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<document ID-DOI="http://dx.doi.org/10.3897/subtbiol.46.105798" ID-Pensoft-Pub="1314-2615-46-1" ID-Pensoft-UUID="73A944AECDDB5130BA73E57C7CB6D83F" ID-ZooBank="1A679D4370004DD1BA0D69DA2057401E" ModsDocID="1314-2615-46-1" checkinTime="1690300902025" checkinUser="pensoft" docAuthor="Huber, Bernhard A., Meng, Guanliang, Clark, Huon L. &amp; Cazanove, Gregory" docDate="2023" docId="74ABF9BFEDB95E2E9F1104932BE7F15E" docLanguage="en" docName="SubterBiol 46: 1-19" docOrigin="Subterranean Biology 46" docPubDate="2023-07-24" docSource="http://dx.doi.org/10.3897/subtbiol.46.105798" docTitle="Buitinga ifrit Huber &amp; Cazanove 2023, sp. nov." docType="treatment" docUuid="094155C6-26E5-47A3-8D88-AE84E1D7873D" docUuidSource="ZooBank" docVersion="1" id="73A944AECDDB5130BA73E57C7CB6D83F" lastPageNumber="1" masterDocId="73A944AECDDB5130BA73E57C7CB6D83F" masterDocTitle="First blind daddy long-legs spiders from Australia and Reunion (Araneae, Pholcidae)" masterLastPageNumber="19" masterPageNumber="1" pageNumber="1" updateTime="1690300902025" updateUser="pensoft">
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<mods:title id="2B1B3F80CADDE1E6C9E12D8F9EE63364">First blind daddy long-legs spiders from Australia and Reunion (Araneae, Pholcidae)</mods:title>
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<mods:name id="53653C119C91097E452A278780B316D3" type="personal">
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<mods:namePart id="2409F3B74DB777ADEE8885C9B24941C1">Huber, Bernhard A.</mods:namePart>
<mods:nameIdentifier id="3A7EADAECCA9C697FC3CFCABEC93C110" type="ORCID">https://orcid.org/0000-0002-7566-5424</mods:nameIdentifier>
<mods:affiliation id="B5C22BDB07FB1FC218DBC51BEF92B572">Zoological Research Museum Alexander Koenig, LIB, Bonn, Germany</mods:affiliation>
<mods:nameIdentifier id="9712F9FD749C65F3E559BFC3D9CC8613" type="email">b.huber@leibniz-zfmk.de</mods:nameIdentifier>
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<mods:namePart id="81BA24DC3BB303C0E7AB35A04B43B2B5">Meng, Guanliang</mods:namePart>
<mods:affiliation id="B46FDAEAB1446A49E00C89AAD8228125">Zoological Research Museum Alexander Koenig, LIB, Bonn, Germany</mods:affiliation>
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<mods:namePart id="95AD86654F8272C612EF19F51B2D045F">Clark, Huon L.</mods:namePart>
<mods:affiliation id="B4038824EA84930A36F9627B995D7C80">Bennelongia Environmental Consultants, 5 Bishop Street, Jolimont WA 6014, Australia &amp; Collections and Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia</mods:affiliation>
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<mods:namePart id="F4249BC92BCEB8BF3350C81306387979">Cazanove, Gregory</mods:namePart>
<mods:affiliation id="62ED5B6CDE32AF00E7071FB034A2A61A">Museum d'Histoire Naturelle, Saint Denis, Reunion</mods:affiliation>
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<mods:typeOfResource id="C7A579A1AFF6E690B55D3EE27C7BAD45">text</mods:typeOfResource>
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<mods:title id="65FEDD34730FA811DFA568619FA84CA4">Subterranean Biology</mods:title>
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<mods:part id="A568716A3F50BF92DB801C5098FF8A90">
<mods:date id="29C6AE2C9A37F258492D3BAC354EBB06">2023</mods:date>
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<mods:number id="654558F8A9518EA144EA972C53037AA9">2023-07-24</mods:number>
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<mods:number id="913AE79FE2F71110FAFD081813370290">46</mods:number>
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<mods:classification id="DC390625BB33D8FB5751990A2A742864">journal article</mods:classification>
<mods:identifier id="E46B391C1C2CDEB24CC003D7A5F039F1" type="DOI">http://dx.doi.org/10.3897/subtbiol.46.105798</mods:identifier>
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<subSection id="3D5C858A7BDF38F7ACB1F2477FE6242D" pageId="0" pageNumber="1" type="results">
<treatment id="74ABF9BFEDB95E2E9F1104932BE7F15E" LSID="urn:lsid:zoobank.org:act:094155C6-26E5-47A3-8D88-AE84E1D7873D" httpUri="http://treatment.plazi.org/id/74ABF9BFEDB95E2E9F1104932BE7F15E" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="5FD777659B138F869231302BEAC2DF88" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="31B5B1B139DF822216544556C90F7B5C" pageId="0" pageNumber="1">
<taxonomicName id="E51294746BDD79F48E262895A93C2BA0" LSID="https://zoobank.org/094155C6-26E5-47A3-8D88-AE84E1D7873D" authority="Huber &amp; Cazanove" authorityName="Huber &amp; Cazanove" authorityYear="2023" class="Arachnida" family="Pholcidae" genus="Buitinga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buitinga ifrit" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ifrit" status="sp. nov.">Buitinga ifrit Huber &amp; Cazanove</taxonomicName>
<taxonomicNameLabel id="3C15BB164B7B69CF06765CE1702C85D4" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="3DDB180B444A3F4B99EFD1C477D580AC" pageId="0" pageNumber="1" type="description">
<paragraph id="04BC786F756A7DB0C676FE086291FC8D" pageId="0" pageNumber="1">
<figureCitation id="26286DBB49BF97E573410F37B507BD9C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A male habitus, dorsal view B female abdomen, ventral view (arrow: darker median area) C, D cleared female genitalia, ventral and dorsal views. Abbreviations: ls, dark lateral structure; mp, median posterior process; sp, spinnerets. Scale bars: 0.5 mm (A); 0.2 mm (B); 0.1 mm (C, D)." figureDoi="10.3897/subtbiol.46.105798.figure3" httpUri="https://binary.pensoft.net/fig/883299" pageId="0" pageNumber="1">Figs 3</figureCitation>
<figureCitation id="01F78CF10320EBC892BB2709F0009A0F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A, B left male palp, prolateral and retrolateral views C, D male chelicerae, frontal view, and frontal male cheliceral apophyses in more detail E, F left tarsus and procursus, prolateral and retrolateral views G-I cleared female genitalia, dorsal view, and posterior median structure in two cleared specimens in more detail. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; f, femur; pa, patella; pr, procursus; ta, tarsus; ti, tibia; tr, trochanter. Scale bars: 0.2 mm (A, B, G); 0.1 mm (C, E, F); 0.05 mm (D, H, I)." figureDoi="10.3897/subtbiol.46.105798.figure4" httpUri="https://binary.pensoft.net/fig/883300" pageId="0" pageNumber="1">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F7DA1D4740CF58D3CC278ED124080677" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="E0D9142C83BE17097F688C8FB0111B85" pageId="0" pageNumber="1">
<taxonomicName id="8096690F33F5059B4E69669FF8831583" authority="Huber &amp; Meng &amp; Clark &amp; Cazanove, 2023" authorityName="Huber &amp; Meng &amp; Clark &amp; Cazanove" authorityYear="2023" class="Arachnida" family="Pholcidae" genus="Buitinga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buitinga" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus" status="sp. nov.">Buitinga</taxonomicName>
? sp. nov.
<normalizedToken id="6251807C54258E107DCBB3016F609D18" originalValue="“Reun1”">&quot;Reun1&quot;</normalizedToken>
:
<bibRefCitation id="C053FD09236B75BA7FDBA1EDFA30766C" DOI="http://doi.org/10.3897/subtbiol.26.26430" author="Huber, BA" journalOrPublisher="Subterranean Biology" pageId="0" pageNumber="1" pagination="1 - 18" refId="B30" refString="Huber, BA, 2018. Cave-dwelling pholcid spiders (Araneae, Pholcidae): a review. Subterranean Biology 26: 1 - 18, DOI: http://doi.org/10.3897/subtbiol.26.26430" title="Cave-dwelling pholcid spiders (Araneae, Pholcidae): a review." url="http://doi.org/10.3897/subtbiol.26.26430" volume="26" year="2018">Huber 2018</bibRefCitation>
: 4.
</paragraph>
</subSubSection>
<subSubSection id="20AE5FC801B3C2F9193106AA0DA94E39" pageId="0" pageNumber="1" type="type material">
<paragraph id="DB49553EBE5BB7973E92C8918BF1C1BE" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="24B95E46C8065C045F3080F4CC644DE6" pageId="0" pageNumber="1">
<normalizedToken id="27958D388C2DFAC67053A78C8479D031" originalValue="Réunion">Reunion</normalizedToken>
- Saint-Paul • ♂ holotype; Grotte (Caverne) de la Tortue;
<geoCoordinate id="DEDCA9872C45990028EFB0FF39D2D76C" degrees="21.0743" direction="south" orientation="latitude" precision="5" value="-21.0743">21.0743°S</geoCoordinate>
,
<geoCoordinate id="A3780BB3C52E8B16CBCD3FAAB561C82B" degrees="55.2491" direction="east" orientation="longitude" precision="5" value="55.2491">55.2491°E</geoCoordinate>
; ~220 m a.s.l.; 9 Mar. 1997; J.-C. Ledoux leg.; ZFMK (Ar 24187).
</paragraph>
</subSubSection>
<subSubSection id="F1FCBD033546D9759191868AEC9FBBAD" pageId="0" pageNumber="1" type="type locality">
<paragraph id="1DBB2C7B43B0258912DA8590B4377DFE" pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph id="2FA505BECAE8EB7F1D6E4E4AD3B40327" pageId="0" pageNumber="1">
<normalizedToken id="1AB663987C0D7DE7A3AB9AC535E28966" originalValue="Réunion">Reunion</normalizedToken>
, Saint-Paul, Grotte (Caverne) de la Tortue,
<geoCoordinate id="6BBFC313A542A5A6AEDF08C54598CFA6" degrees="21.0743" direction="south" orientation="latitude" precision="5" value="-21.0743">21.0743°S</geoCoordinate>
,
<geoCoordinate id="DCC0B8C96DE292B646488760EF7B477D" degrees="55.2491" direction="east" orientation="longitude" precision="5" value="55.2491">55.2491°E</geoCoordinate>
, ca 220 m a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="01D1B14711E92AB4FA9C081A3709A28F" pageId="0" pageNumber="1" type="type specimen">
<paragraph id="D72117A6D3414D7987E3094BBD2A65C4" pageId="0" pageNumber="1">Type specimen.</paragraph>
<paragraph id="F35110074178B8DDB914FD4880222D94" pageId="0" pageNumber="1">
Holotype male, in ethanol. Original label: &quot;St. Paul, grotte de la Tortue,
<taxonomicName id="D32DB79EBA43428C36EA2263AD9C8F5B" class="Arachnida" family="Pholcidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Pholcidae</taxonomicName>
(sur toile) 9/3/1997&quot;.
</paragraph>
</subSubSection>
<subSubSection id="F0CA3AA3DBCAB56C85C9DA0E291D2E22" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="14151260951DB9C053D36A5B9C826EEA" pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph id="D44A0A58D5C3DDDA776E0631ED879F4E" pageId="0" pageNumber="1">
<materialsCitation id="3559ADAEC7DFDCABA0148ED47992B906" collectingDate="1996-07-13" collectingDateMax="2007-02-24" collectingDateMin="1996-07-13" collectorName="Salle du Museum, C. Guillermet, S. Gasnier" country="Reunion" location="Saint-Paul" municipality="Salle du Museum" specimenCount="11" specimenCount-female="8" specimenCount-juvenile="2" specimenCount-male="1" typeStatus="paratypes">
<normalizedToken id="301AE3E57D18C1D8FE57DBCAC5155373" originalValue="Réunion">
<collectingCountry id="124BFD030193293EF841FD043CC0B2D8" name="Réunion">Reunion</collectingCountry>
</normalizedToken>
-
<location id="268446C54A6F60E388D4CDFD6EFCEBCC" LSID="urn:lsid:plazi:treatment:74ABF9BFEDB95E2E9F1104932BE7F15E:268446C54A6F60E388D4CDFD6EFCEBCC" country="Reunion" municipality="Salle du Museum" name="Saint-Paul">Saint-Paul</location>
<specimenCount id="61E71182BB9C0E24AD9DE1F368A63750" type="male">1 ♂</specimenCount>
<specimenCount id="14377AAA502953FE1F25D32B0B8F902A" type="female">6 ♀</specimenCount>
(+
<specimenCount id="69AB039E3A38A5F2FD26755DF25AE7AF" type="generic">1 juv.</specimenCount>
)
<typeStatus id="FB3AD9A915616957D1C982A99C2937C0">paratypes</typeStatus>
; same data as for holotype; MHNR •
<specimenCount id="4E43A8DFCBAF29F902606982130F4A18" type="female">1 ♀</specimenCount>
<typeStatus id="0284E2E485559E5278B592FBACB15FAF">paratype</typeStatus>
(+
<specimenCount id="7D5BFF3676FF59BF359FCB97A1341F03" type="generic">1 juv.</specimenCount>
); same locality as for
<typeStatus id="45A7A057CAD5A632A2B23FCE0DF90BAC">holotype</typeStatus>
,
<collectorName id="34D48767ABDB1CB801A5520D90F04F2E">
<collectingMunicipality id="C92C52B6E58BF8BDA81E79CE808E2659">
Salle du
<normalizedToken id="0A7E6C4708B108D65683F0FF61BB6C87" originalValue="Muséum">Museum</normalizedToken>
</collectingMunicipality>
</collectorName>
;
<collectingDate id="AE1E5F0D7536A64E634CC50D7D75E6B7" value="1996-07-13">13 Jul. 1996</collectingDate>
;
<collectorName id="2580CE4FE81C48EF110DFA6A3F53B4F5">C. Guillermet</collectorName>
leg.; between stones from scree which obstruct the bottom of the gallery,
<normalizedToken id="F75FCD8619DA315F7F2B35BCDBEC4AD0" originalValue="“2022.E.6.2”">&quot;2022.E.6.2&quot;</normalizedToken>
; ZFMK (Ar 24188) •
<specimenCount id="E81B645372E1D1E54CF80B96295E6D4B" type="female">1 ♀</specimenCount>
<typeStatus id="FBADACC266091E68D0CB4976EAB38ACF">paratype</typeStatus>
; same locality as for
<typeStatus id="69A9921AF507F403EC3E65B7953B92E0">holotype</typeStatus>
;
<collectorName id="FC85CBBE9C547C3300C2881222F1FF61">
Salle du
<normalizedToken id="7B8FA5C18A023A832167C38023F140E9" originalValue="Muséum">Museum</normalizedToken>
</collectorName>
;
<collectingDate id="EB9799398BF63AEF8900469CB9C3231B" value="2007-02-24">24 Feb. 2007</collectingDate>
;
<collectorName id="8E77E694A4EAA209B2A2B40A9077D0DA">S. Gasnier</collectorName>
leg.; on the ground,
<normalizedToken id="35CDAFE626D1F359FD2DE29A5D9C9012" originalValue="“CT/070124.004”">&quot;CT/070124.004&quot;</normalizedToken>
<normalizedToken id="B3F5D10181C1608AE7A064517EF79B98" originalValue="“2011.E.65.1”">&quot;2011.E.65.1&quot;</normalizedToken>
; MHNR
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="371C5E41DCE384D817524F798BC8103C" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="957311A99367E604E32139376D666881" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="94D7C756191CD6F93E5C7D449ED9DE51" pageId="0" pageNumber="1">
Distinguished from known congeners (and other
<taxonomicName id="729D3CA50DCFD00146E75CE4C28EE765" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
) by strong bifid process on male palpal trochanter (Fig.
<figureCitation id="CDA8DF979C7DDE222CF0E4A7BBEC4E5C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A, B left male palp, prolateral and retrolateral views C, D male chelicerae, frontal view, and frontal male cheliceral apophyses in more detail E, F left tarsus and procursus, prolateral and retrolateral views G-I cleared female genitalia, dorsal view, and posterior median structure in two cleared specimens in more detail. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; f, femur; pa, patella; pr, procursus; ta, tarsus; ti, tibia; tr, trochanter. Scale bars: 0.2 mm (A, B, G); 0.1 mm (C, E, F); 0.05 mm (D, H, I)." figureDoi="10.3897/subtbiol.46.105798.figure4" httpUri="https://binary.pensoft.net/fig/883300" pageId="0" pageNumber="1">4B</figureCitation>
), by details of procursus (Fig.
<figureCitation id="E4AA355DA2C5726A0593E6CB5923CA5F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A, B left male palp, prolateral and retrolateral views C, D male chelicerae, frontal view, and frontal male cheliceral apophyses in more detail E, F left tarsus and procursus, prolateral and retrolateral views G-I cleared female genitalia, dorsal view, and posterior median structure in two cleared specimens in more detail. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; f, femur; pa, patella; pr, procursus; ta, tarsus; ti, tibia; tr, trochanter. Scale bars: 0.2 mm (A, B, G); 0.1 mm (C, E, F); 0.05 mm (D, H, I)." figureDoi="10.3897/subtbiol.46.105798.figure4" httpUri="https://binary.pensoft.net/fig/883300" pageId="0" pageNumber="1">4E, F</figureCitation>
; short main branch curved towards dorsal and with deeply bifid tip, large and mostly membranous side branch originating from ventral side of main branch, with complex tip), genital bulb (Fig.
<figureCitation id="8FC75E67BAD7FF30112080783B847F62" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A, B left male palp, prolateral and retrolateral views C, D male chelicerae, frontal view, and frontal male cheliceral apophyses in more detail E, F left tarsus and procursus, prolateral and retrolateral views G-I cleared female genitalia, dorsal view, and posterior median structure in two cleared specimens in more detail. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; f, femur; pa, patella; pr, procursus; ta, tarsus; ti, tibia; tr, trochanter. Scale bars: 0.2 mm (A, B, G); 0.1 mm (C, E, F); 0.05 mm (D, H, I)." figureDoi="10.3897/subtbiol.46.105798.figure4" httpUri="https://binary.pensoft.net/fig/883300" pageId="0" pageNumber="1">4A</figureCitation>
; bulbal apophysis slender and weakly curved, with small process at basis, embolus cylindrical), male chelicerae (Fig.
<figureCitation id="B89906E63D4FC18E547A2BE9C6602E8A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A, B left male palp, prolateral and retrolateral views C, D male chelicerae, frontal view, and frontal male cheliceral apophyses in more detail E, F left tarsus and procursus, prolateral and retrolateral views G-I cleared female genitalia, dorsal view, and posterior median structure in two cleared specimens in more detail. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; f, femur; pa, patella; pr, procursus; ta, tarsus; ti, tibia; tr, trochanter. Scale bars: 0.2 mm (A, B, G); 0.1 mm (C, E, F); 0.05 mm (D, H, I)." figureDoi="10.3897/subtbiol.46.105798.figure4" httpUri="https://binary.pensoft.net/fig/883300" pageId="0" pageNumber="1">4C, D</figureCitation>
; pair of light proximal processes; pair of distal frontal apophyses with four modified hairs each), epigynum (Fig.
<figureCitation id="5119AD41679FFF8AF3A6793159AC9609" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A male habitus, dorsal view B female abdomen, ventral view (arrow: darker median area) C, D cleared female genitalia, ventral and dorsal views. Abbreviations: ls, dark lateral structure; mp, median posterior process; sp, spinnerets. Scale bars: 0.5 mm (A); 0.2 mm (B); 0.1 mm (C, D)." figureDoi="10.3897/subtbiol.46.105798.figure3" httpUri="https://binary.pensoft.net/fig/883299" pageId="0" pageNumber="1">3B</figureCitation>
; barely protruding, without pockets, distinctive median posterior process only visible in cleared specimens), and female internal genitalia (Figs
<figureCitation id="C164865B7E9D85E4A4D19F71D720A41B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A male habitus, dorsal view B female abdomen, ventral view (arrow: darker median area) C, D cleared female genitalia, ventral and dorsal views. Abbreviations: ls, dark lateral structure; mp, median posterior process; sp, spinnerets. Scale bars: 0.5 mm (A); 0.2 mm (B); 0.1 mm (C, D)." figureDoi="10.3897/subtbiol.46.105798.figure3" httpUri="https://binary.pensoft.net/fig/883299" pageId="0" pageNumber="1">3C, D</figureCitation>
,
<figureCitation id="E11D837C8351802805AD1F750275319E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A, B left male palp, prolateral and retrolateral views C, D male chelicerae, frontal view, and frontal male cheliceral apophyses in more detail E, F left tarsus and procursus, prolateral and retrolateral views G-I cleared female genitalia, dorsal view, and posterior median structure in two cleared specimens in more detail. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; f, femur; pa, patella; pr, procursus; ta, tarsus; ti, tibia; tr, trochanter. Scale bars: 0.2 mm (A, B, G); 0.1 mm (C, E, F); 0.05 mm (D, H, I)." figureDoi="10.3897/subtbiol.46.105798.figure4" httpUri="https://binary.pensoft.net/fig/883300" pageId="0" pageNumber="1">4G</figureCitation>
; oval pore plates; pair of dark lateral structures).
</paragraph>
<caption id="0CF659C7D280F4831D19F6100249D075" doi="10.3897/subtbiol.46.105798.figure3" httpUri="https://binary.pensoft.net/fig/883299" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
<paragraph id="B154D68BAB7DF1313BB78FBC5AC58A7C" pageId="0" pageNumber="1">
<emphasis id="6F37103C4287DB105D1AA722E91E92FF" bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
<taxonomicName id="D3C46C2085CB6F154A807886DD33D9B5" authorityName="Huber &amp; Cazanove" authorityYear="2023" class="Arachnida" family="Pholcidae" genus="Buitinga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buitinga ifrit" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ifrit">
<emphasis id="C0818C988923285EDC6180E4B6BBCD4E" italics="true" pageId="0" pageNumber="1">Buitinga ifrit</emphasis>
</taxonomicName>
Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187-88)
<emphasis id="723AC114940FD91E5ECFA93D13FCC1C1" bold="true" pageId="0" pageNumber="1">A</emphasis>
male habitus, dorsal view
<emphasis id="EEDB6DDCF4A9271895D549EF3A79D78B" bold="true" pageId="0" pageNumber="1">B</emphasis>
female abdomen, ventral view (arrow: darker median area)
<emphasis id="074BD44384479D9FE9588D5F640997D3" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
cleared female genitalia, ventral and dorsal views. Abbreviations: ls, dark lateral structure; mp, median posterior process; sp, spinnerets. Scale bars: 0.5 mm (
<emphasis id="8F6D8CE940E97E89078A0D94DCD5F569" bold="true" pageId="0" pageNumber="1">A</emphasis>
); 0.2 mm (
<emphasis id="5FD1B9BFB91903AD36BDE05F663E82A9" bold="true" pageId="0" pageNumber="1">B</emphasis>
); 0.1 mm (
<emphasis id="1ECA81CEC62CB5FE2C5592AA5BC9B32F" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
).
</paragraph>
</caption>
<caption id="C47802D18F78C43B6F1ADC858D07BA31" doi="10.3897/subtbiol.46.105798.figure4" httpUri="https://binary.pensoft.net/fig/883300" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph id="CACB10CC9453C1952A91FF451B785576" pageId="0" pageNumber="1">
<emphasis id="C7EA1CB1FEF8E66A81A1187ACAEC3557" bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
<taxonomicName id="D07AA0FCBF3E3304D252F650FC6FE1BE" authorityName="Huber &amp; Cazanove" authorityYear="2023" class="Arachnida" family="Pholcidae" genus="Buitinga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buitinga ifrit" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ifrit">
<emphasis id="DE1AF1E575BA68DBDEF39FA1BF08632C" italics="true" pageId="0" pageNumber="1">Buitinga ifrit</emphasis>
</taxonomicName>
Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187-88)
<emphasis id="ABC8C10A50B6956BEEB1902BFC13E1E4" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
left male palp, prolateral and retrolateral views
<emphasis id="AF4BDB32AAF98642EE0D9466AFD54BF8" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
male chelicerae, frontal view, and frontal male cheliceral apophyses in more detail
<emphasis id="3D7749FCF777C766A6EC1F3E61E93E98" bold="true" pageId="0" pageNumber="1">E, F</emphasis>
left tarsus and procursus, prolateral and retrolateral views
<emphasis id="7DCB1A2485E94D3E007038BC796F8BF5" bold="true" pageId="0" pageNumber="1">G-I</emphasis>
cleared female genitalia, dorsal view, and posterior median structure in two cleared specimens in more detail. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; f, femur; pa, patella; pr, procursus; ta, tarsus; ti, tibia; tr, trochanter. Scale bars: 0.2 mm (
<emphasis id="C65FB1303827CB7E1F5F6B3DB70DD31C" bold="true" pageId="0" pageNumber="1">A, B, G</emphasis>
); 0.1 mm (
<emphasis id="32CEB303250E66F735762783BB6FED98" bold="true" pageId="0" pageNumber="1">C, E, F</emphasis>
); 0.05 mm (
<emphasis id="D8854AE880A1C513E2B9F47C9CB0075E" bold="true" pageId="0" pageNumber="1">D, H, I</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="07094BA62A8993CBA7024573947107FE" pageId="0" pageNumber="1" type="description">
<paragraph id="7C0FBA667F645F51AF4FB59B7FA71273" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="D488408E6C874CBB7F7D9423406B86FB" pageId="0" pageNumber="1">
<emphasis id="CCEAB0D151221B5F966A204B115A20C6" bold="true" pageId="0" pageNumber="1">Male</emphasis>
(holotype).
<emphasis id="14664412EAEAA0B62FCA9129814E7F81" bold="true" pageId="0" pageNumber="1">
<emphasis id="8FCA72E567BCCCF6B412090C1D6BAAEE" italics="true" pageId="0" pageNumber="1">Measurements</emphasis>
.
</emphasis>
Total body length 2.0, carapace width 0.70. Leg 1: 18.5 (4.6 + 0.3 + 5.1 + 7.3 + 1.2); tibia 2: 3.1; tibia 3: 1.9; tibia 4: 2.8; tibia 1 L/d: 85.
</paragraph>
<paragraph id="EA4C58C9B8DB28BA3C44E0BCC6EAE525" pageId="0" pageNumber="1">
<emphasis id="3B735E5E6FB7A65ACC864AF7510A6E27" bold="true" italics="true" pageId="0" pageNumber="1">Color</emphasis>
(in ethanol). Entire specimen pale ochre to whitish (Fig.
<figureCitation id="335EE82BB92274B65C312C89C905B772" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A male habitus, dorsal view B female abdomen, ventral view (arrow: darker median area) C, D cleared female genitalia, ventral and dorsal views. Abbreviations: ls, dark lateral structure; mp, median posterior process; sp, spinnerets. Scale bars: 0.5 mm (A); 0.2 mm (B); 0.1 mm (C, D)." figureDoi="10.3897/subtbiol.46.105798.figure3" httpUri="https://binary.pensoft.net/fig/883299" pageId="0" pageNumber="1">3A</figureCitation>
).
</paragraph>
<paragraph id="099AB6D97DD78C55741D622B968C9089" pageId="0" pageNumber="1">
<emphasis id="30B2CC62E8F3EDDCE747CE757B87F258" bold="true" pageId="0" pageNumber="1">
<emphasis id="1C6B162D2D6BF619C5B8B2B9252F1C6D" italics="true" pageId="0" pageNumber="1">Body</emphasis>
.
</emphasis>
Habitus as in Fig.
<figureCitation id="168C6902F7D6306D95E72D72D28CAC4C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A male habitus, dorsal view B female abdomen, ventral view (arrow: darker median area) C, D cleared female genitalia, ventral and dorsal views. Abbreviations: ls, dark lateral structure; mp, median posterior process; sp, spinnerets. Scale bars: 0.5 mm (A); 0.2 mm (B); 0.1 mm (C, D)." figureDoi="10.3897/subtbiol.46.105798.figure3" httpUri="https://binary.pensoft.net/fig/883299" pageId="0" pageNumber="1">3A</figureCitation>
. Ocular area slightly raised, without any remnants of eye pigment and lenses. Without thoracic groove. Clypeus unmodified. Sternum wider than long (0.50/0.40), unmodified. Abdomen oval, conical at spinnerets.
</paragraph>
<paragraph id="1FF2365FEC85C8C7CB8409251BF304C6" pageId="0" pageNumber="1">
<emphasis id="AEA07C68C36ED76169D9801A0C0C7DB9" bold="true" pageId="0" pageNumber="1">
<emphasis id="0B6E27F4D71F052BE412D39E4821A746" italics="true" pageId="0" pageNumber="1">Chelicerae</emphasis>
.
</emphasis>
As in Fig.
<figureCitation id="0B27C09BB71041F39ADDF9BF34D189AE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A, B left male palp, prolateral and retrolateral views C, D male chelicerae, frontal view, and frontal male cheliceral apophyses in more detail E, F left tarsus and procursus, prolateral and retrolateral views G-I cleared female genitalia, dorsal view, and posterior median structure in two cleared specimens in more detail. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; f, femur; pa, patella; pr, procursus; ta, tarsus; ti, tibia; tr, trochanter. Scale bars: 0.2 mm (A, B, G); 0.1 mm (C, E, F); 0.05 mm (D, H, I)." figureDoi="10.3897/subtbiol.46.105798.figure4" httpUri="https://binary.pensoft.net/fig/883300" pageId="0" pageNumber="1">4C</figureCitation>
, with pair of light proximal processes and pair of distal frontal apophyses with four modified hairs each (Fig.
<figureCitation id="C0339ECE311C87C10C1750AFDB58F16A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A, B left male palp, prolateral and retrolateral views C, D male chelicerae, frontal view, and frontal male cheliceral apophyses in more detail E, F left tarsus and procursus, prolateral and retrolateral views G-I cleared female genitalia, dorsal view, and posterior median structure in two cleared specimens in more detail. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; f, femur; pa, patella; pr, procursus; ta, tarsus; ti, tibia; tr, trochanter. Scale bars: 0.2 mm (A, B, G); 0.1 mm (C, E, F); 0.05 mm (D, H, I)." figureDoi="10.3897/subtbiol.46.105798.figure4" httpUri="https://binary.pensoft.net/fig/883300" pageId="0" pageNumber="1">4D</figureCitation>
); without stridulatory files.
</paragraph>
<paragraph id="8B2E1C4950EBF0F4E68F3B4DD30F8532" pageId="0" pageNumber="1">
<emphasis id="A1F72779078CB916B901C67B99DB99FC" bold="true" pageId="0" pageNumber="1">
<emphasis id="BE28629B88A81A7CE0E0FB3095A27736" italics="true" pageId="0" pageNumber="1">Palps</emphasis>
.
</emphasis>
As in Fig.
<figureCitation id="24F67FB7EDE55D13B2914D89700B03B8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A, B left male palp, prolateral and retrolateral views C, D male chelicerae, frontal view, and frontal male cheliceral apophyses in more detail E, F left tarsus and procursus, prolateral and retrolateral views G-I cleared female genitalia, dorsal view, and posterior median structure in two cleared specimens in more detail. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; f, femur; pa, patella; pr, procursus; ta, tarsus; ti, tibia; tr, trochanter. Scale bars: 0.2 mm (A, B, G); 0.1 mm (C, E, F); 0.05 mm (D, H, I)." figureDoi="10.3897/subtbiol.46.105798.figure4" httpUri="https://binary.pensoft.net/fig/883300" pageId="0" pageNumber="1">4A, B</figureCitation>
; coxa unmodified, trochanter with strong bifid process, femur short, slightly widening distally, without process; tibia with two trichobothria; procursus (Fig.
<figureCitation id="A3E5DABC5C81D508921FAD379D0BA234" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A, B left male palp, prolateral and retrolateral views C, D male chelicerae, frontal view, and frontal male cheliceral apophyses in more detail E, F left tarsus and procursus, prolateral and retrolateral views G-I cleared female genitalia, dorsal view, and posterior median structure in two cleared specimens in more detail. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; f, femur; pa, patella; pr, procursus; ta, tarsus; ti, tibia; tr, trochanter. Scale bars: 0.2 mm (A, B, G); 0.1 mm (C, E, F); 0.05 mm (D, H, I)." figureDoi="10.3897/subtbiol.46.105798.figure4" httpUri="https://binary.pensoft.net/fig/883300" pageId="0" pageNumber="1">4E, F</figureCitation>
) complex, with short main branch curved towards dorsal and with deeply bifid tip, large and mostly membranous side branch originating from ventral side of main branch, with complex tip; genital bulb with slender and weakly curved bulbal apophysis with small process at basis, and cylindrical embolus with transparent fringes at tip.
</paragraph>
<paragraph id="6EEB6B50BC2F0223D89A180AA41E5A20" pageId="0" pageNumber="1">
<emphasis id="05CE083009C27AF6EF9E33833319486A" bold="true" pageId="0" pageNumber="1">
<emphasis id="67E19E257328D14801C35EA61901B964" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
.
</emphasis>
Apparently without spines and curved hairs (many hairs missing); few short vertical hairs; leg trichobothria and pseudosegments not seen.
</paragraph>
<paragraph id="E67E6CE8B5C92761FBDFE175064C3BBE" pageId="0" pageNumber="1">
<emphasis id="D7CD948AAB55C00A60695BD05776727C" bold="true" pageId="0" pageNumber="1">Variation</emphasis>
(male). Second male apparently indistinguishable; legs 1 missing.
</paragraph>
<paragraph id="44B13E208C766D709B6159A33DE56A41" pageId="0" pageNumber="1">
<emphasis id="6E0872E33412654D1A8FFE9A5260BC5A" bold="true" pageId="0" pageNumber="1">Female.</emphasis>
In general similar to male; total body length 1.8-2.0; carapace width 0.68-0.72; tibia 1 in seven females: 3.7-4.7 (mean 4.1). Epigynum (Fig.
<figureCitation id="AE7D44F89908AAF36AFA0FCCFBB5D85C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A male habitus, dorsal view B female abdomen, ventral view (arrow: darker median area) C, D cleared female genitalia, ventral and dorsal views. Abbreviations: ls, dark lateral structure; mp, median posterior process; sp, spinnerets. Scale bars: 0.5 mm (A); 0.2 mm (B); 0.1 mm (C, D)." figureDoi="10.3897/subtbiol.46.105798.figure3" httpUri="https://binary.pensoft.net/fig/883299" pageId="0" pageNumber="1">3B</figureCitation>
) main anterior plate slightly protruding, barely visible in ventral view, with darker median area, without pockets, with distinctive median posterior process (Fig.
<figureCitation id="CF984585C789F21201E63E0E1B45634D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A, B left male palp, prolateral and retrolateral views C, D male chelicerae, frontal view, and frontal male cheliceral apophyses in more detail E, F left tarsus and procursus, prolateral and retrolateral views G-I cleared female genitalia, dorsal view, and posterior median structure in two cleared specimens in more detail. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; f, femur; pa, patella; pr, procursus; ta, tarsus; ti, tibia; tr, trochanter. Scale bars: 0.2 mm (A, B, G); 0.1 mm (C, E, F); 0.05 mm (D, H, I)." figureDoi="10.3897/subtbiol.46.105798.figure4" httpUri="https://binary.pensoft.net/fig/883300" pageId="0" pageNumber="1">4H, I</figureCitation>
) only visible in cleared specimens. Internal genitalia (Figs
<figureCitation id="61E563DDDE7D977A24C677D92092B99E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A male habitus, dorsal view B female abdomen, ventral view (arrow: darker median area) C, D cleared female genitalia, ventral and dorsal views. Abbreviations: ls, dark lateral structure; mp, median posterior process; sp, spinnerets. Scale bars: 0.5 mm (A); 0.2 mm (B); 0.1 mm (C, D)." figureDoi="10.3897/subtbiol.46.105798.figure3" httpUri="https://binary.pensoft.net/fig/883299" pageId="0" pageNumber="1">3C, D</figureCitation>
,
<figureCitation id="094639D8F8A50F1098E3360CFC06477C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Buitinga ifrit Huber &amp; Cazanove, sp. nov., male holotype and female paratype, ZFMK (Ar 24187 - 88) A, B left male palp, prolateral and retrolateral views C, D male chelicerae, frontal view, and frontal male cheliceral apophyses in more detail E, F left tarsus and procursus, prolateral and retrolateral views G-I cleared female genitalia, dorsal view, and posterior median structure in two cleared specimens in more detail. Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; f, femur; pa, patella; pr, procursus; ta, tarsus; ti, tibia; tr, trochanter. Scale bars: 0.2 mm (A, B, G); 0.1 mm (C, E, F); 0.05 mm (D, H, I)." figureDoi="10.3897/subtbiol.46.105798.figure4" httpUri="https://binary.pensoft.net/fig/883300" pageId="0" pageNumber="1">4G</figureCitation>
) with barely visible oval pore plates and pair of dark lateral structures. Each ALS apparently with only two spigots (large widened spigot and long pointed spigot; checked in cleared abdomen).
</paragraph>
</subSubSection>
<subSubSection id="F7C779AA7455F0C160AC2AD9688E992C" pageId="0" pageNumber="1" type="etymology">
<paragraph id="0299A9C3BFFBA038D1F176AFAAC8BB8B" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="D60C9B97ACD86587BD788B0433D5C2D9" pageId="0" pageNumber="1">The species epithet refers to a demon in Islamic mythology that is often associated with the underworld; noun in apposition.</paragraph>
</subSubSection>
<subSubSection id="CE9312554C9DAF2035B12F2C8D131466" pageId="0" pageNumber="1" type="distribution">
<paragraph id="E642DEE389C51FC6BF80AFC4914C5DDE" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="F74D4EB7FDEDFD4B5D4E1D413F25FB29" pageId="0" pageNumber="1">Known from type locality only.</paragraph>
</subSubSection>
<subSubSection id="4D742B4C047C8F50F1EAA2C15FD7EA3A" pageId="0" pageNumber="1" type="natural history">
<paragraph id="2162B063AB5BFF56EDE96A21D4765232" pageId="0" pageNumber="1">Natural history.</paragraph>
<paragraph id="226BCF41EE9655D1178136F19B5C11B4" pageId="0" pageNumber="1">
The spiders were collected deep within the Grotte de la Tortue lava tube (Fig.
<figureCitation id="A434CD999F1B42118219DDE28674BE56" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epigean and hypogean habitats and collecting method A Hamersley Range in Western Australia; the area of the sampled boreholes B weighted net (arrow) above a borehole C weighted net used for scrape sampling D main entrance of the Grotte de La Tortue (photo T. Percheron, 2015) E gallery of the lost goat in Caverne de La Tortue (photo T. Percheron, 2022)." figureDoi="10.3897/subtbiol.46.105798.figure5" httpUri="https://binary.pensoft.net/fig/883301" pageId="0" pageNumber="1">5D, E</figureCitation>
). Some of the specimens were collected in the Salle du
<normalizedToken id="1A82B3381B5FAD34EF4B65EF0B3E1F45" originalValue="Muséum">Museum</normalizedToken>
, a section that is about 150 m from the cave entrance (see topographic map in
<bibRefCitation id="5F30CEDF6FFA7533781402A82A7AFD81" DOI="https://doi.org/10.1080/00379271.2003.10697384" author="Hoch, H" journalOrPublisher="Annales de la Societe entomologique de France" pageId="0" pageNumber="1" pagination="265 - 270" refId="B24" refString="Hoch, H, Bonfils, J, Reynaud, B, Attie, M, 2003. First record of troglobitic Hemiptera (Fulgoromorpha: Cixiidae) from La Reunion Island. Annales de la Societe entomologique de France 39: 265 - 270, DOI: https://doi.org/10.1080/00379271.2003.10697384" title="First record of troglobitic Hemiptera (Fulgoromorpha: Cixiidae) from La Reunion Island." url="https://doi.org/10.1080/00379271.2003.10697384" volume="39" year="2003">Hoch et al. 2003</bibRefCitation>
). They were found among rocks on the ground. For the other specimens, cave section and microhabitat are unknown. For a general characterization of the caves and cave fauna of
<normalizedToken id="BAF677F5A957C800B28CCD6B31FF75C9" originalValue="Réunion">Reunion</normalizedToken>
, see Discussion.
</paragraph>
<caption id="8AF126AC67592169977D45B4C1ECA18A" doi="10.3897/subtbiol.46.105798.figure5" httpUri="https://binary.pensoft.net/fig/883301" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
<paragraph id="F3FA1F1EE8D17A6F2E77127AC6732537" pageId="0" pageNumber="1">
<emphasis id="FAAFC63AED28C1698E478F32E7CAB1C0" bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
Epigean and hypogean habitats and collecting method
<emphasis id="D9317B21322BD69AE327C475B2AAAE63" bold="true" pageId="0" pageNumber="1">A</emphasis>
Hamersley Range in Western Australia; the area of the sampled boreholes
<emphasis id="BCCD13FF576ABCB48535F5A6D72F2FCA" bold="true" pageId="0" pageNumber="1">B</emphasis>
weighted net (arrow) above a borehole
<emphasis id="BB4FEDE9A11894B2233AE101B7B51AD3" bold="true" pageId="0" pageNumber="1">C</emphasis>
weighted net used for scrape sampling
<emphasis id="E6DBAAD1D203B56D231CA948943FCFD7" bold="true" pageId="0" pageNumber="1">D</emphasis>
main entrance of the Grotte de La Tortue (photo T. Percheron, 2015)
<emphasis id="216211EE1DD96158BC4C7B4118A17611" bold="true" pageId="0" pageNumber="1">E</emphasis>
gallery of the lost goat in Caverne de La Tortue (photo T. Percheron, 2022).
</paragraph>
</caption>
</subSubSection>
</treatment>
</subSection>
</document>