226 lines
27 KiB
XML
226 lines
27 KiB
XML
<document id="BA1B22F95D0B2D5DAC0FA1CA32D24A80" ID-DOI="10.11646/zootaxa.5319.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8182108" ID-ZooBank="45CD8E29-3320-42E2-A8A8-3A36BC0F41CE" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1690276500814" checkinUser="plazi" docAuthor="Hashizume, Takuto, Yamamoto, Shûhei & Maruyama, Munetoshi" docDate="2023" docId="090C879DFFE28E1C899AFE3CA97FFA93" docLanguage="en" docName="zootaxa.5319.1.2.pdf" docOrigin="Zootaxa 5319 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5319.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudatheta Cameron 1920" docType="treatment" docVersion="1" lastPageNumber="29" masterDocId="F535FFE5FFE08E1E890DFFDAA87BFFDA" masterDocTitle="Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan" masterLastPageNumber="47" masterPageNumber="27" pageNumber="29" updateTime="1690313925549" updateUser="jonas" zenodo-license-document="CLOSED">
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<mods:title id="FAE16DF34991936FEDA2D6D6FF2C8EBA">Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan</mods:title>
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<mods:name id="EAD656D596CFD4DC0D7726BD450C50A5" type="personal">
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<mods:roleTerm id="E7F5A4F7C526E3D7E05149EA8408433E">Author</mods:roleTerm>
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<mods:namePart id="ABA50D9BC4C8B9188F6D55838A854FD0">Hashizume, Takuto</mods:namePart>
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<mods:affiliation id="3DD674413DE9345D6FE9009D9C08B4E2">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, 819 - 0395 Japan.</mods:affiliation>
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<mods:name id="04504BEFF13D60234A09857FEE31CAF3" type="personal">
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<mods:roleTerm id="F5039AB352A4AC3E6F9A3FCB062198E3">Author</mods:roleTerm>
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<mods:namePart id="B9FCBAD107B611CE361B6F9C35D50AED">Yamamoto, Shûhei</mods:namePart>
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<mods:nameIdentifier id="69E44762EF165AAC4A59CA4D161855FE" type="ORCID">0000-0002-4162-8457</mods:nameIdentifier>
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<mods:affiliation id="05AFEA9CF37F0FF345CC6C5653CACD70">The Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo, 060 - 0810 Japan. s. yamamoto. 64 @ gmail. com; https: // orcid. org / 0000 - 0002 - 4162 - 8457</mods:affiliation>
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<mods:nameIdentifier id="C3C8037160F8C15B58036AC916C3CF17" type="email">s.yamamoto.64@gmail.com</mods:nameIdentifier>
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<mods:namePart id="E2E3DA123A2E5D35EFFBB841A9F09E7C">Maruyama, Munetoshi</mods:namePart>
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<mods:affiliation id="A733665DB4F12CA6DE9B6D9346513E29">The Kyushu University Museum, Fukuoka, 812 - 8581 Japan.</mods:affiliation>
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<mods:title id="75675B9CDDFF8975CE19E2172B93DEAA">Zootaxa</mods:title>
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<mods:date id="E1EFE43192EC8E93F3C0C9C3A205DAF1">2023</mods:date>
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<mods:number id="A2462E7BEBE7DA16A73C160471EB0BE8">2023-07-24</mods:number>
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<mods:classification id="7EF56F393C0BD9EF57F4682AB584C2D5">journal article</mods:classification>
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<treatment id="090C879DFFE28E1C899AFE3CA97FFA93" ID-DOI="http://doi.org/10.5281/zenodo.8182110" ID-Zenodo-Dep="8182110" LSID="urn:lsid:plazi:treatment:090C879DFFE28E1C899AFE3CA97FFA93" httpUri="http://treatment.plazi.org/id/090C879DFFE28E1C899AFE3CA97FFA93" lastPageNumber="29" pageId="2" pageNumber="29">
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<subSubSection id="C9BF6500FFE28E1C899AFE3CAA33FDDA" box="[151,584,486,513]" pageId="2" pageNumber="29" type="nomenclature">
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<paragraph id="811A368BFFE28E1C899AFE3CAA33FDDA" blockId="2.[151,832,486,549]" box="[151,584,486,513]" pageId="2" pageNumber="29">
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<heading id="DA5281E7FFE28E1C899AFE3CAA33FDDA" bold="true" box="[151,584,486,513]" fontSize="11" level="1" pageId="2" pageNumber="29" reason="1">
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<emphasis id="B3D1EA99FFE28E1C899AFE3CAA33FDDA" bold="true" box="[151,584,486,513]" pageId="2" pageNumber="29">
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Genus
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<taxonomicName id="46A54D08FFE28E1C89E3FE3CAA33FDDA" ID-CoL="8W2RP" authority="Cameron, 1920" authorityName="Cameron" authorityYear="1920" box="[238,584,486,513]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis id="B3D1EA99FFE28E1C89E3FE3CA9F9FDDB" bold="true" box="[238,386,486,513]" italics="true" pageId="2" pageNumber="29">Pseudatheta</emphasis>
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<bibRefCitation id="E5344B7AFFE28E1C8887FE3CAA33FDDA" author="Cameron, M." box="[394,584,486,512]" pageId="2" pageNumber="29" pagination="212 - 284" refId="ref15189" refString="Cameron, M. (1920) New species of Staphylinidae from Singapore. Part III. Transactions of the Entomological Society of London, 1920, 212 - 284. https: // doi. org / 10.1111 / j. 1365 - 2311.1920. tb 00214. x" type="journal article" year="1920">Cameron, 1920</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</paragraph>
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<subSubSection id="C9BF6500FFE28E1C899AFDD1AB3BFDFF" box="[151,832,523,549]" pageId="2" pageNumber="29" type="vernacular_names">
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<paragraph id="811A368BFFE28E1C899AFDD1AB3BFDFF" blockId="2.[151,832,486,549]" box="[151,832,523,549]" pageId="2" pageNumber="29">[Japanese name: Hime-kinokotsuyakeshi-hanekakushi-zoku]</paragraph>
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</subSubSection>
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<subSubSection id="C9BF6500FFE28E1C899AFD90AABDFDB8" box="[151,710,586,610]" pageId="2" pageNumber="29" type="reference_group">
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<paragraph id="811A368BFFE28E1C899AFD90AABDFDB8" blockId="2.[151,1240,586,610]" box="[151,710,586,610]" pageId="2" pageNumber="29">
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<treatmentCitationGroup id="A1B511A5FFE28E1C899AFD90AABDFDB8" box="[151,710,586,610]" pageId="2" pageNumber="29">
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<treatmentCitation id="0004109AFFE28E1C899AFD90A991FDB8" author="Cameron, M." box="[151,490,586,610]" page="224" pageId="2" pageNumber="29" year="1920">
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<taxonomicName id="46A54D08FFE28E1C899AFD90A991FDB8" ID-CoL="8W2RP" authority="Cameron, 1920: 224" authorityName="Cameron" authorityPageNumber="224" authorityYear="1920" box="[151,490,586,610]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis id="B3D1EA99FFE28E1C899AFD90A96FFDB8" box="[151,276,586,610]" italics="true" pageId="2" pageNumber="29">Pseudatheta</emphasis>
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<bibRefCitation id="E5344B7AFFE28E1C8816FD90A991FDB8" author="Cameron, M." box="[283,490,586,610]" pageId="2" pageNumber="29" pagination="212 - 284" refId="ref15189" refString="Cameron, M. (1920) New species of Staphylinidae from Singapore. Part III. Transactions of the Entomological Society of London, 1920, 212 - 284. https: // doi. org / 10.1111 / j. 1365 - 2311.1920. tb 00214. x" type="journal article" year="1920">Cameron, 1920: 224</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(original description;
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</treatmentCitationGroup>
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</paragraph>
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<subSubSection id="C9BF6500FFE28E1C8BC1FD90ACA3FDB8" box="[716,1240,586,610]" pageId="2" pageNumber="29" type="type_taxon">
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<paragraph id="811A368BFFE28E1C8BC1FD90ACA3FDB8" blockId="2.[151,1240,586,610]" box="[716,1240,586,610]" pageId="2" pageNumber="29">
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<typeStatus id="5E1E8829FFE28E1C8BC1FD90AA83FDB8" box="[716,760,586,610]" pageId="2" pageNumber="29">type</typeStatus>
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species:
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<taxonomicName id="46A54D08FFE28E1C8A58FD90ACB7FDB8" authority="Cameron, 1920" authorityName="Cameron" authorityYear="1920" box="[853,1228,586,610]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="elegans">
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<emphasis id="B3D1EA99FFE28E1C8A58FD90AC5EFDB8" box="[853,1061,586,610]" italics="true" pageId="2" pageNumber="29">Pseudatheta elegans</emphasis>
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<bibRefCitation id="E5344B7AFFE28E1C8D21FD90ACB7FDB8" author="Cameron, M." box="[1068,1228,586,610]" pageId="2" pageNumber="29" pagination="212 - 284" refId="ref15189" refString="Cameron, M. (1920) New species of Staphylinidae from Singapore. Part III. Transactions of the Entomological Society of London, 1920, 212 - 284. https: // doi. org / 10.1111 / j. 1365 - 2311.1920. tb 00214. x" type="journal article" year="1920">Cameron, 1920</bibRefCitation>
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</taxonomicName>
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).
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||
</paragraph>
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<subSubSection id="C9BF6500FFE28E1C899AFD51AB6BFCEF" pageId="2" pageNumber="29" type="discussion">
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<paragraph id="811A368BFFE28E1C899AFD51AB6BFCEF" blockId="2.[151,1437,651,821]" pageId="2" pageNumber="29">
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<emphasis id="B3D1EA99FFE28E1C899AFD51A970FD7F" bold="true" box="[151,267,651,677]" pageId="2" pageNumber="29">Remarks.</emphasis>
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Species of this genus often have intraspecific variation in body coloration, the ratio of the length and width of each antennomere, and the processes on the posterior margin of the tergite VIII of males, which do not necessarily correspond to those described or illustrated in this paper. The genus
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<taxonomicName id="46A54D08FFE28E1C8D2EFD09ACD6FD37" authorityName="Cameron" authorityYear="1920" box="[1059,1197,723,749]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis id="B3D1EA99FFE28E1C8D2EFD09ACD6FD37" box="[1059,1197,723,749]" italics="true" pageId="2" pageNumber="29">Pseudatheta</emphasis>
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</taxonomicName>
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can be distinguished from the genus
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<taxonomicName id="46A54D08FFE28E1C884FFD2DA9C5FCCB" box="[322,446,759,785]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis id="B3D1EA99FFE28E1C884FFD2DA9C5FCCB" box="[322,446,759,785]" italics="true" pageId="2" pageNumber="29">Phymatura</emphasis>
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</taxonomicName>
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by following character states: mesoventrite without longitudinal carina; number of setose lobes on flabellum of hind wing varies from zero to one.
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</paragraph>
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</subSubSection>
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<subSubSection id="C9BF6500FFE28E1C899AFC5DAA8FFC7B" box="[151,756,902,929]" pageId="2" pageNumber="29" type="multiple">
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<paragraph id="811A368BFFE28E1C899AFC5DAA8FFC7B" blockId="2.[151,756,902,929]" box="[151,756,902,929]" pageId="2" pageNumber="29">
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<heading id="DA5281E7FFE28E1C899AFC5DAA8FFC7B" bold="true" box="[151,756,902,929]" fontSize="11" level="1" pageId="2" pageNumber="29" reason="1">
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<emphasis id="B3D1EA99FFE28E1C899AFC5DAA8FFC7B" bold="true" box="[151,756,902,929]" pageId="2" pageNumber="29">
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Key to species of the genus
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||
<taxonomicName id="46A54D08FFE28E1C88E1FC5CAAFBFC7B" ID-CoL="8W2RP" authorityName="Cameron" authorityYear="1920" box="[492,640,902,929]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis id="B3D1EA99FFE28E1C88E1FC5CAAFBFC7B" bold="true" box="[492,640,902,929]" italics="true" pageId="2" pageNumber="29">Pseudatheta</emphasis>
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</taxonomicName>
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in
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<collectingCountry id="F9B2761BFFE28E1C8BAAFC5DAA8FFC7B" box="[679,756,903,929]" name="Japan" pageId="2" pageNumber="29">Japan</collectingCountry>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C9BF6500FFE28E1C899AFC1FADE7FB38" pageId="2" pageNumber="29" type="key">
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<keyStep id="3A512E2EFFE28E1C899AFC1FADE7FBE8" pageId="2" pageNumber="29">
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<paragraph id="811A368BFFE28E1C899AFC1FADE7FC2D" blockId="2.[151,1437,965,1251]" pageId="2" pageNumber="29">
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<keyLead id="3A5495BEFFE28E1C899AFC1FADE7FC2D" pageId="2" pageNumber="29">
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1. Body larger, forebody length 1.00–
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<quantity id="465D9B6EFFE28E1C8B34FC1FAAE9FC00" box="[569,658,965,987]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="2" pageNumber="29" unit="mm" value="1.25">1.25 mm</quantity>
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; male sternite VI with small medial lobe on posterior margin (
|
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<figureCitation id="199E2A0EFFE28E1C8DF8FC1FAD38FC00" box="[1269,1347,965,987]" captionStart="FIGURE 4" captionStartId="7.[151,250,627,651]" captionTargetBox="[164,1419,309,603]" captionTargetId="figure-90@7.[151,1436,309,603]" captionTargetPageId="7" captionText="FIGURE 4. Pseudatheta taiwanensis Pace, 2008. A. male elytra (white arrow indicates granules); B. male tergite VII (white arrow indicates granule); C. male sternite VI (white arrow indicates medial lobe)." figureDoi="http://doi.org/10.5281/zenodo.8182120" httpUri="https://zenodo.org/record/8182120/files/figure.png" pageId="2" pageNumber="29">Fig. 4C</figureCitation>
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); median lobe of aedeagus without ventral process; distal portion of spermatheca long, strongly curved (
|
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<figureCitation id="199E2A0EFFE28E1C8D60FC38ACABFC2D" box="[1133,1232,994,1016]" captionStart="FIGURE 5" captionStartId="8.[151,250,1626,1650]" captionTargetBox="[161,1430,250,1592]" captionTargetId="figure-73@8.[151,1436,237,1603]" captionTargetPageId="8" captionText="FIGURE 5. Pseudatheta spp. A–C: Pseudatheta crenulicauda (Bernhauer, 1907), comb. nov., D–F: Pseudatheta taiwanensis Pace, 2008. A, D. male tergite VIII in dorsal view; B, E. median lobe of aedeagus in lateral view; C, F. spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182126" httpUri="https://zenodo.org/record/8182126/files/figure.png" pageId="2" pageNumber="29">Fig. 5C, F</figureCitation>
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).............. 2
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</keyLead>
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</paragraph>
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<paragraph id="811A368BFFE28E1C899AFBDAADE7FBE8" blockId="2.[151,1437,965,1251]" pageId="2" pageNumber="29">
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<keyLead id="3A5495BEFFE28E1C899AFBDAADE7FBE8" pageId="2" pageNumber="29">
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- Body smaller, forebody length
|
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<quantity id="465D9B6EFFE28E1C8B14FBDAAAD0FBCF" box="[537,683,1024,1046]" metricMagnitude="-4" metricUnit="m" metricValue="9.35" metricValueMax="10.1" metricValueMin="8.6" pageId="2" pageNumber="29" unit="mm" value="0.935" valueMax="1.01" valueMin="0.86">0.86–1.01 mm</quantity>
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; male sternite VI without medial lobe on posterior margin; median lobe of aedeagus with ventral process; distal portion of spermatheca short, not curved or slightly curved (
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||
<figureCitation id="199E2A0EFFE28E1C8D84FBC7AC89FBE8" box="[1161,1266,1053,1075]" captionStart="FIGURE 7" captionStartId="12.[151,250,1890,1914]" captionTargetBox="[161,1422,497,1858]" captionTargetId="figure-190@12.[151,1436,489,1866]" captionTargetPageId="12" captionText="FIGURE 7. Pseudatheta spp. A–D: Ps. cooteri Pace, 1998, E–G: Ps. elegans Cameron, 1920. A, E. male tergite VIII in dorsal view; B, F. median lobe of aedeagus in lateral view; D, G. spermatheca; C. ventral process of median lobe of aedeagus in lateral view, a: Yaku-shima Is., b: Iriomote-jima Is. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182134" httpUri="https://zenodo.org/record/8182134/files/figure.png" pageId="2" pageNumber="29">Fig. 7D, G</figureCitation>
|
||
)............ 3
|
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</keyLead>
|
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</paragraph>
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</keyStep>
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||
<paragraph id="811A368BFFE28E1C899AFBE0ADE7FB95" blockId="2.[151,1437,965,1251]" box="[151,1436,1082,1104]" pageId="2" pageNumber="29">
|
||
<keyLead id="3A5495BEFFE28E1C899AFBE0ADE7FB95" box="[151,1436,1082,1104]" pageId="2" pageNumber="29">
|
||
2. Apical process of aedeagus dilated apically in lateral view (
|
||
<figureCitation id="199E2A0EFFE28E1C8A10FBE0AB1CFB8A" box="[797,871,1082,1104]" captionStart="FIGURE 5" captionStartId="8.[151,250,1626,1650]" captionTargetBox="[161,1430,250,1592]" captionTargetId="figure-73@8.[151,1436,237,1603]" captionTargetPageId="8" captionText="FIGURE 5. Pseudatheta spp. A–C: Pseudatheta crenulicauda (Bernhauer, 1907), comb. nov., D–F: Pseudatheta taiwanensis Pace, 2008. A, D. male tergite VIII in dorsal view; B, E. median lobe of aedeagus in lateral view; C, F. spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182126" httpUri="https://zenodo.org/record/8182126/files/figure.png" pageId="2" pageNumber="29">Fig. 5B</figureCitation>
|
||
)...........
|
||
<taxonomicName id="46A54D08FFE28E1C8D0AFBE1AD5EFB8A" authority="(Bernhauer)" authorityName="Hashizume & Yamamoto & Maruyama" authorityYear="2023" baseAuthorityName="Bernhauer" baseAuthorityYear="1907" box="[1031,1317,1082,1104]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="crenulicauda" status="comb. nov.">
|
||
<emphasis id="B3D1EA99FFE28E1C8D0AFBE1ACD2FB95" box="[1031,1193,1082,1104]" italics="true" pageId="2" pageNumber="29">Ps. crenulicauda</emphasis>
|
||
(Bernhauer)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B3D1EA99FFE28E1C8C23FBE0ADE7FB95" bold="true" box="[1326,1436,1082,1103]" pageId="2" pageNumber="29">
|
||
<taxonomicNameLabel id="A8E257E2FFE28E1C8C23FBE0ADE7FB95" box="[1326,1436,1082,1103]" pageId="2" pageNumber="29" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="811A368BFFE28E1C899AFB82ADE7FBB7" blockId="2.[151,1437,965,1251]" box="[151,1436,1111,1134]" pageId="2" pageNumber="29">
|
||
<keyLead id="3A5495BEFFE28E1C899AFB82ADE7FBB7" box="[151,1436,1111,1134]" pageId="2" pageNumber="29">
|
||
- Apical process of aedeagus parallel to near apex, not dilated in lateral view (
|
||
<figureCitation id="199E2A0EFFE28E1C8AB2FB82AC73FBB7" box="[959,1032,1111,1134]" captionStart="FIGURE 5" captionStartId="8.[151,250,1626,1650]" captionTargetBox="[161,1430,250,1592]" captionTargetId="figure-73@8.[151,1436,237,1603]" captionTargetPageId="8" captionText="FIGURE 5. Pseudatheta spp. A–C: Pseudatheta crenulicauda (Bernhauer, 1907), comb. nov., D–F: Pseudatheta taiwanensis Pace, 2008. A, D. male tergite VIII in dorsal view; B, E. median lobe of aedeagus in lateral view; C, F. spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182126" httpUri="https://zenodo.org/record/8182126/files/figure.png" pageId="2" pageNumber="29">Fig. 5E</figureCitation>
|
||
)...............
|
||
<taxonomicName id="46A54D08FFE28E1C8DD9FB82ADE7FBB7" authority="Pace" authorityName="Pace" authorityYear="2008" box="[1236,1436,1112,1133]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="taiwanensis">
|
||
<emphasis id="B3D1EA99FFE28E1C8DD9FB82AD12FBB7" box="[1236,1385,1112,1133]" italics="true" pageId="2" pageNumber="29">Ps. taiwanensis</emphasis>
|
||
Pace
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="3A512E2EFFE28E1C899AFBAFADE7FB38" pageId="2" pageNumber="29">
|
||
<paragraph id="811A368BFFE28E1C899AFBAFADE7FB72" blockId="2.[151,1437,965,1251]" pageId="2" pageNumber="29">
|
||
<keyLead id="3A5495BEFFE28E1C899AFBAFADE7FB72" pageId="2" pageNumber="29">
|
||
3. Apical process of median lobe of aedeagus simply curved (
|
||
<figureCitation id="199E2A0EFFE28E1C8A1BFBAFAB19FB50" box="[790,866,1141,1163]" captionStart="FIGURE 7" captionStartId="12.[151,250,1890,1914]" captionTargetBox="[161,1422,497,1858]" captionTargetId="figure-190@12.[151,1436,489,1866]" captionTargetPageId="12" captionText="FIGURE 7. Pseudatheta spp. A–D: Ps. cooteri Pace, 1998, E–G: Ps. elegans Cameron, 1920. A, E. male tergite VIII in dorsal view; B, F. median lobe of aedeagus in lateral view; D, G. spermatheca; C. ventral process of median lobe of aedeagus in lateral view, a: Yaku-shima Is., b: Iriomote-jima Is. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182134" httpUri="https://zenodo.org/record/8182134/files/figure.png" pageId="2" pageNumber="29">Fig. 7B</figureCitation>
|
||
); ventral process of median lobe of aedeagus with a distinct angle on dorsal side (
|
||
<figureCitation id="199E2A0EFFE28E1C88A3FB48A981FB7D" box="[430,506,1170,1192]" captionStart="FIGURE 7" captionStartId="12.[151,250,1890,1914]" captionTargetBox="[161,1422,497,1858]" captionTargetId="figure-190@12.[151,1436,489,1866]" captionTargetPageId="12" captionText="FIGURE 7. Pseudatheta spp. A–D: Ps. cooteri Pace, 1998, E–G: Ps. elegans Cameron, 1920. A, E. male tergite VIII in dorsal view; B, F. median lobe of aedeagus in lateral view; D, G. spermatheca; C. ventral process of median lobe of aedeagus in lateral view, a: Yaku-shima Is., b: Iriomote-jima Is. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182134" httpUri="https://zenodo.org/record/8182134/files/figure.png" pageId="2" pageNumber="29">Fig. 7C</figureCitation>
|
||
); distal portion of spermatheca nearly conical shape (
|
||
<figureCitation id="199E2A0EFFE28E1C8AF7FB48AC3EFB7D" box="[1018,1093,1170,1192]" captionStart="FIGURE 7" captionStartId="12.[151,250,1890,1914]" captionTargetBox="[161,1422,497,1858]" captionTargetId="figure-190@12.[151,1436,489,1866]" captionTargetPageId="12" captionText="FIGURE 7. Pseudatheta spp. A–D: Ps. cooteri Pace, 1998, E–G: Ps. elegans Cameron, 1920. A, E. male tergite VIII in dorsal view; B, F. median lobe of aedeagus in lateral view; D, G. spermatheca; C. ventral process of median lobe of aedeagus in lateral view, a: Yaku-shima Is., b: Iriomote-jima Is. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182134" httpUri="https://zenodo.org/record/8182134/files/figure.png" pageId="2" pageNumber="29">Fig. 7D</figureCitation>
|
||
)..............
|
||
<taxonomicName id="46A54D08FFE28E1C8C0CFB49ADE7FB72" authority="Pace" authorityName="Pace" authorityYear="1998" box="[1281,1436,1171,1192]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="cooteri">
|
||
<emphasis id="B3D1EA99FFE28E1C8C0CFB49AD12FB72" box="[1281,1385,1171,1192]" italics="true" pageId="2" pageNumber="29">Ps. cooteri</emphasis>
|
||
Pace
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="811A368BFFE28E1C899AFB6AADE7FB38" blockId="2.[151,1437,965,1251]" pageId="2" pageNumber="29">
|
||
<keyLead id="3A5495BEFFE28E1C899AFB6AADE7FB38" pageId="2" pageNumber="29">
|
||
- Apical process of median lobe of aedeagus sinuate (
|
||
<figureCitation id="199E2A0EFFE28E1C8BDCFB6AAB60FB1F" box="[721,795,1200,1222]" captionStart="FIGURE 7" captionStartId="12.[151,250,1890,1914]" captionTargetBox="[161,1422,497,1858]" captionTargetId="figure-190@12.[151,1436,489,1866]" captionTargetPageId="12" captionText="FIGURE 7. Pseudatheta spp. A–D: Ps. cooteri Pace, 1998, E–G: Ps. elegans Cameron, 1920. A, E. male tergite VIII in dorsal view; B, F. median lobe of aedeagus in lateral view; D, G. spermatheca; C. ventral process of median lobe of aedeagus in lateral view, a: Yaku-shima Is., b: Iriomote-jima Is. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182134" httpUri="https://zenodo.org/record/8182134/files/figure.png" pageId="2" pageNumber="29">Fig. 7F</figureCitation>
|
||
); ventral process of median lobe of aedeagus sinuate; distal portion of spermatheca nearly reniform (
|
||
<figureCitation id="199E2A0EFFE28E1C8B10FB17AA12FB38" box="[541,617,1229,1251]" captionStart="FIGURE 7" captionStartId="12.[151,250,1890,1914]" captionTargetBox="[161,1422,497,1858]" captionTargetId="figure-190@12.[151,1436,489,1866]" captionTargetPageId="12" captionText="FIGURE 7. Pseudatheta spp. A–D: Ps. cooteri Pace, 1998, E–G: Ps. elegans Cameron, 1920. A, E. male tergite VIII in dorsal view; B, F. median lobe of aedeagus in lateral view; D, G. spermatheca; C. ventral process of median lobe of aedeagus in lateral view, a: Yaku-shima Is., b: Iriomote-jima Is. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182134" httpUri="https://zenodo.org/record/8182134/files/figure.png" pageId="2" pageNumber="29">Fig. 7G</figureCitation>
|
||
)..................................................
|
||
<taxonomicName id="46A54D08FFE28E1C8DDCFB14ADE7FB38" authority="Cameron" authorityName="Cameron" authorityYear="1920" box="[1233,1436,1229,1251]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="elegans">
|
||
<emphasis id="B3D1EA99FFE28E1C8DDCFB14AD45FB38" box="[1233,1342,1229,1251]" italics="true" pageId="2" pageNumber="29">Ps. elegans</emphasis>
|
||
Cameron
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
<subSubSection id="C9BF6500FFE28E1C899AFAD1A97FFA93" pageId="2" pageNumber="29" type="discussion">
|
||
<paragraph id="811A368BFFE28E1C899AFAD1A97FFA93" blockId="2.[151,1436,1291,1353]" pageId="2" pageNumber="29">
|
||
<emphasis id="B3D1EA99FFE28E1C899AFAD1A970FAFF" bold="true" box="[151,267,1291,1317]" pageId="2" pageNumber="29">Remarks.</emphasis>
|
||
<taxonomicName id="46A54D08FFE28E1C881FFAD1AA32FAFF" authority="(Sharp)" authorityName="Hashizume & Yamamoto & Maruyama" authorityYear="2023" baseAuthorityName="Sharp" baseAuthorityYear="1888" box="[274,585,1291,1317]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="hilaris" status="comb. nov.">
|
||
<emphasis id="B3D1EA99FFE28E1C881FFAD1A990FAFF" box="[274,491,1291,1317]" italics="true" pageId="2" pageNumber="29">Pseudatheta hilaris</emphasis>
|
||
(Sharp)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B3D1EA99FFE28E1C8B59FAD1AAAFFAFF" bold="true" box="[596,724,1291,1317]" pageId="2" pageNumber="29">
|
||
<taxonomicNameLabel id="A8E257E2FFE28E1C8B59FAD1AAAFFAFF" box="[596,724,1291,1317]" pageId="2" pageNumber="29" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, is excluded from this key because only female specimens were available.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |