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<document id="991910EE030162501DEAD893EEA5E3A3" ID-DOI="10.5281/zenodo.213349" ID-GBIF-Dataset="e39fee2a-d7cb-4f2c-99f1-3a3b07ca6b9f" ID-ISSN="1175-5326" ID-Zenodo-Dep="213349" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460556320027" checkinUser="plazi" docAuthor="Shear, William A." docDate="2012" docId="136F879E905FFFF0FF00FE40FEF7FBC7" docLanguage="en" docName="zt03560p086.pdf" docOrigin="Zootaxa 3560" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pyrgeuma Shear, new genus" docType="treatment" docVersion="8" lastPageNumber="80" masterDocId="EF56FFE6905EFFF1FF97FFFEFFB0FFC8" masterDocTitle="Pyrgeuma pyrgodesmoides, n. gen., n. sp., a new millipede from Malaysia with unusual surface structures (Diplopoda, Chordeumatida, Heterochordeumatidae)" masterLastPageNumber="86" masterPageNumber="79" pageNumber="80" updateTime="1698319305980" updateUser="plazi">
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<mods:title id="EE53C7B544B5096FD79B4034B95F5BB5">Pyrgeuma pyrgodesmoides, n. gen., n. sp., a new millipede from Malaysia with unusual surface structures (Diplopoda, Chordeumatida, Heterochordeumatidae)</mods:title>
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<mods:namePart id="64904502680F27A9EBCD2FDD494ED0BC">Shear, William A.</mods:namePart>
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<mods:date id="F952ADC0ADEA67B76DFE41C4F60A39A4">2012</mods:date>
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<subSubSection id="D3DC6503905FFFF0FF00FE40FE41FE10" box="[151,497,446,472]" pageId="1" pageNumber="80" type="nomenclature">
<paragraph id="9B793688905FFFF0FF00FE40FE41FE10" blockId="1.[151,497,446,472]" box="[151,497,446,472]" pageId="1" pageNumber="80">
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<emphasis id="A9B2EA9A905FFFF0FF00FE40FE41FE10" bold="true" box="[151,497,446,472]" pageId="1" pageNumber="80">
<taxonomicName id="5CC64D0B905FFFF0FF00FE40FED2FE10" ID-CoL="7PX6K" authority="Shear" authorityName="Shear" box="[151,354,446,472]" class="Diplopoda" family="Heterochordeumatidae" genus="Pyrgeuma" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="80" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A9B2EA9A905FFFF0FF00FE40FEA3FE10" bold="true" box="[151,275,446,472]" italics="true" pageId="1" pageNumber="80">Pyrgeuma</emphasis>
Shear
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,
<taxonomicNameLabel id="B28157E1905FFFF0FEF8FE40FE41FE10" box="[367,497,446,472]" pageId="1" pageNumber="80" rank="genus">new genus</taxonomicNameLabel>
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<subSubSection id="D3DC6503905FFFF0FF00FDFEFD49FDDF" box="[151,761,511,536]" pageId="1" pageNumber="80" type="materials_examined">
<paragraph id="9B793688905FFFF0FF00FDFEFD49FDDF" blockId="1.[151,1436,511,1039]" box="[151,761,511,536]" pageId="1" pageNumber="80">
<emphasis id="A9B2EA9A905FFFF0FF00FDFEFE84FDD0" bold="true" box="[151,308,511,536]" pageId="1" pageNumber="80">
<typeStatus id="447D882A905FFFF0FF00FDFEFF62FDD0" box="[151,210,512,536]" pageId="1" pageNumber="80">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="5CC64D0B905FFFF0FEACFDFFFD02FDDF" authority="Shear" authorityName="Shear" box="[315,690,511,536]" class="Diplopoda" family="Heterochordeumatidae" genus="Pyrgeuma" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="80" phylum="Arthropoda" rank="species" species="pyrgodesmoides" status="sp. nov.">
<emphasis id="A9B2EA9A905FFFF0FEACFDFFFDD9FDD0" box="[315,617,512,536]" italics="true" pageId="1" pageNumber="80">Pyrgeuma pyrgodesmoides</emphasis>
Shear
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,
<taxonomicNameLabel id="B28157E1905FFFF0FD2AFDFEFD49FDDF" box="[701,761,512,535]" pageId="1" pageNumber="80" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="D3DC6503905FFFF0FF50FDDDFE3CFC0F" pageId="1" pageNumber="80" type="diagnosis">
<paragraph id="9B793688905FFFF0FF50FDDDFE3CFC0F" blockId="1.[151,1436,511,1039]" pageId="1" pageNumber="80">
<emphasis id="A9B2EA9A905FFFF0FF50FDDDFEF3FDF4" bold="true" box="[199,323,547,572]" pageId="1" pageNumber="80">Diagnosis:</emphasis>
Males of this genus, like
<taxonomicName id="5CC64D0B905FFFF0FD34FDDAFCABFDF4" box="[675,795,548,572]" class="Diplopoda" family="Heterochordeumatidae" genus="Infulathrix" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="A9B2EA9A905FFFF0FD34FDDAFCABFDF4" box="[675,795,548,572]" italics="true" pageId="1" pageNumber="80">Infulathrix</emphasis>
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males, have 30 rings, compared to the 32 rings of
<taxonomicName id="5CC64D0B905FFFF0FF00FDB6FED1FDA8" box="[151,353,584,608]" class="Diplopoda" family="Heterochordeumatidae" genus="Heterochordeuma" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="A9B2EA9A905FFFF0FF00FDB6FED1FDA8" box="[151,353,584,608]" italics="true" pageId="1" pageNumber="80">Heterochordeuma</emphasis>
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species males (females of all heterochordeumatids have 32 rings). Unlike species of
<emphasis id="A9B2EA9A905FFFF0FF00FD92FED7FD4C" box="[151,359,620,644]" italics="true" pageId="1" pageNumber="80">
<taxonomicName id="5CC64D0B905FFFF0FF00FD92FED3FD4C" box="[151,355,620,644]" class="Diplopoda" family="Heterochordeumatidae" genus="Heterochordeuma" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="80" phylum="Arthropoda" rank="genus">Heterochordeuma</taxonomicName>
,
</emphasis>
the anterior gonopods do not pass lateral and posterior to the posterior gonopod colpocoxites, and unlike
<taxonomicName id="5CC64D0B905FFFF0FE84FD6EFDE0FD6F" authority="Shear 2000" authorityName="Shear" authorityYear="2000" box="[275,592,655,680]" class="Diplopoda" family="Heterochordeumatidae" genus="Infulathrix" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="80" phylum="Arthropoda" rank="species" species="siam">
<emphasis id="A9B2EA9A905FFFF0FE84FD6EFE77FD60" box="[275,455,656,680]" italics="true" pageId="1" pageNumber="80">Infulathrix siam</emphasis>
<bibRefCitation id="FF574B79905FFFF0FE59FD71FDE0FD6F" author="Shear" box="[462,592,655,679]" pageId="1" pageNumber="80" refString="Shear, W. A. (2000) On the milliped family Heterochordeumatidae, with comments on the higher classification of the order Chordeumatida (Diplopoda). Invertebrate Taxonomy, 14, 363 - 376." type="journal article" year="2000">Shear 2000</bibRefCitation>
</taxonomicName>
, the male tenth coxae are not conspicuously enlarged and modified, nor are the third male femora enlarged. The integument of at least some species of heterochordeumatids appears slightly roughened, or velvety, due to small microtrichia. In
<emphasis id="A9B2EA9A905FFFF0FD71FD27FCEFFD38" box="[742,863,729,752]" italics="true" pageId="1" pageNumber="80">Pyrgeuma,</emphasis>
these microtrichia are larger, curved and spatulate, and give the dorsal surface a coarsely rough appearance. Beginning on the first segment after the collum, the metazonites bear a pair of elevations that become more prominent posteriorly. The six metazonital setae characteristic of chordeumatidans are not detectable at dissecting microscope magnification; they are no larger than the exaggerated microtrichia on midbody segments. Under the scanning electron microscope (SEM) they can be seen to be blunt, with 57 prominent longitudinal ridges. Further details are given in the description below of the single known species.
</paragraph>
</subSubSection>
<subSubSection id="D3DC6503905FFFF0FF50FC2AFEF7FBC7" pageId="1" pageNumber="80" type="etymology">
<paragraph id="9B793688905FFFF0FF50FC2AFEF7FBC7" blockId="1.[151,1436,511,1039]" pageId="1" pageNumber="80">
<emphasis id="A9B2EA9A905FFFF0FF50FC2AFEE1FC24" bold="true" box="[199,337,980,1004]" pageId="1" pageNumber="80">Etymology:</emphasis>
The name of the genus (neuter) draws attention to the resemblance of the species to pyrgodesmid polydesmidans.
</paragraph>
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