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<document id="162E28761AD073BFB4FE9F76374FE78B" ID-DOI="10.11646/zootaxa.4816.4.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3954685" ID-ZooBank="5154FD08-0AED-4826-A275-37AA9FBF4874" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1595340302449" checkinUser="plazi" docAuthor="Akhil, S. V., Divya, M. &amp; Sabu, K. Thomas" docDate="2020" docId="6B0687C5A662FFF3ADD3FD73E4869DB1" docLanguage="en" docName="zootaxa.4816.4.7.pdf" docOrigin="Zootaxa 4816 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Brachinus eucosmus Andrewes 1937" docType="treatment" docVersion="6" lastPageNumber="581" masterDocId="973FFFBDA667FFF6AD44FFAAE6649856" masterDocTitle="Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India" masterLastPageNumber="600" masterPageNumber="576" pageNumber="581" updateTime="1698783441694" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="215299B09408C9AEFFA822CF3AEF3890">Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India</mods:title>
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<mods:namePart id="F5392BF9546100E9ABF7BE84078D15FA">Akhil, S. V.</mods:namePart>
<mods:affiliation id="CB0E43AEC26B54CF097610084CEF2AEC">Entomology Research Lab, P. G. &amp; Research Department of Zoology, St. Josephs College (Autonomous), Devagiri, Calicut, Kerala, 673008, India</mods:affiliation>
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<mods:namePart id="FB48E3FB391EE7E3484FFB138972A7C9">Divya, M.</mods:namePart>
<mods:affiliation id="48EFDCA79F566ACC46EAE5FD1901E37F">Entomology Research Lab, P. G. &amp; Research Department of Zoology, St. Josephs College (Autonomous), Devagiri, Calicut, Kerala, 673008, India &amp; divyamanikandan 123 @ gmail. com; https: // orcid. org / 0000 - 0003 - 4594 - 5991</mods:affiliation>
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<mods:namePart id="EEB50747B3746FD60606D342920B513F">Sabu, K. Thomas</mods:namePart>
<mods:affiliation id="4D9ED2F2CEDC9D98BA4D1A1D380864B6">Entomology Research Lab, P. G. &amp; Research Department of Zoology, St. Josephs College (Autonomous), Devagiri, Calicut, Kerala, 673008, India &amp; sabukthomas @ gmail. com; https: // orcid. org / 0000 - 0002 - 8701 - 0484</mods:affiliation>
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<mods:date id="82F958AB008162B989231B506BAE2025">2020</mods:date>
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<paragraph id="E31036D3A662FFF3ADD3FD73E4059AA5" blockId="5.[151,609,729,791]" box="[151,609,729,756]" pageId="5" pageNumber="581">
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<taxonomicName id="24AF4D50A662FFF3ADD3FD73E4059AA5" ID-CoL="MT9D" authority="Andrewes, 1937" authorityName="Andrewes" authorityYear="1937" box="[151,609,729,756]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="eucosmus">
<emphasis id="D1DBEAC1A662FFF3ADD3FD73E4059AA5" bold="true" box="[151,609,729,756]" pageId="5" pageNumber="581">
<emphasis id="D1DBEAC1A662FFF3ADD3FD73E7F19AA5" bold="true" box="[151,405,729,756]" italics="true" pageId="5" pageNumber="581">Brachinus eucosmus</emphasis>
<bibRefCitation id="873E4B22A662FFF3ACDEFD73E4059AA5" author="Andrewes, H. E." box="[410,609,729,755]" pageId="5" pageNumber="581" pagination="561 - 576" refId="ref17835" refString="Andrewes, H. E. (1937) Papers on Oriental Carabidae-XXXII. Annals and Magazine of Natural History, Series 10, 20, 561 - 576. https: // doi. org / 10.1080 / 00222933708655387" type="journal article" year="1937">Andrewes, 1937</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="ABB56558A662FFF3ADD3FD57E6849B41" box="[151,224,765,791]" pageId="5" pageNumber="581" type="description">
<paragraph id="E31036D3A662FFF3ADD3FD57E6849B41" blockId="5.[151,609,729,791]" box="[151,224,765,791]" pageId="5" pageNumber="581">
<figureCitation id="7B942A56A662FFF3ADD3FD57E6849B41" box="[151,224,765,791]" captionStart="FIGURE 1" captionStartId="14.[151,250,1885,1910]" captionTargetBox="[241,1348,192,1860]" captionTargetId="figure@14.[230,1356,181,1860]" captionTargetPageId="14" captionText="FIGURE 1. Dorsal habitus of Indian Brachinus species: a) B. dryas Andrewes, 1936 (holotype, BMNH); b) B. inops Andrewes, 1932 (holotype, BMNH); c) B. hexagrammus Chaudoir, 1876 (syntype, MNHN); d) B. sexmaculatus Dejean, 1825 (syntype, MNHN); e) B. tetracolon Chuadoir, 1876 (syntype, MNHN); f) B. tetragrammus Chaudoir, 1876 (syntype, MNHN); g) B. pictus (Hope, 1833) (ZSIM); h) B. cinctellus Chaudoir, 1876 (syntype, MNHN); i) B. eucosmus Andrewes, 1937 (holotype, BMNH)" figureDoi="http://doi.org/10.5281/zenodo.3954687" httpUri="https://zenodo.org/record/3954687/files/figure.png" pageId="5" pageNumber="581">Fig. 1i</figureCitation>
</paragraph>
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<paragraph id="E31036D3A662FFF3ADD3FCEFE58F9B09" blockId="5.[151,1003,836,863]" box="[151,1003,836,863]" pageId="5" pageNumber="581">
<treatmentCitationGroup id="C3BF11FDA662FFF3ADD3FCEFE58F9B09" box="[151,1003,836,863]" pageId="5" pageNumber="581">
<treatmentCitation id="620E10C2A662FFF3ADD3FCEFE4139B09" author="Andrewes, H. E." box="[151,631,836,863]" page="571" pageId="5" pageNumber="581" year="1937">
<taxonomicName id="24AF4D50A662FFF3ADD3FCEFE4139B09" ID-CoL="MT9D" authority="Andrewes, 1937: 571" authorityName="Andrewes" authorityPageNumber="571" authorityYear="1937" box="[151,631,836,863]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="eucosmus">
<emphasis id="D1DBEAC1A662FFF3ADD3FCEFE71A9B09" box="[151,382,837,863]" italics="true" pageId="5" pageNumber="581">Brachinus eucosmus</emphasis>
<bibRefCitation id="873E4B22A662FFF3ACC0FCEFE4139B09" author="Andrewes, H. E." box="[388,631,836,863]" pageId="5" pageNumber="581" pagination="561 - 576" refId="ref17835" refString="Andrewes, H. E. (1937) Papers on Oriental Carabidae-XXXII. Annals and Magazine of Natural History, Series 10, 20, 561 - 576. https: // doi. org / 10.1080 / 00222933708655387" type="journal article" year="1937">Andrewes, 1937: 571</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="620E10C2A662FFF3AFC7FCEFE5029B09" author="Hrdlicka, J." box="[643,870,836,863]" page="215" pageId="5" pageNumber="581" year="2003">
<bibRefCitation id="873E4B22A662FFF3AFC7FCEFE5029B09" author="Hrdlicka, J." box="[643,870,836,863]" pageId="5" pageNumber="581" pagination="212 - 218" refId="ref18818" refString="Hrdlicka, J. (2003) Brachininae. In: Lobl I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata- Myxophaga-Adephaga. Apollo Books, Stenstrup, pp. 212 - 218." type="book chapter" year="2003">Hrdlička, 2003: 215</bibRefCitation>
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;
<treatmentCitation id="620E10C2A662FFF3AE36FCEFE5839B09" author="Hrdlicka, J." box="[882,999,837,863]" page="476" pageId="5" pageNumber="581" year="2017">
<bibRefCitation id="873E4B22A662FFF3AE36FCEFE5839B09" author="Hrdlicka, J." box="[882,999,837,863]" pageId="5" pageNumber="581" pagination="471 - 480" refId="ref18865" refString="Hrdlicka, J. (2017) Brachininae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata- Myxophaga-Adephaga. Revised and Updated Edition. Brill, Leiden, pp. 471 - 480." type="book chapter" year="2017">2017: 476</bibRefCitation>
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.
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</paragraph>
</subSubSection>
<subSubSection id="ABB56558A662FFF3ADD3FC27E59A9C45" pageId="5" pageNumber="581" type="materials_examined">
<paragraph id="E31036D3A662FFF3ADD3FC27E2089BF1" blockId="5.[151,1437,908,1511]" box="[151,1132,908,935]" pageId="5" pageNumber="581">
<materialsCitation id="53C73C8EA662FFF3ADD3FC27E2089BF1" box="[151,1132,908,935]" collectorName="J. C. M. Gardner &amp; G. D. Bhasin." country="India" location="Uttarakhand" pageId="5" pageNumber="581" specimenCount="1" stateProvince="Uttarakhand" typeStatus="holotype">
<emphasis id="D1DBEAC1A662FFF3ADD3FC27E7529BF1" bold="true" box="[151,310,908,935]" pageId="5" pageNumber="581">
<typeStatus id="3C148871A662FFF3ADD3FC27E6B69BF1" box="[151,210,909,935]" pageId="5" pageNumber="581">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="9BB87643A662FFF3AC78FC27E71E9BF1" box="[316,378,909,935]" name="India" pageId="5" pageNumber="581">India</collectingCountry>
:
<collectingRegion id="216BF831A662FFF3ACC1FC27E4709BF1" box="[389,532,909,935]" country="India" name="Uttarakhand" pageId="5" pageNumber="581">Uttarakhand</collectingRegion>
: Dehra Dun, coll.
<collectorName id="4E5A5305A662FFF3AF9BFC27E5F89BF1" box="[735,924,909,935]" pageId="5" pageNumber="581">J. C. M. Gardner</collectorName>
and
<collectorName id="4E5A5305A662FFF3AE96FC27E2089BF1" box="[978,1132,909,935]" pageId="5" pageNumber="581">G. D. Bhasin.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="E31036D3A662FFF3AD83FC1BE78B9B9D" blockId="5.[151,1437,908,1511]" box="[199,495,944,971]" pageId="5" pageNumber="581">
<materialsCitation id="53C73C8EA662FFF3AD83FC1BE78F9B9D" box="[199,491,944,971]" collectionCode="BMNH" pageId="5" pageNumber="581" specimenCount="1" typeStatus="holotype">
<emphasis id="D1DBEAC1A662FFF3AD83FC1BE7E89B9D" bold="true" box="[199,396,944,971]" pageId="5" pageNumber="581">
<typeStatus id="3C148871A662FFF3AD83FC1BE7669B9D" box="[199,258,945,971]" pageId="5" pageNumber="581">Type</typeStatus>
depository.
</emphasis>
<collectionCode id="85BEAE16A662FFF3ACD7FC1BE78F9B9D" box="[403,491,945,971]" pageId="5" pageNumber="581">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="E31036D3A662FFF3AD83FC7EE59A9C45" blockId="5.[151,1437,908,1511]" pageId="5" pageNumber="581">
<emphasis id="D1DBEAC1A662FFF3AD83FC7EE7D79BB9" bold="true" box="[199,435,980,1007]" pageId="5" pageNumber="581">Specimen examined</emphasis>
(n=1).
<typeStatus id="3C148871A662FFF3AF41FC7FE4099BB9" box="[517,621,981,1007]" pageId="5" pageNumber="581" type="holotype">Holotype</typeStatus>
labelled “Type, Dehra
<collectingRegion id="216BF831A662FFF3AE29FC7FE5C49BB9" box="[877,928,981,1007]" country="United Kingdom" name="Down" pageId="5" pageNumber="581">Dun</collectingRegion>
, U.P., coll. J. C. M. Gardner,
<date id="97111013A662FFF3A9A6FC7FE3069BB9" box="[1250,1378,981,1007]" pageId="5" pageNumber="581" value="1933-03-20">20.III.1933</date>
, Ex. Descented termite mound, 2328, det. H. E. Andrewes, Brit.Mus.1937.548.”
</paragraph>
</subSubSection>
<subSubSection id="ABB56558A662FFF3AD83FBB6E7099D95" pageId="5" pageNumber="581" type="description">
<paragraph id="E31036D3A662FFF3AD83FBB6E3629C0D" blockId="5.[151,1437,908,1511]" pageId="5" pageNumber="581">
<emphasis id="D1DBEAC1A662FFF3AD83FBB6E73D9C61" bold="true" box="[199,345,1052,1079]" pageId="5" pageNumber="581">Description.</emphasis>
Length 18.019.0 mm. Head and pronotum brownish red; base and lateral margin of pronotum brownish black; palpi, antennae and legs reddish brown; elytra black with three reddish yellow spots.
</paragraph>
<paragraph id="E31036D3A662FFF3AD83FBCFE7869CF5" blockId="5.[151,1437,908,1511]" pageId="5" pageNumber="581">Head convex; wide, shallow frontal foveae; surface finely punctate; eyes prominent; antennae stout, almost reaching the middle of elytra.</paragraph>
<paragraph id="E31036D3A662FFF3AD83FB07E43E9D65" blockId="5.[151,1437,908,1511]" pageId="5" pageNumber="581">Pronotum convex, cordate, wider than head, wider than long; surface finely punctate, becoming coarse along base; base as wide as apex; lateral bead narrow; disc laterally with anterior two-thirds strongly convex, posterior third strongly sinuate, hind angles acute, sharp, projecting laterally; median groove fine, deeper near base; transverse impression present, shallow, clear.</paragraph>
<paragraph id="E31036D3A662FFF3AD83FA97E7099D95" blockId="5.[151,1437,908,1511]" pageId="5" pageNumber="581">Elytra convex, dilated behind, longer than broad, humerus rounded; apex with a re-entrant angle and glabrous membranous border; intervals convex with fine scattered punctures; three reddish yellow spots present, first fairly large, placed near to humerus and rounded, second irregularly rounded and just behind mid region, third small, oblique, near apex.</paragraph>
</subSubSection>
<subSubSection id="ABB56558A662FFF3AD83FA66E4869DB1" box="[199,738,1484,1511]" pageId="5" pageNumber="581" type="distribution">
<paragraph id="E31036D3A662FFF3AD83FA66E4869DB1" blockId="5.[151,1437,908,1511]" box="[199,738,1484,1511]" pageId="5" pageNumber="581">
<emphasis id="D1DBEAC1A662FFF3AD83FA66E7059DB1" bold="true" box="[199,353,1484,1511]" pageId="5" pageNumber="581">Distribution.</emphasis>
<collectingCountry id="9BB87643A662FFF3AC23FA67E7DF9DB1" box="[359,443,1485,1511]" name="India" pageId="5" pageNumber="581">INDIA</collectingCountry>
:
<collectingRegion id="216BF831A662FFF3AC82FA67E4319DB1" box="[454,597,1485,1511]" country="India" name="Uttarakhand" pageId="5" pageNumber="581">Uttarakhand</collectingRegion>
: Dehra Dun.
</paragraph>
</subSubSection>
</treatment>
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