415 lines
54 KiB
XML
415 lines
54 KiB
XML
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<mods:title id="30C40987BA42AD09D0FB2316634D8FD0">Revision of the genus Exaesiopus Reichardt, 1926 (Coleoptera, Histeridae, Saprininae)</mods:title>
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<mods:namePart id="3C4FE72FDC44E9775BCBBC93002F380E">Lackner, Tomas</mods:namePart>
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<mods:date id="5ADFEEE5BED94B00BDA463B9927BC8F0">2015</mods:date>
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<mods:number id="82360CE7EDA207BEFCB72692272F8ECF">479</mods:number>
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<mods:classification id="C176664215302E20E12EEC784BDF6A1D">journal article</mods:classification>
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<mods:identifier id="B3B91C9CB026ED73B8A8C1E94A7C6935" type="DOI">http://dx.doi.org/10.3897/zookeys.479.8738</mods:identifier>
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<treatment id="C0972B290BCF50C160B085B724DC685D" ID-GBIF-Taxon="152058089" LSID="urn:lsid:plazi:treatment:C0972B290BCF50C160B085B724DC685D" httpUri="http://treatment.plazi.org/id/C0972B290BCF50C160B085B724DC685D" lastPageId="10" lastPageNumber="75" pageId="3" pageNumber="68">
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<subSubSection id="795577BE33C884A6E7A503216B3D0475" pageId="3" pageNumber="68" type="multiple">
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<paragraph id="F1B19B03352EF1DC6C5412463D50171B" pageId="3" pageNumber="68">Taxon classification Animalia Coleoptera Histeridae</paragraph>
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</subSubSection>
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<subSubSection id="605DEA49AD25F0E15A51A748EF7FDED0" pageId="3" pageNumber="68" type="nomenclature">
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<paragraph id="EF5960F364A2588209173F1BAB73BA82" pageId="3" pageNumber="68">
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<taxonomicName id="50DD0019CDBF0F3E85008780677081CB" ID-CoL="3DJJH" authority="Marseul, 1855" authorityName="Marseul" authorityYear="1855" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes (Marseul, 1855)</taxonomicName>
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Figs 1, 2-9, 10-13, 14-16, 17-25, 26-34
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</paragraph>
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</subSubSection>
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<subSubSection id="A375493BCF792B4AF8C4B6BFD3368BC8" pageId="3" pageNumber="68" type="reference_group">
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<paragraph id="F1D0F305019D41A63367C89F7F8C1BE7" pageId="3" pageNumber="68">
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<taxonomicName id="D0B1C14FB6836D5EFFF7C829AFF08245" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus grossipes" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="grossipes">Saprinus grossipes</taxonomicName>
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Marseul 1855: 718, t. XX, fig. 153; Schmidt (1885): 315.
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</paragraph>
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<paragraph id="4BD043D61E5D68CED42D210C3B8CB982" pageId="3" pageNumber="68">
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<taxonomicName id="0614244C4412288A52E9B20B58BAF24B" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus rugicollis" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="rugicollis">Saprinus rugicollis</taxonomicName>
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Schmidt 1890: 19 (nomen nudum, given as synonym).
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</paragraph>
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<paragraph id="B55508DE12A3F4CBB007614315B9481C" pageId="3" pageNumber="68">
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<taxonomicName id="3D3DDBE8E95861E4C7CCA0C8927F136A" class="Insecta" family="Histeridae" genus="Pachylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylopus grossipes" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="grossipes">Pachylopus grossipes</taxonomicName>
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:
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<bibRefCitation id="C430D7B6CF33D217C55ADA33425C8C5B" author="Schmidt, J" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="33" pageNumber="98" pagination="205 - 207" title="Description d'un Histeridae algerien du genre Pachylopus Er. et remarques sur la composition de ce genre (Col.)." volume="1896" year="1896">Schmidt (1896)</bibRefCitation>
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: 296;
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<bibRefCitation id="C36D18B603B5B25684A0258306EFD592" author="Mueller, G" journalOrPublisher="Memoire de la Societa Entomologica Italiana" pageId="33" pageNumber="98" pagination="93 - 104" title="Note sugli Histeridae della Tripolitania e Tunisia raccolti durante i viaggi organizzati da S. A. Serenissima il Principe Alessandro della torre tasso." volume="10" year="1931">
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G.
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<normalizedToken id="62FCDAD64EAA530EBEBC5165626F6171" originalValue="Müller">Mueller</normalizedToken>
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||
(1931)
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||
</bibRefCitation>
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: 102.
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</paragraph>
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<paragraph id="6518A62EAEF6E413A1199794D28294DE" pageId="3" pageNumber="68">
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<taxonomicName id="E9C27C0E38E9E3529E9760420427B131" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus grossipes" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="grossipes">Hypocaccus grossipes</taxonomicName>
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:
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<bibRefCitation id="D42DC7C381E1C3E918210E60D601F92A" author="Ganglbauer, L" journalOrPublisher="Carl Gerolds Sohn, Wien" pageId="32" pageNumber="97" title="Die Kaefer von Mitteleuropa. Die Kaefer der oesterreichisch-ungarischen Monarchie, Deutschland, der Schweiz, sowie des franzoesischen und italienischen Alpengebiets. III, 1. Familienreihe Staphylinoidea. 2 Theil: Scydmaenidae, Silphidae, Clambidae, Leptinidae, Platypsyllidae, Corylophidae, Sphaeritidae, Trichopterigydae, Hydroscaphidae, Scaphidiidae, Histeridae." year="1899">Ganglbauer (1899)</bibRefCitation>
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: 393.
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</paragraph>
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<paragraph id="8B75762F7B3BF8A8CCC40BEE23824FE1" pageId="3" pageNumber="68">
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<taxonomicName id="66825B37BA9D0F3911A1878A954AB114" class="Insecta" family="Histeridae" genus="Styphrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Styphrus grossipes" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="grossipes">Styphrus grossipes</taxonomicName>
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:
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<bibRefCitation id="FB7CA40E964898F164CF55E9D19E93C7" author="Jakobson, GG" journalOrPublisher="A. F. Devrien, St. Petersburg" pageId="32" pageNumber="97" title="Zhuki Rosii i zapadnoy Evropy. Rukovodstvo k opredeleniyu zhukov" year="1911">Jakobson (1911)</bibRefCitation>
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: 651.
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</paragraph>
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<paragraph id="1EAF2EDCD91B027634BF252445065952" pageId="3" pageNumber="68">
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<taxonomicName id="111A7C20B5A63E7C9C0CEBC6F1F02ABF" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
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<bibRefCitation id="D81FE0C459A7EC504C29B45AE49E153A" author="Reichardt, A" journalOrPublisher="Entomologische Blaetter" pageId="33" pageNumber="98" pagination="12 - 18" title="Ueber die mit Pachylopus verwandten Arten." volume="22" year="1926">Reichardt (1926)</bibRefCitation>
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: 16;
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<bibRefCitation id="F93F87A88F9723508F19363D226CBF54" author="Reichardt, A" journalOrPublisher="Nauka, Moskva-Leningrad" pageId="33" pageNumber="98" title="Semeystva Sphaeritidae i Histeridae" volumeTitle="Fauna SSSR, Nasekomye Zhestokrylye" year="1941">Reichardt (1941)</bibRefCitation>
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||
: 329, 330, fig. 117;
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<bibRefCitation id="02456A8BA9301AB24ED00FEFBED320B9" author="Peyerimhoff, P" journalOrPublisher="Bulletin de la Societe Entomologique d'Egypte" pageId="33" pageNumber="98" pagination="213 - 228" title="Les Saprinines Sabulicoles du Nord de l'Afrique (Coleoptera: Histeridae)." volume="20" year="1936">Peyerimhoff (1936)</bibRefCitation>
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: 227;
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<bibRefCitation id="5ABAA62E3956E1D7BA7F0ACC946878D2" author="Kryzhanovskij, OL" journalOrPublisher="Nauka, Leningrad" pageId="32" pageNumber="97" title="Zhuki Nadsemeystva Histeroidea (semeystva Sphaeritidae, Histeridae, Synteliidae)." volumeTitle="Fauna SSSR, Zhestokrylye" year="1976">Kryzhanovskij and Reichardt (1976)</bibRefCitation>
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: 232, Figs 455-458;
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<bibRefCitation id="5938BDD56AA1EF34ED2100F168BB536D" author="Vienna, P" journalOrPublisher="Calderini Press, Bologna" pageId="33" pageNumber="98" title="Fauna d'Italia. Vol. XVI. ColeopteraHisteridae" year="1980">Vienna (1980)</bibRefCitation>
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: 196, fig. 69;
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<bibRefCitation id="B09C70DA6981485E0EDEE29AD453D340" author="Mazur, S" journalOrPublisher="Academic Press, Budapest" pageId="33" pageNumber="98" title="Sutabogarak - Histeridae. 14. Fuezet. In: Fauna Hungariae. 138. VII Koetet." year="1980">Mazur and Kaszab (1980)</bibRefCitation>
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||
: 61, Figs 31, 34 D, E, F;
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<bibRefCitation id="D2F15F16EB631236EEB510D9722ED2F0" author="Mazur, S" journalOrPublisher="Polskie Pismo Entomologiczne" pageId="33" pageNumber="98" pagination="1 - 376" title="A world catalogue of Histeridae." volume="54" year="1984">Mazur (1984)</bibRefCitation>
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||
: 101;
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||
<bibRefCitation id="1F0AD921F7BCB61F894F0A9A1BA7A9BB" pageId="3" pageNumber="68">Mazur (1997)</bibRefCitation>
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||
: 263;
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<bibRefCitation id="EDD5795D580F094B320AA8FE35D8A0AF" author="Yelamos, T" editor="Ramos, MA" journalOrPublisher="Museo Nacional de Ciencias Naturales, CSCI, Madrid" pageId="33" pageNumber="98" title="Fauna Iberica" year="2002">
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||
<normalizedToken id="EA93D4AFB1C3A3FAC1FE6617285B4C70" originalValue="Yélamos">Yelamos</normalizedToken>
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||
(2002)
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||
</bibRefCitation>
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||
: 338, Figs 12E, 161G, 169, 170A;
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<bibRefCitation id="D45D62E053EC23215336680955686573" author="Mazur, S" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="33" pageNumber="98" pagination="68 - 102" title="Family Histeridae." volumeTitle="Catalogue of Palaearctic Coleoptera, Vol. 2, Hydrophiloidea-Histeroidea-Staphylinoidea" year="2004">Mazur (2004)</bibRefCitation>
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: 92;
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||
<bibRefCitation id="AF80432A9DD5CFAEFA8C78A6BDDC07A9" author="Lackner, T" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="32" pageNumber="97" pagination="1 - 254" title="Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae)." volume="50" year="2010">Lackner (2010)</bibRefCitation>
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||
: 112, Figs 19, 54, 89, 118, 339-359;
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<bibRefCitation id="4E6473B59626CB33A6CE990081484DE3" author="Mazur, S" journalOrPublisher="Warsaw University of Life Sciences, SGGW Press, Warsaw" pageId="33" pageNumber="98" title="A concise catalogue of the Histeridae (Coleoptera)." year="2011">Mazur (2011)</bibRefCitation>
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: 210.
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</paragraph>
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<paragraph id="D08CD85148E733BE0BA4F02845856F28" pageId="3" pageNumber="68">
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<taxonomicName id="0CB6575CCB97DDE3C0AD6A16F3D37E11" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="subspecies" species="grossipes" subspecies="berberus">Exaesiopus grossipes berberus</taxonomicName>
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<bibRefCitation id="862151AB86BE06F749BC1BF0335D85B4" author="Peyerimhoff, P" journalOrPublisher="Bulletin de la Societe Entomologique d'Egypte" pageId="33" pageNumber="98" pagination="213 - 228" title="Les Saprinines Sabulicoles du Nord de l'Afrique (Coleoptera: Histeridae)." volume="20" year="1936">Peyerimhoff 1936</bibRefCitation>
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: 227 - syn. n.
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</paragraph>
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</subSubSection>
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<subSubSection id="A6818C835FC3B7CFB7F05DF9DB68AF38" pageId="3" pageNumber="68" type="type locality">
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<paragraph id="83D78B7CFA7EB50971D246ABA93259E1" pageId="3" pageNumber="68">Type locality.</paragraph>
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<paragraph id="4BA8B02B46BB00C5174F59C39BB33EE1" pageId="3" pageNumber="68">Spain, France: Bayeux, Marseille.</paragraph>
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</subSubSection>
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<subSubSection id="AAA2DA2482598003100EE3A742B3DCAC" pageId="3" pageNumber="68" type="type material examined">
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<paragraph id="B2C760AA37A5F40AD04D11600D82E945" pageId="3" pageNumber="68">Type material examined.</paragraph>
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<paragraph id="DC45082BCDA958E05AFA2AE0F6253825" pageId="3" pageNumber="68">
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<taxonomicName id="3609908C0F7715D8B5FF2C978DF220AE" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus grossipes" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="grossipes">Saprinus grossipes</taxonomicName>
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: Lectotype, present designation, sex undetermined, pinned, right mesotibia, left mesotarus, both hind legs missing, with the following labels: "153 /
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<taxonomicName id="36E561F335D73A30CC98EE0AA14F2504" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
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/
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<taxonomicName id="E810F3459FF0E48C4E8ADECC61681BE0" lsidName="grossipes" pageId="3" pageNumber="68" rank="species" species="grossipes">grossipes</taxonomicName>
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/ m / Marseille / Barage?" (round illegible label, written); followed by: "MUSEUM PARIS / COLL. / DE MARSEUL 1890" (printed); followed by:
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<normalizedToken id="99068A876E3080583DD4A8247245B3DC" originalValue="“TYPE”">"TYPE"</normalizedToken>
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(red-printed label); followed by: "
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<taxonomicName id="D603BCD08EC9159473C273A4798B15CA" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus grossipes" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="grossipes">Saprinus grossipes</taxonomicName>
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/ Marseul, 1855 / LECTOTYPE 2014 / des. T. Lackner" (red label, written).
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</paragraph>
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<paragraph id="4A92CA22507C3A840A9AA5A897055A01" pageId="3" pageNumber="68">
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<taxonomicName id="59D2C91A415F1855F921656A2AFC60D8" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="subspecies" species="grossipes" subspecies="berberus">Exaesiopus grossipes berberus</taxonomicName>
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: Lectotype, ♀, side-mounted on a triangular point, final two metatarsomeres on right hind leg missing, with the following labels:
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<normalizedToken id="A21636E9985A314A4A5C487EF0A98028" originalValue="“Laghouat”">"Laghouat"</normalizedToken>
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||
(written); followed by: "Coll.
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<normalizedToken id="3656CB1569B321F4B8E0BD69774F75E8" originalValue="Hénon">Henon</normalizedToken>
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||
/ T
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<normalizedToken id="AC22EAA0A4E6C3E2256595F3157D6559" originalValue="Théry”">Thery"</normalizedToken>
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(written); followed by: "
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<taxonomicName id="19AEDEBF70D26FAA4229769A5444F4E7" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
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/
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<taxonomicName id="6628024825A86E99E521C9520DA60D57" lsidName="grossipes" pageId="3" pageNumber="68" rank="subspecies" species="grossipes" subspecies="berberus">grossipes berberus</taxonomicName>
|
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/ Peyerimhoff / TYPE" (written); followed by: "
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||
<taxonomicName id="AD46B13A3E0AD6F650D23ADA8388B4AB" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="genus">Exaesiopus</taxonomicName>
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/
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<taxonomicName id="37CD0194B4130BEEBB1E411AA1C2F03A" lsidName="grossipes" pageId="3" pageNumber="68" rank="subspecies" species="grossipes" subspecies="berberus">grossipes berberus</taxonomicName>
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/ Peyerimhoff, 1936 / LECTOTYPE / des. T. Lackner 2014" (red label, written) (MNHN). Paralectotypes, 2 ♂♂, both mounted on a single pin on triangular mounting points with extracted genitalia, with the following label:
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<normalizedToken id="6883F42D45F02F0DE902960589AEDAA2" originalValue="“Bône”">"Bone"</normalizedToken>
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||
(written); followed by: "Coll.
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||
<normalizedToken id="2EEB4B86D10B4B451A279288633C0EBF" originalValue="Hénon">Henon</normalizedToken>
|
||
/ T
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||
<normalizedToken id="2D998823513D1EC61DDE3A2F0633676E" originalValue="Théry”">Thery"</normalizedToken>
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(written); followed by: "
|
||
<taxonomicName id="F39C60096077B2A8F16D9512811BACD4" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus grossipes" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="grossipes">Saprinus grossipes</taxonomicName>
|
||
/
|
||
<taxonomicName id="DA63BCBC941C2149C672C158924282AA" lsidName="berberus" pageId="3" pageNumber="68" rank="subspecies" species="berberus" subspecies="berberus">berberus</taxonomicName>
|
||
Peyerimh / TYPE" (written); followed by: "
|
||
<taxonomicName id="87400D49660F89314B85DEF843321CB2" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="genus">Exaesiopus</taxonomicName>
|
||
/
|
||
<taxonomicName id="2E420162076D93EC072A0D39C7FD6D8E" lsidName="grossipes" pageId="3" pageNumber="68" rank="subspecies" species="grossipes" subspecies="berberus">grossipes berberus</taxonomicName>
|
||
/ Peyerimhoff, 1936 / PARALECTOTYPE / des. T. Lackner 2014" (red label, written) (MNHN).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1589A7A3A6743D8C39B57E21E121BF3" pageId="3" pageNumber="68" type="note">
|
||
<paragraph id="31FE45995C44ECEB3DA9354895E4A772" pageId="3" pageNumber="68">Note.</paragraph>
|
||
<paragraph id="C2476208FF333BC7F5A61DFA14E8FB39" pageId="3" pageNumber="68">
|
||
<bibRefCitation id="DBFA279E5971A1A2929246BA2EBE8A61" author="Peyerimhoff, P" journalOrPublisher="Bulletin de la Societe Entomologique d'Egypte" pageId="33" pageNumber="98" pagination="213 - 228" title="Les Saprinines Sabulicoles du Nord de l'Afrique (Coleoptera: Histeridae)." volume="20" year="1936">Peyerimhoff (1936</bibRefCitation>
|
||
: 227) distinguished the subspecies
|
||
<taxonomicName id="220731EE443FC986E02707519C0C7F22" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="subspecies" species="grossipes" subspecies="berberus">Exaesiopus grossipes berberus</taxonomicName>
|
||
from the nominotypical one based on elytral punctation that should cover almost the entire elytral disc basally and laterally up to the second dorsal elytral stria. Among the three specimens that he furnished with type labels, however, only the female from Laghouat (Algeria) exhibits these characteristics; the two males from
|
||
<normalizedToken id="47125BAAC49F000B9615B05CA01BC17B" originalValue="Bône">Bone</normalizedToken>
|
||
[=Annaba, Algeria] have their apical half (approximately) impunctate and the punctation of their elytral discs terminates in third elytral stria. Therefore, a female from Laghouat has been selected for the Lectotype.
|
||
<bibRefCitation id="E388000C7C855F3A964357F8E7B2FCA3" author="Peyerimhoff, P" journalOrPublisher="Bulletin de la Societe Entomologique d'Egypte" pageId="33" pageNumber="98" pagination="213 - 228" title="Les Saprinines Sabulicoles du Nord de l'Afrique (Coleoptera: Histeridae)." volume="20" year="1936">Peyerimhoff (1936</bibRefCitation>
|
||
: 227) himself did not specify which specimen(s) belonged to the type series; he listed several localities with his extremely brief diagnosis of the new subspecies. Both Laghouat and
|
||
<normalizedToken id="8D81C02C0CBB70C5B5B9E3B68933BB98" originalValue="Bône">Bone</normalizedToken>
|
||
[=Annaba] were among the listed localities.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="47C9A52449879182E9359C1CFDF2C359" lastPageId="5" lastPageNumber="70" pageId="3" pageNumber="68" type="additional material examined">
|
||
<paragraph id="224A6346D8CFBC5ACE15C64C6AD089F6" pageId="3" pageNumber="68">Additional material examined.</paragraph>
|
||
<paragraph id="0ECD24CF9B2CD11AAEC490589E766F73" lastPageId="5" lastPageNumber="70" pageId="3" pageNumber="68">
|
||
Bulgaria: 1 ♀, Asenovgrad, vi.1963, A. Olexa leg. (TLAN); 1 spec., Nessebar, 30.v.1996, O. Majzlan leg. (TLAN); 1 spec., Kazanlak, vi. 1963, A. Olexa leg. (TLAN); 1 spec., Plovdiv,
|
||
<normalizedToken id="D34014026BCFCBE91F1F03D3F22874FE" originalValue="Rektořík">Rektorik</normalizedToken>
|
||
leg. (TLAN); 1 ♂, Newrokop, 19.vi.1938, leg. Hlinikowski (TLAN); 1 spec., SW Bulgaria, 2 km N Gara Pirin, 11.-12.vi. 1983, leg. Hieke (NCB). BOSNIA-HERZEGOVINA: 1 ♀, Mostar, V. Zoufal leg. (ZMHUB). MACEDONIA: 2 specs., 5 km E of Velandovo, 31.v.1992, P.
|
||
<normalizedToken id="EA26840C9E640D1ABF4F6970B0FCF5A8" originalValue="Zahradník">Zahradnik</normalizedToken>
|
||
leg. (TLAN). Slovakia: 1 ♂, 1 ♀ + 1 spec.,
|
||
<normalizedToken id="98CCDB74C12A5938E2908CDCD16893B1" originalValue="Čenkov">Cenkov</normalizedToken>
|
||
, 24.vi. 1987, V.
|
||
<normalizedToken id="D450E058C0B20E862460646BA46C17DD" originalValue="Kubáň">Kubaň</normalizedToken>
|
||
lgt. (TLAN); 1 spec., ibid, but, 29.v.1993, T.
|
||
<normalizedToken id="33CC633F6A1A60DE9BAB7C371B565CB9" originalValue="Růžička">Ruzicka</normalizedToken>
|
||
leg. (TLAN); 2 specs.,
|
||
<normalizedToken id="72845E6541ABAEFBB832C43433403551" originalValue="Kameničná-Balvany">Kamenicna-Balvany</normalizedToken>
|
||
8174c, 13.vii.2000, O. Majzlan leg. (TLAN). Serbia: 1 spec., Veliko
|
||
<normalizedToken id="15C4C1E08478FDD6862051A175FEA876" originalValue="Gradište">Gradiste</normalizedToken>
|
||
, 2.ix.1955,
|
||
<normalizedToken id="8B78C992F9DB471A4CDD90C0D581CD9A" originalValue="Stanćić">Stancic</normalizedToken>
|
||
leg. (TLAN); 1 spec., Vranje, 4.vi.1968, collector unknown (TLAN). ITALY: 1 ♀, Piemont, coll. Bickhardt (MNHN); 1 ♂, Torino, coll. Fea (MNHN); 1 ♂ +1 ♀, Lagnola (?), xi. 1910, Sekera leg. (TLAN); 1 spec., PO, Fiume, Piacenza, 2.vi.1963, leg. P. Ratti (NCB); 1 spec., Veneto, Caorle, v. 1999, Clereau leg. (CAS); 2 ♂♂ + 1 ♀, Ponferrada, Paganetti (ZMHUB). FRANCE: 1 ♂,
|
||
<normalizedToken id="5671F9F079FC87A8BE0EA3F066AED83D" originalValue="Bouches-du-Rhône">Bouches-du-Rhone</normalizedToken>
|
||
, Les Saintes-Maries-de-la-Mer, 18.iv.1978, P. Queney leg. (CYG); 1 ♀, Charente Maritime,
|
||
<normalizedToken id="1D6320A34E3FBB3029CFED0BC0AC1EF9" originalValue="île">ile</normalizedToken>
|
||
de
|
||
<normalizedToken id="B194F5F6D0A631E26D16FE29059D2B11" originalValue="Ré">Re</normalizedToken>
|
||
, viii.1978 (P. Queney leg. (CYG); 1 ♂+1 ♀, Gard, Le Grau-du-Roi, 31.viii.1947, J.
|
||
<normalizedToken id="62423C1CEF30ED2E8F5A6EABD9DB1DA0" originalValue="Thérond">Therond</normalizedToken>
|
||
leg. (CYG); 4 ♂♂+1 ♀, Gironde, Soulac, 12.iv.1890, E. Giraud leg. (CYG); 1 ♀, idem, but 6.vii.1975, G.
|
||
<normalizedToken id="31BB79497823F098445E519ECE95F75C" originalValue="Tempère">Tempere</normalizedToken>
|
||
leg. (CYG); 1 ♀, Manche, Portbail, beach, 22.vi.1955, H. Chevin leg. (CYG); 1 ♂,
|
||
<normalizedToken id="C127A99D544EB3A4C73E4A7204F28FE4" originalValue="Pyrénées">Pyrenees</normalizedToken>
|
||
, (CYG); 1 ♂, Montelimar, 7.v.1912, Laboissere leg. (MNHN); 1 ♀, Plouharnel, 1878, no further data (MNHN); 5 specs., Bretagne, Nicolas, no further data (MNHN); 1 ex., Var, Nice, flooding, v. 1951 (MNHN); 3 specs., Grande
|
||
<normalizedToken id="080C85E6143E21D5C3A17D050E9C49AC" originalValue="Coté">Cote</normalizedToken>
|
||
, Royau, v. 1918, Chobaut & R. Lebon (MNHN); 1 spec., Aveyron, Millau, 6.iv.1960, Fages lgt. (MNHN); 2 specs., Ile de
|
||
<normalizedToken id="2C7F3C357567D5BB4230D60250C6CF77" originalValue="Ré">Re</normalizedToken>
|
||
, coll. Bonnaire, no further data (MNHN); 3 specs., La Rochelle
|
||
<normalizedToken id="110FB748C4447C1DCD3268936EA0686A" originalValue="d’Orbigny">d'Orbigny</normalizedToken>
|
||
, no further data (MNHN); 4 specs., Carcassonne, no further
|
||
<pageBreakToken id="B3716E8E364C5693DEDFBE1D30C30AD6" pageId="4" pageNumber="69" start="start">data</pageBreakToken>
|
||
(MNHN); 2 specs., Pluharnel, dept. de Morbihan, no further data (MNHN); 2 specs., St. Jean de Monts, P. Sirguey (MNHN); 1 spec., Agen, 30.v.1908, G. Nicolas (MNHN); 2 specs., Toulouse, Col. D. Grenier (MNHN); 1 spec., Grau du Roi, 11.iv.1955, J.
|
||
<normalizedToken id="6D5ED1C540658C2F4267DE698048AFA9" originalValue="Thérond">Therond</normalizedToken>
|
||
leg. (MNHN); 2 specs., ibid, but 2.ii.1938, J.
|
||
<normalizedToken id="86761249E774C1E3070E37161B22BC96" originalValue="Thérond">Therond</normalizedToken>
|
||
leg. (MNHN); 1 spec., ibid, but 10.ix.1948, J.
|
||
<normalizedToken id="DBC9CD524C330749932653BC70B1089A" originalValue="Thérond">Therond</normalizedToken>
|
||
leg. (MNHN); 1 spec., ibid, but 19.v.1970, J.
|
||
<normalizedToken id="5DEAC9F35E63BA0488DF0C1008DB9F82" originalValue="Thérond">Therond</normalizedToken>
|
||
leg. (MNHN); 1 spec., ibid, but 15.v.1951, J.
|
||
<normalizedToken id="03FDC66548AB79D7CD43C9B0DCA86778" originalValue="Thérond">Therond</normalizedToken>
|
||
leg. (MNHN); 1 spec., ibid, but 1.x.1949, J.
|
||
<normalizedToken id="D4D27962C969ED40570BE6CFD66C8D87" originalValue="Thérond">Therond</normalizedToken>
|
||
leg. (MNHN); 2 specs., ibid, but 9.iii.1940, J.
|
||
<normalizedToken id="B8BDED61AB7F292BA8B34D33FC6F63B8" originalValue="Thérond">Therond</normalizedToken>
|
||
leg. (MNHN); 2 specs., Pont du Gard, 3.iii.1927, J.
|
||
<normalizedToken id="A91F5299440CA8B72ABA1288CB033DC4" originalValue="Thérond">Therond</normalizedToken>
|
||
leg. (MNHN); 2 specs., ibid, but 29.v.1928, J.
|
||
<normalizedToken id="129DA4E11A220210DB170F08830D1952" originalValue="Thérond">Therond</normalizedToken>
|
||
leg. (MNHN); 2 specs., ibid, but 22.ix.1931, J.
|
||
<normalizedToken id="7F16C25F1D68C55E06E4AB7372081ED7" originalValue="Thérond">Therond</normalizedToken>
|
||
leg. (MNHN); 1 spec., Camargue, St. Maries, 9.x.1928, L. Puel leg. (MNHN); 5 specs.,
|
||
<normalizedToken id="DC45D00D10B432C5033F6361787647F8" originalValue="Vendée">Vendee</normalizedToken>
|
||
, St. Jean de Monts, vi.1926, P. Sirguey leg. (NCB); 2 ♂♂ + 1 ♀, ibid, but MNHN; 1 spec., Morgat, Brittany, no further data (BMNH); 2 specs., France, no further data (BMNH); 1 spec., Erqny,
|
||
<normalizedToken id="F41909140D99B47D1A57B8E153C7B895" originalValue="Côtes">Cotes</normalizedToken>
|
||
du Nord, H.D. Preston leg. (BMNH); 1 spec., Provence, no further data (BMNH); 3 specs., St. Jean de Monts (
|
||
<normalizedToken id="EBC255F745E949F921BA967570B79752" originalValue="Vendée">Vendee</normalizedToken>
|
||
), P. Sirguey leg., 1926 (BMNH); 1 spec., Manche, Utah Beach, 6.vi.[19]64; 1 spec., Beziers, no further data (BMNH) 1 spec., ibid but ZMHUB; 1 spec., Grau du Roi, 29.iii.1943, J.
|
||
<normalizedToken id="DE714893682751286EC55C584B1972C7" originalValue="Thérond">Therond</normalizedToken>
|
||
leg. (ZIN); 1 spec., Lyon, in
|
||
<normalizedToken id="E8CD59F64BF55AC41E5D1620A01F8036" originalValue="Rhône">Rhone</normalizedToken>
|
||
, no data or collector (ZIN). HUNGARY: 1 spec., Hungary, no further data (BMNH); 1 ♂,
|
||
<normalizedToken id="3A198705CB6EF6BAE4CA95AFF80D40E0" originalValue="Jarabszállás">Jarabszallas</normalizedToken>
|
||
, 30.v.1971, P.
|
||
<normalizedToken id="0015A8FDEE77A5A1DAD0707A4C695AE4" originalValue="Polák">Polak</normalizedToken>
|
||
leg. (TLAN); 1 ♂, Dunakeszi, no further data (ZMHUB). SPAIN: 1 spec., Valencia, no further data (MNHN). IRAQ: 1 spec., Mesopotamia, Millingen, no further data (BMNH); 1 ♀, Mesopotamia, no further data (ZMHUB). Russia: 1 spec., Volgogradskaya obl., Tsimlya, 27.vii.1894, collector unknown (ZIN). Ukraine: 1 spec., Stan. Luganskaya, Lugansk okr., 17.vi.1928, collector unknown (ZIN); 2 ♀♀, Khersonskaya oblast, Alyoshki, Dneprovskij uezd, 26.v.1926, D. Znojko leg.; 2 specs., ibid, but 19.v.1929, N. Kostenko leg.; 1 spec., Khersonskaya oblast, Burkutskie plavni [zapovednik], 17.v.1929, N. Kostenko leg.; 1 spec., ditto, but Kazach village, 7.vi.1928, N. Kostenko leg. (all exs. ZIN). Greece: 1 spec., Peloponese occid., Epitalion, Alfios River, nr. Pyrgos, 13.iv.1995, T.
|
||
<normalizedToken id="428826BECD085C4BFCF91C24CF95B2BB" originalValue="Kopecký">Kopecky</normalizedToken>
|
||
leg. (TLAN); 1 ♀, Pirgos, 1.v.1971, leg. Wewalka leg. (TLAN); 1 spec., Peloponesus, Xylokatron, 22.v.1962, H. Pochon leg. (MNHN); 1 spec., Thessalia, no further data (ZMHUB). MOROCCO: 1 ♀, Tauorirt, 10.iii.1993, G. Chavanon leg. (CYG); 1 ♂, Morocco centr., Moyen Atlas, Aguelmame Azegza lake, 22.-26.vi.1998, T. Lackner leg. (TLAN); 1 spec., Ouarzazate prov., Oued Draa River valley, Agdz env., N 30.40.52 W 006.25.08, 29.iii.2011, in human faeces, A. Gusakov leg. (CAS); 1 ♂, Beni Ounif near Figuig, 11.v.1944, Barbier leg. (MNHN). TUNISIA: 1 ♂, Medjez el Bab, v.1935, R. Demoflys leg. (CYG); 1 ♂,
|
||
<normalizedToken id="397A53182339B530EDE5CC7A482CEA54" originalValue="Gabès">Gabes</normalizedToken>
|
||
, v. 1944, R. Demoflys leg. (CYG); 1 spec., Zarzia, 5.-11.v.1977, M.A. Hielkema leg. (NCB). 1 ♀, Tunis,
|
||
<normalizedToken id="5AB9E249CB57966E872B2C0FA384F67F" originalValue="i–ii">i-ii</normalizedToken>
|
||
.1882, G. & L. Doria leg. (ZMHUB); 1 spec., Hammamet mer., 25.iii.-4.iv.1992, A.
|
||
<normalizedToken id="0DBB0E65325078B3F4173EAA71AC561F" originalValue="Pütz">Puetz</normalizedToken>
|
||
leg.; 2 ♀♀, 1 ♂ & 1 spec., 6-11.vi.1982, Kairuan, A. Olexa leg. (TLAN). ALGERIA: 1 ♀, Bona [=
|
||
<normalizedToken id="7CEB2C41AD36213D617A80FAD52D12A7" originalValue="Bône">Bone</normalizedToken>
|
||
?], Desbr., no further data (ZMHUB); 2 ♂ + 1 ♀, Oued Sebaou near Tizi-Ouzou, 25.vi.1908, collector unknown (MNHN); 2 ♀♀,
|
||
<normalizedToken id="0B3256AFE617EE84657A7C64F7FFE8A2" originalValue="Aïn">Ain</normalizedToken>
|
||
Sefra, v. 1936, collector unknown (MNHN); 1 ♂, ibid, but coll. Bonnaire (MNHN); 1 ♀, Bou-Ktoub, S of Oran,
|
||
<normalizedToken id="DF5756833B8DF277DBA3FF2206424B82" originalValue="Déchoguat">Dechoguat</normalizedToken>
|
||
leg. (MNHN); 1 ♀, Biskra, v. 1885, L. Bleuse leg. (MNHN);
|
||
<pageBreakToken id="04022C53EFFA7328E804FB07EBF6B27E" pageId="5" pageNumber="70" start="start">1</pageBreakToken>
|
||
♀, Bou-Saada, no further data (MNHN); 1 ♀, south of Oran, no further data (MNHN) 1 ♂ + 1 ♀,
|
||
<normalizedToken id="1DE24C4E129901C2B9094BD243F24D4B" originalValue="Colomb-Béchar">Colomb-Bechar</normalizedToken>
|
||
, 1912, P. Germain leg. (MNHN).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="795626E4382F4A5668653B051DB011DF" lastPageId="10" lastPageNumber="75" pageId="5" pageNumber="70" type="redescription">
|
||
<paragraph id="49BDC407D6434A607AFB54C998C0E9B3" pageId="5" pageNumber="70">Redescription.</paragraph>
|
||
<paragraph id="B2EE2747559D6E8979CA2C592323250C" pageId="5" pageNumber="70">
|
||
Although this species has been recently re-described by the author (
|
||
<bibRefCitation id="D5671E751BEB57862CDCD4DA5D95E405" author="Lackner, T" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="32" pageNumber="97" pagination="1 - 254" title="Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae)." volume="50" year="2010">Lackner 2010</bibRefCitation>
|
||
: 112-116), I prefer to repeat this re-description here for the reason that the following species are morphologically rather similar to
|
||
<taxonomicName id="C5FCA5A9E8E3154A46881EDC5FA06A69" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="5" pageNumber="70" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
|
||
. Those species are consequently provided only with diagnostic descriptions outlining their respective differences.
|
||
</paragraph>
|
||
<paragraph id="7167529DB492529D180B3B06E7ABE879" pageId="5" pageNumber="70">Body length: PEL: 2.10-2.75 mm; APW: 0.825-1.00 mm; PPW: 1.625-2.25 mm; EL: 1.25-2.00 mm; EW: 1.875-2.50 mm.</paragraph>
|
||
<paragraph id="4A32203DD22B92057DC9E107987C9341" pageId="5" pageNumber="70">Body (Fig. 1) oval, convex, cuticle light to dark brown, sometimes with feeble bronze or greenish metallic tinge; legs, mouthparts and antennae rufopiceous.</paragraph>
|
||
<caption id="84F671E403079BF0617C951DC3D094A7" pageId="5" pageNumber="70">
|
||
<paragraph id="3607727F68F2D34E460EB59AEF63B18B" pageId="5" pageNumber="70">
|
||
Figure 1.
|
||
<taxonomicName id="84781AB42C9A94E568307BAFB11AF57E" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="5" pageNumber="70" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
|
||
(Marseul, 1855) habitus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="421978E5C993490C7ADEBBEF4A4E4399" pageId="5" pageNumber="70">Antennal scape with few short setae; club (Fig. 2) round, entire surface with thick short yellow sensilla intermingled with sparse slightly longer setae; sensory structures of the antennal club (Fig. 14) in form of stipe-shaped vesicle situated under circular sensory area on internal distal margin of the ventral side of antennal club.</paragraph>
|
||
<caption id="85D8BCDE5EABC56100562F1B94E583AC" pageId="5" pageNumber="70">
|
||
<paragraph id="618B218A3F6E29E19ED230C5BC22B765" pageId="5" pageNumber="70">
|
||
Figures 2-9.
|
||
<taxonomicName id="E4A2CAB27FEB6560E27333D1BFAB305A" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="5" pageNumber="70" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
|
||
(Marseul, 1855) 2 head, dorsal view 3 mentum, ventral view 4 clypeus 5 propygidium + pygidium 6 prosternum 7 mesoventrite 8 lateral disc od metaventrite + metepisternum 9 protibia, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4DC824710263CBB11A4E7E26DB5C95A6" pageId="5" pageNumber="70">Mouthparts: mandibles (Fig. 15) stout, outer margin slightly curved; mandibular apex bluntly pointed; sub-apical tooth of left mandible large, almost perpendicular; labrum (Fig. 16) sparsely punctate, shallowly depressed medially, two labral pits present, two labral setae arising from each; epipharynx almost completely hidden under labral fold; terminal labial palpomere elongated, its width less than half its length; mentum (Fig. 3) square-shaped, with deep antero-median notch; anterior margin with few long setae, lateral margins with single row of sparse shorter ramose setae; cardo of maxilla with few short setae on lateral margin; stipes triangular, with three much longer setae; terminal maxillary palpomere somewhat thickened, its width less than half its length, about twice as long as penultimate.</paragraph>
|
||
<paragraph id="26426AC64CD0862FA09B828896028CD3" pageId="5" pageNumber="70">Clypeus (Fig. 4) rectangular, almost smooth, can be slightly rugose, anterior margin elevated, clypeus depressed medially; frontal stria well impressed, almost straight (sometimes somewhat curved outwardly), carinate, continued as carinate supraorbital and postorbital striae; frons with two to several irregularly shaped carinate transverse rugae or chevrons; eyes flattened, inconspicuous from above.</paragraph>
|
||
<paragraph id="AD8255F8B52C2FB1C498D2C232CF1970" pageId="5" pageNumber="70">Pronotal sides slightly convergent forwards; apical angles blunt; marginal stria complete; pronotal disc convex, with round dense punctation, forming transverse rugae laterally, postero-median part of disc usually smooth, at times entire disc punctate (punctation can also stop short of lateral pronotal margin); pronotal base with a double row of round dense punctures; pronotal hypomeron with amber setae; scutellum small, visible.</paragraph>
|
||
<paragraph id="BDDE9094F2E4B94B5288D5709BD3202F" lastPageId="6" lastPageNumber="71" pageId="5" pageNumber="70">
|
||
Elytral humeri slightly prominent, elytra broad, almost as broad as long at its widest point; elytral epipleura with microscopic punctures, almost smooth; marginal epipleural stria complete; marginal elytral stria deeply impressed, continued as well impressed apical elytral stria; regular row of round punctures present along elytral marginal stria. Humeral elytral stria weakly impressed on basal third, sometimes doubled; inner subhumeral stria present medially, deep and rather long, rarely joining marginal elytral stria; elytra with four dorsal punctate elytral striae 1-4, all striae approximately reaching elytral half
|
||
<pageBreakToken id="7648F208765EEB2A34D14FFC24E8189E" pageId="6" pageNumber="71" start="start">apically</pageBreakToken>
|
||
(occasionally slightly surpassing it), fourth elytral stria basally connected with sutural elytral stria; sutural stria deeply punctured, apically joining apical elytral stria. Elytral punctation variable, often confined to apical half of elytra, along elytral suture reaching almost anterior third of elytral disc, punctures regular and deep, separated by about half to their own diameter, occasionally (often in specimens from North Africa) covering most part of elytral disc (elytral flanks and humeri almost always smooth).
|
||
</paragraph>
|
||
<paragraph id="AEA62D1333BC1AFC678087D24F4F58F7" pageId="6" pageNumber="71">Propygidium (Fig. 5) almost completely exposed, long, covered with coarse and dense regular punctation; punctation of pygidium (Fig. 5) sparser and finer, punctures separated by about 1-3 times their diameter.</paragraph>
|
||
<paragraph id="57C93300E08F906F1A035EF3EAC13E8C" pageId="6" pageNumber="71">Anterior margin of median portion of prosternum (Fig. 6) regularly rounded; prosternal foveae weakly to well impressed, small and often indiscernible under conventional binocular microscope; prosternal process slightly to deeply concave, dorsally impunctate, laterally substrigulate-punctate, few microscopic setae present; carinal prosternal striae divergent between procoxae, subparallel, vaguely united in front, at times obliterated on their anterior third; lateral prosternal striae well impressed, carinate, convergent anteriorly, united in front of apices of carinal prosternal striae.</paragraph>
|
||
<paragraph id="1C2328C4498344C3F41AE8CAD8EF7FBF" lastPageId="7" lastPageNumber="72" pageId="6" pageNumber="71">
|
||
Mesoventral disc (Fig. 7) somewhat convex, almost smooth, slightly wider than long; meso-metaventral sutural stria well impressed, with several accompanying punc
|
||
<pageBreakToken id="D6320ADFA4F9EA6BB2F43467F87D3493" pageId="7" pageNumber="72" start="start">tures</pageBreakToken>
|
||
; intercoxal disc of metaventrite with longitudinal depression in male, smooth, basally with irregular sparse shallow fine punctures; lateral metaventral stria (Fig. 8) well impressed, carinate, obliquely arcuate, apically almost reaching metacoxa; lateral disc of metaventrite concave, with shallow setiferous punctures of various sizes, separated by approximately their own diameter; metepisternum with even denser and coarser punctation and setae, on apical third + metepimeron punctation much finer and sparser; metepisternal stria deeply impressed, present on metepimeron and approximately apical third of metepisternum.
|
||
</paragraph>
|
||
<paragraph id="B19E5D1B079D69D0938C4901FC6962D1" pageId="7" pageNumber="72">Intercoxal disc of first abdominal sternite almost completely striate laterally; disc almost smooth, with sparse punctures along apical margin; lateral portion of disc of all visible abdominal sternites with short setae.</paragraph>
|
||
<paragraph id="0C01831F5FE640173F2E91D134EF1AC0" lastPageId="8" lastPageNumber="73" pageId="7" pageNumber="72">
|
||
Protibia (Fig. 9) on outer margin with two to three low teeth, topped with triangular to rounded (blunt, if worn) denticle followed by two inconspicuous rounded denticles; setae of outer row sparse, moderately long; setae of median row shorter than those of outer row, sparse; anterior protibial stria shortened apically; protibial groove shallow; protibial spur (Fig. 10) minuscule, growing out from apical margin of protibia; outer part of posterior surface of protibia (Fig. 10) obscurely variolate, vaguely separated from comparatively narrower median part, posterior protibial stria complete,
|
||
<pageBreakToken id="6A749ED09D936D042F3836458ED9716E" pageId="8" pageNumber="73" start="start">terminating</pageBreakToken>
|
||
in two minute inner posterior denticles; inner margin of protibia with double row of short dense ramose setae.
|
||
</paragraph>
|
||
<paragraph id="18C4118EEA18DCF5BD0D1B7D4E48E592" lastPageId="9" lastPageNumber="74" pageId="8" pageNumber="73">
|
||
Mesotibia (Fig. 11) moderately dilated and thickened, outer margin with two rows of sparse short denticles; setae of outer row well sclerotized, comparatively short; setae of median row shorter and sparser, covering most of posterior surface;
|
||
<pageBreakToken id="475EFD3D16FC7BE453378C2649ECFC01" pageId="9" pageNumber="74" start="start">posterior</pageBreakToken>
|
||
mesotibial stria vaguely impressed, shortened apically; mesotibial spur stout, prominent and long; anterior face of mesotibia (Fig. 12) smooth; anterior mesotibial stria shortened apically; claws of last tarsomere bent, shortened, shorter than half its length.
|
||
</paragraph>
|
||
<caption id="BCED6228A7335E8D74C7C3A3ABBA20B2" pageId="9" pageNumber="74">
|
||
<paragraph id="70B5B6B81865AE978FDED98C7C07195E" pageId="9" pageNumber="74">
|
||
Figures 10-13.
|
||
<taxonomicName id="A29415F5512A46E2D074CA827C2896AD" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="9" pageNumber="74" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
|
||
(Marseul, 1855) 10 protibia, ventral view 11 mesotibia, dorsal view 12 ditto, ventral view 13 metatibia, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="D90CCC365AC1580E4DF3CBEB92902A28" pageId="9" pageNumber="74">
|
||
<paragraph id="DACE2081326A1C2DF0BF8F2D3AF1C93C" pageId="9" pageNumber="74">
|
||
Figure 14-16. 14
|
||
<taxonomicName id="59940CFADBE17A0D292147E242E8404B" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="9" pageNumber="74" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
|
||
(Marseul, 1855) sensory structures of the antenna 15 mandibles, dorsal view 16 labrum, left half depicting dorsum; right half depicting underside.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B476FD64101EFFEA190432E6EBA8FC1A" pageId="10" pageNumber="75">
|
||
<pageBreakToken id="5FBBC8CE6FC211B4C1DF2DA5D1A7D1D0" pageId="10" pageNumber="75" start="start">Metatibia</pageBreakToken>
|
||
(Fig. 13) triangularly dilated and thickened apically; outer margin with four widely-spaced short rounded denticles, a single row of tiny sparse rounded denticles present dorsally on thickened anterior face of metatibia; setae of intermedian row shorter and denser, cover almost the entire posterior face of metatibia; otherwise metatibia similar to mesotibia.
|
||
</paragraph>
|
||
<paragraph id="D878564530FCC8CD7AD6DF374FC7A2B8" pageId="10" pageNumber="75">
|
||
Male genitalia. Eighth sternite (Figs 17-18) on apical half longitudinally separated medially, with tiny asetose vela, eighth sternite and tergite fused laterally (Fig. 19). Ninth tergite (Figs 20-21) apically with faint emargination; basally deeply emarginated; tenth tergite (Fig. 20) apically outwardly arcuate, basally faintly inwardly arcuate. Spiculum gastrale (Figs 24-25) typical for the subfamily representing the most common type with
|
||
<normalizedToken id="D474B589EFB8B6E3F5C9ED447AE88450" originalValue="‘head’">'head'</normalizedToken>
|
||
and
|
||
<normalizedToken id="2B2A091966B56A2D87A57FA2949F89A8" originalValue="‘tail’">'tail'</normalizedToken>
|
||
(sensu Caterino & Tishechkin, 2013);
|
||
<normalizedToken id="AE913755D79D37D4CB878E3CC271A9F7" originalValue="‘tail’">'tail'</normalizedToken>
|
||
cordate,
|
||
<normalizedToken id="CCD503F16637E89C3ACE722376322160" originalValue="‘head’">'head'</normalizedToken>
|
||
with two narrow, curved arms. Aedeagus (Figs 22-23) tube-like, slender, basal piece of aedeagus rather short, ratio of its length: length of parameres 1:3.5; parameres fused along their basal two-thirds, aedeagus slightly curved ventrad (Fig. 23).
|
||
</paragraph>
|
||
<caption id="7B8A79019E2B12B92536EAC905C4E473" pageId="10" pageNumber="75">
|
||
<paragraph id="77A747DEB3D8C38DED162E061CC6D2EB" pageId="10" pageNumber="75">
|
||
Figures 17-25.
|
||
<taxonomicName id="FE8B03A99655223EB5BA363431DB6E00" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
|
||
(Marseul, 1855) from Bulgaria, 8th sternite and tergite, 17 ventral view 18 ditto, dorsal view 19 ditto, lateral view 20 9th + 10th tergites, dorsal view 21 ditto, lateral view 22 aedeagus, dorsal view 23 ditto, lateral view 24 spiculum gastrale, ventral view 25 ditto, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="CAD4B5738BB32629AACA1B174F777D53" pageId="10" pageNumber="75" type="differential diagnosis">
|
||
<paragraph id="7EFF6AE8AEED6986A03A5C27D9B0B524" pageId="10" pageNumber="75">Differential diagnosis.</paragraph>
|
||
<paragraph id="78305D07153868029EBDD18BC25D3CA7" pageId="10" pageNumber="75">
|
||
<taxonomicName id="1045E8FEFB413E2D12762959297409B6" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
|
||
differs from the three species
|
||
<taxonomicName id="94D01492296815FFF7A37BF37034F470" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus henoni" order="Coleoptera" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="henoni">Exaesiopus henoni</taxonomicName>
|
||
,
|
||
<taxonomicName id="4D7F12F658377C59884993253678C81D" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus therondi" order="Coleoptera" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="therondi">Exaesiopus therondi</taxonomicName>
|
||
and
|
||
<taxonomicName id="EE199EA8E3981AE5982B0C786380FB34" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus laevis" order="Coleoptera" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="laevis">Exaesiopus laevis</taxonomicName>
|
||
chiefly by the shape of its protibia, which is on its outer margin furnished with three low teeth topped by triangular or rounded denticles (Figs 9, 10), whereas the three other mentioned species have their protibia furnished with two large teeth topped by triangular denticles on outer margin (Figs 61, 79 & 114). From
|
||
<taxonomicName id="B35F9BDC2D58B41B9CA0CBF29B890300" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus atrovirens" order="Coleoptera" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="atrovirens">Exaesiopus atrovirens</taxonomicName>
|
||
and
|
||
<taxonomicName id="8EDD3020F00AD6102871757D15930613" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus glaucus" order="Coleoptera" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="glaucus">Exaesiopus glaucus</taxonomicName>
|
||
it differs chiefly by the absence of a green metallic hue of the dorsum (compare Figs 1 with 73 & 98); from
|
||
<taxonomicName id="1B4BE6BBCFB8313A2EC95583158A3D23" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus glaucus" order="Coleoptera" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="glaucus">Exaesiopus glaucus</taxonomicName>
|
||
it differs furthermore by thickened and dilated metatibia (compare Figs 13 with 106). On the other hand, some specimens of
|
||
<taxonomicName id="008860BF3F690228944017BA01B35163" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
|
||
(especially from N. Africa that formerly belonged to the subspecies
|
||
<taxonomicName id="AE244FCB127F3D30D4F0EB2E4A7B474D" lsidName="berberus" pageId="10" pageNumber="75" rank="subspecies" species="berberus" subspecies="berberus">berberus</taxonomicName>
|
||
) can resemble the specimens of Middle-Asian
|
||
<taxonomicName id="20E0EA6FC198D1E9BE0B276A546634AB" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus torvus" order="Coleoptera" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="torvus">Exaesiopus torvus</taxonomicName>
|
||
by their densely punctate dorsum. These specimens differ, however, from
|
||
<taxonomicName id="E3C9D993D636205F4F809BC9C3B2D50E" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus torvus" order="Coleoptera" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="torvus">Exaesiopus torvus</taxonomicName>
|
||
by their respective male genitalia (compare Figs 17-34 with 64-72) and the less punctate pronotal disc (see also Key to the species for details, below). Most specimens of
|
||
<taxonomicName id="EC5772220ABEDFDD43B4EAE15E2D2112" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
|
||
, however (especially those from the northern shore of the Mediterranean Sea and South Europe) have distinctly less punctate dorsum than the specimens of
|
||
<taxonomicName id="E549F489F6A43947DB4ED51C30AE586C" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus torvus" order="Coleoptera" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="torvus">Exaesiopus torvus</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="345E8CAB4F8D3522D9322C6792315A01" pageId="10" pageNumber="75">
|
||
<paragraph id="075E51C2A82432F0A7866A228CAEBCE9" pageId="10" pageNumber="75">
|
||
Figures 26-34.
|
||
<taxonomicName id="C74C3858D16855819C0211D7A06852C3" class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus grossipes" order="Coleoptera" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="grossipes">Exaesiopus grossipes</taxonomicName>
|
||
(Marseul, 1855) from Tunisia, 8th sternite and tergite, 26 ventral view 27 ditto, dorsal view 28 ditto, lateral view 29 9th + 10th tergites, dorsal view 30 ditto, lateral view 31 spiculum gastrale, ventral view 32 ditto, lateral view 33 aedeagus, dorsal view 34 ditto, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="8F9DA91FDAB3FC435DBB438F9961DF64" pageId="10" pageNumber="75" type="biology">
|
||
<paragraph id="10706630E3F6A6C92CFD8FAA491C8FF1" pageId="10" pageNumber="75">Biology.</paragraph>
|
||
<paragraph id="15A7C2F18B640C2419BF70777FC5000D" pageId="10" pageNumber="75">This species is found on the beach under coastal wrack as well as further away from the waterfront, almost exclusively on sandy soil. Beetles can be found under rotting fish, excrements or buried under vegetation.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="21E8AAC32113564FD8DA88D8335852FB" pageId="10" pageNumber="75" type="distribution">
|
||
<paragraph id="3035C9F385321B37C82EE78DB8AB517D" pageId="10" pageNumber="75">Distribution.</paragraph>
|
||
<paragraph id="1013C3B380287C3BED060B02676429C1" pageId="10" pageNumber="75">Known from the Canary Islands, Morocco, Algeria, Tunisia, Libya, Spain, France, Italy, Greece, Bosnia and Herzegovina, Macedonia, Bulgaria, Russia, Serbia, Slovenia, Ukraine, Slovakia, Hungary, Austria, Iraq.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="536132FB939C69C486A4B671615A41E1" pageId="10" pageNumber="75" type="remarks">
|
||
<paragraph id="A2E4D335697C37ECAA0E955D4278E9F5" pageId="10" pageNumber="75">Remarks.</paragraph>
|
||
<paragraph id="8E8A6DB945EDF1409C02EA57BE05FABA" pageId="10" pageNumber="75">A variable species, covering vast area from the Canary Islands in the west to Iraq in the east. Its external morphology as well as male genitalia exhibit a certain degree of variation (compare Figs 17-25 and 26-34), but I find it difficult to discern discrete states among the variation and prefer to lump all examined specimens under the same species.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |