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<document id="BD284DDF71A614C23D2F53F2B72D23EB" ID-CLB-Dataset="30205" ID-DOI="10.11646/zootaxa.4413.2.1" ID-GBIF-Dataset="fa65180b-5ae2-4eb7-8616-1c131cfe7251" ID-ISSN="1175-5326" ID-Zenodo-Dep="1227343" ID-ZooBank="495CE6EB-4A83-4A05-8E42-A92889F1C1C4" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1524517321172" checkinUser="plazi" docAuthor="Givens, Donald R." docDate="2018" docId="652FF8637631433AFF79FCAC1AA15341" docLanguage="en" docName="zootaxa.4413.2.1.pdf" docOrigin="Zootaxa 4413 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ecclisomyia maculosa Banks 1907" docType="treatment" docVersion="7" lastPageNumber="240" masterDocId="9916801B762C4312FFEEFFDA19455079" masterDocTitle="The Nearctic Ecclisomyia species (Trichoptera: Limnephilidae)" masterLastPageNumber="259" masterPageNumber="201" pageNumber="230" updateTime="1698540793590" updateUser="ExternalLinkService">
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<mods:title id="7DB63797D736FA4C6E6F86C4FF610E16">The Nearctic Ecclisomyia species (Trichoptera: Limnephilidae)</mods:title>
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<treatment id="652FF8637631433AFF79FCAC1AA15341" ID-DOI="http://doi.org/10.5281/zenodo.5975597" ID-GBIF-Taxon="143653038" ID-Zenodo-Dep="5975597" LSID="urn:lsid:plazi:treatment:652FF8637631433AFF79FCAC1AA15341" httpUri="http://treatment.plazi.org/id/652FF8637631433AFF79FCAC1AA15341" lastPageId="40" lastPageNumber="240" pageId="29" pageNumber="230">
<subSubSection id="A59C1AFE7631430FFF79FCAC1B7953E8" box="[151,572,886,913]" pageId="29" pageNumber="230" type="nomenclature">
<paragraph id="ED3949757631430FFF79FCAC1B7953E8" blockId="29.[151,1273,886,946]" box="[151,572,886,913]" pageId="29" pageNumber="230">
<heading id="B671FE197631430FFF79FCAC1B7953E8" bold="true" box="[151,572,886,913]" fontSize="11" level="1" pageId="29" pageNumber="230" reason="1">
<taxonomicName id="2A8632F67631430FFF79FCAC1B7953E8" ID-CoL="6DTQ6" authority="Banks 1907" authorityName="Banks" authorityYear="1907" box="[151,572,886,913]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF295677631430FFF79FCAC1B7953E8" bold="true" box="[151,572,886,913]" pageId="29" pageNumber="230">
<emphasis id="DFF295677631430FFF79FCAC18E053E9" bold="true" box="[151,421,886,912]" italics="true" pageId="29" pageNumber="230">Ecclisomyia maculosa</emphasis>
<bibRefCitation id="891734847631430FFE42FCAD1B7953E8" author="Banks, N." box="[428,572,887,913]" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="29" pageNumber="230" pagination="117 - 133" part="8" refId="ref35730" refString="Banks, N. (1907) Descriptions of new Trichoptera. Proceedings of the Entomological Society of Washington, 8, 117 - 133." title="Descriptions of new Trichoptera" type="journal article" year="1907">Banks 1907</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A59C1AFE7631430FFF79FC431DBC53CB" box="[151,1273,921,946]" pageId="29" pageNumber="230" type="description">
<paragraph id="ED3949757631430FFF79FC431DBC53CB" blockId="29.[151,1273,886,946]" box="[151,1273,921,946]" pageId="29" pageNumber="230">
<figureCitation id="75BD55F07631430FFF79FC43187253CB" box="[151,311,921,946]" captionStart="FIGURES22" captionStartId="30.[151,264,1920,1942]" captionTargetBox="[288,1334,357,1861]" captionTargetId="figure@30.[725,1341,1089,1765]" captionTargetPageId="30" captionText="FIGURES22a22c. Ecclisomyiamaculosa Banks, male genitalia.22a, left lateral; 22b,ventral; 22c,dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt= tergum X." httpUri="https://zenodo.org/record/1227383/files/figure.png" pageId="29" pageNumber="230">Figs. 22a22c</figureCitation>
; 23a23c; 24a24c; 25a25c; 26a, 26b; 27a, 27b; 28a, 28b; 29a29d; 39; 41; 42; 43c.
</paragraph>
</subSubSection>
<subSubSection id="A59C1AFE7631430FFF79FC3818FC54C9" pageId="29" pageNumber="230" type="reference_group">
<paragraph id="ED3949757631430FFF79FC3818FC54C9" blockId="29.[151,1436,994,1200]" pageId="29" pageNumber="230">
<taxonomicName id="2A8632F67631430FFF79FC38182B5381" box="[151,366,994,1016]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="maculos">
<emphasis id="DFF295677631430FFF79FC38182B5381" box="[151,366,994,1016]" italics="true" pageId="29" pageNumber="230">Ecclisomyia maculos</emphasis>
</taxonomicName>
a
<bibRefCitation id="891734847631430FFE6FFC3818B85381" author="Banks, N." box="[385,509,994,1016]" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="29" pageNumber="230" pagination="117 - 133" part="8" refId="ref35730" refString="Banks, N. (1907) Descriptions of new Trichoptera. Proceedings of the Entomological Society of Washington, 8, 117 - 133." title="Descriptions of new Trichoptera" type="journal article" year="1907">Banks 1907</bibRefCitation>
, 123, fig. 18, female, Boulder, Colorado, July 31, 1904 (Oslar), (description);
<bibRefCitation id="891734847631430FFACCFC391874544C" author="Ross, H. H." journalOrPublisher="Psyche" pageId="29" pageNumber="230" pagination="1 - 61" part="45" publicationUrl="https://doi.org/10.1155/1938/25928" refId="ref36863" refString="Ross, H. H. (1938) Lectotypes of North American caddisflies in the Museum of Comparative Zoology. Psyche, 45 (1), 1 - 61. https: // doi. org / 10.1155 / 1938 / 25928" title="Lectotypes of North American caddisflies in the Museum of Comparative Zoology" type="journal article" year="1938">Ross (1938, 31, 57, figs. 50, 50a, designation of Lectotype female, Boulder, Colorado and Allotype male, Slate Creek, Summit County, Colorado)</bibRefCitation>
;
<bibRefCitation id="891734847631430FFED4FBC51890544C" author="Denning, D. G." box="[314,469,1055,1077]" journalOrPublisher="Psyche" pageId="29" pageNumber="230" pagination="16 - 27" part="55" publicationUrl="https://doi.org/10.1155/1948/71426" refId="ref35958" refString="Denning, D. G. (1948) New and little known species of Nearctic Trichoptera. Psyche, 55 (1), 16 - 27. https: // doi. org / 10.1155 / 1948 / 71426" title="New and little known species of Nearctic Trichoptera" type="journal article" year="1948">Denning (1948</bibRefCitation>
, 18, figs. 3a, b, description male and female, bionomics);
<bibRefCitation id="891734847631430FFBF7FBFA1DCB544C" author="Ross, H. H." box="[1049,1166,1055,1077]" journalOrPublisher="American Midland Naturalist" pageId="29" pageNumber="230" pagination="410 - 429" part="43" publicationUrl="https://doi.org/10.2307/2421910" refId="ref36907" refString="Ross, H. H. (1950) Synoptic notes on some Nearctic limnephilid caddisflies (Trichoptera: Limnephilidae). American Midland Naturalist, 43 (2), 410 - 429. https: // doi. org / 10.2307 / 2421910" title="Synoptic notes on some Nearctic limnephilid caddisflies (Trichoptera: Limnephilidae)" type="journal article" year="1950">Ross (1950</bibRefCitation>
, 423, figs. 14, 14a, b. male genitalia, distribution);
<bibRefCitation id="891734847631430FFE56FBE71A50542A" author="Denning, D. G." box="[440,789,1085,1107]" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="29" pageNumber="230" pagination="157 - 162" part="24" refId="ref35999" refString="Denning, D. G. (1951) Records and descriptions of Nearctic caddis flies, Part III. Journal of the Kansas Entomological Society, 24 (4), 157 - 162." title="Records and descriptions of Nearctic caddis flies, Part III" type="journal article" year="1951">Denning (1951, 161, distribution)</bibRefCitation>
;
<bibRefCitation id="891734847631430FFCCEFBE71AF6542A" author="Nimmo, A." box="[800,947,1085,1107]" journalOrPublisher="Quaestiones Entomologicae" pageId="29" pageNumber="230" pagination="3 - 234" part="7" refId="ref36774" refString="Nimmo, A. (1971) The adult Rhyacophilidae and Limnephilidae (Trichoptera) of Alberta and Eastern British Columbia and their post-glacial origin. Quaestiones Entomologicae, 7 (1), 3 - 234." title="The adult Rhyacophilidae and Limnephilidae (Trichoptera) of Alberta and Eastern British Columbia and their post-glacial origin" type="journal article" year="1971">Nimmo (1971</bibRefCitation>
, 61, figs. 128a, b, 188193, 598, description of male and female genitalia, bionomics, distribution);
<bibRefCitation id="891734847631430FFD06FB861D02540B" author="Anderson, N. H." bookContentInfo="152 pp." box="[744,1095,1116,1138]" journalOrPublisher="Technical Bulletin 134. Agricultural Experiment Station, Oregon State University, Corvallis, Oregon" pageId="29" pageNumber="230" refId="ref35693" refString="Anderson, N. H. (1976) The distribution and biology of the Oregon Trichoptera. Technical Bulletin 134. Agricultural Experiment Station, Oregon State University, Corvallis, Oregon, 152 pp." title="The distribution and biology of the Oregon Trichoptera" type="book" year="1976">Anderson (1976, 80, distribution)</bibRefCitation>
;
<bibRefCitation id="891734847631430FFBBAFB86180254E8" author="Newell, R. L. &amp; Minshall, G. W." journalOrPublisher="Great Basin Naturalist" pageId="29" pageNumber="230" pagination="253 - 257" part="37" refId="ref36691" refString="Newell, R. L. &amp; Minshall, G. W. (1977) An annotated list of the aquatic insects of southeastern Idaho, part III Trichoptera. Great Basin Naturalist, 37 (2), 253 - 257." title="An annotated list of the aquatic insects of southeastern Idaho, part III Trichoptera" type="journal article" year="1977">Newell &amp; Minshall (1977, 255, distribution)</bibRefCitation>
;
<bibRefCitation id="891734847631430FFEBFFBA11A7A54E8" author="Swegman, B. G. &amp; Ferrington, L. C. Jr." box="[337,831,1147,1169]" journalOrPublisher="Great Basin Naturalist" pageId="29" pageNumber="230" pagination="287 - 291" part="40" refId="ref37188" refString="Swegman, B. G. &amp; Ferrington, L. C. Jr. (1980) New records of Western Trichoptera with notes on their biology. Great Basin Naturalist, 40 (3), 287 - 291." title="New records of Western Trichoptera with notes on their biology" type="journal article" year="1980">Swegman &amp; Ferrington (1980, 289, distribution)</bibRefCitation>
;
<bibRefCitation id="891734847631430FFCA7FBA61D6E54E8" author="Herrmann, S. J. &amp; Ruiter, D. E. &amp; Unzicker, J. D." box="[841,1067,1147,1169]" journalOrPublisher="The Southwestern Naturalist" pageId="29" pageNumber="230" pagination="4" part="31" publicationUrl="https://doi.org/10.2307/3671699" refId="ref36243" refString="Herrmann, S. J., Ruiter, D. E. &amp; Unzicker, J. D. (1986) Distribution and records of Colorado Trichoptera. The Southwestern Naturalist, 31 (4), 421 - - 457. https: // doi. org / 10.2307 / 3671699" title="Distribution and records of Colorado Trichoptera" type="journal article" year="1986">
Herrmann
<emphasis id="DFF295677631430FFC5BFBA61AAD54E8" box="[949,1000,1147,1169]" italics="true" pageId="29" pageNumber="230">et al.</emphasis>
(1986
</bibRefCitation>
, distribution);
<bibRefCitation id="891734847631430FFB51FBA618F054C9" author="Baumann, R. W. &amp; Unzicker, J. D." journalOrPublisher="Encyclia" pageId="29" pageNumber="230" pagination="27 - 29" part="58" refId="ref35796" refString="Baumann, R. W. &amp; Unzicker, J. D. (1981) Preliminary checklist of Utah caddisflies (Trichoptera). Encyclia, 58, 27 - 29." title="Preliminary checklist of Utah caddisflies (Trichoptera)" type="journal article" year="1981">Baumann &amp; Unzicker (1981, 27, distribution)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A59C1AFE7631430FFF79FB061D8E55D0" pageId="29" pageNumber="230" type="diagnosis">
<paragraph id="ED3949757631430FFF79FB061D8E55D0" blockId="29.[151,1437,1244,2025]" pageId="29" pageNumber="230">
<emphasis id="DFF295677631430FFF79FB061817548C" bold="true" box="[151,338,1244,1269]" pageId="29" pageNumber="230">Male diagnosis.</emphasis>
Each inferior appendages has 2 large, black, stout spines arising from its base (
<figureCitation id="75BD55F07631430FFB09FB061C10548C" box="[1255,1365,1244,1269]" captionStart="FIGURES22" captionStartId="30.[151,264,1920,1942]" captionTargetBox="[288,1334,357,1861]" captionTargetId="figure@30.[725,1341,1089,1765]" captionTargetPageId="30" captionText="FIGURES22a22c. Ecclisomyiamaculosa Banks, male genitalia.22a, left lateral; 22b,ventral; 22c,dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt= tergum X." httpUri="https://zenodo.org/record/1227383/files/figure.png" pageId="29" pageNumber="230">Figs. 22b</figureCitation>
,
<figureCitation id="75BD55F07631430FFA8CFB061CCA548C" box="[1378,1423,1244,1269]" captionStart="FIGURES23" captionStartId="31.[151,264,1918,1940]" captionTargetBox="[277,1313,188,1862]" captionTargetId="figure@31.[277,1295,188,1054]" captionTargetPageId="31" captionText="FIGURES23a23c. Ecclisomyiamaculosa Banks, male genitalia.23a, left lateral; 23b, ventral; 23c, dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); ph = phallus; phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt = tergum X." httpUri="https://zenodo.org/record/1227385/files/figure.png" pageId="29" pageNumber="230">23b</figureCitation>
), separating
<taxonomicName id="2A8632F67631430FFEFBFADB18E15561" authorityName="Banks" authorityYear="1907" box="[277,420,1280,1304]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF295677631430FFEFBFADB18E15561" box="[277,420,1280,1304]" italics="true" pageId="29" pageNumber="230">E. maculosa</emphasis>
</taxonomicName>
from
<taxonomicName id="2A8632F67631430FFE01FADB1B105561" authorityName="Denning" authorityYear="1951" box="[495,597,1280,1304]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="bilera">
<emphasis id="DFF295677631430FFE01FADB1B105561" box="[495,597,1280,1304]" italics="true" pageId="29" pageNumber="230">E. bilera</emphasis>
</taxonomicName>
, which lacks spines on the inferior appendages, and from
<taxonomicName id="2A8632F67631430FFAE9FADB1CD95561" box="[1287,1436,1281,1304]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="conspersa">
<emphasis id="DFF295677631430FFAE9FADB1CD95561" box="[1287,1436,1281,1304]" italics="true" pageId="29" pageNumber="230">E. conspersa</emphasis>
</taxonomicName>
which has a single black, stout spine on each inferior appendage (
<figureCitation id="75BD55F07631430FFC4AFAFE1D4D5544" box="[932,1032,1316,1341]" captionStart="FIGURES 10" captionStartId="15.[151,264,1918,1940]" captionTargetBox="[188,1383,239,1855]" captionTargetId="figure@15.[171,833,1146,1760]" captionTargetPageId="15" captionText="FIGURES 10a10c. Ecclisomyia conspersa Banks, male genitalia. 10a, left lateral; 10b, ventral; 10c, dorsal. Abbreviations: ifa= inferior appendage (paired); ima = intermediate appendage; IX= segment IX; par = parameres (paired); ph = phallus; phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt = tergum X." httpUri="https://zenodo.org/record/1227363/files/figure.png" pageId="29" pageNumber="230">Fig. 10a</figureCitation>
). The inferior appendages, viewed laterally, extend further caudad than the triangular inferior appendages of
<emphasis id="DFF295677631430FFC3DFA931D1F5519" box="[979,1114,1352,1376]" italics="true" pageId="29" pageNumber="230">
<taxonomicName id="2A8632F67631430FFC3DFA931D135519" baseAuthorityName="Givens" baseAuthorityYear="2014" box="[979,1110,1352,1376]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="simulata">E. simulata</taxonomicName>
.
</emphasis>
The shallow U- to V-shaped notch at the distal ends of the fused parameres of
<taxonomicName id="2A8632F67631430FFD3AFAB71A2555FC" authorityName="Banks" authorityYear="1907" box="[724,864,1389,1413]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF295677631430FFD3AFAB71A2555FC" box="[724,864,1389,1413]" italics="true" pageId="29" pageNumber="230">E. maculosa</emphasis>
</taxonomicName>
(
<figureCitation id="75BD55F07631430FFC9FFAB61A9A55FC" box="[881,991,1388,1413]" captionStart="FIGURES22" captionStartId="30.[151,264,1920,1942]" captionTargetBox="[288,1334,357,1861]" captionTargetId="figure@30.[725,1341,1089,1765]" captionTargetPageId="30" captionText="FIGURES22a22c. Ecclisomyiamaculosa Banks, male genitalia.22a, left lateral; 22b,ventral; 22c,dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt= tergum X." httpUri="https://zenodo.org/record/1227383/files/figure.png" pageId="29" pageNumber="230">Figs. 22b</figureCitation>
,
<figureCitation id="75BD55F07631430FFC02FAB61D5255FC" box="[1004,1047,1388,1413]" captionStart="FIGURES23" captionStartId="31.[151,264,1918,1940]" captionTargetBox="[277,1313,188,1862]" captionTargetId="figure@31.[277,1295,188,1054]" captionTargetPageId="31" captionText="FIGURES23a23c. Ecclisomyiamaculosa Banks, male genitalia.23a, left lateral; 23b, ventral; 23c, dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); ph = phallus; phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt = tergum X." httpUri="https://zenodo.org/record/1227385/files/figure.png" pageId="29" pageNumber="230">23b</figureCitation>
) distinguishes it from
<taxonomicName id="2A8632F67631430FFAF5FAB71CD955FC" baseAuthorityName="Givens" baseAuthorityYear="2014" box="[1307,1436,1389,1413]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="simulata">
<emphasis id="DFF295677631430FFAF5FAB71CD955FC" box="[1307,1436,1389,1413]" italics="true" pageId="29" pageNumber="230">E. simulata</emphasis>
</taxonomicName>
which has a deep U- to V-shaped notch at the distal end of the fused parameres (
<figureCitation id="75BD55F07631430FFBF7FA4A1DC055D0" box="[1049,1157,1424,1449]" captionStart="FIGURES 30" captionStartId="41.[151,264,1898,1920]" captionTargetBox="[245,1357,272,1795]" captionTargetId="figure@41.[275,1251,237,1043]" captionTargetPageId="41" captionText="FIGURES 30a30c. Ecclisomyia simulata Banks, male genitalia. 30a, left lateral; 30b, ventral; 30c, dorsal. Abbreviations: ifa = inferior appendage (paired); ima = intermediate appendage;IX = segment IX; par = paramere (fusedpair); ph = phallus; phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt = tergum X." httpUri="https://zenodo.org/record/1227399/files/figure.png" pageId="29" pageNumber="230">Figs. 30b</figureCitation>
,
<figureCitation id="75BD55F07631430FFB7FFA4A1DFB55D0" box="[1169,1214,1424,1449]" captionStart="FIGURES 31" captionStartId="42.[151,264,1934,1956]" captionTargetBox="[187,1384,241,1910]" captionTargetId="figure@42.[273,1348,241,1163]" captionTargetPageId="42" captionText="FIGURES 31a31c. Ecclisomyia simulata Banks, male genitalia. 31a, left lateral; 31b, ventral; 31c, dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); ph = phallus; phcr = phallocrypt; sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt= tergum VIII; Xt = tergum X." httpUri="https://zenodo.org/record/1227401/files/figure.png" pageId="29" pageNumber="230">31b</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A59C1AFE76314331FF29FA6E1BEE57BB" lastPageId="35" lastPageNumber="236" pageId="29" pageNumber="230" type="description">
<paragraph id="ED3949757631430CFF29FA6E1D8A5164" blockId="29.[151,1437,1244,2025]" lastBlockId="30.[151,1436,151,285]" lastPageId="30" lastPageNumber="231" pageId="29" pageNumber="230">
<emphasis id="DFF295677631430FFF29FA6E18DD55B4" bold="true" box="[199,408,1460,1485]" pageId="29" pageNumber="230">Male description.</emphasis>
See
<bibRefCitation id="891734847631430FFE20FA6E1B2F55B4" author="Banks, N." box="[462,618,1460,1485]" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="29" pageNumber="230" pagination="117 - 133" part="8" refId="ref35730" refString="Banks, N. (1907) Descriptions of new Trichoptera. Proceedings of the Entomological Society of Washington, 8, 117 - 133." title="Descriptions of new Trichoptera" type="journal article" year="1907">Banks (1907)</bibRefCitation>
;
<bibRefCitation id="891734847631430FFD9BFA6E1A6E55B4" author="Denning, D. G." box="[629,811,1460,1485]" journalOrPublisher="Psyche" pageId="29" pageNumber="230" pagination="16 - 27" part="55" publicationUrl="https://doi.org/10.1155/1948/71426" refId="ref35958" refString="Denning, D. G. (1948) New and little known species of Nearctic Trichoptera. Psyche, 55 (1), 16 - 27. https: // doi. org / 10.1155 / 1948 / 71426" title="New and little known species of Nearctic Trichoptera" type="journal article" year="1948">Denning (1948)</bibRefCitation>
; and
<bibRefCitation id="891734847631430FFC88FA6E1D5455B4" author="Nimmo, A." box="[870,1041,1460,1485]" journalOrPublisher="Quaestiones Entomologicae" pageId="29" pageNumber="230" pagination="3 - 234" part="7" refId="ref36774" refString="Nimmo, A. (1971) The adult Rhyacophilidae and Limnephilidae (Trichoptera) of Alberta and Eastern British Columbia and their post-glacial origin. Quaestiones Entomologicae, 7 (1), 3 - 234." title="The adult Rhyacophilidae and Limnephilidae (Trichoptera) of Alberta and Eastern British Columbia and their post-glacial origin" type="journal article" year="1971">Nimmo (1971)</bibRefCitation>
. Supplemental description follows: Length 811 mm (N = 29). Head, thorax, legs brown. Wings brown, hyaline with scattered pale areas, giving wings spotted appearance. Ocelli midway between eyes and midline. Antennae with about 4245 segments, dorsal surface of each segment with 4 stout black to reddish spines, except segment 1 (scape) and segment 2 with none. Antennal segments with numerous fine black setae; scape large, pedicel short, third antennal segment 2 X length of pedicel. Tergum VIII with row of transparent to reddish-brown setae across posterior area of tergum (
<figureCitation id="75BD55F07631430FFAC2F9B21CCA56F8" box="[1324,1423,1640,1665]" captionStart="FIGURES23" captionStartId="31.[151,264,1918,1940]" captionTargetBox="[277,1313,188,1862]" captionTargetId="figure@31.[277,1295,188,1054]" captionTargetPageId="31" captionText="FIGURES23a23c. Ecclisomyiamaculosa Banks, male genitalia.23a, left lateral; 23b, ventral; 23c, dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); ph = phallus; phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt = tergum X." httpUri="https://zenodo.org/record/1227385/files/figure.png" pageId="29" pageNumber="230">Fig. 23c</figureCitation>
). Sternum VIII brown, with long reddish setae on distal ventral margin and distal lateral surfaces; in lateral view narrow, longer than wide (
<figureCitation id="75BD55F07631430FFE2FF96A1B6856B0" box="[449,557,1712,1737]" captionStart="FIGURES22" captionStartId="30.[151,264,1920,1942]" captionTargetBox="[288,1334,357,1861]" captionTargetId="figure@30.[725,1341,1089,1765]" captionTargetPageId="30" captionText="FIGURES22a22c. Ecclisomyiamaculosa Banks, male genitalia.22a, left lateral; 22b,ventral; 22c,dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt= tergum X." httpUri="https://zenodo.org/record/1227383/files/figure.png" pageId="29" pageNumber="230">Figs. 22a</figureCitation>
,
<figureCitation id="75BD55F07631430FFDD7F96B1B2056B0" box="[569,613,1713,1737]" captionStart="FIGURES23" captionStartId="31.[151,264,1918,1940]" captionTargetBox="[277,1313,188,1862]" captionTargetId="figure@31.[277,1295,188,1054]" captionTargetPageId="31" captionText="FIGURES23a23c. Ecclisomyiamaculosa Banks, male genitalia.23a, left lateral; 23b, ventral; 23c, dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); ph = phallus; phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt = tergum X." httpUri="https://zenodo.org/record/1227385/files/figure.png" pageId="29" pageNumber="230">23a</figureCitation>
). Pleura well developed, membranous. Tergum IX nearly semicircular in lateral view, longest mesolaterally, longitudinally short dorsal and ventrally; dorsal, ventral, and anterior margins heavily sclerotized (
<figureCitation id="75BD55F07631430FFE94F922189A5768" box="[378,479,1784,1809]" captionStart="FIGURES22" captionStartId="30.[151,264,1920,1942]" captionTargetBox="[288,1334,357,1861]" captionTargetId="figure@30.[725,1341,1089,1765]" captionTargetPageId="30" captionText="FIGURES22a22c. Ecclisomyiamaculosa Banks, male genitalia.22a, left lateral; 22b,ventral; 22c,dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt= tergum X." httpUri="https://zenodo.org/record/1227383/files/figure.png" pageId="29" pageNumber="230">Figs 22a</figureCitation>
,
<figureCitation id="75BD55F07631430FFE02F9231B5D5768" box="[492,536,1785,1809]" captionStart="FIGURES23" captionStartId="31.[151,264,1918,1940]" captionTargetBox="[277,1313,188,1862]" captionTargetId="figure@31.[277,1295,188,1054]" captionTargetPageId="31" captionText="FIGURES23a23c. Ecclisomyiamaculosa Banks, male genitalia.23a, left lateral; 23b, ventral; 23c, dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); ph = phallus; phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt = tergum X." httpUri="https://zenodo.org/record/1227385/files/figure.png" pageId="29" pageNumber="230">23a</figureCitation>
); left and right lateral portions connected dorsally and ventrally with transverse sclerotized bridges (straps) (
<figureCitation id="75BD55F07631430FFE0FF8C61BCE574C" box="[481,651,1820,1845]" captionStart="FIGURES22" captionStartId="30.[151,264,1920,1942]" captionTargetBox="[288,1334,357,1861]" captionTargetId="figure@30.[725,1341,1089,1765]" captionTargetPageId="30" captionText="FIGURES22a22c. Ecclisomyiamaculosa Banks, male genitalia.22a, left lateral; 22b,ventral; 22c,dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt= tergum X." httpUri="https://zenodo.org/record/1227383/files/figure.png" pageId="29" pageNumber="230">Figs. 22b, 22c</figureCitation>
,
<figureCitation id="75BD55F07631430FFD72F8C61A41574C" box="[668,772,1820,1845]" captionStart="FIGURES23" captionStartId="31.[151,264,1918,1940]" captionTargetBox="[277,1313,188,1862]" captionTargetId="figure@31.[277,1295,188,1054]" captionTargetPageId="31" captionText="FIGURES23a23c. Ecclisomyiamaculosa Banks, male genitalia.23a, left lateral; 23b, ventral; 23c, dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); ph = phallus; phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt = tergum X." httpUri="https://zenodo.org/record/1227385/files/figure.png" pageId="29" pageNumber="230">23b, 23c</figureCitation>
). Superior appendages divergent, nearly parallel-sided in lateral and dorsal views, rounded distally; with hair-like setae along lateral and distal margins (
<figureCitation id="75BD55F07631430FFB06F89A1C105720" box="[1256,1365,1856,1881]" captionStart="FIGURES22" captionStartId="30.[151,264,1920,1942]" captionTargetBox="[288,1334,357,1861]" captionTargetId="figure@30.[725,1341,1089,1765]" captionTargetPageId="30" captionText="FIGURES22a22c. Ecclisomyiamaculosa Banks, male genitalia.22a, left lateral; 22b,ventral; 22c,dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt= tergum X." httpUri="https://zenodo.org/record/1227383/files/figure.png" pageId="29" pageNumber="230">Figs. 22a</figureCitation>
,
<figureCitation id="75BD55F07631430FFA8DF89B1CD55720" box="[1379,1424,1857,1881]" captionStart="FIGURES23" captionStartId="31.[151,264,1918,1940]" captionTargetBox="[277,1313,188,1862]" captionTargetId="figure@31.[277,1295,188,1054]" captionTargetPageId="31" captionText="FIGURES23a23c. Ecclisomyiamaculosa Banks, male genitalia.23a, left lateral; 23b, ventral; 23c, dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); ph = phallus; phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt = tergum X." httpUri="https://zenodo.org/record/1227385/files/figure.png" pageId="29" pageNumber="230">23a</figureCitation>
). Intermediate appendage ventrad of tergum X sclerotized; distal margin with several elongate setae (
<figureCitation id="75BD55F07631430FFB00F8BE1C1D5704" box="[1262,1368,1892,1917]" captionStart="FIGURES22" captionStartId="30.[151,264,1920,1942]" captionTargetBox="[288,1334,357,1861]" captionTargetId="figure@30.[725,1341,1089,1765]" captionTargetPageId="30" captionText="FIGURES22a22c. Ecclisomyiamaculosa Banks, male genitalia.22a, left lateral; 22b,ventral; 22c,dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt= tergum X." httpUri="https://zenodo.org/record/1227383/files/figure.png" pageId="29" pageNumber="230">Figs. 22a</figureCitation>
,
<figureCitation id="75BD55F07631430FFA8DF8BF1CD55704" box="[1379,1424,1893,1917]" captionStart="FIGURES23" captionStartId="31.[151,264,1918,1940]" captionTargetBox="[277,1313,188,1862]" captionTargetId="figure@31.[277,1295,188,1054]" captionTargetPageId="31" captionText="FIGURES23a23c. Ecclisomyiamaculosa Banks, male genitalia.23a, left lateral; 23b, ventral; 23c, dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); ph = phallus; phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt = tergum X." httpUri="https://zenodo.org/record/1227385/files/figure.png" pageId="29" pageNumber="230">23a</figureCitation>
). Tergum X with short spike-like setae along lateral margins (
<figureCitation id="75BD55F07631430FFCDBF8521A9D57D8" box="[821,984,1928,1953]" captionStart="FIGURES22" captionStartId="30.[151,264,1920,1942]" captionTargetBox="[288,1334,357,1861]" captionTargetId="figure@30.[725,1341,1089,1765]" captionTargetPageId="30" captionText="FIGURES22a22c. Ecclisomyiamaculosa Banks, male genitalia.22a, left lateral; 22b,ventral; 22c,dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt= tergum X." httpUri="https://zenodo.org/record/1227383/files/figure.png" pageId="29" pageNumber="230">Figs. 22b, 22c</figureCitation>
); apicolateral margins flanged ventrally; in dorsal view shaped somewhat like a bowling pin (
<figureCitation id="75BD55F07631430FFD07F8761A0E57BC" box="[745,843,1964,1989]" captionStart="FIGURES22" captionStartId="30.[151,264,1920,1942]" captionTargetBox="[288,1334,357,1861]" captionTargetId="figure@30.[725,1341,1089,1765]" captionTargetPageId="30" captionText="FIGURES22a22c. Ecclisomyiamaculosa Banks, male genitalia.22a, left lateral; 22b,ventral; 22c,dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt= tergum X." httpUri="https://zenodo.org/record/1227383/files/figure.png" pageId="29" pageNumber="230">Fig. 22c</figureCitation>
). Apices of parameres fused, with shallow U-shaped notch apically forming 2 short prong-like caudal extensions (
<figureCitation id="75BD55F07631430FFCA2F80A1AFC5790" box="[844,953,2000,2025]" captionStart="FIGURES22" captionStartId="30.[151,264,1920,1942]" captionTargetBox="[288,1334,357,1861]" captionTargetId="figure@30.[725,1341,1089,1765]" captionTargetPageId="30" captionText="FIGURES22a22c. Ecclisomyiamaculosa Banks, male genitalia.22a, left lateral; 22b,ventral; 22c,dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt= tergum X." httpUri="https://zenodo.org/record/1227383/files/figure.png" pageId="29" pageNumber="230">Figs. 22b</figureCitation>
,
<figureCitation id="75BD55F07631430FFC28F80A1AB15790" box="[966,1012,2000,2025]" captionStart="FIGURES23" captionStartId="31.[151,264,1918,1940]" captionTargetBox="[277,1313,188,1862]" captionTargetId="figure@31.[277,1295,188,1054]" captionTargetPageId="31" captionText="FIGURES23a23c. Ecclisomyiamaculosa Banks, male genitalia.23a, left lateral; 23b, ventral; 23c, dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); ph = phallus; phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt = tergum X." httpUri="https://zenodo.org/record/1227385/files/figure.png" pageId="29" pageNumber="230">23b</figureCitation>
). Phallocrypt narrower anteriorly and caudally, wider mesally in lateral view (
<figureCitation id="75BD55F07632430CFDBDFF4D1BF850C9" box="[595,701,151,176]" captionStart="FIGURES22" captionStartId="30.[151,264,1920,1942]" captionTargetBox="[288,1334,357,1861]" captionTargetId="figure@30.[725,1341,1089,1765]" captionTargetPageId="30" captionText="FIGURES22a22c. Ecclisomyiamaculosa Banks, male genitalia.22a, left lateral; 22b,ventral; 22c,dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt= tergum X." httpUri="https://zenodo.org/record/1227383/files/figure.png" pageId="30" pageNumber="231">Figs. 22a</figureCitation>
,
<figureCitation id="75BD55F07632430CFD26FF421BB050C9" box="[712,757,152,176]" captionStart="FIGURES23" captionStartId="31.[151,264,1918,1940]" captionTargetBox="[277,1313,188,1862]" captionTargetId="figure@31.[277,1295,188,1054]" captionTargetPageId="31" captionText="FIGURES23a23c. Ecclisomyiamaculosa Banks, male genitalia.23a, left lateral; 23b, ventral; 23c, dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); ph = phallus; phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt = tergum X." httpUri="https://zenodo.org/record/1227385/files/figure.png" pageId="30" pageNumber="231">23a</figureCitation>
). Inferior appendages divergent, elongate, apices rounded; in ventral view lateral margins nearly parallel; distal areas with numerous long reddish setae (
<figureCitation id="75BD55F07632430CFB5AFF661C1050AC" box="[1204,1365,188,213]" captionStart="FIGURES22" captionStartId="30.[151,264,1920,1942]" captionTargetBox="[288,1334,357,1861]" captionTargetId="figure@30.[725,1341,1089,1765]" captionTargetPageId="30" captionText="FIGURES22a22c. Ecclisomyiamaculosa Banks, male genitalia.22a, left lateral; 22b,ventral; 22c,dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt= tergum X." httpUri="https://zenodo.org/record/1227383/files/figure.png" pageId="30" pageNumber="231">Figs 22a, 22b</figureCitation>
,
<figureCitation id="75BD55F07632430CFA8CFF661CCA50AD" box="[1378,1423,188,212]" captionStart="FIGURES23" captionStartId="31.[151,264,1918,1940]" captionTargetBox="[277,1313,188,1862]" captionTargetId="figure@31.[277,1295,188,1054]" captionTargetPageId="31" captionText="FIGURES23a23c. Ecclisomyiamaculosa Banks, male genitalia.23a, left lateral; 23b, ventral; 23c, dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); ph = phallus; phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt = tergum X." httpUri="https://zenodo.org/record/1227385/files/figure.png" pageId="30" pageNumber="231">23a</figureCitation>
); each with 2 large stout black to reddish-brown spines arising from base and extending caudad (
<figureCitation id="75BD55F07632430CFB3FFF051C7A5081" box="[1233,1343,223,248]" captionStart="FIGURES22" captionStartId="30.[151,264,1920,1942]" captionTargetBox="[288,1334,357,1861]" captionTargetId="figure@30.[725,1341,1089,1765]" captionTargetPageId="30" captionText="FIGURES22a22c. Ecclisomyiamaculosa Banks, male genitalia.22a, left lateral; 22b,ventral; 22c,dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt= tergum X." httpUri="https://zenodo.org/record/1227383/files/figure.png" pageId="30" pageNumber="231">Figs. 22b</figureCitation>
,
<figureCitation id="75BD55F07632430CFAA5FF051C3C5081" box="[1355,1401,223,248]" captionStart="FIGURES23" captionStartId="31.[151,264,1918,1940]" captionTargetBox="[277,1313,188,1862]" captionTargetId="figure@31.[277,1295,188,1054]" captionTargetPageId="31" captionText="FIGURES23a23c. Ecclisomyiamaculosa Banks, male genitalia.23a, left lateral; 23b, ventral; 23c, dorsal. Abbreviations:ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); ph = phallus; phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt = tergum X." httpUri="https://zenodo.org/record/1227385/files/figure.png" pageId="30" pageNumber="231">23b</figureCitation>
); 3 shorter, stout reddish to black spines along posterior mesal margin lateral of these 2 stout spines.
</paragraph>
<caption id="B9F919FD7632430CFF79F85A1CD957AD" httpUri="https://zenodo.org/record/1227383/files/figure.png" pageId="30" pageNumber="231" startId="30.[151,264,1920,1942]" targetBox="[288,1334,357,1861]" targetPageId="30">
<paragraph id="ED3949757632430CFF79F85A1CD957AD" blockId="30.[151,1436,1920,2004]" pageId="30" pageNumber="231">
<emphasis id="DFF295677632430CFF79F85A182E57EF" bold="true" box="[151,363,1920,1942]" pageId="30" pageNumber="231">FIGURES 22a22c.</emphasis>
<taxonomicName id="2A8632F67632430CFE9FF85A1BDE57EF" authority="Banks" authorityName="Banks" authorityYear="1907" box="[369,667,1920,1942]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF295677632430CFE9FF85A1B1657EF" box="[369,595,1920,1942]" italics="true" pageId="30" pageNumber="231">Ecclisomyia maculosa</emphasis>
Banks
</taxonomicName>
, male genitalia. 22a, left lateral; 22b, ventral; 22c, dorsal. Abbreviations: ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt = tergum X.
</paragraph>
</caption>
<caption id="B9F919FD7633430DFF79F8A419BB5789" httpUri="https://zenodo.org/record/1227385/files/figure.png" pageId="31" pageNumber="232" startId="31.[151,264,1918,1940]" targetBox="[277,1313,188,1862]" targetPageId="31">
<paragraph id="ED3949757633430DFF79F8A419BB5789" blockId="31.[151,1436,1918,2032]" pageId="31" pageNumber="232">
<emphasis id="DFF295677633430DFF79F8A4182E57ED" bold="true" box="[151,363,1918,1940]" pageId="31" pageNumber="232">FIGURES 23a23c.</emphasis>
<taxonomicName id="2A8632F67633430DFE9FF8A41BDE57ED" authority="Banks" authorityName="Banks" authorityYear="1907" box="[369,667,1918,1940]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF295677633430DFE9FF8A41B1657ED" box="[369,595,1918,1940]" italics="true" pageId="31" pageNumber="232">Ecclisomyia maculosa</emphasis>
Banks
</taxonomicName>
, male genitalia. 23a, left lateral; 23b, ventral; 23c, dorsal. Abbreviations: ifa = inferior appendage (paired); ima = intermediate appendage; IX = segment IX; par = parameres (fused pair); ph = phallus; phcr = phallocrypt; pl = pleuron (paired); sa = superior appendage (paired); VIIIs = sternum VIII; VIIIt = tergum VIII; Xt = tergum X.
</paragraph>
</caption>
<caption id="B9F919FD760C4332FF79F8E018A157D5" httpUri="https://zenodo.org/record/1227387/files/figure.png" pageId="32" pageNumber="233" startId="32.[151,264,1850,1872]" targetBox="[196,1404,269,1789]" targetPageId="32">
<paragraph id="ED394975760C4332FF79F8E018A157D5" blockId="32.[151,1437,1850,1964]" pageId="32" pageNumber="233">
<emphasis id="DFF29567760C4332FF79F8E0182E5729" bold="true" box="[151,363,1850,1872]" pageId="32" pageNumber="233">FIGURES 24a24c.</emphasis>
<taxonomicName id="2A8632F6760C4332FE9EF8E01BDE5729" authority="Banks" authorityName="Banks" authorityYear="1907" box="[368,667,1850,1872]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF29567760C4332FE9EF8E01B165729" box="[368,595,1850,1872]" italics="true" pageId="32" pageNumber="233">Ecclisomyia maculosa</emphasis>
Banks
</taxonomicName>
, female terminalia. 24a, left lateral; 24b, dorsal; 24c, ventral. Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; IXs = sternum IX; IXt = tergum IX; pl = pleuron (paired); spt.sc = spermathecal sclerite; ssc.o = spermathecal sclerite opening; VIIIs = sternum VIII; IXs = sternum IX; IXt = tergum IX; Xt = tergum X.
</paragraph>
</caption>
<caption id="B9F919FD760D4333FF79F8FC1AB25703" httpUri="https://zenodo.org/record/1227389/files/figure.png" pageId="33" pageNumber="234" startId="33.[151,262,1830,1852]" targetBox="[181,1427,253,1779]" targetPageId="33">
<paragraph id="ED394975760D4333FF79F8FC1AB25703" blockId="33.[151,1437,1830,1914]" pageId="33" pageNumber="234">
<emphasis id="DFF29567760D4333FF79F8FC18275745" bold="true" box="[151,354,1830,1852]" pageId="33" pageNumber="234">FIGURES 25a25c</emphasis>
.
<taxonomicName id="2A8632F6760D4333FE81F8FC1BD35745" authority="Banks" authorityName="Banks" authorityYear="1907" box="[367,662,1830,1852]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF29567760D4333FE81F8FC1B0B5745" box="[367,590,1830,1852]" italics="true" pageId="33" pageNumber="234">Ecclisomyia maculosa</emphasis>
Banks
</taxonomicName>
, female terminalia. 25a, left lateral; 25b, ventral; 25c, ventral. Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; spt.sc = spermathecal sclerite; ssc.o = spermathecal sclerite opening; VIIIs = sternum VIII; IXs = sternum IX; Xt = tergum X.
</paragraph>
</caption>
<caption id="B9F919FD760E4330FF79F8F41DF2573D" box="[151,1207,1838,1860]" httpUri="https://zenodo.org/record/1227391/files/figure.png" pageId="34" pageNumber="235" startId="34.[151,264,1838,1860]" targetBox="[279,1271,253,886]" targetPageId="34">
<paragraph id="ED394975760E4330FF79F8F41DF2573D" blockId="34.[151,1207,1838,1860]" box="[151,1207,1838,1860]" pageId="34" pageNumber="235">
<emphasis id="DFF29567760E4330FF79F8F41822573D" bold="true" box="[151,359,1838,1860]" pageId="34" pageNumber="235">FIGURES 26a, 26b</emphasis>
.
<taxonomicName id="2A8632F6760E4330FE9AF8F41BE5573D" authority="Banks" authorityName="Banks" authorityYear="1907" box="[372,672,1838,1860]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF29567760E4330FE9AF8F41B12573D" box="[372,599,1838,1860]" italics="true" pageId="34" pageNumber="235">Ecclisomyia maculosa</emphasis>
Banks
</taxonomicName>
, male pupa, apical process. 26a, dorsal; 26b, ventral.
</paragraph>
</caption>
<caption id="B9F919FD760E4330FF79F8971DF2571A" box="[151,1207,1869,1891]" httpUri="https://zenodo.org/record/1227393/files/figure.png" pageId="34" pageNumber="235" startId="34.[151,264,1869,1891]" targetBox="[239,1359,1036,1765]" targetPageId="34">
<paragraph id="ED394975760E4330FF79F8971DF2571A" blockId="34.[151,1207,1869,1891]" box="[151,1207,1869,1891]" pageId="34" pageNumber="235">
<emphasis id="DFF29567760E4330FF79F897182B571A" bold="true" box="[151,366,1869,1891]" pageId="34" pageNumber="235">FIGURES 27a, 27b.</emphasis>
<taxonomicName id="2A8632F6760E4330FE9AF8971BE5571A" authority="Banks" authorityName="Banks" authorityYear="1907" box="[372,672,1869,1891]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF29567760E4330FE9AF8971B12571A" box="[372,599,1869,1891]" italics="true" pageId="34" pageNumber="235">Ecclisomyia maculosa</emphasis>
Banks
</taxonomicName>
, male pupa, apical process. 27a, dorsal; 27b, ventral.
</paragraph>
</caption>
<caption id="B9F919FD760F4331FF79FB941D8F541D" box="[151,1226,1102,1124]" httpUri="https://zenodo.org/record/1227395/files/figure.png" pageId="35" pageNumber="236" targetBox="[222,1350,257,1012]" targetPageId="35">
<paragraph id="ED394975760F4331FF79FB941D8F541D" blockId="35.[151,1226,1102,1124]" box="[151,1226,1102,1124]" pageId="35" pageNumber="236">
<emphasis id="DFF29567760F4331FF79FB941822541D" bold="true" box="[151,359,1102,1124]" pageId="35" pageNumber="236">FIGURES 28a, 28b</emphasis>
.
<taxonomicName id="2A8632F6760F4331FE9AFB941BE5541D" authority="Banks" authorityName="Banks" authorityYear="1907" box="[372,672,1102,1124]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF29567760F4331FE9AFB941B12541D" box="[372,599,1102,1124]" italics="true" pageId="35" pageNumber="236">Ecclisomyia maculosa</emphasis>
Banks
</taxonomicName>
, female pupa, apical process. 28a, dorsal; 28b, ventral.
</paragraph>
</caption>
<paragraph id="ED394975760F4331FF29FB4B1B565620" blockId="35.[151,1436,1169,1986]" pageId="35" pageNumber="236">
<emphasis id="DFF29567760F4331FF29FB4B18DE54D3" bold="true" box="[199,411,1169,1194]" pageId="35" pageNumber="236">Female diagnosis.</emphasis>
The distal margin of tergum X has a V-shaped incision extending about 1/23/4 the length of the segment in dorsal view (
<figureCitation id="75BD55F0760F4331FDECFB6F1B3454B7" box="[514,625,1205,1230]" captionStart="FIGURES24" captionStartId="32.[151,264,1850,1872]" captionTargetBox="[196,1404,269,1789]" captionTargetId="figure@32.[181,1313,944,1693]" captionTargetPageId="32" captionText="FIGURES24a24c.Ecclisomyia maculosa Banks, female terminalia.24a, leftlateral; 24b, dorsal;24c, ventral.Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; IXs = sternum IX; IXt = tergum IX; pl = pleuron (paired); spt.sc = spermathecal sclerite; ssc.o = spermathecal sclerite opening; VIIIs = sternumVIII; IXs = sternum IX; IXt = tergum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227387/files/figure.png" pageId="35" pageNumber="236">Figs. 24b</figureCitation>
,
<figureCitation id="75BD55F0760F4331FD91FB6F1BE954B7" box="[639,684,1205,1230]" captionStart="FIGURES 25" captionStartId="33.[151,262,1830,1852]" captionTargetBox="[181,1427,253,1779]" captionTargetId="figure@33.[375,1427,934,1683]" captionTargetPageId="33" captionText="FIGURES 25a25c. Ecclisomyia maculosa Banks, female terminalia. 25a, left lateral; 25b, ventral; 25c, ventral. Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; spt.sc = spermathecal sclerite; ssc.o = spermathecalsclerite opening; VIIIs = sternum VIII; IXs =sternum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227389/files/figure.png" pageId="35" pageNumber="236">25b</figureCitation>
), whereas the distal margin of tergum X of
<taxonomicName id="2A8632F6760F4331FB42FB6C1C6B54B4" baseAuthorityName="Givens" baseAuthorityYear="2014" box="[1196,1326,1205,1229]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="simulata">
<emphasis id="DFF29567760F4331FB42FB6C1C6B54B4" box="[1196,1326,1205,1229]" italics="true" pageId="35" pageNumber="236">E. simulata</emphasis>
</taxonomicName>
has a Ushaped incision that divides the tergum into 2 lobes, the incision about 3/4 the length of the tergum. The external part of gonopod VIII in both species is large; however, in
<taxonomicName id="2A8632F6760F4331FCCBFB241AF5556C" authorityName="Banks" authorityYear="1907" box="[805,944,1277,1301]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF29567760F4331FCCBFB241AF5556C" box="[805,944,1277,1301]" italics="true" pageId="35" pageNumber="236">E. maculosa</emphasis>
</taxonomicName>
it is nearly as wide as long (
<figureCitation id="75BD55F0760F4331FB1AFB271C1A556F" box="[1268,1375,1277,1302]" captionStart="FIGURES24" captionStartId="32.[151,264,1850,1872]" captionTargetBox="[196,1404,269,1789]" captionTargetId="figure@32.[181,1313,944,1693]" captionTargetPageId="32" captionText="FIGURES24a24c.Ecclisomyia maculosa Banks, female terminalia.24a, leftlateral; 24b, dorsal;24c, ventral.Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; IXs = sternum IX; IXt = tergum IX; pl = pleuron (paired); spt.sc = spermathecal sclerite; ssc.o = spermathecal sclerite opening; VIIIs = sternumVIII; IXs = sternum IX; IXt = tergum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227387/files/figure.png" pageId="35" pageNumber="236">Figs. 24c</figureCitation>
,
<figureCitation id="75BD55F0760F4331FA85FB2719855543" captionStart="FIGURES 25" captionStartId="33.[151,262,1830,1852]" captionTargetBox="[181,1427,253,1779]" captionTargetId="figure@33.[375,1427,934,1683]" captionTargetPageId="33" captionText="FIGURES 25a25c. Ecclisomyia maculosa Banks, female terminalia. 25a, left lateral; 25b, ventral; 25c, ventral. Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; spt.sc = spermathecal sclerite; ssc.o = spermathecalsclerite opening; VIIIs = sternum VIII; IXs =sternum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227389/files/figure.png" pageId="35" pageNumber="236">25b, 25c</figureCitation>
) whereas in
<taxonomicName id="2A8632F6760F4331FEBBFAF818935540" baseAuthorityName="Givens" baseAuthorityYear="2014" box="[341,470,1313,1337]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="simulata">
<emphasis id="DFF29567760F4331FEBBFAF818935540" box="[341,470,1313,1337]" italics="true" pageId="35" pageNumber="236">E. simulata</emphasis>
</taxonomicName>
it is longer than wide (
<figureCitation id="75BD55F0760F4331FD08FAFB1AC75543" box="[742,898,1313,1338]" captionStart="FIGURES 32" captionStartId="43.[151,264,1816,1838]" captionTargetBox="[223,1358,246,1768]" captionTargetId="figure@43.[348,1154,223,1010]" captionTargetPageId="43" captionText="FIGURES 32a32c. Ecclisomyia simulata, female genitalia. 32a, ventral; 32b, ventral; 32c, left lateral. Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; spt.sc = spermathecal sclerite; ssc.o = spermathecal sclerite opening; VIIIs = sternum VIII; IXs = sternum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227403/files/figure.png" pageId="35" pageNumber="236">Fig. 32a, 32b</figureCitation>
); the caudal margin may be convex (
<figureCitation id="75BD55F0760F4331FAC3FAFB1CCA5543" box="[1325,1423,1313,1338]" captionStart="FIGURES24" captionStartId="32.[151,264,1850,1872]" captionTargetBox="[196,1404,269,1789]" captionTargetId="figure@32.[181,1313,944,1693]" captionTargetPageId="32" captionText="FIGURES24a24c.Ecclisomyia maculosa Banks, female terminalia.24a, leftlateral; 24b, dorsal;24c, ventral.Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; IXs = sternum IX; IXt = tergum IX; pl = pleuron (paired); spt.sc = spermathecal sclerite; ssc.o = spermathecal sclerite opening; VIIIs = sternumVIII; IXs = sternum IX; IXt = tergum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227387/files/figure.png" pageId="35" pageNumber="236">Fig. 24c</figureCitation>
), straight (
<figureCitation id="75BD55F0760F4331FF17FA9F18135527" box="[249,342,1349,1374]" captionStart="FIGURES 25" captionStartId="33.[151,262,1830,1852]" captionTargetBox="[181,1427,253,1779]" captionTargetId="figure@33.[375,1427,934,1683]" captionTargetPageId="33" captionText="FIGURES 25a25c. Ecclisomyia maculosa Banks, female terminalia. 25a, left lateral; 25b, ventral; 25c, ventral. Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; spt.sc = spermathecal sclerite; ssc.o = spermathecalsclerite opening; VIIIs = sternum VIII; IXs =sternum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227389/files/figure.png" pageId="35" pageNumber="236">Fig. 25c</figureCitation>
) or with a V-shaped concavity (
<figureCitation id="75BD55F0760F4331FD56FA9F1A5F5527" box="[696,794,1349,1374]" captionStart="FIGURES 25" captionStartId="33.[151,262,1830,1852]" captionTargetBox="[181,1427,253,1779]" captionTargetId="figure@33.[375,1427,934,1683]" captionTargetPageId="33" captionText="FIGURES 25a25c. Ecclisomyia maculosa Banks, female terminalia. 25a, left lateral; 25b, ventral; 25c, ventral. Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; spt.sc = spermathecal sclerite; ssc.o = spermathecalsclerite opening; VIIIs = sternum VIII; IXs =sternum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227389/files/figure.png" pageId="35" pageNumber="236">Fig. 25b</figureCitation>
). The external gonopod of segment IX sclerotized, caudal margin with deep V-shaped declivity extending posterad about 2 X its width beyond the caudal margin of the extended gonopod of segment VIII (
<figureCitation id="75BD55F0760F4331FDD4FA571BED55DF" box="[570,680,1421,1446]" captionStart="FIGURES24" captionStartId="32.[151,264,1850,1872]" captionTargetBox="[196,1404,269,1789]" captionTargetId="figure@32.[181,1313,944,1693]" captionTargetPageId="32" captionText="FIGURES24a24c.Ecclisomyia maculosa Banks, female terminalia.24a, leftlateral; 24b, dorsal;24c, ventral.Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; IXs = sternum IX; IXt = tergum IX; pl = pleuron (paired); spt.sc = spermathecal sclerite; ssc.o = spermathecal sclerite opening; VIIIs = sternumVIII; IXs = sternum IX; IXt = tergum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227387/files/figure.png" pageId="35" pageNumber="236">Figs. 24c</figureCitation>
,
<figureCitation id="75BD55F0760F4331FD5BFA571A5B55DF" box="[693,798,1421,1446]" captionStart="FIGURES 25" captionStartId="33.[151,262,1830,1852]" captionTargetBox="[181,1427,253,1779]" captionTargetId="figure@33.[375,1427,934,1683]" captionTargetPageId="33" captionText="FIGURES 25a25c. Ecclisomyia maculosa Banks, female terminalia. 25a, left lateral; 25b, ventral; 25c, ventral. Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; spt.sc = spermathecal sclerite; ssc.o = spermathecalsclerite opening; VIIIs = sternum VIII; IXs =sternum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227389/files/figure.png" pageId="35" pageNumber="236">25b, 25b</figureCitation>
); in
<taxonomicName id="2A8632F6760F4331FCBDFA541A9355DC" baseAuthorityName="Givens" baseAuthorityYear="2014" box="[851,982,1421,1445]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="simulata">
<emphasis id="DFF29567760F4331FCBDFA541A9355DC" box="[851,982,1421,1445]" italics="true" pageId="35" pageNumber="236">E. simulata</emphasis>
</taxonomicName>
it is wide, not as deeply cleft and only extends 1X its width posterad beyond the external part of gonopod VIII (
<figureCitation id="75BD55F0760F4331FC10FA6B1DE455B3" box="[1022,1185,1457,1482]" captionStart="FIGURES 32" captionStartId="43.[151,264,1816,1838]" captionTargetBox="[223,1358,246,1768]" captionTargetId="figure@43.[348,1154,223,1010]" captionTargetPageId="43" captionText="FIGURES 32a32c. Ecclisomyia simulata, female genitalia. 32a, ventral; 32b, ventral; 32c, left lateral. Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; spt.sc = spermathecal sclerite; ssc.o = spermathecal sclerite opening; VIIIs = sternum VIII; IXs = sternum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227403/files/figure.png" pageId="35" pageNumber="236">Figs 32a, 32b</figureCitation>
).The external part of gonopod VIII is massive (
<figureCitation id="75BD55F0760F4331FE27FA0F1B695597" box="[457,556,1493,1518]" captionStart="FIGURES 25" captionStartId="33.[151,262,1830,1852]" captionTargetBox="[181,1427,253,1779]" captionTargetId="figure@33.[375,1427,934,1683]" captionTargetPageId="33" captionText="FIGURES 25a25c. Ecclisomyia maculosa Banks, female terminalia. 25a, left lateral; 25b, ventral; 25c, ventral. Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; spt.sc = spermathecal sclerite; ssc.o = spermathecalsclerite opening; VIIIs = sternum VIII; IXs =sternum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227389/files/figure.png" pageId="35" pageNumber="236">Fig. 25b</figureCitation>
) whereas it is short and narrow in
<taxonomicName id="2A8632F6760F4331FC22FA0C1D775594" authorityName="Denning" authorityYear="1951" box="[972,1074,1493,1517]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="bilera">
<emphasis id="DFF29567760F4331FC22FA0C1D775594" box="[972,1074,1493,1517]" italics="true" pageId="35" pageNumber="236">E. bilera</emphasis>
</taxonomicName>
(
<figureCitation id="75BD55F0760F4331FBABFA0F1DD95597" box="[1093,1180,1493,1518]" captionStart="FIGURES 3" captionStartId="7.[151,264,1446,1468]" captionTargetBox="[169,1385,224,1392]" captionTargetId="figure@7.[509,1046,818,1298]" captionTargetPageId="7" captionText="FIGURES 3a3c. Ecclisomyia bilera Denning, female genitalia. 3a, left lateral; 3b, ventral; 3c, dorsal. Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; IXt = tergum IX; pl = pleuron (paired); spt.sc = spermathecal sclerite;ssc.o = spermathecal sclerite opening;VIIIs = sternumVIII; VIIIt= tergum VIII;Xt = tergum X." httpUri="https://zenodo.org/record/1227349/files/figure.png" pageId="35" pageNumber="236">Fig. 3b</figureCitation>
). The external part of gonopod IX has a deep mesal division (
<figureCitation id="75BD55F0760F4331FDBFFA231BF7566B" box="[593,690,1529,1554]" captionStart="FIGURES24" captionStartId="32.[151,264,1850,1872]" captionTargetBox="[196,1404,269,1789]" captionTargetId="figure@32.[181,1313,944,1693]" captionTargetPageId="32" captionText="FIGURES24a24c.Ecclisomyia maculosa Banks, female terminalia.24a, leftlateral; 24b, dorsal;24c, ventral.Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; IXs = sternum IX; IXt = tergum IX; pl = pleuron (paired); spt.sc = spermathecal sclerite; ssc.o = spermathecal sclerite opening; VIIIs = sternumVIII; IXs = sternum IX; IXt = tergum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227387/files/figure.png" pageId="35" pageNumber="236">Fig. 24c</figureCitation>
); in
<taxonomicName id="2A8632F6760F4331FD0DFA201A005668" authorityName="Denning" authorityYear="1951" box="[739,837,1529,1553]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="bilera">
<emphasis id="DFF29567760F4331FD0DFA201A005668" box="[739,837,1529,1553]" italics="true" pageId="35" pageNumber="236">E. bilera</emphasis>
</taxonomicName>
it is broadly concave as viewed from a ventral aspect (
<figureCitation id="75BD55F0760F4331FF71F9C71845564F" box="[159,256,1565,1590]" captionStart="FIGURES 3" captionStartId="7.[151,264,1446,1468]" captionTargetBox="[169,1385,224,1392]" captionTargetId="figure@7.[509,1046,818,1298]" captionTargetPageId="7" captionText="FIGURES 3a3c. Ecclisomyia bilera Denning, female genitalia. 3a, left lateral; 3b, ventral; 3c, dorsal. Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; IXt = tergum IX; pl = pleuron (paired); spt.sc = spermathecal sclerite;ssc.o = spermathecal sclerite opening;VIIIs = sternumVIII; VIIIt= tergum VIII;Xt = tergum X." httpUri="https://zenodo.org/record/1227349/files/figure.png" pageId="35" pageNumber="236">Figs. 3b</figureCitation>
,
<figureCitation id="75BD55F0760F4331FEE0F9C41869564F" box="[270,300,1566,1590]" captionStart="FIGURES 4" captionStartId="8.[151,264,1706,1728]" captionTargetBox="[211,1402,191,1677]" captionTargetId="figure@8.[221,1394,191,998]" captionTargetPageId="8" captionText="FIGURES 4a4c. Ecclisomyia bilera Denning, female genitalia. 4a, ventral; 4b, left lateral; 4c, dorsal. Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; spt.sc = spermathecal sclerite; ssc.o = spermathecal sclerite opening; VIIIs = sternum VIII; Xt = tergum X." httpUri="https://zenodo.org/record/1227351/files/figure.png" pageId="35" pageNumber="236">4a</figureCitation>
). The spermathecal sclerite is not as long as in
<taxonomicName id="2A8632F6760F4331FCB5F9C41AAB564C" box="[859,1006,1566,1589]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="conspersa">
<emphasis id="DFF29567760F4331FCB5F9C41AAB564C" box="[859,1006,1566,1589]" italics="true" pageId="35" pageNumber="236">E. conspersa</emphasis>
</taxonomicName>
; the spermathecal sclerite opening is relatively longer than in
<taxonomicName id="2A8632F6760F4331FE44F9981B495620" authorityName="Denning" authorityYear="1951" box="[426,524,1601,1625]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="bilera">
<emphasis id="DFF29567760F4331FE44F9981B495620" box="[426,524,1601,1625]" italics="true" pageId="35" pageNumber="236">E. bilera</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="ED394975760F4331FF29F9BF1BEE57BB" blockId="35.[151,1436,1169,1986]" pageId="35" pageNumber="236">
<emphasis id="DFF29567760F4331FF29F9BF18EB5607" bold="true" box="[199,430,1637,1662]" pageId="35" pageNumber="236">Female description</emphasis>
. See
<bibRefCitation id="89173484760F4331FE1DF9BF1BE95607" author="Denning, D. G." box="[499,684,1637,1662]" journalOrPublisher="Psyche" pageId="35" pageNumber="236" pagination="16 - 27" part="55" publicationUrl="https://doi.org/10.1155/1948/71426" refId="ref35958" refString="Denning, D. G. (1948) New and little known species of Nearctic Trichoptera. Psyche, 55 (1), 16 - 27. https: // doi. org / 10.1155 / 1948 / 71426" title="New and little known species of Nearctic Trichoptera" type="journal article" year="1948">Denning (1948)</bibRefCitation>
;
<bibRefCitation id="89173484760F4331FD55F9BF1A2C5607" author="Nimmo, A." box="[699,873,1637,1662]" journalOrPublisher="Quaestiones Entomologicae" pageId="35" pageNumber="236" pagination="3 - 234" part="7" refId="ref36774" refString="Nimmo, A. (1971) The adult Rhyacophilidae and Limnephilidae (Trichoptera) of Alberta and Eastern British Columbia and their post-glacial origin. Quaestiones Entomologicae, 7 (1), 3 - 234." title="The adult Rhyacophilidae and Limnephilidae (Trichoptera) of Alberta and Eastern British Columbia and their post-glacial origin" type="journal article" year="1971">Nimmo (1971)</bibRefCitation>
. Supplemental description follows: Length 810 mm (N = 14). Head, thorax, legs reddish brown. Antennae each with approximately 3842 segments; antennal segments with numerous translucent setae and short spines as in males, but less visible. Tergum X fused with tergum IX (
<figureCitation id="75BD55F0760F4331FEF2F90B18385693" box="[284,381,1745,1770]" captionStart="FIGURES24" captionStartId="32.[151,264,1850,1872]" captionTargetBox="[196,1404,269,1789]" captionTargetId="figure@32.[181,1313,944,1693]" captionTargetPageId="32" captionText="FIGURES24a24c.Ecclisomyia maculosa Banks, female terminalia.24a, leftlateral; 24b, dorsal;24c, ventral.Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; IXs = sternum IX; IXt = tergum IX; pl = pleuron (paired); spt.sc = spermathecal sclerite; ssc.o = spermathecal sclerite opening; VIIIs = sternumVIII; IXs = sternum IX; IXt = tergum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227387/files/figure.png" pageId="35" pageNumber="236">Fig. 24a</figureCitation>
); distal margin of tergum X with V-shaped incision dividing tergum into 2 lobes for about 1/ 23/4 length of tergum. External part of gonopod VIII large, extending beyond genital opening; in ventral aspect lateral margins straight, caudal margin nearly straight (
<figureCitation id="75BD55F0760F4331FD15F8C31A1D574B" box="[763,856,1817,1842]" captionStart="FIGURES 25" captionStartId="33.[151,262,1830,1852]" captionTargetBox="[181,1427,253,1779]" captionTargetId="figure@33.[375,1427,934,1683]" captionTargetPageId="33" captionText="FIGURES 25a25c. Ecclisomyia maculosa Banks, female terminalia. 25a, left lateral; 25b, ventral; 25c, ventral. Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; spt.sc = spermathecal sclerite; ssc.o = spermathecalsclerite opening; VIIIs = sternum VIII; IXs =sternum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227389/files/figure.png" pageId="35" pageNumber="236">Fig. 25c</figureCitation>
) to moderately convex (
<figureCitation id="75BD55F0760F4331FB87F8C31D8F574B" box="[1129,1226,1817,1842]" captionStart="FIGURES24" captionStartId="32.[151,264,1850,1872]" captionTargetBox="[196,1404,269,1789]" captionTargetId="figure@32.[181,1313,944,1693]" captionTargetPageId="32" captionText="FIGURES24a24c.Ecclisomyia maculosa Banks, female terminalia.24a, leftlateral; 24b, dorsal;24c, ventral.Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; IXs = sternum IX; IXt = tergum IX; pl = pleuron (paired); spt.sc = spermathecal sclerite; ssc.o = spermathecal sclerite opening; VIIIs = sternumVIII; IXs = sternum IX; IXt = tergum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227387/files/figure.png" pageId="35" pageNumber="236">Fig. 24c</figureCitation>
), or may have a Vshaped mesal indentation (
<figureCitation id="75BD55F0760F4331FE51F8E71B67572F" box="[447,546,1853,1878]" captionStart="FIGURES 25" captionStartId="33.[151,262,1830,1852]" captionTargetBox="[181,1427,253,1779]" captionTargetId="figure@33.[375,1427,934,1683]" captionTargetPageId="33" captionText="FIGURES 25a25c. Ecclisomyia maculosa Banks, female terminalia. 25a, left lateral; 25b, ventral; 25c, ventral. Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; spt.sc = spermathecal sclerite; ssc.o = spermathecalsclerite opening; VIIIs = sternum VIII; IXs =sternum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227389/files/figure.png" pageId="35" pageNumber="236">Fig. 25b</figureCitation>
). External gonopod of segment IX sclerotized, narrow forming dorsal surface to vaginal opening; caudal margin with deep V-shaped declivity, extending posterad about 2 X its width beyond caudal margin of extended gonopod of segment VIII (
<figureCitation id="75BD55F0760F4331FD19F85F1A2757E7" box="[759,866,1925,1950]" captionStart="FIGURES24" captionStartId="32.[151,264,1850,1872]" captionTargetBox="[196,1404,269,1789]" captionTargetId="figure@32.[181,1313,944,1693]" captionTargetPageId="32" captionText="FIGURES24a24c.Ecclisomyia maculosa Banks, female terminalia.24a, leftlateral; 24b, dorsal;24c, ventral.Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; IXs = sternum IX; IXt = tergum IX; pl = pleuron (paired); spt.sc = spermathecal sclerite; ssc.o = spermathecal sclerite opening; VIIIs = sternumVIII; IXs = sternum IX; IXt = tergum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227387/files/figure.png" pageId="35" pageNumber="236">Figs. 24c</figureCitation>
,
<figureCitation id="75BD55F0760F4331FC83F85F1A9757E7" box="[877,978,1925,1950]" captionStart="FIGURES 25" captionStartId="33.[151,262,1830,1852]" captionTargetBox="[181,1427,253,1779]" captionTargetId="figure@33.[375,1427,934,1683]" captionTargetPageId="33" captionText="FIGURES 25a25c. Ecclisomyia maculosa Banks, female terminalia. 25a, left lateral; 25b, ventral; 25c, ventral. Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; spt.sc = spermathecal sclerite; ssc.o = spermathecalsclerite opening; VIIIs = sternum VIII; IXs =sternum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227389/files/figure.png" pageId="35" pageNumber="236">25b, 25c</figureCitation>
). Spermathecal sclerite nearly as long as wide (
<figureCitation id="75BD55F0760F4331FF33F873180257BB" box="[221,327,1961,1986]" captionStart="FIGURES24" captionStartId="32.[151,264,1850,1872]" captionTargetBox="[196,1404,269,1789]" captionTargetId="figure@32.[181,1313,944,1693]" captionTargetPageId="32" captionText="FIGURES24a24c.Ecclisomyia maculosa Banks, female terminalia.24a, leftlateral; 24b, dorsal;24c, ventral.Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; IXs = sternum IX; IXt = tergum IX; pl = pleuron (paired); spt.sc = spermathecal sclerite; ssc.o = spermathecal sclerite opening; VIIIs = sternumVIII; IXs = sternum IX; IXt = tergum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227387/files/figure.png" pageId="35" pageNumber="236">Figs. 24c</figureCitation>
,
<figureCitation id="75BD55F0760F4331FEBCF873183A57BB" box="[338,383,1961,1986]" captionStart="FIGURES 25" captionStartId="33.[151,262,1830,1852]" captionTargetBox="[181,1427,253,1779]" captionTargetId="figure@33.[375,1427,934,1683]" captionTargetPageId="33" captionText="FIGURES 25a25c. Ecclisomyia maculosa Banks, female terminalia. 25a, left lateral; 25b, ventral; 25c, ventral. Abbreviations: e.gon.VIII = external part of gonopod VIII; e.gon.IX = external part of gonopod IX; spt.sc = spermathecal sclerite; ssc.o = spermathecalsclerite opening; VIIIs = sternum VIII; IXs =sternum IX; Xt = tergum X." httpUri="https://zenodo.org/record/1227389/files/figure.png" pageId="35" pageNumber="236">25b</figureCitation>
), anterior margin rounded.
</paragraph>
</subSubSection>
<caption id="B9F919FD76084336FF79F9541A3856BA" httpUri="https://zenodo.org/record/1227397/files/figure.png" pageId="36" pageNumber="237" targetBox="[283,1275,227,1577]" targetPageId="36">
<paragraph id="ED39497576084336FF79F9541A3856BA" blockId="36.[151,1436,1678,1731]" pageId="36" pageNumber="237">
<emphasis id="DFF2956776084336FF79F954182C56DD" bold="true" box="[151,361,1678,1700]" pageId="36" pageNumber="237">FIGURES 29a29d</emphasis>
.
<taxonomicName id="2A8632F676084336FE99F9541BE056DD" authority="Banks" authorityName="Banks" authorityYear="1907" box="[375,677,1678,1700]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF2956776084336FE99F9541B1956DD" box="[375,604,1678,1700]" italics="true" pageId="36" pageNumber="237">Ecclisomyia maculosa</emphasis>
Banks
</taxonomicName>
, pupa. 29a, labrum, dorsal; 29b, mandibles, ventral; 29c, abdominal setae and hook plates, dorsal; 29d, dorsal process abdominal segment I, dorsal.
</paragraph>
</caption>
<subSubSection id="A59C1AFE76084336FF29F9351C0157C5" pageId="36" pageNumber="237" type="diagnosis">
<paragraph id="ED39497576084336FF29F9351C0157C5" blockId="36.[151,1436,1775,2016]" pageId="36" pageNumber="237">
<emphasis id="DFF2956776084336FF29F93518CF5771" bold="true" box="[199,394,1775,1800]" pageId="36" pageNumber="237">Pupal diagnosis.</emphasis>
The
<taxonomicName id="2A8632F676084336FE2AF92B1B0A5771" authorityName="Banks" authorityYear="1907" box="[452,591,1776,1800]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF2956776084336FE2AF92B1B0A5771" box="[452,591,1776,1800]" italics="true" pageId="36" pageNumber="237">E. maculosa</emphasis>
</taxonomicName>
pupa may be separated from that of
<taxonomicName id="2A8632F676084336FC05F92B1D0B5771" authorityName="Denning" authorityYear="1951" box="[1003,1102,1776,1800]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="bilera">
<emphasis id="DFF2956776084336FC05F92B1D0B5771" box="[1003,1102,1776,1800]" italics="true" pageId="36" pageNumber="237">E. bilera</emphasis>
</taxonomicName>
by the absence of dorsal gills on abdominal segment VIIa and presence on
<taxonomicName id="2A8632F676084336FD4FF8CF1A405755" authorityName="Denning" authorityYear="1951" box="[673,773,1812,1836]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="bilera">
<emphasis id="DFF2956776084336FD4FF8CF1A405755" box="[673,773,1812,1836]" italics="true" pageId="36" pageNumber="237">E. bilera</emphasis>
</taxonomicName>
. The
<taxonomicName id="2A8632F676084336FCA4F8CF1A9D5755" authorityName="Banks" authorityYear="1907" box="[842,984,1812,1836]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF2956776084336FCA4F8CF1A9D5755" box="[842,984,1812,1836]" italics="true" pageId="36" pageNumber="237">E. maculosa</emphasis>
</taxonomicName>
pupa may be separated from that of
<taxonomicName id="2A8632F676084336FA6AF8CF184C5729" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="conspersa">
<emphasis id="DFF2956776084336FA6AF8CF184C5729" italics="true" pageId="36" pageNumber="237">E. conspersa</emphasis>
</taxonomicName>
by the absence of lateral line gills; they are present on abdominal segments IIpIVa of
<taxonomicName id="2A8632F676084336FB3EF8E31C255729" box="[1232,1376,1849,1872]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="conspersa">
<emphasis id="DFF2956776084336FB3EF8E31C255729" box="[1232,1376,1849,1872]" italics="true" pageId="36" pageNumber="237">E. conspersa</emphasis>
</taxonomicName>
(
<figureCitation id="75BD55F076084336FA80F8ED19F2570D" captionStart="FIGURE 42" captionStartId="51.[151,250,1456,1478]" captionTargetBox="[202,1382,260,1371]" captionTargetPageId="51" captionText="FIGURE 42. Pupal and larval gill schematic diagrams, left lateral, for Ecclisomyia bilera Denning, Ecclisomyia conspersa Banks, Ecclisomyia maculosa Banks, Ecclisomyia simulata Banks. Abbreviations: IVIII = dorsal abdominal segments, with Arabic numerals representing the number of gills per segment in anterior and posterior positions at dorsal, lateral or lateral line (dorsolateral and ventrolateral), and ventral levels." httpUri="https://zenodo.org/record/1227419/files/figure.svg" pageId="36" pageNumber="237">Fig. 42</figureCitation>
). The apical processes of
<taxonomicName id="2A8632F676084336FE09F8871B30570D" authorityName="Banks" authorityYear="1907" box="[487,629,1884,1908]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF2956776084336FE09F8871B30570D" box="[487,629,1884,1908]" italics="true" pageId="36" pageNumber="237">E. maculosa</emphasis>
</taxonomicName>
each have 2 stout, long, black setae at the apex, whereas those of
<taxonomicName id="2A8632F676084336FA6AF887184C57E1" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="conspersa">
<emphasis id="DFF2956776084336FA6AF887184C57E1" italics="true" pageId="36" pageNumber="237">E. conspersa</emphasis>
</taxonomicName>
each have 3 stout, long, black setae. The
<taxonomicName id="2A8632F676084336FD36F85B1A2657E1" authorityName="Banks" authorityYear="1907" box="[728,867,1920,1944]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF2956776084336FD36F85B1A2657E1" box="[728,867,1920,1944]" italics="true" pageId="36" pageNumber="237">E. maculosa</emphasis>
</taxonomicName>
pupa may be separated from that of
<taxonomicName id="2A8632F676084336FB14F85B1C3F57E1" baseAuthorityName="Givens" baseAuthorityYear="2014" box="[1274,1402,1920,1944]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="simulata">
<emphasis id="DFF2956776084336FB14F85B1C3F57E1" box="[1274,1402,1920,1944]" italics="true" pageId="36" pageNumber="237">E. simulata</emphasis>
</taxonomicName>
by the absence of single filament dorsal gills on abdominal segment IIa and presence on
<taxonomicName id="2A8632F676084336FBBBF87F1D9057C5" baseAuthorityName="Givens" baseAuthorityYear="2014" box="[1109,1237,1956,1980]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="simulata">
<emphasis id="DFF2956776084336FBBBF87F1D9057C5" box="[1109,1237,1956,1980]" italics="true" pageId="36" pageNumber="237">E. simulata</emphasis>
</taxonomicName>
(
<figureCitation id="75BD55F076084336FB0AF87E1C7D57C5" box="[1252,1336,1956,1981]" captionStart="FIGURE 42" captionStartId="51.[151,250,1456,1478]" captionTargetBox="[202,1382,260,1371]" captionTargetPageId="51" captionText="FIGURE 42. Pupal and larval gill schematic diagrams, left lateral, for Ecclisomyia bilera Denning, Ecclisomyia conspersa Banks, Ecclisomyia maculosa Banks, Ecclisomyia simulata Banks. Abbreviations: IVIII = dorsal abdominal segments, with Arabic numerals representing the number of gills per segment in anterior and posterior positions at dorsal, lateral or lateral line (dorsolateral and ventrolateral), and ventral levels." httpUri="https://zenodo.org/record/1227419/files/figure.svg" pageId="36" pageNumber="237">Fig. 42</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A59C1AFE76084337FF29F81D1A4E552D" lastPageId="37" lastPageNumber="238" pageId="36" pageNumber="237" type="description">
<paragraph id="ED39497576084337FF29F81D1B815139" blockId="36.[151,1436,1775,2016]" lastBlockId="37.[151,1437,151,2013]" lastPageId="37" lastPageNumber="238" pageId="36" pageNumber="237">
<emphasis id="DFF2956776084336FF29F81D18E15799" bold="true" box="[199,420,1991,2016]" pageId="36" pageNumber="237">Pupal description.</emphasis>
Length 810 mm (N = 8). Head: Light brown. Male antennae extending to or slightly caudad of abdominal apical processes; antennae in females extending to abdominal segment V. Labrum dorsal margin convex with 10 long black setae on dorsal surface, setae directed dorsad, distal end of each seta hooked (
<figureCitation id="75BD55F076094337FF71FF0518445081" box="[159,257,223,248]" captionStart="FIGURES 29" captionStartId="36.[151,264,1678,1700]" captionTargetBox="[283,1275,227,1577]" captionTargetPageId="36" captionText="FIGURES 29a29d. Ecclisomyia maculosa Banks, pupa. 29a, labrum, dorsal; 29b, mandibles, ventral; 29c, abdominal setae and hook plates, dorsal;29d, dorsal process abdominal segment I, dorsal." httpUri="https://zenodo.org/record/1227397/files/figure.png" pageId="37" pageNumber="238">Fig. 29a</figureCitation>
); posterior area lateral of meson with 2 short reddish setae. Mandibles black to reddish-brown, bases of mandibles broad, each tapering into single triangular apical tooth; mesal margin with minute serrations; ventrolateral surface with 2 long, black setae (
<figureCitation id="75BD55F076094337FDBAFEFD1BF25139" box="[596,695,295,320]" captionStart="FIGURES 29" captionStartId="36.[151,264,1678,1700]" captionTargetBox="[283,1275,227,1577]" captionTargetPageId="36" captionText="FIGURES 29a29d. Ecclisomyia maculosa Banks, pupa. 29a, labrum, dorsal; 29b, mandibles, ventral; 29c, abdominal setae and hook plates, dorsal;29d, dorsal process abdominal segment I, dorsal." httpUri="https://zenodo.org/record/1227397/files/figure.png" pageId="37" pageNumber="238">Fig. 29b</figureCitation>
).
</paragraph>
<paragraph id="ED39497576094337FF29FE961B5A5244" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="238">Thorax: Pro-, meso-, and metanota light brown. Pro-, meso-, and metathoracic legs brown to yellow-brown. In mature pupa, tarsal and tibial segments with both single and paired reddish-black stout spines. Spines absent in immature pupa. Procoxae of both mature and immature pupae frequently each with 1 long black seta near lateral margin and 35 shorter black setae on ventral surface. Mesocoxae may each have 1 long black seta near lateral margin and 13 short black setae on ventral surface. Profemora each with single short black spine on ventral surface near distal margin. Mesotarsal segments 14 each with 2 lateral fringes of long reddish to opaque (whitish) silk-like hairs, directed posteriorly.</paragraph>
<paragraph id="ED39497576094337FF29FD9D1DE154BD" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="238">
Abdomen: Lateral fringe on each side consisting of long feathery reddish hairs directed dorsad; originating from posterior area of abdominal segment V, with line of fringe arching ventrad, extending to anterior margin of abdominal segment VIII. Dorsal and ventral abdominal gills present, each single-filament; present dorsoposteriorly and ventroposteriorly on segments IIVII; present dorsoanteriorly and ventroanteriorly on segments IIIVI; segments II and VII without gills dorsoanteriorly and ventroanteriorly; without lateral-line gills (
<figureCitation id="75BD55F076094337FB08FD0D1C795289" box="[1254,1340,727,752]" captionStart="FIGURE 42" captionStartId="51.[151,250,1456,1478]" captionTargetBox="[202,1382,260,1371]" captionTargetPageId="51" captionText="FIGURE 42. Pupal and larval gill schematic diagrams, left lateral, for Ecclisomyia bilera Denning, Ecclisomyia conspersa Banks, Ecclisomyia maculosa Banks, Ecclisomyia simulata Banks. Abbreviations: IVIII = dorsal abdominal segments, with Arabic numerals representing the number of gills per segment in anterior and posterior positions at dorsal, lateral or lateral line (dorsolateral and ventrolateral), and ventral levels." httpUri="https://zenodo.org/record/1227419/files/figure.svg" pageId="37" pageNumber="238">Fig. 42</figureCitation>
). Dorsal process of abdominal segment I with 2 black
<emphasis id="DFF2956776094337FD2AFD271B98536D" box="[708,733,765,788]" italics="true" pageId="37" pageNumber="238">sa</emphasis>
2 setae, each positioned lateral of meson; posterior margin sclerotized, light brown, forming pair of denticulate lobes lateral of meson, each lobe with 810 small spicules (
<figureCitation id="75BD55F076094337FF71FC9E18445324" box="[159,257,836,861]" captionStart="FIGURES 29" captionStartId="36.[151,264,1678,1700]" captionTargetBox="[283,1275,227,1577]" captionTargetPageId="36" captionText="FIGURES 29a29d. Ecclisomyia maculosa Banks, pupa. 29a, labrum, dorsal; 29b, mandibles, ventral; 29c, abdominal setae and hook plates, dorsal;29d, dorsal process abdominal segment I, dorsal." httpUri="https://zenodo.org/record/1227397/files/figure.png" pageId="37" pageNumber="238">Fig. 29d</figureCitation>
); shape of dorsal process varying with maturity of pupa. Hook plates on segments IIIVII (
<figureCitation id="75BD55F076094337FAEDFC9E1C2B5325" box="[1283,1390,836,861]" captionStart="FIGURES 29" captionStartId="36.[151,264,1678,1700]" captionTargetBox="[283,1275,227,1577]" captionTargetPageId="36" captionText="FIGURES 29a29d. Ecclisomyia maculosa Banks, pupa. 29a, labrum, dorsal; 29b, mandibles, ventral; 29c, abdominal setae and hook plates, dorsal;29d, dorsal process abdominal segment I, dorsal." httpUri="https://zenodo.org/record/1227397/files/figure.png" pageId="37" pageNumber="238">Figs. 29c</figureCitation>
,
<figureCitation id="75BD55F076094337FA97FC9E1CD25325" box="[1401,1431,836,860]" captionStart="FIGURES 39, 40" captionStartId="49.[151,264,1684,1706]" captionTargetBox="[206,1332,475,1559]" captionTargetPageId="49" captionText="FIGURES 39, 40. Ecclisomyia spp., pupal hook plates. 39, Ecclisomyia maculosa Banks; 40, Ecclisomyia simulata Banks. Abbreviations: IIIVII = abdominal terga III through VII; a = anterior; p = posterior." httpUri="https://zenodo.org/record/1227415/files/figure.png" pageId="37" pageNumber="238">39</figureCitation>
,
<figureCitation id="75BD55F076094337FF79FCB219F353F9" box="[151,182,872,896]" captionStart="FIGURE 41" captionStartId="50.[151,250,1928,1950]" captionTargetBox="[214,1335,746,1881]" captionTargetPageId="50" captionText="FIGURE 41. Pupal schematic hook plate diagrams, dorsal, for Ecclisomyia bilera Denning, Ecclisomyia conspersa Banks, Ecclisomyia maculosa Banks, Ecclisomyia simulata Banks. Abbreviations: IVIII = dorsal abdominal segments, with Arabic numerals representing the average number of hooks per plate." httpUri="https://zenodo.org/record/1227417/files/figure.svg" pageId="37" pageNumber="238">41</figureCitation>
). Pairs of hook plates present anteriorly on segments IIIVII, posteriorly on segment V. Hook plate IIIa with 27 hooks; IVa with 28; Va with 38; Vp with 917; VIa with 26; and VIIa with 37. Hook plates reddish brown, hooks directed posterad on anterior hook plates of segments III-VII and anterad on circular to elliptical hook plates Vp. Dorsal abdominal segments each with 1 pair short black
<emphasis id="DFF2956776094337FCBEFC0F1A2C5395" box="[848,873,981,1004]" italics="true" pageId="37" pageNumber="238">sa</emphasis>
2 setae on each of abdominal segments IIV; 23 pairs on abdominal segments V and VII (
<figureCitation id="75BD55F076094337FD9CFC2D1B905469" box="[626,725,1015,1040]" captionStart="FIGURES 29" captionStartId="36.[151,264,1678,1700]" captionTargetBox="[283,1275,227,1577]" captionTargetPageId="36" captionText="FIGURES 29a29d. Ecclisomyia maculosa Banks, pupa. 29a, labrum, dorsal; 29b, mandibles, ventral; 29c, abdominal setae and hook plates, dorsal;29d, dorsal process abdominal segment I, dorsal." httpUri="https://zenodo.org/record/1227397/files/figure.png" pageId="37" pageNumber="238">Fig. 29c</figureCitation>
); Dorsal abdominal segments VI sometimes with 1 pair short, black
<emphasis id="DFF2956776094337FF34FBC719B6544D" box="[218,243,1053,1076]" italics="true" pageId="37" pageNumber="238">sa</emphasis>
2 setae; segments VVII sometimes with 1 pair short, black
<emphasis id="DFF2956776094337FC7EFBC61AF2544D" box="[912,951,1052,1076]" italics="true" pageId="37" pageNumber="238">sa1</emphasis>
setae. Dorsal abdominal segment VIII with 2 to 3 pairs of black to translucent setae lateral of meson. Distal area formed into 2 caudal lobes (apical processes); apical processes of male with light covering of spicules; spicules at apices of caudal lobes very difficult to discern with light microscope. Caudal lobes each with 2 long fine reddish-black to black setae on distal margin. Lobes of male and female slanted mesad; in ventral view lateral margins sinuate (
<figureCitation id="75BD55F076094337FC51FB761D6C54BC" box="[959,1065,1196,1221]" captionStart="FIGURES 26" captionStartId="34.[151,264,1838,1860]" captionTargetBox="[279,1271,253,886]" captionTargetId="figure@34.[771,1359,1036,1670]" captionTargetPageId="34" captionText="FIGURES 26a, 26b. Ecclisomyia maculosa Banks, male pupa, apical process.26a, dorsal; 26b, ventral." httpUri="https://zenodo.org/record/1227391/files/figure.png" pageId="37" pageNumber="238">Figs. 26a</figureCitation>
,
<figureCitation id="75BD55F076094337FBDBFB761D2554BD" box="[1077,1120,1196,1220]" captionStart="FIGURES 27" captionStartId="34.[151,264,1869,1891]" captionTargetBox="[239,1359,1036,1765]" captionTargetId="figure@34.[771,1359,1036,1670]" captionTargetPageId="34" captionText="FIGURES 27a, 27b. Ecclisomyia maculosa Banks, male pupa, apical process.27a, dorsal; 27b, ventral." httpUri="https://zenodo.org/record/1227393/files/figure.png" pageId="37" pageNumber="238">27a</figureCitation>
, 28a).
</paragraph>
<paragraph id="ED39497576094337FF29FB151A4E552D" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="238">
<emphasis id="DFF2956776094337FF29FB15180B5491" bold="true" box="[199,334,1231,1256]" pageId="37" pageNumber="238">Pupal case.</emphasis>
Length 912 mm (N = 19). Width 2.53.0 mm (N = 6). Case long, narrow, constructed of small pebbles and grains of sand, with tight interstitial spaces, over an inner lining of silk surrounding pupa. Case nearly uniform in diameter, anterior end slightly wider than posterior end (
<figureCitation id="75BD55F076094337FC47FACD1D485549" box="[937,1037,1303,1328]" captionStart="FIGURES 43" captionStartId="52.[151,264,1972,1994]" captionTargetBox="[504,1116,273,1884]" captionTargetId="figure@52.[492,1124,1018,1268]" captionTargetPageId="52" captionText="FIGURES 43a43d. Pupal cases. 43a-1 and 43a-2, Ecclisomyia bilera Denning (811 mm); 43b, Ecclisomyia conspersa Banks (16 mm); 43c, Ecclisomyia maculosa Banks (10.5 mm); 43d, Ecclisomyia simulata Banks (11 mm)." httpUri="https://zenodo.org/record/1227421/files/figure.png" pageId="37" pageNumber="238">Fig. 43c</figureCitation>
). Ends of case rounded, each with small open area to allow for flow of water through case.
</paragraph>
</subSubSection>
<subSubSection id="A59C1AFE76094337FF29FA851DDB5629" pageId="37" pageNumber="238" type="diagnosis">
<paragraph id="ED39497576094337FF29FA851DDB5629" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="238">
<emphasis id="DFF2956776094337FF29FA8518DD5501" bold="true" box="[199,408,1375,1400]" pageId="37" pageNumber="238">Larval diagnosis.</emphasis>
Of the four Nearctic
<taxonomicName id="2A8632F676094337FD76FABA1A645501" authorityName="Banks" authorityYear="1907" box="[664,801,1376,1400]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="genus">
<emphasis id="DFF2956776094337FD76FABA1A645501" box="[664,801,1376,1400]" italics="true" pageId="37" pageNumber="238">Ecclisomyia</emphasis>
</taxonomicName>
, only
<taxonomicName id="2A8632F676094337FC80FABB1AB85501" authorityName="Banks" authorityYear="1907" box="[878,1021,1376,1400]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF2956776094337FC80FABB1AB85501" box="[878,1021,1376,1400]" italics="true" pageId="37" pageNumber="238">E. maculosa</emphasis>
</taxonomicName>
lacks a prosternal sclerite, or small sclerotized area on the venter of the prothorax. The parietals have distinct muscle scars ventral and posterior of the eyes.
<taxonomicName id="2A8632F676094337FF38FA72189455B9" authorityName="Banks" authorityYear="1907" box="[214,465,1448,1472]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF2956776094337FF38FA72189455B9" box="[214,465,1448,1472]" italics="true" pageId="37" pageNumber="238">Ecclisomyia maculosa</emphasis>
</taxonomicName>
lacks both anterodorsal and anteroventral gills on abdominal segment VII, but gills are present in the other Nearctic
<taxonomicName id="2A8632F676094337FE31FA161B2D559D" authorityName="Banks" authorityYear="1907" box="[479,616,1484,1508]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="genus">
<emphasis id="DFF2956776094337FE31FA161B2D559D" box="[479,616,1484,1508]" italics="true" pageId="37" pageNumber="238">Ecclisomyia</emphasis>
</taxonomicName>
(
<figureCitation id="75BD55F076094337FD96FA161B88559D" box="[632,717,1484,1509]" captionStart="FIGURE 42" captionStartId="51.[151,250,1456,1478]" captionTargetBox="[202,1382,260,1371]" captionTargetPageId="51" captionText="FIGURE 42. Pupal and larval gill schematic diagrams, left lateral, for Ecclisomyia bilera Denning, Ecclisomyia conspersa Banks, Ecclisomyia maculosa Banks, Ecclisomyia simulata Banks. Abbreviations: IVIII = dorsal abdominal segments, with Arabic numerals representing the number of gills per segment in anterior and posterior positions at dorsal, lateral or lateral line (dorsolateral and ventrolateral), and ventral levels." httpUri="https://zenodo.org/record/1227419/files/figure.svg" pageId="37" pageNumber="238">Fig. 42</figureCitation>
). In addition, the
<taxonomicName id="2A8632F676094337FC7BFA171D65559D" authorityName="Banks" authorityYear="1907" box="[917,1056,1484,1508]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF2956776094337FC7BFA171D65559D" box="[917,1056,1484,1508]" italics="true" pageId="37" pageNumber="238">E. maculosa</emphasis>
</taxonomicName>
larva may be separated from that of
<taxonomicName id="2A8632F676094337FF5BFA2B18705671" baseAuthorityName="Givens" baseAuthorityYear="2014" box="[181,309,1520,1544]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="simulata">
<emphasis id="DFF2956776094337FF5BFA2B18705671" box="[181,309,1520,1544]" italics="true" pageId="37" pageNumber="238">E. simulata</emphasis>
</taxonomicName>
by the lack of dorsoanterior gills on abdominal segment II, and the presence of these gills on the
<taxonomicName id="2A8632F676094337FA6AFA2B19BD5655" baseAuthorityName="Givens" baseAuthorityYear="2014" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="simulata">
<emphasis id="DFF2956776094337FA6AFA2B19BD5655" italics="true" pageId="37" pageNumber="238">E. simulata</emphasis>
</taxonomicName>
larva. The
<taxonomicName id="2A8632F676094337FE99F9CF1B445655" authorityName="Banks" authorityYear="1907" box="[375,513,1556,1580]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF2956776094337FE99F9CF1B445655" box="[375,513,1556,1580]" italics="true" pageId="37" pageNumber="238">E. maculosa</emphasis>
</taxonomicName>
larva is without lateral line gills, whereas the
<taxonomicName id="2A8632F676094337FBE8F9CF1DD25655" box="[1030,1175,1557,1580]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="conspersa">
<emphasis id="DFF2956776094337FBE8F9CF1DD25655" box="[1030,1175,1557,1580]" italics="true" pageId="37" pageNumber="238">E. conspersa</emphasis>
</taxonomicName>
larva has 2 gills with 1 dorsolateral gill and 1 ventrolateral gill on each side of abdominal segments IIIV (
<figureCitation id="75BD55F076094337FBD3F9ED1DD75629" box="[1085,1170,1591,1616]" captionStart="FIGURE 42" captionStartId="51.[151,250,1456,1478]" captionTargetBox="[202,1382,260,1371]" captionTargetPageId="51" captionText="FIGURE 42. Pupal and larval gill schematic diagrams, left lateral, for Ecclisomyia bilera Denning, Ecclisomyia conspersa Banks, Ecclisomyia maculosa Banks, Ecclisomyia simulata Banks. Abbreviations: IVIII = dorsal abdominal segments, with Arabic numerals representing the number of gills per segment in anterior and posterior positions at dorsal, lateral or lateral line (dorsolateral and ventrolateral), and ventral levels." httpUri="https://zenodo.org/record/1227419/files/figure.svg" pageId="37" pageNumber="238">Fig. 42</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A59C1AFE76094335FF29F986185D51D4" lastPageId="39" lastPageNumber="240" pageId="37" pageNumber="238" type="description">
<paragraph id="ED39497576094334FF29F9861AED5245" blockId="37.[151,1437,151,2013]" lastBlockId="38.[151,1437,151,2013]" lastPageId="38" lastPageNumber="239" pageId="37" pageNumber="238">
<emphasis id="DFF2956776094337FF29F98618EB560C" bold="true" box="[199,430,1628,1653]" pageId="37" pageNumber="238">Larval description.</emphasis>
Length (mature instar) 811 mm (N = 18). Head: Reddish brown. Parietals, genae and post genae glabrous, reddish brown to dark brown; area about coronal suture with faint yellow strip; muscle scars distinct posterior and lateral of eyes. Mandibles black, each with 4 small blunt apical teeth; posterolateral area with 12 short translucent setae; mesal margin with tuft of translucent feather-like setae (
<figureCitation id="75BD55F076094337FB90F91D1DA65699" box="[1150,1251,1735,1760]" captionStart="FIGURES 19" captionStartId="21.[151,264,1984,2006]" captionTargetBox="[365,1241,234,1912]" captionTargetId="figure@21.[351,1259,211,1133]" captionTargetPageId="21" captionText="FIGURES 19a, 19b. Ecclisomyia conspersa Banks, larval head. 19a, dorsal, with primary setae numbered; 19b, ventral, with primary setae numbered." httpUri="https://zenodo.org/record/1227377/files/figure.png" pageId="37" pageNumber="238">Fig. 19a</figureCitation>
). Frontoclypeus smooth, without texture; posterior half without evident muscle scars, or muscle scars indistinct. Ventral apodome triangular, dark brown, 1.142.40 X longer than ventral ecdysical line. Early instar larva with ventral apodome shorter than ventral ecdysical line. Primary setae in positions 7 and 917 (
<figureCitation id="75BD55F076094337FC00F8EE1D145735" box="[1006,1105,1844,1869]" captionStart="FIGURES 19" captionStartId="21.[151,264,1984,2006]" captionTargetBox="[365,1241,234,1912]" captionTargetId="figure@21.[351,1259,211,1133]" captionTargetPageId="21" captionText="FIGURES 19a, 19b. Ecclisomyia conspersa Banks, larval head. 19a, dorsal, with primary setae numbered; 19b, ventral, with primary setae numbered." httpUri="https://zenodo.org/record/1227377/files/figure.png" pageId="37" pageNumber="238">Fig. 19a</figureCitation>
). Primary setae in position 7 long, translucent; primary setae in position 9 black to reddish; primary setae in position 10 short, appressed, and transparent; primary setae in position 11 short, black to translucent; primary setae in position 12, short to long, black to transparent, fine, may be difficult to discern; primary setae in position 13 short, translucent, difficult to discern, may be missing; primary setae in position 14 long, black, and stout; primary setae in position 15 black; primary setae in position 16 appressed, translucent, hard to discern, may be absent; primary setae in position 17 black. Venter of head with primary setae in positions 8 and 18 (
<figureCitation id="75BD55F0760A4334FCB2FF661AFB50AC" box="[860,958,188,213]" captionStart="FIGURES 19" captionStartId="21.[151,264,1984,2006]" captionTargetBox="[365,1241,234,1912]" captionTargetId="figure@21.[351,1259,211,1133]" captionTargetPageId="21" captionText="FIGURES 19a, 19b. Ecclisomyia conspersa Banks, larval head. 19a, dorsal, with primary setae numbered; 19b, ventral, with primary setae numbered." httpUri="https://zenodo.org/record/1227377/files/figure.png" pageId="38" pageNumber="239">Fig. 19b</figureCitation>
). Primary setae in position 8 short, may be translucent, difficult to discern; primary setae in position 18 translucent, short, difficult to discern, best seen in lateral view. Frontoclypeus smooth, without texture; wide anteriorly, narrow posteriorly, triangular; glabrous except for primary setae in positions 16 (
<figureCitation id="75BD55F0760A4334FD92FEFD1B9B5139" box="[636,734,295,320]" captionStart="FIGURES 19" captionStartId="21.[151,264,1984,2006]" captionTargetBox="[365,1241,234,1912]" captionTargetId="figure@21.[351,1259,211,1133]" captionTargetPageId="21" captionText="FIGURES 19a, 19b. Ecclisomyia conspersa Banks, larval head. 19a, dorsal, with primary setae numbered; 19b, ventral, with primary setae numbered." httpUri="https://zenodo.org/record/1227377/files/figure.png" pageId="38" pageNumber="239">Fig. 19a</figureCitation>
). Primary setae in position 1 short or long, translucent, semiappressed; primary setae in positions 2 and 3 long and black, sometimes short and translucent; primary setae in position 4 appressed, short or long, black or translucent; primary setae in position 5 long, black, sometimes translucent, difficult to discern; primary setae in position 6 translucent, difficult to discern. Labrum brown to dark brown, sometimes brown posteriorly, yellowish anteriorly. Primary setae in positions 16 (
<figureCitation id="75BD55F0760A4334FB0EFE6D1C0D51A9" box="[1248,1352,439,464]" captionStart="FIGURES 19" captionStartId="21.[151,264,1984,2006]" captionTargetBox="[365,1241,234,1912]" captionTargetId="figure@21.[351,1259,211,1133]" captionTargetPageId="21" captionText="FIGURES 19a, 19b. Ecclisomyia conspersa Banks, larval head. 19a, dorsal, with primary setae numbered; 19b, ventral, with primary setae numbered." httpUri="https://zenodo.org/record/1227377/files/figure.png" pageId="38" pageNumber="239">Fig. 19a</figureCitation>
); setae translucent, setal positions may be difficult to determine or setae absent. Primary setae in positions 1 and 2 translucent, short, best seen from ventral aspect. Cardo sclerites elongate, narrow, rectangular, and brown to dark brown. Submental sclerite with 1 short black seta set lateral of meson.
</paragraph>
<paragraph id="ED394975760A4334FF29FD9D1A0F5501" blockId="38.[151,1437,151,2013]" pageId="38" pageNumber="239">
Thorax: Pro- and mesonota brown to reddish brown, nota with mesal yellow strip; metanotal sclerites brown. Anterior margins of pronota each with 1013 long, black slender setae set equidistant from each other along anterior margin; short translucent setae positioned between these longer setae along anterior margin. Each half of pronotum with transverse line of 711 black
<emphasis id="DFF29567760A4334FD4FFD6E1BF152B5" box="[673,692,692,716]" italics="true" pageId="38" pageNumber="239">ls</emphasis>
setae posterior to mid-dorsal transverse groove;
<emphasis id="DFF29567760A4334FB04FD6F1C4652B5" box="[1258,1283,693,716]" italics="true" pageId="38" pageNumber="239">sa</emphasis>
2 setae short, black. Mesonotum with black
<emphasis id="DFF29567760A4334FE01FD021B475289" box="[495,514,728,752]" italics="true" pageId="38" pageNumber="239">ls</emphasis>
setae set along anterior margin. Mesonotum with black setae in 3 distinct groups roughly corresponding to
<emphasis id="DFF29567760A4334FE57FD271897536D" box="[441,466,765,788]" italics="true" pageId="38" pageNumber="239">sa</emphasis>
1,
<emphasis id="DFF29567760A4334FE03FD271B43536D" box="[493,518,765,788]" italics="true" pageId="38" pageNumber="239">sa</emphasis>
2, and
<emphasis id="DFF29567760A4334FDBCFD271B2E536D" box="[594,619,765,788]" italics="true" pageId="38" pageNumber="239">sa</emphasis>
3 positions; muscle scars not evident or indistinct. Metanota
<emphasis id="DFF29567760A4334FAFCFD271C6E536D" box="[1298,1323,765,788]" italics="true" pageId="38" pageNumber="239">sa</emphasis>
1 sclerites with 47 black
<emphasis id="DFF29567760A4334FEA0FCFA18245341" box="[334,353,800,824]" italics="true" pageId="38" pageNumber="239">ls</emphasis>
setae, setal sockets prominent; metanotal
<emphasis id="DFF29567760A4334FCA3FCFB1A235341" box="[845,870,801,824]" italics="true" pageId="38" pageNumber="239">sa</emphasis>
2 sclerites with 49 black
<emphasis id="DFF29567760A4334FB73FCFA1DF55341" box="[1181,1200,800,824]" italics="true" pageId="38" pageNumber="239">ls</emphasis>
setae; metanota
<emphasis id="DFF29567760A4334FA9BFCFB1CCB5341" box="[1397,1422,801,824]" italics="true" pageId="38" pageNumber="239">sa</emphasis>
3 sclerites parenthesis-shaped, each with 617
<emphasis id="DFF29567760A4334FD61FC9E1BE75325" box="[655,674,836,860]" italics="true" pageId="38" pageNumber="239">ls</emphasis>
black setae on anterior area of sclerites; metanotal sclerites in some specimens with setae translucent, not black. Prosternal horn short, sclerotized. Prosternal sclerite or sclerotized area absent. In prepupae examined, there is no prosternal sclerite or sclerotized area. Legs yellowbrown; foretrochantins dark brown; forming tent-like structure over procoxae. Procoxae with primary setae in positions 1 and 2; primary setae in position 2 black to translucent, setal sockets prominent (
<figureCitation id="75BD55F0760A4334FBB4FC0E1DFA5395" box="[1114,1215,980,1005]" captionStart="FIGURES 21" captionStartId="23.[151,264,1964,1986]" captionTargetBox="[427,1246,189,1907]" captionTargetId="figure@23.[445,1265,774,1216]" captionTargetPageId="23" captionText="FIGURES 21a21c. Ecclisomyia conspersa Banks, larva, right legs. 21a, prothoracic leg; 21b, mesothoracic leg; 21c, metathoracic leg.Abbreviations:ls = long slender setae; sl = spike-like setae; tb = trochanteral brush." httpUri="https://zenodo.org/record/1227381/files/figure.png" pageId="38" pageNumber="239">Fig. 21a</figureCitation>
). Pro-, meso-, and metacoxae each with black primary setae in position 2; ventral margins with 47 black
<emphasis id="DFF29567760A4334FB6EFC221DD65469" box="[1152,1171,1016,1040]" italics="true" pageId="38" pageNumber="239">ls</emphasis>
setae, sometimes short and translucent; dorsal margins of pro-, meso-, and metacoxae each with 1517 long black setae, all coxal setae with prominent setal sockets (
<figureCitation id="75BD55F0760A4334FE1EFBE51B945421" box="[496,721,1087,1112]" captionStart="FIGURES 21" captionStartId="23.[151,264,1964,1986]" captionTargetBox="[427,1246,189,1907]" captionTargetId="figure@23.[445,1265,774,1216]" captionTargetPageId="23" captionText="FIGURES 21a21c. Ecclisomyia conspersa Banks, larva, right legs. 21a, prothoracic leg; 21b, mesothoracic leg; 21c, metathoracic leg.Abbreviations:ls = long slender setae; sl = spike-like setae; tb = trochanteral brush." httpUri="https://zenodo.org/record/1227381/files/figure.png" pageId="38" pageNumber="239">Figs. 21a, 21b, 21c</figureCitation>
); some specimens with only 46 short translucent setae. Pro-, meso-, and metatrochanters each with black primary
<emphasis id="DFF29567760A4334FD07FBBE1BB95405" box="[745,764,1124,1148]" italics="true" pageId="38" pageNumber="239">ls</emphasis>
setae in positions 2, 3, and 5; seta in position 5 often absent or difficult to discern (
<figureCitation id="75BD55F0760A4334FE7DFB5D1B2B54D9" box="[403,622,1159,1184]" captionStart="FIGURES 21" captionStartId="23.[151,264,1964,1986]" captionTargetBox="[427,1246,189,1907]" captionTargetId="figure@23.[445,1265,774,1216]" captionTargetPageId="23" captionText="FIGURES 21a21c. Ecclisomyia conspersa Banks, larva, right legs. 21a, prothoracic leg; 21b, mesothoracic leg; 21c, metathoracic leg.Abbreviations:ls = long slender setae; sl = spike-like setae; tb = trochanteral brush." httpUri="https://zenodo.org/record/1227381/files/figure.png" pageId="38" pageNumber="239">Figs. 21a, 21b, 21c</figureCitation>
). Ventral margins of pro- and mesotrochanters and pro- and mesofemora each with trochanteral brush, feather-like, translucent, extending to proximal ventral area of each femur (
<figureCitation id="75BD55F0760A4334FAC3FB7619815491" captionStart="FIGURES 21" captionStartId="23.[151,264,1964,1986]" captionTargetBox="[427,1246,189,1907]" captionTargetId="figure@23.[445,1265,774,1216]" captionTargetPageId="23" captionText="FIGURES 21a21c. Ecclisomyia conspersa Banks, larva, right legs. 21a, prothoracic leg; 21b, mesothoracic leg; 21c, metathoracic leg.Abbreviations:ls = long slender setae; sl = spike-like setae; tb = trochanteral brush." httpUri="https://zenodo.org/record/1227381/files/figure.png" pageId="38" pageNumber="239">Figs. 21a, 21b</figureCitation>
). Protrochanters each with single reddish elongate
<emphasis id="DFF29567760A4334FCE8FB0A1A5C5491" box="[774,793,1232,1256]" italics="true" pageId="38" pageNumber="239">sl</emphasis>
seta on ventral margin. Profemora with primary setae in position 2 and with 2 elongate long reddish-brown to translucent stout
<emphasis id="DFF29567760A4334FC56FB2E1A8E5575" box="[952,971,1268,1292]" italics="true" pageId="38" pageNumber="239">sl</emphasis>
seta on ventral margin (
<figureCitation id="75BD55F0760A4334FB31FB2E1C055575" box="[1247,1344,1268,1293]" captionStart="FIGURES 21" captionStartId="23.[151,264,1964,1986]" captionTargetBox="[427,1246,189,1907]" captionTargetId="figure@23.[445,1265,774,1216]" captionTargetPageId="23" captionText="FIGURES 21a21c. Ecclisomyia conspersa Banks, larva, right legs. 21a, prothoracic leg; 21b, mesothoracic leg; 21c, metathoracic leg.Abbreviations:ls = long slender setae; sl = spike-like setae; tb = trochanteral brush." httpUri="https://zenodo.org/record/1227381/files/figure.png" pageId="38" pageNumber="239">Fig. 21a</figureCitation>
). Meso- and metafemora each with black
<emphasis id="DFF29567760A4334FDFFFAC21B615549" box="[529,548,1304,1328]" italics="true" pageId="38" pageNumber="239">ls</emphasis>
primary seta in positions 2, 3, and 4. Pro-, meso-, and metafemora with short reddish to translucent
<emphasis id="DFF29567760A4334FE7CFAE618E0552D" box="[402,421,1340,1364]" italics="true" pageId="38" pageNumber="239">sl</emphasis>
peg-like setae along ventral margins. Protibiae with primary setae in position 3. Pro- and mesopleura, and pro- and mesoepimera brown to dark brown.
</paragraph>
<paragraph id="ED394975760A4335FF29FA5E185D51D4" blockId="38.[151,1437,151,2013]" lastBlockId="39.[151,1437,151,2013]" lastPageId="39" lastPageNumber="240" pageId="38" pageNumber="239">
Abdomen: Abdominal segment I with 45 black setae on dorsal hump; dorsal hump sometimes with sclerites; ventral of dorsal hump 413 black
<emphasis id="DFF29567760A4334FDC8FA721B7C55B9" box="[550,569,1448,1472]" italics="true" pageId="38" pageNumber="239">ls</emphasis>
setae generally set on sclerotized areas, however setae sometimes with only basal sclerotization; these setae may also be short, reddish, without sclerotization. Dorsal hump sometimes with black
<emphasis id="DFF29567760A4334FF35FA2A19AB5671" box="[219,238,1520,1544]" italics="true" pageId="38" pageNumber="239">ls</emphasis>
setae lateral of meson, setal sockets prominent, setae possibly with sclerotized bases. Lateral humps each with 11
<emphasis id="DFF29567760A4334FF18F9CE184C5655" box="[246,265,1556,1580]" italics="true" pageId="38" pageNumber="239">ls</emphasis>
setae; dorsal areas may have 12 small sclerotized areas with 45 short black setae. Dorsal of lateral hump 6 black
<emphasis id="DFF29567760A4334FED9F9E2180F5629" box="[311,330,1592,1616]" italics="true" pageId="38" pageNumber="239">ls</emphasis>
setae with sclerotized bases and prominent setal sockets. Each lateral hump with 1 long black seta ventrally. Ventral of lateral hump 39
<emphasis id="DFF29567760A4334FD83F9871BC6560D" box="[621,643,1629,1652]" italics="true" pageId="38" pageNumber="239">ss</emphasis>
to
<emphasis id="DFF29567760A4334FD5BF9861B8D560D" box="[693,712,1628,1652]" italics="true" pageId="38" pageNumber="239">ls</emphasis>
black setae sometimes with sclerotized bases; setal sockets prominent. Venter of abdominal segment I with 618 black
<emphasis id="DFF29567760A4334FCBBF95A1A2D56E1" box="[853,872,1664,1688]" italics="true" pageId="38" pageNumber="239">ls</emphasis>
setae forming an elliptical pattern; setal sockets prominent. Venter of abdominal segment I may have pair of sclerites or lightly sclerotized areas, each with 23 black setae, often with basal sclerotization. Ventral abdominal segments IIVIII glabrous. Dorsal and ventral abdominal gills present, each single-filament; present dorsoanteriorly and ventroanteriorly on segments IIIVI; present dorsoposteriorly and ventroposteriorly on segments IIVII (
<figureCitation id="75BD55F0760A4334FC7EF8D51AA05751" box="[912,997,1807,1832]" captionStart="FIGURE 42" captionStartId="51.[151,250,1456,1478]" captionTargetBox="[202,1382,260,1371]" captionTargetPageId="51" captionText="FIGURE 42. Pupal and larval gill schematic diagrams, left lateral, for Ecclisomyia bilera Denning, Ecclisomyia conspersa Banks, Ecclisomyia maculosa Banks, Ecclisomyia simulata Banks. Abbreviations: IVIII = dorsal abdominal segments, with Arabic numerals representing the number of gills per segment in anterior and posterior positions at dorsal, lateral or lateral line (dorsolateral and ventrolateral), and ventral levels." httpUri="https://zenodo.org/record/1227419/files/figure.svg" pageId="38" pageNumber="239">Fig. 42</figureCitation>
). Dorsum of abdomen with short black primary
<emphasis id="DFF29567760A4334FF16F8EF18545735" box="[248,273,1845,1868]" italics="true" pageId="38" pageNumber="239">sa</emphasis>
2 setae on abdominal segments IIVII; primary setae sometimes absent or difficult to discern, may be black or translucent. Dorsum of abdominal segments VII and VIII with short black setae adjacent to
<emphasis id="DFF29567760A4334FAC2F8831C005709" box="[1324,1349,1881,1904]" italics="true" pageId="38" pageNumber="239">sa</emphasis>
2 setae. Tergum IX sclerite brown to yellow brown, elliptical, with 6 black, stout
<emphasis id="DFF29567760A4334FC29F8A61A9F57ED" box="[967,986,1916,1940]" italics="true" pageId="38" pageNumber="239">ls</emphasis>
setae spaced equidistant along posterior margin; surface of sclerite with 56 shorter black setae. Venter of abdominal segment IX with 24 short black setae in
<emphasis id="DFF29567760A4334FF5AF81F198857A5" box="[180,205,1989,2012]" italics="true" pageId="38" pageNumber="239">sa</emphasis>
2 position. Lateral fringe of fine reddish hairs extending on each side from anterior of segment III to anterior of segment VIII. Chloride epithelia present on venter of abdominal segments IIIVII; may not be evident in early instars. Anal prolegs each with lateral sclerite brown; narrowing ventrally and forming light brown sclerotized strap (band or projection) delineating sternum IX from sternum X; with 13 black
<emphasis id="DFF29567760B4335FBD2FF3A1D0A5081" box="[1084,1103,224,248]" italics="true" pageId="39" pageNumber="240">ls</emphasis>
seta on ventrocaudal margin; setal sockets prominent; with 16 black
<emphasis id="DFF29567760B4335FD8CFEDE1B305165" box="[610,629,260,284]" italics="true" pageId="39" pageNumber="240">ls</emphasis>
setae on dorsal surface and with basal tuft of 45 long brown to black setae, of which inner 2 setae longer than shorter outer setae. Sole plates brown to dark brown; heavily sclerotized, each without setae or only bearing single short black seta near distoventral margin. Caudal lobes glabrous except for minute
<emphasis id="DFF29567760B4335FEF2FEAA186A51F1" box="[284,303,368,392]" italics="true" pageId="39" pageNumber="240">sl</emphasis>
reddish spicules on either side of anal opening. Anal hooks each bearing small accessory hook (denticule).
</paragraph>
</subSubSection>
<subSubSection id="A59C1AFE760B4335FF29FE6D1A0852FC" pageId="39" pageNumber="240" type="distribution">
<paragraph id="ED394975760B4335FF29FE6D1A0852FC" blockId="39.[151,1437,151,2013]" pageId="39" pageNumber="240">
<emphasis id="DFF29567760B4335FF29FE6D181C51A9" bold="true" box="[199,345,439,464]" pageId="39" pageNumber="240">Distribution</emphasis>
.
<taxonomicName id="2A8632F6760B4335FE87FE621B2051A9" authorityName="Banks" authorityYear="1907" box="[361,613,440,464]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF29567760B4335FE87FE621B2051A9" box="[361,613,440,464]" italics="true" pageId="39" pageNumber="240">Ecclisomyia maculosa</emphasis>
</taxonomicName>
is confined to the Intermountain Region and the Rocky Mountains, and has been recorded from Alberta, British Columbia, Colorado, Montana, and Wyoming (
<bibRefCitation id="89173484760B4335FB7BFE061C51518C" author="Ross, H. H." box="[1173,1300,476,501]" journalOrPublisher="American Midland Naturalist" pageId="39" pageNumber="240" pagination="410 - 429" part="43" publicationUrl="https://doi.org/10.2307/2421910" refId="ref36907" refString="Ross, H. H. (1950) Synoptic notes on some Nearctic limnephilid caddisflies (Trichoptera: Limnephilidae). American Midland Naturalist, 43 (2), 410 - 429. https: // doi. org / 10.2307 / 2421910" title="Synoptic notes on some Nearctic limnephilid caddisflies (Trichoptera: Limnephilidae)" type="journal article" year="1950">Ross 1950</bibRefCitation>
).
<bibRefCitation id="89173484760B4335FAC5FE06180B5261" author="Newell, R. L. &amp; Minshall, G. W." journalOrPublisher="Great Basin Naturalist" pageId="39" pageNumber="240" pagination="253 - 257" part="37" refId="ref36691" refString="Newell, R. L. &amp; Minshall, G. W. (1977) An annotated list of the aquatic insects of southeastern Idaho, part III Trichoptera. Great Basin Naturalist, 37 (2), 253 - 257." title="An annotated list of the aquatic insects of southeastern Idaho, part III Trichoptera" type="journal article" year="1977">Newell &amp; Minshall (1977)</bibRefCitation>
recorded
<taxonomicName id="2A8632F6760B4335FE2DFDDB1B155261" authorityName="Banks" authorityYear="1907" box="[451,592,512,536]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF29567760B4335FE2DFDDB1B155261" box="[451,592,512,536]" italics="true" pageId="39" pageNumber="240">E. maculosa</emphasis>
</taxonomicName>
from Idaho. The distribution record for
<taxonomicName id="2A8632F6760B4335FBCCFDDB1DEA5261" authorityName="Banks" authorityYear="1907" box="[1058,1199,512,536]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF29567760B4335FBCCFDDB1DEA5261" box="[1058,1199,512,536]" italics="true" pageId="39" pageNumber="240">E. maculosa</emphasis>
</taxonomicName>
reported for Oregon (
<bibRefCitation id="89173484760B4335FF70FDFE18005244" author="Denning, D. G." box="[158,325,548,573]" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="39" pageNumber="240" pagination="157 - 162" part="24" refId="ref35999" refString="Denning, D. G. (1951) Records and descriptions of Nearctic caddis flies, Part III. Journal of the Kansas Entomological Society, 24 (4), 157 - 162." title="Records and descriptions of Nearctic caddis flies, Part III" type="journal article" year="1951">Denning 1951</bibRefCitation>
) is
<taxonomicName id="2A8632F6760B4335FE98FDFF18BF5245" baseAuthorityName="Givens" baseAuthorityYear="2014" box="[374,506,548,572]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="simulata">
<emphasis id="DFF29567760B4335FE98FDFF18BF5245" box="[374,506,548,572]" italics="true" pageId="39" pageNumber="240">E. simulata</emphasis>
</taxonomicName>
, as reported by
<bibRefCitation id="89173484760B4335FD50FDFF1A0B5244" author="Ross, H. H." box="[702,846,548,573]" journalOrPublisher="American Midland Naturalist" pageId="39" pageNumber="240" pagination="410 - 429" part="43" publicationUrl="https://doi.org/10.2307/2421910" refId="ref36907" refString="Ross, H. H. (1950) Synoptic notes on some Nearctic limnephilid caddisflies (Trichoptera: Limnephilidae). American Midland Naturalist, 43 (2), 410 - 429. https: // doi. org / 10.2307 / 2421910" title="Synoptic notes on some Nearctic limnephilid caddisflies (Trichoptera: Limnephilidae)" type="journal article" year="1950">Ross (1950)</bibRefCitation>
. I have examined specimens of
<taxonomicName id="2A8632F6760B4335FB22FDFF1C1E5245" authorityName="Banks" authorityYear="1907" box="[1228,1371,548,572]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF29567760B4335FB22FDFF1C1E5245" box="[1228,1371,548,572]" italics="true" pageId="39" pageNumber="240">E. maculosa</emphasis>
</taxonomicName>
from Alberta, British Columbia, Colorado, Idaho, Montana, and Wyoming. The specimens of
<taxonomicName id="2A8632F6760B4335FB90FD931C4F5219" authorityName="Banks" authorityYear="1907" box="[1150,1290,584,608]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF29567760B4335FB90FD931C4F5219" box="[1150,1290,584,608]" italics="true" pageId="39" pageNumber="240">E. maculosa</emphasis>
</taxonomicName>
from British Columbia have been from the southeastern Rocky Mountains.
</paragraph>
</subSubSection>
<subSubSection id="A59C1AFE760B4335FF29FD5518245394" pageId="39" pageNumber="240" type="biology_ecology">
<paragraph id="ED394975760B4335FF29FD5518245394" blockId="39.[151,1437,151,2013]" pageId="39" pageNumber="240">
<emphasis id="DFF29567760B4335FF29FD55180652D1" bold="true" box="[199,323,655,680]" pageId="39" pageNumber="240">Bionomics</emphasis>
. In Colorado,
<taxonomicName id="2A8632F6760B4335FE17FD4B1BCE52D1" authorityName="Banks" authorityYear="1907" box="[505,651,656,680]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF29567760B4335FE17FD4B1BCE52D1" box="[505,651,656,680]" italics="true" pageId="39" pageNumber="240">E. maculosa</emphasis>
</taxonomicName>
appears to be primarily a subalpine to alpine species, frequently collected at altitudes of 24384314 m.
<bibRefCitation id="89173484760B4335FD8DFD6F1A3452B4" author="Herrmann, S. J. &amp; Ruiter, D. E. &amp; Unzicker, J. D." box="[611,881,692,717]" journalOrPublisher="The Southwestern Naturalist" pageId="39" pageNumber="240" pagination="4" part="31" publicationUrl="https://doi.org/10.2307/3671699" refId="ref36243" refString="Herrmann, S. J., Ruiter, D. E. &amp; Unzicker, J. D. (1986) Distribution and records of Colorado Trichoptera. The Southwestern Naturalist, 31 (4), 421 - - 457. https: // doi. org / 10.2307 / 3671699" title="Distribution and records of Colorado Trichoptera" type="journal article" year="1986">
Herrmann
<emphasis id="DFF29567760B4335FD31FD6F1A5952B5" box="[735,796,692,716]" italics="true" pageId="39" pageNumber="240">et al.</emphasis>
(1986)
</bibRefCitation>
reported an altitude range of 25583377 m. In Alberta,
<taxonomicName id="2A8632F6760B4335FF16FD0318C75289" authorityName="Banks" authorityYear="1907" box="[248,386,728,752]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF29567760B4335FF16FD0318C75289" box="[248,386,728,752]" italics="true" pageId="39" pageNumber="240">E. maculosa</emphasis>
</taxonomicName>
has been recorded from an altitudinal range of 13712134 m (
<bibRefCitation id="89173484760B4335FBD4FD0D1D915289" author="Nimmo, A." box="[1082,1236,727,752]" journalOrPublisher="Quaestiones Entomologicae" pageId="39" pageNumber="240" pagination="3 - 234" part="7" refId="ref36774" refString="Nimmo, A. (1971) The adult Rhyacophilidae and Limnephilidae (Trichoptera) of Alberta and Eastern British Columbia and their post-glacial origin. Quaestiones Entomologicae, 7 (1), 3 - 234." title="The adult Rhyacophilidae and Limnephilidae (Trichoptera) of Alberta and Eastern British Columbia and their post-glacial origin" type="journal article" year="1971">Nimmo 1971</bibRefCitation>
). This species has been collected from a shallow, narrow, swiftly flowing mountain stream (
<bibRefCitation id="89173484760B4335FC26FD261D28536D" author="Denning, D. G." box="[968,1133,764,789]" journalOrPublisher="Psyche" pageId="39" pageNumber="240" pagination="16 - 27" part="55" publicationUrl="https://doi.org/10.1155/1948/71426" refId="ref35958" refString="Denning, D. G. (1948) New and little known species of Nearctic Trichoptera. Psyche, 55 (1), 16 - 27. https: // doi. org / 10.1155 / 1948 / 71426" title="New and little known species of Nearctic Trichoptera" type="journal article" year="1948">Denning 1948</bibRefCitation>
).
<bibRefCitation id="89173484760B4335FB6EFD261C63536D" author="Nimmo, A." box="[1152,1318,764,789]" journalOrPublisher="Quaestiones Entomologicae" pageId="39" pageNumber="240" pagination="3 - 234" part="7" refId="ref36774" refString="Nimmo, A. (1971) The adult Rhyacophilidae and Limnephilidae (Trichoptera) of Alberta and Eastern British Columbia and their post-glacial origin. Quaestiones Entomologicae, 7 (1), 3 - 234." title="The adult Rhyacophilidae and Limnephilidae (Trichoptera) of Alberta and Eastern British Columbia and their post-glacial origin" type="journal article" year="1971">Nimmo (1971)</bibRefCitation>
noted that this species occurs in “small, riffled mountain creeks, running gently over fine gravel bottoms.”
<taxonomicName id="2A8632F6760B4335FAFDFCFA18465325" authorityName="Banks" authorityYear="1907" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF29567760B4335FAFDFCFA18465325" italics="true" pageId="39" pageNumber="240">Ecclisomyia maculosa</emphasis>
</taxonomicName>
may be widely sympatric with
<taxonomicName id="2A8632F6760B4335FD87FC9F1BBE5325" box="[617,763,837,860]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="conspersa">
<emphasis id="DFF29567760B4335FD87FC9F1BBE5325" box="[617,763,837,860]" italics="true" pageId="39" pageNumber="240">E. conspersa</emphasis>
</taxonomicName>
. I have examined one female of
<taxonomicName id="2A8632F6760B4335FB9FFC9F1DB85325" authorityName="Banks" authorityYear="1907" box="[1137,1277,836,860]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF29567760B4335FB9FFC9F1DB85325" box="[1137,1277,836,860]" italics="true" pageId="39" pageNumber="240">E. maculosa</emphasis>
</taxonomicName>
and one male specimen of
<taxonomicName id="2A8632F6760B4335FEC4FCB318F853F9" box="[298,445,873,896]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="conspersa">
<emphasis id="DFF29567760B4335FEC4FCB318F853F9" box="[298,445,873,896]" italics="true" pageId="39" pageNumber="240">E. conspersa</emphasis>
</taxonomicName>
from Canyon Creek at the USFS trailhead west of Hamilton, Montana, and 1 male
<taxonomicName id="2A8632F6760B4335FA6AFCB3184C53DD" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="conspersa">
<emphasis id="DFF29567760B4335FA6AFCB3184C53DD" italics="true" pageId="39" pageNumber="240">E. conspersa</emphasis>
</taxonomicName>
and 1 male
<taxonomicName id="2A8632F6760B4335FE61FC571B5C53DD" authorityName="Banks" authorityYear="1907" box="[399,537,908,932]" class="Insecta" family="Limnephilidae" genus="Ecclisomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="DFF29567760B4335FE61FC571B5C53DD" box="[399,537,908,932]" italics="true" pageId="39" pageNumber="240">E. maculosa</emphasis>
</taxonomicName>
from Tindall Meadows 12.9 km S of Landmark, Idaho. I am aware of only these two localities where both species are sympatric. Additional collecting may indicate other sites of sympatry for these two species.
</paragraph>
</subSubSection>
<subSubSection id="A59C1AFE760B433AFF29FC2D1AA15341" lastPageId="40" lastPageNumber="241" pageId="39" pageNumber="240" type="materials_examined">
<paragraph id="ED394975760B433AFF29FC2D1AA15341" blockId="39.[151,1437,151,2013]" lastBlockId="40.[151,1437,151,824]" lastPageId="40" lastPageNumber="241" pageId="39" pageNumber="240">
<emphasis id="DFF29567760B4335FF29FC2D1BED5469" bold="true" box="[199,680,1015,1040]" pageId="39" pageNumber="240">Material Examined. CANADA: Alberta,</emphasis>
sedge marsh, 1.6 km S Bow Pass, (Hwy. 93), Banff Natl. Park, 25- vii-1966, 1 M (APN) (SMEC); Lake Agnes, Lake Louise, Banff Natl. Park, 10-vii-1967, 1 M, 1F (APN) (SMEC); 10-viii-1967, 7 M, 1 F (APN) (SMEC); Lynx Creek, Forest Trail Rd., 14-vii-1967, 2 M (APN) (SMEC); Bow Pass, Banff. Natl. Park, 10-viii-1967, 1 M (APN) (SMEC); Snaring River, N of Jasper, (Hwy. 16), Jasper Natl. Park, 15-vii-1968, 1 M (APN) (SMEC); Poboktan Creek, (Hwy. 93), Jasper Natl. Park, 06-viii-1968, 1 M (APN) (SMEC); Yara Creek, Forest Trail Rd., 6.4 km N of Red Deer River Crossing, 13-ix-1968, 1 M (APN) (SMEC); Crowsnest Pass, 19-vii-2003, 3 M (PAO) (CSUC).
<emphasis id="DFF29567760B4335FD30FB151AEA5491" bold="true" box="[734,943,1231,1256]" pageId="39" pageNumber="240">British Columbia</emphasis>
, Fairy Creek, Fernie, (Hwy. 3), Elk Valley, 12-viii-1967, 1 M (APN) (SMEC).
<emphasis id="DFF29567760B4335FDC4FB2E1ACF5574" bold="true" box="[554,906,1268,1293]" pageId="39" pageNumber="240">COLORADO: Boulder Co., t</emphasis>
ributary to N. Fork Boulder Creek, (Forest Rd. 298), 23-vii-2013, 1 F (CJV) (CSUC); Middle Fork Vrain River, Peaceful Valley, 30-vii-2013, 3 F (CJV) (CSUC); Como Creek, (County Forest Rd. 116 and Forest Rd. 248), 04-viii-2013, 1 M, 1 F (CJV) (CSUC); Middle Fork Vrain River, Peaceful Valley, N40.047 W105.877, 09-vii-2015, 1 M (T. McNary) (CSUC).
<emphasis id="DFF29567760B4335FB60FA851C0C5501" bold="true" box="[1166,1353,1375,1400]" pageId="39" pageNumber="240">Clear Creek Co</emphasis>
., Quail Creek, nr. Jct. of Grizzly Gulch and Stevens Gulch Roads, 22-vii-2013, 2 M (S. Fitzgerald) (CSUC).
<emphasis id="DFF29567760B4335FACEFA5E1CCB55E4" bold="true" box="[1312,1422,1412,1437]" pageId="39" pageNumber="240">Eagle Co</emphasis>
., Beaver Creek, Beaver Lake Trail, 20-vii-2014, 3 M 6 F (S. Fitzgerald) (CSUC).
<emphasis id="DFF29567760B4335FBF8FA7D1DE655B9" bold="true" box="[1046,1187,1447,1472]" pageId="39" pageNumber="240">Garfield Co</emphasis>
., stream, below Slide, Mandell Lake, 01-viii-1982, 3 M P, 3 F P, 18 P (B. Brooks) (DERPC).
<emphasis id="DFF29567760B4335FC51FA161D73559C" bold="true" box="[959,1078,1484,1509]" pageId="39" pageNumber="240">Gilpin Co</emphasis>
., Inlet stream, Crater Lake, 29- vii-1991, 2 F P, 4 P (D.E. Rathke) (DERPC).
<emphasis id="DFF29567760B4335FD76FA351A545671" bold="true" box="[664,785,1519,1544]" pageId="39" pageNumber="240">Grand Co</emphasis>
., Tonahutu Creek, 19-vii-1988, 2 M, 4 F (BCK) (CSUC); Tonahutu Creek, above Big Meadows, 23-vii-1988, 7 M (M. Harris) (CSUC); Timber Creek, Back Country Cmpgd., 26-vii-1989, 1 M, 4 F (Barker and M. Harris) (CSUC); Colorado River, trailhead, Rocky Mt. Nat'l. Park, 25-vii-1997, 4 M, 7 F (PAO and E. Buckner) (CSUC).
<emphasis id="DFF29567760B4335FCE2F9861A0C560C" bold="true" box="[780,841,1628,1653]" pageId="39" pageNumber="240">Lake</emphasis>
Co., Lake Creek, 7.2 km below Independence Pass, (Hwy. 82), N39.192 W106.8968, 12-vi-2013, 3 L (BDH) (CSUC); N. Fork Lake Creek, off Rd. 826, 0.8 km S of Hwy. 82, 20-v-2013, 5 L (BDH) (CSUC); Busk Creek, Meadows, 0.9 km W Turquoise Lake, N39.444 W106.742, 01-vi-2013, 1 L (BDH) (CSUC); 16-vi-2013, 1 L (BDH) (CSUC).
<emphasis id="DFF29567760B4335FC6EF91D1D4B5699" bold="true" box="[896,1038,1735,1760]" pageId="39" pageNumber="240">Larimer Co</emphasis>
., Icy Brook above the Loch, Rocky Mt. Nat'l. Park, 03-viii-1986, 2 M, (BCK) (CSUC); 06-vii-1988, 3 M (BCK) (CSUC); Fall River, Endovalley Picnic Grounds, Rocky Mt. Nat'l. Park, 22-vii-1987, 2 M (BCK) (CSUC); Fall River 1.6 km E Hidden Valley Rd., Rocky Mt. Natl. Park, N40.4038 W105.6158, 16-vi-2007, 1 L (BDH) (CSUC); Spring Creek, Rocky Mt. Nat'l. Park, 29-vii-1989, 1 M (M. Harris) (CSUC); Spruce Creek, Rocky Mt. Nat'l. Park, 29-vii-1989, 1 M (M. Harris) (CSUC); Endovalley Picnic Area, Rocky Mt. Nat'l. Park, 13-vii-1993, 5 M (PAO) (CSUC); 07-viii-1995, 21 M, 1 F (D. Katz) (CSUC); Kern Lake Trailhead, Rocky Mt. Nat'l. Park, 22-vii-1998, 1 M (S. Simonson) (CSUC); S. Fork Poudre River, 03-viii-2003, 1 M (D. Finn) (CSUC).
<emphasis id="DFF29567760B4335FD1EF81E1AC857A4" bold="true" box="[752,909,1988,2013]" pageId="39" pageNumber="240">Saguache Co</emphasis>
., Jones Creek, at Jones Creek Cmpgd., 12-vi- 2005, 3 M (BCK and R. Baumann) (CSUC).
<emphasis id="DFF295677604433AFD70FF4D1A6250C9" bold="true" box="[670,807,151,176]" pageId="40" pageNumber="241">Summit Co</emphasis>
., Boulder Creek, 3.2 km N of Rt. 9, 03-viii-1995, 1 M (BCK) (CSUC); 10 Mile Creek, nr. I-70 crossing, 01-viii-2014, 5 M, 3 F (S. Fitzgerald) (CSUC).
<emphasis id="DFF295677604433AFB1DFF661C1750AC" bold="true" box="[1267,1362,188,213]" pageId="40" pageNumber="241">IDAHO</emphasis>
:
<emphasis id="DFF295677604433AFA8CFF6618455081" bold="true" pageId="40" pageNumber="241">Bear Lake Co</emphasis>
., Eight Mile Creek, (Eight Mile Creek Rd.) 24 km S Soda Springs, 30-vi-2007, 1 L (M. Kippenhan) (CSUC).
<emphasis id="DFF295677604433AFEEEFEDE18335164" bold="true" box="[256,374,260,285]" pageId="40" pageNumber="241">Lemhi Co</emphasis>
., 11.3 km E Leadore, 26-vi-1965, 1 M (SDS and E.R. Logan) (DRGPC).
<emphasis id="DFF295677604433AFB47FEDE1C065164" bold="true" box="[1193,1347,260,285]" pageId="40" pageNumber="241">Shoshone Co</emphasis>
., Jct. of Boulder and Marble creeks, 19.3 km E Colder, 04-vii-1964, 8 L (SDS) (DRGPC).
<emphasis id="DFF295677604433AFBA3FEFD1D855139" bold="true" box="[1101,1216,295,320]" pageId="40" pageNumber="241">Valley Co</emphasis>
., Tindall meadows 12.9 km S Landmark, 22-vii-1965, 1 M (SDS and E.R. Logan) (WFBM).
<emphasis id="DFF295677604433AFC1BFE961DC2511C" bold="true" box="[1013,1159,332,357]" pageId="40" pageNumber="241">MONTANA</emphasis>
:
<emphasis id="DFF295677604433AFB76FE961C7B511C" bold="true" box="[1176,1342,332,357]" pageId="40" pageNumber="241">Flat Head Co</emphasis>
., Upper Kintla Cr., Glacier Natl. Park, 01-viii-1979, 3 M (J. Stanford) (FLBS); Upper Kintla Lake, Glacier Natl. Park, 01- viii-1979, 1 M (J. Stanford) (FLBS); Picnic Lakes, Jewel Basin, 07-vii-1987, 4 M (FLBS); Twin Springs, Nyack, 25-vii-2006 to 03-viii-2006, 9 M 44 F (M. Anderson) (FLBS); stream along Hwy. 2, at milemarker 193.5 (km 311.4), 03-vi-1996, 3 PP (DER) (DRGPC).
<emphasis id="DFF295677604433AFD6DFE061A4A518C" bold="true" box="[643,783,476,501]" pageId="40" pageNumber="241">Gallatin Co</emphasis>
., Swann Creek, Swann Creek Rd., N45.3730 W111.1710, 06-viii-2015, 2 F (BCK and CJV) (CSUC).
<emphasis id="DFF295677604433AFD68FE251BBA5261" bold="true" box="[646,767,511,536]" pageId="40" pageNumber="241">Ravilli Co</emphasis>
., Canyon Creek, at USFS trailhead W of Hamilton, 30-vii- 2008, 1 F (R. Durfee) (CSUC); Daly Creek, 3.2 km upstream from Jct. with Skalkaho Creek, (Route 38), 05-viii- 2015 1 F (BCK, R. Durfee, and CJV) (CSUC).
<emphasis id="DFF295677604433AFD2EFD9D1A105219" bold="true" box="[704,853,583,608]" pageId="40" pageNumber="241">WYOMING</emphasis>
:
<emphasis id="DFF295677604433AFC89FD9D1AAF5219" bold="true" box="[871,1002,583,608]" pageId="40" pageNumber="241">Albany Co</emphasis>
., Meadow Creek, West Glacier Lake, Medicine Bow Nat'l. Forest, 15-vii-1987, 1 M (H. Copeland) (CSUC); West Glacier Lake, Medicine Bow Nat'l. Forest, 21-vii-1987, 4 F (BCK and W. Painter) (CSUC); Glacier Lakes, Medicine Bow Nat'l. Forest, 25-vii-1990, 9 M, 6 F (L. Bjostad, W. Painter, and D. Jewett) (CSUC).
<emphasis id="DFF295677604433AFCE0FD6E1ADA52B4" bold="true" box="[782,927,692,717]" pageId="40" pageNumber="241">Fremont Co</emphasis>
., McKenzie Highland Ranch NW of Dubois, 24-vi-1999, 1 M (K. Bagdonis) (CSUC).
<emphasis id="DFF295677604433AFD8DFD0D1BAA5289" bold="true" box="[611,751,727,752]" pageId="40" pageNumber="241">Johnson Co</emphasis>
., Middle Fork Clear Creek, (Forest Service Rd. 323 off Hwy. 16) (Middle Fork Cmpgd.), 30-vii-2010, 2 M, 3 F (BCK) (CSUC).
<emphasis id="DFF295677604433AFC67FD261AB0536C" bold="true" box="[905,1013,764,789]" pageId="40" pageNumber="241">Teton Co</emphasis>
., N. Fork Trail Creek, (Hwy. 22, E of Teton Pass), 09-vi-1997, 4 M, 2 M P, 1 F P, 7 P, 7 PP, 2 L (DER) (DERPC).
</paragraph>
</subSubSection>
</treatment>
</document>