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<document id="729515BEAE1FF43394D3EA88705BB5D7" ID-CLB-Dataset="3385" ID-DOI="10.11646/zootaxa.4958.1.38" ID-GBIF-Dataset="47fb68b4-6fa6-4898-a9a7-adbb9fe77c42" ID-ISSN="1175-5326" ID-Zenodo-Dep="4692683" ID-ZooBank="05DDD729-45BF-4BC5-B1A0-399403609A93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618477958168" checkinUser="plazi" docAuthor="Eger, Joseph E." docDate="2021" docId="03B64A2DFFC7FFBCFF3EFDD8FED9434C" docLanguage="en" docName="zootaxa.4958.1.38.pdf" docOrigin="Zootaxa 4958 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Graziaedessa Eger 2021, n. gen." docType="treatment" docVersion="6" lastPageNumber="645" masterDocId="FF8F3255FFC6FFBEFFA9FF9EFFA5470B" masterDocTitle="Graziaedessa anastrephae (Heteroptera: Pentatomidae: Edessinae) a new genus and species collected in Multilure fruit fly traps baited with ammonium acetate and putrescine" masterLastPageNumber="648" masterPageNumber="643" pageNumber="644" updateTime="1698936951072" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F88790D8CE6865F28162BD7DC68D5DF5">Graziaedessa anastrephae (Heteroptera: Pentatomidae: Edessinae) a new genus and species collected in Multilure fruit fly traps baited with ammonium acetate and putrescine</mods:title>
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<mods:namePart id="2C34F5B183233DA7E8B5AAEEF34055CB">Eger, Joseph E.</mods:namePart>
<mods:affiliation id="4030AAE44DBEE03079A7F9DA7E75D17D">Florida State Collection of Arthropods, 2606 South Dundee Street, Tampa, FL, 33629, USA.</mods:affiliation>
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<mods:date id="8F6F92E2983AA93997F0C8F1F72BD195">2021</mods:date>
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<treatment id="03B64A2DFFC7FFBCFF3EFDD8FED9434C" ID-DOI="http://doi.org/10.5281/zenodo.4692685" ID-GBIF-Taxon="190310634" ID-Zenodo-Dep="4692685" LSID="urn:lsid:plazi:treatment:03B64A2DFFC7FFBCFF3EFDD8FED9434C" httpUri="http://treatment.plazi.org/id/03B64A2DFFC7FFBCFF3EFDD8FED9434C" lastPageId="2" lastPageNumber="645" pageId="1" pageNumber="644">
<subSubSection id="C305A8B0FFC7FFBFFF3EFDD8FE35456A" box="[151,400,582,609]" pageId="1" pageNumber="644" type="nomenclature">
<paragraph id="8BA0FB3BFFC7FFBFFF3EFDD8FE35456A" blockId="1.[151,400,582,645]" box="[151,400,582,609]" pageId="1" pageNumber="644">
<heading id="D0E84C57FFC7FFBFFF3EFDD8FE35456A" bold="true" box="[151,400,582,609]" fontSize="11" level="1" pageId="1" pageNumber="644" reason="1">
<emphasis id="B96B2729FFC7FFBFFF3EFDD8FE35456A" bold="true" box="[151,400,582,609]" pageId="1" pageNumber="644">
<taxonomicName id="4C1F80B8FFC7FFBFFF3EFDD8FE92456A" authority="Eger, 2021" authorityName="Eger" authorityYear="2021" box="[151,311,582,609]" class="Insecta" family="Tephritidae" genus="Graziaedessa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="644" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96B2729FFC7FFBFFF3EFDD8FE92456A" bold="true" box="[151,311,582,609]" italics="true" pageId="1" pageNumber="644">Graziaedessa</emphasis>
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<taxonomicNameLabel id="A2589A52FFC7FFBFFE97FDD9FE35456A" box="[318,400,583,609]" pageId="1" pageNumber="644" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C305A8B0FFC7FFBFFF3EFDF5FE85458E" box="[151,288,619,645]" pageId="1" pageNumber="644" type="description">
<paragraph id="8BA0FB3BFFC7FFBFFF3EFDF5FE85458E" blockId="1.[151,400,582,645]" box="[151,288,619,645]" pageId="1" pageNumber="644">
(
<figureCitation id="1324E7BEFFC7FFBFFF36FDF5FEBD458E" box="[159,280,619,645]" captionStart-0="FIGURES 12" captionStart-1="FIGURES 37" captionStart-2="FIGURES 810" captionStartId-0="1.[151,264,1655,1679]" captionStartId-1="3.[151,264,1499,1523]" captionStartId-2="4.[151,264,1634,1658]" captionTargetBox-0="[153,1435,977,1632]" captionTargetBox-1="[174,1432,181,1465]" captionTargetBox-2="[156,1431,921,1611]" captionTargetId-0="figure-226@1.[153,1435,977,1632]" captionTargetId-1="figure-15@3.[156,1432,181,1474]" captionTargetId-2="figure-308@4.[156,1431,921,1611]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 12. Graziaedessa anastrephae. 1 dorsal habitus; 2 ventral habitus. Dimensional line equals 1.0 mm." captionText-1="FIGURES 37. Graziaedessa anastrephae. 35 male pygophore; 3 ventral view, 4 dorsal view, 5 caudal view; 6 thoracic sterna; 7 female genital plates (dr—dorsal rim of genital cup, gc8—gonocoxite 8, gc9—gonocoxites 9, gp—genital cup process, la8—laterotergite 8, la9—laterotergite 9, msc—mesosternal carina, mtp—metasternal process, pa—paramere, pgc—projection of the floor of the genital cup, pr—proctiger, vr—ventral rim of genital cup). Dimensional lines equal 1.0 mm." captionText-2="FIGURES 810. Graziaedessa anastrephae. 89 paramere, 8 anterior face, 9 posterior face; 10 phallus, lateral view (llh—lateral lobe of the head of the paramere, mlh—mesial lobe of the head of the paramere, ph—phallotheca, sh—shank of paramere). Dimensional lines equal 0.25 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4692687" figureDoi-1="http://doi.org/10.5281/zenodo.4692689" figureDoi-2="http://doi.org/10.5281/zenodo.4692691" httpUri-0="https://zenodo.org/record/4692687/files/figure.png" httpUri-1="https://zenodo.org/record/4692689/files/figure.png" httpUri-2="https://zenodo.org/record/4692691/files/figure.png" pageId="1" pageNumber="644">Figs. 110</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C305A8B0FFC7FFBFFF3EFD34FCAB45C9" box="[151,782,682,706]" pageId="1" pageNumber="644" type="type_taxon">
<paragraph id="8BA0FB3BFFC7FFBFFF3EFD34FCAB45C9" blockId="1.[151,782,682,706]" box="[151,782,682,706]" pageId="1" pageNumber="644">
<emphasis id="B96B2729FFC7FFBFFF3EFD34FE8345C9" bold="true" box="[151,294,682,706]" pageId="1" pageNumber="644">
<typeStatus id="54A44599FFC7FFBFFF3EFD34FF6945C9" box="[151,204,682,706]" pageId="1" pageNumber="644">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C1F80B8FFC7FFBFFE85FD34FD9D45C9" authority="Eger, 2021" authorityName="Eger" authorityYear="2021" box="[300,568,682,706]" class="Insecta" family="Tephritidae" genus="Graziaedessa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="644" phylum="Arthropoda" rank="species" species="anastrephae" status="sp. nov.">
<emphasis id="B96B2729FFC7FFBFFE85FD34FD9D45C9" box="[300,568,682,706]" italics="true" pageId="1" pageNumber="644">Graziaedessa anastrephae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2589A52FFC7FFBFFD96FD34FDD245CA" box="[575,631,682,705]" pageId="1" pageNumber="644" rank="species">n. sp.</taxonomicNameLabel>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C305A8B0FFC7FFBFFF3EFD75FA8F4446" pageId="1" pageNumber="644" type="etymology">
<paragraph id="8BA0FB3BFFC7FFBFFF3EFD75FA8F4446" blockId="1.[151,1437,746,953]" pageId="1" pageNumber="644">
<emphasis id="B96B2729FFC7FFBFFF3EFD75FE87440E" bold="true" box="[151,290,747,773]" pageId="1" pageNumber="644">Etymology:</emphasis>
Gender feminine. It is a pleasure to dedicate this genus to Jocélia Grazia, on the occasion of her 75
<superScript id="7C6A5673FFC7FFBFFA39FD74FA3845F3" attach="none" box="[1424,1437,746,760]" fontSize="6" pageId="1" pageNumber="644">th</superScript>
birthday, in recognition of her numerous contributions to the systematics of the Pentatomoidea, not only through her own work, but also through the many students she educated and developed over the course of her career.
</paragraph>
</subSubSection>
<subSubSection id="C305A8B0FFC7FFBCFF6EFCC8FAD845EF" lastPageId="2" lastPageNumber="645" pageId="1" pageNumber="644" type="description">
<paragraph id="8BA0FB3BFFC7FFBFFF6EFCC8FC0444B2" blockId="1.[151,1437,746,953]" pageId="1" pageNumber="644">
<emphasis id="B96B2729FFC7FFBFFF6EFCC8FEFC447B" bold="true" box="[199,345,854,880]" pageId="1" pageNumber="644">Description.</emphasis>
Medium sized (11.712.6), body shape oval, dorsum predominately light brown with darker markings; posterior 1/3 of pronotum with dark brown fascia (
<figureCitation id="1324E7BEFFC7FFBFFCDDFCE5FC1E449E" box="[884,955,891,917]" captionStart="FIGURES 12" captionStartId="1.[151,264,1655,1679]" captionTargetBox="[153,1435,977,1632]" captionTargetId="figure-226@1.[153,1435,977,1632]" captionTargetPageId="1" captionText="FIGURES 12. Graziaedessa anastrephae. 1 dorsal habitus; 2 ventral habitus. Dimensional line equals 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4692687" httpUri="https://zenodo.org/record/4692687/files/figure.png" pageId="1" pageNumber="644">Fig. 1</figureCitation>
); venter pale brown with intersegmental abdominal sutures and extreme lateral margins darker brown (
<figureCitation id="1324E7BEFFC7FFBFFCE7FC01FC3144B2" box="[846,916,927,953]" captionStart="FIGURES 12" captionStartId="1.[151,264,1655,1679]" captionTargetBox="[153,1435,977,1632]" captionTargetId="figure-226@1.[153,1435,977,1632]" captionTargetPageId="1" captionText="FIGURES 12. Graziaedessa anastrephae. 1 dorsal habitus; 2 ventral habitus. Dimensional line equals 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4692687" httpUri="https://zenodo.org/record/4692687/files/figure.png" pageId="1" pageNumber="644">Fig. 2</figureCitation>
).
</paragraph>
<caption id="DF60ABB3FFC7FFBFFF3EF9E9FAA84184" ID-DOI="http://doi.org/10.5281/zenodo.4692687" ID-Zenodo-Dep="4692687" box="[151,1293,1655,1680]" httpUri="https://zenodo.org/record/4692687/files/figure.png" pageId="1" pageNumber="644" startId="1.[151,264,1655,1679]" targetBox="[153,1435,977,1632]" targetPageId="1">
<paragraph id="8BA0FB3BFFC7FFBFFF3EF9E9FAA84184" blockId="1.[151,1293,1655,1680]" box="[151,1293,1655,1680]" pageId="1" pageNumber="644">
<emphasis id="B96B2729FFC7FFBFFF3EF9E9FE9F4184" bold="true" box="[151,314,1655,1679]" pageId="1" pageNumber="644">FIGURES 12.</emphasis>
<taxonomicName id="4C1F80B8FFC7FFBFFEE8F9E9FDE84184" authorityName="Eger" authorityYear="2021" box="[321,589,1655,1679]" class="Insecta" family="Tephritidae" genus="Graziaedessa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="644" phylum="Arthropoda" rank="species" species="anastrephae">
<emphasis id="B96B2729FFC7FFBFFEE8F9E9FDE84184" box="[321,589,1655,1679]" italics="true" pageId="1" pageNumber="644">Graziaedessa anastrephae</emphasis>
</taxonomicName>
. 1 dorsal habitus; 2 ventral habitus. Dimensional line equals 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BA0FB3BFFC7FFBFFF6EF920FA2A4063" blockId="1.[151,1437,1725,2040]" pageId="1" pageNumber="644">Head 1.6 times wider than long; ¾ of eyes protruding beyond lateral margin of mandibular plates; mandibular plates much longer than clypeus, converging and contiguous in front of clypeus. Bucculae well-developed, short, flap-like; first rostral segment exceeding posterior margin of bucculae. Rostrum surpassing procoxae, reaching anterior margin of metasternal process, not reaching mesocoxae; relative length of segments: 2&gt;1&gt;3=4. Antennal segment 1 relatively thick, short; remaining segments elongate and thin; relative length of segments: 4&gt;5&gt;3&gt;2&gt;1.</paragraph>
<paragraph id="8BA0FB3BFFC7FFBCFF6EF8ECFE3C466A" blockId="1.[151,1437,1725,2040]" lastBlockId="2.[151,1437,151,1095]" lastPageId="2" lastPageNumber="645" pageId="1" pageNumber="644">
Pronotum light brown with scattered darker punctation on anterior 2/3, posterior 1/3 with dark brown fascia, fascia notably more densely punctate than remainder of pronotum; humeral angles little developed, narrowly rounded; anterior angles with small tooth; anterolateral margins straight, finely carinate. Scutellum subtriangular, post-frenal portion occupying about 1/3 of scutellar length. Coria mostly dark brown with pale markings. Ostiolar peritreme ruga-shaped, extending about ¾ of distance from ostiole to lateral margin of metapleuron. Evaporatoria shagreened, concolorous with remainder of pleura, impunctate; lateral margin of metapleural evaporatorium almost straight. Legs light brown with small dark macules, mostly at base of setae. Metasternum produced, bilobed posteriorly, simple and broadly rounded anteriorly, extending for short distance onto mesosternum (
<figureCitation id="1324E7BEFFC4FFBCFBE9FE9EFB2D4610" box="[1088,1160,256,283]" captionStart="FIGURES 37" captionStartId="3.[151,264,1499,1523]" captionTargetBox="[174,1432,181,1465]" captionTargetId="figure-15@3.[156,1432,181,1474]" captionTargetPageId="3" captionText="FIGURES 37. Graziaedessa anastrephae. 35 male pygophore; 3 ventral view, 4 dorsal view, 5 caudal view; 6 thoracic sterna; 7 female genital plates (dr—dorsal rim of genital cup, gc8—gonocoxite 8, gc9—gonocoxites 9, gp—genital cup process, la8—laterotergite 8, la9—laterotergite 9, msc—mesosternal carina, mtp—metasternal process, pa—paramere, pgc—projection of the floor of the genital cup, pr—proctiger, vr—ventral rim of genital cup). Dimensional lines equal 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4692689" httpUri="https://zenodo.org/record/4692689/files/figure.png" pageId="2" pageNumber="645">Fig. 6</figureCitation>
). Mesosternum carinate mesially, thinner anteriorly, reaching between procoxae, enlarged and divided posteriorly, outlining anterior margin of metasternal process.
</paragraph>
<paragraph id="8BA0FB3BFFC4FFBCFF6EFEF4FB3446E5" blockId="2.[151,1437,151,1095]" pageId="2" pageNumber="645">
Abdominal urosternites 36 with small dark tooth at posterolateral limit of segment, segment 7 moderately and acutely produced posteriorly, not reaching posterior margin of laterotergites
<quantity id="4CE756DEFFC4FFBCFC55FE13FB8C46AC" box="[1020,1065,397,423]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="2" pageNumber="645" unit="in" value="8.0">8 in</quantity>
females. Mesial trichobothria in line with spiracles, lateral trichobothria just laterad of this line. Urosternite 3 produced into a distinct mesial spine on anterior margin, this spine fitting into void between bifurcations of metasternal process.
</paragraph>
<paragraph id="8BA0FB3BFFC4FFBCFF6EFE69FCD04571" blockId="2.[151,1437,151,1095]" pageId="2" pageNumber="645">
Male genitalia: pygophore about as long as wide (
<figureCitation id="1324E7BEFFC4FFBCFD5DFE69FCC5451A" box="[756,864,503,529]" captionStart="FIGURES 37" captionStartId="3.[151,264,1499,1523]" captionTargetBox="[174,1432,181,1465]" captionTargetId="figure-15@3.[156,1432,181,1474]" captionTargetPageId="3" captionText="FIGURES 37. Graziaedessa anastrephae. 35 male pygophore; 3 ventral view, 4 dorsal view, 5 caudal view; 6 thoracic sterna; 7 female genital plates (dr—dorsal rim of genital cup, gc8—gonocoxite 8, gc9—gonocoxites 9, gp—genital cup process, la8—laterotergite 8, la9—laterotergite 9, msc—mesosternal carina, mtp—metasternal process, pa—paramere, pgc—projection of the floor of the genital cup, pr—proctiger, vr—ventral rim of genital cup). Dimensional lines equal 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4692689" httpUri="https://zenodo.org/record/4692689/files/figure.png" pageId="2" pageNumber="645">Figs. 3, 4</figureCitation>
); superior processes of genital cup present (
<figureCitation id="1324E7BEFFC4FFBCFAE0FE69FA28451A" box="[1353,1421,502,529]" captionStart="FIGURES 37" captionStartId="3.[151,264,1499,1523]" captionTargetBox="[174,1432,181,1465]" captionTargetId="figure-15@3.[156,1432,181,1474]" captionTargetPageId="3" captionText="FIGURES 37. Graziaedessa anastrephae. 35 male pygophore; 3 ventral view, 4 dorsal view, 5 caudal view; 6 thoracic sterna; 7 female genital plates (dr—dorsal rim of genital cup, gc8—gonocoxite 8, gc9—gonocoxites 9, gp—genital cup process, la8—laterotergite 8, la9—laterotergite 9, msc—mesosternal carina, mtp—metasternal process, pa—paramere, pgc—projection of the floor of the genital cup, pr—proctiger, vr—ventral rim of genital cup). Dimensional lines equal 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4692689" httpUri="https://zenodo.org/record/4692689/files/figure.png" pageId="2" pageNumber="645">Fig. 5</figureCitation>
); proctiger with distinct dorsal and posterior faces; shank of parameres relatively broad and short, head of parameres large and transversely oriented (
<figureCitation id="1324E7BEFFC4FFBCFDB7FDA3FD2B455C" box="[542,654,573,599]" captionStart="FIGURES 810" captionStartId="4.[151,264,1634,1658]" captionTargetBox="[156,1431,921,1611]" captionTargetId="figure-308@4.[156,1431,921,1611]" captionTargetPageId="4" captionText="FIGURES 810. Graziaedessa anastrephae. 89 paramere, 8 anterior face, 9 posterior face; 10 phallus, lateral view (llh—lateral lobe of the head of the paramere, mlh—mesial lobe of the head of the paramere, ph—phallotheca, sh—shank of paramere). Dimensional lines equal 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4692691" httpUri="https://zenodo.org/record/4692691/files/figure.png" pageId="2" pageNumber="645">Fig. 8, 9</figureCitation>
). Phallotheca cylindrical, strongly sclerotized, apparently lacking conjunctiva and vessica, its apex with a simple opening (
<figureCitation id="1324E7BEFFC4FFBCFCBDFDFEFCC24571" box="[788,871,608,634]" captionStart="FIGURES 810" captionStartId="4.[151,264,1634,1658]" captionTargetBox="[156,1431,921,1611]" captionTargetId="figure-308@4.[156,1431,921,1611]" captionTargetPageId="4" captionText="FIGURES 810. Graziaedessa anastrephae. 89 paramere, 8 anterior face, 9 posterior face; 10 phallus, lateral view (llh—lateral lobe of the head of the paramere, mlh—mesial lobe of the head of the paramere, ph—phallotheca, sh—shank of paramere). Dimensional lines equal 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4692691" httpUri="https://zenodo.org/record/4692691/files/figure.png" pageId="2" pageNumber="645">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="8BA0FB3BFFC4FFBCFF6EFD1DFAD845EF" blockId="2.[151,1437,151,1095]" pageId="2" pageNumber="645">
Female genitalia: gonocoxites 8 subtrapezoidal (
<figureCitation id="1324E7BEFFC4FFBCFD49FD1DFC864595" box="[736,803,643,670]" captionStart="FIGURES 37" captionStartId="3.[151,264,1499,1523]" captionTargetBox="[174,1432,181,1465]" captionTargetId="figure-15@3.[156,1432,181,1474]" captionTargetPageId="3" captionText="FIGURES 37. Graziaedessa anastrephae. 35 male pygophore; 3 ventral view, 4 dorsal view, 5 caudal view; 6 thoracic sterna; 7 female genital plates (dr—dorsal rim of genital cup, gc8—gonocoxite 8, gc9—gonocoxites 9, gp—genital cup process, la8—laterotergite 8, la9—laterotergite 9, msc—mesosternal carina, mtp—metasternal process, pa—paramere, pgc—projection of the floor of the genital cup, pr—proctiger, vr—ventral rim of genital cup). Dimensional lines equal 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4692689" httpUri="https://zenodo.org/record/4692689/files/figure.png" pageId="2" pageNumber="645">Fig. 7</figureCitation>
, gonocoxites 8 artificially opened in figure); gonocoxites 9 broadly exposed; apices of laterotergites 9 slightly surpassing band uniting laterotergites 8, the latter longitudinally convex, moderately developed posteriorly. Internal female genitalia not examined in the single female
<typeStatus id="54A44599FFC4FFBCFABCFD54FADC45EF" box="[1301,1401,714,740]" pageId="2" pageNumber="645" type="paratype">paratype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C305A8B0FFC4FFBCFF6EFD73FED9434C" pageId="2" pageNumber="645" type="discussion">
<paragraph id="8BA0FB3BFFC4FFBCFF6EFD73FED9434C" blockId="2.[151,1437,151,1095]" pageId="2" pageNumber="645">
<emphasis id="B96B2729FFC4FFBCFF6EFD73FEF7440C" bold="true" box="[199,338,749,775]" pageId="2" pageNumber="645">Comments:</emphasis>
<taxonomicName id="4C1F80B8FFC4FFBCFEF3FD73FE57440C" authorityName="Eger" authorityYear="2021" box="[346,498,749,775]" class="Insecta" family="Tephritidae" genus="Graziaedessa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="645" phylum="Arthropoda" rank="genus">
<emphasis id="B96B2729FFC4FFBCFEF3FD73FE57440C" box="[346,498,749,775]" italics="true" pageId="2" pageNumber="645">Graziaedessa</emphasis>
</taxonomicName>
is readily distinguished from all other genera of
<taxonomicName id="4C1F80B8FFC4FFBCFBB6FD73FB2A440C" baseAuthorityName="Barcellos &amp; Grazia" baseAuthorityYear="2003" box="[1055,1167,749,775]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="645" phylum="Arthropoda" rank="subFamily" subFamily="Edessinae">Edessinae</taxonomicName>
by the unique structure of the mesosternum and metasternum. It bears a superficial resemblance to
<taxonomicName id="4C1F80B8FFC4FFBCFC5BFC8FFBF74421" box="[1010,1106,785,810]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="645" phylum="Arthropoda" rank="genus">
<emphasis id="B96B2729FFC4FFBCFC5BFC8FFBF74421" box="[1010,1106,785,810]" italics="true" pageId="2" pageNumber="645">Hypoxys</emphasis>
</taxonomicName>
in overall appearance and in the slightly developed humeral angles and the black fascia on posterior 1/3 of the pronotum, dark markings on coria, female segment 7 not reaching posterior margin of laterotergites 8, the presence of a superior process of the genital cup, the structure of the proctiger, and the simple, cylindrical phallotheca. The carinate anterolateral pronotal margins distinguish this genus from
<taxonomicName id="4C1F80B8FFC4FFBCFD8FFC00FD2344BC" box="[550,646,926,951]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="645" phylum="Arthropoda" rank="genus">
<emphasis id="B96B2729FFC4FFBCFD8FFC00FD2344BC" box="[550,646,926,951]" italics="true" pageId="2" pageNumber="645">Hypoxys</emphasis>
</taxonomicName>
and most other edessines which have the anterolateral pronotal margins smoothly rounded.
<bibRefCitation id="EF8E86CAFFC4FFBCFED8FC5EFDC044D1" author="Almeida, F. R. A. &amp; Nunes, B. M. &amp; Fernandes, J. A. M." box="[369,613,960,986]" pageId="2" pageNumber="645" pagination="254 - 268" refId="ref3288" refString="Almeida, F. R. A., Nunes, B. M. &amp; Fernandes, J. A. M. (2018) A new genus and new species of Edessinae (Hemiptera: Heteroptera: Pentatomidae). Zootaxa, 4377 (2), 254 - 268. https: // doi. org / 10.11646 / zootaxa. 4377.2.6" type="journal article" year="2018">
Almeida
<emphasis id="B96B2729FFC4FFBCFE70FC5FFDB744D1" box="[473,530,960,986]" italics="true" pageId="2" pageNumber="645">et al.</emphasis>
(2018)
</bibRefCitation>
presented a key to genera of
<taxonomicName id="4C1F80B8FFC4FFBCFC1CFC5EFB8044D1" baseAuthorityName="Barcellos &amp; Grazia" baseAuthorityYear="2003" box="[949,1061,960,986]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="645" phylum="Arthropoda" rank="subFamily" subFamily="Edessinae">Edessinae</taxonomicName>
and subgenera of
<taxonomicName id="4C1F80B8FFC4FFBCFB5FFC5EFAE044D1" authorityName="Fabricius" authorityYear="1803" box="[1270,1349,960,986]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="645" phylum="Arthropoda" rank="genus">
<emphasis id="B96B2729FFC4FFBCFB5FFC5EFAE044D1" box="[1270,1349,960,986]" italics="true" pageId="2" pageNumber="645">Edessa</emphasis>
</taxonomicName>
. In this key,
<taxonomicName id="4C1F80B8FFC4FFBCFF62FC7AFEC644F5" authorityName="Eger" authorityYear="2021" box="[203,355,996,1022]" class="Insecta" family="Tephritidae" genus="Graziaedessa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="645" phylum="Arthropoda" rank="genus">
<emphasis id="B96B2729FFC4FFBCFF62FC7AFEC644F5" box="[203,355,996,1022]" italics="true" pageId="2" pageNumber="645">Graziaedessa</emphasis>
</taxonomicName>
keys to couplet 3 (metasternal process projected anteriorly in a simple process) but does not fit either of the choices of couplet 3, both genera in this couplet having the metasternal process extending over much of the mesosternum.
</paragraph>
</subSubSection>
</treatment>
</document>