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<document ID-CLB-Dataset="54510" ID-DOI="http://dx.doi.org/10.3897/dez.70.98873" ID-GBIF-Dataset="7e00d827-34bb-4253-9012-18cc783e1176" ID-Pensoft-Pub="1860-1324-1-113" ID-Pensoft-UUID="5A06769429B25374A83F6CAF37C454B1" ID-ZooBank="420300CAA42A45D19F572F8017534959" ModsDocID="1860-1324-70-1-113" checkinTime="1679031782851" checkinUser="pensoft" docAuthor="Nakamine, Hiroshi, Yamamoto, Shuhei, Takahashi, Yui &amp; Liu, Xingyue" docDate="2023" docId="78C00204201A547B84BDA59E80043721" docLanguage="en" docName="DeutEntomolZeit 70(1): 113-120" docOrigin="Deutsche Entomologische Zeitschrift 70 (1)" docPubDate="2023-03-16" docSource="http://dx.doi.org/10.3897/dez.70.98873" docTitle="Sisyroneurorthus Nakamine &amp; Yamamoto &amp; Takahashi &amp; Liu 2023, gen. nov." docType="treatment" docUuid="AA446547-70EC-4549-A734-76E156C4445B" docUuidSource="ZooBank" docVersion="5" id="5A06769429B25374A83F6CAF37C454B1" lastPageNumber="113" masterDocId="5A06769429B25374A83F6CAF37C454B1" masterDocTitle="A remarkable new genus of Nevrorthidae (Neuroptera, Osmyloidea) from mid-Cretaceous Kachin amber of northern Myanmar" masterLastPageNumber="120" masterPageNumber="113" pageNumber="113" updateTime="1732812803750" updateUser="ExternalLinkService">
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<mods:title id="5340F8433D75D66B25E42ED6717F2F92">A remarkable new genus of Nevrorthidae (Neuroptera, Osmyloidea) from mid-Cretaceous Kachin amber of northern Myanmar</mods:title>
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<mods:namePart id="F5025D8E6017F810FDDD7F8579290AD7">Nakamine, Hiroshi</mods:namePart>
<mods:nameIdentifier id="3E94B8A07AAC0BE37D41A2595F38A39F" type="ORCID">https://orcid.org/0000-0002-4640-2203</mods:nameIdentifier>
<mods:affiliation id="1F66840EFA64313C1A6A940BCCB6D99E">Minoh Park Insect Museum, Minoh Park 1 - 18, Minoh City, Osaka 562 - 0002, Japan</mods:affiliation>
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<mods:namePart id="A91FEB4D5D209416521DA4AC464AC1AE">Yamamoto, Shuhei</mods:namePart>
<mods:nameIdentifier id="7861AC82CCEDCBB44FA77C1BA37EFA9C" type="ORCID">https://orcid.org/0000-0002-4162-8457</mods:nameIdentifier>
<mods:affiliation id="320C5E802BE4C83ADE8CFD21F803C3AF">The Hokkaido University Museum, Hokkaido University, Kita 10 - jo, Nishi 8 - chome, Kita-ku, Sapporo 060 - 0810, Japan</mods:affiliation>
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<mods:namePart id="C262918C4D21A7F95449F8964089C2DC">Takahashi, Yui</mods:namePart>
<mods:nameIdentifier id="9E1D9596F5E8118026F47743A305659D" type="ORCID">https://orcid.org/0000-0003-2861-9940</mods:nameIdentifier>
<mods:affiliation id="5E1C505A31A5A7142818268DE23834B9">Keio Yochisha Elementary School, Ebisu 2 - 35 - 1, Shibuya-ku, Tokyo 150 - 0013, Japan</mods:affiliation>
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<mods:namePart id="C6C0F0EE9642909588BBA78AAD772DF9">Liu, Xingyue</mods:namePart>
<mods:nameIdentifier id="E88F089E1E877DDAFFECC1BBA2624951" type="ORCID">https://orcid.org/0000-0002-9168-0659</mods:nameIdentifier>
<mods:affiliation id="AC087BBFE0DDA880DFC1F52E04CBD80F">Department of Entomology, China Agricultural University, Beijing 100193, China</mods:affiliation>
<mods:nameIdentifier id="7E7970A4C0A85030DE4C1C1608517DDF" type="email">xingyue_liu@yahoo.com</mods:nameIdentifier>
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<mods:date id="44B01938837B8F8469C576F5FEA40EE6">2023</mods:date>
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<treatment id="78C00204201A547B84BDA59E80043721" ID-GBIF-Taxon="209451371" LSID="urn:lsid:zoobank.org:act:AA446547-70EC-4549-A734-76E156C4445B" httpUri="http://treatment.plazi.org/id/78C00204201A547B84BDA59E80043721" lastPageNumber="113" pageId="0" pageNumber="113" scope_family="Nevrorthidae" scope_order="Neuroptera" scope_superFamily="Osmyloidea">
<subSubSection id="09F1F1B63CCD63EF8B7E63035BC98900" pageId="0" pageNumber="113" type="nomenclature">
<paragraph id="664836A4ADF3EA4C2250FC941CA6B922" pageId="0" pageNumber="113">
Genus
<taxonomicName id="AEE6714BEA1A95A3C0415F4CCCC0835D" LSID="https://zoobank.org/AA446547-70EC-4549-A734-76E156C4445B" authority="Nakamine &amp; Yamamoto &amp; Takahashi &amp; Liu, 2023" authorityName="Nakamine &amp; Yamamoto &amp; Takahashi &amp; Liu" authorityYear="2023" class="Insecta" family="Nevrorthidae" genus="Sisyroneurorthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sisyroneurorthus" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="genus" status="gen. nov.">Sisyroneurorthus</taxonomicName>
<taxonomicNameLabel id="1E143CDC265E241988BFD689B97B0DA7" pageId="0" pageNumber="113">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="0F13225098B6F9985032A0D8D9EF2E39" pageId="0" pageNumber="113" type="description">
<paragraph id="9AC8B06B5526709023600BE5B0BE5DC3" pageId="0" pageNumber="113">
<figureCitation id="C2BF8F95CADBD786427E8208FC55410E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Sisyroneurorthus aspoeckorum gen. et sp. nov., holotype female (CAU-BA-NH- 22001) in mid-Cretaceous Kachin amber. A. Dorsal view; B. Left lateral view. Scale bars: 1.0 mm." figureDoi="10.3897/dez.70.98873.figure1" httpUri="https://binary.pensoft.net/fig/826132" pageId="0" pageNumber="113">Figs 1</figureCitation>
<figureCitation id="B9AADBDC1A1FFAA32F348DBFDE805C04" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Wing venation of Sisyroneurorthus aspoeckorum gen. et sp. nov., holotype female (CAU-BA-NH- 22001) in mid-Cretaceous Kachin amber. A. Right forewing; B. Right hind-wing. Abbreviations. A (a) - anal vein; A 1 - A 3 - first to third anal vein; Cu (cu) - cubitus; CuA - cubitus anterior; CuP - cubitus posterior; M (m) - media: MA - media anterior; MP - media posterior; R (r) - radius; RA - radius anterior; RP - radius posterior; Sc (sc) - subcosta; ScP - subcosta posterior. Scale bar: 0.5 mm." figureDoi="10.3897/dez.70.98873.figure2" httpUri="https://binary.pensoft.net/fig/826133" pageId="0" pageNumber="113">, 2</figureCitation>
<figureCitation id="13814FC3DD9394C20924ADF393397F3F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Sisyroneurorthus aspoeckorum gen. et sp. nov., holotype female (CAU-BA-NH- 22001) in mid-Cretaceous Kachin amber. A. Left anterolateral view of head and antenna; B. Left anterolateral view of mouthparts; C. Right anterolateral view of mouthparts; D. Distinct trichosors present on apical wing margin in right forewing; E. Obscured trichoshors on posterior margin in right forewing. Abbreviations. lp - labial palpus; mp - maxillary palpus. Scale bars: 0.2 mm (A-C); 0.1 mm (D, E)." figureDoi="10.3897/dez.70.98873.figure3" httpUri="https://binary.pensoft.net/fig/826134" pageId="0" pageNumber="113">, 3</figureCitation>
<figureCitation id="E18FB7AC258254F64E70079DD56636AA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Terminalia of Sisyroneurorthus aspoeckorum gen. et sp. nov., holotype female (CAU-BA-NH- 22001) in mid-Cretaceous Kachin amber. A. Left lateral view of terminalia; B. Left lateral view of terminalia with white line indicating each segment, gonapophyses and gonocoxite; C. Right ventrolateral view of terminalia; D. Right ventrolateral view of terminalia with white line indicating each segment and gonapophyses. Abbreviations. e - ectoproct; gp - gonapophyses; gx - gonocoxite; S - sternum; T - tergum. Scale bars: 0.2 mm." figureDoi="10.3897/dez.70.98873.figure4" httpUri="https://binary.pensoft.net/fig/826135" pageId="0" pageNumber="113">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="49DE52659DD011AB51F64370171C9BAB" pageId="0" pageNumber="113" type="type species">
<paragraph id="DA345D073F9205504DCEE788427840E8" pageId="0" pageNumber="113">Type species.</paragraph>
<paragraph id="6547F72C0B47B2929602CD5C434B3A20" pageId="0" pageNumber="113">
<taxonomicName id="3E97FDDDB4C9D9AA200168A929883BA2" authorityName="Nakamine &amp; Yamamoto &amp; Takahashi &amp; Liu" authorityYear="2023" class="Insecta" family="Nevrorthidae" genus="Sisyroneurorthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sisyroneurorthus aspoeckorum" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="aspoeckorum">
<emphasis id="E7D69CDDFA7E714C59A825767A0BE270" italics="true" pageId="0" pageNumber="113">Sisyroneurorthus aspoeckorum</emphasis>
</taxonomicName>
sp. nov., here designated.
</paragraph>
</subSubSection>
<subSubSection id="D0876187067D053EE741FE47154866AE" pageId="0" pageNumber="113" type="diagnosis">
<paragraph id="1D86EA72380CBB63FE30FD84362416D0" pageId="0" pageNumber="113">Diagnosis.</paragraph>
<paragraph id="D31F95F64A29399F45644F74976FA977" pageId="0" pageNumber="113">
<taxonomicName id="70B50B6FEE020BA91C011A12D98C9538" authorityName="Nakamine &amp; Yamamoto &amp; Takahashi &amp; Liu" authorityYear="2023" class="Insecta" family="Nevrorthidae" genus="Sisyroneurorthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sisyroneurorthus" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="CDA0281FED6FAAFA94DBF5E07D1E33E1" italics="true" pageId="0" pageNumber="113">Sisyroneurorthus</emphasis>
</taxonomicName>
gen. nov. can be distinguished from the other nevrorthid genera by the combination of the following characters. Forewing: costal crossveins simple; ScP fused with RA at distal fourth; CuA and CuP diverging well distad 1m-cu; cua-cup crossvein absent; a long oblique crossvein present between CuP and A1. Hind-wing: CuA with pectinate branching on distal half, all branches simple.
</paragraph>
</subSubSection>
<subSubSection id="BB5E38AD7ACD0FD25EC80B070932C8EF" pageId="0" pageNumber="113" type="species included">
<paragraph id="7EDF3CEDA7A2CD2DE6B264FF32105134" pageId="0" pageNumber="113">Species included.</paragraph>
<paragraph id="82CB9C207D30E70D4F8C82AF35E28528" pageId="0" pageNumber="113">
<taxonomicName id="4896F6B4BE874AB72809974C2F67E2EA" authorityName="Nakamine &amp; Yamamoto &amp; Takahashi &amp; Liu" authorityYear="2023" class="Insecta" family="Nevrorthidae" genus="Sisyroneurorthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sisyroneurorthus aspoeckorum" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="aspoeckorum">
<emphasis id="73E6E09AE73FC407721EF1D3F2ABB962" italics="true" pageId="0" pageNumber="113">Sisyroneurorthus aspoeckorum</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection id="FF7DB397311BAB5519387A6599D0DB0F" pageId="0" pageNumber="113" type="etymology">
<paragraph id="EFA50178FC9A25EC6F358360513990AF" pageId="0" pageNumber="113">Etymology.</paragraph>
<paragraph id="D00404BD559A4DF806F7314EFD37CA51" pageId="0" pageNumber="113">
The new generic name is a combination of &quot;
<taxonomicName id="18EB5547008EAF814FC427D4EA4DB129" authorityName="Burmeister" authorityYear="1839" class="Insecta" family="Sisyridae" genus="Sisyra" kingdom="Animalia" lsidName="Sisyra" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="85B1539D583568AD80489C9002ABAA78" italics="true" pageId="0" pageNumber="113">Sisyra</emphasis>
</taxonomicName>
&quot; (Greek, the name of the type genus of
<taxonomicName id="7F062F887DBF2718148D157C544652AC" class="Insecta" family="Sisyridae" kingdom="Animalia" lsidName="" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="family">Sisyridae</taxonomicName>
) and &quot;
<taxonomicName id="0EC3C02FC4936DEC9F952EBD76EF1E25" authorityName="Esben-Petersen" authorityYear="1929" class="Insecta" genus="Neurorthus" kingdom="Animalia" lsidName="Neurorthus" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="C9B426EBE39B8C3BEB4740771AEEEE06" italics="true" pageId="0" pageNumber="113">Neurorthus</emphasis>
</taxonomicName>
&quot; (Greek, the common suffix of the generic name of
<taxonomicName id="F18B5C9E37B3190EE9DF21849C3A0DA8" class="Insecta" family="Nevrorthidae" kingdom="Animalia" lsidName="" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="family">Nevrorthidae</taxonomicName>
) with reference to the mosaic characters with
<taxonomicName id="EACDA4A33C3289F085183461040B7F0E" class="Insecta" family="Sisyridae" kingdom="Animalia" lsidName="" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="family">Sisyridae</taxonomicName>
and
<taxonomicName id="BF200A4F0F49FDC18CE42FC8865FCDC6" class="Insecta" family="Nevrorthidae" kingdom="Animalia" lsidName="" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="family">Nevrorthidae</taxonomicName>
in the new genus.
</paragraph>
</subSubSection>
<subSubSection id="28771C1A3F8D55980A379C0A4068D152" pageId="0" pageNumber="113" type="remarks">
<paragraph id="7863EB8E405FFC8CE4D48DA2D5AE0889" pageId="0" pageNumber="113">Remarks.</paragraph>
<paragraph id="F6E5187EFC9F860595675D449B8986A8" pageId="0" pageNumber="113">
This new genus is placed in
<taxonomicName id="8783E39FD32501748FFF07F5ACCA62DB" class="Insecta" family="Nevrorthidae" kingdom="Animalia" lsidName="" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="family">Nevrorthidae</taxonomicName>
by the presence of three radial crossveins and the long distal half of CuA with 10 branches and nearly parallel with the posterior wing margin in the hind-wing. Interestingly, the new genus shares the distal fusion of the forewing ScP and RA due to the terminal curve of ScP with many species of
<taxonomicName id="184B8F001C901751B558A5F0496327EC" class="Insecta" family="Sisyridae" kingdom="Animalia" lsidName="" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="family">Sisyridae</taxonomicName>
.
<bibRefCitation id="27E32D1EEA8F8E9B0CA8CC7C87CE1A97" author="Wichard, W" journalOrPublisher="Kessel, Remagen" pageId="0" pageNumber="113" refId="B20" refString="Wichard, W, Groehn, C, Seredszus, F, 2009. Aquatic Insects in Baltic Amber. Kessel, Remagen" title="Aquatic Insects in Baltic Amber." year="2009">Wichard et al. (2009)</bibRefCitation>
use the fusion/separation of ScP and RA to distinguish
<taxonomicName id="D0ADDE68C0975A489D06887E788616D0" class="Insecta" family="Sisyridae" kingdom="Animalia" lsidName="" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="family">Sisyridae</taxonomicName>
and
<taxonomicName id="ABC8599177FA5A8985E6F9E575305092" class="Insecta" family="Nevrorthidae" kingdom="Animalia" lsidName="" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="family">Nevrorthidae</taxonomicName>
in their key to the aquatic
<taxonomicName id="36989DC43CEC5BE2EB263EDA2AB92C1C" authorityName="Esben-Petersen" authorityYear="1929" class="Insecta" kingdom="Animalia" lsidName="" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="order">Neuroptera</taxonomicName>
in the Eocene Baltic amber. However, the fusion of ScP and RA in some sisyrids, such as
<taxonomicName id="3B57866D4E5FAC7FAE60F34D2552C46F" authorityName="Banks" authorityYear="1939" class="Insecta" family="Sisyridae" genus="Sisyrina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sisyrina" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="A9E144C86157C1E715EB6B97F7018CB2" italics="true" pageId="0" pageNumber="113">Sisyrina</emphasis>
</taxonomicName>
Banks, 1939,
<taxonomicName id="18F687665A14BCF75992281D1CAD0BE9" authorityName="Perkovsky &amp; Makarkin" authorityYear="2015" class="Insecta" family="Sisyridae" genus="Prosisyrina" kingdom="Animalia" lsidName="Prosisyrina" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="CCCD838E479C7A299A180AC973477754" italics="true" pageId="0" pageNumber="113">Prosisyrina</emphasis>
</taxonomicName>
Perkovsky &amp; Makarkin, 2015 and
<taxonomicName id="3AA6EF96DF752646DF33B337687A158A" class="Insecta" family="Sisyridae" genus="Paleosisyra" kingdom="Animalia" lsidName="Paleosisyra electrobaltica" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="electrobaltica">
<emphasis id="9C8303A7491B738654F5AC8DC1665229" italics="true" pageId="0" pageNumber="113">Paleosisyra electrobaltica</emphasis>
</taxonomicName>
Wichard,
<normalizedToken id="F5098C93354FB8920E981707DADBB1A6" originalValue="Gröhn">Groehn</normalizedToken>
&amp; Seredszus, 2009 is atypical because the apical part of ScP is abruptly curved towards RA and the short terminal section connecting to RA can be also interpreted as a crossvein (e.g. 2scp-r in
<bibRefCitation id="60A11E009BCF13803C7D8CC9ABDC379C" DOI="https://doi.org/10.1016/j.cretres.2016.03.010" author="Makarkin, VN" journalOrPublisher="Cretaceous Research" pageId="0" pageNumber="113" pagination="170 - 176" refId="B14" refString="Makarkin, VN, Perkovsky, EE, 2016. An interesting new species of Sisyridae (Neuroptera) from the Upper Cretaceous Taimyr amber. Cretaceous Research 63: 170 - 176, DOI: https://doi.org/10.1016/j.cretres.2016.03.010" title="An interesting new species of Sisyridae (Neuroptera) from the Upper Cretaceous Taimyr amber." url="https://doi.org/10.1016/j.cretres.2016.03.010" volume="63" year="2016">Makarkin and Perkovsky (2016)</bibRefCitation>
or 2scp-ra in
<bibRefCitation id="30694F56C8C6632A63593BF3E481F943" DOI="https://doi.org/10.11646/zootaxa.5052.4.5" author="Yang, Y" journalOrPublisher="Zootaxa" pageId="0" pageNumber="113" pagination="552 - 566" refId="B23" refString="Yang, Y, Liu, X, 2021. New spongillaflies of the genus Sisyrina Banks, 1939 (Neuroptera: Sisyridae) from the Oriental faunal region. Zootaxa 5052 (4): 552 - 566, DOI: https://doi.org/10.11646/zootaxa.5052.4.5" title="New spongillaflies of the genus Sisyrina Banks, 1939 (Neuroptera: Sisyridae) from the Oriental faunal region." url="https://doi.org/10.11646/zootaxa.5052.4.5" volume="5052" year="2021">Yang and Liu (2021)</bibRefCitation>
). This feature is just the state present in
<taxonomicName id="8893647AE20E0AC4FCD26FFD7C2316E3" authorityName="Nakamine &amp; Yamamoto &amp; Takahashi &amp; Liu" authorityYear="2023" class="Insecta" family="Nevrorthidae" genus="Sisyroneurorthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sisyroneurorthus" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="872E030661C9ED502B10077EE79B00D6" italics="true" pageId="0" pageNumber="113">Sisyroneurorthus</emphasis>
</taxonomicName>
gen. nov. and should not be used to place the new genus in
<taxonomicName id="6227A1712F3D79C9ADD8CEDD148BDBA5" class="Insecta" family="Sisyridae" kingdom="Animalia" lsidName="" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="family">Sisyridae</taxonomicName>
. Moreover, all known sisyrids have only two radial crossveins, but lack 1ra-rp in the hind-wing. They also have fewer (usually five or six) branches of hind-wing CuA.
</paragraph>
<paragraph id="3129EA1081308A534C7BA46455990B55" pageId="0" pageNumber="113">
The new genus is most similar to the contemporary genus
<taxonomicName id="A92478E81563C5846CD2EE86DF761ADE" authorityName="Wichard" authorityYear="2017" class="Insecta" family="Nevrorthidae" genus="Cretarophalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cretarophalis" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="6C4E4D7EF49110E0E25AA0D88231EF58" italics="true" pageId="0" pageNumber="113">Cretarophalis</emphasis>
</taxonomicName>
Wichard, 2017 by having simple forewing costal crossveins and simple hind-wing CuA branches. However, it differs from the latter genus by the configuration of ScP, which is terminally curved towards RA (in
<taxonomicName id="C52E73285D432F97CFA48487446D9B36" authorityName="Wichard" authorityYear="2017" class="Insecta" family="Nevrorthidae" genus="Cretarophalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cretarophalis" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="DF0846EB5D50827DBBC6BC8259D8DD34" italics="true" pageId="0" pageNumber="113">Cretarophalis</emphasis>
</taxonomicName>
, ScP runs straight terminally), the position of the crossvein between Cu and A1 (2cu-a1), which is distad separation of CuA and CuP (in
<taxonomicName id="CBD5F1E3F14C6237DDCF39ED58561341" authorityName="Wichard" authorityYear="2017" class="Insecta" family="Nevrorthidae" genus="Cretarophalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cretarophalis" order="Neuroptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="2B19BA1CA983921CDE6ABF0B00D55BAE" italics="true" pageId="0" pageNumber="113">Cretarophalis</emphasis>
</taxonomicName>
2cu-a1 proximad separation of CuA and CuP).
</paragraph>
</subSubSection>
</treatment>
</document>
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