treatments-xml/data/B8/28/19/B8281932FFA4FFBAFF96FA25FD143E70.xml

<document id="886D81C68157ED81EEFB414C6EDBC38B" ID-DOI="10.11646/zootaxa.5353.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8426702" ID-ZooBank="F6738CF9-1A17-4227-9B38-0562FA67DAB9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1696942587025" checkinUser="plazi" docAuthor="Lo, Ying-Yuan, Wei, Chi &amp; Cheng, Ren-Chung" docDate="2023" docId="B8281932FFA4FFBAFF96FA25FD143E70" docLanguage="en" docName="zootaxa.5353.1.2.pdf" docOrigin="Zootaxa 5353 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5353.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hogna arborea Lo &amp; Wei &amp; Cheng 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="54" masterDocId="4411614AFFA0FFBDFF01FF86FFAE3D49" masterDocTitle="Tree-dwelling wolves: a new arboreal Hogna species (Araneae: Lycosidae) from Taiwan" masterLastPageNumber="59" masterPageNumber="47" pageNumber="51" updateTime="1696943054205" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B31F37099AE44ED7FD782939CF430E57">Tree-dwelling wolves: a new arboreal Hogna species (Araneae: Lycosidae) from Taiwan</mods:title>
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<mods:namePart id="D4A5455C9DF5603D2B02DE5464E3567D">Lo, Ying-Yuan</mods:namePart>
<mods:nameIdentifier id="F8C8D41A95A9ACB70B378B82D66CC053" type="ORCID">0000-0002-2862-5155</mods:nameIdentifier>
<mods:affiliation id="3AD21FCAC0F8D4C14CC2729932CE70B9">Department of Life Science, National Taiwan Normal University, No. 88, Sec. 4, Tingzhou Rd., Taipei 11677, Taiwan yylo. robert @ gamil. com; https: // orcid. org / 0000 - 0002 - 2862 - 5155 &amp; Zoology Division, Taiwan Endemic Species Research Institute, No. 1, Minsheng E. Rd., Jiji Township, Nantou 55244, Taiwan Department of Biological Sciences, Old Dominion University, Norfolk, Virginia 23529, USA</mods:affiliation>
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<mods:namePart id="48A961AA2F29E1568DE12AAE8D9F7A6F">Wei, Chi</mods:namePart>
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<mods:namePart id="052CCD26B925943BD8BCF205AC369F9D">Cheng, Ren-Chung</mods:namePart>
<mods:affiliation id="0D6EBB46E696AD4182DEB33B47C33CB7">Department of Life Sciences, National Chung Hsing University, No. 145, Xingda Rd., South Dist., Taichung 40227, Taiwan Research Center for Global Change Biology, National Chung Hsing University, No. 145, Xingda Rd., South Dist., Taichung 40227, Taiwan</mods:affiliation>
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<treatment id="B8281932FFA4FFBAFF96FA25FD143E70" ID-DOI="http://doi.org/10.5281/zenodo.8426722" ID-Zenodo-Dep="8426722" LSID="urn:lsid:plazi:treatment:B8281932FFA4FFBAFF96FA25FD143E70" httpUri="http://treatment.plazi.org/id/B8281932FFA4FFBAFF96FA25FD143E70" lastPageId="7" lastPageNumber="54" pageId="4" pageNumber="51">
<subSubSection id="789BFBAFFFA4FFB9FF96FA25FE1B38F4" box="[151,437,1442,1469]" pageId="4" pageNumber="51" type="nomenclature">
<paragraph id="303EA824FFA4FFB9FF96FA25FE1B38F4" blockId="4.[151,437,1442,1506]" box="[151,437,1442,1469]" pageId="4" pageNumber="51">
<heading id="6B761F48FFA4FFB9FF96FA25FE1B38F4" bold="true" box="[151,437,1442,1469]" fontSize="11" level="1" pageId="4" pageNumber="51" reason="1">
<emphasis id="02F57436FFA4FFB9FF96FA25FE1B38F4" bold="true" box="[151,437,1442,1469]" pageId="4" pageNumber="51">
<taxonomicName id="F781D3A7FFA4FFB9FF96FA25FEFF38F4" authority="Lo &amp; Wei &amp; Cheng, 2023" authorityName="Lo &amp; Wei &amp; Cheng" authorityYear="2023" box="[151,337,1442,1469]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="arborea" status="sp. nov.">
<emphasis id="02F57436FFA4FFB9FF96FA25FEFF38F4" bold="true" box="[151,337,1442,1469]" italics="true" pageId="4" pageNumber="51">Hogna arborea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C6C94DFFA4FFB9FE56FA25FE1B38F4" box="[343,437,1443,1469]" pageId="4" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="789BFBAFFFA4FFB9FF96FA41FF5638AB" box="[151,248,1479,1506]" pageId="4" pageNumber="51" type="description">
<paragraph id="303EA824FFA4FFB9FF96FA41FF5638AB" blockId="4.[151,437,1442,1506]" box="[151,248,1479,1506]" pageId="4" pageNumber="51">
<figureCitation id="A8BAB4A1FFA4FFB9FF96FA41FF5638AB" box="[151,248,1479,1506]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="5.[151,250,951,975]" captionStartId-1="5.[151,250,1827,1851]" captionStartId-2="6.[151,250,1449,1473]" captionTargetBox-0="[151,1432,276,925]" captionTargetBox-1="[158,1431,1146,1802]" captionTargetBox-2="[155,1435,705,1421]" captionTargetId-0="figure-71@5.[151,1435,273,927]" captionTargetId-1="figure-166@5.[151,1436,1143,1804]" captionTargetId-2="figure-277@6.[151,1435,705,1425]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 3 A–E. Epigyne and left palp organ of Hogna arborea sp. nov. (A–B, Holotype, TESRI- Ar3045; C–E, Paratype, TESRI-Ar5507). A, ventral view; B, dorsal view; C, prolateral view; D, ventral view; E, retrolateral view. Scale bar = 0.05mm. Abbreviations: BS- base of spermatheca, C- conductor, CD- copulatory duct, Em- embolus, H- hood, HS- head of spermatheca, MA- median apophysis, MF- massive fold, P- palea, S- septum, SB- septum base, St- subtegulum, T- tegulum, TA- terminal apophysis." captionText-1="FIGURE 4 A–E. Epigyne and left palp organ of Hogna arborea sp. nov. (A–B, Holotype, TESRI- Ar3045; C–E, Paratype, TESRI-Ar5507). A, ventral view; B, dorsal view; C, prolateral view; D, ventral view; E, retrolateral view. Scale bar = 0.05mm. Abbreviations: BS- base of spermatheca, C- conductor, CD- copulatory duct, Em- embolus, H- hood, HS- head of spermatheca, MA- median apophysis, MF- massive fold, P- palea, S- septum, SB- septum base, St- subtegulum, T- tegulum, TA- terminal apophysis." captionText-2="FIGURE 5 A–G. Habitus of Hogna arborea sp. nov. (A, C, E, G, Holotype, TESRI- Ar3045; B, D, F, Paratype, TESRI- Ar5507). A–B, general appearance, dorsal view; C–D, prosoma, lateral view; E–F, eyes; G, metatarsus I and tarsus I, prolateral view. Scale bar = 5mm (A–D); 0.5mm (E–F), 1mm (G)." figureDoi-0="http://doi.org/10.5281/zenodo.8426710" figureDoi-1="http://doi.org/10.5281/zenodo.8426712" figureDoi-2="http://doi.org/10.5281/zenodo.8426716" httpUri-0="https://zenodo.org/record/8426710/files/figure.png" httpUri-1="https://zenodo.org/record/8426712/files/figure.png" httpUri-2="https://zenodo.org/record/8426716/files/figure.png" pageId="4" pageNumber="51">Figs 3–5</figureCitation>
</paragraph>
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<emphasis id="02F57436FFA4FFB9FF96F989FEE53B60" bold="true" box="[151,331,1551,1577]" pageId="4" pageNumber="51">Type material.</emphasis>
<materialsCitation id="80E9A279FFA4FFB9FE56F989FE2A3B04" collectingDate="2018-12-04" collectorName="Ying-Yuan Lo" latitude="24.53" location="Yilan County" longLatPrecision="750" longitude="121.83" pageId="4" pageNumber="51" specimenCount="1" stateProvince="Yilan" typeStatus="holotype">
<emphasis id="02F57436FFA4FFB9FE56F989FE693B60" box="[343,455,1551,1577]" italics="true" pageId="4" pageNumber="51">
<typeStatus id="EF3A1686FFA4FFB9FE56F989FE6F3B60" box="[343,449,1551,1577]" pageId="4" pageNumber="51">Holotype</typeStatus>
:
</emphasis>
female (TESRI-Ar3045), 
<date id="443F8EE4FFA4FFB9FC01F989FC393B60" box="[768,919,1551,1577]" pageId="4" pageNumber="51" value="2018-12-04">
<collectingDate id="547B770CFFA4FFB9FC01F989FC393B60" box="[768,919,1551,1577]" pageId="4" pageNumber="51" value="2018-12-04">4 Dec. 2018</collectingDate>
</date>
, 
<collectorName id="9D74CDF2FFA4FFB9FCA7F989FBE53B60" box="[934,1099,1551,1577]" pageId="4" pageNumber="51">Ying-Yuan Lo</collectorName>
<emphasis id="02F57436FFA4FFB9FB59F989FB2F3B60" box="[1112,1153,1551,1577]" italics="true" pageId="4" pageNumber="51">leg.</emphasis>
, Dongao, 
<collectingRegion id="F24566C6FFA4FFB9FA01F989FA323B60" box="[1280,1436,1551,1577]" country="Taiwan" name="Yilan" pageId="4" pageNumber="51">Yilan County</collectingRegion>
(
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, 
<geoCoordinate id="55B5CEE3FFA4FFB9FE0FF9B5FED23B04" box="[270,380,1587,1613]" degrees="121.83" direction="east" orientation="longitude" pageId="4" pageNumber="51" precision="555" value="121.83">121.83°E</geoCoordinate>
)
</materialsCitation>
. 
<materialsCitation id="80E9A279FFA4FFB9FE8EF9B2FBB33B3B" collectingDate="2020-02-10" collectingDateMax="2020-08-24" collectingDateMin="2020-02-10" collectorName="Chi-Chun Liao" latitude="24.01" location="County" longLatPrecision="752" longitude="121.58" municipality="Saguer Trail" pageId="4" pageNumber="51" specimenCount="1" specimenCount-male="1" stateProvince="Hualien" typeStatus="paratype">
<emphasis id="02F57436FFA4FFB9FE8EF9B2FDA43B04" box="[399,522,1588,1613]" italics="true" pageId="4" pageNumber="51">
<typeStatus id="EF3A1686FFA4FFB9FE8EF9B2FDAA3B04" box="[399,516,1588,1613]" pageId="4" pageNumber="51">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="268763ADFFA4FFB9FD13F9B5FDD63B04" box="[530,632,1587,1613]" pageId="4" pageNumber="51" type="male">one male</specimenCount>
(TESRI-Ar5507), 
<date id="443F8EE4FFA4FFB9FC4EF9B5FC483B04" box="[847,998,1587,1613]" pageId="4" pageNumber="51" value="2020-02-10">
<collectingDate id="547B770CFFA4FFB9FC4EF9B5FC483B04" box="[847,998,1587,1613]" pageId="4" pageNumber="51" value="2020-02-10">10 Feb. 2020</collectingDate>
</date>
, 
<collectorName id="9D74CDF2FFA4FFB9FCF3F9B5FB323B04" box="[1010,1180,1587,1613]" pageId="4" pageNumber="51">Chi-Chun Liao</collectorName>
<emphasis id="02F57436FFA4FFB9FBA5F9B5FB633B04" box="[1188,1229,1587,1613]" italics="true" pageId="4" pageNumber="51">leg.</emphasis>
(reared in lab and matured on 
<date id="443F8EE4FFA4FFB9FE20F9D1FE6F3B38" box="[289,449,1623,1649]" pageId="4" pageNumber="51" value="2020-08-24">
<collectingDate id="547B770CFFA4FFB9FE20F9D1FE6F3B38" box="[289,449,1623,1649]" pageId="4" pageNumber="51" value="2020-08-24">24 Aug. 2020</collectingDate>
</date>
), 
<collectingMunicipality id="D05A325EFFA4FFB9FED7F9D1FDCA3B38" box="[470,612,1623,1649]" pageId="4" pageNumber="51">Saguer Trail</collectingMunicipality>
, 
<collectingRegion id="F24566C6FFA4FFB9FD73F9D1FD653B38" box="[626,715,1623,1649]" country="Taiwan" name="Hualien" pageId="4" pageNumber="51">Hualian</collectingRegion>
<location id="355EFEFFFFA4FFB9FDD5F9D1FC883B38" LSID="urn:lsid:plazi:treatment:B8281932FFA4FFBAFF96FA25FD143E70:355EFEFFFFA4FFB9FDD5F9D1FC883B38" box="[724,806,1623,1649]" latitude="24.01" longLatPrecision="752" longitude="121.58" municipality="Saguer Trail" name="County" pageId="4" pageNumber="51" stateProvince="Hualien">County</location>
(
<geoCoordinate id="55B5CEE3FFA4FFB9FC36F9D1FC373B38" box="[823,921,1623,1649]" degrees="24.01" direction="north" orientation="latitude" pageId="4" pageNumber="51" precision="555" value="24.01">24.01°N</geoCoordinate>
, 
<geoCoordinate id="55B5CEE3FFA4FFB9FCA6F9D1FBBB3B3B" box="[935,1045,1623,1650]" degrees="121.58" direction="east" orientation="longitude" pageId="4" pageNumber="51" precision="555" value="121.58">121.58°E</geoCoordinate>
)
</materialsCitation>
; 
<materialsCitation id="80E9A279FFA4FFB9FB2BF9D1FCEF3BDC" collectingDate="2019-12-23" collectorName="Chi-Chun Liao" latitude="24.53" location="Yilan County" longLatPrecision="750" longitude="121.83" pageId="4" pageNumber="51" specimenCount="1" specimenCount-female="1" stateProvince="Yilan" typeStatus="paratype">
<specimenCount id="268763ADFFA4FFB9FB2BF9D1FB063B38" box="[1066,1192,1623,1649]" pageId="4" pageNumber="51" type="female">one female</specimenCount>
(TESRI-Ar3549), 
<date id="443F8EE4FFA4FFB9FA81F9D1FEA23BDC" pageId="4" pageNumber="51" value="2019-12-23">
<collectingDate id="547B770CFFA4FFB9FA81F9D1FEA23BDC" pageId="4" pageNumber="51" value="2019-12-23">23 Dec. 2019</collectingDate>
</date>
, 
<collectorName id="9D74CDF2FFA4FFB9FE16F9FDFE103BDC" box="[279,446,1659,1685]" pageId="4" pageNumber="51">Chi-Chun Liao</collectorName>
<emphasis id="02F57436FFA4FFB9FEC4F9FDFE403BDC" box="[453,494,1659,1685]" italics="true" pageId="4" pageNumber="51">leg.</emphasis>
, same locality as the holotype
</materialsCitation>
; 
<materialsCitation id="80E9A279FFA4FFB9FC4AF9FDFCC93BF3" collectingDate="2016-03-26" collectorName="Hsu" latitude="22.8" location="Lijia Forest" longLatPrecision="755" longitude="121.05" pageId="4" pageNumber="51" specimenCount="1" specimenCount-female="1" stateProvince="Taitung" typeStatus="paratype">
<specimenCount id="268763ADFFA4FFB9FC4AF9FDFC683BDC" box="[843,966,1659,1685]" pageId="4" pageNumber="51" type="female">one female</specimenCount>
(TESRI-Ar5523), 
<date id="443F8EE4FFA4FFB9FB99F9FDFA9E3BDC" box="[1176,1328,1659,1685]" pageId="4" pageNumber="51" value="2016-03-26">
<collectingDate id="547B770CFFA4FFB9FB99F9FDFA9E3BDC" box="[1176,1328,1659,1685]" pageId="4" pageNumber="51" value="2016-03-26">26 Mar. 2016</collectingDate>
</date>
, Chen-Fu 
<collectorName id="9D74CDF2FFA4FFB9FF96F919FF6A3BF0" box="[151,196,1695,1721]" pageId="4" pageNumber="51">Hsu</collectorName>
<emphasis id="02F57436FFA4FFB9FFCBF919FF423BF0" box="[202,236,1695,1721]" italics="true" pageId="4" pageNumber="51">leg</emphasis>
., 
<location id="355EFEFFFFA4FFB9FE00F919FE2F3BF0" LSID="urn:lsid:plazi:treatment:B8281932FFA4FFBAFF96FA25FD143E70:355EFEFFFFA4FFB9FE00F919FE2F3BF0" box="[257,385,1695,1721]" latitude="22.8" longLatPrecision="755" longitude="121.05" name="Lijia Forest" pageId="4" pageNumber="51" stateProvince="Taitung">Lijia Forest</location>
Trail, 
<collectingRegion id="F24566C6FFA4FFB9FEC8F919FDD83BF0" box="[457,630,1695,1721]" country="Taiwan" name="Taitung" pageId="4" pageNumber="51">Taitung County</collectingRegion>
(
<geoCoordinate id="55B5CEE3FFA4FFB9FD85F919FD493BF0" box="[644,743,1695,1721]" degrees="22.80" direction="north" orientation="latitude" pageId="4" pageNumber="51" precision="555" value="22.8">22.80°N</geoCoordinate>
, 
<geoCoordinate id="55B5CEE3FFA4FFB9FDF0F919FCF13BF3" box="[753,863,1695,1722]" degrees="121.05" direction="east" orientation="longitude" pageId="4" pageNumber="51" precision="555" value="121.05">121.05°E</geoCoordinate>
)
</materialsCitation>
; 
<materialsCitation id="80E9A279FFA4FFB9FC73F919FBA73B94" collectingDate="2021-01-03" collectorName="Chang-Lin Chung" latitude="21.95" location="Kenting National Park" longLatPrecision="757" longitude="120.81" pageId="4" pageNumber="51" specimenCount="1" specimenCount-female="1" stateProvince="Pingtung" typeStatus="paratype">
<specimenCount id="268763ADFFA4FFB9FC73F919FC423BF0" box="[882,1004,1695,1721]" pageId="4" pageNumber="51" type="female">one female</specimenCount>
(TESRI-Ar5753), 
<date id="443F8EE4FFA4FFB9FBBFF919FAEE3BF0" box="[1214,1344,1695,1721]" pageId="4" pageNumber="51" value="2021-01-03">
<collectingDate id="547B770CFFA4FFB9FBBFF919FAEE3BF0" box="[1214,1344,1695,1721]" pageId="4" pageNumber="51" value="2021-01-03">3 Jan. 2021</collectingDate>
</date>
, 
<collectorName id="9D74CDF2FFA4FFB9FA4BF919FEBE3B94" pageId="4" pageNumber="51">Chang-Lin Chung</collectorName>
<emphasis id="02F57436FFA4FFB9FE16F945FE973B94" box="[279,313,1731,1757]" italics="true" pageId="4" pageNumber="51">leg</emphasis>
., 
<location id="355EFEFFFFA4FFB9FE4FF945FDE33B94" LSID="urn:lsid:plazi:treatment:B8281932FFA4FFBAFF96FA25FD143E70:355EFEFFFFA4FFB9FE4FF945FDE33B94" box="[334,589,1731,1757]" latitude="21.95" longLatPrecision="757" longitude="120.81" name="Kenting National Park" pageId="4" pageNumber="51" stateProvince="Pingtung">Kenting National Park</location>
, 
<collectingRegion id="F24566C6FFA4FFB9FD59F945FCB83B94" box="[600,790,1731,1757]" country="Taiwan" name="Pingtung" pageId="4" pageNumber="51">Pingtung County</collectingRegion>
(
<geoCoordinate id="55B5CEE3FFA4FFB9FC27F945FC263B97" box="[806,904,1731,1758]" degrees="21.95" direction="north" orientation="latitude" pageId="4" pageNumber="51" precision="555" value="21.95">21.95°N</geoCoordinate>
, 
<geoCoordinate id="55B5CEE3FFA4FFB9FC92F945FBAF3B94" box="[915,1025,1731,1757]" degrees="120.81" direction="east" orientation="longitude" pageId="4" pageNumber="51" precision="555" value="120.81">120.81°E</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
<paragraph id="303EA824FFA4FFB9FFC6F961FD203A6C" blockId="4.[151,1437,1551,2009]" pageId="4" pageNumber="51">
<emphasis id="02F57436FFA4FFB9FFC6F961FE5E3A48" bold="true" box="[199,496,1767,1793]" pageId="4" pageNumber="51">Other material examined</emphasis>
: 
<materialsCitation id="80E9A279FFA4FFB9FEFDF961FD263A6C" collectingDate="2001-10-10" collectorName="Ren-Jie Chen &amp; Kenting National" latitude="21.95" location="Kenting National Park" longLatPrecision="757" longitude="120.81" pageId="4" pageNumber="51" specimenCount="1" specimenCount-male="1" stateProvince="Pingtung">
<specimenCount id="268763ADFFA4FFB9FEFDF961FDC23A48" box="[508,620,1767,1793]" pageId="4" pageNumber="51" type="male">two males</specimenCount>
(TESRI-Ar3562/3563), 
<date id="443F8EE4FFA4FFB9FC7DF961FBA23A48" box="[892,1036,1767,1793]" pageId="4" pageNumber="51" value="2001-10-10">
<collectingDate id="547B770CFFA4FFB9FC7DF961FBA23A48" box="[892,1036,1767,1793]" pageId="4" pageNumber="51" value="2001-10-10">10 Oct. 2001</collectingDate>
</date>
, 
<collectorName id="9D74CDF2FFA4FFB9FB14F961FB043A48" box="[1045,1194,1767,1793]" pageId="4" pageNumber="51">Ren-Jie Chen</collectorName>
<emphasis id="02F57436FFA4FFB9FBAEF961FB763A48" box="[1199,1240,1767,1793]" italics="true" pageId="4" pageNumber="51">leg.</emphasis>
<location id="355EFEFFFFA4FFB9FBDCF961FF633A6C" LSID="urn:lsid:plazi:treatment:B8281932FFA4FFBAFF96FA25FD143E70:355EFEFFFFA4FFB9FBDCF961FF633A6C" latitude="21.95" longLatPrecision="757" longitude="120.81" name="Kenting National Park" pageId="4" pageNumber="51" stateProvince="Pingtung">
<collectorName id="9D74CDF2FFA4FFB9FBDCF961FA323A48" box="[1245,1436,1767,1793]" pageId="4" pageNumber="51">Kenting National</collectorName>
Park
</location>
, 
<collectingRegion id="F24566C6FFA4FFB9FFD8F88DFE393A6C" box="[217,407,1803,1829]" country="Taiwan" name="Pingtung" pageId="4" pageNumber="51">Pingtung County</collectingRegion>
(
<geoCoordinate id="55B5CEE3FFA4FFB9FEA7F88DFDA73A6F" box="[422,521,1803,1830]" degrees="21.95" direction="north" orientation="latitude" pageId="4" pageNumber="51" precision="555" value="21.95">21.95°N</geoCoordinate>
, 
<geoCoordinate id="55B5CEE3FFA4FFB9FD15F88DFDD33A6C" box="[532,637,1803,1829]" degrees="120.81" direction="east" orientation="longitude" pageId="4" pageNumber="51" precision="555" value="120.81">120.81°E</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="789BFBAFFFA4FFB8FFC6F8A9FC763DB0" lastPageId="5" lastPageNumber="52" pageId="4" pageNumber="51" type="diagnosis">
<paragraph id="303EA824FFA4FFB8FFC6F8A9FC763DB0" blockId="4.[151,1437,1551,2009]" lastBlockId="5.[151,1436,151,249]" lastPageId="5" lastPageNumber="52" pageId="4" pageNumber="51">
<emphasis id="02F57436FFA4FFB9FFC6F8A9FEEC3A00" bold="true" box="[199,322,1839,1865]" pageId="4" pageNumber="51">Diagnosis.</emphasis>
<taxonomicName id="F781D3A7FFA4FFB9FE4AF8B6FE573A00" authority="Lo &amp; Wei &amp; Cheng, 2023" authorityName="Lo &amp; Wei &amp; Cheng" authorityYear="2023" box="[331,505,1839,1865]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="arborea" status="sp. nov.">
<emphasis id="02F57436FFA4FFB9FE4AF8B6FE573A00" box="[331,505,1839,1865]" italics="true" pageId="4" pageNumber="51">Hogna arborea</emphasis>
</taxonomicName>
<emphasis id="02F57436FFA4FFB9FD03F8A9FDF03A00" bold="true" box="[514,606,1839,1865]" pageId="4" pageNumber="51">
<taxonomicNameLabel id="19C6C94DFFA4FFB9FD03F8A9FDF03A00" box="[514,606,1839,1865]" pageId="4" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, is similar to 
<taxonomicName id="F781D3A7FFA4FFB9FC01F8B6FB3A3A00" authority="Yin, Bao &amp; Zhang, 1996" authorityName="Yin, Bao &amp; Zhang" authorityYear="1996" box="[768,1172,1839,1865]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="trunca">
<emphasis id="02F57436FFA4FFB9FC01F8B6FCC03A00" box="[768,878,1840,1865]" italics="true" pageId="4" pageNumber="51">H. trunca</emphasis>
<bibRefCitation id="5410D5D5FFA4FFB9FC77F8A9FB3A3A00" author="Yin, C. &amp; Bao, Y. &amp; Zhang, Y." box="[886,1172,1839,1865]" pageId="4" pageNumber="51" pagination="5 - 9" refId="ref7990" refString="Yin, C., Bao, Y. &amp; Zhang, Y. (1996) On two new species of wolf spiders from Zhejiang province (Araneae: Lycosidae). Acta Archnologica Sinica, 5, 5 - 9." type="journal article" year="1996">
Yin, Bao &amp; Zhang, 
<quantity id="F77905C1FFA4FFB9FB5DF8A9FB3A3A00" box="[1116,1172,1839,1865]" metricMagnitude="1" metricUnit="m" metricValue="5.06984" pageId="4" pageNumber="51" unit="in" value="1996.0">1996</quantity>
</bibRefCitation>
</taxonomicName>
in the female epigyne, the shape of inversed T-shaped septal pedicel, and in the median apophysis of male palp organ with a triangular prominence toward the ventral side. However, 
<taxonomicName id="F781D3A7FFA4FFB9FDA0F8FEFCB23AD8" authority="Lo &amp; Wei &amp; Cheng, 2023" authorityName="Lo &amp; Wei &amp; Cheng" authorityYear="2023" box="[673,796,1911,1937]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="arborea" status="sp. nov.">
<emphasis id="02F57436FFA4FFB9FDA0F8FEFCB23AD8" box="[673,796,1911,1937]" italics="true" pageId="4" pageNumber="51">H. arborea</emphasis>
</taxonomicName>
<emphasis id="02F57436FFA4FFB9FC22F8F1FCD23AD8" bold="true" box="[803,892,1911,1937]" pageId="4" pageNumber="51">
<taxonomicNameLabel id="19C6C94DFFA4FFB9FC22F8F1FCD23AD8" box="[803,892,1911,1937]" pageId="4" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, can be distinguished from the latter by its vulva with a pair of large, leaf-like massive folds, septum pedicel of epigyne narrower and conspicuously constricted anteriorly (
<figureCitation id="A8BAB4A1FFA4FFB9FE13F839FE213A90" box="[274,399,1983,2009]" captionStart="FIGURE 3" captionStartId="5.[151,250,951,975]" captionTargetBox="[151,1432,276,925]" captionTargetId="figure-71@5.[151,1435,273,927]" captionTargetPageId="5" captionText="FIGURE 3 A–E. Epigyne and left palp organ of Hogna arborea sp. nov. (A–B, Holotype, TESRI- Ar3045; C–E, Paratype, TESRI-Ar5507). A, ventral view; B, dorsal view; C, prolateral view; D, ventral view; E, retrolateral view. Scale bar = 0.05mm. Abbreviations: BS- base of spermatheca, C- conductor, CD- copulatory duct, Em- embolus, H- hood, HS- head of spermatheca, MA- median apophysis, MF- massive fold, P- palea, S- septum, SB- septum base, St- subtegulum, T- tegulum, TA- terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.8426710" httpUri="https://zenodo.org/record/8426710/files/figure.png" pageId="4" pageNumber="51">Figs 3A–B</figureCitation>
, 
<figureCitation id="A8BAB4A1FFA4FFB9FE98F839FE4A3A90" box="[409,484,1983,2009]" captionStart="FIGURE 4" captionStartId="5.[151,250,1827,1851]" captionTargetBox="[158,1431,1146,1802]" captionTargetId="figure-166@5.[151,1436,1143,1804]" captionTargetPageId="5" captionText="FIGURE 4 A–E. Epigyne and left palp organ of Hogna arborea sp. nov. (A–B, Holotype, TESRI- Ar3045; C–E, Paratype, TESRI-Ar5507). A, ventral view; B, dorsal view; C, prolateral view; D, ventral view; E, retrolateral view. Scale bar = 0.05mm. Abbreviations: BS- base of spermatheca, C- conductor, CD- copulatory duct, Em- embolus, H- hood, HS- head of spermatheca, MA- median apophysis, MF- massive fold, P- palea, S- septum, SB- septum base, St- subtegulum, T- tegulum, TA- terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.8426712" httpUri="https://zenodo.org/record/8426712/files/figure.png" pageId="4" pageNumber="51">4 A–B</figureCitation>
), apex of the cymbium of male pedipalp not hook-shaped and without a bent ventrally (
<figureCitation id="A8BAB4A1FFA5FFB8FF9EFF11FF593DF8" box="[159,247,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,951,975]" captionTargetBox="[151,1432,276,925]" captionTargetId="figure-71@5.[151,1435,273,927]" captionTargetPageId="5" captionText="FIGURE 3 A–E. Epigyne and left palp organ of Hogna arborea sp. nov. (A–B, Holotype, TESRI- Ar3045; C–E, Paratype, TESRI-Ar5507). A, ventral view; B, dorsal view; C, prolateral view; D, ventral view; E, retrolateral view. Scale bar = 0.05mm. Abbreviations: BS- base of spermatheca, C- conductor, CD- copulatory duct, Em- embolus, H- hood, HS- head of spermatheca, MA- median apophysis, MF- massive fold, P- palea, S- septum, SB- septum base, St- subtegulum, T- tegulum, TA- terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.8426710" httpUri="https://zenodo.org/record/8426710/files/figure.png" pageId="5" pageNumber="52">Figs 3E</figureCitation>
, 
<figureCitation id="A8BAB4A1FFA5FFB8FE00FF11FE8D3DF8" box="[257,291,151,177]" captionStart="FIGURE 4" captionStartId="5.[151,250,1827,1851]" captionTargetBox="[158,1431,1146,1802]" captionTargetId="figure-166@5.[151,1436,1143,1804]" captionTargetPageId="5" captionText="FIGURE 4 A–E. Epigyne and left palp organ of Hogna arborea sp. nov. (A–B, Holotype, TESRI- Ar3045; C–E, Paratype, TESRI-Ar5507). A, ventral view; B, dorsal view; C, prolateral view; D, ventral view; E, retrolateral view. Scale bar = 0.05mm. Abbreviations: BS- base of spermatheca, C- conductor, CD- copulatory duct, Em- embolus, H- hood, HS- head of spermatheca, MA- median apophysis, MF- massive fold, P- palea, S- septum, SB- septum base, St- subtegulum, T- tegulum, TA- terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.8426712" httpUri="https://zenodo.org/record/8426712/files/figure.png" pageId="5" pageNumber="52">4E</figureCitation>
; ventrally bent apex in 
<emphasis id="02F57436FFA5FFB8FD1FFF11FD0B3DF9" box="[542,677,151,176]" italics="true" pageId="5" pageNumber="52">
<taxonomicName id="F781D3A7FFA5FFB8FD1FFF11FD293DF9" authorityName="Yin, Bao &amp; Zhang" authorityYear="1996" box="[542,647,151,176]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="trunca">H. trunca</taxonomicName>
—
</emphasis>
<bibRefCitation id="5410D5D5FFA5FFB8FDABFF11FC853DF8" author="Shida, R." box="[682,811,151,177]" pageId="5" pageNumber="52" pagination="129 - 133" refId="ref7336" refString="Shida, R. (2022) First records of the arboreal wolf spider species Hogna trunca Yin, Bao &amp; Zhang 1996 (Araneae: Lycosidae) from Japan. Acta Arachnologica, 71, 129 - 133. https: // doi. org / 10.2476 / asjaa. 71.129" type="journal article" year="2022">Shida 2022</bibRefCitation>
: 
<figureCitation id="A8BAB4A1FFA5FFB8FC35FF11FC273DF8" box="[820,905,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,951,975]" captionTargetBox="[151,1432,276,925]" captionTargetId="figure-71@5.[151,1435,273,927]" captionTargetPageId="5" captionText="FIGURE 3 A–E. Epigyne and left palp organ of Hogna arborea sp. nov. (A–B, Holotype, TESRI- Ar3045; C–E, Paratype, TESRI-Ar5507). A, ventral view; B, dorsal view; C, prolateral view; D, ventral view; E, retrolateral view. Scale bar = 0.05mm. Abbreviations: BS- base of spermatheca, C- conductor, CD- copulatory duct, Em- embolus, H- hood, HS- head of spermatheca, MA- median apophysis, MF- massive fold, P- palea, S- septum, SB- septum base, St- subtegulum, T- tegulum, TA- terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.8426710" httpUri="https://zenodo.org/record/8426710/files/figure.png" pageId="5" pageNumber="52">Fig. 3D</figureCitation>
), and a palea proportionally larger.Additionally, specimens of 
<taxonomicName id="F781D3A7FFA5FFB8FE2FFF3DFE063D9C" authority="Lo &amp; Wei &amp; Cheng, 2023" authorityName="Lo &amp; Wei &amp; Cheng" authorityYear="2023" box="[302,424,187,213]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="arborea" status="sp. nov.">
<emphasis id="02F57436FFA5FFB8FE2FFF3DFE063D9C" box="[302,424,187,213]" italics="true" pageId="5" pageNumber="52">H. arborea</emphasis>
</taxonomicName>
<emphasis id="02F57436FFA5FFB8FEACFF3DFDAB3D9C" bold="true" box="[429,517,187,213]" pageId="5" pageNumber="52">
<taxonomicNameLabel id="19C6C94DFFA5FFB8FEACFF3DFDAB3D9C" box="[429,517,187,213]" pageId="5" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(females 
<quantity id="F77905C1FFA5FFB8FD70FF3DFCBB3D9C" box="[625,789,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.09" metricValueMax="2.12" metricValueMin="2.06" pageId="5" pageNumber="52" unit="mm" value="20.9" valueMax="21.2" valueMin="20.6">20.6–21.2 mm</quantity>
; male 
<quantity id="F77905C1FFA5FFB8FC5AFF3CFC103D9C" box="[859,958,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="5" pageNumber="52" unit="mm" value="19.5">19.5 mm</quantity>
) are larger than those of 
<taxonomicName id="F781D3A7FFA5FFB8FBCCFF3DFA983D9D" authorityName="Yin, Bao &amp; Zhang" authorityYear="1996" box="[1229,1334,187,212]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="trunca">
<emphasis id="02F57436FFA5FFB8FBCCFF3DFA983D9D" box="[1229,1334,187,212]" italics="true" pageId="5" pageNumber="52">H. trunca</emphasis>
</taxonomicName>
(females 
<quantity id="F77905C1FFA5FFB8FF96FF59FE933DB0" box="[151,317,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.8049999999999997" metricValueMax="2.1399999999999997" metricValueMin="1.47" pageId="5" pageNumber="52" unit="mm" value="18.049999999999997" valueMax="21.4" valueMin="14.7">14.7–21.4 mm</quantity>
; males 13.0– 
<quantity id="F77905C1FFA5FFB8FED0FF58FD993DB0" box="[465,567,222,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="5" pageNumber="52" unit="mm" value="15.6">15.6 mm</quantity>
; see 
<bibRefCitation id="5410D5D5FFA5FFB8FD6DFF59FCBC3DB0" author="Yin, C. &amp; Bao, Y. &amp; Zhang, Y." box="[620,786,223,249]" pageId="5" pageNumber="52" pagination="5 - 9" refId="ref7990" refString="Yin, C., Bao, Y. &amp; Zhang, Y. (1996) On two new species of wolf spiders from Zhejiang province (Araneae: Lycosidae). Acta Archnologica Sinica, 5, 5 - 9." type="journal article" year="1996">
Yin 
<emphasis id="02F57436FFA5FFB8FD9AFF59FD633DB0" box="[667,717,223,249]" italics="true" pageId="5" pageNumber="52">et al</emphasis>
. 1996
</bibRefCitation>
and 
<bibRefCitation id="5410D5D5FFA5FFB8FC48FF59FC633DB0" author="Shida, R." box="[841,973,223,249]" pageId="5" pageNumber="52" pagination="129 - 133" refId="ref7336" refString="Shida, R. (2022) First records of the arboreal wolf spider species Hogna trunca Yin, Bao &amp; Zhang 1996 (Araneae: Lycosidae) from Japan. Acta Arachnologica, 71, 129 - 133. https: // doi. org / 10.2476 / asjaa. 71.129" type="journal article" year="2022">Shida 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="64FEF8ACFFA5FFB8FF96FC31FEAA3916" ID-DOI="http://doi.org/10.5281/zenodo.8426710" ID-Zenodo-Dep="8426710" httpUri="https://zenodo.org/record/8426710/files/figure.png" pageId="5" pageNumber="52" startId="5.[151,250,951,975]" targetBox="[151,1432,276,925]" targetPageId="5" targetType="figure">
<paragraph id="303EA824FFA5FFB8FF96FC31FEAA3916" blockId="5.[151,1437,951,1119]" pageId="5" pageNumber="52">
<emphasis id="02F57436FFA5FFB8FF96FC31FEE23E86" bold="true" box="[151,332,951,975]" pageId="5" pageNumber="52">FIGURE 3 A–E.</emphasis>
Epigyne and left palp organ of 
<taxonomicName id="F781D3A7FFA5FFB8FD9BFC3EFC963E86" authority="Lo &amp; Wei &amp; Cheng, 2023" authorityName="Lo &amp; Wei &amp; Cheng" authorityYear="2023" box="[666,824,951,975]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="arborea" status="sp. nov.">
<emphasis id="02F57436FFA5FFB8FD9BFC3EFC963E86" box="[666,824,951,975]" italics="true" pageId="5" pageNumber="52">Hogna arborea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C6C94DFFA5FFB8FC41FC31FC3A3E87" box="[832,916,951,975]" pageId="5" pageNumber="52" rank="species">
<emphasis id="02F57436FFA5FFB8FC41FC31FC233E86" bold="true" box="[832,909,951,975]" pageId="5" pageNumber="52">sp. nov</emphasis>
.
</taxonomicNameLabel>
(A–B, Holotype, TESRI- Ar3045; C–E, Paratype, TESRI-Ar5507). 
<emphasis id="02F57436FFA5FFB8FE49FC5DFECE3EBA" bold="true" box="[328,352,987,1011]" pageId="5" pageNumber="52">A,</emphasis>
ventral view; 
<emphasis id="02F57436FFA5FFB8FEF0FC5DFDA63EBA" bold="true" box="[497,520,987,1011]" pageId="5" pageNumber="52">B,</emphasis>
dorsal view; 
<emphasis id="02F57436FFA5FFB8FD91FC5DFD073EBA" bold="true" box="[656,681,987,1011]" pageId="5" pageNumber="52">C,</emphasis>
prolateral view; 
<emphasis id="02F57436FFA5FFB8FC55FC5DFCC23EBA" bold="true" box="[852,876,987,1011]" pageId="5" pageNumber="52">D,</emphasis>
ventral view; 
<emphasis id="02F57436FFA5FFB8FCFCFC5DFBBA3EBA" bold="true" box="[1021,1044,987,1011]" pageId="5" pageNumber="52">E,</emphasis>
retrolateral view. Scale bar = 0.05mm. Abbreviations: 
<emphasis id="02F57436FFA5FFB8FE32FC79FEFC395E" bold="true" box="[307,338,1023,1047]" pageId="5" pageNumber="52">BS</emphasis>
- base of spermatheca, 
<emphasis id="02F57436FFA5FFB8FD36FC79FDE7395E" bold="true" box="[567,585,1023,1047]" pageId="5" pageNumber="52">C</emphasis>
- conductor, 
<emphasis id="02F57436FFA5FFB8FDC6FC79FD42395E" bold="true" box="[711,748,1023,1047]" pageId="5" pageNumber="52">CD</emphasis>
- copulatory duct, 
<emphasis id="02F57436FFA5FFB8FCA2FC79FC67395E" bold="true" box="[931,969,1023,1047]" pageId="5" pageNumber="52">Em</emphasis>
- embolus, 
<emphasis id="02F57436FFA5FFB8FB39FC79FBE2395E" bold="true" box="[1080,1100,1023,1047]" pageId="5" pageNumber="52">H</emphasis>
- hood, 
<emphasis id="02F57436FFA5FFB8FB99FC79FB14395E" bold="true" box="[1176,1210,1023,1047]" pageId="5" pageNumber="52">HS</emphasis>
- head of spermatheca, 
<emphasis id="02F57436FFA5FFB8FF96FBA5FF6F3972" bold="true" box="[151,193,1059,1083]" pageId="5" pageNumber="52">MA</emphasis>
- median apophysis, 
<emphasis id="02F57436FFA5FFB8FE97FBA5FE133972" bold="true" box="[406,445,1059,1083]" pageId="5" pageNumber="52">MF</emphasis>
- massive fold, 
<emphasis id="02F57436FFA5FFB8FD5DFBA5FDC53972" bold="true" box="[604,619,1059,1083]" pageId="5" pageNumber="52">P</emphasis>
- palea, 
<emphasis id="02F57436FFA5FFB8FDBFFBA5FD623972" bold="true" box="[702,716,1059,1083]" pageId="5" pageNumber="52">S</emphasis>
- septum, 
<emphasis id="02F57436FFA5FFB8FC33FBA5FCFF3972" bold="true" box="[818,849,1059,1083]" pageId="5" pageNumber="52">SB</emphasis>
- septum base, 
<emphasis id="02F57436FFA5FFB8FCEDFBA5FBAD3972" bold="true" box="[1004,1027,1059,1083]" pageId="5" pageNumber="52">St</emphasis>
- subtegulum, 
<emphasis id="02F57436FFA5FFB8FB97FBA5FB093972" bold="true" box="[1174,1191,1059,1083]" pageId="5" pageNumber="52">T</emphasis>
- tegulum, 
<emphasis id="02F57436FFA5FFB8FA16FBA5FA963972" bold="true" box="[1303,1336,1059,1083]" pageId="5" pageNumber="52">TA</emphasis>
- terminal apophysis.
</paragraph>
</caption>
<caption id="64FEF8ACFFA5FFB8FF96F8A5FEAA3A85" ID-DOI="http://doi.org/10.5281/zenodo.8426712" ID-Zenodo-Dep="8426712" httpUri="https://zenodo.org/record/8426712/files/figure.png" pageId="5" pageNumber="52" startId="5.[151,250,1827,1851]" targetBox="[158,1431,1146,1802]" targetPageId="5" targetType="figure">
<paragraph id="303EA824FFA5FFB8FF96F8A5FEAA3A85" blockId="5.[151,1437,1827,1996]" pageId="5" pageNumber="52">
<emphasis id="02F57436FFA5FFB8FF96F8A5FEE23A72" bold="true" box="[151,332,1827,1851]" pageId="5" pageNumber="52">FIGURE 4 A–E.</emphasis>
Epigyne and left palp organ of 
<taxonomicName id="F781D3A7FFA5FFB8FD9BF8A2FC963A75" authority="Lo &amp; Wei &amp; Cheng, 2023" authorityName="Lo &amp; Wei &amp; Cheng" authorityYear="2023" box="[666,824,1828,1852]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="arborea" status="sp. nov.">
<emphasis id="02F57436FFA5FFB8FD9BF8A2FC963A75" box="[666,824,1828,1852]" italics="true" pageId="5" pageNumber="52">Hogna arborea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C6C94DFFA5FFB8FC41F8A2FC3A3A72" box="[832,916,1828,1852]" pageId="5" pageNumber="52" rank="species">
<emphasis id="02F57436FFA5FFB8FC41F8A2FC233A75" bold="true" box="[832,909,1828,1852]" pageId="5" pageNumber="52">sp. nov</emphasis>
.
</taxonomicNameLabel>
(A–B, Holotype, TESRI- Ar3045; C–E, Paratype, TESRI-Ar5507). 
<emphasis id="02F57436FFA5FFB8FE49F8C1FECE3A16" bold="true" box="[328,352,1863,1887]" pageId="5" pageNumber="52">A,</emphasis>
ventral view; 
<emphasis id="02F57436FFA5FFB8FEF0F8CEFDA63A29" bold="true" box="[497,520,1864,1888]" pageId="5" pageNumber="52">B,</emphasis>
dorsal view; 
<emphasis id="02F57436FFA5FFB8FD91F8C1FD073A16" bold="true" box="[656,681,1863,1887]" pageId="5" pageNumber="52">C,</emphasis>
prolateral view; 
<emphasis id="02F57436FFA5FFB8FC55F8CEFCC23A29" bold="true" box="[852,876,1864,1888]" pageId="5" pageNumber="52">D,</emphasis>
ventral view; 
<emphasis id="02F57436FFA5FFB8FCFCF8CEFBBA3A29" bold="true" box="[1021,1044,1864,1888]" pageId="5" pageNumber="52">E,</emphasis>
retrolateral view. Scale bar = 0.05mm. Abbreviations: 
<emphasis id="02F57436FFA5FFB8FE32F8EDFEFC3ACA" bold="true" box="[307,338,1899,1923]" pageId="5" pageNumber="52">BS</emphasis>
- base of spermatheca, 
<emphasis id="02F57436FFA5FFB8FD36F8EDFDE73ACA" bold="true" box="[567,585,1899,1923]" pageId="5" pageNumber="52">C</emphasis>
- conductor, 
<emphasis id="02F57436FFA5FFB8FDC6F8EDFD423ACA" bold="true" box="[711,748,1899,1923]" pageId="5" pageNumber="52">CD</emphasis>
- copulatory duct, 
<emphasis id="02F57436FFA5FFB8FCA2F8EAFC673ACD" bold="true" box="[931,969,1900,1924]" pageId="5" pageNumber="52">Em</emphasis>
- embolus, 
<emphasis id="02F57436FFA5FFB8FB39F8EAFBE23ACD" bold="true" box="[1080,1100,1900,1924]" pageId="5" pageNumber="52">H</emphasis>
- hood, 
<emphasis id="02F57436FFA5FFB8FB99F8EDFB143ACA" bold="true" box="[1176,1210,1899,1923]" pageId="5" pageNumber="52">HS</emphasis>
- head of spermatheca, 
<emphasis id="02F57436FFA5FFB8FF96F809FF6F3AEE" bold="true" box="[151,193,1935,1959]" pageId="5" pageNumber="52">MA</emphasis>
- median apophysis, 
<emphasis id="02F57436FFA5FFB8FE97F816FE133AE1" bold="true" box="[406,445,1936,1960]" pageId="5" pageNumber="52">MF</emphasis>
- massive fold, 
<emphasis id="02F57436FFA5FFB8FD5DF816FDC53AE1" bold="true" box="[604,619,1936,1960]" pageId="5" pageNumber="52">P</emphasis>
- palea, 
<emphasis id="02F57436FFA5FFB8FDBFF809FD623AEE" bold="true" box="[702,716,1935,1959]" pageId="5" pageNumber="52">S</emphasis>
- septum, 
<emphasis id="02F57436FFA5FFB8FC33F809FCFF3AEE" bold="true" box="[818,849,1935,1959]" pageId="5" pageNumber="52">SB</emphasis>
- septum base, 
<emphasis id="02F57436FFA5FFB8FCEDF809FBAD3AEE" bold="true" box="[1004,1027,1935,1959]" pageId="5" pageNumber="52">St</emphasis>
- subtegulum, 
<emphasis id="02F57436FFA5FFB8FB97F816FB093AE1" bold="true" box="[1174,1191,1936,1960]" pageId="5" pageNumber="52">T</emphasis>
- tegulum, 
<emphasis id="02F57436FFA5FFB8FA16F809FA963AEE" bold="true" box="[1303,1336,1935,1959]" pageId="5" pageNumber="52">TA</emphasis>
- terminal apophysis.
</paragraph>
</caption>
<subSubSection id="789BFBAFFFA6FFBAFFC6FF10FB223C98" lastPageId="7" lastPageNumber="54" pageId="6" pageNumber="53" type="description">
<paragraph id="303EA824FFA6FFBBFFC6FF10FD543FE0" blockId="6.[151,1437,150,681]" pageId="6" pageNumber="53">
<emphasis id="02F57436FFA6FFBBFFC6FF10FEFF3DF9" bold="true" box="[199,337,150,176]" pageId="6" pageNumber="53">Description</emphasis>
. 
<emphasis id="02F57436FFA6FFBBFE5FFF11FD4B3DF8" box="[350,741,151,177]" italics="true" pageId="6" pageNumber="53">
Female (TESRI-Ar3045, 
<typeStatus id="EF3A1686FFA6FFBBFD75FF11FD773DF8" box="[628,729,151,177]" pageId="6" pageNumber="53">holotype</typeStatus>
).
</emphasis>
Total length 20.6, carapace length 10.0, width 8.3; Opistosoma length 10.6, width 8.0. Prosoma in lateral view with gradually descending thoracic region, ocular area not elevated (
<figureCitation id="A8BAB4A1FFA6FFBBFF9EFF59FF593DB0" box="[159,247,222,249]" captionStart="FIGURE 5" captionStartId="6.[151,250,1449,1473]" captionTargetBox="[155,1435,705,1421]" captionTargetId="figure-277@6.[151,1435,705,1425]" captionTargetPageId="6" captionText="FIGURE 5 A–G. Habitus of Hogna arborea sp. nov. (A, C, E, G, Holotype, TESRI- Ar3045; B, D, F, Paratype, TESRI- Ar5507). A–B, general appearance, dorsal view; C–D, prosoma, lateral view; E–F, eyes; G, metatarsus I and tarsus I, prolateral view. Scale bar = 5mm (A–D); 0.5mm (E–F), 1mm (G)." figureDoi="http://doi.org/10.5281/zenodo.8426716" httpUri="https://zenodo.org/record/8426716/files/figure.png" pageId="6" pageNumber="53">Fig. 5C</figureCitation>
). Carapace yellowish with a distinct brown central band from PME to posterior margin, and a pair of black, irregular inner edge stripes on lateral region. Fovea longitudinal, cervical and radical grooves indistinct. Clypeus, AER and PME region black, AER straight and PER strongly recurved. AME slightly larger than ALE, and width of AER approximately equal to those of SER (
<figureCitation id="A8BAB4A1FFA6FFBBFDACFECDFCAB3C2C" box="[685,773,330,357]" captionStart="FIGURE 5" captionStartId="6.[151,250,1449,1473]" captionTargetBox="[155,1435,705,1421]" captionTargetId="figure-277@6.[151,1435,705,1425]" captionTargetPageId="6" captionText="FIGURE 5 A–G. Habitus of Hogna arborea sp. nov. (A, C, E, G, Holotype, TESRI- Ar3045; B, D, F, Paratype, TESRI- Ar5507). A–B, general appearance, dorsal view; C–D, prosoma, lateral view; E–F, eyes; G, metatarsus I and tarsus I, prolateral view. Scale bar = 5mm (A–D); 0.5mm (E–F), 1mm (G)." figureDoi="http://doi.org/10.5281/zenodo.8426716" httpUri="https://zenodo.org/record/8426716/files/figure.png" pageId="6" pageNumber="53">Fig. 5E</figureCitation>
). Eye diameters and interdistances: AME 0.50, ALE 0.40, PME 1.0, PLE 0.9; MOA-L 1.70, MOA-AW 1.30, MOA-PW 2.60; AME-AME 0.20, AME-ALE 0.20, PME-PME 0.90, PME-PLE 1.00; clypeus height 0.50. Chelicerae massive, black with several reddish setae at the frontside. Chelicera furrow with one promarginal tooth and three retromarginal teeth. Endites and labium black. Sternum black, heart-like, densely covered by setae. Legs strong, yellowish with black annular patches and armed with several macrosetae. Venter of coxa black while alive. Metatarsus and tarsus I + II with well-developed scopulae (
<figureCitation id="A8BAB4A1FFA6FFBBFF9EFDA5FF543F74" box="[159,250,546,573]" captionStart="FIGURE 5" captionStartId="6.[151,250,1449,1473]" captionTargetBox="[155,1435,705,1421]" captionTargetId="figure-277@6.[151,1435,705,1425]" captionTargetPageId="6" captionText="FIGURE 5 A–G. Habitus of Hogna arborea sp. nov. (A, C, E, G, Holotype, TESRI- Ar3045; B, D, F, Paratype, TESRI- Ar5507). A–B, general appearance, dorsal view; C–D, prosoma, lateral view; E–F, eyes; G, metatarsus I and tarsus I, prolateral view. Scale bar = 5mm (A–D); 0.5mm (E–F), 1mm (G)." figureDoi="http://doi.org/10.5281/zenodo.8426716" httpUri="https://zenodo.org/record/8426716/files/figure.png" pageId="6" pageNumber="53">Fig. 5G</figureCitation>
). Measurements of pedipalps and legs: pedipalp 11.5 (4.0, 1.9, 2.4, 3.2), leg I 32.4 (8.7, 3.5, 7.0, 9.4, 3.8), leg II 25.0 (7.1, 3.3, 5.3, 6.1, 3.2), leg III 26.1 (7.5, 3.5, 5.8, 6.0, 3.3), leg IV 28.2 (8.1, 4.0, 6.4, 6.2, 3.5). Opistosoma oval, dark brown, with a dark cardiac mark and four black transverse lines behind the cardiac mark (
<figureCitation id="A8BAB4A1FFA6FFBBFBF2FDEDFAE33FCC" box="[1267,1357,618,645]" captionStart="FIGURE 5" captionStartId="6.[151,250,1449,1473]" captionTargetBox="[155,1435,705,1421]" captionTargetId="figure-277@6.[151,1435,705,1425]" captionTargetPageId="6" captionText="FIGURE 5 A–G. Habitus of Hogna arborea sp. nov. (A, C, E, G, Holotype, TESRI- Ar3045; B, D, F, Paratype, TESRI- Ar5507). A–B, general appearance, dorsal view; C–D, prosoma, lateral view; E–F, eyes; G, metatarsus I and tarsus I, prolateral view. Scale bar = 5mm (A–D); 0.5mm (E–F), 1mm (G)." figureDoi="http://doi.org/10.5281/zenodo.8426716" httpUri="https://zenodo.org/record/8426716/files/figure.png" pageId="6" pageNumber="53">Fig. 5A</figureCitation>
). Sides black with scattered brown spots. Venter almost black.
</paragraph>
<caption id="64FEF8ACFFA6FFBBFF96FA2FFD613B40" ID-DOI="http://doi.org/10.5281/zenodo.8426716" ID-Zenodo-Dep="8426716" httpUri="https://zenodo.org/record/8426716/files/figure.png" pageId="6" pageNumber="53" startId="6.[151,250,1449,1473]" targetBox="[155,1435,705,1421]" targetPageId="6" targetType="figure">
<paragraph id="303EA824FFA6FFBBFF96FA2FFD613B40" blockId="6.[151,1436,1449,1545]" pageId="6" pageNumber="53">
<emphasis id="02F57436FFA6FFBBFF96FA2FFEFF3888" bold="true" box="[151,337,1449,1473]" pageId="6" pageNumber="53">FIGURE 5 A–G.</emphasis>
Habitus of 
<taxonomicName id="F781D3A7FFA6FFBBFED0FA2CFDDE3888" authority="Lo &amp; Wei &amp; Cheng, 2023" authorityName="Lo &amp; Wei &amp; Cheng" authorityYear="2023" box="[465,624,1449,1473]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="arborea" status="sp. nov.">
<emphasis id="02F57436FFA6FFBBFED0FA2CFDDE3888" box="[465,624,1449,1473]" italics="true" pageId="6" pageNumber="53">Hogna arborea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C6C94DFFA6FFBBFD78FA2FFD613888" box="[633,719,1449,1473]" pageId="6" pageNumber="53" rank="species">
<emphasis id="02F57436FFA6FFBBFD78FA2FFD663888" bold="true" box="[633,712,1449,1473]" pageId="6" pageNumber="53">sp. nov</emphasis>
.
</taxonomicNameLabel>
(A, C, E, G, Holotype, TESRI- Ar3045; B, D, F, Paratype, TESRIAr5507). 
<emphasis id="02F57436FFA6FFBBFFF8FA4BFE8138AC" bold="true" box="[249,303,1485,1509]" pageId="6" pageNumber="53">A–B,</emphasis>
general appearance, dorsal view; 
<emphasis id="02F57436FFA6FFBBFD87FA4BFD1338AC" bold="true" box="[646,701,1485,1509]" pageId="6" pageNumber="53">C–D,</emphasis>
prosoma, lateral view; 
<emphasis id="02F57436FFA6FFBBFCABFA4BFC7538AC" bold="true" box="[938,987,1485,1509]" pageId="6" pageNumber="53">E–F,</emphasis>
eyes; 
<emphasis id="02F57436FFA6FFBBFB1AFA4BFB9A38AC" bold="true" box="[1051,1076,1485,1509]" pageId="6" pageNumber="53">G,</emphasis>
metatarsus I and tarsus I, prolateral view. Scale bar = 5mm (A–D); 0.5mm (E–F), 1mm (G).
</paragraph>
</caption>
<paragraph id="303EA824FFA6FFBBFFC6F9DDFABE3BF4" blockId="6.[151,1437,1627,2014]" pageId="6" pageNumber="53">
Epigyne (
<figureCitation id="A8BAB4A1FFA6FFBBFE32F9DDFE213B3C" box="[307,399,1627,1653]" captionStart="FIGURE 3" captionStartId="5.[151,250,951,975]" captionTargetBox="[151,1432,276,925]" captionTargetId="figure-71@5.[151,1435,273,927]" captionTargetPageId="5" captionText="FIGURE 3 A–E. Epigyne and left palp organ of Hogna arborea sp. nov. (A–B, Holotype, TESRI- Ar3045; C–E, Paratype, TESRI-Ar5507). A, ventral view; B, dorsal view; C, prolateral view; D, ventral view; E, retrolateral view. Scale bar = 0.05mm. Abbreviations: BS- base of spermatheca, C- conductor, CD- copulatory duct, Em- embolus, H- hood, HS- head of spermatheca, MA- median apophysis, MF- massive fold, P- palea, S- septum, SB- septum base, St- subtegulum, T- tegulum, TA- terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.8426710" httpUri="https://zenodo.org/record/8426710/files/figure.png" pageId="6" pageNumber="53">Figs 3A</figureCitation>
, 
<figureCitation id="A8BAB4A1FFA6FFBBFE9AF9DDFE103B3C" box="[411,446,1627,1653]" captionStart="FIGURE 4" captionStartId="5.[151,250,1827,1851]" captionTargetBox="[158,1431,1146,1802]" captionTargetId="figure-166@5.[151,1436,1143,1804]" captionTargetPageId="5" captionText="FIGURE 4 A–E. Epigyne and left palp organ of Hogna arborea sp. nov. (A–B, Holotype, TESRI- Ar3045; C–E, Paratype, TESRI-Ar5507). A, ventral view; B, dorsal view; C, prolateral view; D, ventral view; E, retrolateral view. Scale bar = 0.05mm. Abbreviations: BS- base of spermatheca, C- conductor, CD- copulatory duct, Em- embolus, H- hood, HS- head of spermatheca, MA- median apophysis, MF- massive fold, P- palea, S- septum, SB- septum base, St- subtegulum, T- tegulum, TA- terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.8426712" httpUri="https://zenodo.org/record/8426712/files/figure.png" pageId="6" pageNumber="53">4A</figureCitation>
) with a pair of anterior hoods. Median septum inverted T-shaped, middle part widest and anterior constricted. Length of septum longer than width of septal base. Internal genitalia with a pair of massive, leaf-like and wiggly folds (
<figureCitation id="A8BAB4A1FFA6FFBBFEC6F925FD8C3BF4" box="[455,546,1699,1725]" captionStart="FIGURE 3" captionStartId="5.[151,250,951,975]" captionTargetBox="[151,1432,276,925]" captionTargetId="figure-71@5.[151,1435,273,927]" captionTargetPageId="5" captionText="FIGURE 3 A–E. Epigyne and left palp organ of Hogna arborea sp. nov. (A–B, Holotype, TESRI- Ar3045; C–E, Paratype, TESRI-Ar5507). A, ventral view; B, dorsal view; C, prolateral view; D, ventral view; E, retrolateral view. Scale bar = 0.05mm. Abbreviations: BS- base of spermatheca, C- conductor, CD- copulatory duct, Em- embolus, H- hood, HS- head of spermatheca, MA- median apophysis, MF- massive fold, P- palea, S- septum, SB- septum base, St- subtegulum, T- tegulum, TA- terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.8426710" httpUri="https://zenodo.org/record/8426710/files/figure.png" pageId="6" pageNumber="53">Figs 3B</figureCitation>
, 
<figureCitation id="A8BAB4A1FFA6FFBBFD2FF925FDFC3BF4" box="[558,594,1699,1725]" captionStart="FIGURE 4" captionStartId="5.[151,250,1827,1851]" captionTargetBox="[158,1431,1146,1802]" captionTargetId="figure-166@5.[151,1436,1143,1804]" captionTargetPageId="5" captionText="FIGURE 4 A–E. Epigyne and left palp organ of Hogna arborea sp. nov. (A–B, Holotype, TESRI- Ar3045; C–E, Paratype, TESRI-Ar5507). A, ventral view; B, dorsal view; C, prolateral view; D, ventral view; E, retrolateral view. Scale bar = 0.05mm. Abbreviations: BS- base of spermatheca, C- conductor, CD- copulatory duct, Em- embolus, H- hood, HS- head of spermatheca, MA- median apophysis, MF- massive fold, P- palea, S- septum, SB- septum base, St- subtegulum, T- tegulum, TA- terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.8426712" httpUri="https://zenodo.org/record/8426712/files/figure.png" pageId="6" pageNumber="53">4B</figureCitation>
). Spermathecae columnar with irregular projections on the tip.
</paragraph>
<paragraph id="303EA824FFA6FFBBFFC6F941FD783AF0" blockId="6.[151,1437,1627,2014]" pageId="6" pageNumber="53">
<emphasis id="02F57436FFA6FFBBFFC6F941FD933BAB" box="[199,573,1735,1762]" italics="true" pageId="6" pageNumber="53">
Male (TESRI-Ar5507, 
<typeStatus id="EF3A1686FFA6FFBBFEC9F94EFD9E3BAB" box="[456,560,1736,1762]" pageId="6" pageNumber="53">paratype</typeStatus>
).
</emphasis>
Body shape and color pattern similar to females (
<figureCitation id="A8BAB4A1FFA6FFBBFB77F941FB613BAB" box="[1142,1231,1735,1762]" captionStart="FIGURE 5" captionStartId="6.[151,250,1449,1473]" captionTargetBox="[155,1435,705,1421]" captionTargetId="figure-277@6.[151,1435,705,1425]" captionTargetPageId="6" captionText="FIGURE 5 A–G. Habitus of Hogna arborea sp. nov. (A, C, E, G, Holotype, TESRI- Ar3045; B, D, F, Paratype, TESRI- Ar5507). A–B, general appearance, dorsal view; C–D, prosoma, lateral view; E–F, eyes; G, metatarsus I and tarsus I, prolateral view. Scale bar = 5mm (A–D); 0.5mm (E–F), 1mm (G)." figureDoi="http://doi.org/10.5281/zenodo.8426716" httpUri="https://zenodo.org/record/8426716/files/figure.png" pageId="6" pageNumber="53">Fig. 5B</figureCitation>
), but the prosoma in lateral view is obviously inclined behind fovea (
<figureCitation id="A8BAB4A1FFA6FFBBFDE5F96DFCEF3A4F" box="[740,833,1771,1798]" captionStart="FIGURE 5" captionStartId="6.[151,250,1449,1473]" captionTargetBox="[155,1435,705,1421]" captionTargetId="figure-277@6.[151,1435,705,1425]" captionTargetPageId="6" captionText="FIGURE 5 A–G. Habitus of Hogna arborea sp. nov. (A, C, E, G, Holotype, TESRI- Ar3045; B, D, F, Paratype, TESRI- Ar5507). A–B, general appearance, dorsal view; C–D, prosoma, lateral view; E–F, eyes; G, metatarsus I and tarsus I, prolateral view. Scale bar = 5mm (A–D); 0.5mm (E–F), 1mm (G)." figureDoi="http://doi.org/10.5281/zenodo.8426716" httpUri="https://zenodo.org/record/8426716/files/figure.png" pageId="6" pageNumber="53">Fig. 5D</figureCitation>
). Total length 19.5, carapace length 10.0, width 8.0; opistosoma length 9.5, width 7.2. AER approximately equal to SER (
<figureCitation id="A8BAB4A1FFA6FFBBFCB0F889FBA73A63" box="[945,1033,1807,1834]" captionStart="FIGURE 5" captionStartId="6.[151,250,1449,1473]" captionTargetBox="[155,1435,705,1421]" captionTargetId="figure-277@6.[151,1435,705,1425]" captionTargetPageId="6" captionText="FIGURE 5 A–G. Habitus of Hogna arborea sp. nov. (A, C, E, G, Holotype, TESRI- Ar3045; B, D, F, Paratype, TESRI- Ar5507). A–B, general appearance, dorsal view; C–D, prosoma, lateral view; E–F, eyes; G, metatarsus I and tarsus I, prolateral view. Scale bar = 5mm (A–D); 0.5mm (E–F), 1mm (G)." figureDoi="http://doi.org/10.5281/zenodo.8426716" httpUri="https://zenodo.org/record/8426716/files/figure.png" pageId="6" pageNumber="53">Fig. 5F</figureCitation>
). Eye diameters and interdistances: AME 0.50, ALE 0.40, PME 0.9, PLE 0.9; MOA-L 1.60, MOA-AW 1.30, MOA-PW 2.30; AME-AME 0.30, AME-ALE 0.20, PME-PME 0.90, PME-PLE 0.80; clypeus height 0.30. Measurements of pedipalps and legs: pedipalp 11.9 (4.1, 1.9, 2.4, 3.5), leg I 33.1 (9.3, 4.1, 7.4, 8.3, 4.0), leg II 31.7 (9.2, 4.0, 8.6, 7.7, 4.0), leg III 29.2 (8.7, 3.6, 5.9, 7.4, 3.6), leg IV 36.3 (10.3, 3.8, 7.7, 10.5, 4.0).
</paragraph>
<paragraph id="303EA824FFA6FFBAFFC6F845FA333DB0" blockId="6.[151,1437,1627,2014]" lastBlockId="7.[151,1437,151,827]" lastPageId="7" lastPageNumber="54" pageId="6" pageNumber="53">
Palp organ (
<figureCitation id="A8BAB4A1FFA6FFBBFE53F845FE613A94" box="[338,463,1987,2013]" captionStart="FIGURE 3" captionStartId="5.[151,250,951,975]" captionTargetBox="[151,1432,276,925]" captionTargetId="figure-71@5.[151,1435,273,927]" captionTargetPageId="5" captionText="FIGURE 3 A–E. Epigyne and left palp organ of Hogna arborea sp. nov. (A–B, Holotype, TESRI- Ar3045; C–E, Paratype, TESRI-Ar5507). A, ventral view; B, dorsal view; C, prolateral view; D, ventral view; E, retrolateral view. Scale bar = 0.05mm. Abbreviations: BS- base of spermatheca, C- conductor, CD- copulatory duct, Em- embolus, H- hood, HS- head of spermatheca, MA- median apophysis, MF- massive fold, P- palea, S- septum, SB- septum base, St- subtegulum, T- tegulum, TA- terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.8426710" httpUri="https://zenodo.org/record/8426710/files/figure.png" pageId="6" pageNumber="53">Figs 3C–E</figureCitation>
, 
<figureCitation id="A8BAB4A1FFA6FFBBFEDDF845FD8F3A94" box="[476,545,1987,2013]" captionStart="FIGURE 4" captionStartId="5.[151,250,1827,1851]" captionTargetBox="[158,1431,1146,1802]" captionTargetId="figure-166@5.[151,1436,1143,1804]" captionTargetPageId="5" captionText="FIGURE 4 A–E. Epigyne and left palp organ of Hogna arborea sp. nov. (A–B, Holotype, TESRI- Ar3045; C–E, Paratype, TESRI-Ar5507). A, ventral view; B, dorsal view; C, prolateral view; D, ventral view; E, retrolateral view. Scale bar = 0.05mm. Abbreviations: BS- base of spermatheca, C- conductor, CD- copulatory duct, Em- embolus, H- hood, HS- head of spermatheca, MA- median apophysis, MF- massive fold, P- palea, S- septum, SB- septum base, St- subtegulum, T- tegulum, TA- terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.8426712" httpUri="https://zenodo.org/record/8426712/files/figure.png" pageId="6" pageNumber="53">4C–E</figureCitation>
). Cymbium 2.2 times longer than wide, with two claws and a bunch of apical macrosetae. Palea with two sickle-shaped terminal apophysis, one of which is parallel and behind embolus. Subtegulum much smaller than tegulum. Median apophysis large and transverse, with a retrolateral pointed tip and a triangular prominence toward the ventral side. Embolus curved, tapering on the tip behind the median apophysis.
</paragraph>
<paragraph id="303EA824FFA7FFBAFFC6FE84FB223C98" blockId="7.[151,1437,151,827]" pageId="7" pageNumber="54">
<emphasis id="02F57436FFA7FFBAFFC6FE84FE963C55" bold="true" box="[199,312,258,284]" pageId="7" pageNumber="54">Variation</emphasis>
. As male specimens were damaged except 
<specimenCount id="268763ADFFA7FFBAFC23FE85FC1B3C54" box="[802,949,259,285]" pageId="7" pageNumber="54" type="generic" typeStatus="paratype">one paratype</specimenCount>
, measures of morphological variation were based solely on female specimens. Values indicate mean ± standard deviation. Total length 20.9 ± 0.3, carapace length 10.2 ± 0.2, width 8.7 ± 0.5; opisthosoma length 10.6 ± 0.2, width 8.1 ± 0.6. Diameters of AME 0.53 ± 0.06, ALE 0.40 ± 0.00, PME 1.00 ± 0.00, PLE 0.9 ± 0.00, MOA-AW 1.37 ± 0.12, MOA-PW 2.63 ± 0.06, MOA-L 1.77 ± 0.12, AME-I 0.23 ± 0.06, PME-I 0.87 ± 0.06, AML-I 0.20 ± 0.00, PML-I 1.03 ± 0.06; Clypeus height 0.53 ± 0.06. Pedipalp 12.3 ± 0.8, leg I 31.1 ± 1.3, leg II 27.5 ± 2.2, leg III 26.7 ± 0.7, leg IV 32.5 ± 3.8.
</paragraph>
</subSubSection>
<subSubSection id="789BFBAFFFA7FFBAFFC6FE5CFC763CBC" box="[199,984,474,501]" pageId="7" pageNumber="54" type="distribution">
<paragraph id="303EA824FFA7FFBAFFC6FE5CFC763CBC" blockId="7.[151,1437,151,827]" box="[199,984,474,501]" pageId="7" pageNumber="54">
<emphasis id="02F57436FFA7FFBAFFC6FE5CFEF73CBD" bold="true" box="[199,345,474,500]" pageId="7" pageNumber="54">Distribution</emphasis>
. Known only from southern and eastern 
<collectingCountry id="4896E8B4FFA7FFBAFC25FE5DFCD83CBC" box="[804,886,475,501]" name="Taiwan" pageId="7" pageNumber="54">Taiwan</collectingCountry>
(
<figureCitation id="A8BAB4A1FFA7FFBAFC84FE5DFC683CBC" box="[901,966,475,501]" captionStart="FIGURE 1" captionStartId="1.[151,250,1910,1934]" captionTargetBox="[151,1435,346,1887]" captionTargetId="figure-90@1.[151,1435,345,1887]" captionTargetPageId="1" captionText="FIGURE 1. Collection localities of Hogna arborea sp. nov. from Taiwan in the present study." figureDoi="http://doi.org/10.5281/zenodo.8426704" httpUri="https://zenodo.org/record/8426704/files/figure.png" pageId="7" pageNumber="54">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="789BFBAFFFA7FFBAFFC6FE79FD143E70" pageId="7" pageNumber="54" type="etymology">
<paragraph id="303EA824FFA7FFBAFFC6FE79FE993F74" blockId="7.[151,1437,151,827]" pageId="7" pageNumber="54">
<emphasis id="02F57436FFA7FFBAFFC6FE79FEE03F50" bold="true" box="[199,334,511,537]" pageId="7" pageNumber="54">Etymology.</emphasis>
The specific epithet “ 
<emphasis id="02F57436FFA7FFBAFD41FE79FD073F50" box="[576,681,511,537]" italics="true" pageId="7" pageNumber="54">
<taxonomicName id="F781D3A7FFA7FFBAFD41FE79FD313F50" authorityName="Lo &amp; Wei &amp; Cheng" authorityYear="2023" box="[576,671,511,537]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="arborea">arborea</taxonomicName>
”
</emphasis>
is a Latinized adjective referring to the tree-dwelling natural history of the species.
</paragraph>
<paragraph id="303EA824FFA7FFBAFFC6FDC0FD143E70" blockId="7.[151,1437,151,827]" pageId="7" pageNumber="54">
<emphasis id="02F57436FFA7FFBAFFC6FDC0FD8B3F28" bold="true" box="[199,549,582,609]" pageId="7" pageNumber="54">DNA Barcode and phylogeny</emphasis>
. The COI alignment in our analysis consists of 669 base pairs and includes 41 terminals. The ML phylogenetic tree showed that 
<taxonomicName id="F781D3A7FFA7FFBAFDD5FDEDFC2C3FCC" authority="Lo &amp; Wei &amp; Cheng, 2023" authorityName="Lo &amp; Wei &amp; Cheng" authorityYear="2023" box="[724,898,619,645]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="arborea" status="sp. nov.">
<emphasis id="02F57436FFA7FFBAFDD5FDEDFC2C3FCC" box="[724,898,619,645]" italics="true" pageId="7" pageNumber="54">Hogna arborea</emphasis>
</taxonomicName>
<emphasis id="02F57436FFA7FFBAFC8AFDEDFC483FCC" bold="true" box="[907,998,619,645]" pageId="7" pageNumber="54">
<taxonomicNameLabel id="19C6C94DFFA7FFBAFC8AFDEDFC483FCC" box="[907,998,619,645]" pageId="7" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were placed within a clade with most other 
<taxonomicName id="F781D3A7FFA7FFBAFFDBFD09FE2D3FE0" box="[218,387,655,681]" pageId="7" pageNumber="54">
<emphasis id="02F57436FFA7FFBAFFDBFD09FE883FE1" box="[218,294,655,680]" italics="true" pageId="7" pageNumber="54">Hogna</emphasis>
species
</taxonomicName>
, including the 
<typeStatus id="EF3A1686FFA7FFBAFD2FFD09FDF03FE0" box="[558,606,655,681]" pageId="7" pageNumber="54">type</typeStatus>
species, 
<taxonomicName id="F781D3A7FFA7FFBAFDC9FD09FC903FE0" baseAuthorityName="Latreille" baseAuthorityYear="1817" box="[712,830,655,681]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="radiata">
<emphasis id="02F57436FFA7FFBAFDC9FD09FC903FE0" box="[712,830,655,681]" italics="true" pageId="7" pageNumber="54">H. radiata</emphasis>
</taxonomicName>
(
<figureCitation id="A8BAB4A1FFA7FFBAFC51FD09FC383FE0" box="[848,918,655,681]" captionStart="FIGURE 6" captionStartId="7.[151,250,1748,1772]" captionTargetBox="[155,1432,889,1720]" captionTargetId="figure-337@7.[151,1435,885,1724]" captionTargetPageId="7" captionText="FIGURE 6. Maximum likelihood phylogenetic tree of Hogna arborea sp. nov. and related species based on COI gene dataset. Number above branches are ultrafast bootstrap (UFBoot) and SH-like approximate likelihood ratio test (SH-aLRT) values (both values less than 70 were omitted). Scale bar shows substitutions per site. Species marked with an asterisk indicates the type species of the genus." figureDoi="http://doi.org/10.5281/zenodo.8426720" httpUri="https://zenodo.org/record/8426720/files/figure.png" pageId="7" pageNumber="54">Fig. 6</figureCitation>
). On the other hand, the intraspecific genetic distance (K2P) of 
<taxonomicName id="F781D3A7FFA7FFBAFE64FD35FE4F3F84" authorityName="Lo &amp; Wei &amp; Cheng" authorityYear="2023" box="[357,481,691,717]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="arborea">
<emphasis id="02F57436FFA7FFBAFE64FD35FE4F3F84" box="[357,481,691,717]" italics="true" pageId="7" pageNumber="54">H. arborea</emphasis>
</taxonomicName>
was 0.23%, and the minimum interspecific genetic distance was 6.28% compared to congeners (
<tableCitation id="7D039D9FFFA7FFBAFE17FD51FEC33FB8" box="[278,365,727,753]" captionStart="TABLE 2" captionStartId="8.[152,244,803,829]" captionText="TABLE 2. intra- and interspecific K2P genetic distance (%) among species within Hogna clade.The diagonal means intraspecific distances and lower triangular means interspecific distances." pageId="7" pageNumber="54">Table 2</tableCitation>
). Therefore, the result confirmed that 
<taxonomicName id="F781D3A7FFA7FFBAFC10FD51FC253FB8" authority="Lo &amp; Wei &amp; Cheng, 2023" authorityName="Lo &amp; Wei &amp; Cheng" authorityYear="2023" box="[785,907,727,753]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="arborea" status="sp. nov.">
<emphasis id="02F57436FFA7FFBAFC10FD51FC253FB8" box="[785,907,727,753]" italics="true" pageId="7" pageNumber="54">H. arborea</emphasis>
</taxonomicName>
<emphasis id="02F57436FFA7FFBAFC90FD51FC443FB8" bold="true" box="[913,1002,727,753]" pageId="7" pageNumber="54">
<taxonomicNameLabel id="19C6C94DFFA7FFBAFC90FD51FC443FB8" box="[913,1002,727,753]" pageId="7" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from 
<collectingCountry id="4896E8B4FFA7FFBAFB2AFD51FBD33FB8" box="[1067,1149,727,753]" name="Taiwan" pageId="7" pageNumber="54">Taiwan</collectingCountry>
is an independent species according to “10X rule” (
<bibRefCitation id="5410D5D5FFA7FFBAFEB2FD7DFD2A3E5C" author="Hebert, P. D. N. &amp; Stoeckle, M. Y. &amp; Zemlak, T. S. &amp; Francis, C. M." box="[435,644,763,789]" pageId="7" pageNumber="54" pagination="312" refId="ref5986" refString="Hebert, P. D. N., Stoeckle, M. Y., Zemlak, T. S. &amp; Francis, C. M. (2004) Identification of Birds through DNA Barcodes. PLOS Biology, 2, e 312. https: // doi. org / 10.1371 / journal. pbio. 0020312" type="journal article" year="2004">
Hebert 
<emphasis id="02F57436FFA7FFBAFD08FD7DFD943E5C" box="[521,570,763,789]" italics="true" pageId="7" pageNumber="54">et al</emphasis>
. 2004
</bibRefCitation>
; 
<bibRefCitation id="5410D5D5FFA7FFBAFD8CFD7DFC243E5C" author="Yang, Z. &amp; Rannala, B." box="[653,906,763,789]" pageId="7" pageNumber="54" pagination="3028 - 3036" refId="ref7942" refString="Yang, Z. &amp; Rannala, B. (2017) Bayesian species identification under the multispecies coalescent provides significant improvements to DNA barcoding analyses. Molecular Ecology, 26, 3028 - 3036. https: // doi. org / 10.1111 / mec. 14093" type="journal article" year="2017">Yang &amp; Rannala 2017</bibRefCitation>
), and the threshold between 1.2% and 3.6% for lycosids (according to 
<bibRefCitation id="5410D5D5FFA7FFBAFE94FCA6FD013E70" author="Candek, K. &amp; Kuntner, M." box="[405,687,798,827]" pageId="7" pageNumber="54" pagination="268 - 277" refId="ref5565" refString="Candek, K. &amp; Kuntner, M. (2015) DNA barcoding gap: reliable species identification over morphological and geographical scales. Molecular Ecology Resources, 15, 268 - 277. https: // doi. org / 10.1111 / 1755 - 0998.12304" type="journal article" year="2015">Čandek &amp; Kuntner 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>