treatments-xml/data/57/CA/A5/57CAA5C1803E59B988ACEA5926F98E8C.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1030.62544" ID-GBIF-Dataset="91fec855-7fa9-4ad6-84d7-9b9d35e23e8b" ID-PMC="PMC8060247" ID-Pensoft-Pub="1313-2970-1030-1" ID-Pensoft-UUID="875199397EEE5F7898EA1DB25DA62D25" ID-PubMed="33958904" ModsDocID="1313-2970-1030-1" checkinTime="1618371561543" checkinUser="pensoft" docAuthor="Means, Jackson C., Hennen, Derek A. &amp; Marek, Paul E." docDate="2021" docId="57CAA5C1803E59B988ACEA5926F98E8C" docLanguage="en" docName="ZooKeys 1030: 1-180" docOrigin="ZooKeys 1030" docPubDate="2021-04-13" docSource="http://dx.doi.org/10.3897/zookeys.1030.62544" docTitle="Nannaria hippopotamus Means &amp; Hennen &amp; Marek 2021, sp. nov." docType="treatment" docVersion="5" id="875199397EEE5F7898EA1DB25DA62D25" lastPageNumber="1" masterDocId="875199397EEE5F7898EA1DB25DA62D25" masterDocTitle="A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)" masterLastPageNumber="180" masterPageNumber="1" pageNumber="1" updateTime="1673961214978" updateUser="julia">
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<mods:titleInfo>
<mods:title>A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Means, Jackson C.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7377-0696</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
<mods:nameIdentifier type="email">mjacks4@vt.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hennen, Derek A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7005-1151</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marek, Paul E.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7048-2514</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1030</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>180</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1030.62544</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1030.62544</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1030-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">875199397EEE5F7898EA1DB25DA62D25</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="181004740" LSID="urn:lsid:zoobank.org:act:AFE8AC4E-F463-4AA3-A9F6-4D0B4C16DCC9" httpUri="http://treatment.plazi.org/id/57CAA5C1803E59B988ACEA5926F98E8C" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName ID-CoL="8RVTN" LSID="http://zoobank.org/AFE8AC4E-F463-4AA3-A9F6-4D0B4C16DCC9" authority="Means &amp; Hennen &amp; Marek, 2021" authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria hippopotamus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hippopotamus" status="sp. nov.">Nannaria hippopotamus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 100" captionStartId="F100" captionText="Figure 100. Nannaria hippopotamus sp. nov. paratype ♂ (NCSM, NAN 0469) left gonopod A anterior view; red arrow indicates lateral flange; red triangle indicates acropodite medial swelling B medial view C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure100" httpUri="https://binary.pensoft.net/fig/531203" pageId="0" pageNumber="1">Figs 100</figureCitation>
<figureCitation captionStart="Figure 101" captionStartId="F101" captionText="Figure 101. Nannaria hippopotamus sp. nov. non-type ♂ (VTEC, MPE 04141) coloration. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1030.62544.figure101" httpUri="https://binary.pensoft.net/fig/531204" pageId="0" pageNumber="1">, 101 Vernacular name: &quot;The Horse Creek Twisted-Claw Millipede&quot;</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3071355336" accessionNumber="MPE04150" collectingDate="2018-06-16" collectorName="D. Hennen" country="United States" county="Greene County" elevation="581" latitude="36.1059" location="Horse Creek Rec Area" longLatPrecision="6" longitude="-82.6545" municipality="Cherokee National Forest" specimenCount="♂" stateProvince="Tennessee" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
</emphasis>
<collectingCountry name="United States of America">United States</collectingCountry>
- 
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Tennessee">Tennessee</collectingRegion>
</emphasis>
• 
<specimenCount>♂</specimenCount>
; 
<collectingCounty>Greene County</collectingCounty>
, 
<collectingMunicipality>Cherokee National Forest</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:57CAA5C1803E59B988ACEA5926F98E8C:BC19303FB9684ED78C382CF03801417D" country="United States" county="Greene County" latitude="36.1059" longLatPrecision="6" longitude="-82.6545" municipality="Cherokee National Forest" name="Horse Creek Rec Area" stateProvince="Tennessee">Horse Creek Rec Area</location>
, picnic area on east side of 
<location LSID="urn:lsid:plazi:treatment:57CAA5C1803E59B988ACEA5926F98E8C:A1CC0929A09DC28E964AC11D6ABEF01B" country="United States" county="Greene County" latitude="36.1059" longLatPrecision="6" longitude="-82.6545" municipality="Cherokee National Forest" name="Horse Creek" stateProvince="Tennessee">Horse Creek</location>
; 
<geoCoordinate degrees="36.1059" direction="north" orientation="latitude" precision="5" value="36.1059">36.1059°N</geoCoordinate>
, - 
<geoCoordinate degrees="82.6545" direction="west" orientation="longitude" precision="5" value="-82.6545">82.6545°W</geoCoordinate>
; elev. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.81" unit="m" value="581.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.81" unit="m" value="581.0">581 m</elevation>
</quantity>
; 
<collectingDate value="2018-06-16">16 June 2018</collectingDate>
; hand collected; 
<collectorName>D. Hennen</collectorName>
leg.; VTEC 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE04150">MPE04150</accessionNumber>
.
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3071355586" accessionNumber="MPE04151" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="D. Hennen" country="United States" county="Greene County" elevation="581" latitude="36.1059" location="Tennessee" longLatPrecision="6" longitude="-82.6545" municipality="Cherokee National Forest" specimenCount="3" specimenCount-female="3" stateProvince="Tennessee" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">
<typeStatus>Paratypes</typeStatus>
</emphasis>
:
</emphasis>
<collectingCountry name="United States of America">United States</collectingCountry>
- 
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Tennessee">Tennessee</collectingRegion>
</emphasis>
• 
<specimenCount type="female">3 ♀</specimenCount>
; same collection data as holotype; VTEC 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE04151">MPE04151</accessionNumber>
</materialsCitation>
-53 • 
<materialsCitation ID-GBIF-Occurrence="3071355433" accessionNumber="MPE04154" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="D. Hennen" country="United States" county="Greene County" elevation="581" latitude="36.1059" location="Horse Creek Rec Area" longLatPrecision="6" longitude="-82.6545" municipality="Cherokee National Forest" specimenCount="3" specimenCount-female="3" stateProvince="Tennessee" typeStatus="paratype">
<specimenCount type="female">3 ♀♀</specimenCount>
; same collection data as holotype; VMNH 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE04154">MPE04154</accessionNumber>
</materialsCitation>
, 55, 4233 • 
<materialsCitation ID-GBIF-Occurrence="3071355758" accessionNumber="MPE04150" collectingDate="1978-10-16" collectorName="R. Shelley, W. Jones" country="United States" county="Greene County" elevation="581" latitude="36.1059" location="Horse Creek Rec Area" longLatPrecision="6" longitude="-82.6545" municipality="Cherokee National Forest" specimenCount="1" specimenCount-male="1" stateProvince="Tennessee" typeStatus="paratype">
<specimenCount type="male">1 ♂</specimenCount>
; same collection data as holotype; 
<collectingDate value="1978-10-16">16 Oct. 1978</collectingDate>
; 
<collectorName>R. Shelley</collectorName>
, 
<collectorName>W. Jones</collectorName>
leg.; NCSM NAN0469
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3071355388" accessionNumber="MPE04141" collectingDate="2018-06-15" collectorName="D. Hennen" country="United States" county="Greene County" elevation="623" latitude="36.0878" location="Cherokee National Forest" longLatPrecision="6" longitude="-82.6811" municipality="Horse Creek" specimenCount="1" specimenCount-male="1" stateProvince="Tennessee">
<collectingCountry name="United States of America">United States</collectingCountry>
- 
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Tennessee">Tennessee</collectingRegion>
</emphasis>
• 
<specimenCount type="male">1 ♂</specimenCount>
; 
<collectingCounty>Greene County</collectingCounty>
, 8.5 air km south of 
<collectingMunicipality>Horse Creek</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:57CAA5C1803E59B988ACEA5926F98E8C:0D62AE3FCA3A82B29CEC0340F37EBB01" country="United States" county="Greene County" latitude="36.0878" longLatPrecision="6" longitude="-82.6811" municipality="Horse Creek" name="Cherokee National Forest" stateProvince="Tennessee">Cherokee National Forest</location>
, 
<location LSID="urn:lsid:plazi:treatment:57CAA5C1803E59B988ACEA5926F98E8C:C797B466856298FF3BDA02DEEA6CBA0C" country="United States" county="Greene County" latitude="36.0878" longLatPrecision="6" longitude="-82.6811" municipality="Horse Creek" name="Old Forge Recreation Area" stateProvince="Tennessee">Old Forge Recreation Area</location>
; 
<geoCoordinate degrees="36.0878" direction="north" orientation="latitude" precision="5" value="36.0878">36.0878°N</geoCoordinate>
, - 
<geoCoordinate degrees="82.6811" direction="west" orientation="longitude" precision="5" value="-82.6811">82.6811°W</geoCoordinate>
; elev. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.2299999999999995" unit="m" value="623.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.2299999999999995" unit="m" value="623.0">623 m</elevation>
</quantity>
; 
<collectingDate value="2018-06-15">15 June 2018</collectingDate>
; hand collected; 
<collectorName>D. Hennen</collectorName>
leg.; VTEC 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE04141">MPE04141</accessionNumber>
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="3071355351" accessionNumber="MPE04142" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="D. Hennen" country="United States" county="Greene County" elevation="623" latitude="36.0878" location="Cherokee National Forest" longLatPrecision="6" longitude="-82.6811" municipality="Horse Creek" specimenCount="1" specimenCount-female="1" stateProvince="Tennessee">
<specimenCount type="female">1 ♀</specimenCount>
; same collection data as preceding; VTEC 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE04142">MPE04142</accessionNumber>
</materialsCitation>
. For detailed collection data see Suppl. material 7.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Adult males of 
<taxonomicName authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria hippopotamus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hippopotamus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria hippopotamus</emphasis>
</taxonomicName>
sp. nov. are distinct from other 
<taxonomicName authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
</taxonomicName>
, and the nearby 
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. scutellaria" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="scutellaria">
<emphasis italics="true" pageId="0" pageNumber="1">N. scutellaria</emphasis>
</taxonomicName>
, based on the following combination of characters: 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Gonopods</emphasis>
.
</emphasis>
Gonopodal acropodite straight, curving medially at apex. Acropodite tip with prominent lobed lateral flange (Fig. 
<figureCitation captionStart="Figure 100" captionStartId="F100" captionText="Figure 100. Nannaria hippopotamus sp. nov. paratype ♂ (NCSM, NAN 0469) left gonopod A anterior view; red arrow indicates lateral flange; red triangle indicates acropodite medial swelling B medial view C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure100" httpUri="https://binary.pensoft.net/fig/531203" pageId="0" pageNumber="1">100A</figureCitation>
, red arrow), curving dorsomedially, not simple as in 
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. terricola" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="terricola">
<emphasis italics="true" pageId="0" pageNumber="1">N. terricola</emphasis>
</taxonomicName>
, or with tooth-like medial flange as in 
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. scutellaria" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="scutellaria">
<emphasis italics="true" pageId="0" pageNumber="1">N. scutellaria</emphasis>
</taxonomicName>
. Acropodite swollen medially before apex (Fig. 
<figureCitation captionStart="Figure 100" captionStartId="F100" captionText="Figure 100. Nannaria hippopotamus sp. nov. paratype ♂ (NCSM, NAN 0469) left gonopod A anterior view; red arrow indicates lateral flange; red triangle indicates acropodite medial swelling B medial view C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure100" httpUri="https://binary.pensoft.net/fig/531203" pageId="0" pageNumber="1">100A</figureCitation>
, red triangle). Telopodite basal zone ca. 1/2 length of acropodite, not ca. 1/4 as in 
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. scutellaria" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="scutellaria">
<emphasis italics="true" pageId="0" pageNumber="1">N. scutellaria</emphasis>
</taxonomicName>
. Telopodite basal zone with very slight lateral bulge, not prominent as in 
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. terricola" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="terricola">
<emphasis italics="true" pageId="0" pageNumber="1">N. terricola</emphasis>
</taxonomicName>
. Prefemur with straight, acicular prefemoral process, not curving ventrally as in 
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. scutellaria" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="scutellaria">
<emphasis italics="true" pageId="0" pageNumber="1">N. scutellaria</emphasis>
</taxonomicName>
. Prefemoral spine small, sharp, not fused with prefemoral process as in 
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. terricola" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="terricola">
<emphasis italics="true" pageId="0" pageNumber="1">N. terricola</emphasis>
</taxonomicName>
. 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Color</emphasis>
.
</emphasis>
Tergites with orange paranotal spots (Fig. 
<figureCitation captionStart="Figure 101" captionStartId="F101" captionText="Figure 101. Nannaria hippopotamus sp. nov. non-type ♂ (VTEC, MPE 04141) coloration. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1030.62544.figure101" httpUri="https://binary.pensoft.net/fig/531204" pageId="0" pageNumber="1">101</figureCitation>
). Black background. Dorsum of collum smooth with orange margin.
</paragraph>
<caption doi="10.3897/zookeys.1030.62544.figure100" httpUri="https://binary.pensoft.net/fig/531203" pageId="0" pageNumber="1" start="Figure 100" startId="F100">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 100.</emphasis>
<taxonomicName authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria hippopotamus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hippopotamus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria hippopotamus</emphasis>
</taxonomicName>
sp. nov. paratype ♂ (NCSM, NAN0469) left gonopod 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
anterior view; red arrow indicates lateral flange; red triangle indicates acropodite medial swelling 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
medial view 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
posterior view. Scale bar: 0.5 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1030.62544.figure101" httpUri="https://binary.pensoft.net/fig/531204" pageId="0" pageNumber="1" start="Figure 101" startId="F101">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 101.</emphasis>
<taxonomicName authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria hippopotamus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hippopotamus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria hippopotamus</emphasis>
</taxonomicName>
sp. nov. non-type ♂ (VTEC, MPE04141) coloration. Scale bar: 4.0 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="1">♂ holotype (VTEC, MPE04150): BL = 30.4, CW = 4.2, IW = 2.3, ISW = 1.0, B11W = 4.9, B11H = 3.3; ♀ paratype (VTEC, MPE04152): BL = 31.2, CW - 4.1, IW = 2.3, ISW = 1.0, B11W = 5.3, B11H = 4.0.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
<paragraph pageId="0" pageNumber="1">No known variation.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Known only from two campsites in eastern Tennessee (Tennessee: Greene County; Suppl. material 5; Fig. 
<figureCitation captionStart="Figure 126" captionStartId="F126" captionText="Figure 126. Distribution map of Nannaria minor species group collection localities in the central Appalachians." figureDoi="10.3897/zookeys.1030.62544.figure126" httpUri="https://binary.pensoft.net/fig/531229" pageId="0" pageNumber="1">126</figureCitation>
). Distribution area: N/A; status: MRE.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Individuals of 
<taxonomicName authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria hippopotamus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hippopotamus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria hippopotamus</emphasis>
</taxonomicName>
sp. nov. have been found under moist leaf litter in mesic broadleaved forests dominated by sweetgum, hickory, hemlock, oak, rhododendron, and maple.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is named for its type locality, Horse Creek, a national forest recreation area hardwood cove. The specific name is a noun in apposition derived from the Greek 
<emphasis italics="true" pageId="0" pageNumber="1">hippos</emphasis>
-, meaning horse, and 
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="potamós">potamos</normalizedToken>
</emphasis>
, meaning river.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
United States, Tennessee, Greene County, Cherokee National Forest, Horse Creek Rec Area, picnic area on east side, 
<geoCoordinate degrees="36.1059" direction="north" orientation="latitude" precision="5" value="36.1059">36.1059°N</geoCoordinate>
, - 
<geoCoordinate degrees="82.6545" direction="west" orientation="longitude" precision="5" value="-82.6545">82.6545°W</geoCoordinate>
.
</paragraph>
</subSubSection>
</treatment>
</document>