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<document id="D42F8E76FBDE4BE51FB79B44BD48C946" ID-DOI="10.11646/zootaxa.3941.1.7" ID-GBIF-Dataset="31a60fc0-e1e2-43da-a652-a3da2e5143c3" ID-ISSN="1175-5326" ID-Zenodo-Dep="239551" ID-ZooBank="49EC3DD7-10DF-484B-A4D1-4F0880E7F33B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461024441417" checkinUser="plazi" docAuthor="Motyka, Michal &amp; Bocak, Ladislav" docDate="2015" docId="03B63135FFEEA14BFF40FADAFC7BF77F" docLanguage="en" docName="zt03941p130.pdf" docOrigin="Zootaxa 3941 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calochromus Guerin-Meneville" docType="treatment" docVersion="8" lastPageNumber="127" masterDocId="FF8F494DFFEFA149FFD7FFA9FFAEF376" masterDocTitle="The classification of Flabellochromus and Dumbrellia (Coleoptera: Lycidae) with descriptions of two new species of Calochromus from the Sundaland Region" masterLastPageNumber="130" masterPageNumber="125" pageNumber="126" updateTime="1698571534837" updateUser="plazi">
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<mods:title id="ED43E03BCCD42EA5589974C31B67F81F">The classification of Flabellochromus and Dumbrellia (Coleoptera: Lycidae) with descriptions of two new species of Calochromus from the Sundaland Region</mods:title>
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<mods:namePart id="BA3530B312199F8CD4C79DEF931A612B">Motyka, Michal</mods:namePart>
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<mods:namePart id="7554BB301AA4324B7F2FF68516DE9C40">Bocak, Ladislav</mods:namePart>
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<mods:date id="E1D6555781B6188C506738E8BFD4FD1B">2015</mods:date>
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<treatment id="03B63135FFEEA14BFF40FADAFC7BF77F" ID-DOI="http://doi.org/10.5281/zenodo.6112576" ID-GBIF-Taxon="119605949" ID-Zenodo-Dep="6112576" LSID="urn:lsid:plazi:treatment:03B63135FFEEA14BFF40FADAFC7BF77F" httpUri="http://treatment.plazi.org/id/03B63135FFEEA14BFF40FADAFC7BF77F" lastPageId="2" lastPageNumber="127" pageId="1" pageNumber="126">
<subSubSection id="C305D3A8FFEEA148FF40FADAFDB0F6F8" box="[151,542,1395,1422]" pageId="1" pageNumber="126" type="nomenclature">
<paragraph id="8BA08023FFEEA148FF40FADAFDB0F6F8" blockId="1.[151,542,1395,1422]" box="[151,542,1395,1422]" pageId="1" pageNumber="126">
<heading id="D0E8374FFFEEA148FF40FADAFDB0F6F8" bold="true" box="[151,542,1395,1422]" fontSize="11" level="1" pageId="1" pageNumber="126" reason="1">
<taxonomicName id="4C1FFBA0FFEEA148FF40FADAFDB0F6F8" ID-CoL="3FSN" ID-ENA="71196" authority="Guerin-Meneville" authorityName="Guerin-Meneville" box="[151,542,1395,1422]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEEA148FF40FADAFDB0F6F8" bold="true" box="[151,542,1395,1422]" pageId="1" pageNumber="126">
<emphasis id="B96B5C31FFEEA148FF40FADAFE94F6FB" bold="true" box="[151,314,1395,1421]" italics="true" pageId="1" pageNumber="126">Calochromus</emphasis>
Guérin-Méneville
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C305D3A8FFEEA148FF40FA13FDE2F6A6" box="[151,588,1466,1488]" pageId="1" pageNumber="126" type="reference_group">
<paragraph id="8BA08023FFEEA148FF40FA13FDE2F6A6" blockId="1.[151,917,1466,1642]" box="[151,588,1466,1488]" pageId="1" pageNumber="126">
<treatmentCitationGroup id="AB0FA70DFFEEA148FF40FA13FDE2F6A6" box="[151,588,1466,1488]" pageId="1" pageNumber="126">
<taxonomicName id="4C1FFBA0FFEEA148FF40FA13FDE2F6A6" ID-CoL="3FSN" authority="Guerin-Meneville, 1833: 158" authorityName="Guerin-Meneville" authorityPageNumber="158" authorityYear="1833" box="[151,588,1466,1488]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEEA148FF40FA13FEB1F6A6" box="[151,287,1466,1488]" italics="true" pageId="1" pageNumber="126">Calochromus</emphasis>
<treatmentCitation id="0ABEA632FFEEA148FEF3FA13FDE2F6A6" author="Guerin-Meneville" box="[292,588,1466,1488]" page="158" pageId="1" pageNumber="126" year="1833">
<bibRefCitation id="EF8EFDD2FFEEA148FEF3FA13FDB2F6A6" author="Guerin-Meneville" box="[292,540,1466,1488]" pageId="1" pageNumber="126" refString="Guerin-Meneville, F. E. (1833) Memoire sur deux nouveaux genres de lordre des coleopteres, et description des especes qui les composent. Annales de la Societe Entomologique de France, 2, 155 - 160." type="journal article" year="1833">Guérin-Méneville, 1833</bibRefCitation>
: 158
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</paragraph>
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<subSubSection id="C305D3A8FFEEA148FF10FA73FD6FF51C" pageId="1" pageNumber="126" type="materials_examined">
<paragraph id="8BA08023FFEEA148FF10FA73FD4BF578" blockId="1.[151,917,1466,1642]" pageId="1" pageNumber="126">
<typeStatus id="54A43E81FFEEA148FF10FA73FF57F699" box="[199,249,1498,1519]" pageId="1" pageNumber="126">Type</typeStatus>
species.
<taxonomicName id="4C1FFBA0FFEEA148FE82FA70FC3BF699" authority="Guerin-Meneville, 1833: 159" authorityName="Guerin-Meneville" authorityPageNumber="159" authorityYear="1833" box="[341,917,1497,1519]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="glaucopterus">
<emphasis id="B96B5C31FFEEA148FE82FA70FDC9F699" box="[341,615,1497,1519]" italics="true" pageId="1" pageNumber="126">Calochromus glaucopterus</emphasis>
<bibRefCitation id="EF8EFDD2FFEEA148FDBAFA70FCCBF699" author="Guerin-Meneville" box="[621,869,1497,1519]" pageId="1" pageNumber="126" refString="Guerin-Meneville, F. E. (1833) Memoire sur deux nouveaux genres de lordre des coleopteres, et description des especes qui les composent. Annales de la Societe Entomologique de France, 2, 155 - 160." type="journal article" year="1833">Guérin-Méneville, 1833</bibRefCitation>
: 159
</taxonomicName>
=
<taxonomicName id="4C1FFBA0FFEEA148FF7CFA51FD85F578" authority="Motschulsky, 1861: 138" authorityName="Motschulsky" authorityPageNumber="138" authorityYear="1861" box="[171,555,1528,1550]" class="Insecta" family="Lycidae" genus="Micronychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEEA148FF7CFA51FE80F578" box="[171,302,1528,1550]" italics="true" pageId="1" pageNumber="126">Micronychus</emphasis>
<bibRefCitation id="EF8EFDD2FFEEA148FEE3FA51FE57F578" author="Motschulsky" box="[308,505,1528,1550]" pageId="1" pageNumber="126" refString="Motschulsky, V. V. (1861) Essai d'un Catalogue des Insectes de l'ile Ceylan. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 34, 95 - 155." type="book chapter" year="1861">Motschulsky, 1861</bibRefCitation>
: 138
</taxonomicName>
;
<bibRefCitation id="EF8EFDD2FFEEA148FDE1FA51FD18F578" author="Kleine" box="[566,694,1528,1550]" pageId="1" pageNumber="126" refString="Kleine, R. (1933) Lycidae. In: Schenkling, S. (Ed.), Coleopterum catalogus auspiciis et auxilo W. Junk. Vol. 128. W. Junk, Berlin, 145 pp." type="book" year="1933">Kleine 1933</bibRefCitation>
: 106
</paragraph>
<paragraph id="8BA08023FFEEA148FF10F9BEFC80F53D" blockId="1.[151,917,1466,1642]" pageId="1" pageNumber="126">
<typeStatus id="54A43E81FFEEA148FF10F9BEFF57F55A" box="[199,249,1559,1580]" pageId="1" pageNumber="126">Type</typeStatus>
species.
<taxonomicName id="4C1FFBA0FFEEA148FE82F9BFFCFAF55A" authority="Motschulsky, 1861: 138" authorityName="Motschulsky" authorityPageNumber="138" authorityYear="1861" box="[341,852,1558,1580]" class="Insecta" family="Lycidae" genus="Micronychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B96B5C31FFEEA148FE82F9BFFDF5F55A" box="[341,603,1558,1580]" italics="true" pageId="1" pageNumber="126">Micronychus bimaculatus</emphasis>
<bibRefCitation id="EF8EFDD2FFEEA148FDB6F9BFFC8AF55A" author="Motschulsky" box="[609,804,1558,1580]" pageId="1" pageNumber="126" refString="Motschulsky, V. V. (1861) Essai d'un Catalogue des Insectes de l'ile Ceylan. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 34, 95 - 155." type="book chapter" year="1861">Motschulsky, 1861</bibRefCitation>
: 138
</taxonomicName>
=
<taxonomicName id="4C1FFBA0FFEEA148FF7CF99CFE4AF53D" authority="Pic, 1925: 11" authorityName="Pic" authorityPageNumber="11" authorityYear="1925" box="[171,484,1589,1611]" class="Insecta" family="Lycidae" genus="Flabellochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEEA148FF7CF99CFEF9F53D" box="[171,343,1589,1611]" italics="true" pageId="1" pageNumber="126">Flabellochromus</emphasis>
<bibRefCitation id="EF8EFDD2FFEEA148FE89F99CFE6FF53D" author="Pic" box="[350,449,1589,1611]" pageId="1" pageNumber="126" refString="Pic, M. (1925) Malacodermes exotiques. L'Echange, 418 - 426, 1 - 36, [hors-texte]" type="book chapter" year="1925">Pic, 1925</bibRefCitation>
: 11
</taxonomicName>
(as a subgenus of
<taxonomicName id="4C1FFBA0FFEEA148FD48F99CFC88F53D" box="[671,806,1589,1611]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEEA148FD48F99CFC88F53D" box="[671,806,1589,1611]" italics="true" pageId="1" pageNumber="126">Calochromus</emphasis>
</taxonomicName>
)
</paragraph>
<paragraph id="8BA08023FFEEA148FF10F9FCFD6FF51C" blockId="1.[151,917,1466,1642]" box="[199,705,1620,1642]" pageId="1" pageNumber="126">
<typeStatus id="54A43E81FFEEA148FF10F9FCFF57F51C" box="[199,249,1621,1642]" pageId="1" pageNumber="126">Type</typeStatus>
species.
<taxonomicName id="4C1FFBA0FFEEA148FE82F9FDFD6FF51C" authority="Pic, 1925: 11" authorityName="Pic" authorityPageNumber="11" authorityYear="1925" box="[341,705,1620,1642]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="B96B5C31FFEEA148FE82F9FDFD9BF51C" box="[341,565,1620,1642]" italics="true" pageId="1" pageNumber="126">Calochromus pallidus</emphasis>
<bibRefCitation id="EF8EFDD2FFEEA148FDECF9FDFD31F51C" author="Pic" box="[571,671,1620,1642]" pageId="1" pageNumber="126" refString="Pic, M. (1925) Malacodermes exotiques. L'Echange, 418 - 426, 1 - 36, [hors-texte]" type="book chapter" year="1925">Pic, 1925</bibRefCitation>
: 11
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C305D3A8FFEEA148FF40F93FFCC4F46D" pageId="1" pageNumber="126" type="diagnosis">
<paragraph id="8BA08023FFEEA148FF40F93FFCC4F46D" blockId="1.[151,1437,1686,2034]" pageId="1" pageNumber="126">
<emphasis id="B96B5C31FFEEA148FF40F93FFEBFF5D9" bold="true" box="[151,273,1686,1711]" pageId="1" pageNumber="126">Diagnosis.</emphasis>
<taxonomicName id="4C1FFBA0FFEEA148FECCF93FFE1FF5D8" box="[283,433,1686,1710]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEEA148FECCF93FFE1FF5D8" box="[283,433,1686,1710]" italics="true" pageId="1" pageNumber="126">Calochromus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C1FFBA0FFEEA148FD99F93EFD50F5D9" authority="Duges" authorityName="Duges" box="[590,766,1687,1711]" class="Insecta" family="Lycidae" genus="Lucaina" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEEA148FD99F93EFD04F5D8" box="[590,682,1687,1710]" italics="true" pageId="1" pageNumber="126">Lucaina</emphasis>
Dugès
</taxonomicName>
,
<taxonomicName id="4C1FFBA0FFEEA148FCDAF93EFBA5F5D9" authority="Mulsant" authorityName="Mulsant" box="[781,1035,1686,1711]" class="Insecta" family="Lycidae" genus="Lygistopterus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEEA148FCDAF93EFC0AF5D8" box="[781,932,1687,1710]" italics="true" pageId="1" pageNumber="126">Lygistopterus</emphasis>
Mulsant
</taxonomicName>
and
<taxonomicName id="4C1FFBA0FFEEA148FB90F93FFAFEF5D9" authority="Pic" authorityName="Pic" box="[1095,1360,1686,1711]" class="Insecta" family="Lycidae" genus="Macrolygistopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEEA148FB90F93FFA8FF5D8" box="[1095,1313,1686,1710]" italics="true" pageId="1" pageNumber="126">Macrolygistopterus</emphasis>
Pic
</taxonomicName>
in the absence of rostrum and from
<taxonomicName id="4C1FFBA0FFEEA148FE20F912FD7FF5A5" authority="Bourgeois" authorityName="Bourgeois" box="[503,721,1722,1747]" class="Insecta" family="Lycidae" genus="Adoceta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEEA148FE20F912FDFDF5A5" box="[503,595,1723,1747]" italics="true" pageId="1" pageNumber="126">Adoceta</emphasis>
Bourgeois
</taxonomicName>
in the stout, elongate body (
<quantity id="4CE72DC6FFEEA148FBFBF913FB0FF5A5" box="[1068,1185,1722,1747]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="2.8" metricValueMin="0.5" pageId="1" pageNumber="126" unit="mm" value="16.5" valueMax="28.0" valueMin="5.0">528 mm</quantity>
). The Nearctic genus
<taxonomicName id="4C1FFBA0FFEEA148FF40F977FE25F581" authority="Green" authorityName="Green" box="[151,395,1758,1783]" class="Insecta" family="Lycidae" genus="Caloptognatha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEEA148FF40F977FE91F580" box="[151,319,1758,1782]" italics="true" pageId="1" pageNumber="126">Caloptognatha</emphasis>
Green
</taxonomicName>
differs in having short mandibles. All species share the characteristic
<typeStatus id="54A43E81FFEEA148FB71F976FB78F581" box="[1190,1238,1759,1783]" pageId="1" pageNumber="126">type</typeStatus>
of male genitalia with wide parameres and asymmetrical phallobase (
<figureCitation id="13249CA6FFEEA148FD0CF8ABFCF0F46D" box="[731,862,1794,1819]" captionStart="FIGURES 12 15" captionStartId="4.[151,264,675,698]" captionTargetBox="[152,1434,194,649]" captionTargetId="figure@4.[152,1434,193,654]" captionTargetPageId="4" captionText="FIGURES 12 15. Male genitalia. 12 — Calochromus harauensis sp. nov.; 13 — C. lamellatus; 14 — C. kelantanensis sp. nov.; 15 — C. impressicornis. Scale 0.5 mm." httpUri="https://zenodo.org/record/239553/files/figure.png" pageId="1" pageNumber="126">Figs 1215</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C305D3A8FFEEA14BFF10F88FFE56F14C" lastPageId="2" lastPageNumber="127" pageId="1" pageNumber="126" type="description">
<paragraph id="8BA08023FFEEA14BFF10F88FFF4EF38E" blockId="1.[151,1437,1686,2034]" lastBlockId="2.[151,1437,151,1034]" lastPageId="2" lastPageNumber="127" pageId="1" pageNumber="126">
<emphasis id="B96B5C31FFEEA148FF10F88FFEDAF449" bold="true" box="[199,372,1830,1855]" pageId="1" pageNumber="126">Redescription.</emphasis>
Male. Body
<quantity id="4CE72DC6FFEEA148FDDCF88FFDDBF448" box="[523,629,1830,1855]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="2.8" metricValueMin="0.5" pageId="1" pageNumber="126" unit="mm" value="16.5" valueMax="28.0" valueMin="5.0">
5
<emphasis id="B96B5C31FFEEA148FDCEF88EFD8CF448" box="[537,546,1831,1854]" italics="true" pageId="1" pageNumber="126">-</emphasis>
28 mm
</quantity>
, parallel-sided to slightly widened backward. Head small, prognathous to slightly hypognathous, partly retracted in pronotum. Eyes small, hemispherically prominent, frontal interocular distance longer than maximum eye diameter. Labrum sclerotized, transverse, mandibles slender, slightly curved, incisor margin simple, maxilla with long, setose mala, maxillary palpi 4-segmented. Labium with plate-like mentum and shortly setose ligula, labial palpi 3-segmented. Antennal tubercles inconspicuous to deeply separated by narrow groove. Antennae 11-segmented, surpassing humeral third of elytra, filiform, weakly serrate to flabellate, scapus pear-like, robust, pedicel small, antennomere 3 much longer than pedicel, antennomere gradually slenderer to apex, sometimes middle antennomeres modified, antennomeres 3(4)-10 bearing lamellae in flabellate forms.
</paragraph>
<paragraph id="8BA08023FFEDA14BFF10FEADFF47F2DD" blockId="2.[151,1437,151,1034]" pageId="2" pageNumber="127">
Pronotum transverse to subquadrate with two longitudinal obtuse keels (
<figureCitation id="13249CA6FFEDA14BFBC0FEAAFB2FF26D" box="[1047,1153,259,284]" captionStart="FIGURES 1 11" captionStartId="3.[151,264,1881,1904]" captionTargetBox="[168,1431,545,1856]" captionTargetId="figure@3.[151,1436,543,1867]" captionTargetPageId="3" captionText="FIGURES 1 11. General appearance. 1 — Calochromus lamellatus; 2 — C. impressicornis; 3 — C. kelantanensis sp. nov.; 4 — C. harauensis sp. nov. Head. 5 — C. kelantanensis sp. nov.; 6 — C. harauensis sp. nov. Antenna. 7 — C. harauensis sp. nov.; 8 — C. lamellatus. Pronotum, the holotype of: 9 — Plateros brevicornis; 10 — P. melancholicus; 11 — P. barronensis. Photographs 9 11 taken by P. Hudson. Scales 2.0 mm (Figs 1 4), 1.0 mm (Figs 5 8)." httpUri="https://zenodo.org/record/239552/files/figure.png" pageId="2" pageNumber="127">Figs 14</figureCitation>
); anterior angles widely rounded, posterior ones acute to rounded. Scutellum long. Elytra flat, slightly widened backward; longitudinal elytral costae weak, often apically absent, transverse costae absent. Legs slender, femora robust and curved in some species, tibiae slender, tarsi 5-segmented, tarsomeres 3-4 with membranous pads, tarsomere 5 long, slender, claws simple.
</paragraph>
<paragraph id="8BA08023FFEDA14BFF10FE1CFC0CF284" blockId="2.[151,1437,151,1034]" pageId="2" pageNumber="127">
Male genitalia trilobate, phallus and parameres similar in length, parameres wide, phallus stout, almost straight, laterally compressed; phallobase asymmetrical (
<figureCitation id="13249CA6FFEDA14BFCC4FE70FC38F284" box="[787,918,473,498]" captionStart="FIGURES 12 15" captionStartId="4.[151,264,675,698]" captionTargetBox="[152,1434,194,649]" captionTargetId="figure@4.[152,1434,193,654]" captionTargetPageId="4" captionText="FIGURES 12 15. Male genitalia. 12 — Calochromus harauensis sp. nov.; 13 — C. lamellatus; 14 — C. kelantanensis sp. nov.; 15 — C. impressicornis. Scale 0.5 mm." httpUri="https://zenodo.org/record/239553/files/figure.png" pageId="2" pageNumber="127">Figs 1215</figureCitation>
).
</paragraph>
<paragraph id="8BA08023FFEDA14BFF10FE54FE56F14C" blockId="2.[151,1437,151,1034]" pageId="2" pageNumber="127">Female. Similar to male, always with weakly serrate to filiform antennae. Ovipositor very slender, with short coxites and very long valvifers.</paragraph>
</subSubSection>
<subSubSection id="C305D3A8FFEDA14BFF10FDEDFE45F12B" box="[199,491,580,605]" pageId="2" pageNumber="127" type="distribution">
<paragraph id="8BA08023FFEDA14BFF10FDEDFE45F12B" blockId="2.[151,1437,151,1034]" box="[199,491,580,605]" pageId="2" pageNumber="127">
<emphasis id="B96B5C31FFEDA14BFF10FDEDFEF7F12B" bold="true" box="[199,345,580,605]" pageId="2" pageNumber="127">Distribution</emphasis>
. Worldwide.
</paragraph>
</subSubSection>
<subSubSection id="C305D3A8FFEDA14BFF10FDC1FC7BF77F" pageId="2" pageNumber="127" type="discussion">
<paragraph id="8BA08023FFEDA14BFF10FDC1FC7BF77F" blockId="2.[151,1437,151,1034]" pageId="2" pageNumber="127">
<emphasis id="B96B5C31FFEDA14BFF10FDC1FE95F1F7" bold="true" box="[199,315,616,641]" pageId="2" pageNumber="127">Remarks.</emphasis>
<bibRefCitation id="EF8EFDD2FFEDA14BFE93FDC1FE15F1F7" author="Pic" box="[324,443,616,641]" pageId="2" pageNumber="127" refString="Pic, M. (1925) Malacodermes exotiques. L'Echange, 418 - 426, 1 - 36, [hors-texte]" type="book chapter" year="1925">Pic (1925)</bibRefCitation>
described
<taxonomicName id="4C1FFBA0FFEDA14BFDEEFDC1FD59F1F6" box="[569,759,616,640]" class="Insecta" family="Lycidae" genus="Flabellochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEDA14BFDEEFDC1FD59F1F6" box="[569,759,616,640]" italics="true" pageId="2" pageNumber="127">Flabellochromus</emphasis>
</taxonomicName>
as a subgenus of
<taxonomicName id="4C1FFBA0FFEDA14BFC1FFDC1FBF0F1F6" box="[968,1118,616,640]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEDA14BFC1FFDC1FBF0F1F6" box="[968,1118,616,640]" italics="true" pageId="2" pageNumber="127">Calochromus</emphasis>
</taxonomicName>
and based the taxon on the presence of flabellate male antennae.
<bibRefCitation id="EF8EFDD2FFEDA14BFD93FD25FD4FF1D3" author="Kleine" box="[580,737,652,677]" pageId="2" pageNumber="127" refString="Kleine, R. (1926) Neue Lycidae von den Sunda-Inseln. Treubia, 9, 293 - 316." type="journal article" year="1926">Kleine (1926)</bibRefCitation>
described
<taxonomicName id="4C1FFBA0FFEDA14BFC89FD25FAA5F1D3" authority="Kleine, 1926" authorityName="Kleine" authorityYear="1926" box="[862,1291,652,677]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="lamellatus">
<emphasis id="B96B5C31FFEDA14BFC89FD25FBDFF1D2" box="[862,1137,652,676]" italics="true" pageId="2" pageNumber="127">Calochromus lamellatus</emphasis>
<bibRefCitation id="EF8EFDD2FFEDA14BFBAEFD25FAA5F1D3" author="Kleine" box="[1145,1291,652,677]" pageId="2" pageNumber="127" refString="Kleine, R. (1926) Neue Lycidae von den Sunda-Inseln. Treubia, 9, 293 - 316." type="journal article" year="1926">Kleine, 1926</bibRefCitation>
</taxonomicName>
with similar flabellate antennae and noted a high similarity of external morphology and male genitalia of this new species and known
<taxonomicName id="4C1FFBA0FFEDA14BFF3DFD7DFE2EF19A" box="[234,384,724,748]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEDA14BFF3DFD7DFE2EF19A" box="[234,384,724,748]" italics="true" pageId="2" pageNumber="127">Calochromus</emphasis>
</taxonomicName>
with filiform antennae.
<bibRefCitation id="EF8EFDD2FFEDA14BFD47FD7AFC83F19A" author="Kleine" box="[656,813,723,748]" pageId="2" pageNumber="127" refString="Kleine, R. (1933) Lycidae. In: Schenkling, S. (Ed.), Coleopterum catalogus auspiciis et auxilo W. Junk. Vol. 128. W. Junk, Berlin, 145 pp." type="book" year="1933">Kleine (1933)</bibRefCitation>
listed
<taxonomicName id="4C1FFBA0FFEDA14BFCAFFD7DFC56F19A" box="[888,1016,724,748]" class="Insecta" family="Lycidae" genus="Dumbrellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEDA14BFCAFFD7DFC56F19A" box="[888,1016,724,748]" italics="true" pageId="2" pageNumber="127">Dumbrellia</emphasis>
</taxonomicName>
in
<taxonomicName id="4C1FFBA0FFEDA14BFBCBFD7AFB18F19A" box="[1052,1206,723,748]" class="Insecta" family="Lycidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="tribe" tribe="Calochromini">Calochromini</taxonomicName>
and in contrast with his previous opinion transferred both
<taxonomicName id="4C1FFBA0FFEDA14BFDBEFD5EFCC7F079" box="[617,873,759,783]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="B96B5C31FFEDA14BFDBEFD5EFCC7F079" box="[617,873,759,783]" italics="true" pageId="2" pageNumber="127">Calochromus pallidus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1FFBA0FFEDA14BFC65FD51FBFFF079" box="[946,1105,759,783]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="lamellatus">
<emphasis id="B96B5C31FFEDA14BFC65FD51FBFFF079" box="[946,1105,759,783]" italics="true" pageId="2" pageNumber="127">C. lamellatus</emphasis>
</taxonomicName>
to
<taxonomicName id="4C1FFBA0FFEDA14BFB50FD5EFAA9F079" box="[1159,1287,759,783]" class="Insecta" family="Lycidae" genus="Dumbrellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEDA14BFB50FD5EFAA9F079" box="[1159,1287,759,783]" italics="true" pageId="2" pageNumber="127">Dumbrellia</emphasis>
</taxonomicName>
and treated
<taxonomicName id="4C1FFBA0FFEDA14BFF40FCB2FE66F042" authority="Pic, 1925" authorityName="Pic" authorityYear="1925" box="[151,456,795,820]" class="Insecta" family="Lycidae" genus="Flabellochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEDA14BFF40FCB2FEFBF045" box="[151,341,795,819]" italics="true" pageId="2" pageNumber="127">Flabellochromus</emphasis>
<bibRefCitation id="EF8EFDD2FFEDA14BFE8AFCB2FE66F042" author="Pic" box="[349,456,795,820]" pageId="2" pageNumber="127" refString="Pic, M. (1925) Malacodermes exotiques. L'Echange, 418 - 426, 1 - 36, [hors-texte]" type="book chapter" year="1925">Pic, 1925</bibRefCitation>
</taxonomicName>
as a junior synonym of
<taxonomicName id="4C1FFBA0FFEDA14BFD0BFCB2FC76F045" authority="Lea, 1909" authorityName="Lea" authorityYear="1909" box="[732,984,795,819]" class="Insecta" family="Lycidae" genus="Dumbrellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEDA14BFD0BFCB2FCF3F045" box="[732,861,795,819]" italics="true" pageId="2" pageNumber="127">Dumbrellia</emphasis>
<bibRefCitation id="EF8EFDD2FFEDA14BFCB3FCB5FC76F045" author="Lea" box="[868,984,795,819]" pageId="2" pageNumber="127" refString="Lea, A. M. (1909) Revision of the Australian and Tasmanian Malacodermidae. Transactions of the Entomological Society of London, 1909, 45 - 251." type="journal article" year="1909">Lea, 1909</bibRefCitation>
</taxonomicName>
. We compared the external morphology of numerous species of Oriental
<taxonomicName id="4C1FFBA0FFEDA14BFDD8FC96FD0BF021" box="[527,677,831,855]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEDA14BFDD8FC96FD0BF021" box="[527,677,831,855]" italics="true" pageId="2" pageNumber="127">Calochromus</emphasis>
</taxonomicName>
with available species with flabellate antennae and we found that they do not differ in any character. Although many genus group taxa were based on antennal morphology, our research confirms Kleine's opinion and shows that flabellate antennae originated in many net-winged beetle lineages multiple times and that taxa based on this character often are polyphyletic (
<bibRefCitation id="EF8EFDD2FFEDA14BFB93FC03FB56F0B4" author="Bocakova" box="[1092,1272,938,963]" pageId="2" pageNumber="127" refString="Bocakova, M. (2001) Revision and the phylogenetic analysis of the subfamily Platerodinae (Coleoptera: Lycidae). European Journal of Entomology, 98, 53 - 85. http: // dx. doi. org / 10.14411 / eje. 2001.010" type="journal article" year="2001">Bocakova 2001</bibRefCitation>
,
<bibRefCitation id="EF8EFDD2FFEDA14BFAD4FC03FF56F090" author="Sklenarova" pageId="2" pageNumber="127" refString="Sklenarova, K., Kubecek, V. &amp; Bocak, L. (2014) Subtribal classification of Metriorrhynchini (Insecta: Coleoptera: Lycidae): an integrative approach using molecular phylogeny and morphology of adults and larvae. Arthropod Systematics &amp; Phylogeny, 72, 37 - 54." type="journal article" year="2014">
Sklenarova
<emphasis id="B96B5C31FFEDA14BFA5FFC02FF1AF090" italics="true" pageId="2" pageNumber="127">et al.</emphasis>
2014
</bibRefCitation>
). Therefore, we propose to place all species of
<taxonomicName id="4C1FFBA0FFEDA14BFCD1FC64FC0EF090" box="[774,928,973,998]" class="Insecta" family="Lycidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="tribe" tribe="Calochromini">Calochromini</taxonomicName>
with flabellate antennae in
<taxonomicName id="4C1FFBA0FFEDA14BFB01FC67FAC2F090" box="[1238,1388,974,998]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEDA14BFB01FC67FAC2F090" box="[1238,1388,974,998]" italics="true" pageId="2" pageNumber="127">Calochromus</emphasis>
</taxonomicName>
and to consider
<taxonomicName id="4C1FFBA0FFEDA14BFECEFC58FDE6F77C" authority="Pic, 1925" authorityName="Pic" authorityYear="1925" box="[281,584,1009,1034]" class="Insecta" family="Lycidae" genus="Flabellochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEDA14BFECEFC58FE79F77F" box="[281,471,1009,1033]" italics="true" pageId="2" pageNumber="127">Flabellochromus</emphasis>
<bibRefCitation id="EF8EFDD2FFEDA14BFE08FC58FDE6F77C" author="Pic" box="[479,584,1009,1034]" pageId="2" pageNumber="127" refString="Pic, M. (1925) Malacodermes exotiques. L'Echange, 418 - 426, 1 - 36, [hors-texte]" type="book chapter" year="1925">Pic, 1925</bibRefCitation>
</taxonomicName>
as a junior synonym
<taxonomicName id="4C1FFBA0FFEDA14BFCEEFC58FC61F77F" box="[825,975,1009,1033]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="B96B5C31FFEDA14BFCEEFC58FC61F77F" box="[825,975,1009,1033]" italics="true" pageId="2" pageNumber="127">Calochromus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>