treatments-xml/data/03/D3/A7/03D3A756FFC84F5A58CA9E0DB0BF43EF.xml

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<mods:title id="D84D63DA277987637B0AAB74D7C91F07">A new trasandean species of the genus Tigridania Kaye, 1918 (Lepidoptera: Erebidae: Arctiinae: Arctiini: Euchromiina) from Colombia</mods:title>
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<mods:namePart id="A11B3FDD076264C6A963704113477B75">Alzate-Cano, Julian David</mods:namePart>
<mods:affiliation id="5BF65C3AF3C2971E6DA27E6A7F121623">Grupo de Entomología Universidad de Antioquia (GEUA), Universidad de Antioquia, Medellín, Colombia.</mods:affiliation>
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<mods:namePart id="17B45D53125F4DF4B802867DFB3C6A36">Wolff, Marta</mods:namePart>
<mods:affiliation id="62FA6AA6DF19F74F9A01593ABE2C5947">Grupo de Entomología Universidad de Antioquia (GEUA), Universidad de Antioquia, Medellín, Colombia.</mods:affiliation>
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<mods:namePart id="196E9FAA8B0DE7D6CED58E578E10EF49">Laguerre, Michel</mods:namePart>
<mods:affiliation id="00FF4682D3A575D543BA50EF59AF8A5F">31, rue de la Haute-Lande, 33850, Léognan, France.</mods:affiliation>
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<mods:date id="92FECE18156DF96FCBA2677A2F53F388">2025</mods:date>
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<taxonomicName id="4C7A6DC3FFC84F5858CA9E0DB3CF4220" authority="Alzate-Cano &amp; Laguerre" authorityName="Alzate-Cano &amp; Wolff &amp; Laguerre" authorityYear="2025" box="[151,753,1625,1652]" class="Insecta" family="Erebidae" genus="Tigridania" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="597" phylum="Arthropoda" rank="species" species="magdalenae" status="sp. nov.">
<emphasis id="B90ECA52FFC84F5858CA9E0DB08F4220" bold="true" box="[151,433,1625,1652]" italics="true" pageId="0" pageNumber="597">Tigridania magdalenae</emphasis>
Alzate-Cano &amp; Laguerre
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,
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<paragraph id="8BC51640FFC84F5858CA9ED4B54642CC" blockId="0.[151,1437,1590,2012]" box="[151,1144,1664,1688]" pageId="0" pageNumber="597">
https://zoobank.org/
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urn:lsid:zoobank.org:act:
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(
<figureCitation id="13410AC5FFC84F5858C29EF0B1C242E8" box="[159,252,1700,1724]" captionStart="FIGURE 1" captionStartId="1.[151,250,735,759]" captionTargetBox="[164,1426,237,704]" captionTargetId="figure-46@1.[162,1426,235,704]" captionTargetPageId="1" captionText="FIGURE 1. A. Distribution map. B. Pair of Tigridania magdalenae sp. nov. in copulation. They were observed inside a primary forest. (Photo: Camilo Flórez Valencia)." figureDoi="http://doi.org/10.5281/zenodo.14804489" httpUri="https://zenodo.org/record/14804489/files/figure.png" pageId="0" pageNumber="597">Figs. 1B</figureCitation>
,
<figureCitation id="13410AC5FFC84F5859559EF0B02842E8" box="[264,278,1700,1724]" captionStart="FIGURE 2" captionStartId="2.[151,250,1703,1727]" captionTargetBox="[172,1413,199,1666]" captionTargetId="figure-25@2.[153,1434,180,1678]" captionTargetPageId="2" captionText="FIGURE 2. A. Male holotype dorsal view. B. Male holotype ventral view. C. Female paratype dorsal view D. Female paratype ventral view. E. Male genitalia dorsal view. F. Male genitalia lateral view. G. Male genitalia ventral view. H. Aedeagus lateral view. I. Scent pouches signaled with arrows. J. Female genitalia." figureDoi="http://doi.org/10.5281/zenodo.14804493" httpUri="https://zenodo.org/record/14804493/files/figure.png" pageId="0" pageNumber="597">2</figureCitation>
)
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:
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,
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:
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,
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,
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,
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,
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,
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,
<collectorName id="268F7396FFC84F585DD99E9CB1F74357" pageId="0" pageNumber="597">C. Bota</collectorName>
and
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<emphasis id="B90ECA52FFC84F5859309EB8B0AA4350" box="[365,404,1772,1796]" italics="true" pageId="0" pageNumber="597">leg.</emphasis>
,
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. Very humid premountain forest. Collected while copulating.
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.
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<typeStatus id="54C1A8E2FFC84F5858CA9F44B037437C" box="[151,265,1808,1832]" pageId="0" pageNumber="597" type="paratype">Paratypes</typeStatus>
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.
</emphasis>
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<emphasis id="B90ECA52FFC84F58594B9F44B01D437C" bold="true" box="[278,291,1808,1832]" pageId="0" pageNumber="597">2</emphasis>
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.
<specimenCount id="9D7CDDC9FFC84F5859149F44B04E4373" box="[329,368,1808,1832]" count="1" pageId="0" pageNumber="597" type="female">1 ♀</specimenCount>
, same data as holotype. Collected while copulating
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.
<materialsCitation id="3B121C1DFFC84F585C6C9F44B2AA4318" collectingDate="1986-07" collectingMethod="entomological net" collectionCode="CEUA" collectorName="Olga Lucia Z." country="Colombia" location="Rio Claro" pageId="0" pageNumber="597" specimenCode="CEUA 142766" specimenCount="1" specimenCount-female="1" stateProvince="Antioquia" typeStatus="paratype">
<specimenCount id="9D7CDDC9FFC84F585C6C9F44B5664373" box="[1073,1112,1808,1832]" count="1" pageId="0" pageNumber="597" type="female">1 ♀</specimenCount>
,
<collectingCountry id="F36D56D0FFC84F585C389F44B5C6437C" box="[1125,1272,1808,1832]" name="Colombia" pageId="0" pageNumber="597">COLOMBIA</collectingCountry>
:
<collectingRegion id="49BED8A2FFC84F585CA29F44B452437C" box="[1279,1388,1808,1832]" country="Colombia" name="Antioquia" pageId="0" pageNumber="597">Antioquia</collectingRegion>
,
<location id="8EA5409BFFC84F585D2B9F44B1EB4318" LSID="urn:lsid:plazi:treatment:03D3A756FFC84F5A58CA9E0DB0BF43EF:8EA5409BFFC84F585D2B9F44B1EB4318" country="Colombia" name="Rio Claro" pageId="0" pageNumber="597" stateProvince="Antioquia">Rio Claro</location>
,
<date id="FFC43080FFC84F5858BD9F60B0754318" box="[224,331,1844,1868]" pageId="0" pageNumber="597" value="1986-07">
<collectingDate id="EF80C968FFC84F5858BD9F60B0754318" box="[224,331,1844,1868]" pageId="0" pageNumber="597" value="1986-07">July 1986</collectingDate>
</date>
,
<collectorName id="268F7396FFC84F58590B9F60B0D5431F" box="[342,491,1844,1868]" pageId="0" pageNumber="597">Olga Lucía Z.</collectorName>
<emphasis id="B90ECA52FFC84F5859AC9F60B3214318" box="[497,543,1844,1868]" italics="true" pageId="0" pageNumber="597">leg.,</emphasis>
<collectingMethod id="523B6E57FFC84F585A7B9F60B3D7431F" box="[550,745,1844,1868]" pageId="0" pageNumber="597">entomological net</collectingMethod>
.
<specimenCode id="DBDCBE3BFFC84F585AAE9F60B2AA4318" box="[755,916,1844,1868]" collectionCode="CEUA" pageId="0" pageNumber="597">CEUA 142766</specimenCode>
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.
</paragraph>
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<paragraph id="8BC51640FFC84F59589A9F03B3AD4487" blockId="0.[151,1437,1590,2012]" lastBlockId="1.[151,1436,151,211]" lastPageId="1" lastPageNumber="598" pageId="0" pageNumber="597">
<emphasis id="B90ECA52FFC84F58589A9F03B00B433B" bold="true" box="[199,309,1879,1903]" pageId="0" pageNumber="597">Diagnosis</emphasis>
: This species can be easily distinguished from
<taxonomicName id="4C7A6DC3FFC84F585B6A9F0CB5824324" authority="Kaye, 1918" authorityName="Kaye" authorityYear="1918" box="[823,1212,1880,1904]" class="Insecta" family="Erebidae" genus="Tigridania" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="597" phylum="Arthropoda" rank="species" species="quadricincta">
<emphasis id="B90ECA52FFC84F585B6A9F0CB5064324" box="[823,1080,1880,1904]" italics="true" pageId="0" pageNumber="597">Tigridania quadricincta</emphasis>
<bibRefCitation id="EFEB6BB1FFC84F585C1D9F0CB5824324" author="Kaye, W. J." box="[1088,1212,1880,1904]" pageId="0" pageNumber="597" pagination="225 - 232" refId="ref1897" refString="Kaye, W. J. (1918) Descriptions from the Joicey Collection of new Species of Syntomidae, Nymphalidae, and Hesperidae, and Two Genera of Syntomidae. Annals and Magazine of Natural History, Series 9, 2 (9), 225-232. https://doi.org/10.1080/00222931808562370" type="journal article" year="1918">Kaye, 1918</bibRefCitation>
</taxonomicName>
by its entirely black thorax and the presence of red scales on the last abdominal segments. Additionally, the male genitalia exhibit distinctly different valvae, and the caecum of the penis is noticeably shorter. Externally, it resembles
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<emphasis id="B90ECA52FFC84F585C2B9FF4B47D43EC" box="[1142,1347,1952,1976]" italics="true" pageId="0" pageNumber="597">Isanthrene thyestes</emphasis>
(
<bibRefCitation id="EFEB6BB1FFC84F585D0E9FF4B1EE4388" author="Druce, H." pageId="0" pageNumber="597" pagination="372 - 384" refId="ref1730" refString="Druce, H. (1883) Descriptions of new Species of Zygaenidae and Arctiidae. Proceedings of the Zoological Society of London, 1883, 372-384. https://doi.org/10.1111/j.1469-7998.1883.tb06656.x" type="book chapter" year="1883">Druce, 1883</bibRefCitation>
)
</taxonomicName>
, but can be separated by its larger size and the thorax, which is completely covered in black scales, whereas in
<taxonomicName id="4C7A6DC3FFC84F585DD09F90B4A3438F" box="[1421,1437,1988,2011]" pageId="0" pageNumber="597">
<emphasis id="B90ECA52FFC84F585DD09F90B4A3438F" box="[1421,1437,1988,2011]" italics="true" pageId="0" pageNumber="597">I.</emphasis>
</taxonomicName>
<emphasis id="B90ECA52FFC94F5958CA98C3B1D544FB" box="[151,235,151,175]" italics="true" pageId="1" pageNumber="598">thyestes</emphasis>
the thorax is yellow with some black longitudinal stripes. Furthermore,
<taxonomicName id="4C7A6DC3FFC94F595C5598CCB54A44FB" baseAuthorityName="Druce" baseAuthorityYear="1883" box="[1032,1140,151,175]" class="Insecta" family="Erebidae" genus="Isanthrene" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="thyestes">
<emphasis id="B90ECA52FFC94F595C5598CCB54A44FB" box="[1032,1140,151,175]" italics="true" pageId="1" pageNumber="598">I. thyestes</emphasis>
</taxonomicName>
has red antennae, while in
<taxonomicName id="4C7A6DC3FFC94F5958CA98EFB0AE4487" authorityName="Alzate-Cano &amp; Wolff &amp; Laguerre" authorityYear="2025" box="[151,400,187,211]" class="Insecta" family="Erebidae" genus="Tigridania" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="magdalenae">
<emphasis id="B90ECA52FFC94F5958CA98EFB0AE4487" box="[151,400,187,211]" italics="true" pageId="1" pageNumber="598">Tigridania magdalenae</emphasis>
</taxonomicName>
, they are entirely black.
</paragraph>
</subSubSection>
<caption id="DF0546C8FFC94F5958CA9A8BB30C4748" ID-DOI="http://doi.org/10.5281/zenodo.14804489" ID-Zenodo-Dep="14804489" httpUri="https://zenodo.org/record/14804489/files/figure.png" pageId="1" pageNumber="598" startId="1.[151,250,735,759]" targetBox="[164,1426,237,704]" targetPageId="1" targetType="figure">
<paragraph id="8BC51640FFC94F5958CA9A8BB30C4748" blockId="1.[151,1436,735,796]" pageId="1" pageNumber="598">
<emphasis id="B90ECA52FFC94F5958CA9A8BB00E46A3" bold="true" box="[151,304,735,759]" pageId="1" pageNumber="598">FIGURE 1. A.</emphasis>
Distribution map.
<emphasis id="B90ECA52FFC94F5959B09AB4B33A46AC" bold="true" box="[493,516,736,760]" pageId="1" pageNumber="598">B.</emphasis>
Pair of
<taxonomicName id="4C7A6DC3FFC94F595A0F9A8BB20146A3" authorityName="Alzate-Cano &amp; Wolff &amp; Laguerre" authorityYear="2025" box="[594,831,735,759]" class="Insecta" family="Erebidae" genus="Tigridania" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="magdalenae" status="sp. nov.">
<emphasis id="B90ECA52FFC94F595A0F9A8BB20146A3" box="[594,831,735,759]" italics="true" pageId="1" pageNumber="598">Tigridania magdalenae</emphasis>
</taxonomicName>
<emphasis id="B90ECA52FFC94F595B199AB4B2AD46AC" bold="true" box="[836,915,736,760]" pageId="1" pageNumber="598">
<taxonomicNameLabel id="A23D7729FFC94F595B199AB4B2AD46AC" box="[836,915,736,760]" pageId="1" pageNumber="598" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in copulation. They were observed inside a primary forest. (Photo: Camilo Flórez Valencia).
</paragraph>
</caption>
<subSubSection id="C36045CBFFC94F59589A9B3BB461437B" pageId="1" pageNumber="598" type="description">
<paragraph id="8BC51640FFC94F59589A9B3BB40B47D3" blockId="1.[151,1437,879,1983]" box="[199,1333,879,904]" pageId="1" pageNumber="598">
<emphasis id="B90ECA52FFC94F59589A9B3BB3CB47DC" box="[199,757,879,904]" inLineHeading="true" pageId="1" pageNumber="598" reason="1">
<emphasis id="B90ECA52FFC94F59589A9B3BB30647DC" bold="true" box="[199,568,879,904]" pageId="1" pageNumber="598">
Description of the
<typeStatus id="54C1A8E2FFC94F5959C49B24B0C547DC" box="[409,507,880,904]" pageId="1" pageNumber="598" type="holotype">holotype</typeStatus>
male
</emphasis>
(figure 2 on top).
</emphasis>
Forewing length male =
<quantity id="4C82BBA5FFC94F595C5F9B24B57147D3" box="[1026,1103,880,904]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="598" unit="mm" value="30.0">30 mm</quantity>
, female =
<quantity id="4C82BBA5FFC94F595CE29B24B40F47D3" box="[1215,1329,880,904]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" metricValueMax="3.4" metricValueMin="3.2" pageId="1" pageNumber="598" unit="mm" value="33.0" valueMax="34.0" valueMin="32.0">32-34 mm</quantity>
.
</paragraph>
<paragraph id="8BC51640FFC94F59589A9BC7B0104163" blockId="1.[151,1437,879,1983]" pageId="1" pageNumber="598">
<emphasis id="B90ECA52FFC94F59589A9BC7B03C47FF" bold="true" box="[199,258,915,939]" italics="true" pageId="1" pageNumber="598">Head</emphasis>
: Antennae bipectinate, entirely black. Head entirely black, ocelli present, proboscis brown. Palpi relatively long, reaching the forehead. Second segment straight and longer than 1 or 3. Segment 3 projected forward, at the level of the forehead.
<emphasis id="B90ECA52FFC94F59591A9B8FB0A647A7" bold="true" box="[327,408,987,1011]" italics="true" pageId="1" pageNumber="598">Thorax</emphasis>
: Completely plain black, covered with long black scales.
<emphasis id="B90ECA52FFC94F595C5B9B88B50447A7" bold="true" box="[1030,1082,988,1011]" italics="true" pageId="1" pageNumber="598">Legs</emphasis>
: Coxae inner face white, the rest completely covered with short black scales.
<emphasis id="B90ECA52FFC94F595A2B9BABB3E14043" bold="true" box="[630,735,1023,1047]" italics="true" pageId="1" pageNumber="598">Abdomen</emphasis>
: Tergite I entirely black, tergites II-V black on the apical margin and golden yellow on the rest of each. Tergite VI-X black on the apical margin and carmine on the rest with some yellow scales on the tergite VII.
<emphasis id="B90ECA52FFC94F5959F99C13B327400B" bold="true" box="[420,537,1095,1119]" italics="true" pageId="1" pageNumber="598">Forewings</emphasis>
: Mainly transparent with black veins. Basal region of the wing dotted with whitishyellowish scales, forming spots on Sc + M and A1. Antemedial region slightly translucent, with a smoky blue hue. Wing apex with a triangular black spot covering about
<quantity id="4C82BBA5FFC94F595AC29CDBB3DE40F3" box="[671,736,1167,1191]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="598" unit="mm" value="5.0">5 mm</quantity>
. Discal cell slightly opaque between veins C and Sc. Basal region dotted with cream-colored scales. It has a spurious vein that divides the disc cell and projects to the apical spot of the wing, running between veins M1 and M2.
<emphasis id="B90ECA52FFC94F595A099C83B3EC40BB" bold="true" box="[596,722,1239,1263]" italics="true" pageId="1" pageNumber="598">Hind wings</emphasis>
: Mainly transparent with black veins. First third of the costal region cream colored, the rest of the edges completely black. Black spot in the region of the tornus and black anal area. Fimbriae grayish black.
</paragraph>
<paragraph id="8BC51640FFC94F59589A9D10B37E41B8" blockId="1.[151,1437,879,1983]" pageId="1" pageNumber="598">
<emphasis id="B90ECA52FFC94F59589A9D10B053410F" bold="true" box="[199,365,1347,1372]" pageId="1" pageNumber="598">Male genitalia</emphasis>
(figure 2E to H): genitalia symmetrical. Uncus long and narrow, bent downward with a slightly bifid extremity and a short pointed tip. On the tegumen, above the uncus and on each side, a pointed triangular process projecting outside. Valvae outreaching the uncus tip, large at base, then narrowed before a subrectangular last third with a straight border. Vinculum evenly bent with a very small saccus. Aedeagus short, cylindrical and straight with a moderate caecum penis. Vesica smooth (
<figureCitation id="13410AC5FFC94F5959839D87B30A41B8" box="[478,564,1491,1516]" captionStart="FIGURE 2" captionStartId="2.[151,250,1703,1727]" captionTargetBox="[172,1413,199,1666]" captionTargetId="figure-25@2.[153,1434,180,1678]" captionTargetPageId="2" captionText="FIGURE 2. A. Male holotype dorsal view. B. Male holotype ventral view. C. Female paratype dorsal view D. Female paratype ventral view. E. Male genitalia dorsal view. F. Male genitalia lateral view. G. Male genitalia ventral view. H. Aedeagus lateral view. I. Scent pouches signaled with arrows. J. Female genitalia." figureDoi="http://doi.org/10.5281/zenodo.14804493" httpUri="https://zenodo.org/record/14804493/files/figure.png" pageId="1" pageNumber="598">Fig. 2H</figureCitation>
).
</paragraph>
<paragraph id="8BC51640FFC94F59589A9DACB4004203" blockId="1.[151,1437,879,1983]" pageId="1" pageNumber="598">
On the abdomen underside it is possible to see the secondary sexual characters as an elongated and ovoid pouch filled with flocculent material (
<figureCitation id="13410AC5FFC94F5959F79E4FB0C94260" box="[426,503,1563,1588]" captionStart="FIGURE 2" captionStartId="2.[151,250,1703,1727]" captionTargetBox="[172,1413,199,1666]" captionTargetId="figure-25@2.[153,1434,180,1678]" captionTargetPageId="2" captionText="FIGURE 2. A. Male holotype dorsal view. B. Male holotype ventral view. C. Female paratype dorsal view D. Female paratype ventral view. E. Male genitalia dorsal view. F. Male genitalia lateral view. G. Male genitalia ventral view. H. Aedeagus lateral view. I. Scent pouches signaled with arrows. J. Female genitalia." figureDoi="http://doi.org/10.5281/zenodo.14804493" httpUri="https://zenodo.org/record/14804493/files/figure.png" pageId="1" pageNumber="598">Fig. 2I</figureCitation>
).
<bibRefCitation id="EFEB6BB1FFC94F595A519E4FB2044260" author="Grados, J. &amp; Ramirez, J. J." box="[524,826,1563,1588]" pageId="1" pageNumber="598" refId="ref1819" refString="Grados, J. &amp; Ramirez, J. J. (2021) Lista anotada de los Euchromiina &quot; polillas avispas &quot; (Lepidoptera: Erebidae: Arctiinae: Arctiini) del departamento de Loreto (Peru), con el reporte de nuevos registros y sinonimos. Revista Peruana de Biologia, 28 (Especial), e 21906. https://doi.org/10.15381/rpb.v28iespecial.21906" type="journal volume" year="2021">Grados and Ramirez (2021)</bibRefCitation>
qualified this pouch as &quot;subabdominal pouch
<typeStatus id="54C1A8E2FFC94F595D6E9E48B4564260" box="[1331,1384,1564,1588]" pageId="1" pageNumber="598">Type</typeStatus>
III&quot;, however, unlike that present in
<taxonomicName id="4C7A6DC3FFC94F5959BB9E14B3B64203" authorityName="Kaye" authorityYear="1918" box="[486,648,1599,1623]" class="Insecta" family="Erebidae" genus="Tigridania" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="quadricincta">
<emphasis id="B90ECA52FFC94F5959BB9E14B3B64203" box="[486,648,1599,1623]" italics="true" pageId="1" pageNumber="598">T. quadricincta</emphasis>
</taxonomicName>
, the pouch present in
<taxonomicName id="4C7A6DC3FFC94F595B299E14B52E4203" authorityName="Alzate-Cano &amp; Wolff &amp; Laguerre" authorityYear="2025" box="[884,1040,1599,1623]" class="Insecta" family="Erebidae" genus="Tigridania" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="magdalenae">
<emphasis id="B90ECA52FFC94F595B299E14B52E4203" box="[884,1040,1599,1623]" italics="true" pageId="1" pageNumber="598">T. magdalenae</emphasis>
</taxonomicName>
is divided into two brushes.
</paragraph>
<paragraph id="8BC51640FFC94F59589A9E30B5EE4297" blockId="1.[151,1437,879,1983]" pageId="1" pageNumber="598">
<emphasis id="B90ECA52FFC94F59589A9E30B0244228" bold="true" box="[199,282,1636,1660]" pageId="1" pageNumber="598">Female</emphasis>
(figure 2, second row): Similar to the male, but slightly larger. Serrated antennae. Tergite VI covered with golden yellow scales. Sternites II-V with a black triangular patch. First third of the dorsal margin, before the separation of Cu2 and A1 with a cream-colored spot. Coxae inner face almost entirely black, just white at the base.
</paragraph>
<paragraph id="8BC51640FFC94F59589A9E84B461437B" blockId="1.[151,1437,879,1983]" pageId="1" pageNumber="598">
<emphasis id="B90ECA52FFC94F59589A9E84B0BD42B3" bold="true" box="[199,387,1743,1768]" pageId="1" pageNumber="598">Female genitalia</emphasis>
(figure 2J): Relatively small, occupying only tergites IX-XI. Papillae analis large and sclerotized, with short apophyses, the anteriors clearly shorter than the posteriors. Ductus bursae relatively elongated. Bursa copulatrix spheroidal, without appendix bursae or signum. Ductus seminalis helical, detaching near the lamella antevaginalis.
</paragraph>
</subSubSection>
<subSubSection id="C36045CBFFC94F59589A9F6FB2E14323" pageId="1" pageNumber="598" type="distribution">
<paragraph id="8BC51640FFC94F59589A9F6FB2E14323" blockId="1.[151,1437,879,1983]" pageId="1" pageNumber="598">
<emphasis id="B90ECA52FFC94F59589A9F6FB06C4307" bold="true" box="[199,338,1851,1875]" pageId="1" pageNumber="598">Distribution</emphasis>
: Humid forests of the east hillside of the Central Cordillera in the
<collectingRegion id="49BED8A2FFC94F595C7C9F6FB5A94307" box="[1057,1175,1851,1875]" country="Colombia" name="Magdalena" pageId="1" pageNumber="598">Magdalena</collectingRegion>
valley. It is only known from two localities. Found during the day inside secondary and primary forest.
</paragraph>
</subSubSection>
<subSubSection id="C36045CBFFC94F59589A9FD0B03643EB" pageId="1" pageNumber="598" type="etymology">
<paragraph id="8BC51640FFC94F59589A9FD0B03643EB" blockId="1.[151,1437,879,1983]" pageId="1" pageNumber="598">
<emphasis id="B90ECA52FFC94F59589A9FD0B07543C8" bold="true" box="[199,331,1924,1948]" pageId="1" pageNumber="598">Etymology:</emphasis>
Named in honor of the
<collectingRegion id="49BED8A2FFC94F595A019FD7B3EC43CF" box="[604,722,1923,1947]" country="Colombia" name="Magdalena" pageId="1" pageNumber="598">Magdalena</collectingRegion>
River, the most economically and historically important river in
<collectingCountry id="F36D56D0FFC94F5958CA9FF3B03A43EB" box="[151,260,1959,1983]" name="Colombia" pageId="1" pageNumber="598">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF0546C8FFCA4F5A58CA9EF3B2144353" ID-DOI="http://doi.org/10.5281/zenodo.14804493" ID-Zenodo-Dep="14804493" httpUri="https://zenodo.org/record/14804493/files/figure.png" pageId="2" pageNumber="599" startId="2.[151,250,1703,1727]" targetBox="[172,1413,199,1666]" targetPageId="2" targetType="figure">
<paragraph id="8BC51640FFCA4F5A58CA9EF3B2144353" blockId="2.[151,1436,1703,1800]" pageId="2" pageNumber="599">
<emphasis id="B90ECA52FFCA4F5A58CA9EF3B00E42EB" bold="true" box="[151,304,1703,1727]" pageId="2" pageNumber="599">FIGURE 2. A.</emphasis>
Male holotype dorsal view.
<emphasis id="B90ECA52FFCA4F5A5A129EFCB3584294" bold="true" box="[591,614,1704,1728]" pageId="2" pageNumber="599">B.</emphasis>
Male holotype ventral view.
<emphasis id="B90ECA52FFCA4F5A5BD39EF3B29842EB" bold="true" box="[910,934,1703,1727]" pageId="2" pageNumber="599">C.</emphasis>
Female paratype dorsal view
<emphasis id="B90ECA52FFCA4F5A5C8B9EFCB5D04294" bold="true" box="[1238,1262,1704,1728]" pageId="2" pageNumber="599">D.</emphasis>
Female paratype ventral view.
<emphasis id="B90ECA52FFCA4F5A597C9E98B00642B0" bold="true" box="[289,312,1740,1764]" pageId="2" pageNumber="599">E.</emphasis>
Male genitalia dorsal view.
<emphasis id="B90ECA52FFCA4F5A5A049E98B35242B0" bold="true" box="[601,620,1740,1764]" pageId="2" pageNumber="599">F.</emphasis>
Male genitalia lateral view.
<emphasis id="B90ECA52FFCA4F5A5BD39E9FB29642B7" bold="true" box="[910,936,1739,1763]" pageId="2" pageNumber="599">G.</emphasis>
Male genitalia ventral view.
<emphasis id="B90ECA52FFCA4F5A5C8F9E98B5D242B0" bold="true" box="[1234,1260,1740,1764]" pageId="2" pageNumber="599">H.</emphasis>
Aedeagus lateral view.
<emphasis id="B90ECA52FFCA4F5A588E9EA4B1DD435C" bold="true" box="[211,227,1776,1800]" pageId="2" pageNumber="599">I.</emphasis>
Scent pouches signaled with arrows.
<emphasis id="B90ECA52FFCA4F5A5A3F9EA4B34B435C" bold="true" box="[610,629,1776,1800]" pageId="2" pageNumber="599">J.</emphasis>
Female genitalia.
</paragraph>
</caption>
<subSubSection id="C36045CBFFCA4F5A589A9F08B0BF43EF" pageId="2" pageNumber="599" type="biology_ecology">
<paragraph id="8BC51640FFCA4F5A589A9F08B0BF43EF" blockId="2.[151,1436,1883,1979]" pageId="2" pageNumber="599">
<emphasis id="B90ECA52FFCA4F5A589A9F08B0154320" bold="true" box="[199,299,1884,1908]" pageId="2" pageNumber="599">Ecology:</emphasis>
Some pairs were seen in the forest copulating during the month of March. One pair collected while copulating (
<figureCitation id="13410AC5FFCA4F5A594B9F2BB0B843CC" box="[278,390,1919,1944]" captionStart="FIGURE 1" captionStartId="1.[151,250,735,759]" captionTargetBox="[164,1426,237,704]" captionTargetId="figure-46@1.[162,1426,235,704]" captionTargetPageId="1" captionText="FIGURE 1. A. Distribution map. B. Pair of Tigridania magdalenae sp. nov. in copulation. They were observed inside a primary forest. (Photo: Camilo Flórez Valencia)." figureDoi="http://doi.org/10.5281/zenodo.14804489" httpUri="https://zenodo.org/record/14804489/files/figure.png" pageId="2" pageNumber="599">Figure 1B</figureCitation>
). Additionally, one individual was found during the month of July. It seems that this species is at least partially diurnal.
</paragraph>
</subSubSection>
</treatment>
</document>