243 lines
30 KiB
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243 lines
30 KiB
XML
<document ID-CLB-Dataset="72036" ID-DOI="http://dx.doi.org/10.3897/dez.69.83840" ID-GBIF-Dataset="eafc7326-93e2-4d55-b708-2da4b77e2e5e" ID-Pensoft-Pub="1860-1324-1-71" ID-Pensoft-UUID="DF59A16543B05799A4FD7B66E721FC76" ID-ZooBank="7DEB0587A3734B06BAADCA6102EC6581" ModsDocID="1860-1324-69-1-71" checkinTime="1655205803791" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav, Wrase, David W., Assmann, Thorsten, Muilwijk, Jan & Machard, Patrice" docDate="2022" docId="EA2B83521F5A560B9F375CE2836D3A15" docLanguage="en" docName="DeutEntomolZeit 69(1): 71-122" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-06-13" docSource="http://dx.doi.org/10.3897/dez.69.83840" docTitle="Platyderus (Eremoderus) irakensis Guéorguiev & Wrase & Assmann & Muilwijk & Machard 2022, sp. nov." docType="treatment" docUuid="95E89306-626A-468D-BA68-F16489ABB9F3" docUuidSource="ZooBank" docVersion="6" id="DF59A16543B05799A4FD7B66E721FC76" lastPageNumber="71" masterDocId="DF59A16543B05799A4FD7B66E721FC76" masterDocTitle="Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)" masterLastPageNumber="122" masterPageNumber="71" pageNumber="71" updateTime="1732679132351" updateUser="ExternalLinkService">
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<mods:title id="D0EC95A52A0A64A65345452A1F4C0A4B">Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)</mods:title>
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<mods:namePart id="B46EE9098E768824A00CA1B41C39737F">Gueorguiev, Borislav</mods:namePart>
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<mods:nameIdentifier id="01BA725F9850A3AE163C81B819EA1737" type="ORCID">https://orcid.org/0000-0002-8532-0657</mods:nameIdentifier>
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<mods:affiliation id="99B85C739947F745B2D237844D8BF6F0">National Museum of Natural History, Bulgarian Academy of Sciences, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
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<mods:nameIdentifier id="F9C79C40397C49BDABAD782AF0E59CF5" type="email">gueorguiev@nmnhs.com</mods:nameIdentifier>
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<mods:namePart id="90BC554A496B7F5D45DDFE9F02599C6B">Wrase, David W.</mods:namePart>
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<mods:nameIdentifier id="6B0C2991BABA2CEC1B6FFF6CED88998F" type="ORCID">https://orcid.org/0000-0003-1779-2684</mods:nameIdentifier>
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<mods:affiliation id="F2160884B6708801216DEEB7A8AE7AA3">Oderstr. 2, 15306, Gusow-Platkow, Germany</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="C0C68CB19B8721A41340E4F388F81105">Author</mods:roleTerm>
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<mods:namePart id="78391722ECC19E907B63E948C091A5BA">Assmann, Thorsten</mods:namePart>
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<mods:nameIdentifier id="1F14AB8F4D8C0113D2478A4E6B4510E9" type="ORCID">https://orcid.org/0000-0002-9203-769X</mods:nameIdentifier>
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<mods:affiliation id="8C3C6C88C149E456345B86F7E1251ABC">Leuphana University Lueneburg, Institute of Ecology, Universitaetsallee 1, 21335 Lueneburg, Germany</mods:affiliation>
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<mods:namePart id="300F11A77353CA843B413BBC46E3A8D9">Muilwijk, Jan</mods:namePart>
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<mods:affiliation id="7F89AFA820D4A3EF473699667519FBD2">Department of Entomology, Naturalis Biodiversity Centre, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
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<mods:namePart id="41861E7918D2ECE5B03C50EB2B455E93">Machard, Patrice</mods:namePart>
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<mods:affiliation id="84C566489DBDB9545E378A91274FA490">29 Chemin de Champigny, Molineuf, F- 41190 Valencisse, France</mods:affiliation>
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<mods:title id="4E22FE6EEECD831914C580ABCB15F25D">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="6D8F8DD2DB919B6E6F0BC78042F6BE91">2022</mods:date>
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<mods:number id="F70CB17BB891724E8E2406462D3B1E7C">2022-06-13</mods:number>
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<mods:number id="3E22D5C91B60D48FAFE5431E9F833193">69</mods:number>
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<mods:number id="77C1C279FF50241600FCA09E835A889E">1</mods:number>
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<mods:start id="C082F261FD61D10EF19B08003AD785FC">71</mods:start>
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<mods:classification id="56BF274A6DEE27E27DDE8F37B475B7EA">journal article</mods:classification>
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<mods:identifier id="85FAEBF1445406308F03B464EC9C8C5C" type="DOI">http://dx.doi.org/10.3897/dez.69.83840</mods:identifier>
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<treatment id="EA2B83521F5A560B9F375CE2836D3A15" ID-GBIF-Taxon="195868739" LSID="urn:lsid:zoobank.org:act:95E89306-626A-468D-BA68-F16489ABB9F3" httpUri="http://treatment.plazi.org/id/EA2B83521F5A560B9F375CE2836D3A15" lastPageNumber="71" pageId="0" pageNumber="71" scope_family="Carabidae" scope_order="Coleoptera" scope_subGenus="Platyderus" scope_tribe="Sphodrini">
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<subSubSection id="20F202DDA3F8AE4C71B4FF424AF13B1B" pageId="0" pageNumber="71" type="nomenclature">
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<paragraph id="637389988E6C022166BCBDFD54E03D33" pageId="0" pageNumber="71">
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17.
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<taxonomicName id="E4112E0CF492F61CA339EE26AC18FC84" LSID="http://zoobank.org/95E89306-626A-468D-BA68-F16489ABB9F3" authority="Guéorguiev & Wrase & Assmann & Muilwijk & Machard, 2022" authorityName="Guéorguiev & Wrase & Assmann & Muilwijk & Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) irakensis</taxonomicName>
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<taxonomicNameLabel id="21625B9AAEE70C42714932BAE911AA34" pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="465AD97BAA664265E0BB803CEB1026CF" pageId="0" pageNumber="71" type="description">
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<paragraph id="BC6170DB3CA790D38FF71BEE7FD4BFA8" pageId="0" pageNumber="71">
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<figureCitation id="29C53515C5AB5054054B3D6D9D9942E8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, " Jerusalem Syria ", Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">Figs 4B</figureCitation>
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<figureCitation id="667C57D771F485AC97CD149E1C81237E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche & Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">, 6C</figureCitation>
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<figureCitation id="A01A569C519E89C11F4C795BCD55EE6C" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche & Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15D</figureCitation>
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<figureCitation id="A81588F11A12216DDB4FEC33E1C46A8D" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">, 19</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="0A211E92523452922724E4E20BC916A1" pageId="0" pageNumber="71" type="type locality">
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<paragraph id="698953688EA00DD9616814168F77A57A" pageId="0" pageNumber="71">Type locality.</paragraph>
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<paragraph id="D67A7663EC33556FB489C66EB19B4064" pageId="0" pageNumber="71">Iraq, Al Anbar Governorate, Ar Rutba District, ca. 115 km east of Ar-Rutbah Town.</paragraph>
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</subSubSection>
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<subSubSection id="678553EF50BF2DB955FC12827D996ADE" pageId="0" pageNumber="71" type="type material">
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<paragraph id="F0C038F468DD8358F35D24090031E519" pageId="0" pageNumber="71">Type material.</paragraph>
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<paragraph id="A74EAF94713797DD0DBD6665600C74C5" pageId="0" pageNumber="71">
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<emphasis id="DD414B1198D1750EF4CA8937B04C0E25" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
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♀, 'IRAQ, Al-Anbar / Rutba, singled / 115 km. E of town / 8.I.1978 [w, h] // No. 350 /
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<normalizedToken id="8D3289290655D56D6598CE4C7810C5A2" originalValue="Topál">Topal</normalizedToken>
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& Zilahy [w, h]' (HNHM); paratype ♀, 'IRAQ, Al-Anbar / Rutba, singled / at 30 km W of / town, 9.I.1978 [w, h] // No. 351 /
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<normalizedToken id="DA3F61A1D021D2A0BE18E82D71F4E1F5" originalValue="Topál">Topal</normalizedToken>
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& Zilahy [w, h]' (NMNHS).
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</paragraph>
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<paragraph id="6944EEDFEFA22911CA3F3412908BF33D" pageId="0" pageNumber="71">TME: 2 specimens. TGE: 1♀.</paragraph>
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</subSubSection>
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<subSubSection id="617F52C03D6618EFFC7D5FB14AC7CFAB" pageId="0" pageNumber="71" type="etymology">
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<paragraph id="3CE51B53ED1B8F51873102A918EE1CA8" pageId="0" pageNumber="71">Etymology.</paragraph>
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<paragraph id="7108342DBB61E68AF15A777D2CD526FA" pageId="0" pageNumber="71">The specific epithet is a Latinized adjective, referring to the name of the country in which this new species was found.</paragraph>
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</subSubSection>
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<subSubSection id="DB84E0B3A940937786BCB9A61843CA5A" pageId="0" pageNumber="71" type="diagnosis">
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<paragraph id="E0BC7FC199737C9E979B4D126AE31DEA" pageId="0" pageNumber="71">Diagnosis.</paragraph>
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<paragraph id="DFCB230D97FA96DA92117D36E08EA0FA" pageId="0" pageNumber="71">
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<taxonomicName id="804433C3D97D8B14C4C095379761DDC4" authorityName="Guéorguiev & Wrase & Assmann & Muilwijk & Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
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<emphasis id="B6D7CDF8DAC38F9DE9DA1080113DEE39" italics="true" pageId="0" pageNumber="71">Platyderus irakensis</emphasis>
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</taxonomicName>
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is distinct from all other species of "
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<taxonomicName id="3AF36286D66FA9C65C944454CE73A970" baseAuthorityName="Reiche & Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
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<emphasis id="15C833457F17BA99A147C9AE011B98B9" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
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</taxonomicName>
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" group by the orange-brown color of integument (Fig.
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<figureCitation id="D8600DC906CA1C41933553F99776C99F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, " Jerusalem Syria ", Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4B</figureCitation>
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), mesotarsomeres and metatarsomeres dorsally flattened and slightly grooved (Fig.
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<figureCitation id="D730B85777CDCA0BD3613201B0D13C64" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche & Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">6C</figureCitation>
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), pronotum with sides to base slightly convex and posterior angles laterally not prominent, and distal enlargement of bursa copulatrix (Fig.
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<figureCitation id="2BF97CDBCD9DADA70DA5DACE9B7AE01C" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche & Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15D</figureCitation>
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); the last character may be an autapomorphy. Among Asian representatives of species group, it is the form with the highest mean values for EL/PL (= 2.69) and EL/EW (= 1.63), and, together with
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<taxonomicName id="32784EE0CD294C907FAC63E3FBC18B79" class="Insecta" kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
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<emphasis id="ECC97239E52B255954EAAE8D37F6B003" italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
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</taxonomicName>
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sp. nov., the one possessing lowest index for PW/PB (= 1.10).
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</paragraph>
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</subSubSection>
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<subSubSection id="A874EDF4BEE9E4940C18ECA211DE1E7B" pageId="0" pageNumber="71" type="description">
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<paragraph id="B8FC0AD9E057F59725693AE186F076B4" pageId="0" pageNumber="71">Description.</paragraph>
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<paragraph id="3793E59906F5E06FC5A8A6693F30A0C8" pageId="0" pageNumber="71">
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<emphasis id="65CBF4F76681606F8C1BA4FCF955241A" bold="true" pageId="0" pageNumber="71">
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<emphasis id="F06C283F9CA6C8292F58A8CDC9644B52" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
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.
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</emphasis>
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Specimens of moderate size for
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<taxonomicName id="1C8972CBDB4CDD4F0EC5EAB9498785E7" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
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<emphasis id="DF41C6AE90981F97C3A16C2556C9502F" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
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</taxonomicName>
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species (BL: 8.30-9.00 mm; BW: 2.90-3.20 mm), with elongate and fairly convex body (Fig.
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<figureCitation id="4EE4BE8E93BFD76B5412FBF39708EE02" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, " Jerusalem Syria ", Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4B</figureCitation>
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).
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<emphasis id="EEE27C7779F685ACD9A00D97682E159F" bold="true" pageId="0" pageNumber="71">
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<emphasis id="E09AD9D4236B50FE1DEC67755F5C6135" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
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.
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</emphasis>
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See Table
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<tableCitation id="A1C3FAC50812411B85D248BCF5BC904B" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
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.
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<emphasis id="3DAE9103AB93A296FAD8AA952D16B3F4" bold="true" pageId="0" pageNumber="71">
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<emphasis id="BF81F19B2389E453602BFF29A1017226" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
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.
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</emphasis>
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Body and appendages orange-brown, with head somewhat darker than the rest of the body and ventral surface lighter than dorsal surface. Integument slightly to moderately shiny, head and pronotum as shiny as elytra, ventral surface somewhat shinier than dorsal surface.
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<emphasis id="2EE59135DDC7FD1464DB79E5A373F5C8" bold="true" pageId="0" pageNumber="71">
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<emphasis id="F5180657761EB2D636FB458A62E37F4A" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
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.
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</emphasis>
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Pronotum with evident microreticulation, sculpticells regular isodiametric. Elytral intervals, scutellum and lateral gutter with distinct isodiametric sculpticells, sculpticells of basal margin more or less reduced. Ventral surface with isodiametric or slight transverse sculpticells, less apparent on epipleura and middle coxa. Head impunctate and smooth, only frontal furrows and lateral ends of clypeus, with very fine and short wrinkles. Pronotum surface mostly smooth, only apical part medially in front of anterior transverse impression and basal area medially behind posterior transverse impression longitudinally wrinkled; basal foveae and adjacent lateral areas shallowly punctate. Elytral intervals without evident punctures. Abdominal ventrites 1-3 finely wrinkled at sides, smooth medially.
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<emphasis id="8DB26D5B128ED40CB4794CE3F35663E5" bold="true" pageId="0" pageNumber="71">
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<emphasis id="D37F0E030BC3B0EAB7D64325170E2BFB" italics="true" pageId="0" pageNumber="71">Head</emphasis>
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.
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</emphasis>
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About one time and a third as wide as pronotum (PW/HW= 1.30-1.32). Eyes long, moderately convex. Labrum subrectangular, barely shorter than clypeus, with anterior margin concave medially. Frontoclypeal suture distinct in middle, reduced at sides. Frontal furrows punctiform, shallow. Paraorbital sulci straight, fine, backward surpassing posterior margin of eye, not reaching level of posterior supraorbital pore.
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<emphasis id="0F3A6EE49F48E95107FD6964CEC06B1D" bold="true" pageId="0" pageNumber="71">
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<emphasis id="2B1A971807F0F21443F426890C7CDEB1" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
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.
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</emphasis>
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Pronotum about a tenth wider than long (PW/PL= 1.13), with widest point at medial third. Anterior transverse impression indistinct; posterior transverse impression indistinct to barely distinct. Sides not sinuate, shallowly convex anteriorly, barely convex posteriorly; anterior bead present laterally, lacking or present in medial ninth to tenth; basal bead present laterally, reduced to absent in medial third. Metepisternum slightly longer than wide, MA/MI= 0.91-0.95.
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<emphasis id="136A1AE4A0388A9DDC22E7C95FB25A47" bold="true" pageId="0" pageNumber="71">
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<emphasis id="59B3B44CA1B5BA33CC5CE78213F90B0A" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
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.
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</emphasis>
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Elongate, about one and two thirds as long as wide (EL/EW= 1.62-1.63), two and two thirds as long as pronotum (EL/PL= 2.65-2.72), and almost one and a half times as wide as pronotum (EW/PW= 1.45-1.47), with widest point at first half of third quarter. Parascutellar striole and striae 1-8 shallowly impressed, superficially to indistinctly punctate; striole short, not joining stria 1; bases of striae not reaching basal bead (except for stria 2 in paratype). Interval 3 with three discal setiferous punctures (posterior puncture lacking on left elytron of holotype), all adjoining stria 2; one additional discal puncture adjoining stria 4 at anterior quarter of right elytron of paratype. Umbilicate setiferous series with 16 punctures on left elytron and 17 on right elytron in holotype, respectively, with 16 and 15 punctures in paratype.
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<emphasis id="A51D94C2ABD3D262530668F5078DA5F5" bold="true" pageId="0" pageNumber="71">
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<emphasis id="581451617B75624752E3AF54719B2697" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
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.
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</emphasis>
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Posterior side of profemur with one or two setae in basal third and one in medial third. Mesofemur mostly with 4 or 5 setiferous punctures on anterior side ventrally. Anterior side of metafemur ventrally with a few long setae, one in basal third and one-two in apical half.
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<emphasis id="185E476C2436FCC266757707EBDFDEA3" bold="true" pageId="0" pageNumber="71">
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<emphasis id="94289AF9AB5F31D3ED0000355626B899" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
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.
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</emphasis>
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|
Unknown.
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<emphasis id="F3B77E091BD521A1828AC9D995A8BA39" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
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|
(Fig.
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|
<figureCitation id="1199D535025DE4E745A1E841280ED1FC" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche & Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15D</figureCitation>
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). Apical gonocoxite with rounded apex and one dorsolateral ensiform seta. Bursa copulatrix two-chambered, with a shorter, but wider basal part and a longer, but narrower apical enlargement. Spermathecal canal connected in basal third of receptaculum.
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</paragraph>
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</subSubSection>
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|
<subSubSection id="23D5A6A598F79EDAD9DF32810EA017E4" pageId="0" pageNumber="71" type="comparisons">
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<paragraph id="26D5ADBF8ECBEDD9629DDF1BF38BF5DD" pageId="0" pageNumber="71">Comparisons.</paragraph>
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<paragraph id="3C7D2B011FDF0D81F0D862323B9825FA" pageId="0" pageNumber="71">
|
|
In addition to characters mentioned in
|
|
<normalizedToken id="8ED3017C9B4ADD3A66E84F303AE46475" originalValue="“Diagnosis”">"Diagnosis"</normalizedToken>
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|
, the new species differs from
|
|
<taxonomicName id="B95645334534393F864E514F1C0997C7" class="Insecta" kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
|
|
<emphasis id="1B2809E79BB636A6E281070F6A8D362F" italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
|
|
</taxonomicName>
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|
sp. nov., by: (1) head wider, compared to pronotum (PW/HW: 1.30-1.32, vs. PW/HW: 1.35-1.42); (2) pronotum with sides less constricted anteriorly and posteriorly (PW/PA: 1.34-1.38 and PW/PB: 1.09-1.11, vs. PW/PA: 1.39-1.52 and PW/PB: 1.15-1.22); (3) elytra longer compared to their width and the length of the pronotum (EL/EW: 1.62-1.63 and EL/PL: 2.65-2.72, vs. EL/EW: 1.56-1.60 and EL/PL: 2.41-2.60).
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</paragraph>
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<paragraph id="B8901E7F04899D0D2AB0DA119C770E57" pageId="0" pageNumber="71">
|
|
For differences between
|
|
<taxonomicName id="0DDA9139A917FF18BCB9A851F7E8AD91" class="Insecta" kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
|
|
<emphasis id="AAAA4A03A2962E6510368A78027DFD57" italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. and
|
|
<taxonomicName id="BB53620A2F9559B4D5BCB97C412DA7AE" class="Insecta" kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
|
|
<emphasis id="EDE7CB25D5B34E9B6C68A51D2084278C" italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. and such between
|
|
<taxonomicName id="75687745B76E9CF9ED74D7994D210A08" class="Insecta" kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
|
|
<emphasis id="F8FB7F833C7EA3A347FED0F8A0F44BDE" italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. and
|
|
<taxonomicName id="A8BBB69C4087893F0766E73F4C13D301" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
|
|
<emphasis id="065F1E20E294702C9EB086AAA9E12BEA" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
|
|
</taxonomicName>
|
|
, see
|
|
<normalizedToken id="58DDD8AB9DF29E9D3F68423A828BD207" originalValue="“Comparisons”">"Comparisons"</normalizedToken>
|
|
under the latter species.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="3ED63E3CE282185B862FF9134883C411" pageId="0" pageNumber="71" type="habitat">
|
|
<paragraph id="896A593812174DD8E72CA93FA8F70956" pageId="0" pageNumber="71">Habitat.</paragraph>
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|
<paragraph id="079B4F48D946D69A2B2D6A472BCF4E75" pageId="0" pageNumber="71">Nothing is known about the bionomics of this species.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="120463BBC54C65B991CD14F12530480F" pageId="0" pageNumber="71" type="distribution">
|
|
<paragraph id="482638913415734BAE8097A09A23F31D" pageId="0" pageNumber="71">Distribution.</paragraph>
|
|
<paragraph id="FF0F012F4829C8D882167533A593F5F8" pageId="0" pageNumber="71">
|
|
It is currently known from two localities situated in the Ar Rutba District, which is the largest district of Al Anbar Governorate, Western Iraq (Fig.
|
|
<figureCitation id="3CD49227915598816927E24DA782003D" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">19</figureCitation>
|
|
). The holotype was collected about 115 km east of Ar-Rutbah Town whilst the paratype was found about 30 km west of the same town. Most of the area of the Ar Rutba District is a high plateau.
|
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</paragraph>
|
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</subSubSection>
|
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</treatment>
|
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</document> |