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<document id="060EC2FAB97565F03E2AE3B3DB2DBC59" ID-DOI="10.5852/ejt.2016.238" ID-GBIF-Dataset="8b57d1e6-b2b7-4229-912a-c042457efaaf" ID-ISSN="2118-9773" ID-Zenodo-Dep="3849874" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1590439951474" checkinUser="valdenar" docAuthor="Doczkal, Dieter, Radenković, Snežana, Lyneborg, Leif &amp; Pape, Thomas" docDate="2016" docId="03EA327EFFE3EC13FF27EFF1FB5BFE93" docLanguage="en" docName="ejt-2016-238.pdf.imf" docOrigin="European Journal of Taxonomy 238" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Megatrigon Johnson 1898" docType="treatment" docVersion="2" lastPageNumber="4" masterDocId="FFD34A06FFE1EC17FF9AEA71FFC7FFFF" masterDocTitle="Taxonomic revision of the Afrotropical genus Megatrigon Johnson, 1898 (Diptera: Syrphidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="3" updateTime="1734461804313" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B54536E313876C2574516B5A26EC681E">Taxonomic revision of the Afrotropical genus Megatrigon Johnson, 1898 (Diptera: Syrphidae)</mods:title>
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<mods:namePart id="7F9F62AC53C7F5BE7A5E10DCB95693FF">Doczkal, Dieter</mods:namePart>
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<mods:affiliation id="1D2A86FB84855C4DC280DF7AA266E831">Bavarian State Collection of Zoology, Zoologische Staatssammlung, Munich, Germany.</mods:affiliation>
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<mods:namePart id="F08E6E9FA1671B8487F5129465F96500">Radenković, Snežana</mods:namePart>
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<mods:affiliation id="05651E26378A60DD45587A2B3FFF902C">University of Novi Sad, Department of Biology and Ecology, Trg Dositeja Obradovica 2, 21000 Novi Sad, Serbia.</mods:affiliation>
<mods:nameIdentifier id="103371F3122E0B954215DF9668BCA7B3" type="email">snezana.radenkovic@dbe.uns.ac.rs</mods:nameIdentifier>
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<mods:namePart id="CF08EB636D88AA5E9294D937A0F555E8">Lyneborg, Leif</mods:namePart>
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<mods:affiliation id="A91FC5129064484AD744B0AAEF31F5BA">Natural History Museum of Denmark, Universitetsparken 15, 2100 Copenhagen, Denmark.</mods:affiliation>
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<mods:namePart id="4C0A597494C2D08FC60239EF5EA56C92">Pape, Thomas</mods:namePart>
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<mods:affiliation id="E01D28DEE31483EBD3BCC197DDBA2D84">Natural History Museum of Denmark, Universitetsparken 15, 2100 Copenhagen, Denmark.</mods:affiliation>
<mods:nameIdentifier id="1A0A7F31B9B077A8D1CE583743F05D32" type="email">tpape@snm.ku.dk&amp;tpape@snm.ku.dk</mods:nameIdentifier>
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<treatment id="03EA327EFFE3EC13FF27EFF1FB5BFE93" ID-DOI="http://doi.org/10.5281/zenodo.14510416" ID-Zenodo-Dep="14510416" LSID="urn:lsid:plazi:treatment:03EA327EFFE3EC13FF27EFF1FB5BFE93" httpUri="http://treatment.plazi.org/id/03EA327EFFE3EC13FF27EFF1FB5BFE93" lastPageId="4" lastPageNumber="4" pageId="2" pageNumber="3" scope_class="Insecta" scope_family="Syrphidae" scope_order="Diptera" scope_subFamily="Eristalinae" scope_tribe="Merodontini">
<subSubSection id="C359D0E3FFE3EC15FF27EFF1FEB6FA43" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BFC8368FFE3EC15FF27EFF1FEB6FA43" blockId="2.[189,1398,1407,1468]" pageId="2" pageNumber="3">
<taxonomicName id="4C43F8EBFFE3EC15FF27EFF1FDE9FA65" authority="Johnson, 1898: 159" authorityName="Johnson" authorityPageNumber="159" authorityYear="1898" box="[189,558,1407,1434]" class="Insecta" family="Syrphidae" genus="Megatrigon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B9375F7AFFE3EC15FF27EFF1FE8FFA66" box="[189,328,1408,1433]" italics="true" pageId="2" pageNumber="3">Megatrigon</emphasis>
<bibRefCitation id="EFD2FE99FFE3EC15FED7EF0EFDE9FA65" author="Johnson C. W." box="[333,558,1407,1434]" pageId="2" pageNumber="3" pagination="157 - 164" refId="ref17020" refString="Johnson C. W. 1898. Diptera collected by Dr. A. Donaldson Smith in Somaliland, eastern Africa. Proceedings of the Academy of Natural Sciences Philadelphia 50: 157 - 164. Available from http: // biodiversitylibrary. org / page / 26368835 [accessed 6 Sep. 2016]" type="journal article" year="1898">Johnson, 1898: 159</bibRefCitation>
</taxonomicName>
stat. nov. (resurrected as a valid genus from the synonymy with
<taxonomicName id="4C43F8EBFFE3EC15FA94EFF1FEA3FA43" authority="Meigen, 1822" authorityName="Meigen" authorityYear="1822" class="Insecta" family="Syrphidae" genus="Eumerus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B9375F7AFFE3EC15FA94EFF1FAB1FA66" box="[1294,1398,1408,1433]" italics="true" pageId="2" pageNumber="3">Eumerus</emphasis>
Meigen, 1822
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C359D0E3FFE3EC15FF27EF92FD52F9D3" pageId="2" pageNumber="3" type="taxon_list">
<paragraph id="8BFC8368FFE3EC15FF27EF92FDC9FA02" blockId="2.[189,526,1507,1533]" box="[189,526,1507,1533]" pageId="2" pageNumber="3">
<heading id="D0B43404FFE3EC15FF27EF92FDC9FA02" bold="true" box="[189,526,1507,1533]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B9375F7AFFE3EC15FF27EF92FDC9FA02" bold="true" box="[189,526,1507,1533]" pageId="2" pageNumber="3">Originally included species</emphasis>
</heading>
</paragraph>
<paragraph id="8BFC8368FFE3EC15FF27EC62FD52F9D3" blockId="2.[189,661,1554,1580]" box="[189,661,1554,1580]" pageId="2" pageNumber="3">
<taxonomicName id="4C43F8EBFFE3EC15FF27EC62FD56F9D3" authority="Johnson, 1898" authorityName="Johnson" authorityYear="1898" box="[189,657,1554,1580]" class="Insecta" family="Syrphidae" genus="Megatrigon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="sexfasciatus">
<emphasis id="B9375F7AFFE3EC15FF27EC62FE1AF9D3" box="[189,477,1554,1580]" italics="true" pageId="2" pageNumber="3">Megatrigon sexfasciatus</emphasis>
<bibRefCitation id="EFD2FE99FFE3EC15FE7EEC63FD56F9D3" author="Johnson C. W." box="[484,657,1554,1580]" pageId="2" pageNumber="3" pagination="157 - 164" refId="ref17020" refString="Johnson C. W. 1898. Diptera collected by Dr. A. Donaldson Smith in Somaliland, eastern Africa. Proceedings of the Academy of Natural Sciences Philadelphia 50: 157 - 164. Available from http: // biodiversitylibrary. org / page / 26368835 [accessed 6 Sep. 2016]" type="journal article" year="1898">Johnson, 1898</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C359D0E3FFE3EC15FF27EC25FA8BF963" pageId="2" pageNumber="3" type="type_taxon">
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<heading id="D0B43404FFE3EC15FF27EC25FE9FF992" bold="true" box="[189,344,1619,1646]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B9375F7AFFE3EC15FF27EC25FE9FF992" bold="true" box="[189,344,1619,1646]" pageId="2" pageNumber="3">
<typeStatus id="54F83DCAFFE3EC15FF27EC25FF3DF991" box="[189,250,1620,1646]" pageId="2" pageNumber="3">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFC8368FFE3EC15FF27ECF2FA8BF963" blockId="2.[189,1356,1666,1693]" box="[189,1356,1666,1693]" pageId="2" pageNumber="3">
<taxonomicName id="4C43F8EBFFE3EC15FF27ECF2FD49F963" authority="Johnson, 1898" authorityName="Johnson" authorityYear="1898" box="[189,654,1666,1692]" class="Insecta" family="Syrphidae" genus="Megatrigon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="sexfasciatus">
<emphasis id="B9375F7AFFE3EC15FF27ECF2FE1AF963" box="[189,477,1666,1692]" italics="true" pageId="2" pageNumber="3">Megatrigon sexfasciatus</emphasis>
<bibRefCitation id="EFD2FE99FFE3EC15FE7EECF3FD49F963" author="Johnson C. W." box="[484,654,1666,1692]" pageId="2" pageNumber="3" pagination="157 - 164" refId="ref17020" refString="Johnson C. W. 1898. Diptera collected by Dr. A. Donaldson Smith in Somaliland, eastern Africa. Proceedings of the Academy of Natural Sciences Philadelphia 50: 157 - 164. Available from http: // biodiversitylibrary. org / page / 26368835 [accessed 6 Sep. 2016]" type="journal article" year="1898">Johnson, 1898</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54F83DCAFFE3EC15FD07ECF3FCCFF963" box="[669,776,1666,1692]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
,
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(
<collectionCode id="ED521BADFFE3EC15FCA3ECF3FC50F963" box="[825,919,1666,1692]" pageId="2" pageNumber="3">BMNH</collectionCode>
), examined), by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C359D0E3FFE3EC15FF27ECB5FBD7F842" pageId="2" pageNumber="3" type="discussion">
<paragraph id="8BFC8368FFE3EC15FF27ECB5FEE8F921" blockId="2.[189,303,1732,1758]" box="[189,303,1732,1758]" pageId="2" pageNumber="3">
<heading id="D0B43404FFE3EC15FF27ECB5FEE8F921" bold="true" box="[189,303,1732,1758]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B9375F7AFFE3EC15FF27ECB5FEE8F921" bold="true" box="[189,303,1732,1758]" pageId="2" pageNumber="3">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BFC8368FFE3EC15FF27EC83FBD7F842" blockId="2.[189,1399,1778,1981]" pageId="2" pageNumber="3">
The
<typeStatus id="54F83DCAFFE3EC15FF6BEC83FE9FF8F3" box="[241,344,1778,1804]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
is without the posteroventral patch of setae on katepisternum. This is quite unusual among
<taxonomicName id="4C43F8EBFFE3EC15FF27ED67FE8FF8D0" authorityName="Johnson" authorityYear="1898" box="[189,328,1814,1839]" class="Insecta" family="Syrphidae" genus="Megatrigon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B9375F7AFFE3EC15FF27ED67FE8FF8D0" box="[189,328,1814,1839]" italics="true" pageId="2" pageNumber="3">Megatrigon</emphasis>
</taxonomicName>
and all other
<taxonomicName id="4C43F8EBFFE3EC15FE7FED64FDBFF8D0" authorityName="Edwards" authorityYear="1915" box="[485,632,1813,1839]" class="Insecta" family="Syrphidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="tribe" tribe="Merodontini">Merodontini</taxonomicName>
(currently only known from
<taxonomicName id="4C43F8EBFFE3EC15FC52ED64FEF9F8AD" authority="Doczkal &amp; Pape, 2009" authorityName="Doczkal &amp; Pape" authorityYear="2009" class="Insecta" family="Syrphidae" genus="Lyneborgimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="magnifica">
<emphasis id="B9375F7AFFE3EC15FC52ED64FB35F8D0" box="[968,1266,1813,1839]" italics="true" pageId="2" pageNumber="3">Lyneborgimyia magnifica</emphasis>
<bibRefCitation id="EFD2FE99FFE3EC15FB62ED64FEF9F8AD" author="Doczkal D. &amp; Pape T." pageId="2" pageNumber="3" pagination="559 - 573" refId="ref16903" refString="Doczkal D. &amp; Pape T. 2009. Lyneborgimyia magnifica gen. et sp. n. (Diptera: Syrphidae) from Tanzania, with a phylogenetic analysis of the Eumerini using new morphological characters. Systematic Entomology 34: 559 - 573. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.2009.00478. x" type="journal article" year="2009">Doczkal &amp; Pape, 2009</bibRefCitation>
</taxonomicName>
) and probably this is an individual aberration (the number of posteroventral katepisternal setae is very low usually less than 10 in most
<taxonomicName id="4C43F8EBFFE3EC15FD23ED2DFC83F88A" authorityName="Johnson" authorityYear="1898" box="[697,836,1884,1909]" class="Insecta" family="Syrphidae" genus="Megatrigon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B9375F7AFFE3EC15FD23ED2DFC83F88A" box="[697,836,1884,1909]" italics="true" pageId="2" pageNumber="3">Megatrigon</emphasis>
</taxonomicName>
, lower than usually seen in the
<taxonomicName id="4C43F8EBFFE3EC15FB29ED2AFA8CF889" authorityName="Edwards" authorityYear="1915" box="[1203,1355,1883,1910]" class="Insecta" family="Syrphidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="tribe" tribe="Merodontini">Merodontini</taxonomicName>
). In all other characters this specimen fits the diagnosis of
<taxonomicName id="4C43F8EBFFE3EC15FCB5EDF1FC7DF866" authorityName="Johnson" authorityYear="1898" box="[815,954,1920,1945]" class="Insecta" family="Syrphidae" genus="Megatrigon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B9375F7AFFE3EC15FCB5EDF1FC7DF866" box="[815,954,1920,1945]" italics="true" pageId="2" pageNumber="3">Megatrigon</emphasis>
</taxonomicName>
given below. The male of
<taxonomicName id="4C43F8EBFFE3EC15FB76EDF1FE8CF843" authorityName="Johnson" authorityYear="1898" class="Insecta" family="Syrphidae" genus="Megatrigon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="sexfasciatus">
<emphasis id="B9375F7AFFE3EC15FB76EDF1FE8CF843" italics="true" pageId="2" pageNumber="3">Megatrigon sexfasciatus</emphasis>
</taxonomicName>
is unknown (probably not present in the available material).
</paragraph>
</subSubSection>
<subSubSection id="C359D0E3FFE2EC14FF27EB7AFBE6FD4B" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BFC8368FFE2EC14FF27EB7AFEF1FEDA" blockId="3.[189,310,267,293]" box="[189,310,267,293]" pageId="3" pageNumber="4">
<heading id="D0B43404FFE2EC14FF27EB7AFEF1FEDA" bold="true" box="[189,310,267,293]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B9375F7AFFE2EC14FF27EB7AFEF1FEDA" bold="true" box="[189,310,267,293]" pageId="3" pageNumber="4">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BFC8368FFE2EC14FF27EB48FF2EFE65" blockId="3.[189,1399,313,692]" pageId="3" pageNumber="4">
With all autapomorphies of the
<taxonomicName id="4C43F8EBFFE2EC14FDBCEB48FD7EFEAC" authorityName="Edwards" authorityYear="1915" box="[550,697,313,339]" class="Insecta" family="Syrphidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" tribe="Merodontini">Merodontini</taxonomicName>
(
<bibRefCitation id="EFD2FE99FFE2EC14FD52EB48FC00FEAB" author="Doczkal D. &amp; Pape T." box="[712,967,313,340]" pageId="3" pageNumber="4" pagination="559 - 573" refId="ref16903" refString="Doczkal D. &amp; Pape T. 2009. Lyneborgimyia magnifica gen. et sp. n. (Diptera: Syrphidae) from Tanzania, with a phylogenetic analysis of the Eumerini using new morphological characters. Systematic Entomology 34: 559 - 573. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.2009.00478. x" type="journal article" year="2009">Doczkal &amp; Pape 2009</bibRefCitation>
; as
<taxonomicName id="4C43F8EBFFE2EC14FC75EB48FB99FEAB" authorityName="Smirnov" authorityYear="1924" box="[1007,1118,313,340]" class="Insecta" family="Syrphidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" tribe="Eumerini">Eumerini</taxonomicName>
) except for the colour of the pterostigma, which is clear in most species. Unique characters (autapomorphies) within
<taxonomicName id="4C43F8EBFFE2EC14FB7EEB2CFAB0FE88" authorityName="Edwards" authorityYear="1915" box="[1252,1399,349,375]" class="Insecta" family="Syrphidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" tribe="Merodontini">Merodontini</taxonomicName>
are:
</paragraph>
<paragraph id="8BFC8368FFE2EC14FF27EBD2FDF8FE42" blockId="3.[189,1399,313,692]" box="[189,575,419,446]" pageId="3" pageNumber="4">1) lower calypter setose dorsally,</paragraph>
<paragraph id="8BFC8368FFE2EC14FF27EBB7FB80FE1F" blockId="3.[189,1399,313,692]" box="[189,1095,454,481]" pageId="3" pageNumber="4">
2) metafemur posterodorsally on proximal half obliquely striate (
<figureCitation id="13789FEDFFE2EC14FC22EBB7FC39FE1F" box="[952,1022,454,480]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1399,1586,1896]" captionTargetId="figure@9.[189,1399,1586,1898]" captionTargetPageId="9" captionText="Fig. 6. Megatrigon spp., metafemur, ♂ (SEM). A. Megatrigon argentifrons sp. nov., proximal half posterodorsally with striae (arrow). B. M. argentifrons sp. nov., enlarged striae. C. M. magnicornis sp. nov., anteroventral preapical flange." figureDoi="http://doi.org/10.5281/zenodo.3849898" httpUri="https://zenodo.org/record/3849898/files/figure.png" pageId="3" pageNumber="4">Fig. 6</figureCitation>
AB),
</paragraph>
<paragraph id="8BFC8368FFE2EC14FF27EB9BFDECFDD7" blockId="3.[189,1399,313,692]" pageId="3" pageNumber="4">3) stem vein R without setae (if exceptionally one or a few setae are present then they are no longer than 0.5 × the width of the vein),</paragraph>
<paragraph id="8BFC8368FFE2EC14FF27E841FC54FDB5" blockId="3.[189,1399,313,692]" box="[189,915,560,587]" pageId="3" pageNumber="4">4) tergite 1 laterally (lateral to the apex of metepimeron) bare,</paragraph>
<paragraph id="8BFC8368FFE2EC14FF27E822FC21FD91" blockId="3.[189,1399,313,692]" box="[189,998,595,622]" pageId="3" pageNumber="4">
5) tergite 2 anterolaterally with a large triangular bare area (
<figureCitation id="13789FEDFFE2EC14FCE1E822FC12FD92" box="[891,981,595,621]" captionStart="Fig" captionStartId="13.[189,232,1964,1990]" captionTargetBox="[386,1204,1272,1924]" captionTargetId="figure@13.[386,1204,1272,1924]" captionTargetPageId="13" captionText="Fig. 9. Megatrigon argenteus (Walker, 1852) comb. nov., tergites, ♂ (SEM). A. Tergite 2, anterior corner. B. Tergite 4, setae. C. Tergite 3, silver spot. D. Tergite 3, enlarged microtrichia of silver spot." figureDoi="http://doi.org/10.5281/zenodo.3849910" httpUri="https://zenodo.org/record/3849910/files/figure.png" pageId="3" pageNumber="4">Fig. 9A</figureCitation>
),
</paragraph>
<paragraph id="8BFC8368FFE2EC14FF27E807FCCBFD6E" blockId="3.[189,1399,313,692]" box="[189,780,630,657]" pageId="3" pageNumber="4">
6) hamus characteristically S-shaped (
<figureCitation id="13789FEDFFE2EC14FDE6E807FD23FD6E" box="[636,740,630,657]" captionStart="Fig" captionStartId="17.[189,232,1758,1784]" captionTargetBox="[189,1399,340,1716]" captionTargetId="figure@17.[189,1399,340,1716]" captionTargetPageId="17" captionText="Fig. 11. Megatrigon spp., ♂ terminalia, hypandrium (AM) and phallus (N), left lateral view. A. Megatrigon argenteus (Walker, 1852) comb. nov. B. M. argentifrons sp. nov. C. M. argentimaculatus sp. nov. D. M. magnicornis sp. nov. E. M. natalensis sp. nov. F. M. sexmaculatus sp. nov. G. M. apiformis sp. nov. H. M. cooksoni sp. nov. I. M. immaculatus sp. nov. J. M. ochreatus (Hull, 1964) comb. nov. K. M. tabanoides sp. nov. L. M. sp. from sexfasciatus group. MN. M. nivalis (Hull, 1964) comb. nov. Symbol: x = hamus." figureDoi="http://doi.org/10.5281/zenodo.3849918" httpUri="https://zenodo.org/record/3849918/files/figure.png" pageId="3" pageNumber="4">Fig. 11N</figureCitation>
:x),
</paragraph>
<paragraph id="8BFC8368FFE2EC14FF27E8EBFBE6FD4B" blockId="3.[189,1399,313,692]" box="[189,1057,665,692]" pageId="3" pageNumber="4">
7) subepandrial sclerite with long microtrichia (
<figureCitation id="13789FEDFFE2EC14FD76E8E8FC9DFD4B" box="[748,858,665,692]" captionStart="Fig" captionStartId="16.[189,232,1825,1851]" captionTargetBox="[216,1374,263,1796]" captionTargetId="figure@16.[216,1374,263,1796]" captionTargetPageId="16" captionText="Fig. 10. Megatrigon spp., ♂ terminalia, epandrium, left lateral view. A. Megatrigon argenteus (Walker, 1852) comb. nov. B. M. argentifrons sp. nov. C. M. argentimaculatus sp. nov. D. M. magnicornis sp. nov. E. M. natalensis sp. nov. F. M. sexmaculatus sp. nov. G. M. apiformis sp. nov. H. M. cooksoni sp. nov. I. M. immaculatus sp. nov. J. M. ochreatus (Hull, 1964) comb. nov. K. M. tabanoides sp. nov. L. Megatrigon sp. from sexfasciatus group. M. M. nivalis. Symbols: x = ventral margin of median surstyle; y = posterior surstyle; z = cercus; w = subepandrial sclerite." figureDoi="http://doi.org/10.5281/zenodo.3849914" httpUri="https://zenodo.org/record/3849914/files/figure.png" pageId="3" pageNumber="4">Figs 10B</figureCitation>
:w, 13CD, G:z).
</paragraph>
</subSubSection>
<subSubSection id="C359D0E3FFE2EC13FF27E8AAFB5BFE93" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
<paragraph id="8BFC8368FFE2EC14FF27E8AAFE89FD0A" blockId="3.[189,334,731,757]" box="[189,334,731,757]" pageId="3" pageNumber="4">
<heading id="D0B43404FFE2EC14FF27E8AAFE89FD0A" bold="true" box="[189,334,731,757]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B9375F7AFFE2EC14FF27E8AAFE89FD0A" bold="true" box="[189,334,731,757]" pageId="3" pageNumber="4">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BFC8368FFE2EC14FF27E97BFE0EFC08" blockId="3.[189,1399,777,1015]" pageId="3" pageNumber="4">
HEAD. Lateral corners of mouth edge well developed, anterior edge of mouth slightly produced (except for
<taxonomicName id="4C43F8EBFFE2EC14FF7EE95CFD75FCB8" authority="(Herve-Bazin, 1913)" authorityName="Doczkal &amp; Radenković &amp; Lyneborg &amp; Pape" authorityYear="2016" baseAuthorityName="Herve-Bazin" baseAuthorityYear="1913" box="[228,690,812,839]" class="Insecta" family="Syrphidae" genus="Megatrigon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="jacobi" status="comb. nov.">
<emphasis id="B9375F7AFFE2EC14FF7EE95CFE7AFCB8" box="[228,445,813,839]" italics="true" pageId="3" pageNumber="4">Megatrigon jacobi</emphasis>
(Hervé-Bazin, 1913)
</taxonomicName>
<taxonomicNameLabel id="A204E201FFE2EC14FD22E95DFCFDFCB8" box="[696,826,812,839]" pageId="3" pageNumber="4" rank="species">comb. nov.</taxonomicNameLabel>
), dorsal part of face more or less distinctly convex below antenna, median part of lunula extended posteriorly (similar to
<taxonomicName id="4C43F8EBFFE2EC14FC73E921FB94FC95" authorityName="Meigen" authorityYear="1803" box="[1001,1107,848,874]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9375F7AFFE2EC14FC73E921FB94FC95" box="[1001,1107,848,874]" italics="true" pageId="3" pageNumber="4">Merodon</emphasis>
</taxonomicName>
), both dorsal and ventral part of post-ocular orbit wide, post-ocular ridge present, postcranial carina absent, dichoptic (
<figureCitation id="13789FEDFFE2EC14FB69E902FAFEFC72" box="[1267,1337,883,909]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[189,1399,518,1778]" captionTargetId="figure@6.[189,1399,518,1778]" captionTargetPageId="6" captionText="Fig. 2. Megatrigon spp., head, dorsal view. A. Megatrigon argenteus (Walker, 1852) comb. nov. (♂). B. M. argentifrons sp. nov. (♂). C. M. argentimaculatus sp. nov. (♂). D. M. magnicornis sp. nov. (♂). EF. M.natalensissp. nov.(arrow =postocellarspot)(♂).G.M.sexmaculatus sp. nov.(♂).H. M.apiformis sp. nov. (♂). I. M. cooksoni sp. nov. (♂). J. M. immaculatus sp. nov. (♂). K. M. ochreatus (Hull, 1964) comb. nov. (♂). L. M. tabanoides sp. nov. (♂). M. M. jacobi (Hervé-Bazin, 1913) comb. nov. (♂). N. M. nivalis (Hull, 1964) comb. nov. (♀). Symbol: x = postocellar spot." figureDoi="http://doi.org/10.5281/zenodo.3849882" httpUri="https://zenodo.org/record/3849882/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
), eye setose, without enlarged ommatidia in ♂. Fossette ill-defined, in a lateral position, postpedicel without longitudinal furrows, arista simple. Clypeus slender, postclypeus well sclerotized, labellum elongated (as in many
<taxonomicName id="4C43F8EBFFE2EC14FED7E9ACFE70FC08" authorityName="Meigen" authorityYear="1803" box="[333,439,989,1015]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9375F7AFFE2EC14FED7E9ACFE70FC08" box="[333,439,989,1015]" italics="true" pageId="3" pageNumber="4">Merodon</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BFC8368FFE2EC14FF27EE6EFD9AFB17" blockId="3.[189,1399,1054,1257]" pageId="3" pageNumber="4">
THORAX. Notopleural sulcus absent, notal wing process of
<taxonomicName id="4C43F8EBFFE2EC14FC38EE6EFBCDFBC7" authorityName="Meigen" authorityYear="1822" box="[930,1034,1055,1080]" class="Insecta" family="Syrphidae" genus="Eumerus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9375F7AFFE2EC14FC38EE6EFBCDFBC7" box="[930,1034,1055,1080]" italics="true" pageId="3" pageNumber="4">Eumerus</emphasis>
</taxonomicName>
-
<typeStatus id="54F83DCAFFE2EC14FB88EE6EFB8EFBC6" box="[1042,1097,1055,1081]" pageId="3" pageNumber="4">type</typeStatus>
, supra-alar bristles well developed, their alveoli enlarged, scutal alveoli with raised margins, post-alar callus nearly entirely setose, scutellum&gt; 2 × as wide as long, its marginal rim yellow, scutellum with concave ventral surface, without ventral fringe, probasisternum large, approaching proepisternum, proepimeron moderately extended ventrally [as in
<taxonomicName id="4C43F8EBFFE2EC14FE7DEEDDFC40FB3A" authority="(Fabricius, 1798)" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[487,903,1194,1221]" class="Insecta" family="Syrphidae" genus="Eumerus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="tricolor">
<emphasis id="B9375F7AFFE2EC14FE7DEEDDFD76FB3A" box="[487,689,1195,1221]" italics="true" pageId="3" pageNumber="4">Eumerus tricolor</emphasis>
(Fabricius, 1798)
</taxonomicName>
], katepimeron almost flat, katatergite with a weak carina, metepisternum bare.
</paragraph>
<paragraph id="8BFC8368FFE2EC14FF27EF61FBCDFA02" blockId="3.[189,1399,1295,1533]" pageId="3" pageNumber="4">
WING (
<figureCitation id="13789FEDFFE2EC14FE96EF7EFE96FAD5" box="[268,337,1295,1322]" captionStart="Fig" captionStartId="9.[189,232,1494,1520]" captionTargetBox="[189,1399,610,1456]" captionTargetId="figure@9.[189,1399,609,1457]" captionTargetPageId="9" captionText="Fig. 5. Megatrigon spp., right wing, ♂. A. Megatrigon apiformis sp. nov. B. M. cooksoni sp. nov. C. M. tabanoides sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3849894" httpUri="https://zenodo.org/record/3849894/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
). Cross-vein sc-r well developed, C ending far before apex of wing, R
<subScript id="17C7812DFFE2EC14FBE9EF50FB49FAD0" attach="left" box="[1139,1166,1313,1327]" fontSize="6" pageId="3" pageNumber="4">4+5</subScript>
moderately sinuous (as in
<taxonomicName id="4C43F8EBFFE2EC14FE93EF42FE13FAB2" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[265,468,1331,1357]" class="Insecta" family="Syrphidae" genus="Eumerus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="tricolor">
<emphasis id="B9375F7AFFE2EC14FE93EF42FE13FAB2" box="[265,468,1331,1357]" italics="true" pageId="3" pageNumber="4">Eumerus tricolor</emphasis>
</taxonomicName>
), M
<subScript id="17C7812DFFE2EC14FD92EF34FDD6FAAC" attach="left" box="[520,529,1349,1363]" fontSize="6" pageId="3" pageNumber="4">1</subScript>
without stump veins, dm-cu almost perpendicular to CuA
<subScript id="17C7812DFFE2EC14FB51EF34FB13FAAC" attach="left" box="[1227,1236,1349,1363]" fontSize="6" pageId="3" pageNumber="4">1</subScript>
, usually with stump vein, CuA
<subScript id="17C7812DFFE2EC14FE1DEF19FE57FA89" attach="left" box="[391,400,1384,1398]" fontSize="6" pageId="3" pageNumber="4">1</subScript>
slightly more than 2 x as long as dm-cu, bm-cu about as long as basal part of CuA
<subScript id="17C7812DFFE2EC14FAF4EF19FAB0FA89" attach="right" box="[1390,1399,1384,1398]" fontSize="6" pageId="3" pageNumber="4">1</subScript>
or slightly longer, CuA about 3 x as long as CuA
<subScript id="17C7812DFFE2EC14FC94EFFAFCD0FA66" attach="right" box="[782,791,1419,1433]" fontSize="6" pageId="3" pageNumber="4">2</subScript>
, A
<subScript id="17C7812DFFE2EC14FCA1EFFAFC83FA66" attach="right" box="[827,836,1419,1433]" fontSize="6" pageId="3" pageNumber="4">1</subScript>
+CuA
<subScript id="17C7812DFFE2EC14FC11EFFAFC53FA66" attach="left" box="[907,916,1419,1433]" fontSize="6" pageId="3" pageNumber="4">2</subScript>
smoothly curved, Cup short and ending at about half length of CuA, wing broad, about 2.35 × as long as wide, wing margin with undulated membrane, costagium with long setae at posterior margin (except in
<taxonomicName id="4C43F8EBFFE2EC14FB94EFB1FB35FA26" authorityName="Doczkal &amp; Radenković &amp; Lyneborg &amp; Pape" authorityYear="2016" baseAuthorityName="Hull" baseAuthorityYear="1964" box="[1038,1266,1471,1497]" class="Insecta" family="Syrphidae" genus="Megatrigon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="nivalis">
<emphasis id="B9375F7AFFE2EC14FB94EFB1FB35FA26" box="[1038,1266,1471,1497]" italics="true" pageId="3" pageNumber="4">Megatrigon nivalis</emphasis>
</taxonomicName>
), marginal fringe of upper calypter glued (cf. fig. 32IK in
<bibRefCitation id="EFD2FE99FFE2EC14FC9BEF92FC3AFA02" author="Hippa H. &amp; Stahls G." box="[769,1021,1506,1533]" pageId="3" pageNumber="4" pagination="1 - 72" refId="ref16967" refString="Hippa H. &amp; Stahls G. 2005. Morphological characters of adult Syrphidae: descriptions and phylogenetic utility. Acta Zoologica Fennica 215: 1 - 72." type="journal article" year="2005">Hippa &amp; Ståhls 2005</bibRefCitation>
).
</paragraph>
<paragraph id="8BFC8368FFE2EC14FF27EC54FDF0F8CB" blockId="3.[189,1399,1572,1845]" pageId="3" pageNumber="4">
LEGS. Procoxa short and stout, mesocoxa posteriorly usually bare (except for
<taxonomicName id="4C43F8EBFFE2EC14FBFEEC54FB11F9C1" authorityName="Doczkal &amp; Radenković &amp; Lyneborg &amp; Pape" authorityYear="2016" baseAuthorityName="Herve-Bazin" baseAuthorityYear="1913" box="[1124,1238,1572,1598]" class="Insecta" family="Syrphidae" genus="Megatrigon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="jacobi" status="comb. nov.">
<emphasis id="B9375F7AFFE2EC14FBFEEC54FB11F9C1" box="[1124,1238,1572,1598]" italics="true" pageId="3" pageNumber="4">M. jacobi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204E201FFE2EC14FB7BEC55FAAEF9C0" box="[1249,1385,1572,1599]" pageId="3" pageNumber="4" rank="species">comb. nov.</taxonomicNameLabel>
); metafemur shiny, thickened and curved dorsally, about 2.5 × as long as deep, light yellow setae on metafemur exceed relatively large apical flange, usually with shiny, bare, longitudinal area ventrally; ventral face of metafemur slightly convex or straight, with a low anteroventral preapical flange bearing weak bristles on strong finger-like outgrowths, and with a few weak bristles posteroventrally preapically (
<figureCitation id="13789FEDFFE2EC14FF5CECA5FEE4F910" box="[198,291,1748,1775]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1399,1586,1896]" captionTargetId="figure@9.[189,1399,1586,1898]" captionTargetPageId="9" captionText="Fig. 6. Megatrigon spp., metafemur, ♂ (SEM). A. Megatrigon argentifrons sp. nov., proximal half posterodorsally with striae (arrow). B. M. argentifrons sp. nov., enlarged striae. C. M. magnicornis sp. nov., anteroventral preapical flange." figureDoi="http://doi.org/10.5281/zenodo.3849898" httpUri="https://zenodo.org/record/3849898/files/figure.png" pageId="3" pageNumber="4">Fig. 6C</figureCitation>
); metatibia with a sharp bare (a few isolated setae may occur) anteroventral ridge in proximal half, protibia with irregularly spaced anterolateral apical setae; protarsus short and stout, tarsomeres 24 as wide as long; claws enlarged.
</paragraph>
<paragraph id="8BFC8368FFE2EC13FF27ED2DFB5BFE93" blockId="3.[189,1399,1884,2016]" lastBlockId="4.[189,1399,267,364]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
ABDOMEN. Tergites strongly curved ventrally at margin; anterior corners of tergite 2 inflated, border between tergites 1 and 2 not crenulated; lateral margins of tergites microtrichiose; ♂ sternite 4 without projections, with incised posterior margin and a fine desclerotised median line, lateral margins simple, sternite 6 bare, sternite 7 with a basal bump. Anterior surstyle lobe absent, posterior surstyle lobe narrow (
<figureCitation id="13789FEDFFE5EC13FF5CEB7AFE92FED9" box="[198,341,267,294]" captionStart="Fig" captionStartId="30.[189,232,1894,1920]" captionTargetBox="[189,1398,875,1852]" captionTargetId="figure@30.[189,1398,875,1852]" captionTargetPageId="30" captionText="Fig. 13. Megatrigon spp., ♂ genitalia, lateral (A, F) and ventral (BE, G) view (SEM). AC. Megatrigon argenteus (Walker, 1852) comb. nov. D. M. magnicornis sp. nov. E. M. jacobi (Hervé-Bazin, 1913) comb. nov. FG. Megatrigon sp. from the sexfasciatus group. Symbols: w = median surstyle; x = inner accessory lobe of median surstyle; y = posterior surstyle; z = subepandrial sclerite." figureDoi="http://doi.org/10.5281/zenodo.3849926" httpUri="https://zenodo.org/record/3849926/files/figure.png" pageId="4" pageNumber="5">Fig. 13A, E</figureCitation>
:y), subepandrial sclerite microtrichiose (
<figureCitation id="13789FEDFFE5EC13FCD8EB7AFC52FEDA" box="[834,917,267,293]" captionStart="Fig" captionStartId="30.[189,232,1894,1920]" captionTargetBox="[189,1398,875,1852]" captionTargetId="figure@30.[189,1398,875,1852]" captionTargetPageId="30" captionText="Fig. 13. Megatrigon spp., ♂ genitalia, lateral (A, F) and ventral (BE, G) view (SEM). AC. Megatrigon argenteus (Walker, 1852) comb. nov. D. M. magnicornis sp. nov. E. M. jacobi (Hervé-Bazin, 1913) comb. nov. FG. Megatrigon sp. from the sexfasciatus group. Symbols: w = median surstyle; x = inner accessory lobe of median surstyle; y = posterior surstyle; z = subepandrial sclerite." figureDoi="http://doi.org/10.5281/zenodo.3849926" httpUri="https://zenodo.org/record/3849926/files/figure.png" pageId="4" pageNumber="5">Fig. 13</figureCitation>
CD, G:z). Setae on various part of the body serrated (
<figureCitation id="13789FEDFFE5EC13FEF0EB5FFE02FEB6" box="[362,453,302,329]" captionStart="Fig" captionStartId="13.[189,232,1964,1990]" captionTargetBox="[386,1204,1272,1924]" captionTargetId="figure@13.[386,1204,1272,1924]" captionTargetPageId="13" captionText="Fig. 9. Megatrigon argenteus (Walker, 1852) comb. nov., tergites, ♂ (SEM). A. Tergite 2, anterior corner. B. Tergite 4, setae. C. Tergite 3, silver spot. D. Tergite 3, enlarged microtrichia of silver spot." figureDoi="http://doi.org/10.5281/zenodo.3849910" httpUri="https://zenodo.org/record/3849910/files/figure.png" pageId="4" pageNumber="5">Fig. 9B</figureCitation>
), except in
<taxonomicName id="4C43F8EBFFE5EC13FDD1EB5EFD07FEB6" authorityName="Doczkal &amp; Radenković &amp; Lyneborg &amp; Pape" authorityYear="2016" baseAuthorityName="Hull" baseAuthorityYear="1964" box="[587,704,303,329]" class="Insecta" family="Syrphidae" genus="Megatrigon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="nivalis" status="comb. nov.">
<emphasis id="B9375F7AFFE5EC13FDD1EB5EFD07FEB6" box="[587,704,303,329]" italics="true" pageId="4" pageNumber="5">M. nivalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204E201FFE5EC13FD5DEB5FFC8FFEB6" box="[711,840,302,329]" pageId="4" pageNumber="5" rank="species">comb. nov.</taxonomicNameLabel>
The included species can be arranged into three well defined species groups, each defined by several autapomorphic character states.
</paragraph>
</subSubSection>
</treatment>
</document>