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<document id="65DAA3B342D62E8731BDFD6335048CB1" ID-DOI="10.5852/ejt.2019.585" ID-GBIF-Dataset="dca608e8-fccf-4c1c-b8df-ef0c28e1d518" ID-ISSN="2118-9773" ID-Zenodo-Dep="3660259" ID-ZooBank="181B6FBA-ED75-4BB4-84C4-FB512B794749" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1581251975682" checkinUser="plazi" docAuthor="Ebersole, Jun A., Cicimurri, David J. &amp; Stringer, Gary L." docDate="2019" docId="18174D41FF5DFF76FDDE99AE4C4C0A2A" docLanguage="en" docName="ejt-585_ebersole_cicimurri_stinger.pdf.imf" docOrigin="European Journal of Taxonomy 585" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ekokenia eporrecta - Frizzell &amp; Dante 1965" docType="treatment" docVersion="20" lastPageNumber="218" masterDocId="E42E3539FF85FFAFFFC69F344D4B0F37" masterDocTitle="Taxonomy and biostratigraphy of the elasmobranchs and bony fishes (Chondrichthyes and Osteichthyes) of the lower-to-middle Eocene (Ypresian to Bartonian) Claiborne Group in Alabama, USA, including an analysis of otoliths" masterLastPageNumber="274" masterPageNumber="1" pageNumber="217" updateTime="1732819509307" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4D8B928CEE59A7BC902FA4428CB6D6F7">Taxonomy and biostratigraphy of the elasmobranchs and bony fishes (Chondrichthyes and Osteichthyes) of the lower-to-middle Eocene (Ypresian to Bartonian) Claiborne Group in Alabama, USA, including an analysis of otoliths</mods:title>
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<mods:namePart id="98505C16134459A048FCFA65C89C5874">Ebersole, Jun A.</mods:namePart>
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<subSubSection id="D8A4AFDCFF5DFF77FDDE99AE49570982" box="[536,1052,1690,1717]" pageId="216" pageNumber="217" type="nomenclature">
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<taxonomicName id="57BE87D4FF5DFF77FDDE99AE49570982" authority="Frizzell &amp; Dante, 1965" authorityName="- Frizzell &amp; Dante" authorityYear="1965" box="[536,1052,1690,1717]" genus="Ekokenia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="216" pageNumber="217" phylum="Chordata" rank="species" species="eporrecta">
<emphasis id="A2CA2045FF5DFF77FDDE99AE4E4E0983" bold="true" box="[536,773,1690,1717]" italics="true" pageId="216" pageNumber="217">Ekokenia eporrecta</emphasis>
<bibRefCitation id="F42F81A6FF5DFF77FCCB99AE49570982" author="Frizzell D. L. &amp; Dante J. H." box="[781,1052,1690,1717]" pageId="216" pageNumber="217" pagination="368 - 718" refId="ref142032" refString="Frizzell D. L. &amp; Dante J. H. 1965. Otoliths of some early Cenozoic fishes of the Gulf Coast. Journal of Paleontology 39 (4): 368 - 718." type="journal article" year="1965">Frizzell &amp; Dante, 1965</bibRefCitation>
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<figureCitation id="0885E0D2FF5DFF77FD1E99894E6109EF" box="[728,810,1725,1752]" captionStart="Fig" captionStartId="217.[189,232,1894,1920]" captionTargetBox="[201,1386,1389,1862]" captionTargetId="figure@217.[189,1398,1376,1877]" captionTargetPageId="217" captionText="Fig. 72. Claiborne Group, “upper” Lisbon Formation, otoliths. AD. Jefitchia claybornensis Frizzell &amp; Dante, 1965. AB. MSC 39057. A. Inner face (right sagitta, juvenile). B. Dorsal view. CD. MSC 39049. C. Inner face (right sagitta). D. Dorsal view. — EF. Ekokenia eporrecta Frizzell &amp; Dante, 1965, MSC 39048. E. Inner face (left sagitta). F. Dorsal view. Scale bars: AD=2 mm; EF=5 mm." figureDoi="http://doi.org/10.5281/zenodo.3660399" httpUri="https://zenodo.org/record/3660399/files/figure.png" pageId="216" pageNumber="217">Fig. 72</figureCitation>
EF
</paragraph>
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<paragraph id="9001FC57FF5DFF77FF7B99CB4962082E" blockId="216.[189,1065,1791,1818]" box="[189,1065,1791,1818]" pageId="216" pageNumber="217">
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<taxonomicName id="57BE87D4FF5DFF77FF7B99CB4E02082E" authority="Koken, 1888: 282" authorityName=", Koken" authorityPageNumber="282" authorityYear="1888" box="[189,841,1791,1818]" family="Sciaenidae" genus="Otolithus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="216" pageNumber="217" phylum="Chordata" rank="species" species="eporrectus" subGenus="Sciaenidarum">
<emphasis id="A2CA2045FF5DFF77FF7B99CB4C61082E" box="[189,298,1791,1817]" italics="true" pageId="216" pageNumber="217">Otolithus</emphasis>
(
<emphasis id="A2CA2045FF5DFF77FEFD99CB4C95082E" box="[315,478,1791,1817]" italics="true" pageId="216" pageNumber="217">Sciaenidarum</emphasis>
)
<emphasis id="A2CA2045FF5DFF77FE2998344F20082E" box="[495,619,1792,1817]" italics="true" pageId="216" pageNumber="217">eporrectus</emphasis>
<treatmentCitation id="111FDA46FF5DFF77FDB599CB4E02082E" author="Koken E." box="[627,841,1791,1817]" page="282" pageId="216" pageNumber="217" year="1888">
<bibRefCitation id="F42F81A6FF5DFF77FDB599CB4E02082E" author="Koken E." box="[627,841,1791,1817]" pageId="216" pageNumber="217" pagination="274 - 305" refId="ref144486" refString="Koken E. 1888. Neue Untersuchungen an tertiaren Fisch-Otolithen. Zeitschrift der Deutschen Geologischen Gesellschaft 40: 274 - 305." type="journal article" year="1888">Koken, 1888: 282</bibRefCitation>
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, pl. 18, figs 1617.
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</paragraph>
<paragraph id="9001FC57FF5DFF77FF7B9875483C086C" blockId="216.[189,1399,1857,2027]" box="[189,1399,1857,1884]" pageId="216" pageNumber="217">
<treatmentCitationGroup id="B0AEDB79FF5DFF77FF7B9875483C086C" box="[189,1399,1857,1884]" pageId="216" pageNumber="217">
<taxonomicName id="57BE87D4FF5DFF77FF7B98754E38086C" authority="- Posthumus 1924: 27" authorityName="Posthumus" authorityPageNumber="27" authorityYear="1924" box="[189,883,1857,1884]" family="Sciaenidae" genus="Otolithus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="216" pageNumber="217" phylum="Chordata" rank="species" species="eporrectus" subGenus="Sciaenidarum">
<emphasis id="A2CA2045FF5DFF77FF7B98754C61086C" box="[189,298,1857,1883]" italics="true" pageId="216" pageNumber="217">Otolithus</emphasis>
(
<emphasis id="A2CA2045FF5DFF77FEFC98754C96086C" box="[314,477,1857,1883]" italics="true" pageId="216" pageNumber="217">Sciaenidarum</emphasis>
)
<emphasis id="A2CA2045FF5DFF77FE2A98764F23086C" box="[492,616,1858,1883]" italics="true" pageId="216" pageNumber="217">eporrectus</emphasis>
<treatmentCitation id="111FDA46FF5DFF77FD4598754E38086C" author="Posthumus O." box="[643,883,1857,1883]" page="27" pageId="216" pageNumber="217" year="1924">
<bibRefCitation id="F42F81A6FF5DFF77FD4598754E38086C" author="Posthumus O." box="[643,883,1857,1883]" pageId="216" pageNumber="217" refId="ref148841" refString="Posthumus O. 1924. Otolithi piscium. In: Diener C. (ed.) Fossilium Catalogus I, Pars 24, Animalia. W. Junk, Berlin." type="book" year="1924">Posthumus 1924: 27</bibRefCitation>
</treatmentCitation>
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. —
<treatmentCitation id="111FDA46FF5DFF77FC66987549D6086C" author="Campbell R." box="[928,1181,1857,1883]" page="264" pageId="216" pageNumber="217" year="1929">
<bibRefCitation id="F42F81A6FF5DFF77FC66987549D6086C" author="Campbell R." box="[928,1181,1857,1883]" pageId="216" pageNumber="217" pagination="254 - 279" refId="ref137518" refString="Campbell R. 1929 a. Fish otoliths, their occurrence and value as stratigraphic markers. Journal of Paleontology 3 (3): 254 - 279." type="journal article" year="1929">Campbell 1929a: 264</bibRefCitation>
</treatmentCitation>
, pl. 29, figs 1617.
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</paragraph>
<paragraph id="9001FC57FF5DFF77FF7B98514CFA0894" blockId="216.[189,1399,1857,2027]" pageId="216" pageNumber="217">
<treatmentCitationGroup id="B0AEDB79FF5DFF77FF7B98514CFA0894" pageId="216" pageNumber="217">
<taxonomicName id="57BE87D4FF5DFF77FF7B98514FBE0848" authority="- Frizzell &amp; Dante 1965: 704" authorityName="- Frizzell &amp; Dante" authorityPageNumber="704" authorityYear="1965" box="[189,757,1892,1919]" genus="Ekokenia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="216" pageNumber="217" phylum="Chordata" rank="species" species="eporrecta">
<emphasis id="A2CA2045FF5DFF77FF7B98514CEB0849" box="[189,416,1893,1919]" italics="true" pageId="216" pageNumber="217">Ekokenia eporrecta</emphasis>
<treatmentCitation id="111FDA46FF5DFF77FE7F98504FBE0848" author="Frizzell D. L. &amp; Dante J. H." box="[441,757,1892,1919]" page="704" pageId="216" pageNumber="217" year="1965">
<bibRefCitation id="F42F81A6FF5DFF77FE7F98504FBE0848" author="Frizzell D. L. &amp; Dante J. H." box="[441,757,1892,1919]" pageId="216" pageNumber="217" pagination="368 - 718" refId="ref142032" refString="Frizzell D. L. &amp; Dante J. H. 1965. Otoliths of some early Cenozoic fishes of the Gulf Coast. Journal of Paleontology 39 (4): 368 - 718." type="journal article" year="1965">Frizzell &amp; Dante 1965: 704</bibRefCitation>
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, pl. 87, figs 1112, 16; pl. 88, figs 5, 11. —
<treatmentCitation id="111FDA46FF5DFF77FB1A98504C1F0894" author="Schwarzhans W." page="27" pageId="216" pageNumber="217" year="1993">
<bibRefCitation id="F42F81A6FF5DFF77FB1A98504C1F0894" author="Schwarzhans W." pageId="216" pageNumber="217" pagination="1 - 245" refId="ref150082" refString="Schwarzhans W. 1993. A comparative morphological treatise of recent and fossil otoliths of the family Sciaenidae (Perciformes). In: Pfeil F. (ed.) Piscium Catalogus, Otolithi Piscium: 1 - 245. Verlag Dr. Freidrich Pfeil, Munich." type="book chapter" year="1993">Schwarzhans 1993: 27</bibRefCitation>
</treatmentCitation>
, fig. 11.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9001FC57FF5DFF77FF7B9899483D08F0" blockId="216.[189,1399,1857,2027]" box="[189,1398,1964,1991]" pageId="216" pageNumber="217">
“genus
<emphasis id="A2CA2045FF5DFF77FED598994CFF08F0" box="[275,436,1965,1991]" italics="true" pageId="216" pageNumber="217">Sciaenidarum</emphasis>
<taxonomicName id="57BE87D4FF5DFF77FE01989A4EEC08F0" authority="- Breard &amp; Stringer 1999: 135" authorityName="Breard &amp; Stringer" authorityPageNumber="135" authorityYear="1999" box="[455,935,1964,1991]" family="Sciaenidae" genus="Otolithus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="216" pageNumber="217" phylum="Chordata" rank="species" species="eporrectus">
<emphasis id="A2CA2045FF5DFF77FE01989A4F0908F0" box="[455,578,1966,1991]" italics="true" pageId="216" pageNumber="217">eporrectus</emphasis>
<bibRefCitation id="F42F81A6FF5DFF77FD9A98994EEC08F0" author="Breard S. &amp; Stringer G." box="[604,935,1964,1991]" pageId="216" pageNumber="217" pagination="132 - 142" refId="ref137334" refString="Breard S. &amp; Stringer G. 1999. Integrated paleoecology and marine vertebrate fauna of the Stone City Formation (middle Eocene), Brazos River section, Texas. Transactions of the Gulf Coast Association of Geological Societies 49: 132 - 142." type="journal article" year="1999">Breard &amp; Stringer 1999: 135</bibRefCitation>
</taxonomicName>
. —
<bibRefCitation id="F42F81A6FF5DFF77FC12989949D408F0" author="Muller A." box="[980,1183,1965,1991]" pageId="216" pageNumber="217" pagination="1 - 360" refId="ref146931" refString="Muller A. 1999. Ichthyofaunen aus dem atlantischen Tertiar der USA. Leipziger Geowissenschaften 9 - 10: 1 - 360." type="journal article" year="1999">Müller 1999: 160</bibRefCitation>
, pl. 33, figs 1015.
</paragraph>
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<emphasis id="A2CA2045FF5DFF77FF0C98E54C7508DC" box="[202,318,2001,2027]" italics="true" pageId="216" pageNumber="217">Sciaenida</emphasis>
<taxonomicName id="57BE87D4FF5DFF77FE9498E64FDD08DC" authority="- Nolf 2013: 107" authorityName="Nolf" authorityPageNumber="107" authorityYear="2013" box="[338,662,2001,2027]" genus="Ekokenia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="216" pageNumber="217" phylum="Chordata" rank="species" species="eporrecta">
<emphasis id="A2CA2045FF5DFF77FE9498E64C8808DC" box="[338,451,2002,2027]" italics="true" pageId="216" pageNumber="217">eporrecta</emphasis>
<bibRefCitation id="F42F81A6FF5DFF77FE2698E54FDD08DC" author="Nolf D." box="[480,662,2001,2027]" pageId="216" pageNumber="217" refId="ref147755" refString="Nolf D. 2013. The Diversity of Fish Otoliths, Past and Present. Operational directorate &quot; Earth and History of Life &quot; of the Royal Belgian Institute of Natural Sciences, Brussels." type="book" year="2013">Nolf 2013: 107</bibRefCitation>
</taxonomicName>
, pl. 281.
</paragraph>
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<emphasis id="A2CA2045FF5CFF76FF7B9E3F4CE10E12" bold="true" box="[189,426,267,293]" pageId="217" pageNumber="218">Material examined</emphasis>
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<paragraph id="9001FC57FF5CFF76FF7B9E0E4E210E40" blockId="217.[189,1399,313,375]" pageId="217" pageNumber="218">
<materialsCitation id="20D6F60AFF5CFF76FF7B9E0E4E2D0E40" ID-GBIF-Occurrence="2564277922" collectionCode="MSC, WSU" country="United States of America" location="Alabama" pageId="217" pageNumber="218" specimenCode="MSC 7300, CC 473.1" specimenCount="33" stateProvince="Alabama">
<collectingCountry id="E8A9BCC7FF5CFF76FF7B9E0E4F2E0E63" box="[189,613,314,340]" name="United States of America" pageId="217" pageNumber="218">UNITED STATES OF AMERICA</collectingCountry>
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<collectingRegion id="527A32B5FF5CFF76FD569E0D4E490E64" box="[656,770,313,339]" country="United States of America" name="Alabama" pageId="217" pageNumber="218">Alabama</collectingRegion>
</emphasis>
• 36 otoliths; GLS otolith comparative collection (
<specimenCount id="86B837DEFF5CFF76FF009E694C210E40" box="[198,362,348,375]" pageId="217" pageNumber="218" type="generic">33 specimens</specimenCount>
),
<specimenCode id="C018542CFF5CFF76FEB89E694F4A0E40" box="[382,513,349,375]" collectionCode="MSC" pageId="217" pageNumber="218">
<collectionCode id="F6AF6492FF5CFF76FEB89E694CF70E40" box="[382,444,349,375]" pageId="217" pageNumber="218">MSC</collectionCode>
7300
</specimenCode>
,
<collectionCode id="F6AF6492FF5CFF76FDCB9E694F000E40" box="[525,587,349,375]" pageId="217" pageNumber="218">MSC</collectionCode>
39048,
<collectionCode id="F6AF6492FF5CFF76FD6F9E694FA10E40" box="[681,746,349,375]" country="USA" httpUri="http://grbio.org/cool/1raa-g3is" name="Weber State University, Bird and Mammal Collection" pageId="217" pageNumber="218">WSU</collectionCode>
<specimenCode id="C018542CFF5CFF76FD349E694E2D0E40" box="[754,870,349,375]" collectionCode="WSU" country="USA" httpUri="http://grbio.org/cool/1raa-g3is" name="Weber State University, Bird and Mammal Collection" pageId="217" pageNumber="218">CC 473.1</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D8A4AFDCFF5CFF76FF7B9EAA4F030C87" pageId="217" pageNumber="218" type="description">
<paragraph id="9001FC57FF5CFF76FF7B9EAA4C050E8F" blockId="217.[189,334,414,440]" box="[189,334,414,440]" pageId="217" pageNumber="218">
<heading id="CB494B3BFF5CFF76FF7B9EAA4C050E8F" bold="true" box="[189,334,414,440]" fontSize="11" level="3" pageId="217" pageNumber="218" reason="3">
<emphasis id="A2CA2045FF5CFF76FF7B9EAA4C050E8F" bold="true" box="[189,334,414,440]" pageId="217" pageNumber="218">Description</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FF5CFF76FF7B9EF84F030C87" blockId="217.[189,1399,460,945]" pageId="217" pageNumber="218">
Sagittae range from approximately oval (smaller specimens) to elliptical (larger specimens). Inner face fairly smooth, convex. Adult specimens fairly large, exceeding
<quantity id="574651B2FF5CFF76FC3F9EC449040D3D" box="[1017,1103,496,522]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="217" pageNumber="218" unit="mm" value="10.0">10 mm</quantity>
in length. Height/length ratios range from approximately 6274%. Margins irregularly smooth on adult specimens, can be finely crenulate on small specimens.Anterior margin slopes backwards on smaller specimens, nearly vertical on larger specimens. Dorsal margin irregular, dorsal dome usually present. Posterodorsal dome is present on many specimens. Posterior margin straight to slightly incurved. Ventral margin broadly, evenly rounded. Very prominent heterosulcoid-type sulcus occurs on inner face, extending from anterior margin almost to posteroventral margin. Sulcus excavated, posterior slightly flexed downward. Ostium about onehalf the length of cauda, twice as wide. Ostium filled with colliculum. Sides of ostium approximately parallel. Cauda narrow (one-half width of ostium), slightly flexed downward at posterior. Posterior of cauda pointed. Very prominent depressed area, nearly rectangular in shape, above sulcus. Crista superior well-developed, forms distinct raised ridge above cauda. Ventral furrow appears absent. Outer face concave; concavity increases with size. Outer face excavated on dorsal half. Irregular undulations common on the larger specimens.
</paragraph>
</subSubSection>
<subSubSection id="D8A4AFDCFF5CFF76FF7B9CEC4C4C0A2A" pageId="217" pageNumber="218" type="discussion">
<paragraph id="9001FC57FF5CFF76FF7B9CEC4C640CC5" blockId="217.[189,303,984,1010]" box="[189,303,984,1010]" pageId="217" pageNumber="218">
<heading id="CB494B3BFF5CFF76FF7B9CEC4C640CC5" bold="true" box="[189,303,984,1010]" fontSize="11" level="3" pageId="217" pageNumber="218" reason="3">
<emphasis id="A2CA2045FF5CFF76FF7B9CEC4C640CC5" bold="true" box="[189,303,984,1010]" pageId="217" pageNumber="218">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FF5CFF76FF7B9B324E7D0BBD" blockId="217.[189,1399,1030,1162]" pageId="217" pageNumber="218">
<bibRefCitation id="F42F81A6FF5CFF76FF7B9B324C290B17" author="Koken E." box="[189,354,1030,1056]" pageId="217" pageNumber="218" pagination="274 - 305" refId="ref144486" refString="Koken E. 1888. Neue Untersuchungen an tertiaren Fisch-Otolithen. Zeitschrift der Deutschen Geologischen Gesellschaft 40: 274 - 305." type="journal article" year="1888">Koken (1888)</bibRefCitation>
first reported this taxon from Newton,
<collectingRegion id="527A32B5FF5CFF76FCF79B324EF10B17" box="[817,954,1030,1056]" country="United States of America" name="Mississippi" pageId="217" pageNumber="218">Mississippi</collectingRegion>
, and the formation from which it was found is now known to be the middle Eocene Cook Mountain Formation. The species also occurs in the Cook Mountain Formation of
<collectingRegion id="527A32B5FF5CFF76FDDD9B794F140B50" box="[539,607,1101,1127]" country="United States of America" name="Texas" pageId="217" pageNumber="218">Texas</collectingRegion>
(very common in the Stone City Member) and
<collectingRegion id="527A32B5FF5CFF76FB439B7849B70B51" box="[1157,1276,1100,1126]" country="United States of America" name="Louisiana" pageId="217" pageNumber="218">Louisiana</collectingRegion>
, as well as the middle Eocene Piney Point Formation of
<collectingRegion id="527A32B5FF5CFF76FD099B444E790BBD" box="[719,818,1136,1162]" country="United States of America" name="Virginia" pageId="217" pageNumber="218">Virginia</collectingRegion>
.
</paragraph>
<paragraph id="9001FC57FF5CFF76FF7B9B854E5F0BFC" blockId="217.[189,788,1201,1227]" box="[189,788,1201,1227]" pageId="217" pageNumber="218">
<heading id="CB494B3BFF5CFF76FF7B9B854E5F0BFC" bold="true" box="[189,788,1201,1227]" fontSize="11" level="3" pageId="217" pageNumber="218" reason="3">
<emphasis id="A2CA2045FF5CFF76FF7B9B854E5F0BFC" bold="true" box="[189,788,1201,1227]" pageId="217" pageNumber="218">
Stratigraphic and geographic range in
<collectingRegion id="527A32B5FF5CFF76FD649B854E5F0BFC" box="[674,788,1201,1227]" country="United States of America" name="Alabama" pageId="217" pageNumber="218">Alabama</collectingRegion>
</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FF5CFF76FF7B9BD44C4C0A2A" blockId="217.[189,1398,1248,1309]" pageId="217" pageNumber="218">
This species is common in the “upper”
<collectingRegion id="527A32B5FF5CFF76FD5B9BD44FA50BCD" box="[669,750,1248,1274]" country="Portugal" name="Lisboa" pageId="217" pageNumber="218">Lisbon</collectingRegion>
Formation at site ACh-8. Bartonian, zones NP16 and NP17.
</paragraph>
</subSubSection>
</treatment>
</document>