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<mods:title id="55B60EE38E1A3945AB68EF1309EC3503">Redescription of Ryukyua circularis (Pillai, 1954) (Isopoda, Cymothoidae), parasite of the Bleeker smoothbelly sardinella Amblygaster clupeoides Bleeker, 1849 from India</mods:title>
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<treatment id="4B6C2C30FF8EE63CEF8BB0F9FBA1699E" ID-DOI="http://doi.org/10.5281/zenodo.3502143" ID-GBIF-Taxon="154507634" ID-Zenodo-Dep="3502143" LSID="urn:lsid:plazi:treatment:4B6C2C30FF8EE63CEF8BB0F9FBA1699E" httpUri="http://treatment.plazi.org/id/4B6C2C30FF8EE63CEF8BB0F9FBA1699E" lastPageId="2" lastPageNumber="196" pageId="1" pageNumber="196">
<subSubSection id="8BDFCEADFF8EE63FEF8BB0F9FCDE6C6C" box="[151,802,1003,1030]" pageId="1" pageNumber="196" type="nomenclature">
<paragraph id="C37A9D26FF8EE63FEF8BB0F9FCDE6C6C" blockId="1.[151,802,1003,1030]" box="[151,802,1003,1030]" pageId="1" pageNumber="196">
<heading id="98322A4AFF8EE63FEF8BB0F9FCDE6C6C" bold="true" box="[151,802,1003,1030]" fontSize="11" level="1" pageId="1" pageNumber="196" reason="1">
<emphasis id="F1B14134FF8EE63FEF8BB0F9FCDE6C6C" bold="true" box="[151,802,1003,1030]" pageId="1" pageNumber="196">
Genus
<taxonomicName id="04C5E6A5FF8EE63FEFF2B0FEFCDE6C6C" ID-CoL="7PY92" authority="Williams &amp; Bunkley-Williams, 1994" authorityName="Williams &amp; Bunkley-Williams" authorityYear="1994" box="[238,802,1003,1030]" class="Malacostraca" family="Cymothoidae" genus="Ryukyua" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="F1B14134FF8EE63FEFF2B0FEFEA56C6F" bold="true" box="[238,345,1003,1029]" italics="true" pageId="1" pageNumber="196">Ryukyua</emphasis>
<bibRefCitation id="A754E0D7FF8EE63FEE7CB0F9FCDE6C6C" author="Williams, E. H. Jr. &amp; Bunkley-Williams, L." box="[352,802,1004,1030]" pageId="1" pageNumber="196" pagination="155 - 161" refId="ref5511" refString="Williams, E. H. Jr. &amp; Bunkley-Williams, L. (1994 b) Ryukyua globosa n. gen., n. sp., and R. circularis n. comb., parasitic in the opercular chambers of Pacific and Indian Ocean clupeid fishes. Journal of Aquactic animal Health, 6, 155 - 161." type="journal article" year="1994">Williams &amp; Bunkley-Williams, 1994</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8BDFCEADFF8EE63FEF8BB727FD496C0D" pageId="1" pageNumber="196" type="reference_group">
<paragraph id="C37A9D26FF8EE63FEF8BB727FD2A6C22" blockId="1.[151,726,1074,1127]" box="[151,726,1074,1096]" pageId="1" pageNumber="196">
<treatmentCitationGroup id="E3D5BA08FF8EE63FEF8BB727FD2A6C22" box="[151,726,1074,1096]" pageId="1" pageNumber="196">
<taxonomicName id="04C5E6A5FF8EE63FEF8BB727FD626C22" ID-CoL="7PY92" authority="Williams &amp; Bunkley-Williams, 1994: 155" authorityName="Williams &amp; Bunkley-Williams" authorityPageNumber="155" authorityYear="1994" box="[151,670,1074,1096]" class="Malacostraca" family="Cymothoidae" genus="Ryukyua" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="F1B14134FF8EE63FEF8BB727FF126C22" box="[151,238,1074,1096]" italics="true" pageId="1" pageNumber="196">Ryukyua</emphasis>
<treatmentCitation id="4264BB37FF8EE63FEFE8B727FD626C22" author="Williams, E. H. Jr. &amp; Bunkley-Williams, L." box="[244,670,1074,1096]" page="155" pageId="1" pageNumber="196" year="1994">
<bibRefCitation id="A754E0D7FF8EE63FEFE8B727FD916C22" author="Williams, E. H. Jr. &amp; Bunkley-Williams, L." box="[244,621,1074,1096]" pageId="1" pageNumber="196" pagination="155 - 161" refId="ref5511" refString="Williams, E. H. Jr. &amp; Bunkley-Williams, L. (1994 b) Ryukyua globosa n. gen., n. sp., and R. circularis n. comb., parasitic in the opercular chambers of Pacific and Indian Ocean clupeid fishes. Journal of Aquactic animal Health, 6, 155 - 161." type="journal article" year="1994">Williams &amp; Bunkley-Williams, 1994</bibRefCitation>
: 155
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</taxonomicName>
161.
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</paragraph>
<paragraph id="C37A9D26FF8EE63FEF8BB747FD496C0D" blockId="1.[151,726,1074,1127]" box="[151,693,1105,1127]" pageId="1" pageNumber="196">
<treatmentCitationGroup id="E3D5BA08FF8EE63FEF8BB747FD496C0D" box="[151,693,1105,1127]" pageId="1" pageNumber="196">
<taxonomicName id="04C5E6A5FF8EE63FEF8BB747FF086C0D" ID-CoL="7PBZK" authorityName="Leach" authorityYear="1818" box="[151,244,1106,1127]" class="Malacostraca" family="Cymothoidae" genus="Livoneca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="F1B14134FF8EE63FEF8BB747FF086C0D" box="[151,244,1106,1127]" italics="true" pageId="1" pageNumber="196">Livoneca</emphasis>
</taxonomicName>
<treatmentCitation id="4264BB37FF8EE63FEE09B744FE4D6C0D" author="Pillai, N. K." box="[277,433,1105,1127]" page="17" pageId="1" pageNumber="196" year="1954">
<bibRefCitation id="A754E0D7FF8EE63FEE09B744FE776C0D" author="Pillai, N. K." box="[277,395,1105,1127]" pageId="1" pageNumber="196" pagination="1 - 22" refId="ref5056" refString="Pillai, N. K. (1954) A preliminary note on the Tanaidacea and Isopoda of Travancore. Bulletin of the Central Research Institute, University of Travancore, Trivandrum, 3, 1 - 22." type="journal article" year="1954">Pillai, 1954</bibRefCitation>
: 17
</treatmentCitation>
; 1964: 218220 [partim].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="8BDFCEADFF8EE63FEF8BB781FE496CBA" pageId="1" pageNumber="196" type="materials_examined">
<paragraph id="C37A9D26FF8EE63FEF8BB781FE496CBA" blockId="1.[151,1437,1172,2025]" pageId="1" pageNumber="196">
<emphasis id="F1B14134FF8EE63FEF8BB781FEC76CC7" bold="true" box="[151,315,1172,1197]" pageId="1" pageNumber="196">
<typeStatus id="1C7E2384FF8EE63FEF8BB781FF2E6CC7" box="[151,210,1172,1197]" pageId="1" pageNumber="196">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="04C5E6A5FF8EE63FEE56B781FC2A6CC6" authority="Williams &amp; Bunkley-Williams, 1994" authorityName="Williams &amp; Bunkley-Williams" authorityYear="1994" box="[330,982,1172,1197]" class="Malacostraca" family="Cymothoidae" genus="Ryukyua" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="F1B14134FF8EE63FEE56B781FDED6CC6" box="[330,529,1172,1196]" italics="true" pageId="1" pageNumber="196">Ryukyua globosa</emphasis>
<bibRefCitation id="A754E0D7FF8EE63FED03B781FC2A6CC6" author="Williams, E. H. Jr. &amp; Bunkley-Williams, L." box="[543,982,1172,1197]" pageId="1" pageNumber="196" pagination="155 - 161" refId="ref5511" refString="Williams, E. H. Jr. &amp; Bunkley-Williams, L. (1994 b) Ryukyua globosa n. gen., n. sp., and R. circularis n. comb., parasitic in the opercular chambers of Pacific and Indian Ocean clupeid fishes. Journal of Aquactic animal Health, 6, 155 - 161." type="journal article" year="1994">Williams &amp; Bunkley-Williams, 1994</bibRefCitation>
</taxonomicName>
, by original designation (
<bibRefCitation id="A754E0D7FF8EE63FEA13B781FE556CBA" author="Williams, E. H. Jr. &amp; Bunkley-Williams, L." pageId="1" pageNumber="196" pagination="155 - 161" refId="ref5511" refString="Williams, E. H. Jr. &amp; Bunkley-Williams, L. (1994 b) Ryukyua globosa n. gen., n. sp., and R. circularis n. comb., parasitic in the opercular chambers of Pacific and Indian Ocean clupeid fishes. Journal of Aquactic animal Health, 6, 155 - 161." type="journal article" year="1994">Williams &amp; Bunkley-Williams 1994</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8BDFCEADFF8EE63FEFDBB7C9FAC06C9E" box="[199,1340,1244,1269]" pageId="1" pageNumber="196" type="diagnosis">
<paragraph id="C37A9D26FF8EE63FEFDBB7C9FAC06C9E" blockId="1.[151,1437,1172,2025]" box="[199,1340,1244,1269]" pageId="1" pageNumber="196">
<emphasis id="F1B14134FF8EE63FEFDBB7C9FEBF6C9F" bold="true" box="[199,323,1244,1269]" pageId="1" pageNumber="196">Diagnosis:</emphasis>
Generic diagnosis of
<taxonomicName id="04C5E6A5FF8EE63FED25B7C9FD656C9E" authorityName="Williams &amp; Bunkley-Williams" authorityYear="1994" box="[569,665,1244,1268]" class="Malacostraca" family="Cymothoidae" genus="Ryukyua" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="F1B14134FF8EE63FED25B7C9FD656C9E" box="[569,665,1244,1268]" italics="true" pageId="1" pageNumber="196">Ryukyua</emphasis>
</taxonomicName>
is here adapted from
<bibRefCitation id="A754E0D7FF8EE63FEC90B7C9FAC46C9E" author="Williams, E. H. Jr. &amp; Bunkley-Williams, L." box="[908,1336,1244,1269]" pageId="1" pageNumber="196" pagination="155 - 161" refId="ref5511" refString="Williams, E. H. Jr. &amp; Bunkley-Williams, L. (1994 b) Ryukyua globosa n. gen., n. sp., and R. circularis n. comb., parasitic in the opercular chambers of Pacific and Indian Ocean clupeid fishes. Journal of Aquactic animal Health, 6, 155 - 161." type="journal article" year="1994">Williams &amp; Bunkley-Williams (1994)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8BDFCEADFF8EE63FEFDBB7EAFDB16E37" pageId="1" pageNumber="196" type="description">
<paragraph id="C37A9D26FF8EE63FEFDBB7EAFDB16E37" blockId="1.[151,1437,1172,2025]" pageId="1" pageNumber="196">
<emphasis id="F1B14134FF8EE63FEFDBB7EAFEDB6D72" bold="true" box="[199,295,1279,1304]" italics="true" pageId="1" pageNumber="196">Female:</emphasis>
Body round, almost spherical in gravid females, compact, widest at pereonites 4 and 5; pereonite 7 shortest, pereonite 6 0.5 as long times as long as pereonite 5. Cephalon immersed in pereonite 1, anterior margin rounded to form blunt rostrum. Eyes 0.3 times cephalon width. Pleonites visible in dorsal view, lateral margin of pereonite 7 produced posteriorly and overlapping lateral margin of pleonites 14; pleon narrower than pereon. Pleotelson wider than long, posterior margin straight to broadly rounded or slightly emarginated. Antennula and antennae with 89 articles, antennula shorter than antennae in length and well-separated. Mandible palp article 3 shorter than article 2. Maxilliped palp article 2 without setae, proximal part of maxilliped not expanded. Maxilla medial lobe partially fused to lateral lobe. Pereopods stout, propodus as wide as long; pereopods 14 short, 56 longer. Pleopods without folds, pockets, setose or protrusions; uropods not reaching posterior margin of pleotelson, rami elongate-oval, subequal in length.
</paragraph>
</subSubSection>
<subSubSection id="8BDFCEADFF8EE63CEFDBB572FBA1699E" lastPageId="2" lastPageNumber="197" pageId="1" pageNumber="196" type="discussion">
<paragraph id="C37A9D26FF8EE63FEFDBB572FE906E87" blockId="1.[151,1437,1172,2025]" pageId="1" pageNumber="196">
<emphasis id="F1B14134FF8EE63FEFDBB572FEC16EEA" bold="true" box="[199,317,1639,1664]" pageId="1" pageNumber="196">Remarks:</emphasis>
<bibRefCitation id="A754E0D7FF8EE63FEE5AB572FD086EEA" author="Williams, E. H. Jr. &amp; Bunkley-Williams, L." box="[326,756,1639,1664]" pageId="1" pageNumber="196" pagination="155 - 161" refId="ref5511" refString="Williams, E. H. Jr. &amp; Bunkley-Williams, L. (1994 b) Ryukyua globosa n. gen., n. sp., and R. circularis n. comb., parasitic in the opercular chambers of Pacific and Indian Ocean clupeid fishes. Journal of Aquactic animal Health, 6, 155 - 161." type="journal article" year="1994">Williams &amp; Bunkley-Williams (1994)</bibRefCitation>
provided a brief diagnosis for the genus
<emphasis id="F1B14134FF8EE63FEBD0B57DFA606EEA" box="[1228,1436,1640,1664]" italics="true" pageId="1" pageNumber="196">
<taxonomicName id="04C5E6A5FF8EE63FEBD0B57DFACC6EEA" authorityName="Williams &amp; Bunkley-Williams" authorityYear="1994" box="[1228,1328,1640,1664]" class="Malacostraca" family="Cymothoidae" genus="Ryukyua" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">Ryukyua</taxonomicName>
.
<taxonomicName id="04C5E6A5FF8EE63FEA20B57DFA606EEA" authorityName="Williams &amp; Bunkley-Williams" authorityYear="1994" box="[1340,1436,1640,1664]" class="Malacostraca" family="Cymothoidae" genus="Ryukyua" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">Ryukyua</taxonomicName>
</emphasis>
can be identified by the strongly compact and spherical body; mandibular palp all articles distinct article 3 shorter than article 2; coxae partially visible dorsally, shorter than respective pereonites; uropod rami elongate-oval, subequal in length.
</paragraph>
<paragraph id="C37A9D26FF8EE63CEFDBB5E2FBA1699E" blockId="1.[151,1437,1172,2025]" lastBlockId="2.[151,1436,151,501]" lastPageId="2" lastPageNumber="197" pageId="1" pageNumber="196">
Other branchial cymothoids differ with
<taxonomicName id="04C5E6A5FF8EE63FED98B5EDFD186F7A" authorityName="Williams &amp; Bunkley-Williams" authorityYear="1994" box="[644,740,1784,1808]" class="Malacostraca" family="Cymothoidae" genus="Ryukyua" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="F1B14134FF8EE63FED98B5EDFD186F7A" box="[644,740,1784,1808]" italics="true" pageId="1" pageNumber="196">Ryukyua</emphasis>
</taxonomicName>
in the following:
<taxonomicName id="04C5E6A5FF8EE63FECB1B5EDFAA56F7A" authority="Schioedte and Meinert, 1884" authorityName="Schioedte and Meinert" authorityYear="1884" box="[941,1369,1783,1808]" class="Malacostraca" family="Cymothoidae" genus="Cterissa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="F1B14134FF8EE63FECB1B5EDFBF66F65" box="[941,1034,1784,1807]" italics="true" pageId="1" pageNumber="196">Cterissa</emphasis>
<bibRefCitation id="A754E0D7FF8EE63FEB0DB5E2FAA56F7A" author="Schioedte, J. C. &amp; Meinert, F." box="[1041,1369,1783,1808]" pageId="1" pageNumber="196" pagination="221 - 454" refId="ref5370" refString="Schioedte, J. C. &amp; Meinert, F. (1884) Symbolae ad monographium Cymothoarum crustaceorum isopodum familiae. IV. Cymothoidae Trib. II. Cymothoinae. Trib. III: Lironecinae. Naturhistorisk Tidsskrift, 14, 221 - 454." type="journal article" year="1884">Schiöedte and Meinert, 1884</bibRefCitation>
</taxonomicName>
: body dorsoventrally flattened, laterally expanded and more flattened on one side, coxae 27 visible, with posteroventral angles expanded at one end rounded; body round, almost spherical, pleonites 13 lateral margins not bilobed (
<bibRefCitation id="A754E0D7FF8EE63FEF82B471FDA96F17" author="Bunkley-Williams, L. &amp; Williams, E. H. Jr." box="[158,597,1892,1917]" pageId="1" pageNumber="196" pagination="213 - 216" refId="ref4672" refString="Bunkley-Williams, L. &amp; Williams, E. H. Jr. (1986) Some parasitic isopods (Crustacea: Cymothoidae) of marine fishes from Thailand. Galaxea, 5, 213 - 216." type="journal article" year="1986">Bunkley-Williams &amp; Williams 1986</bibRefCitation>
);
<taxonomicName id="04C5E6A5FF8EE63FED6EB471FBD76F16" authority="Schiodte and Meinert, 1884" authorityName="Schiodte and Meinert" authorityYear="1884" box="[626,1067,1892,1917]" class="Malacostraca" family="Cymothoidae" genus="Agarna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="F1B14134FF8EE63FED6EB471FD3A6F11" box="[626,710,1892,1915]" italics="true" pageId="1" pageNumber="196">Agarna</emphasis>
<bibRefCitation id="A754E0D7FF8EE63FEDC9B471FBD76F16" author="Schioedte, J. C. &amp; Meinert, F." box="[725,1067,1892,1917]" pageId="1" pageNumber="196" pagination="221 - 454" refId="ref5370" refString="Schioedte, J. C. &amp; Meinert, F. (1884) Symbolae ad monographium Cymothoarum crustaceorum isopodum familiae. IV. Cymothoidae Trib. II. Cymothoinae. Trib. III: Lironecinae. Naturhistorisk Tidsskrift, 14, 221 - 454." type="journal article" year="1884">Schiödte and Meinert, 1884</bibRefCitation>
</taxonomicName>
: pereon elevated mid-dorsally, pereonites 37 are greatly expanded laterally; pereonite 7 posteriorlaterally produced and overlapping pleonites 3 5 (
<bibRefCitation id="A754E0D7FF8EE63FEFA8B4B9FDAC6FAE" author="Williams, E. H. Jr. &amp; Bunkley-Williams, L." box="[180,592,1964,1989]" pageId="1" pageNumber="196" pagination="155 - 161" refId="ref5511" refString="Williams, E. H. Jr. &amp; Bunkley-Williams, L. (1994 b) Ryukyua globosa n. gen., n. sp., and R. circularis n. comb., parasitic in the opercular chambers of Pacific and Indian Ocean clupeid fishes. Journal of Aquactic animal Health, 6, 155 - 161." type="journal article" year="1994">Williams &amp; Bunkley-Williams 1994</bibRefCitation>
;
<bibRefCitation id="A754E0D7FF8EE63FED41B4B9FCC96FAE" author="Aneesh, P. T. &amp; Sudha, K. &amp; Helna, A. K. &amp; Anilkumar, G." box="[605,821,1964,1989]" pageId="1" pageNumber="196" pagination="1 - 22" refId="ref4364" refString="Aneesh, P. T., Sudha, K. Helna, A. K. &amp; Anilkumar, G. (2018) Agarna malayi Tiwari 1952 (Crustacea: Isopoda: Cymothoidae) parasitising the marine fish, Tenualosa toli (Clupeidae) from India: Re-description / description of parasite life cycle and patterns of occurrence. Zoological Studies, 57 (25), 1 - 22." type="journal article" year="2018">
Aneesh
<emphasis id="F1B14134FF8EE63FEDA4B4B8FD0D6FAE" box="[696,753,1964,1988]" italics="true" pageId="1" pageNumber="196">et al.</emphasis>
2018
</bibRefCitation>
);
<taxonomicName id="04C5E6A5FF8EE63FEC54B4B8FB386FAE" authority="Bowman &amp; Tareen, 1983" authorityName="Bowman &amp; Tareen" authorityYear="1983" box="[840,1220,1964,1988]" class="Malacostraca" family="Cymothoidae" genus="Joryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="F1B14134FF8EE63FEC54B4B8FC606FAE" box="[840,924,1965,1988]" italics="true" pageId="1" pageNumber="196">Joryma</emphasis>
<bibRefCitation id="A754E0D7FF8EE63FECB8B4B8FB386FAE" author="Bowman, T. E. &amp; Tareen, I. U." box="[932,1220,1964,1988]" pageId="1" pageNumber="196" pagination="1 - 30" refId="ref4437" refString="Bowman, T. E. &amp; Tareen, I. U. (1983) Cymothoidae from fishes of Kuwait (Arabian Gulf) (Crustacea, Isopoda). Smithsonian Contributions to Zoology, 382, 1 - 30. https: // doi. org / 10.5479 / si. 00810282.382" type="journal article" year="1983">Bowman &amp; Tareen, 1983</bibRefCitation>
</taxonomicName>
: pereonite width to pleotelson width ratio similar (
<emphasis id="F1B14134FF8EE63FEEE8B4C4FDF76F82" box="[500,523,2001,2024]" italics="true" pageId="1" pageNumber="196">vs</emphasis>
wider pereon to pleotelson ratio), mandible simple with less pronounced incisor (
<emphasis id="F1B14134FF8DE63CEFBCB38CFF4B68DA" box="[160,183,153,176]" italics="true" pageId="2" pageNumber="197">vs</emphasis>
incisors acute), mandible palp with weak segmentation, pereopod 7 dactylus slender and reaching beyond carpus, pereopod 7 ischium stout and expanded (
<emphasis id="F1B14134FF8DE63CEDFBB3A8FD0268BE" box="[743,766,189,212]" italics="true" pageId="2" pageNumber="197">vs</emphasis>
slender, without expansion) (
<bibRefCitation id="A754E0D7FF8DE63CEB7EB3A9FA7368BE" author="Bowman, T. E. &amp; Tareen, I. U." box="[1122,1423,188,212]" pageId="2" pageNumber="197" pagination="1 - 30" refId="ref4437" refString="Bowman, T. E. &amp; Tareen, I. U. (1983) Cymothoidae from fishes of Kuwait (Arabian Gulf) (Crustacea, Isopoda). Smithsonian Contributions to Zoology, 382, 1 - 30. https: // doi. org / 10.5479 / si. 00810282.382" type="journal article" year="1983">Bowman &amp; Tareen 1983</bibRefCitation>
);
<taxonomicName id="04C5E6A5FF8DE63CEF8BB3F5FDBC6892" authority=": Schioedte and Meinert, 1884" authorityName="Schioedte and Meinert" authorityYear="1884" box="[151,576,223,248]" class="Malacostraca" family="Cymothoidae" genus="Elthusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="F1B14134FF8DE63CEF8BB3F5FF106892" box="[151,236,224,248]" italics="true" pageId="2" pageNumber="197">Elthusa</emphasis>
:
<bibRefCitation id="A754E0D7FF8DE63CEFE6B3CAFDBC6892" author="Schioedte, J. C. &amp; Meinert, F." box="[250,576,223,248]" pageId="2" pageNumber="197" pagination="221 - 454" refId="ref5370" refString="Schioedte, J. C. &amp; Meinert, F. (1884) Symbolae ad monographium Cymothoarum crustaceorum isopodum familiae. IV. Cymothoidae Trib. II. Cymothoinae. Trib. III: Lironecinae. Naturhistorisk Tidsskrift, 14, 221 - 454." type="journal article" year="1884">Schiöedte and Meinert, 1884</bibRefCitation>
</taxonomicName>
: coxae 47 from dorsal view, eyes width 1/3 cephalon width (
<emphasis id="F1B14134FF8DE63CEBF3B3F4FAFA6892" box="[1263,1286,225,248]" italics="true" pageId="2" pageNumber="197">vs</emphasis>
2/3 cephalon width), maxilliped article 3 slender and with setae (
<bibRefCitation id="A754E0D7FF8DE63CEDEBB211FC7E6976" author="Bruce, N. L." box="[759,898,260,284]" pageId="2" pageNumber="197" pagination="247 - 300" refId="ref4599" refString="Bruce, N. L. (1990) The genera Catoessa, Elthusa, Enispa, Ichthyoxenus, Idusa, Livoneca and Norileca n. gen. (Isopoda, Cymothoidae), crustacean parasites of marine fishes, with descriptions of eastern Australian species. Records of the Australian Museum, 42, 247 - 300. https: // doi. org / 10.3853 / j. 0067 - 1975.42.1990.118" type="journal article" year="1990">Bruce 1990</bibRefCitation>
;
<bibRefCitation id="A754E0D7FF8DE63CEC8DB211FB7E6976" author="Hadfield, K. A. &amp; Tuttle, L. J. &amp; Smit, N. J." box="[913,1154,260,285]" pageId="2" pageNumber="197" pagination="125 - 135" refId="ref4843" refString="Hadfield, K. A., Tuttle, L. J. &amp; Smit, N. J. (2017) Elthusa winstoni sp. n. (Isopoda, Cymothoidae), a new fish parasitic isopod from Hawaii. ZooKeys, 661, 125 - 135. https: // doi. org / 10.3897 / zookeys. 661.11251" type="journal article" year="2017">
Hadfield
<emphasis id="F1B14134FF8DE63CECE2B210FBC76976" box="[1022,1083,260,284]" italics="true" pageId="2" pageNumber="197">et al.</emphasis>
2017
</bibRefCitation>
);
<taxonomicName id="04C5E6A5FF8DE63CEB86B211FA646976" authority="Bruce, 1990" authorityName="Bruce" authorityYear="1990" box="[1178,1432,260,284]" class="Malacostraca" family="Cymothoidae" genus="Norileca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="F1B14134FF8DE63CEB86B211FB006976" box="[1178,1276,260,284]" italics="true" pageId="2" pageNumber="197">Norileca</emphasis>
<bibRefCitation id="A754E0D7FF8DE63CEA1BB210FA646976" author="Bruce, N. L." box="[1287,1432,260,284]" pageId="2" pageNumber="197" pagination="247 - 300" refId="ref4599" refString="Bruce, N. L. (1990) The genera Catoessa, Elthusa, Enispa, Ichthyoxenus, Idusa, Livoneca and Norileca n. gen. (Isopoda, Cymothoidae), crustacean parasites of marine fishes, with descriptions of eastern Australian species. Records of the Australian Museum, 42, 247 - 300. https: // doi. org / 10.3853 / j. 0067 - 1975.42.1990.118" type="journal article" year="1990">Bruce, 1990</bibRefCitation>
</taxonomicName>
: pleopod 5 with folds, pleopods 25 peduncle with laminar lobe, body strongly twisted to one side, mandible palp article 2 greatly expanded, brood pouch with posterior pocket, pleonites 1 and 2 without ventro-lateral processes (
<bibRefCitation id="A754E0D7FF8DE63CEF82B265FED569E2" author="Bruce, N. L." box="[158,297,368,392]" pageId="2" pageNumber="197" pagination="247 - 300" refId="ref4599" refString="Bruce, N. L. (1990) The genera Catoessa, Elthusa, Enispa, Ichthyoxenus, Idusa, Livoneca and Norileca n. gen. (Isopoda, Cymothoidae), crustacean parasites of marine fishes, with descriptions of eastern Australian species. Records of the Australian Museum, 42, 247 - 300. https: // doi. org / 10.3853 / j. 0067 - 1975.42.1990.118" type="journal article" year="1990">Bruce 1990</bibRefCitation>
);
<taxonomicName id="04C5E6A5FF8DE63CEE5DB265FD4C69E2" authority="Costa in Hope, 1851" authorityName="Costa in Hope" authorityYear="1851" box="[321,688,367,392]" class="Malacostraca" family="Cymothoidae" genus="Mothocya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="F1B14134FF8DE63CEE5DB265FE4D69E2" box="[321,433,368,392]" italics="true" pageId="2" pageNumber="197">Mothocya</emphasis>
Costa in Hope, 1851
</taxonomicName>
: cephalon with rostrum, pleon partly overlapped by pereonite 7, pleotelson posterior region rounded, with appendix masculine on pleopod 2 (
<bibRefCitation id="A754E0D7FF8DE63CECE1B281FB7A69C7" author="Bruce, N. L." box="[1021,1158,404,429]" pageId="2" pageNumber="197" pagination="1089 - 1192" refId="ref4542" refString="Bruce, N. L. (1986) Revision of the isopod crustacean genus Mothocya Costa, in Hope, 1851 (Cymothoidae: Flabellifera), parasitic on marine fishes. Journal of Natural History, 20 (5), 1089 - 1192. https: // doi. org / 10.1080 / 00222938600770781" type="journal article" year="1986">Bruce 1986</bibRefCitation>
);
<taxonomicName id="04C5E6A5FF8DE63CEB86B280FA6469C6" authority="Leach, 1818" authorityName="Leach" authorityYear="1818" box="[1178,1432,404,429]" class="Malacostraca" family="Cymothoidae" genus="Livoneca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="F1B14134FF8DE63CEB86B280FAFD69C6" box="[1178,1281,405,428]" italics="true" pageId="2" pageNumber="197">Livoneca</emphasis>
<bibRefCitation id="A754E0D7FF8DE63CEA14B281FA6469C6" author="Leach, W. E." box="[1288,1432,404,429]" pageId="2" pageNumber="197" pagination="338 - 354" refId="ref4946" refString="Leach, W. E. (1818) Cymothoadees. In: Cuvier, F. (Ed.), Dictionnaire des Sciences Naturelles, Paris, 12, pp. 338 - 354." type="journal article" year="1818">Leach, 1818</bibRefCitation>
</taxonomicName>
: coxae longer than respective pereonite; pleon not immersed in pereon, pleonites becoming progressively narrow towards posterior; ventrolateral margin of pleonites 13 weakly bilobed (
<bibRefCitation id="A754E0D7FF8DE63CECD5B2C9FBAD699E" author="Bruce, N. L." box="[969,1105,476,500]" pageId="2" pageNumber="197" pagination="247 - 300" refId="ref4599" refString="Bruce, N. L. (1990) The genera Catoessa, Elthusa, Enispa, Ichthyoxenus, Idusa, Livoneca and Norileca n. gen. (Isopoda, Cymothoidae), crustacean parasites of marine fishes, with descriptions of eastern Australian species. Records of the Australian Museum, 42, 247 - 300. https: // doi. org / 10.3853 / j. 0067 - 1975.42.1990.118" type="journal article" year="1990">Bruce 1990</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>