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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFA0FFE15AB6FD7B4419C885" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus foxworthyanus Craib 1901" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571674" ID-Zenodo-Dep="7571674" LSID="urn:lsid:plazi:treatment:43284262FFA0FFE15AB6FD7B4419C885" httpUri="http://treatment.plazi.org/id/43284262FFA0FFE15AB6FD7B4419C885" pageId="63">
<subSubSection box="[136,1035,734,761]" pageId="63" type="nomenclature">
<paragraph blockId="63.[136,1322,734,796]" box="[136,1035,734,761]" pageId="63">
<emphasis bold="true" box="[136,171,734,760]" pageId="63">25.</emphasis>
<taxonomicName ID-CoL="6NZ7D" authority="Craib (1901: 17)" authorityName="Craib" authorityPageNumber="17" authorityYear="1901" box="[178,681,734,761]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="63" phylum="Tracheophyta" rank="species" species="foxworthyanus">
<emphasis bold="true" box="[178,487,734,761]" italics="true" pageId="63">Lasianthus foxworthyanus</emphasis>
Craib (1901: 17)
</taxonomicName>
<emphasis bold="true" box="[681,709,734,760]" pageId="63"></emphasis>
<figureCitation box="[709,788,734,760]" captionStart="FIGURE 46" captionStartId="64.[136,229,1805,1827]" captionTargetBox="[165,1418,211,1777]" captionTargetId="figure-25@64.[164,1421,209,1781]" captionTargetPageId="64" captionText="FIGURE 46. Lasianthus foxworthyanus Craib (from V6415, FU, VNM and the herbarium of Iriomote Station (Japan)): A. Branch with flower buds, B. Close-up of young branch and leaves, C. Stipules, D. Abaxial leaf surfaces, E. Flower buds, FH. Young fruits (A, F, H: photos by Shuichiro Tagane)." figureDoi="http://doi.org/10.5281/zenodo.7571692" httpUri="https://zenodo.org/record/7571692/files/figure.png" pageId="63">Fig. 46</figureCitation>
(plate),
<figureCitation box="[882,961,734,760]" captionStart="FIGURE 47" captionStartId="65.[136,229,1804,1826]" captionTargetBox="[180,1411,204,1763]" captionTargetId="figure-17@65.[165,1418,193,1780]" captionTargetPageId="65" captionText="FIGURE 47. Distribution maps of Lasianthus foxworthyanus, L. giganteus, L. globularis and L. hatinhensis." figureDoi="http://doi.org/10.5281/zenodo.7571696" httpUri="https://zenodo.org/record/7571696/files/figure.png" pageId="63">Fig. 47</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection pageId="63" type="materials_examined">
<paragraph blockId="63.[136,1322,734,796]" box="[136,1322,774,796]" pageId="63">
<materialsCitation box="[136,1322,774,796]" collectingDate="1930-06-11" collectionCode="SING" collectorName="M. S. Kiah" country="Thailand" location="Bacho" pageId="63" specimenCode="SING0059257" specimenCount="1" stateProvince="Narathiwat" typeStatus="holotype">
<emphasis bold="true" box="[136,192,774,795]" pageId="63">
<typeStatus box="[136,188,774,795]" pageId="63">Type</typeStatus>
.
</emphasis>
<collectingCountry box="[198,325,774,795]" name="Thailand" pageId="63">THAILAND</collectingCountry>
.
<collectingRegion box="[334,443,774,796]" country="Thailand" name="Narathiwat" pageId="63">Narathiwat</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFA0FFE15AB6FD7B4419C885:CE5EA5AFFFA0FFE15BFAFCA34222CF54" box="[452,516,774,796]" country="Thailand" name="Bacho" pageId="63" stateProvince="Narathiwat">Bacho</location>
,
<date box="[526,655,774,795]" pageId="63" value="1930-06-11">
<collectingDate box="[526,655,774,795]" pageId="63" value="1930-06-11">11 June 1930</collectingDate>
</date>
,
<emphasis box="[665,822,774,795]" italics="true" pageId="63">
<collectorName box="[665,756,774,795]" pageId="63">M.S.Kiah</collectorName>
24312
</emphasis>
(isotypes K! SING! [K000777027,
<specimenCode box="[1167,1313,774,796]" collectionCode="SING" country="Singapore" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="63" type="Herbarium">SING0059257</specimenCode>
].
</materialsCitation>
</paragraph>
<paragraph blockId="63.[136,1452,842,1229]" pageId="63">
<emphasis bold="true" box="[136,401,842,868]" pageId="63">
<collectingRegion box="[136,262,842,868]" pageId="63">Specimens</collectingRegion>
examined:
</emphasis>
<materialsCitation collectingDate="2017-02-12" collectorName="Tagane S. &amp; Nagamasu H. &amp; Nguyen" country="Vietnam" elevation="1070" location="Ngoc Linh Nature Reserve" pageId="63" specimenCount="1" stateProvince="Kon Tum">
<collectingCountry box="[430,562,842,868]" name="Vietnam" pageId="63">VIETNAM</collectingCountry>
.
<emphasis bold="true" box="[579,680,842,868]" pageId="63">Central:</emphasis>
<collectingRegion box="[694,924,842,869]" country="Vietnam" name="Kon Tum" pageId="63">Kon Tum Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFA0FFE15AB6FD7B4419C885:CE5EA5AFFFA0FFE15991FCEE44D2CF2D" box="[943,1268,842,869]" country="Vietnam" name="Ngoc Linh Nature Reserve" pageId="63" stateProvince="Kon Tum">Ngoc Linh Nature Reserve</location>
,
<quantity box="[1286,1379,842,869]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="63" unit="m" value="1070.0">
<elevation box="[1286,1379,842,869]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="63" unit="m" value="1070.0">1070 m</elevation>
</quantity>
elev., 15˚1005.07”N, 107˚4523.06E,
<date box="[520,722,878,904]" pageId="63" value="2017-02-12">
<collectingDate box="[520,722,878,904]" pageId="63" value="2017-02-12">12 February 2017</collectingDate>
</date>
,
<emphasis box="[733,1111,879,905]" italics="true" pageId="63">
<collectorName box="[733,844,879,905]" pageId="63">Tagane S.</collectorName>
,
<collectorName box="[857,1013,879,904]" pageId="63">Nagamasu H.</collectorName>
,
<collectorName box="[1025,1111,879,904]" pageId="63">Nguyen</collectorName>
</emphasis>
</materialsCitation>
<materialsCitation collectionCode="DLU, FU, VNM" collectorName="Hoang T. B. &amp; Hoang T. S. &amp; Yang C. J. &amp; Kawakubo A. V" country="Japan" location="Iriomote Station" pageId="63" specimenCount="1" stateProvince="Central">
<emphasis italics="true" pageId="63">
<taxonomicNameLabel box="[1117,1167,879,904]" pageId="63" rank="variety">V.N.</taxonomicNameLabel>
,
<collectorName box="[1178,1308,879,904]" pageId="63">Hoang T.B.</collectorName>
,
<collectorName box="[1320,1447,879,905]" pageId="63">Hoang T.S.</collectorName>
,
<collectorName box="[136,245,915,941]" pageId="63">Yang C.J.</collectorName>
,
<collectorName box="[257,432,914,941]" pageId="63">Kawakubo A. V</collectorName>
6415
</emphasis>
(
<collectionCode box="[502,564,914,940]" pageId="63">DLU</collectionCode>
,
<collectionCode box="[576,615,915,941]" country="China" name="Fudan University, Department of Biology" pageId="63">FU</collectionCode>
,
<collectionCode box="[625,693,915,941]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="63" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFA0FFE15AB6FD7B4419C885:CE5EA5AFFFA0FFE159BCFC37441ECFE4" box="[898,1080,914,940]" country="Japan" name="Iriomote Station" pageId="63" stateProvince="Central">Iriomote Station</location>
(
<collectingCountry box="[1095,1164,914,940]" name="Japan" pageId="63">Japan</collectingCountry>
))
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="63" type="distribution">
<paragraph blockId="63.[136,1452,842,1229]" box="[189,1346,950,977]" pageId="63">
<emphasis bold="true" box="[189,343,950,976]" pageId="63">Distribution.</emphasis>
Southeast Asia region (
<collectingCountry box="[609,716,950,976]" name="Malaysia" pageId="63">Malaysia</collectingCountry>
,
<collectingCountry box="[726,831,950,976]" name="Thailand" pageId="63">Thailand</collectingCountry>
). In
<collectingCountry box="[879,977,950,976]" name="Vietnam" pageId="63">Vietnam</collectingCountry>
, this species occurs in
<collectingRegion box="[1232,1343,951,977]" country="Vietnam" name="Kon Tum" pageId="63">Kon Tum</collectingRegion>
.
</paragraph>
<paragraph blockId="63.[136,1452,842,1229]" box="[189,1185,986,1013]" pageId="63">
<emphasis bold="true" box="[189,419,986,1012]" pageId="63">Habitat &amp; Ecology.</emphasis>
In shaded places of the moist evergreen forests at
<quantity box="[984,1125,986,1013]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="12.0" metricValueMin="3.0" pageId="63" unit="m" value="750.0" valueMax="1200.0" valueMin="300.0">3001200 m</quantity>
elev.
</paragraph>
<paragraph blockId="63.[136,1452,842,1229]" box="[189,860,1022,1049]" pageId="63">
<emphasis bold="true" box="[189,320,1022,1048]" pageId="63">Phenology.</emphasis>
Flowering and fruiting from June to November.
</paragraph>
<paragraph blockId="63.[136,1452,842,1229]" box="[189,652,1058,1087]" pageId="63">
<emphasis bold="true" box="[189,403,1058,1084]" pageId="63">Vernacular name.</emphasis>
Xú hưƠng foxworthy.
</paragraph>
<paragraph blockId="63.[136,1452,842,1229]" pageId="63">
<emphasis bold="true" box="[189,263,1094,1120]" pageId="63">Notes.</emphasis>
<taxonomicName box="[272,567,1094,1120]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="63" phylum="Tracheophyta" rank="species" species="foxworthyanus">
<emphasis box="[272,567,1094,1120]" italics="true" pageId="63">Lasianthus foxworthyanus</emphasis>
</taxonomicName>
is a new record for the flora of
<collectingCountry box="[937,1036,1094,1120]" name="Vietnam" pageId="63">Vietnam</collectingCountry>
. It is easily recognizable in the field from other
<taxonomicName box="[265,388,1130,1156]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="63" phylum="Tracheophyta" rank="genus">
<emphasis box="[265,388,1130,1156]" italics="true" pageId="63">Lasianthus</emphasis>
</taxonomicName>
in having densely appressed-pubescent branches; leaves oblong-lanceolate or oblong-elliptic, 916 ×
<quantity box="[226,314,1166,1193]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="63" unit="cm" value="4.0" valueMax="5.0" valueMin="3.0">35 cm</quantity>
, apex acuminate, base cuneate, secondary veins 69 on each side of midrib; petioles
<quantity box="[1291,1386,1166,1193]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="63" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
long, appressed-pubescent; stipules
<quantity box="[475,568,1202,1229]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="63" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
long, pubescent; fruit globose with 4 pyrenes.
</paragraph>
</subSubSection>
</treatment>
</document>