treatments-xml/data/D1/48/87/D14887D4FF9D2725FF7C41EFB72EE1EC.xml

<document id="2461CBB8829B4C69DC7356A5A27BA011" ID-DOI="10.11865/zs.2023205" ID-ISSN="2095-6827" ID-Zenodo-Dep="10941802" ID-ZooBank="04888FCF-480A-4198-9638-7AD4F8034E9F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712577329921" checkinUser="felipe" docAuthor="Huang, Wenjie, Xie, Xiang, Wang, Antai &amp; Zhang, Yu" docDate="2023" docId="D14887D4FF9D2725FF7C41EFB72EE1EC" docLanguage="en" docName="ZoolSyst.48.2.147-158.pdf" docOrigin="Zoological Systematics 48 (2)" docSource="https://www.mendeley.com/catalogue/0196cf7b-5af1-31de-bf57-0423a961b100/" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Phaenocora shenda Huang &amp; Wang 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="150" masterDocId="2D71FFACFF9F2721FFF4427BB230E01B" masterDocTitle="Two new species of freshwater turbellarians of the family Typhloplanidae (Platyhelminthes: Rhabdocoela) from China" masterLastPageNumber="158" masterPageNumber="147" pageNumber="149" updateTime="1712578319164" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D0E95818172D8479E3A682D0669023AE">Two new species of freshwater turbellarians of the family Typhloplanidae (Platyhelminthes: Rhabdocoela) from China</mods:title>
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<mods:namePart id="A32300D01AF268223FA230A4C8577BC6">Huang, Wenjie</mods:namePart>
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<mods:namePart id="29B0B6533E05A56F83922C983D697E01">Xie, Xiang</mods:namePart>
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<mods:namePart id="9B006863CCF8B55217B875ABDC63BE0F">Wang, Antai</mods:namePart>
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<mods:namePart id="C083B48DDE3BD23A4CF0C0D5A884B02C">Zhang, Yu</mods:namePart>
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<treatment id="D14887D4FF9D2725FF7C41EFB72EE1EC" LSID="urn:lsid:plazi:treatment:D14887D4FF9D2725FF7C41EFB72EE1EC" httpUri="http://treatment.plazi.org/id/D14887D4FF9D2725FF7C41EFB72EE1EC" lastPageId="4" lastPageNumber="150" pageId="2" pageNumber="149">
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3.1 
<taxonomicName id="9EE14D41FF9D2723FF3041E8B06AE3B7" authority="Huang &amp; Wang" authorityName="Huang &amp; Wang" authorityYear="2023" box="[196,602,915,940]" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="2" pageNumber="149" phylum="Platyhelminthes" rank="species" species="shenda" status="sp. nov.">
<emphasis id="6B95EAD0FF9D2723FF3041E8B3AEE3B0" bold="true" box="[196,414,915,939]" italics="true" pageId="2" pageNumber="149">Phaenocora shenda</emphasis>
Huang &amp; Wang
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, 
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</emphasis>
(
<figureCitation id="C1DA2A47FF9D2723FD2941EFB10CE3B7" box="[733,828,916,940]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStartId-0="3.[155,218,1895,1917]" captionStartId-1="4.[155,218,1415,1437]" captionTargetBox-0="[189,1399,499,1857]" captionTargetBox-1="[189,1397,573,1369]" captionTargetId-0="figure-198@3.[189,1399,499,1857]" captionTargetId-1="figure-449@4.[189,1398,569,1370]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figure 1. Phaenocora shenda Huang &amp; Wang, sp. nov. A. Free-living individual with zoochlorellae, dorsal view. B. Free-living individual without zoochlorellae, dorsal view. C. An individual treated with histochemical localization of acetylcholinesterase. D–E. Photomicrographs of histological sections stained with hematoxylin and eosin. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso-intestinal duct; eg—egg; fgc—female genital canal; gp—genital pore; i—intestine; pe—penis; ph—pharynx; t—testis; ut—uterus; vd—vitelloduct; vi—vitellarium; vid—vitelline duct; vs—seminal vesicle. Scale bars: A–C, E = 100 μm; D=20 μm." captionText-1="Figure 2. Phaenocora shenda Huang &amp; Wang, sp. nov., reconstruction of the brain, pharynx and reproductive system. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso intestinal duct; fgc—female genital canal; gp—genital pore; i— intestine; m—mouth; ov—ovary; pe—penis; ph—pharynx; sga—superior genital atrium; ut—uterus; vd—vitelloduct; vde—vas deferens; vs—seminal vesicle. Scale bar=50μm." figureDoi-0="http://doi.org/10.5281/zenodo.10941806" figureDoi-1="http://doi.org/10.5281/zenodo.10941808" httpUri-0="https://zenodo.org/record/10941806/files/figure.png" httpUri-1="https://zenodo.org/record/10941808/files/figure.png" pageId="2" pageNumber="149">Figs 1–2</figureCitation>
)
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</paragraph>
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<subSubSection id="11FB6549FF9D2725FF4941B0B72EE1EC" lastPageId="4" lastPageNumber="151" pageId="2" pageNumber="149" type="etymology">
<paragraph id="595E36C2FF9D2723FF4941B0B74FE3F8" blockId="2.[136,1459,971,2070]" box="[189,1407,971,995]" pageId="2" pageNumber="149">Etymology. The species name is derived from the abbreviated Chinese name of Shenzhen University— “Shenda”.</paragraph>
<paragraph id="595E36C2FF9D2723FF494195B601E475" blockId="2.[136,1459,971,2070]" pageId="2" pageNumber="149">
Diagnosis. Species of 
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<emphasis id="6B95EAD0FF9D2723FE4F4195B00DE41D" box="[443,573,1006,1030]" italics="true" pageId="2" pageNumber="149">Phaenocora</emphasis>
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can be in green color or colorless, depending on whether the endosymbiotic zoochlorellae is present or not. The paired ramified vitellaria extend along both sides of the body, with a reticulate appearance. The male copulatory organ is characterized by the presence of an unarmed penis papilla, while the female copulatory organ is characterized by the presence of both a genito-bursal duct and a burso-intestinal duct.
</paragraph>
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<materialsCitation id="E9893C9FFF9D2723FF494602B04FE4CD" collectingDate="2021-03" collectionCode="IZCAS" collectorName="Wenjie Huang." country="China" latitude="22.529167" location="Xiawenshan Lake" longLatPrecision="21" longitude="113.93111" municipality="Shenzhen University" pageId="2" pageNumber="149" specimenCode="PLA-P001, PLA-P002-007" specimenCount="2" stateProvince="Guangdong" typeStatus="holotype">
Material examined. 
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. 
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(whole mount), a filter pond of the constructed wetland (
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, 
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), 
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, 
<collectingMunicipality id="B93AACB8FF9D2723FE0946E0B0D1E4A8" box="[509,737,1179,1203]" pageId="2" pageNumber="149">Shenzhen University</collectingMunicipality>
, 
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, 
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, 
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<collectingDate id="3D1BE9EAFF9D2723FC3F46E0B662E4AF" box="[971,1106,1179,1204]" pageId="2" pageNumber="149" value="2021-03">March 2021</collectingDate>
</date>
, leg. 
<collectorName id="F4145314FF9D2723FB6546E0B704E4AF" box="[1169,1332,1179,1204]" pageId="2" pageNumber="149">Wenjie Huang.</collectorName>
<typeStatus id="865A8860FF9D2723FAC946E7B798E4AF" box="[1341,1448,1180,1204]" pageId="2" pageNumber="149" type="paratype">Paratypes</typeStatus>
. 
<collectionCode id="3FF0AE07FF9D2723FF7C46C5B2E5E4CD" box="[136,213,1214,1238]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="2" pageNumber="149">IZCAS</collectionCode>
<specimenCode id="09479EB9FF9D2723FF2846C5B3B7E4CD" box="[220,391,1214,1238]" collectionCode="PLA-P" pageId="2" pageNumber="149">PLA-P002–007</specimenCode>
, same data as holotype
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.
</paragraph>
<paragraph id="595E36C2FF9D2723FF49469AB052E5D2" blockId="2.[136,1459,971,2070]" pageId="2" pageNumber="149">
Description. Mature worms body length 3.05 ± 
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(
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= 6), width 1.23 ± 
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(
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= 6). Body dorsoventrally flattened and abruptly narrow at caudal end. Head somewhat rounded anteriorly and broadened posteriorly, while visible eyes absent. Individual in green color or colorless, depending on whether endosymbiotic green algae zoochlorellae present or not (
<figureCitation id="C1DA2A47FF9D2723FF204732B37DE57A" box="[212,333,1353,1377]" captionStart="Figure 1" captionStartId="3.[155,218,1895,1917]" captionTargetBox="[189,1399,499,1857]" captionTargetId="figure-198@3.[189,1399,499,1857]" captionTargetPageId="3" captionText="Figure 1. Phaenocora shenda Huang &amp; Wang, sp. nov. A. Free-living individual with zoochlorellae, dorsal view. B. Free-living individual without zoochlorellae, dorsal view. C. An individual treated with histochemical localization of acetylcholinesterase. D–E. Photomicrographs of histological sections stained with hematoxylin and eosin. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso-intestinal duct; eg—egg; fgc—female genital canal; gp—genital pore; i—intestine; pe—penis; ph—pharynx; t—testis; ut—uterus; vd—vitelloduct; vi—vitellarium; vid—vitelline duct; vs—seminal vesicle. Scale bars: A–C, E = 100 μm; D=20 μm." figureDoi="http://doi.org/10.5281/zenodo.10941806" httpUri="https://zenodo.org/record/10941806/files/figure.png" pageId="2" pageNumber="149">Figs 1A–B</figureCitation>
). Brain located anterior to pharynx. Cerebral ganglion butterfly-shaped, consisting of irregular peripheral glial cells and internal nerve fibers (
<figureCitation id="C1DA2A47FF9D2723FDF34710B06DE59F" box="[519,605,1387,1412]" captionStart="Figure 1" captionStartId="3.[155,218,1895,1917]" captionTargetBox="[189,1399,499,1857]" captionTargetId="figure-198@3.[189,1399,499,1857]" captionTargetPageId="3" captionText="Figure 1. Phaenocora shenda Huang &amp; Wang, sp. nov. A. Free-living individual with zoochlorellae, dorsal view. B. Free-living individual without zoochlorellae, dorsal view. C. An individual treated with histochemical localization of acetylcholinesterase. D–E. Photomicrographs of histological sections stained with hematoxylin and eosin. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso-intestinal duct; eg—egg; fgc—female genital canal; gp—genital pore; i—intestine; pe—penis; ph—pharynx; t—testis; ut—uterus; vd—vitelloduct; vi—vitellarium; vid—vitelline duct; vs—seminal vesicle. Scale bars: A–C, E = 100 μm; D=20 μm." figureDoi="http://doi.org/10.5281/zenodo.10941806" httpUri="https://zenodo.org/record/10941806/files/figure.png" pageId="2" pageNumber="149">Figs 1C</figureCitation>
, 
<figureCitation id="C1DA2A47FF9D2723FD9D4717B049E59F" box="[617,633,1388,1412]" captionStart="Figure 2" captionStartId="4.[155,218,1415,1437]" captionTargetBox="[189,1397,573,1369]" captionTargetId="figure-449@4.[189,1398,569,1370]" captionTargetPageId="4" captionText="Figure 2. Phaenocora shenda Huang &amp; Wang, sp. nov., reconstruction of the brain, pharynx and reproductive system. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso intestinal duct; fgc—female genital canal; gp—genital pore; i— intestine; m—mouth; ov—ovary; pe—penis; ph—pharynx; sga—superior genital atrium; ut—uterus; vd—vitelloduct; vde—vas deferens; vs—seminal vesicle. Scale bar=50μm." figureDoi="http://doi.org/10.5281/zenodo.10941808" httpUri="https://zenodo.org/record/10941808/files/figure.png" pageId="2" pageNumber="149">2</figureCitation>
). Digest system made up of a pharynx, a mouth and an intestine (
<figureCitation id="C1DA2A47FF9D2723FAC54710B797E59F" box="[1329,1447,1387,1412]" captionStart="Figure 1" captionStartId="3.[155,218,1895,1917]" captionTargetBox="[189,1399,499,1857]" captionTargetId="figure-198@3.[189,1399,499,1857]" captionTargetPageId="3" captionText="Figure 1. Phaenocora shenda Huang &amp; Wang, sp. nov. A. Free-living individual with zoochlorellae, dorsal view. B. Free-living individual without zoochlorellae, dorsal view. C. An individual treated with histochemical localization of acetylcholinesterase. D–E. Photomicrographs of histological sections stained with hematoxylin and eosin. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso-intestinal duct; eg—egg; fgc—female genital canal; gp—genital pore; i—intestine; pe—penis; ph—pharynx; t—testis; ut—uterus; vd—vitelloduct; vi—vitellarium; vid—vitelline duct; vs—seminal vesicle. Scale bars: A–C, E = 100 μm; D=20 μm." figureDoi="http://doi.org/10.5281/zenodo.10941806" httpUri="https://zenodo.org/record/10941806/files/figure.png" pageId="2" pageNumber="149">Figs 1D–E</figureCitation>
, 
<figureCitation id="C1DA2A47FF9D2723FF7C47F5B2A8E5BD" box="[136,152,1422,1446]" captionStart="Figure 2" captionStartId="4.[155,218,1415,1437]" captionTargetBox="[189,1397,573,1369]" captionTargetId="figure-449@4.[189,1398,569,1370]" captionTargetPageId="4" captionText="Figure 2. Phaenocora shenda Huang &amp; Wang, sp. nov., reconstruction of the brain, pharynx and reproductive system. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso intestinal duct; fgc—female genital canal; gp—genital pore; i— intestine; m—mouth; ov—ovary; pe—penis; ph—pharynx; sga—superior genital atrium; ut—uterus; vd—vitelloduct; vde—vas deferens; vs—seminal vesicle. Scale bar=50μm." figureDoi="http://doi.org/10.5281/zenodo.10941808" httpUri="https://zenodo.org/record/10941808/files/figure.png" pageId="2" pageNumber="149">2</figureCitation>
). Pharynx 613 ± 64μm (
<emphasis id="6B95EAD0FF9D2723FE6247F4B393E5BD" box="[406,419,1423,1446]" italics="true" pageId="2" pageNumber="149">n</emphasis>
= 4) in length and 405 ± 51 μm (
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= 4) in width, belongs to bulbosus 
<typeStatus id="865A8860FF9D2723FB9947F5B6AEE5BD" box="[1133,1182,1422,1446]" pageId="2" pageNumber="149">type</typeStatus>
. Mouth lies ventrally and can be closed by a sphincter (
<figureCitation id="C1DA2A47FF9D2723FE3647CAB00AE5D2" box="[450,570,1457,1481]" captionStart="Figure 1" captionStartId="3.[155,218,1895,1917]" captionTargetBox="[189,1399,499,1857]" captionTargetId="figure-198@3.[189,1399,499,1857]" captionTargetPageId="3" captionText="Figure 1. Phaenocora shenda Huang &amp; Wang, sp. nov. A. Free-living individual with zoochlorellae, dorsal view. B. Free-living individual without zoochlorellae, dorsal view. C. An individual treated with histochemical localization of acetylcholinesterase. D–E. Photomicrographs of histological sections stained with hematoxylin and eosin. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso-intestinal duct; eg—egg; fgc—female genital canal; gp—genital pore; i—intestine; pe—penis; ph—pharynx; t—testis; ut—uterus; vd—vitelloduct; vi—vitellarium; vid—vitelline duct; vs—seminal vesicle. Scale bars: A–C, E = 100 μm; D=20 μm." figureDoi="http://doi.org/10.5281/zenodo.10941806" httpUri="https://zenodo.org/record/10941806/files/figure.png" pageId="2" pageNumber="149">Figs 1A–C</figureCitation>
, 
<figureCitation id="C1DA2A47FF9D2723FDB147CAB065E5D2" box="[581,597,1457,1481]" captionStart="Figure 2" captionStartId="4.[155,218,1415,1437]" captionTargetBox="[189,1397,573,1369]" captionTargetId="figure-449@4.[189,1398,569,1370]" captionTargetPageId="4" captionText="Figure 2. Phaenocora shenda Huang &amp; Wang, sp. nov., reconstruction of the brain, pharynx and reproductive system. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso intestinal duct; fgc—female genital canal; gp—genital pore; i— intestine; m—mouth; ov—ovary; pe—penis; ph—pharynx; sga—superior genital atrium; ut—uterus; vd—vitelloduct; vde—vas deferens; vs—seminal vesicle. Scale bar=50μm." figureDoi="http://doi.org/10.5281/zenodo.10941808" httpUri="https://zenodo.org/record/10941808/files/figure.png" pageId="2" pageNumber="149">2</figureCitation>
).
</paragraph>
<paragraph id="595E36C2FF9D2723FF4947A8B2C0E6C5" blockId="2.[136,1459,971,2070]" pageId="2" pageNumber="149">
Simultaneously hermaphroditic. Single superior genital atrium and genital pore situated ventrally, while inferior genital atrium absent (
<figureCitation id="C1DA2A47FF9D2723FED8478DB341E615" box="[300,369,1526,1550]" captionStart="Figure 2" captionStartId="4.[155,218,1415,1437]" captionTargetBox="[189,1397,573,1369]" captionTargetId="figure-449@4.[189,1398,569,1370]" captionTargetPageId="4" captionText="Figure 2. Phaenocora shenda Huang &amp; Wang, sp. nov., reconstruction of the brain, pharynx and reproductive system. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso intestinal duct; fgc—female genital canal; gp—genital pore; i— intestine; m—mouth; ov—ovary; pe—penis; ph—pharynx; sga—superior genital atrium; ut—uterus; vd—vitelloduct; vde—vas deferens; vs—seminal vesicle. Scale bar=50μm." figureDoi="http://doi.org/10.5281/zenodo.10941808" httpUri="https://zenodo.org/record/10941808/files/figure.png" pageId="2" pageNumber="149">Fig. 2</figureCitation>
). Female reproductive system consists of a pair of vitellaria, an ovary, an intestine bursa, a burso-intestinal duct, a genito-bursal duct, a female genital canal and a uterus (
<figureCitation id="C1DA2A47FF9D2723FC4F4462B605E62A" box="[955,1077,1561,1585]" captionStart="Figure 1" captionStartId="3.[155,218,1895,1917]" captionTargetBox="[189,1399,499,1857]" captionTargetId="figure-198@3.[189,1399,499,1857]" captionTargetPageId="3" captionText="Figure 1. Phaenocora shenda Huang &amp; Wang, sp. nov. A. Free-living individual with zoochlorellae, dorsal view. B. Free-living individual without zoochlorellae, dorsal view. C. An individual treated with histochemical localization of acetylcholinesterase. D–E. Photomicrographs of histological sections stained with hematoxylin and eosin. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso-intestinal duct; eg—egg; fgc—female genital canal; gp—genital pore; i—intestine; pe—penis; ph—pharynx; t—testis; ut—uterus; vd—vitelloduct; vi—vitellarium; vid—vitelline duct; vs—seminal vesicle. Scale bars: A–C, E = 100 μm; D=20 μm." figureDoi="http://doi.org/10.5281/zenodo.10941806" httpUri="https://zenodo.org/record/10941806/files/figure.png" pageId="2" pageNumber="149">Figs 1C–E</figureCitation>
, 
<figureCitation id="C1DA2A47FF9D2723FBB74462B661E62A" box="[1091,1105,1561,1585]" captionStart="Figure 2" captionStartId="4.[155,218,1415,1437]" captionTargetBox="[189,1397,573,1369]" captionTargetId="figure-449@4.[189,1398,569,1370]" captionTargetPageId="4" captionText="Figure 2. Phaenocora shenda Huang &amp; Wang, sp. nov., reconstruction of the brain, pharynx and reproductive system. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso intestinal duct; fgc—female genital canal; gp—genital pore; i— intestine; m—mouth; ov—ovary; pe—penis; ph—pharynx; sga—superior genital atrium; ut—uterus; vd—vitelloduct; vde—vas deferens; vs—seminal vesicle. Scale bar=50μm." figureDoi="http://doi.org/10.5281/zenodo.10941808" httpUri="https://zenodo.org/record/10941808/files/figure.png" pageId="2" pageNumber="149">2</figureCitation>
). Elliptic-shaped ovary located posterior to pharynx and opens dorsally to genito-bursal duct (
<figureCitation id="C1DA2A47FF9D2723FCDF4440B15EE64F" box="[811,878,1595,1620]" captionStart="Figure 2" captionStartId="4.[155,218,1415,1437]" captionTargetBox="[189,1397,573,1369]" captionTargetId="figure-449@4.[189,1398,569,1370]" captionTargetPageId="4" captionText="Figure 2. Phaenocora shenda Huang &amp; Wang, sp. nov., reconstruction of the brain, pharynx and reproductive system. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso intestinal duct; fgc—female genital canal; gp—genital pore; i— intestine; m—mouth; ov—ovary; pe—penis; ph—pharynx; sga—superior genital atrium; ut—uterus; vd—vitelloduct; vde—vas deferens; vs—seminal vesicle. Scale bar=50μm." figureDoi="http://doi.org/10.5281/zenodo.10941808" httpUri="https://zenodo.org/record/10941808/files/figure.png" pageId="2" pageNumber="149">Fig. 2</figureCitation>
). Genito-bursal duct extends towards intestine bursa, connecting ventrally to female genital canal. Intestine bursa expands and opens towards intestine via burso-intestinal duct (
<figureCitation id="C1DA2A47FF9D2723FF6444FAB337E682" box="[144,263,1665,1689]" captionStart="Figure 1" captionStartId="3.[155,218,1895,1917]" captionTargetBox="[189,1399,499,1857]" captionTargetId="figure-198@3.[189,1399,499,1857]" captionTargetPageId="3" captionText="Figure 1. Phaenocora shenda Huang &amp; Wang, sp. nov. A. Free-living individual with zoochlorellae, dorsal view. B. Free-living individual without zoochlorellae, dorsal view. C. An individual treated with histochemical localization of acetylcholinesterase. D–E. Photomicrographs of histological sections stained with hematoxylin and eosin. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso-intestinal duct; eg—egg; fgc—female genital canal; gp—genital pore; i—intestine; pe—penis; ph—pharynx; t—testis; ut—uterus; vd—vitelloduct; vi—vitellarium; vid—vitelline duct; vs—seminal vesicle. Scale bars: A–C, E = 100 μm; D=20 μm." figureDoi="http://doi.org/10.5281/zenodo.10941806" httpUri="https://zenodo.org/record/10941806/files/figure.png" pageId="2" pageNumber="149">Figs 1D–E</figureCitation>
, 
<figureCitation id="C1DA2A47FF9D2723FEE644FAB312E682" box="[274,290,1665,1689]" captionStart="Figure 2" captionStartId="4.[155,218,1415,1437]" captionTargetBox="[189,1397,573,1369]" captionTargetId="figure-449@4.[189,1398,569,1370]" captionTargetPageId="4" captionText="Figure 2. Phaenocora shenda Huang &amp; Wang, sp. nov., reconstruction of the brain, pharynx and reproductive system. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso intestinal duct; fgc—female genital canal; gp—genital pore; i— intestine; m—mouth; ov—ovary; pe—penis; ph—pharynx; sga—superior genital atrium; ut—uterus; vd—vitelloduct; vde—vas deferens; vs—seminal vesicle. Scale bar=50μm." figureDoi="http://doi.org/10.5281/zenodo.10941808" httpUri="https://zenodo.org/record/10941808/files/figure.png" pageId="2" pageNumber="149">2</figureCitation>
). Female genital canal with a nucleated epithelium (
<figureCitation id="C1DA2A47FF9D2723FCA744FAB199E682" box="[851,937,1665,1689]" captionStart="Figure 1" captionStartId="3.[155,218,1895,1917]" captionTargetBox="[189,1399,499,1857]" captionTargetId="figure-198@3.[189,1399,499,1857]" captionTargetPageId="3" captionText="Figure 1. Phaenocora shenda Huang &amp; Wang, sp. nov. A. Free-living individual with zoochlorellae, dorsal view. B. Free-living individual without zoochlorellae, dorsal view. C. An individual treated with histochemical localization of acetylcholinesterase. D–E. Photomicrographs of histological sections stained with hematoxylin and eosin. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso-intestinal duct; eg—egg; fgc—female genital canal; gp—genital pore; i—intestine; pe—penis; ph—pharynx; t—testis; ut—uterus; vd—vitelloduct; vi—vitellarium; vid—vitelline duct; vs—seminal vesicle. Scale bars: A–C, E = 100 μm; D=20 μm." figureDoi="http://doi.org/10.5281/zenodo.10941806" httpUri="https://zenodo.org/record/10941806/files/figure.png" pageId="2" pageNumber="149">Figs 1E</figureCitation>
, 
<figureCitation id="C1DA2A47FF9D2723FC4144FAB1F5E682" box="[949,965,1665,1689]" captionStart="Figure 2" captionStartId="4.[155,218,1415,1437]" captionTargetBox="[189,1397,573,1369]" captionTargetId="figure-449@4.[189,1398,569,1370]" captionTargetPageId="4" captionText="Figure 2. Phaenocora shenda Huang &amp; Wang, sp. nov., reconstruction of the brain, pharynx and reproductive system. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso intestinal duct; fgc—female genital canal; gp—genital pore; i— intestine; m—mouth; ov—ovary; pe—penis; ph—pharynx; sga—superior genital atrium; ut—uterus; vd—vitelloduct; vde—vas deferens; vs—seminal vesicle. Scale bar=50μm." figureDoi="http://doi.org/10.5281/zenodo.10941808" httpUri="https://zenodo.org/record/10941808/files/figure.png" pageId="2" pageNumber="149">2</figureCitation>
). Paired vitellaria extend on both sides of the body in ramified form, while first half of vitellaria converge as a common vitelloduct before entering female genital canal (
<figureCitation id="C1DA2A47FF9D2723FF6444BDB2D3E6C5" box="[144,227,1734,1758]" captionStart="Figure 1" captionStartId="3.[155,218,1895,1917]" captionTargetBox="[189,1399,499,1857]" captionTargetId="figure-198@3.[189,1399,499,1857]" captionTargetPageId="3" captionText="Figure 1. Phaenocora shenda Huang &amp; Wang, sp. nov. A. Free-living individual with zoochlorellae, dorsal view. B. Free-living individual without zoochlorellae, dorsal view. C. An individual treated with histochemical localization of acetylcholinesterase. D–E. Photomicrographs of histological sections stained with hematoxylin and eosin. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso-intestinal duct; eg—egg; fgc—female genital canal; gp—genital pore; i—intestine; pe—penis; ph—pharynx; t—testis; ut—uterus; vd—vitelloduct; vi—vitellarium; vid—vitelline duct; vs—seminal vesicle. Scale bars: A–C, E = 100 μm; D=20 μm." figureDoi="http://doi.org/10.5281/zenodo.10941806" httpUri="https://zenodo.org/record/10941806/files/figure.png" pageId="2" pageNumber="149">Fig. 1C</figureCitation>
).
</paragraph>
<paragraph id="595E36C2FF9D2723FF494492B14EE75D" blockId="2.[136,1459,971,2070]" pageId="2" pageNumber="149">
Pair testes lie dorsally or embed in vitellaria, extending on both sides of body (
<figureCitation id="C1DA2A47FF9D2723FBFE4492B66DE71A" box="[1034,1117,1769,1793]" captionStart="Figure 1" captionStartId="3.[155,218,1895,1917]" captionTargetBox="[189,1399,499,1857]" captionTargetId="figure-198@3.[189,1399,499,1857]" captionTargetPageId="3" captionText="Figure 1. Phaenocora shenda Huang &amp; Wang, sp. nov. A. Free-living individual with zoochlorellae, dorsal view. B. Free-living individual without zoochlorellae, dorsal view. C. An individual treated with histochemical localization of acetylcholinesterase. D–E. Photomicrographs of histological sections stained with hematoxylin and eosin. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso-intestinal duct; eg—egg; fgc—female genital canal; gp—genital pore; i—intestine; pe—penis; ph—pharynx; t—testis; ut—uterus; vd—vitelloduct; vi—vitellarium; vid—vitelline duct; vs—seminal vesicle. Scale bars: A–C, E = 100 μm; D=20 μm." figureDoi="http://doi.org/10.5281/zenodo.10941806" httpUri="https://zenodo.org/record/10941806/files/figure.png" pageId="2" pageNumber="149">Fig. 1C</figureCitation>
). Paired vas deferentia connect to a spherical-shaped seminal vesicle. Seminal vesicle with a thickened muscular wall, and connects to unarmed penis papilla. Ejaculatory duct opens towards superior genital atrium (
<figureCitation id="C1DA2A47FF9D2723FD2B4555B166E75D" box="[735,854,1838,1862]" captionStart="Figure 1" captionStartId="3.[155,218,1895,1917]" captionTargetBox="[189,1399,499,1857]" captionTargetId="figure-198@3.[189,1399,499,1857]" captionTargetPageId="3" captionText="Figure 1. Phaenocora shenda Huang &amp; Wang, sp. nov. A. Free-living individual with zoochlorellae, dorsal view. B. Free-living individual without zoochlorellae, dorsal view. C. An individual treated with histochemical localization of acetylcholinesterase. D–E. Photomicrographs of histological sections stained with hematoxylin and eosin. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso-intestinal duct; eg—egg; fgc—female genital canal; gp—genital pore; i—intestine; pe—penis; ph—pharynx; t—testis; ut—uterus; vd—vitelloduct; vi—vitellarium; vid—vitelline duct; vs—seminal vesicle. Scale bars: A–C, E = 100 μm; D=20 μm." figureDoi="http://doi.org/10.5281/zenodo.10941806" httpUri="https://zenodo.org/record/10941806/files/figure.png" pageId="2" pageNumber="149">Figs 1D–E</figureCitation>
, 
<figureCitation id="C1DA2A47FF9D2723FC954555B141E75D" box="[865,881,1838,1862]" captionStart="Figure 2" captionStartId="4.[155,218,1415,1437]" captionTargetBox="[189,1397,573,1369]" captionTargetId="figure-449@4.[189,1398,569,1370]" captionTargetPageId="4" captionText="Figure 2. Phaenocora shenda Huang &amp; Wang, sp. nov., reconstruction of the brain, pharynx and reproductive system. Abbreviations: b—brain; bi—intestine bursa; dgb—genito-bursal duct; dgi—burso intestinal duct; fgc—female genital canal; gp—genital pore; i— intestine; m—mouth; ov—ovary; pe—penis; ph—pharynx; sga—superior genital atrium; ut—uterus; vd—vitelloduct; vde—vas deferens; vs—seminal vesicle. Scale bar=50μm." figureDoi="http://doi.org/10.5281/zenodo.10941808" httpUri="https://zenodo.org/record/10941808/files/figure.png" pageId="2" pageNumber="149">2</figureCitation>
).
</paragraph>
<paragraph id="595E36C2FF9D2723FF49452AB1A0E7CA" blockId="2.[136,1459,971,2070]" pageId="2" pageNumber="149">
Habitat. The worms live in an environment where 
<emphasis id="6B95EAD0FF9D2723FD07452AB1B1E772" box="[755,897,1873,1897]" italics="true" pageId="2" pageNumber="149">Chladophora</emphasis>
sp. is abundant. The free-living individuals move slowly by ciliary movement. They respond to light slowly. When feeding with 
<taxonomicName id="9EE14D41FF9D2723FBF6450FB693E790" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1026,1187,1907,1931]" class="Branchiopoda" family="Artemiidae" genus="Artemia" kingdom="Animalia" order="Anostraca" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="salma">
<emphasis id="6B95EAD0FF9D2723FBF6450FB693E790" box="[1026,1187,1907,1931]" italics="true" pageId="2" pageNumber="149">Artemia salma</emphasis>
</taxonomicName>
under artificial culture, individuals of 
<taxonomicName id="9EE14D41FF9D2723FED545ECB004E7B5" authority="Huang &amp; Wang" authorityName="Huang &amp; Wang" authorityYear="2023" box="[289,564,1942,1966]" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="2" pageNumber="149" phylum="Platyhelminthes" rank="species" species="shenda" status="sp. nov.">
<emphasis id="6B95EAD0FF9D2723FED545ECB3B7E7B5" box="[289,391,1942,1966]" italics="true" pageId="2" pageNumber="149">P. shenda</emphasis>
Huang &amp; Wang
</taxonomicName>
, 
<emphasis id="6B95EAD0FF9D2723FDCA45EDB0A3E7B5" bold="true" box="[574,659,1942,1966]" pageId="2" pageNumber="149">
<taxonomicNameLabel id="70A657ABFF9D2723FDCA45EDB0A3E7B5" box="[574,659,1942,1966]" pageId="2" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
usually swim towards the prey. Once encountering the prey, it will quickly wrap the food, stretch out the throat, and swallow the prey into the body.
</paragraph>
<paragraph id="595E36C2FF9D2723FF4945A0B1D6E7E8" blockId="2.[136,1459,971,2070]" box="[189,998,2011,2036]" pageId="2" pageNumber="149">
Distribution. 
<collectingCountry id="21F67652FF9D2723FEB845A0B3BCE7E8" box="[332,396,2011,2035]" name="China" pageId="2" pageNumber="149">China</collectingCountry>
(
<collectingRegion id="9B25F820FF9D2723FE6D45A0B02CE7E8" box="[409,540,2011,2035]" country="China" name="Guangdong" pageId="2" pageNumber="149">Guangdong</collectingRegion>
). So far only known from the 
<typeStatus id="865A8860FF9D2723FCA845A7B1BAE7EF" box="[860,906,2012,2036]" pageId="2" pageNumber="149">type</typeStatus>
locality.
</paragraph>
<paragraph id="595E36C2FF9D2723FF494585B79CE80D" blockId="2.[136,1459,971,2070]" box="[189,1452,2046,2070]" pageId="2" pageNumber="149">Remarks. The male copulatory organ of the new species is characterized by the presence of an unarmed penis papilla.</paragraph>
<paragraph id="595E36C2FF9C2722FF7C4297B2FEE151" blockId="3.[136,1452,236,434]" pageId="3" pageNumber="150">
The male genital system belongs to Duplex-type IIIA. The female copulatory organ is characterized by the presence of both genito-bursal duct and burso-intestinal duct. The female genital system belongs to AGLOBULATA-type (
<bibRefCitation id="3D704B33FF9C2722FAF94374B2F2E151" author="Houben" pageId="3" pageNumber="150" pagination="301 - 354" refId="ref6656" refString="Houben, A. M., van Steenkiste, N., Artois, T. J. 2014. Revision of Phaenocora Ehrenberg, 1836 (Rhabditophora, Typhloplanidae, Phaenocorinae) with the description of two new species. Zootaxa, 3889 (3): 301 - 354. doi: 10.11646 / zootaxa. 3889.3.1" type="journal article" year="2014">
Houben 
<emphasis id="6B95EAD0FF9C2722FA99436BB795E13C" box="[1389,1445,271,295]" italics="true" pageId="3" pageNumber="150">et al.</emphasis>
, 2014
</bibRefCitation>
).
</paragraph>
<paragraph id="595E36C2FF9C2725FF49432EB72EE1EC" blockId="3.[136,1452,236,434]" lastBlockId="4.[136,1458,236,504]" lastPageId="4" lastPageNumber="151" pageId="3" pageNumber="150">
A morphological comparison of the copulatory organ of the new species and three other similar species within the genus is shown in Table 3. In 
<taxonomicName id="9EE14D41FF9C2722FE7A4303B097E194" authority="Ruebush, 1939" authorityName="Ruebush" authorityYear="1939" box="[398,679,375,399]" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="150" phylum="Platyhelminthes" rank="species" species="alticola">
<emphasis id="6B95EAD0FF9C2722FE7A4303B3CDE194" box="[398,509,375,399]" italics="true" pageId="3" pageNumber="150">P. alticola</emphasis>
Ruebush, 1939
</taxonomicName>
and 
<taxonomicName id="9EE14D41FF9C2722FD144303B1D8E194" authority="Marcus, 1959" authorityName="Marcus" authorityYear="1959" box="[736,1000,375,399]" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="150" phylum="Platyhelminthes" rank="species" species="brincki">
<emphasis id="6B95EAD0FF9C2722FD144303B178E194" box="[736,840,375,399]" italics="true" pageId="3" pageNumber="150">P. brincki</emphasis>
<bibRefCitation id="3D704B33FF9C2722FCA44303B1D8E194" author="Marcus" box="[848,1000,375,399]" pageId="3" pageNumber="150" pagination="15 - 42" refId="ref7349" refString="Marcus, E., Marcus, E. 1959. Turbellaria from Madeira and the Azores. Boletim do Museu Municipal do Funchal, 12: 15 - 42." type="journal article" year="1959">Marcus, 1959</bibRefCitation>
</taxonomicName>
, the male and female genital systems are similar to those of 
<taxonomicName id="9EE14D41FF9C2722FEA743E0B05AE1A9" authority="Huang &amp; Wang" authorityName="Huang &amp; Wang" authorityYear="2023" box="[339,618,410,434]" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="150" phylum="Platyhelminthes" rank="species" species="shenda" status="sp. nov.">
<emphasis id="6B95EAD0FF9C2722FEA743E0B389E1A9" box="[339,441,410,434]" italics="true" pageId="3" pageNumber="150">P. shenda</emphasis>
Huang &amp; Wang
</taxonomicName>
, 
<emphasis id="6B95EAD0FF9C2722FD8143E1B0FBE1A9" bold="true" box="[629,715,410,434]" pageId="3" pageNumber="150">
<taxonomicNameLabel id="70A657ABFF9C2722FD8143E1B0FBE1A9" box="[629,715,410,434]" pageId="3" pageNumber="150" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
The body length of 
<taxonomicName id="9EE14D41FF9C2722FC5343E0B623E1A9" authorityName="Ruebush" authorityYear="1939" box="[935,1043,410,434]" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="150" phylum="Platyhelminthes" rank="species" species="alticola">
<emphasis id="6B95EAD0FF9C2722FC5343E0B623E1A9" box="[935,1043,410,434]" italics="true" pageId="3" pageNumber="150">P. alticola</emphasis>
</taxonomicName>
(
<quantity id="9E199B27FF9C2722FBD643E1B692E1A9" box="[1058,1186,410,434]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.8" metricValueMin="1.2" pageId="3" pageNumber="150" unit="mm" value="1.5" valueMax="1.8" valueMin="1.2">1.2–1.8 mm</quantity>
) and 
<taxonomicName id="9EE14D41FF9C2722FB2A43E0B774E1A9" authorityName="Marcus" authorityYear="1959" box="[1246,1348,410,434]" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="150" phylum="Platyhelminthes" rank="species" species="brincki">
<emphasis id="6B95EAD0FF9C2722FB2A43E0B774E1A9" box="[1246,1348,410,434]" italics="true" pageId="3" pageNumber="150">P. brincki</emphasis>
</taxonomicName>
(
<quantity id="9E199B27FF9C2722FAA743E1B794E1A9" box="[1363,1444,410,434]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="150" unit="mm" value="2.2">2.2 mm</quantity>
) is much shorter than that of 
<taxonomicName id="9EE14D41FF9B2725FE464296B0F7E11E" authority="Huang &amp; Wang" authorityName="Huang &amp; Wang" authorityYear="2023" box="[434,711,236,261]" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="4" pageNumber="151" phylum="Platyhelminthes" rank="species" species="shenda" status="sp. nov.">
<emphasis id="6B95EAD0FF9B2725FE464296B028E11F" box="[434,536,236,260]" italics="true" pageId="4" pageNumber="151">P. shenda</emphasis>
Huang &amp; Wang
</taxonomicName>
, 
<emphasis id="6B95EAD0FF9B2725FD244296B116E11E" bold="true" box="[720,806,237,261]" pageId="4" pageNumber="151">
<taxonomicNameLabel id="70A657ABFF9B2725FD244296B116E11E" box="[720,806,237,261]" pageId="4" pageNumber="151" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(3.05 ± 
<quantity id="9E199B27FF9B2725FC8C4296B1E8E11E" box="[888,984,237,261]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="4" pageNumber="151" unit="mm" value="0.18">0.18 mm</quantity>
). Moreover, in 
<taxonomicName id="9EE14D41FF9B2725FB8E4296B6D6E11F" authorityName="Ruebush" authorityYear="1939" box="[1146,1254,236,260]" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="4" pageNumber="151" phylum="Platyhelminthes" rank="species" species="alticola">
<emphasis id="6B95EAD0FF9B2725FB8E4296B6D6E11F" box="[1146,1254,236,260]" italics="true" pageId="4" pageNumber="151">P. alticola</emphasis>
</taxonomicName>
and 
<taxonomicName id="9EE14D41FF9B2725FAEC4296B74EE11F" authorityName="Marcus" authorityYear="1959" box="[1304,1406,236,260]" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="4" pageNumber="151" phylum="Platyhelminthes" rank="species" species="brincki">
<emphasis id="6B95EAD0FF9B2725FAEC4296B74EE11F" box="[1304,1406,236,260]" italics="true" pageId="4" pageNumber="151">P. brincki</emphasis>
</taxonomicName>
, the seminal vesicle is connected to the prostate glands, while the prostate glands converge to form the prostate vesicle. In contrast, prostate glands are absent in 
<taxonomicName id="9EE14D41FF9B2725FE204348B0CBE151" authority="Huang &amp; Wang" authorityName="Huang &amp; Wang" authorityYear="2023" box="[468,763,306,330]" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="4" pageNumber="151" phylum="Platyhelminthes" rank="species" species="shenda" status="sp. nov.">
<emphasis id="6B95EAD0FF9B2725FE204348B00FE151" box="[468,575,306,330]" italics="true" pageId="4" pageNumber="151">P. shenda</emphasis>
Huang &amp; Wang
</taxonomicName>
, 
<emphasis id="6B95EAD0FF9B2725FCFF4349B155E151" bold="true" box="[779,869,306,330]" pageId="4" pageNumber="151">
<taxonomicNameLabel id="70A657ABFF9B2725FCFF4349B155E151" box="[779,869,306,330]" pageId="4" pageNumber="151" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
By comparing the vas deferentia, and overall body configuration and structure, the new species is most similar to 
<taxonomicName id="9EE14D41FF9B2725FCD9432EB60FE177" authority="Marcus, 1946" authorityName="Marcus" authorityYear="1946" box="[813,1087,340,364]" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="4" pageNumber="151" phylum="Platyhelminthes" rank="species" species="evelinae">
<emphasis id="6B95EAD0FF9B2725FCD9432EB191E177" box="[813,929,340,364]" italics="true" pageId="4" pageNumber="151">P. evelinae</emphasis>
<bibRefCitation id="3D704B33FF9B2725FC5C432EB60FE177" author="Marcus" box="[936,1087,340,364]" pageId="4" pageNumber="151" pagination="5 - 254" refId="ref7313" refString="Marcus, E. 1946. Sobre Turbellaria limnieos Brasileiros. Boletim / Faculdade de Filosofia, Letras e Ciencias. Humanas da Universidade de Sao Paulo, Zoologia, 11: 5 - 254." type="journal article" year="1946">Marcus, 1946</bibRefCitation>
</taxonomicName>
. In 
<taxonomicName id="9EE14D41FF9B2725FB9C432EB6ECE177" authorityName="Marcus" authorityYear="1946" box="[1128,1244,340,364]" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="4" pageNumber="151" phylum="Platyhelminthes" rank="species" species="evelinae">
<emphasis id="6B95EAD0FF9B2725FB9C432EB6ECE177" box="[1128,1244,340,364]" italics="true" pageId="4" pageNumber="151">P. evelinae</emphasis>
</taxonomicName>
, each of the paired vas deferentia connects to a seminal vesicle respectively, which is the same as 
<taxonomicName id="9EE14D41FF9B2725FC3E4303B6EEE18B" authority="Huang &amp; Wang" authorityName="Huang &amp; Wang" authorityYear="2023" box="[970,1246,375,400]" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="4" pageNumber="151" phylum="Platyhelminthes" rank="species" species="shenda" status="sp. nov.">
<emphasis id="6B95EAD0FF9B2725FC3E4303B61FE194" box="[970,1071,375,399]" italics="true" pageId="4" pageNumber="151">P. shenda</emphasis>
Huang &amp; Wang
</taxonomicName>
, 
<emphasis id="6B95EAD0FF9B2725FB1D430CB70EE194" bold="true" box="[1257,1342,375,399]" pageId="4" pageNumber="151">
<taxonomicNameLabel id="70A657ABFF9B2725FB1D430CB70EE194" box="[1257,1342,375,399]" pageId="4" pageNumber="151" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
However, the female genital system of 
<taxonomicName id="9EE14D41FF9B2725FE4043E0B015E1A9" authorityName="Marcus" authorityYear="1946" box="[436,549,410,434]" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="4" pageNumber="151" phylum="Platyhelminthes" rank="species" species="evelinae">
<emphasis id="6B95EAD0FF9B2725FE4043E0B015E1A9" box="[436,549,410,434]" italics="true" pageId="4" pageNumber="151">P. evelinae</emphasis>
</taxonomicName>
is EVELINAE-type, with the presence of burso-intestinal duct, but absence of genito-bursal duct, whilst they are both present in 
<taxonomicName id="9EE14D41FF9B2725FDBE43C6B16AE1CE" authority="Huang &amp; Wang" authorityName="Huang &amp; Wang" authorityYear="2023" box="[586,858,444,469]" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="4" pageNumber="151" phylum="Platyhelminthes" rank="species" species="shenda" status="sp. nov.">
<emphasis id="6B95EAD0FF9B2725FDBE43C6B09EE1CF" box="[586,686,444,468]" italics="true" pageId="4" pageNumber="151">P. shenda</emphasis>
Huang &amp; Wang
</taxonomicName>
, 
<emphasis id="6B95EAD0FF9B2725FC9043C6B188E1CE" bold="true" box="[868,952,445,469]" pageId="4" pageNumber="151">
<taxonomicNameLabel id="70A657ABFF9B2725FC9043C6B188E1CE" box="[868,952,445,469]" pageId="4" pageNumber="151" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<bibRefCitation id="3D704B33FF9B2725FC3143C7B6AAE1CE" author="Houben" box="[965,1178,444,469]" pageId="4" pageNumber="151" pagination="301 - 354" refId="ref6656" refString="Houben, A. M., van Steenkiste, N., Artois, T. J. 2014. Revision of Phaenocora Ehrenberg, 1836 (Rhabditophora, Typhloplanidae, Phaenocorinae) with the description of two new species. Zootaxa, 3889 (3): 301 - 354. doi: 10.11646 / zootaxa. 3889.3.1" type="journal article" year="2014">
Houben 
<emphasis id="6B95EAD0FF9B2725FBD443C6B665E1CF" box="[1056,1109,444,468]" italics="true" pageId="4" pageNumber="151">et al.</emphasis>
, 2014
</bibRefCitation>
). Taking these differences into account, we described 
<taxonomicName id="9EE14D41FF9B2725FE5F439BB0F1E1E3" authority="Huang &amp; Wang" authorityName="Huang &amp; Wang" authorityYear="2023" box="[427,705,479,504]" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="4" pageNumber="151" phylum="Platyhelminthes" rank="species" species="shenda" status="sp. nov.">
<emphasis id="6B95EAD0FF9B2725FE5F439BB021E1EC" box="[427,529,479,503]" italics="true" pageId="4" pageNumber="151">P.shenda</emphasis>
Huang &amp; Wang
</taxonomicName>
, 
<emphasis id="6B95EAD0FF9B2725FD3F43A4B111E1EC" bold="true" box="[715,801,479,503]" pageId="4" pageNumber="151">
<taxonomicNameLabel id="70A657ABFF9B2725FD3F43A4B111E1EC" box="[715,801,479,503]" pageId="4" pageNumber="151" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
as a new species within the genus 
<taxonomicName id="9EE14D41FF9B2725FB6043A4B726E1EC" authorityName="Ehrenberg" authorityYear="1837" box="[1172,1302,479,503]" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="4" pageNumber="151" phylum="Platyhelminthes" rank="genus">
<emphasis id="6B95EAD0FF9B2725FB6043A4B726E1EC" box="[1172,1302,479,503]" italics="true" pageId="4" pageNumber="151">Phaenocora</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>