813 lines
69 KiB
XML
813 lines
69 KiB
XML
<document id="E9E2D073073B4762ACD691C5EC2A1A3F" ID-DOI="10.3897/zookeys.1220.130885" ID-ZooBank="7A7FE0E0-7BAA-4DB2-8061-A51BE183924F" ID-publisher-id="130885" URI-arpha="21435FA6-2D5B-5E72-AE9D-E32EF9111E8C" URI-zoobank="http://zoobank.org/7A7FE0E0-7BAA-4DB2-8061-A51BE183924F" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1733757385579" checkinUser="pensoft" docAuthor="Hibino, Yusuke & Ho, Hsuan-Ching" docDate="2024" docId="50B54A9DF01856E5956A358C9A0EC9E2" docLanguage="en" docName="ZooKeys 1220: 45-62" docOrigin="ZooKeys 1220" docSource="https://zookeys.pensoft.net/article/130885/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Yirrkala nkust Hibino & Ho, 2024, sp. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="62" masterDocId="21435FA62D5B5E72AE9DE32EF9111E8C" masterDocTitle="Species of the snake eel genus Yirrkala Whitley, 1940 from Taiwan, with descriptions of a new species and two new records (Anguilliformes, Ophichthidae)" masterLastPageNumber="62" masterPageNumber="45" pageNumber="45" updateTime="1733772390322" updateUser="admin">
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<mods:title id="1ED2082B78CE4A3643CBF92E59FDA651">Species of the snake eel genus Yirrkala Whitley, 1940 from Taiwan, with descriptions of a new species and two new records (Anguilliformes, Ophichthidae)</mods:title>
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<mods:name id="7C63B68950915D6EC47C56552E31036E" type="personal">
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<mods:roleTerm id="E9DC15FD0590DF9A67380C73478AC5EF">Author</mods:roleTerm>
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<mods:namePart id="81ACD16CF23B251952F7B6AFCAAD6B34">Hibino, Yusuke</mods:namePart>
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<mods:nameIdentifier id="B101D1F695C4A821BE08ACA2D2C9E9E4" type="ORCID">0000-0002-5670-3851</mods:nameIdentifier>
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<mods:affiliation id="40D4600619ECB32F452791A355AD6B5C">Kitakyushu Museum of Natural History and Human History, Fukuoka 805 - 0071, Japan</mods:affiliation>
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<mods:roleTerm id="823501F191D3749BAD6F5390DBA4C533">Author</mods:roleTerm>
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<mods:namePart id="310216EB590A41A7587E81EB0BDDA699">Ho, Hsuan-Ching</mods:namePart>
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<mods:nameIdentifier id="1B9B3CA9F21F1F8B3AC1C73838DB24BD" type="ORCID">0000-0003-1154-601X</mods:nameIdentifier>
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<mods:affiliation id="3A3AE0356A39A4322DBA983F5D04EDF8">Department and Graduate Institute of Aquaculture, National Kaohsiung University of Science and Technology, Kaohsiung 81157, Taiwan & Australian Museum, Sydney 2010, Australia</mods:affiliation>
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<mods:title id="308FB4BBF85DEA1A33617ED62A853B0B">ZooKeys</mods:title>
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<mods:part id="2E7AAD31F4E385F42FB3C67D1BDA2146">
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<mods:date id="C9258D6440DCC357C6269A963AA5D8E3">2024</mods:date>
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<mods:detail id="F9CA4AB52C01C5B4A99AC3CE486FEFEC" type="pubDate">
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<mods:number id="1E9BC11D17EA9DA422D24C21F8E1B9A4">2024-12-09</mods:number>
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<mods:number id="F652C376A238C0886BF48E854554A18B">1220</mods:number>
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<mods:identifier id="B4E8B4580A3AB18DD13B1AC71D9C716C" type="DOI">10.3897/zookeys.1220.130885</mods:identifier>
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<mods:identifier id="E1CD7667C30AA98B932918E87C66F914" type="ZooBank">7A7FE0E0-7BAA-4DB2-8061-A51BE183924F</mods:identifier>
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<treatment id="50B54A9DF01856E5956A358C9A0EC9E2" ID-arpha="50B54A9D-F018-56E5-956A-358C9A0EC9E2" ID-zoobank="https://zoobank.org/8EAF8FC7-BEC3-41B8-9F98-3EBEADD0DBF1" LSID="urn:lsid:plazi:treatment:50B54A9DF01856E5956A358C9A0EC9E2" LSID-ZBK="urn:lsid:zoobank.org:act:8EAF8FC7-BEC3-41B8-9F98-3EBEADD0DBF1" httpUri="http://treatment.plazi.org/id/50B54A9DF01856E5956A358C9A0EC9E2" scope_family="Ophichthidae" scope_order="Anguilliformes">
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<subSubSection id="C2885E3E6F6789D9D2EDDA5C47F808D1" type="nomenclature">
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<paragraph id="C4EF1013048AF07F87A6D5CBE073D942">
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<taxonomicName id="99579625A7A0701F16168E5452458230" authorityName="Hibino & Ho" authorityYear="2024" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="nkust" status="sp. nov.">
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<emphasis id="1293845F0401B640B333FF44B5DF5812" italics="true">Yirrkala nkust</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1617100E4567E03D6AD4A07F104CDF38" rank="species">sp. nov.</taxonomicNameLabel>
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||
</paragraph>
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</subSubSection>
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||
<subSubSection id="E0115BBC0003753AA2A52F999C5A7290" type="description">
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<paragraph id="A6DAB7AE9B1206892456E4190A1585CB">
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<figureCitation id="58DA717D40E55277A30D85B28BC78C46" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Fresh condition of Yirrkala nkust sp. nov., holotype, NMMB-P 38652, 496 mm TL A whole body B lateral view of head C dorsal view of head. AG anterior edge of gill opening AN anus DO dorsal-fin origin." figureDoi="10.3897/zookeys.1220.130885.figure1" httpUri="https://binary.pensoft.net/fig/1194273">Figs 1</figureCitation>
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,
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<figureCitation id="FF8B2009F789CE8294514904E9B38D8A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Fresh condition (after freezing) of Yirrkala nkust sp. nov., paratype, KMNH VR 100650, 297 mm TL, photographed by Y. - C. Hsu A whole body B lateral view of head C dorsal view of head D ventral view of head. AG anterior edge of gill opening AN anus DO dorsal-fin origin." figureDoi="10.3897/zookeys.1220.130885.figure2" httpUri="https://binary.pensoft.net/fig/1194274">2</figureCitation>
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,
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<figureCitation id="656B0C56C980A13F20E51DE6E3FA379F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawing of lateral head A Yirrkala nkust sp. nov., holotype, NMMB-P 38652, 496 mm TL B Yirrkala kaupii, NMMB-P 38424, 328 mm TL C Yirrkala omanensis, KMNH VR 100616, 216 mm TL. Arrows indicate ethmoid = 1 st supraorbital (lefts), interorbital (middles) and mid-temporal (rights) pores." figureDoi="10.3897/zookeys.1220.130885.figure3" httpUri="https://binary.pensoft.net/fig/1194275">3 A</figureCitation>
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,
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<tableCitation id="64647ACA27E3181AF9192E7370C9A12D" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Counts and measurements of three Yirrkala species newly collected from Taiwan, with type information." httpUri="http://table.plazi.org/id/BB17082F8F822513CD451454E60E62F5" tableUuid="BB17082F8F822513CD451454E60E62F5">Table 1</tableCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="SECID0ELOAC" type="material">
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||
<paragraph id="45F4E68247A99DB491FF009B6639C355">
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||
<heading id="A93DF7F8372D3B7CCEE57B024F47AD30" reason="title">Materials examined.</heading>
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||
</paragraph>
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<paragraph id="43876C99231919FA242E8599B8BD6280">
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<materialsCitation id="51DE90F21C6484A0D1430C7572608511" collectingDate="2021-07-12" collectionCode="NMMB-P" country="Taiwan" county="Taiwan Strait" elevation="40" latitude="23.666666" location="Chi-kan" longLatPrecision="929" longitude="119.61" municipality="Bai-sha" specimenCode="NMMB-P 38652" specimenCount="1" stateProvince="Penghu" typeStatus="holotype">
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||
<emphasis id="C5855F00C530117F98DD36CFDA8BF1AF" bold="true" italics="true">
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<typeStatus id="6F78B78FDF7E4A1A6FA00A95F82A6E50" type="holotype">Holotype</typeStatus>
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</emphasis>
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•
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<specimenCode id="5E2D9AB3239DDC410A640F9A7A8DEBBE">NMMB-P 38652</specimenCode>
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, 496 mm
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<abbrev id="ABBRID0EWOAC" xlink_title="Total length">TL</abbrev>
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, ca
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0E4OAC" specific-use="{"type":"Point","coordinates":[119.610000,23.666667]}">
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<geoCoordinate id="8D0A315480179A7C6E7FD3DFA79F3317" degrees="23" direction="north" minutes="40" orientation="latitude" precision="925" value="23.666666">23°40'N</geoCoordinate>
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,
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<geoCoordinate id="938B508862576C354B9EFD99647A7F50" degrees="119" direction="east" minutes="36.6" orientation="longitude" precision="92" value="119.61">119°36.6'E</geoCoordinate>
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</named-content>
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||
,
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<location id="BC9605814538EF02C4CBF72A132D051E" LSID="urn:lsid:plazi:treatment:50B54A9DF01856E5956A358C9A0EC9E2:BC9605814538EF02C4CBF72A132D051E" country="Taiwan" county="Taiwan Strait" latitude="23.666666" longLatPrecision="929" longitude="119.61" municipality="Bai-sha" name="Chi-kan" stateProvince="Penghu">Chi-kan</location>
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,
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<collectingMunicipality id="D2DB73DB6772C0397671D773EE5E757E">Bai-sha</collectingMunicipality>
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, northern
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<collectingRegion id="26A1E69B104A3A56AF0E27957105294D" country="Taiwan" name="Penghu">Penghu</collectingRegion>
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, western
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<collectingCountry id="BF3E8FB99521281D5AA1DE95FCB6F24B" name="Taiwan">Taiwan</collectingCountry>
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,
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<collectingCounty id="284CD16238B72481779BEC6ECACB8AB9">Taiwan Strait</collectingCounty>
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, ca
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<elevation id="1DB8FFB072E39E841639E4B9DD6FEBC1" metricMagnitude="1" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" unit="m" value="40.0" valueMax="50.0" valueMin="30.0">
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||
<quantity id="44B441EDE7A90A660ACF9F538349E5AD" metricMagnitude="1" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" unit="m" value="40.0" valueMax="50.0" valueMin="30.0">30–50 m</quantity>
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depth
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</elevation>
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,
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<date id="B75F7B62FF5C0179B193525051ADE894" value="2021-07-12">
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<collectingDate id="69EE9B90ED3BB6D8F3247C2B8382C8DC" value="2021-07-12">12 July 2021</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
.
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<materialsCitation id="8FF18446C5C48E4E76265486AAE6C048" collectingDate="2021-07-12" collectingDateMax="2024-07-23" collectingDateMin="2021-07-12" collectionCode="NMMB-P, VR" country="Taiwan" county="Taiwan Strait" elevation="40" latitude="23.666666" location="Chi-kan" longLatPrecision="929" longitude="119.61" municipality="Bai-sha" specimenCode="NMMB-P 38645, NMMB-P 39317, KMNH VR 100650" specimenCount="1" stateProvince="Penghu" typeStatus="paratype">
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||
<emphasis id="6B3EB19E8E545DD9920BF82EE8CAB94B" bold="true" italics="true">
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||
<typeStatus id="6985DBC0633649EB3E93E3FEA19EEC8C" type="paratype">Paratypes</typeStatus>
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</emphasis>
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•
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<specimenCode id="65BDA866E9F80E03512CC0571CDB6DD9">NMMB-P 38645</specimenCode>
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, 315 mm
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<abbrev id="ABBRID0EFPAC" xlink_title="Total length">TL</abbrev>
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,
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||
<date id="746267EEC03600A447B0549B3C3362C3" value="2021-07-12">
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<collectingDate id="9A614B82191CBCCD56D919C8FCC5F029" value="2021-07-12">12 July 2021</collectingDate>
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</date>
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;
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<specimenCode id="43ED142A42FE7367312EC2A8C40D8ADA">NMMB-P 39317</specimenCode>
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, 462 mm
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<abbrev id="ABBRID0EJPAC" xlink_title="Total length">TL</abbrev>
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,
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<date id="D92CA7F8FA5B67612F3771E035F80157" value="2022-08-16">
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<collectingDate id="1740A0B79151A728C34D84BFEEC5BBC1" value="2022-08-16">16 August 2022</collectingDate>
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</date>
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;
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<specimenCode id="48A924621B6417F95B633518CAAF388D">
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<named-content id="03B0053D167C2DE9392750823E34280C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kitakyushu-museum-and-institute-natural-history" xlink_title="Kitakyushu Museum of Natural History and Human History">KMNH VR</named-content>
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100650
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</specimenCode>
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, 297 mm
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<abbrev id="ABBRID0ERPAC" xlink_title="Total length">TL</abbrev>
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,
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<date id="D6C02FD8E266E45F01AE0DA02705949E" value="2024-07-23">
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<collectingDate id="70FE501AF2B6CB69B76EB53A658E7784" value="2024-07-23">23 July 2024</collectingDate>
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</date>
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; all collected from near the type locality
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</materialsCitation>
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.
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</paragraph>
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<caption id="BB17082F8F822513CD451454E60E62F5" ID-Table-UUID="BB17082F8F822513CD451454E60E62F5" httpUri="http://table.plazi.org/id/BB17082F8F822513CD451454E60E62F5" startId="T1" targetIsTable="true">
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<paragraph id="904752EE52F89E5210A612FFA71B99E6">
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<label id="182F186FC2BF1F0FCE641BE2A1FCEC5D">Table 1.</label>
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</paragraph>
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<paragraph id="E34372D0F8590569A5F3D29D8F0EE015">
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Counts and measurements of three
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<taxonomicName id="DE2F36DDE73BAEDDC5026C6ACBC3F1BE">
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<emphasis id="2FF00E775B32DD03860D2F727A6CB786" italics="true">Yirrkala</emphasis>
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species
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</taxonomicName>
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newly collected from Taiwan, with type information.
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</paragraph>
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</caption>
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<paragraph id="95F32F439CD9ABBE7BFFDDB9B92184B6">
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||
<table id="TID0EHRAG" rules="all">
|
||
<tbody id="F64D47712A5C0A2D94FBBA09D8C858B3">
|
||
<tr id="06E3474823FCCF16BFA353979D3F94D5">
|
||
<th id="470CA6BA52D164FC68FCAF04AA4AB40A" colspan="1" isEmpty="true" rowspan="2">-</th>
|
||
<th id="C2508C9B2ECD0EA3CEDA737D26FFE893" colspan="2" rowspan="1">
|
||
<taxonomicName id="AFB89B11A0D53988E8A0B5C361699C5F" authorityName="Hibino & Ho" authorityYear="2024" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="nkust" status="sp. nov.">
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<emphasis id="BD1715699967C93F29CCA0FB4A4F1813" italics="true">Y. nkust</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="9459FB9BC2DE7F9B513C922DB59F2517" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
<emphasis id="0BEB43F50E68F7D53AED1BEF1B89E42E" italics="true">
|
||
<th id="C55924EBFFF235648381F9A03367B545" colspan="2" rowspan="1">
|
||
<taxonomicName id="A39E412148B2BC73113983816176BC73" baseAuthorityName="Bleeker" baseAuthorityYear="1858" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="kaupii">Y. kaupii</taxonomicName>
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</th>
|
||
<th id="2D0A9995575B91EBF896EDCD79C15DB8" colspan="2" rowspan="1">
|
||
<taxonomicName id="5A5AB0BD085BAEC10B564EFC75A8BEB7" baseAuthorityName="Norman" baseAuthorityYear="1939" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="omanensis">Y. omanensis</taxonomicName>
|
||
</th>
|
||
</emphasis>
|
||
</tr>
|
||
<tr id="F4E29841B7DD94ABEAD7AE1E711E0ECB">
|
||
<th id="263C8C90C8846BC97BBD6EB1DA91B22E" colspan="1" rowspan="1">Holotype</th>
|
||
<th id="98966165072AA04C80DC2097F23F6391" colspan="1" rowspan="1">Paratypes</th>
|
||
<th id="0109E158950A135982FFF83DB9337793" colspan="1" rowspan="1">Present materials</th>
|
||
<th id="CD3CC34A930CF93CF0A8F717D00379AA" colspan="1" rowspan="1">Holotype</th>
|
||
<th id="1B196B0A2FFBBE7ABAD3DAFF42FE9786" colspan="1" rowspan="1">Present material</th>
|
||
<th id="BE758F27B5F4998BE3DC1DBFD144A896" colspan="1" rowspan="1">Holotype</th>
|
||
</tr>
|
||
<tr id="3CDCAF7AD2C0D1E16AF37BB0E415B12C">
|
||
<td id="1A1BA76E7F5F41EDF3C9BD7BAB9D6B5E" colspan="1" rowspan="1">Total length (mm)</td>
|
||
<td id="FBC75F735993A5BD943F0C6F5C5365C4" colspan="1" rowspan="1">496</td>
|
||
<td id="B2D6FCBF566D7AB281BFF47D0998EC55" colspan="1" rowspan="1">
|
||
297–462 (
|
||
<emphasis id="DFAA89F7F7E33850E447D7086C73BB29" italics="true">n</emphasis>
|
||
= 3)
|
||
</td>
|
||
<td id="CED6145273090489EF5504C64219B08C" colspan="1" rowspan="1">
|
||
238–331 (
|
||
<emphasis id="3411B8C43F48326810B8DB35F064AF8A" italics="true">n</emphasis>
|
||
= 3)
|
||
</td>
|
||
<td id="12AC95AF4EA450487DB8AD19EDBF8C0F" colspan="1" rowspan="1">342</td>
|
||
<td id="2F7DED9A4EA9893EB35D285F65FF2A06" colspan="1" rowspan="1">216</td>
|
||
<td id="1336CD5A5B21BB7EDC469F5C7633A3D8" colspan="1" rowspan="1">
|
||
230
|
||
<superScript id="F1D975E54AC351A14288482D673E319E">b</superScript>
|
||
</td>
|
||
</tr>
|
||
<tr id="F47B8F3E3483973405C892FD5B0A2012">
|
||
<td id="F63CD9F156B2CE464B1EF83876DA64D6" colspan="7" rowspan="1">
|
||
As % of
|
||
<abbrev id="ABBRID0ERDAE" xlink_title="Total length">TL</abbrev>
|
||
</td>
|
||
</tr>
|
||
<tr id="3B11C8F31856E1F9F359A9F2D42EEDC1">
|
||
<td id="A97B410079DF0C18B0CB4B6949CAC4E5" colspan="1" rowspan="1">Head length</td>
|
||
<td id="BA0AB2CD07A8DC616283537DAACA58A5" colspan="1" rowspan="1">5.9</td>
|
||
<td id="43AEBDB95BD6B95B159A7F03C857434D" colspan="1" rowspan="1">6.2–6.4</td>
|
||
<td id="1EFA2403FE36FD68283654551E48529E" colspan="1" rowspan="1">7.3–8.4</td>
|
||
<td id="DF02348FDA793702F6AEFE9BB25E4F11" colspan="1" rowspan="1">
|
||
6.5
|
||
<superScript id="33C7D64E846CF9234A586E282B160621">a</superScript>
|
||
</td>
|
||
<td id="6E39B3A3CD6791D7B51250D38BD8D916" colspan="1" rowspan="1">8.1</td>
|
||
<td id="07E75ED7F555DFF12DE1159417C5BBDD" colspan="1" rowspan="1">
|
||
7.7
|
||
<superScript id="01CFA0E46163DDB6183E9240AFE1C21A">b</superScript>
|
||
</td>
|
||
</tr>
|
||
<tr id="74348D3CBE3A2A6AA3E300845350DC33">
|
||
<td id="C3805B3EB786B9940F2CE7E5D00EA66A" colspan="1" rowspan="1">Preanal length</td>
|
||
<td id="C19004B8334A9973A9F4EBCA38ABB545" colspan="1" rowspan="1">51.9</td>
|
||
<td id="CB2D1554A8158E0F82C240F7CF316892" colspan="1" rowspan="1">47.4–52.6</td>
|
||
<td id="C2B42C95DD5F0741588B44B65449255E" colspan="1" rowspan="1">47.1–47.3</td>
|
||
<td id="66CE57FD025A2BC46ABBBFA7D0DA56C4" colspan="1" rowspan="1">N / A</td>
|
||
<td id="47911588096C86ABE3228AF6C39ABBDA" colspan="1" rowspan="1">57.9</td>
|
||
<td id="04A96FF1DD35AE73710BEA653A805EC7" colspan="1" rowspan="1">
|
||
58.3
|
||
<superScript id="5354EA8E34EA2C451EDA5D2806AC10D2">c</superScript>
|
||
</td>
|
||
</tr>
|
||
<tr id="41B66AF4F708D0A7B790D6BEDF834231">
|
||
<td id="844555AE2681776F5F5319AA3E11F0F4" colspan="1" rowspan="1">Tail length</td>
|
||
<td id="B35251686BE0626C43A59046343D9E13" colspan="1" rowspan="1">48.1</td>
|
||
<td id="1D5386ECAD812A5EA549893B259559A6" colspan="1" rowspan="1">47.4–52.6</td>
|
||
<td id="B021E5382FAE3B4C41BA0415C433AEE4" colspan="1" rowspan="1">52.7–52.9</td>
|
||
<td id="A17AF494C96E8EC66B293B25E1EA8CD9" colspan="1" rowspan="1">N / A</td>
|
||
<td id="F05CC14558B304E65DC55252CDFC26CB" colspan="1" rowspan="1">42.1</td>
|
||
<td id="08F8F81E1D3E84B665E0CD30F61629B3" colspan="1" rowspan="1">
|
||
41.7
|
||
<superScript id="DB530225F4E5B6BDE713D6CA48BCD368">c</superScript>
|
||
</td>
|
||
</tr>
|
||
<tr id="EA06AF2F4E6C5BF4E68D073AAC3B4BB7">
|
||
<td id="08594F6153A4879E2EE6B280963DE8E2" colspan="1" rowspan="1">Predorsal length</td>
|
||
<td id="62E1A6DA955DBB84496A31430AC546D6" colspan="1" rowspan="1">5.1</td>
|
||
<td id="E5E3C70F8930C8FF63F6B27A83788C25" colspan="1" rowspan="1">5.5–6.1</td>
|
||
<td id="3750A6BB7FBE20BBE7DCD5614CC86067" colspan="1" rowspan="1">12.5–13.5</td>
|
||
<td id="E4DB06A39AFD8DD78449696AA908D16F" colspan="1" rowspan="1">N / A</td>
|
||
<td id="9E806613E542FC1BFFCB419F0D1FD8C9" colspan="1" rowspan="1">59.8</td>
|
||
<td id="295DFA497D672E68DE4B9EB56AB1CD82" colspan="1" rowspan="1">
|
||
ca 59
|
||
<superScript id="32B129D9B4835A5913CEBBA3F1FE8C75">d</superScript>
|
||
</td>
|
||
</tr>
|
||
<tr id="6B224B6643AC146C9F96F0FF55D0530D">
|
||
<td id="5D9E2D389CB1B6A20272FFDB4F102CB3" colspan="1" rowspan="1">Body depth at gill opening</td>
|
||
<td id="2A767A7104F7986D363DAD13FC7CC6EA" colspan="1" rowspan="1">1.7</td>
|
||
<td id="7B876903E378D2861D22906D87CB5271" colspan="1" rowspan="1">1.6–1.8</td>
|
||
<td id="06D6B0BE8D63C838B7195A18E7D2A23F" colspan="1" rowspan="1">2.1–2.4</td>
|
||
<td id="4EC1B541501B4823EC449903287D2B5B" colspan="1" rowspan="1">
|
||
(1.7)
|
||
<superScript id="087E616181195D4838B4D3F04100024E">a</superScript>
|
||
</td>
|
||
<td id="DC4E5B513F087E9BE95D60816323B2C5" colspan="1" rowspan="1">2.3</td>
|
||
<td id="1790A083BCC38400A918EB8E99A58C06" colspan="1" rowspan="1">
|
||
(1.8)
|
||
<superScript id="735082CC5E5066AF893FC9C06FD0E7A6">c</superScript>
|
||
</td>
|
||
</tr>
|
||
<tr id="4A3F85E5272B8AD9D0249211E375A1AF">
|
||
<td id="D6DAECC0DFBE8552A1E91439ABF4A339" colspan="1" rowspan="1">Body width at gill opening</td>
|
||
<td id="9779928989FF7A770103DBDD1BC202DA" colspan="1" rowspan="1">1.5</td>
|
||
<td id="6AC75D8ACBF7FDF398B2182A586DDA07" colspan="1" rowspan="1">1.5–1.6</td>
|
||
<td id="8C811E1143D711CEAC6212F37C2F2416" colspan="1" rowspan="1">1.4–1.7</td>
|
||
<td id="F0EED50A423C3D3241ACB90D9B93EB50" colspan="1" rowspan="1">N / A</td>
|
||
<td id="8959A883FEA402BC044555B55BE7983E" colspan="1" rowspan="1">2.1</td>
|
||
<td id="3666992A5DEA802F45F5FCDBF761DCFA" colspan="1" rowspan="1">N / A</td>
|
||
</tr>
|
||
<tr id="0D46F6B03E4CA5FC396C186B1FED1E5A">
|
||
<td id="43B4798D3A361CCC8294A6EEC639A4B5" colspan="1" rowspan="1">Body depth at midanus</td>
|
||
<td id="DCB770F0483C6E75593F679C7F10A2DF" colspan="1" rowspan="1">1.7</td>
|
||
<td id="368F8756117D80266C2CDB4CD27E1050" colspan="1" rowspan="1">1.3–1.5</td>
|
||
<td id="A1EAA89A274666627A3DBFF5BC8BC0AB" colspan="1" rowspan="1">2.4–2.5</td>
|
||
<td id="E0403D129CC841ADFB1781A986F36F9A" colspan="1" rowspan="1">N / A</td>
|
||
<td id="E742EF35199F85E042A26A60434D2296" colspan="1" rowspan="1">2.1</td>
|
||
<td id="26E44B62024874DCE796E63C61CBDC87" colspan="1" rowspan="1">N / A</td>
|
||
</tr>
|
||
<tr id="FCDD762060BA81423891F2E92D20BD55">
|
||
<td id="07F0DB81662790500AFF2F2B71718F61" colspan="1" rowspan="1">Body width at midanus</td>
|
||
<td id="0717D14634E4F908170548DE8C874DBB" colspan="1" rowspan="1">1.6</td>
|
||
<td id="C88106A1A824DD2C122966A45992F2AA" colspan="1" rowspan="1">1.4–1.6</td>
|
||
<td id="402F2E59066E2D85C50EF623A2679446" colspan="1" rowspan="1">1.5–2.1</td>
|
||
<td id="41E73B47C417379C8CCC46FFDD629542" colspan="1" rowspan="1">N / A</td>
|
||
<td id="11388A8609564A0E75ED69BCF08B88DF" colspan="1" rowspan="1">2.0</td>
|
||
<td id="E114FEF8592683ED9C4DF69046F8C79E" colspan="1" rowspan="1">N / A</td>
|
||
</tr>
|
||
<tr id="15D63C155AD85CD0383E31423E358BFB">
|
||
<td id="8875D6B3EE9A12E32FA0F1157F8EDA13" colspan="7" rowspan="1">As % of head length</td>
|
||
</tr>
|
||
<tr id="F70962A39C23BA932A465D7B5D069357">
|
||
<td id="5EC87C5FDA8DADF8475A68F3A1DD8C98" colspan="1" rowspan="1">Snout length</td>
|
||
<td id="EA51FCB033C478C0CFBC98D743E4516A" colspan="1" rowspan="1">15.1</td>
|
||
<td id="0DD21626C28C58F876A5E3BC391957BF" colspan="1" rowspan="1">14.7–16.5</td>
|
||
<td id="033563C96C546AF55356BE35DCAE3959" colspan="1" rowspan="1">13.9–14.5</td>
|
||
<td id="C217A6434C0434ED6220B0F4258DEFAD" colspan="1" rowspan="1">
|
||
ca 16
|
||
<superScript id="F68E6D86D85FBF739FB4FB6C862FCC7D">a</superScript>
|
||
</td>
|
||
<td id="218E86A902FC1E126F1223CE365EAF0E" colspan="1" rowspan="1">11.4</td>
|
||
<td id="F85651C0392E93EA83A5FDD8725A2363" colspan="1" rowspan="1">
|
||
ca 11
|
||
<superScript id="352D848677DAC4BBB77695D2206E43F3">b</superScript>
|
||
</td>
|
||
</tr>
|
||
<tr id="6CC95F8C069EF538E7537D78F59FD9E5">
|
||
<td id="434B409CBA711057504EB8C382DDA51F" colspan="1" rowspan="1">Eye diameter</td>
|
||
<td id="71C11E6EDEA0CA8EF0432B0ED17016AD" colspan="1" rowspan="1">4.5</td>
|
||
<td id="68C94928DF9D5C3035B9D22937B6C483" colspan="1" rowspan="1">3.6–4.5</td>
|
||
<td id="4169593D20D75081B85605A2EDD1475F" colspan="1" rowspan="1">6.1–6.5</td>
|
||
<td id="101E83CBC1F17B3FE443F4478FB7AC3A" colspan="1" rowspan="1">
|
||
ca 8
|
||
<superScript id="D582496FA6E11FEDD610DBA38CD7A5CB">a</superScript>
|
||
</td>
|
||
<td id="F723100BD8089CB9119EC839BBFAC094" colspan="1" rowspan="1">5.7</td>
|
||
<td id="08C6EBB71A59F53BF683A159DB14546B" colspan="1" rowspan="1">N / A</td>
|
||
</tr>
|
||
<tr id="47CF5973C4192651D09763EA81E06140">
|
||
<td id="21D9541F1E47A716324D0B036A8EB757" colspan="1" rowspan="1">Upper-jaw length</td>
|
||
<td id="A5E5AAB7ECC680B59201330B7BBD0E82" colspan="1" rowspan="1">28.0</td>
|
||
<td id="2224CD7DB97F5D7C326198BD03F85EA2" colspan="1" rowspan="1">28.0–30.9</td>
|
||
<td id="9723DB15A7D7A0E2EE1E6F898F508391" colspan="1" rowspan="1">22.4–23.4</td>
|
||
<td id="172338DCED203670E598353D76DD3328" colspan="1" rowspan="1">
|
||
ca 25
|
||
<superScript id="67119A55742D165B2BCA240EACEC5F86">a</superScript>
|
||
</td>
|
||
<td id="938431574B1F0A9E677F1200D45CDDA9" colspan="1" rowspan="1">28.6</td>
|
||
<td id="ACBA914166D4E60B58A3F85D69396C4A" colspan="1" rowspan="1">N / A</td>
|
||
</tr>
|
||
<tr id="9AEE9E4E163C87A977321A337486C5DC">
|
||
<td id="A7BA4FD181608F608E0E7B7C91F3B3D1" colspan="1" rowspan="1">Gill-opening length</td>
|
||
<td id="571425150CBFF5E187FAA79BFAA992C8" colspan="1" rowspan="1">12.7</td>
|
||
<td id="5DBC98F330AF7C8226B136D3E2040C60" colspan="1" rowspan="1">11.4–15.7</td>
|
||
<td id="2D337BAB348313799A745CB31E83BA52" colspan="1" rowspan="1">8.2–13.4</td>
|
||
<td id="FBF16A56B6B6271BE978DBE950B1E888" colspan="1" rowspan="1">N / A</td>
|
||
<td id="FC36E2F927CB7E5524E0D78C90A8BA18" colspan="1" rowspan="1">9.7</td>
|
||
<td id="02ABF5915548AB883345FC41431D0396" colspan="1" rowspan="1">N / A</td>
|
||
</tr>
|
||
<tr id="9CEF4510D6502D94AA66279EB462E420">
|
||
<td id="C38554BE66BE7638E8EFCF4EA82BCC53" colspan="1" rowspan="1">Interorbital width</td>
|
||
<td id="7A41AF076E884918C411FBA8E7D469EA" colspan="1" rowspan="1">7.9</td>
|
||
<td id="68233AE74F37BF447081FDE943E8A1D8" colspan="1" rowspan="1">6.6–9.8</td>
|
||
<td id="F4CFF262F060AFECE68A4A63DBE3C3C0" colspan="1" rowspan="1">6.5–8.2</td>
|
||
<td id="E1D36B02C07497F62873F94888C122F5" colspan="1" rowspan="1">N / A</td>
|
||
<td id="BDF0DEE3FB19EB7EA67E8F5C2F20E347" colspan="1" rowspan="1">5.1</td>
|
||
<td id="2D208704F9F4B8FFF02D6D4C6B1E0619" colspan="1" rowspan="1">N / A</td>
|
||
</tr>
|
||
<tr id="9490CD118405BF7626ACB58E5CF7474D">
|
||
<td id="FB3B7C2146460E73A655FD77F0066EA7" colspan="1" rowspan="1">Isthmus width</td>
|
||
<td id="2D9569A4E274553812A40A6BD16DC31C" colspan="1" rowspan="1">5.1</td>
|
||
<td id="04840051344C03D55296B3456BD7D277" colspan="1" rowspan="1">3.9–5.5</td>
|
||
<td id="CD0DED3DB379AD203ADB24FC7920D1D8" colspan="1" rowspan="1">8.5–15.6</td>
|
||
<td id="EF67C6A8C206DEEC5F55394ADF6A43F9" colspan="1" rowspan="1">N / A</td>
|
||
<td id="997B918C2E0D081C57909022BADA6759" colspan="1" rowspan="1">6.9</td>
|
||
<td id="F5DC9B869A7E92DE76B7BCEA2BCDC7A3" colspan="1" rowspan="1">N / A</td>
|
||
</tr>
|
||
<tr id="BBAC40A12DA828C51FF7BFC7B082C6B9">
|
||
<td id="2215F7C445286307ECD80D4E1D4954EF" colspan="7" rowspan="1">Counts</td>
|
||
</tr>
|
||
<tr id="04D32A2B15CCA3B2F97072F0F2443618">
|
||
<td id="2814618E0CE850F33203D0215719D4E8" colspan="1" rowspan="1">Predorsal vertebrae</td>
|
||
<td id="2FB6150D7E09ADA2108934D4B026346A" colspan="1" rowspan="1">8</td>
|
||
<td id="B094A722F0E27660FE33B251E822D677" colspan="1" rowspan="1">7–8</td>
|
||
<td id="7685D0FC3F351560EAB4BFFC15D5E976" colspan="1" rowspan="1">17–18</td>
|
||
<td id="E4D32B563CC9C8DEDFA9048A7EF664C0" colspan="1" rowspan="1">17</td>
|
||
<td id="A523B10D6FEC150C5F30CC8E5020BFC8" colspan="1" rowspan="1">78</td>
|
||
<td id="75957C3864CE3A11EAF480CC2DE00516" colspan="1" rowspan="1">76</td>
|
||
</tr>
|
||
<tr id="34F9E8F78AB29B1A110E6D4E41E8038C">
|
||
<td id="1AE23DD8D1A774D308CF8DD3CC1F5995" colspan="1" rowspan="1">Preanal vertebrae</td>
|
||
<td id="E7E9163AFDD32B0DB98FC5368710E830" colspan="1" rowspan="1">72</td>
|
||
<td id="2246FEAA4131E81A115FBCDBCCB80584" colspan="1" rowspan="1">72–75</td>
|
||
<td id="53FD61942E2505F6E9981EFF82BBD4AF" colspan="1" rowspan="1">64–66</td>
|
||
<td id="F5B0672241B90AD3FAD133504DDB6037" colspan="1" rowspan="1">65</td>
|
||
<td id="E8D7C4F639CD8C8B5C054885106ACA66" colspan="1" rowspan="1">76</td>
|
||
<td id="4BD2007760B6FF18B2DE0238CADC3417" colspan="1" rowspan="1">76</td>
|
||
</tr>
|
||
<tr id="0A039CFD1DA481AF4C5B79E04243B0D3">
|
||
<td id="E2EFF444397DE07752A765DFAD540BD3" colspan="1" rowspan="1">Total vertebrae</td>
|
||
<td id="929BA6541019F785BBDB602C65C9381B" colspan="1" rowspan="1">147</td>
|
||
<td id="7300A92730A311E5BA188AC79A9ED839" colspan="1" rowspan="1">150–152</td>
|
||
<td id="7BFA83309223FC2DA2388E1E95C83E2B" colspan="1" rowspan="1">151–155</td>
|
||
<td id="7FB84B454DA215A84F0F006ED599B300" colspan="1" rowspan="1">156</td>
|
||
<td id="4B44423A51F2918B89296AE715C5AD1B" colspan="1" rowspan="1">141</td>
|
||
<td id="42DA95B13550C0AD36F0AD83DCAEC2FF" colspan="1" rowspan="1">140</td>
|
||
</tr>
|
||
</tbody>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="A3332643B230ED8DCD4B11662F6FDE67">
|
||
<tableNote id="FE1C90A34B97D2E7E04792796634368C">
|
||
Body depths in parentheses are possibly the maximum depth.
|
||
<superScript id="7ED73DF0C7739D83915E7DA3EA23526C">a</superScript>
|
||
from
|
||
<bibRefCitation id="EF35000EF5B6C579FD1DB321D1206101" DOI="10.5962/bhl.title.144153" author="Bleeker P" firstAuthor="Bleeker" issue="8" journalOrPublisher="Acta Societatis Regiae Scientiarum Indo-Neêrlandicae" pagination="1-4" refId="B1" refString="Bleeker P (1858) Twaalfde bijdrage tot de kennis der vischfauna van Celebes. Visschen van Manado. Acta Societatis Regiae Scientiarum Indo-Neêrlandicae 5 (8): 1–4. https://doi.org/10.5962/bhl.title.144153" title="Twaalfde bijdrage tot de kennis der vischfauna van Celebes. Visschen van Manado." volume="5" year="1858">Bleeker (1858)</bibRefCitation>
|
||
;
|
||
<superScript id="B52123157E12C6FA7929E51DA70CA94C">b</superScript>
|
||
from
|
||
<bibRefCitation id="FA52669E2374F5FC35F802EA3E7814A0" author="Norman JR" firstAuthor="Norman" journalOrPublisher="Scientific Reports, John Murray Expedition" pagination="1-116" refId="B27" refString="Norman JR (1939) Fishes. The John Murray Expedition 1933–34. Scientific Reports, John Murray Expedition 7: 1–116." title="Fishes. The John Murray Expedition 1933-34." volume="7" year="1939">Norman (1939)</bibRefCitation>
|
||
;
|
||
<superScript id="CB0C68473A3E40E30AF90763DC5CF8F0">c</superScript>
|
||
from
|
||
<bibRefCitation id="E2BEC508C8430E8606B9DA54B1B9128E" author="McCosker JE" editor="Heemstra PC & Heemstra E & Ebert DA & Holleman W & Randall JE" firstAuthor="McCosker" journalOrPublisher="Vol. 2. South African Institute for Aquatic Biodiversity, Makhanda" pagination="101-137" refId="B21" refString="McCosker JE (2022) Family Ophichthidae, Snake-eels, sand-eels and worm-eels. In: Heemstra PC, Heemstra E, Ebert DA, Holleman W, Randall JE (Eds) Coastal Fishes of the Western Indian Ocean. Vol. 2. South African Institute for Aquatic Biodiversity, Makhanda, 101–137." title="Family Ophichthidae, Snake-eels, sand-eels and worm-eels." volumeTitle="Coastal Fishes of the Western Indian Ocean." year="2022">McCosker (2022)</bibRefCitation>
|
||
; and
|
||
<superScript id="5FDA4C32E1ACE4F32B9482735B09977B">d</superScript>
|
||
estimated from
|
||
<bibRefCitation id="F42068AA76B0E33B1924B96052AADA7B" author="Norman JR" firstAuthor="Norman" journalOrPublisher="Scientific Reports, John Murray Expedition" pagination="1-116" refId="B27" refString="Norman JR (1939) Fishes. The John Murray Expedition 1933–34. Scientific Reports, John Murray Expedition 7: 1–116." title="Fishes. The John Murray Expedition 1933-34." volume="7" year="1939">Norman (1939)</bibRefCitation>
|
||
.
|
||
</tableNote>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EERAE" type="diagnosis">
|
||
<paragraph id="25774A2F95593888B182345FF167EADE">
|
||
<heading id="A13174FD144464D7E23DE267569562E2" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="DAB1EFAFDD16F1B559F237BDF3CFD295">
|
||
An elongate species of
|
||
<taxonomicName id="25252DCA78448746900E0C760DD6F00B" authorityName="Whitley" authorityYear="1940" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="genus">
|
||
<emphasis id="62F18EC2033CC7ABB44B02976DDB1C99" italics="true">Yirrkala</emphasis>
|
||
</taxonomicName>
|
||
with the following combination of characters: body reddish brown without speckles, snout paler; supraorbital pores (
|
||
<abbrev id="ABBRID0ERRAE" xlink_title="supraorbital pores">SO</abbrev>
|
||
) 1 + 3; dorsal-fin origin just above gill opening; lateral-line pores before anus 72–78; predorsal vertebrae 7–8, preanal 72–75, and total 147–152;
|
||
<abbrev id="ABBRID0EVRAE" xlink_title="Mean vertebral formula">MVF</abbrev>
|
||
8-73 - 150.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EZRAE" type="description">
|
||
<paragraph id="113F1202C06E3A72325C487044F00DAB">
|
||
<heading id="BAAC8707FC8076F81BCBC2B204179AC5" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="E3284C92869B47EEC8ACA94933B491AB">
|
||
Body elongate, subcylindrical, tip of tail laterally compressed and extremely pointed (Figs
|
||
<figureCitation id="55AF5F09FA70386B8A5D7D6A0C9FE07E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Fresh condition of Yirrkala nkust sp. nov., holotype, NMMB-P 38652, 496 mm TL A whole body B lateral view of head C dorsal view of head. AG anterior edge of gill opening AN anus DO dorsal-fin origin." figureDoi="10.3897/zookeys.1220.130885.figure1" httpUri="https://binary.pensoft.net/fig/1194273">1 A</figureCitation>
|
||
,
|
||
<figureCitation id="3119379646FC5D3233D1DFE9607182ED" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Fresh condition (after freezing) of Yirrkala nkust sp. nov., paratype, KMNH VR 100650, 297 mm TL, photographed by Y. - C. Hsu A whole body B lateral view of head C dorsal view of head D ventral view of head. AG anterior edge of gill opening AN anus DO dorsal-fin origin." figureDoi="10.3897/zookeys.1220.130885.figure2" httpUri="https://binary.pensoft.net/fig/1194274">2 A</figureCitation>
|
||
). Head moderate in size, 15.7–17.0 in
|
||
<abbrev id="ABBRID0EHSAE" xlink_title="Total length">TL</abbrev>
|
||
(17.0 in
|
||
<typeStatus id="F12EC95C7B86E96BB1F7F20F771C60D1" type="holotype">holotype</typeStatus>
|
||
); preanal length mostly equal to tail,
|
||
<quantity id="F7672CC22B716841ED245F2856342D13" metricMagnitude="-2" metricUnit="m" metricValue="2.54" metricValueMax="2.794" metricValueMin="2.286" unit="in" value="1.0" valueMax="1.1" valueMin="0.9">0.9–1.1 in</quantity>
|
||
tail length and
|
||
<quantity id="A2B66709F1971A88AF578EC1F6B9BCB1" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="5.334" metricValueMin="4.826" unit="in" value="2.0" valueMax="2.1" valueMin="1.9">1.9–2.1 in</quantity>
|
||
<abbrev id="ABBRID0ELSAE" xlink_title="Total length">TL</abbrev>
|
||
(0.9 and
|
||
<quantity id="88C731BDDBF7E4B6016CC82BA030931D" metricMagnitude="-2" metricUnit="m" metricValue="4.826" unit="in" value="1.9">1.9 in</quantity>
|
||
<typeStatus id="E367D461DEED4CDEF276DB63D86AF14A" type="holotype">holotype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
<caption id="2E614ABE7D1D6FBA9EE8EFF3172B5D72" ID-DOI="10.3897/zookeys.1220.130885.figure1" ID-arpha="FB0F031B-FF5C-5621-85C4-D8D05554B1AA" httpUri="https://binary.pensoft.net/fig/1194273" startId="F1">
|
||
<paragraph id="DC63C3D6A687D42BD090B4B517AC9397">
|
||
<label id="5C3DCBDCC26A9947E104FAAF6E9DB850">Figure 1.</label>
|
||
</paragraph>
|
||
<paragraph id="97C9608889E7671E818E7831FF421B01">
|
||
Fresh condition of
|
||
<taxonomicName id="E4A59A165C8F1FBE46D94F20024C3708" authorityName="Hibino & Ho" authorityYear="2024" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="nkust" status="sp. nov.">
|
||
<emphasis id="EB50D01F23437735923A436C76D10B3F" italics="true">Yirrkala nkust</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="749B36E899F97A8F7AA760A9512DDCB4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype,
|
||
<specimenCode id="FC01703F781FB17149F19179CCBFD362">NMMB-P 38652</specimenCode>
|
||
, 496 mm
|
||
<abbrev id="ABBRID0ECTAE" xlink_title="Total length">TL</abbrev>
|
||
<emphasis id="142B1BD9F128550DCE1FBEA5D395A738" bold="true">A</emphasis>
|
||
whole body
|
||
<emphasis id="CE3BE4C132D0571973BA14B5BB75BBE5" bold="true">B</emphasis>
|
||
lateral view of head
|
||
<emphasis id="4086CC634CF9F0B34F8F883742C78D92" bold="true">C</emphasis>
|
||
dorsal view of head.
|
||
<emphasis id="B9CFE9D76D1078E77E2915347A37FFB1" bold="true">AG</emphasis>
|
||
anterior edge of gill opening
|
||
<emphasis id="61CBFA607B77FC4DC9096B6DC249CC43" bold="true">AN</emphasis>
|
||
anus
|
||
<emphasis id="1BFCE73840DDC75CD5003677CC70743E" bold="true">DO</emphasis>
|
||
dorsal-fin origin.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="E16681967BE8AFC23A76DE274BD2AF09" ID-DOI="10.3897/zookeys.1220.130885.figure2" ID-arpha="477FCBB7-F2AA-5D20-B1B4-BC5F53047A53" httpUri="https://binary.pensoft.net/fig/1194274" startId="F2">
|
||
<paragraph id="20B8B9F7C2D0AAA3701F6C5148EFC79D">
|
||
<label id="B9881002A1F19B0E6579146942208847">Figure 2.</label>
|
||
</paragraph>
|
||
<paragraph id="D8369713FB396A7063293A4C860D8827">
|
||
Fresh condition (after freezing) of
|
||
<taxonomicName id="23E685A6BEB002C6948896558F7DF420" authorityName="Hibino & Ho" authorityYear="2024" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="nkust" status="sp. nov.">
|
||
<emphasis id="44185A839F128F40CD98403031385197" italics="true">Yirrkala nkust</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DAD2E405A98E533AE63BC350E94A12E5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype,
|
||
<specimenCode id="EAF4D27EFB705F1C87C781467C529F59">
|
||
<named-content id="2AAC08C06585ABDFFC3FD2D7FAA809AB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kitakyushu-museum-and-institute-natural-history" xlink_title="Kitakyushu Museum of Natural History and Human History">KMNH VR</named-content>
|
||
100650
|
||
</specimenCode>
|
||
, 297 mm
|
||
<abbrev id="ABBRID0EOUAE" xlink_title="Total length">TL</abbrev>
|
||
, photographed by Y. - C. Hsu
|
||
<emphasis id="E389A22CE9BDF2CB12E5F4098AB7655D" bold="true">A</emphasis>
|
||
whole body
|
||
<emphasis id="23F0661CB02966609F569A50A0337163" bold="true">B</emphasis>
|
||
lateral view of head
|
||
<emphasis id="C154B3B9FEE1074B68B6EB52ECDBF62F" bold="true">C</emphasis>
|
||
dorsal view of head
|
||
<emphasis id="0E4245F8008D12593A62946501D73009" bold="true">D</emphasis>
|
||
ventral view of head.
|
||
<emphasis id="038E9580108F4CBF6C1D222ACF70C023" bold="true">AG</emphasis>
|
||
anterior edge of gill opening
|
||
<emphasis id="EE1ADF167C1903168E9B29124BC9504D" bold="true">AN</emphasis>
|
||
anus
|
||
<emphasis id="DE10913CFAD5979E6E518A856100CD4D" bold="true">DO</emphasis>
|
||
dorsal-fin origin.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="AE88517835DFB5A3984E316DA234C813">
|
||
Snout relatively longer, 3.4–3.7 times eye diameter (
|
||
<quantity id="C062BC681345839E941E5D3BACA2CF9A" metricMagnitude="-2" metricUnit="m" metricValue="9.398" unit="in" value="3.7">3.7 in</quantity>
|
||
<typeStatus id="E6EC19784C38A68F4CD29DA65F82C7F8" type="holotype">holotype</typeStatus>
|
||
), its tip pointed; distinct median groove ventrally on snout, its anterior tip reaching to midpoint of anterior-nostril-tube base; slope of dorsal surface of snout smooth, without notch or distinct hump. Anterior nostril tubular, short, tube length about equal to pupil diameter; posterior nostril oval in shape with an inner valve, located on anteroventral margin of eye, opening ventrally, covered by a flap extending slightly below edge of mouth gape. Eye small, covered by a transparent skin; center of eye anterior to mid-jaw (Figs
|
||
<figureCitation id="7C519377BE5AF12361F4A039DF3A1638" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Fresh condition of Yirrkala nkust sp. nov., holotype, NMMB-P 38652, 496 mm TL A whole body B lateral view of head C dorsal view of head. AG anterior edge of gill opening AN anus DO dorsal-fin origin." figureDoi="10.3897/zookeys.1220.130885.figure1" httpUri="https://binary.pensoft.net/fig/1194273">1 B</figureCitation>
|
||
,
|
||
<figureCitation id="2230A027D19C924F0E1FA5F2F1E6A3AE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Fresh condition (after freezing) of Yirrkala nkust sp. nov., paratype, KMNH VR 100650, 297 mm TL, photographed by Y. - C. Hsu A whole body B lateral view of head C dorsal view of head D ventral view of head. AG anterior edge of gill opening AN anus DO dorsal-fin origin." figureDoi="10.3897/zookeys.1220.130885.figure2" httpUri="https://binary.pensoft.net/fig/1194274">2 B</figureCitation>
|
||
). Interorbital region wide, weakly convex (Figs
|
||
<figureCitation id="AB8BE3578FBA136C355F0C391CD20995" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Fresh condition of Yirrkala nkust sp. nov., holotype, NMMB-P 38652, 496 mm TL A whole body B lateral view of head C dorsal view of head. AG anterior edge of gill opening AN anus DO dorsal-fin origin." figureDoi="10.3897/zookeys.1220.130885.figure1" httpUri="https://binary.pensoft.net/fig/1194273">1 C</figureCitation>
|
||
,
|
||
<figureCitation id="11F8EAF20F0459499391037FFE388E62" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Fresh condition (after freezing) of Yirrkala nkust sp. nov., paratype, KMNH VR 100650, 297 mm TL, photographed by Y. - C. Hsu A whole body B lateral view of head C dorsal view of head D ventral view of head. AG anterior edge of gill opening AN anus DO dorsal-fin origin." figureDoi="10.3897/zookeys.1220.130885.figure2" httpUri="https://binary.pensoft.net/fig/1194274">2 C</figureCitation>
|
||
). Mouth inferior, distance from tip of snout to anterior tip of lower jaw 3.0–3.3 times eye diameter (
|
||
<quantity id="C7B17AB27CF112B41E20F37E9DC2AC1E" metricMagnitude="-2" metricUnit="m" metricValue="8.382" unit="in" value="3.3">3.3 in</quantity>
|
||
<typeStatus id="CC63171E75E3B5DCBEB5A9736A0FE99C" type="holotype">holotype</typeStatus>
|
||
); lower jaw short, its tip relatively pointed, not reaching anterior-nostril tube (Figs
|
||
<figureCitation id="8195712C3221779CE7097E1BC511BDD7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Fresh condition of Yirrkala nkust sp. nov., holotype, NMMB-P 38652, 496 mm TL A whole body B lateral view of head C dorsal view of head. AG anterior edge of gill opening AN anus DO dorsal-fin origin." figureDoi="10.3897/zookeys.1220.130885.figure1" httpUri="https://binary.pensoft.net/fig/1194273">1 B</figureCitation>
|
||
,
|
||
<figureCitation id="5FE515CD255AF40084C38AE2C504936D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Fresh condition (after freezing) of Yirrkala nkust sp. nov., paratype, KMNH VR 100650, 297 mm TL, photographed by Y. - C. Hsu A whole body B lateral view of head C dorsal view of head D ventral view of head. AG anterior edge of gill opening AN anus DO dorsal-fin origin." figureDoi="10.3897/zookeys.1220.130885.figure2" httpUri="https://binary.pensoft.net/fig/1194274">2 B, D</figureCitation>
|
||
); rictus short, posterior end of gape slightly behind a vertical through posterior margin of eye; lips smooth with a fold along upper lip, extending from second infraorbital pore to postorbital pore or rictus. Gill openings positioned ventrolaterally of breast, relatively close each side; shape of opening slightly curved, diameter more than twice eye diameter.
|
||
</paragraph>
|
||
<paragraph id="3A8696A99D6A8AB4C629574C22BE769B">
|
||
Sensory pores on head developed, arrangement of those pores as follows (Fig.
|
||
<figureCitation id="3887255D37AF6A64B3ACDAC26FEFBF80" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawing of lateral head A Yirrkala nkust sp. nov., holotype, NMMB-P 38652, 496 mm TL B Yirrkala kaupii, NMMB-P 38424, 328 mm TL C Yirrkala omanensis, KMNH VR 100616, 216 mm TL. Arrows indicate ethmoid = 1 st supraorbital (lefts), interorbital (middles) and mid-temporal (rights) pores." figureDoi="10.3897/zookeys.1220.130885.figure3" httpUri="https://binary.pensoft.net/fig/1194275">3 A</figureCitation>
|
||
): 1 (ethmoid) + 3 on supraorbital, 3 + 3 on infraorbital, 4 on lower jaw, 2 on preopercle, and 5 on supratemporal, one of those on mid-temporal; a single median interorbital pore.
|
||
</paragraph>
|
||
<caption id="B30665469146A425271FA99A0E6663CB" ID-DOI="10.3897/zookeys.1220.130885.figure3" ID-arpha="FDCE85CE-9693-543D-AE97-CE3CA483DB3D" httpUri="https://binary.pensoft.net/fig/1194275" startId="F3">
|
||
<paragraph id="A23954AC98041BDA1124D705E9EA0F5E">
|
||
<label id="046CE38AC8C862A1FB578D40C3EA0856">Figure 3.</label>
|
||
</paragraph>
|
||
<paragraph id="A8C77A72DAD784EA0830128E2F3BF2D4">
|
||
Line drawing of lateral head
|
||
<emphasis id="86189F8D1B1CC894CF6AF32558907545" bold="true">A</emphasis>
|
||
<taxonomicName id="67EEC0579750140E7235F4428246895E" authorityName="Hibino & Ho" authorityYear="2024" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="nkust" status="sp. nov.">
|
||
<emphasis id="99BC7D9BB2FAD25462FCF6D8B85BDBDF" italics="true">Yirrkala nkust</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8398A2888B71973CA5AEAFDEE281F6E" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype,
|
||
<specimenCode id="9EFF50CE12582CBCC0338B4AA4A812C0">NMMB-P 38652</specimenCode>
|
||
, 496 mm
|
||
<abbrev id="ABBRID0E1WAE" xlink_title="Total length">TL</abbrev>
|
||
<emphasis id="7BB0CFAF8C31B5907B6DFDF03318D353" bold="true">B</emphasis>
|
||
<taxonomicName id="A7B277E52D1E994D91BC85757BED74DF" baseAuthorityName="Bleeker" baseAuthorityYear="1858" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="kaupii">
|
||
<emphasis id="B00B7FCDF80D935F012FE8DAEB69DD92" italics="true">Yirrkala kaupii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<specimenCode id="C19C3DED05176992F4E12751AA184439">NMMB-P 38424</specimenCode>
|
||
, 328 mm
|
||
<abbrev id="ABBRID0ELXAE" xlink_title="Total length">TL</abbrev>
|
||
<emphasis id="F6517D99303D5335C91D0C2641355343" bold="true">C</emphasis>
|
||
<taxonomicName id="306BDC5CF6EE320ED217BB44DCA49183" authority=", KMNH VR" authorityName="KMNH VR" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="omanensis">
|
||
<emphasis id="5FD69186DB3D52B0688EB0305A302EFC" italics="true">Yirrkala omanensis</emphasis>
|
||
,
|
||
<specimenCode id="D458F9B8F2A9C6EE9D9F69161B57D61F">
|
||
<named-content id="20872327AE413E35BB0DE59280E9A9EF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kitakyushu-museum-and-institute-natural-history" xlink_title="Kitakyushu Museum of Natural History and Human History">KMNH VR</named-content>
|
||
100616
|
||
</specimenCode>
|
||
</taxonomicName>
|
||
, 216 mm
|
||
<abbrev id="ABBRID0EAYAE" xlink_title="Total length">TL</abbrev>
|
||
. Arrows indicate ethmoid = 1
|
||
<superScript id="6F955A94A0C16771893772AB443BF92C">st</superScript>
|
||
supraorbital (lefts), interorbital (middles) and mid-temporal (rights) pores.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B56B57F66FC2D40EFE114DDA5650364B">
|
||
Lateral-line pores small but obvious. Lateral line almost complete except for near tip of tail, nine anterior to a vertical through gill opening, seven or eight (eight in
|
||
<typeStatus id="58469CB12D5BF35F8ACE3A7D7BF8DCAF" type="holotype">holotype</typeStatus>
|
||
) anterior to dorsal-fin origin, 72–78 (77) anterior to mid anus, and total 149–152 (151).
|
||
</paragraph>
|
||
<paragraph id="EDE4A6F578E5A3822455943CDD6CC495">Teeth pointed, slightly recurved posteriorly; 12–15 teeth on maxilla and 12–13 dentary uniserial; 10–12 vomerine teeth biserial anteriorly and uniserial posteriorly; intermaxillary with four teeth arranged in two rows.</paragraph>
|
||
<paragraph id="F2436FCE3EDDC63BB583CFC7C349C3E8">Median fins low but obvious; origin of dorsal fin slightly behind a vertical through anterior edge of gill opening but not behind posterior edge, ending anterior to tip of tail about 1 eye diameter; origin of anal fin slightly behind anus, ending same as dorsal fin; caudal and pectoral fins absent.</paragraph>
|
||
<paragraph id="150BF7BF06EFBB8ECCBE4C8D4E2390A1">
|
||
<emphasis id="C7E61E0FAB74662ECCA13F80196F106B" bold="true">
|
||
<emphasis id="B6EE2457EAB1CBEDF5FB17465318932B" bold="true" italics="true">Coloration</emphasis>
|
||
.
|
||
</emphasis>
|
||
Just after captured (Fig.
|
||
<figureCitation id="A0B3C2DE11B626B04A3709D169B31612" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Fresh condition of Yirrkala nkust sp. nov., holotype, NMMB-P 38652, 496 mm TL A whole body B lateral view of head C dorsal view of head. AG anterior edge of gill opening AN anus DO dorsal-fin origin." figureDoi="10.3897/zookeys.1220.130885.figure1" httpUri="https://binary.pensoft.net/fig/1194273">1</figureCitation>
|
||
), body greenish brown anteriorly, reddish brown posteriorly; lateral-line pore not margined; head generally greenish brown, postorbital region pinkish; snout with extremely pale brown transverse band dorsally in
|
||
<typeStatus id="6AFAA5D5419F8A339193715F499E5914" type="holotype">holotype</typeStatus>
|
||
,
|
||
<typeStatus id="1C860F4BE4E3639DBBDE28E2D2A2773B" type="paratype">paratype</typeStatus>
|
||
also paler but tip of snout not darker; fins similar color of body. After preservation, body color faded, changed to brown to pale brown, trunk weakly bicolored, melanophores remaining dorsally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EZYAE" type="etymology">
|
||
<paragraph id="2FFAFADFB05AE429A997CC3147D83F51">
|
||
<heading id="0075B7D645B57A4FF91DB45BCC974A6A" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="8B69F16E4B0D8015CC9FEA2C6CA8E82A">
|
||
The scientific name
|
||
<taxonomicName id="366AF044F0BCF0396E6857D41C3AE74B" authorityName="Hibino & Ho" authorityYear="2024" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="nkust">
|
||
<emphasis id="0BE1324AAA4C98906DDCD5C94A33BC17" italics="true">nkust</emphasis>
|
||
</taxonomicName>
|
||
is the acronym of the National
|
||
<collectingRegion id="ABC30319F7702FD76C47DB27FA6A9C1E" country="Taiwan" name="Kaohsiung">Kaohsiung</collectingRegion>
|
||
University of Science and Technology,
|
||
<collectingRegion id="0723D870D49A26C2A50D98E1AA5528FD" country="Taiwan" name="Kaohsiung">Kaohsiung</collectingRegion>
|
||
,
|
||
<collectingCountry id="06C7EE38A6A057098FBE11EEC2186C91" name="Taiwan">Taiwan</collectingCountry>
|
||
, which supports our research work. Used as a noun.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EJZAE" type="distribution">
|
||
<paragraph id="21135E294B8549C7B71EFAC64BBCAFC4">
|
||
<heading id="5877BC00D1535EDD8108040F36A4C4B9" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="4263D6C61AD3742C469FD5D0E268EA88">
|
||
Known from the northern regions of
|
||
<collectingRegion id="72A488FE2946E8731605615B60FC8EE8" country="Taiwan" name="Penghu">Penghu</collectingRegion>
|
||
Islands, an archipelago in the
|
||
<collectingCountry id="B50B13783181F9213A5B01C6EB633252" name="Taiwan">Taiwan</collectingCountry>
|
||
Strait off western
|
||
<collectingCountry id="DF2D3C8BA23631E626B0A3F43EA468FE" name="Taiwan">Taiwan</collectingCountry>
|
||
. The type series was collected from bycatches of the Silver-stripe round herring (
|
||
<taxonomicName id="7C031803B9B56C16C0204D2FA8E7C035" baseAuthorityName="Temminck & Schlegel" baseAuthorityYear="1846" family="Clupeidae" genus="Spratelloides" kingdom="Animalia" order="Clupeiformes" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis id="78D9542DBCA5FEEF721663F661333CAB" italics="true">Spratelloides gracilis</emphasis>
|
||
</taxonomicName>
|
||
) at depths about
|
||
<quantity id="280EC995702EE3F34B59DDF355B7B91E" metricMagnitude="1" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" unit="m" value="40.0" valueMax="50.0" valueMin="30.0">
|
||
30–
|
||
<specimenCount id="A1CB5DBF14E7E058A498129555E4B40C" count="50" type="generic">50 m.</specimenCount>
|
||
</quantity>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E1ZAE" type="remarks">
|
||
<paragraph id="BEA7DFE46B53BA93F624870C0B5CE877">
|
||
<heading id="87E8FF802033F2B7306250939FD8A16B" reason="title">Remarks.</heading>
|
||
</paragraph>
|
||
<paragraph id="AF1E73B3764F269EC494AD7E12881DE9">
|
||
The position of the dorsal-fin origin compared to that of gill opening is an important character for diagnosing
|
||
<taxonomicName id="84420BDAEB5D922902E77B787C9DC91D" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="D901590CD5FB1D1340992A3CCAAC39FE" italics="true">Yirrkala</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
Based on the morphological information of several species (i. e.
|
||
<taxonomicName id="EDD0D06B60AC9C5D399E1805F3501D4B" authority="McCosker, 2011" authorityName="McCosker" authorityYear="2011" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="ori">
|
||
<emphasis id="38555F51F18DA61515D7497217C7B78E" italics="true">Yirrkala ori</emphasis>
|
||
<bibRefCitation id="CCEBAD9ED6BE62A969B1F27B7B3A10C5" author="McCosker JE" journalOrPublisher="Smithiana, Publications in Aquatic Biodiversity, Bulletin" pagination="45-50" refId="B19" refString="McCosker JE (2011) Two new Indo-Pacific species of the sand-eel genus Yirrkala (Anguilliformes: Ophichthidae). Smithiana, Publications in Aquatic Biodiversity, Bulletin 13: 45 - 50." title="Two new Indo-Pacific species of the sand-eel genus Yirrkala (Anguilliformes: Ophichthidae)." volume="13" year="2011">McCosker, 2011</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="91AD8E11E77F6FE777E1B50C765272B2" baseAuthorityName="Gunther" baseAuthorityYear="1872" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="misolensis">
|
||
<emphasis id="3D987BAAE0E084FDF7BCE9DEE836D977" italics="true">Y. misolensis</emphasis>
|
||
</taxonomicName>
|
||
), the origin of dorsal fin quite consistent, showing less intraspecific variation.
|
||
</paragraph>
|
||
<paragraph id="E4212FE7D4BEA7E533BB87C43E49AE67">
|
||
The new species has the dorsal-fin origin situated above gill opening (verticals between anterior and posterior edges of the opening). The character is shared by
|
||
<taxonomicName id="A1B3C2957048F3BC84206616B513FBB2" authorityName="McCosker" authorityYear="2011" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="ori">
|
||
<emphasis id="EA7C1531C7620E4283A68562EE1E09BA" italics="true">Y. ori</emphasis>
|
||
</taxonomicName>
|
||
only, as other 13 species, including
|
||
<taxonomicName id="F3A194A1FEED4257A681255B0F26BCD8" authorityName="Whitley" authorityYear="1940" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="chaselingi">
|
||
<emphasis id="0DC4E5F10B89FCA24100FEE4E6488C7C" italics="true">Y. chaselingi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="71B3B42F6B887BEFAF125C140AA12C28" baseAuthorityName="Bleeker" baseAuthorityYear="1864" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="lumbricoides">
|
||
<emphasis id="CB94EB4A34C228CC3CFBA62D2EDE46D7" italics="true">Y. lumbricoides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="CC0278FFF5275F2B2FF6D12925486886" authority="McCosker, 2006" authorityName="McCosker" authorityYear="2006" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="moorei">
|
||
<emphasis id="BD83121A5CC4B1C704EB59F6628F296B" italics="true">Yirrkala moorei</emphasis>
|
||
McCosker, 2006
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="CAA7789C5342167581637935AAAE8199" authority="(Gunther, 1870)" baseAuthorityName="Gunther" baseAuthorityYear="1870" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="tenuis">
|
||
<emphasis id="EC3EE472776377908A5CA329A35FD50C" italics="true">Yirrkala tenuis</emphasis>
|
||
(
|
||
<bibRefCitation id="D72DEC2CF62D6CD544D650AAE45876D9" author="Günther A" journalOrPublisher="Taylor and Francis, London" refId="B7" refString="Gunther A (1870) Catalogue of the Fishes in the British Museum, Vol. 8. Taylor and Francis, London, 549 pp." title="Catalogue of the Fishes in the British Museum, Vol. 8." year="1870">Günther, 1870</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
before;
|
||
<taxonomicName id="4DD4CB87321B2B27EB9CA134BF1C90D0" authority="McCosker, 2011" authorityName="McCosker" authorityYear="2011" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="calyptra">
|
||
<emphasis id="AB1DD3E55BDC16CCE73EC8C67F4D8CC5" italics="true">Yirrkala calyptra</emphasis>
|
||
<bibRefCitation id="353090B8F6228D21D6DC86FD92C2E554" author="McCosker JE" journalOrPublisher="Smithiana, Publications in Aquatic Biodiversity, Bulletin" pagination="45-50" refId="B19" refString="McCosker JE (2011) Two new Indo-Pacific species of the sand-eel genus Yirrkala (Anguilliformes: Ophichthidae). Smithiana, Publications in Aquatic Biodiversity, Bulletin 13: 45 - 50." title="Two new Indo-Pacific species of the sand-eel genus Yirrkala (Anguilliformes: Ophichthidae)." volume="13" year="2011">McCosker, 2011</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8007858A63D1238BC5331F8535455E25" baseAuthorityName="Weber & de Beaufort" baseAuthorityYear="1916" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="gjellerupi">
|
||
<emphasis id="4BB16BA1EB5BCC9CC72F5BABA05CF658" italics="true">Y. gjellerupi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8C1F77BFB3E0F344169E26174BE1F433" authority="McCosker, 1999" authorityName="McCosker" authorityYear="1999" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="insolitus">
|
||
<emphasis id="6803E8C6C8AE9EAB382AF59C96EA82F4" italics="true">Yirrkala insolitus</emphasis>
|
||
<bibRefCitation id="E2B24F3C4A3510BC5FFF9398A89DF202" author="McCosker JE" journalOrPublisher="Zoologie" pagination="571-588" refId="B17" refString="McCosker JE (1999) Pisces Anguilliformes: deepwater snake eels (Ophichthidae) from the New Caledonia region, southwest Pacific Ocean. Memoires du Museum National d'Histoire Naturelle, Paris (N. S.) (Serie A). Zoologie 180: 571 - 588." title="Pisces Anguilliformes: deepwater snake eels (Ophichthidae) from the New Caledonia region, southwest Pacific Ocean. Mémoires du Muséum National d’Histoire Naturelle, Paris (N. S.) (Série A)." volume="180" year="1999">McCosker, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2735748BC9D035725852B6AC3C152874" baseAuthorityName="Bleeker" baseAuthorityYear="1858" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="kaupii">
|
||
<emphasis id="8659B2A124A16FB782E4074211EFB638" italics="true">Y. kaupii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DAC32ABEFE285B405C900B6ED7ADDD8C" authority="(Bleeker, 1863)" baseAuthorityName="Bleeker" baseAuthorityYear="1863" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="macrodon">
|
||
<emphasis id="FD3930D55F17D6589CD334C2019BC40A" italics="true">Yirrkala macrodon</emphasis>
|
||
(
|
||
<bibRefCitation id="9CE20392E9B3135C38FC24BC7668646C" author="Bleeker P" journalOrPublisher="Nederlandsch Tijdschrift voor de Dierkunde" pagination="179-186" refId="B2" refString="Bleeker P (1863) Description de quelques especes nouvelles ou peu connues d'Ophisuroides de l'Inde archipelagique. Nederlandsch Tijdschrift voor de Dierkunde 1: 179 - 186." title="Description de quelques espèces nouvelles ou peu connues d’Ophisuroïdes de l’Inde archipélagique." volume="1" year="1863">Bleeker, 1863</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="651C2FC6450BF43E6A93AF8E792C3CBB" authority="(Klausewitz, 1964)" baseAuthorityName="Klausewitz" baseAuthorityYear="1964" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="maculata">
|
||
<emphasis id="527C583FDC6E85CA712164D8F888E6B6" italics="true">Yirrkala maculata</emphasis>
|
||
(Klausewitz, 1964)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="FE24CCF730A463A9A0F6FADBC3D9DB3D" baseAuthorityName="Gunther" baseAuthorityYear="1872" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="misolensis">
|
||
<emphasis id="B565DC5CD2CA378CA802405D7BCCB4F3" italics="true">Y. misolensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="293A10B485BA50C23459C2923533A028" authority="(Norman, 1939)" baseAuthorityName="Norman" baseAuthorityYear="1939" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="omanensis">
|
||
<emphasis id="D202EAA9000A89B39EA0B02433C6DEF4" italics="true">Yirrkala omanensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EB1EA4B787351CB13CA0CBCE121A047D" author="Norman JR" journalOrPublisher="Scientific Reports, John Murray Expedition" pagination="1-116" refId="B27" refString="Norman JR (1939) Fishes. The John Murray Expedition 1933 - 34. Scientific Reports, John Murray Expedition 7: 1 - 116." title="Fishes. The John Murray Expedition 1933-34." volume="7" year="1939">Norman, 1939</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A8B6DBA4BCA81FF1E7C1D1849E2F3E77" authority="(Herre, 1936)" baseAuthorityName="Herre" baseAuthorityYear="1936" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="philippinensis">
|
||
<emphasis id="6BDD3EB713142B9987BD4A9B11E2C735" italics="true">Yirrkala philippinensis</emphasis>
|
||
(Herre, 1936)
|
||
</taxonomicName>
|
||
, have their fin origins clearly behind the gill opening (
|
||
<bibRefCitation id="6CF86C2ACDADC1A3BEF5AA054DDC2118" author="Bleeker" firstAuthor="Bleeker" journalOrPublisher="Nederlandsch Tijdschrift voor de Dierkunde" pagination="179-186" refId="B2" refString="Bleeker P (1863) Description de quelques espèces nouvelles ou peu connues d’Ophisuroïdes de l’Inde archipélagique. Nederlandsch Tijdschrift voor de Dierkunde 1: 179–186." title="Description de quelques espèces nouvelles ou peu connues d’Ophisuroïdes de l’Inde archipélagique." volume="1" year="1863">Bleeker 1863</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="F8F524753D9F3FDE8F841E2B0AD87DF0" author="Whitley" firstAuthor="Whitley" refId="B31" refString="Whitley GP (1940) Illustrations of some Australian fishes. Australian Zoologist 9: 397–428 [pls. 30–31]." year="1940">Whitley 1940</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D44F22DA630CB61AB4FF95C84BB79D4E" author="McCosker" firstAuthor="McCosker" journalOrPublisher="Zoologie" pagination="571-588" refId="B17" refString="McCosker JE (1999) Pisces Anguilliformes: deepwater snake eels (Ophichthidae) from the New Caledonia region, southwest Pacific Ocean. Mémoires du Muséum National d’Histoire Naturelle, Paris (N. S.) (Série A). Zoologie 180: 571–588." title="Pisces Anguilliformes: deepwater snake eels (Ophichthidae) from the New Caledonia region, southwest Pacific Ocean. Mémoires du Muséum National d’Histoire Naturelle, Paris (N. S.) (Série A)." volume="180" year="1999">McCosker 1999</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="D780F778A0DC28D003D0480116896EC7" author="McCosker" firstAuthor="McCosker" refId="B18" refString="McCosker JE (2006) A new species of sand eel, Yirrkala moorei (Anguilliformes: Ophichthidae), from the South Pacific. Proceedings of the California Academy of Sciences (Series 4) 57: 373–377." year="2006">2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="F2B6CAA21AAF3F55B1F30838CB95C1DC" DOI="10.1353/psc.2007.0009" author="McCosker" etAl="et al." firstAuthor="McCosker" journalOrPublisher="Pacific Science" pagination="141-144" refId="B23" refString="McCosker JE, Boseto D, Jenkins AP (2007) Redescription of Yirrkala gjellerupi, a poorly known freshwater Indo-Pacific snake eel (Anguilliformes: Ophichthidae). Pacific Science 61: 141–144. https://doi.org/10.1353/psc.2007.0009" title="Redescription of Yirrkala gjellerupi, a poorly known freshwater Indo-Pacific snake eel (Anguilliformes: Ophichthidae)." volume="61" year="2007">McCosker et al. 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="5382D04BFCC7B6E7E44F4FC902BBF442" author="McCosker" firstAuthor="McCosker" journalOrPublisher="Smithiana, Publications in Aquatic Biodiversity, Bulletin" pagination="45-50" refId="B19" refString="McCosker JE (2011) Two new Indo-Pacific species of the sand-eel genus Yirrkala (Anguilliformes: Ophichthidae). Smithiana, Publications in Aquatic Biodiversity, Bulletin 13: 45–50." title="Two new Indo-Pacific species of the sand-eel genus Yirrkala (Anguilliformes: Ophichthidae)." volume="13" year="2011">McCosker 2011</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="947DA28ABC84DE06FE352D5403B7F4CB" DOI="10.11646/zootaxa.5189.1.13" author="Chiu" etAl="et al." firstAuthor="Chiu" journalOrPublisher="Zootaxa" pagination="114-121" refId="B4" refString="Chiu Y-C, Chen H-M, Shao K-T (2022) Additional description on morphology of the Misol snake eel from Taiwan, with four verified barcodes of life sequences. Zootaxa 5189: 114–121. https://doi.org/10.11646/zootaxa.5189.1.13" title="Additional description on morphology of the Misol snake eel from Taiwan, with four verified barcodes of life sequences." volume="5189" year="2022">Chiu et al. 2022</bibRefCitation>
|
||
; this study).
|
||
</paragraph>
|
||
<paragraph id="76F1B4FEC179FB72D8C7D78785A53BC6">
|
||
<taxonomicName id="C408C1879E6060358580F14F0B5E154E" authorityName="Hibino & Ho" authorityYear="2024" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="nkust" status="sp. nov.">
|
||
<emphasis id="45C32FD6B02E9E59DEA99705B3866579" italics="true">Yirrkala nkust</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="93A75061C9DF479D2152B85188D309EC" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="7B8B1E2F03407BF2768A13840402FB0F" authorityName="McCosker" authorityYear="2011" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="ori">
|
||
<emphasis id="78D7BD46161826268330968480C5A2FE" italics="true">Y. ori</emphasis>
|
||
</taxonomicName>
|
||
in having 1 + 3 supraorbital pores (vs 1 + 4), and the position of the lower-jaw tip (not reaching base of the anterior-nostril tube vs beyond posterior edge of the base). The new species has similar vertebral counts of
|
||
<taxonomicName id="20C9E523F95A33FEDC9929624C5BE93B" baseAuthorityName="Bleeker" baseAuthorityYear="1864" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="lumbricoides">
|
||
<emphasis id="B1B84FF0F35B8C86CC760A1FA03D7047" italics="true">Y. lumbricoides</emphasis>
|
||
</taxonomicName>
|
||
but not overlapped in the count of the predorsal vertebrae (7 or 8 vs
|
||
<quantity id="570F876098CEE666CA9771FA043B76E2" metricMagnitude="-1" metricUnit="m" metricValue="1.27" unit="in" value="5.0">5 in</quantity>
|
||
<taxonomicName id="F1E2801805436FF79609C04C6458E87B" baseAuthorityName="Bleeker" baseAuthorityYear="1864" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="lumbricoides">
|
||
<emphasis id="5DD3659D248C56EE4DACB8DF160BCCCA" italics="true">Y. lumbricoides</emphasis>
|
||
</taxonomicName>
|
||
), and the total vertebrae (147–152 vs 150–159) is available (
|
||
<bibRefCitation id="05AF07DAE1C56FA151819B0BEF055C6C" author="McCosker" editor="Heemstra PC & Heemstra E & Ebert DA & Holleman W & Randall JE" firstAuthor="McCosker" journalOrPublisher="Vol. 2. South African Institute for Aquatic Biodiversity, Makhanda" pagination="101-137" refId="B21" refString="McCosker JE (2022) Family Ophichthidae, Snake-eels, sand-eels and worm-eels. In: Heemstra PC, Heemstra E, Ebert DA, Holleman W, Randall JE (Eds) Coastal Fishes of the Western Indian Ocean. Vol. 2. South African Institute for Aquatic Biodiversity, Makhanda, 101–137." title="Family Ophichthidae, Snake-eels, sand-eels and worm-eels." volumeTitle="Coastal Fishes of the Western Indian Ocean." year="2022">McCosker 2022</bibRefCitation>
|
||
; this study).
|
||
</paragraph>
|
||
<paragraph id="78CBC480209FBA6B5078487923B8C01E">
|
||
In addition, although
|
||
<taxonomicName id="EF432FD6A0C9DD45AC9D7534EA6F7A16" authority="Zuiwe, 1793" authorityName="Zuiwe" authorityYear="1793" class="Amphibia" family="Muraenidae" genus="Muraena" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="fusca">
|
||
<emphasis id="5DA571B226EA8697088D05530894F48F" italics="true">Muraena fusca</emphasis>
|
||
Zuiwe, 1793
|
||
</taxonomicName>
|
||
was regarded as a valid species of
|
||
<taxonomicName id="E19D846878D890A0E56425B769120D9D" authorityName="Whitley" authorityYear="1940" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="genus">
|
||
<emphasis id="A8D720C856BA2266C52C0F7675CB1CF6" italics="true">Yirrkala</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="4AF8500C0398B119C53BF265492BF9DC" author="Fricke R & Mahafina J & Behivoke F & Jaonalison H & Léopold M & Ponton D" etAl="et al." firstAuthor="Fricke" journalOrPublisher="FishTaxa" pagination="1-132" refId="B6" refString="Fricke R, Mahafina J, Behivoke F, Jaonalison H, Léopold M, Ponton D (2018) Annotated checklist of the fishes of Madagascar, southwestern Indian Ocean, with 158 new records. FishTaxa 3: 1–132." title="Annotated checklist of the fishes of Madagascar, southwestern Indian Ocean, with 158 new records." volume="3" year="2018">Fricke et al. (2018)</bibRefCitation>
|
||
, this name should be treated as a
|
||
<emphasis id="9A74052357FFEA8808FF5AB1F61EBD4F" italics="true">nomen dubium</emphasis>
|
||
. Its original description is insufficient which lacks direct evidence to identify it to any ophichthid and whereabouts of its
|
||
<typeStatus id="5AFDFEEA835694E1F394E7F070B4148F" type="holotype">holotype</typeStatus>
|
||
is unknown.
|
||
<taxonomicName id="FB714D81604DCEEE51A26BB508B076D1" authority="Peters, 1855" authorityName="Peters" authorityYear="1855" family="Ophichthidae" genus="Sphagebranchus" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="EC641999A24A86EF4634F08E79411718" italics="true">Sphagebranchus brevirostris</emphasis>
|
||
Peters, 1855
|
||
</taxonomicName>
|
||
, which has been regarded as conspecific with
|
||
<taxonomicName id="1E2A1AFAF5B30DF568A91DAC14C43B1E" authority="(Fricke et al. 2018)" baseAuthorityName="Fricke" baseAuthorityYear="2018" class="Amphibia" family="Muraenidae" genus="Muraena" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="fusca">
|
||
<emphasis id="1FCC411C1A7CC0379C777789E21A89B7" italics="true">M. fusca</emphasis>
|
||
(
|
||
<bibRefCitation id="55F2D04F89D6B9D390819BB9E983EB2B" author="Fricke" etAl="et al." firstAuthor="Fricke" journalOrPublisher="FishTaxa" pagination="1-132" refId="B6" refString="Fricke R, Mahafina J, Behivoke F, Jaonalison H, Léopold M, Ponton D (2018) Annotated checklist of the fishes of Madagascar, southwestern Indian Ocean, with 158 new records. FishTaxa 3: 1–132." title="Annotated checklist of the fishes of Madagascar, southwestern Indian Ocean, with 158 new records." volume="3" year="2018">Fricke et al. 2018</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, should be treated in a future work.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |