taxonomicName id="26AB3B977936046AFB764EFE14DFFD27" authority="(Frey, 1970)" baseAuthorityName="Frey" baseAuthorityYear="1970" box="[1193,1437,711,733]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="parvulus">
|
||
<emphasis id="D3DF9C067936046AFB764EFE14DFFD27" bold="true" box="[1193,1437,711,733]" pageId="2" pageNumber="141">
|
||
<emphasis id="D3DF9C067936046AFB764EFE145EFD27" bold="true" box="[1193,1308,711,733]" italics="true" pageId="2" pageNumber="141">C. parvulus</emphasis>
|
||
(Frey, 1970)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |