treatments-xml/data/03/D0/F9/03D0F946FFD5FFDB42ABFC36FBA0F877.xml

<document id="CC6E5429643935CDB2DAA6553BB238D7" ID-CLB-Dataset="30967" ID-DOI="10.11646/zootaxa.4374.2.7" ID-GBIF-Dataset="8377a8d1-29d2-46b3-b0a5-2651e79a1178" ID-ISSN="1175-5326" ID-Zenodo-Dep="1154478" ID-ZooBank="CDC1F5C4-0F3D-4042-9045-BA3C62883386" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1516263969504" checkinUser="plazi" docAuthor="Dow, Rory A., Kompier, Tom &amp; Phan, Quoc Toan" docDate="2018" docId="03D0F946FFD5FFDB42ABFC36FBA0F877" docLanguage="en" docName="zootaxa.4374.2.7.pdf" docOrigin="Zootaxa 4374 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Drepanosticta emtrai Dow &amp; Kompier &amp; Phan 2018, sp. nov." docType="treatment" docUuid="BDC699DA-0DE6-4C10-9CB0-2B394DE345EB" docUuidSource="ZooBank" docVersion="8" lastPageNumber="278" masterDocId="FFE9813EFFD4FFDE423CFFC4FFD4FFCF" masterDocTitle="Drepanosticta emtrai sp. nov. from Vietnam with a discussion of Drepanosticta vietnamica Asahina, 1997 (Odonata: Zygoptera: Platystictidae)" masterLastPageNumber="282" masterPageNumber="273" pageNumber="274" updateTime="1698515973834" updateUser="ExternalLinkService">
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<mods:title id="025A17546DF595C93468F4D4E68C169E">Drepanosticta emtrai sp. nov. from Vietnam with a discussion of Drepanosticta vietnamica Asahina, 1997 (Odonata: Zygoptera: Platystictidae)</mods:title>
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<mods:namePart id="5D395690BB21E93DC620ED73DB5FA238">Dow, Rory A.</mods:namePart>
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<mods:namePart id="77E1304AAED6F8CF63AFEBA25F79775E">Kompier, Tom</mods:namePart>
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<mods:namePart id="82F2E5A3A674D1D568EF0073BD7E879A">Phan, Quoc Toan</mods:namePart>
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<taxonomicName id="4C7933D3FFD5FFDF42ABFC36FE41FBC3" ID-CoL="9X2JQ" authority="Dow &amp; Kompier &amp; Phan, 2018" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[151,405,1010,1036]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="274" phylum="Arthropoda" rank="species" species="emtrai" status="sp. nov.">
<emphasis id="B90D9442FFD5FFDF42ABFC36FE41FBC3" bold="true" box="[151,405,1010,1036]" italics="true" pageId="1" pageNumber="274">Drepanosticta emtrai</emphasis>
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(
<figureCitation id="134254D5FFD5FFDF42A3FBD1FEC5FBE1" box="[159,273,1045,1070]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–12" captionStart-2="FIGURES 13–16" captionStartId-0="2.[151,264,1969,1992]" captionStartId-1="3.[151,264,1376,1399]" captionStartId-2="4.[151,264,1061,1084]" captionTargetBox-0="[151,1435,450,1948]" captionTargetBox-1="[151,1435,193,1355]" captionTargetBox-2="[150,1412,193,1026]" captionTargetId-0="figure@2.[151,1435,450,1948]" captionTargetId-1="figure@3.[151,1435,193,1355]" captionTargetId-2="figure@4.[150,1436,193,1041]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1–6. Drepanosticta emtrai. Head: (1) holotype male; (2) paratype female.Prothorax, dorsal: (3) holotype male; (4) paratype female. Prothorax lateral:(5) holotype male; (6) paratype female." captionText-1="FIGURES 7–12. Drepanosticta emtrai. Synthorax, lateral: (7) holotype male; (8) paratype female. Terminal abdominal segments, dorsal: (9) holotype male; (10) paratype female. Terminal abdominal segments, lateral: (11) holotype male; (12) paratype female." captionText-2="FIGURES 13–16. Drepanosticta emtrai. Anal appendages and genital ligula of holotype male. Anal appendages: (13) dorsal; (14) lateral; (15) left paraproct in ventral view.(16) Genitalligula, ventral view." httpUri-0="https://zenodo.org/record/1154480/files/figure.png" httpUri-1="https://zenodo.org/record/1154482/files/figure.png" httpUri-2="https://zenodo.org/record/1154484/files/figure.png" pageId="1" pageNumber="274">Figs 1–16</figureCitation>
, 
<figureCitation id="134254D5FFD5FFDF4321FBD2FEEEFBE1" box="[285,314,1046,1070]" captionStart="FIGURE 21" captionStartId="7.[151,250,1933,1955]" captionTargetBox="[151,1435,663,1910]" captionTargetId="figure@7.[151,1435,663,1910]" captionTargetPageId="7" captionText="FIGURE 21. Distributions of Drepanosticta emtrai and D. vietnamica. Yellow circles: D. emtrai type series; red square: specimen excluded from the type series of D. emtrai (see remarks section under D. emtrai); black circle: D. vietnamica (approximate position of type locality)." httpUri="https://zenodo.org/record/1154488/files/figure.png" pageId="1" pageNumber="274">21</figureCitation>
)
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.
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, 
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, 
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Ha 
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, 
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(ca 
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, leg. 
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, to be deposited in 
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.
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.
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, in tandem with 
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, to be deposited in 
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; 
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, 
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, 
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(
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), 
<date id="FFC76E90FFD5FFDF4323FB0EFE4FFB2D" box="[287,411,1225,1250]" pageId="1" pageNumber="274" value="2015-05-10">10 v 2015</date>
, leg. T. Kompier, in coll. Dow; 
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, Ba Vi National Park, 
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Province, 
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(
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), 
<date id="FFC76E90FFD5FFDF43A0FB2AFDF4FAC9" box="[412,544,1261,1286]" pageId="1" pageNumber="274" value="2017-06-18">18 vi 2017</date>
, leg. T. Kompier; 
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(semi-teneral), Huay Bordop, Kaew Neua pass area, 
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(
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, 
<date id="FFC76E90FFD5FFDF40C4FAD6FCA7FAE5" box="[760,883,1297,1322]" pageId="1" pageNumber="274" value="2012-04-30">30 iv 2012</date>
, leg. M. Hämäläinen, in coll. Dow.
</paragraph>
<paragraph id="8BC64850FFD5FFDF42FBFAF1FE18FABD" blockId="1.[151,1437,1117,2006]" pageId="1" pageNumber="274">
<emphasis id="B90D9442FFD5FFDF42FBFAF1FE56FA81" bold="true" box="[199,386,1333,1358]" pageId="1" pageNumber="274">Other material.</emphasis>
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, 
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(
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, 
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), 
<date id="FFC76E90FFD5FFDF41A1FAF2FBCCFA81" box="[925,1048,1333,1358]" pageId="1" pageNumber="274" value="2014-06-27">27 vi 2014</date>
, leg. T. Kompier, in coll. Dow, see the Remarks section below.
</paragraph>
</subSubSection>
<subSubSection id="C3631BDBFFD5FFDF42FBFAB9FB5AFA11" pageId="1" pageNumber="274" type="etymology">
<paragraph id="8BC64850FFD5FFDF42FBFAB9FB5AFA11" blockId="1.[151,1437,1117,2006]" pageId="1" pageNumber="274">
<emphasis id="B90D9442FFD5FFDF42FBFAB9FE9AFA59" bold="true" box="[199,334,1405,1430]" pageId="1" pageNumber="274">Etymology.</emphasis>
<taxonomicName id="4C7933D3FFD5FFDF4369FABAFE4AFA5A" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[341,414,1406,1429]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="274" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD5FFDF4369FABAFE4AFA5A" box="[341,414,1406,1429]" italics="true" pageId="1" pageNumber="274">emtrai</emphasis>
</taxonomicName>
, a noun in apposition; this word means younger brother in Vietnamese. It is a reference to the slightly smaller size and apparently slightly later date of emergence of the species compared to 
<taxonomicName id="4C7933D3FFD5FFDF47BDFA66FEE9FA11" authorityName="Wilson" authorityYear="1997" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="274" phylum="Arthropoda" rank="species" species="hongkongensis">
<emphasis id="B90D9442FFD5FFDF47BDFA66FEE9FA11" italics="true" pageId="1" pageNumber="274">D. hongkongensis</emphasis>
</taxonomicName>
, to which it is very similar and with which it co-occurs on Ba Vi Mountain.
</paragraph>
</subSubSection>
<subSubSection id="C3631BDBFFD5FFDA42FBFA2DFF11F8BF" lastPageId="4" lastPageNumber="277" pageId="1" pageNumber="274" type="description">
<paragraph id="8BC64850FFD5FFDF42FBFA2DFAB5F915" blockId="1.[151,1437,1117,2006]" pageId="1" pageNumber="274">
<emphasis id="B90D9442FFD5FFDF42FBFA2DFDFFF9CE" bold="true" box="[199,555,1513,1538]" pageId="1" pageNumber="274">
Description of 
<typeStatus id="54C2F6F2FFD5FFDF4347FA2DFE35F9CD" box="[379,481,1513,1538]" pageId="1" pageNumber="274" type="holotype">holotype</typeStatus>
male.
</emphasis>
<emphasis id="B90D9442FFD5FFDF4008FA2DFDA5F9CD" box="[564,625,1513,1538]" italics="true" pageId="1" pageNumber="274">Head</emphasis>
(
<figureCitation id="134254D5FFD5FFDF40BEFA2DFD11F9CD" box="[642,709,1513,1538]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1969,1992]" captionTargetBox="[151,1435,450,1948]" captionTargetId="figure@2.[151,1435,450,1948]" captionTargetPageId="2" captionText="FIGURES 1–6. Drepanosticta emtrai. Head: (1) holotype male; (2) paratype female.Prothorax, dorsal: (3) holotype male; (4) paratype female. Prothorax lateral:(5) holotype male; (6) paratype female." httpUri="https://zenodo.org/record/1154480/files/figure.png" pageId="1" pageNumber="274">Fig. 1</figureCitation>
). Labium brown and pale greyish. Labrum pale blue, narrowly black along free margin. Mandible bases pale blue, narrowly black along free margin and part way beside genae. Anteclypeus pale blue, postclypeus black. Vertex, frons, occiput and underside of head black, small brownish marks behind antennae bases, immediately anterior of median ocellus (not clearly visible in 
<figureCitation id="134254D5FFD5FFDF46EEF991FAC3F9A1" box="[1234,1303,1621,1646]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1969,1992]" captionTargetBox="[151,1435,450,1948]" captionTargetId="figure@2.[151,1435,450,1948]" captionTargetPageId="2" captionText="FIGURES 1–6. Drepanosticta emtrai. Head: (1) holotype male; (2) paratype female.Prothorax, dorsal: (3) holotype male; (4) paratype female. Prothorax lateral:(5) holotype male; (6) paratype female." httpUri="https://zenodo.org/record/1154480/files/figure.png" pageId="1" pageNumber="274">Fig. 1</figureCitation>
because of angle), immediately exterior to lateral ocelli. Vertex relatively broad, ratio of width of compound eye to width of vertex measured at level of lateral ocelli slightly more than ca 0.7. Transverse occipital carina moderately developed. Ocelli whitish. Antenna with scape and pedicel grey, pedicel brown at top, remainder dark brown.
</paragraph>
<paragraph id="8BC64850FFD5FFDC42FBF921FBC5FE41" blockId="1.[151,1437,1117,2006]" lastBlockId="2.[151,1437,151,398]" lastPageId="2" lastPageNumber="275" pageId="1" pageNumber="274">
<emphasis id="B90D9442FFD5FFDF42FBF921FECCF931" box="[199,280,1765,1790]" italics="true" pageId="1" pageNumber="274">Thorax</emphasis>
. Propleuron mostly black and dark brown, paler areas along lower margin (
<figureCitation id="134254D5FFD5FFDF4646F921FB14F931" box="[1146,1216,1765,1790]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1969,1992]" captionTargetBox="[151,1435,450,1948]" captionTargetId="figure@2.[151,1435,450,1948]" captionTargetPageId="2" captionText="FIGURES 1–6. Drepanosticta emtrai. Head: (1) holotype male; (2) paratype female.Prothorax, dorsal: (3) holotype male; (4) paratype female. Prothorax lateral:(5) holotype male; (6) paratype female." httpUri="https://zenodo.org/record/1154480/files/figure.png" pageId="1" pageNumber="274">Fig. 5</figureCitation>
). Anterior pronotal lobe low, largely blue, dark brown to rear and on lateral extremities (
<figureCitation id="134254D5FFD5FFDF418FF8CDFBC8F8ED" box="[947,1052,1801,1826]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1969,1992]" captionTargetBox="[151,1435,450,1948]" captionTargetId="figure@2.[151,1435,450,1948]" captionTargetPageId="2" captionText="FIGURES 1–6. Drepanosticta emtrai. Head: (1) holotype male; (2) paratype female.Prothorax, dorsal: (3) holotype male; (4) paratype female. Prothorax lateral:(5) holotype male; (6) paratype female." httpUri="https://zenodo.org/record/1154480/files/figure.png" pageId="1" pageNumber="274">Figs 3, 5</figureCitation>
). Middle pronotal lobe black and dark brown with pair of dorsal blue markings (
<figureCitation id="134254D5FFD5FFDF4096F8E9FCC5F889" box="[682,785,1837,1862]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1969,1992]" captionTargetBox="[151,1435,450,1948]" captionTargetId="figure@2.[151,1435,450,1948]" captionTargetPageId="2" captionText="FIGURES 1–6. Drepanosticta emtrai. Head: (1) holotype male; (2) paratype female.Prothorax, dorsal: (3) holotype male; (4) paratype female. Prothorax lateral:(5) holotype male; (6) paratype female." httpUri="https://zenodo.org/record/1154480/files/figure.png" pageId="1" pageNumber="274">Figs 3, 5</figureCitation>
). Posterior lobe black centrally, dark brown with obscure paler areas to sides, where somewhat expanded (
<figureCitation id="134254D5FFD5FFDF40FEF895FCD3F8A5" box="[706,775,1873,1898]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1969,1992]" captionTargetBox="[151,1435,450,1948]" captionTargetId="figure@2.[151,1435,450,1948]" captionTargetPageId="2" captionText="FIGURES 1–6. Drepanosticta emtrai. Head: (1) holotype male; (2) paratype female.Prothorax, dorsal: (3) holotype male; (4) paratype female. Prothorax lateral:(5) holotype male; (6) paratype female." httpUri="https://zenodo.org/record/1154480/files/figure.png" pageId="1" pageNumber="274">Fig. 3</figureCitation>
). Synthorax: Mesepisternum, mesepimeron metallic black with greenish cast except mid dorsal carina where bronzy black, antealar triangles black, mesostigmal plates greyish brown. Metepisternum and metepimeron mostly dark brown, blue marking on metepisternum, well defined towards wing bases but obscure towards legs (
<figureCitation id="134254D5FFD5FFDF40A3F879FD30F819" box="[671,740,1981,2006]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1376,1399]" captionTargetBox="[151,1435,193,1355]" captionTargetId="figure@3.[151,1435,193,1355]" captionTargetPageId="3" captionText="FIGURES 7–12. Drepanosticta emtrai. Synthorax, lateral: (7) holotype male; (8) paratype female. Terminal abdominal segments, dorsal: (9) holotype male; (10) paratype female. Terminal abdominal segments, lateral: (11) holotype male; (12) paratype female." httpUri="https://zenodo.org/record/1154482/files/figure.png" pageId="1" pageNumber="274">Fig. 7</figureCitation>
), small yellowish mark immediately above spiracle. Venter of synthorax pale, gentle swelling visible near abdomen (
<figureCitation id="134254D5FFD6FFDC4131FF53FC80FF7F" box="[781,852,151,176]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1376,1399]" captionTargetBox="[151,1435,193,1355]" captionTargetId="figure@3.[151,1435,193,1355]" captionTargetPageId="3" captionText="FIGURES 7–12. Drepanosticta emtrai. Synthorax, lateral: (7) holotype male; (8) paratype female. Terminal abdominal segments, dorsal: (9) holotype male; (10) paratype female. Terminal abdominal segments, lateral: (11) holotype male; (12) paratype female." httpUri="https://zenodo.org/record/1154482/files/figure.png" pageId="2" pageNumber="275">Fig. 7</figureCitation>
). Mesokatepisternum dark brown, pale adjacent to coxa, metakatepisternum mostly pale. Legs each with coxa, trochanter and femur pale yellowish, femur with narrow, faint, irregular dark stripe along much of lateral extensor surface, dark immediately above joint with tibia. Tibia and tarsi mostly pale with indistinct darker areas, tarsal claws pale brown. Wings: 14 Px (left), 16 Px (right) in Fw, 
<quantity id="4C81E5B5FFD6FFDC42D9FEECFECBFE8F" box="[229,287,295,320]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="2" pageNumber="275" unit="in" value="14.0">14 in</quantity>
Hw. Vein ab present, bifurcated. Arculus slightly distal to Ax2. R4 arising slightly distal to subnodus, IR3 joined to it by short stalk. Pterostigma ca trapezoidal with costal side shorter than anal side, dark brownish black with narrow pale border, covering slightly more than one underlying cell.
</paragraph>
<caption id="DF0618D8FFD6FFDC42ABF875FC5FF829" httpUri="https://zenodo.org/record/1154480/files/figure.png" pageId="2" pageNumber="275" targetBox="[151,1435,450,1948]" targetPageId="2">
<paragraph id="8BC64850FFD6FFDC42ABF875FC5FF829" blockId="2.[151,1436,1969,2022]" pageId="2" pageNumber="275">
<emphasis id="B90D9442FFD6FFDC42ABF875FEEEF809" bold="true" box="[151,314,1969,1992]" pageId="2" pageNumber="275">FIGURES 1–6.</emphasis>
<taxonomicName id="4C7933D3FFD6FFDC437DF876FDCDF808" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[321,537,1970,1991]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="275" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD6FFDC437DF876FDCDF808" box="[321,537,1970,1991]" italics="true" pageId="2" pageNumber="275">Drepanosticta emtrai</emphasis>
</taxonomicName>
. Head: (1) holotype male; (2) paratype female. Prothorax, dorsal: (3) holotype male; (4) paratype female. Prothorax lateral: (5) holotype male; (6) paratype female.
</paragraph>
</caption>
<caption id="DF0618D8FFD7FFDD42ABFAA4FE95FA7B" httpUri="https://zenodo.org/record/1154482/files/figure.png" pageId="3" pageNumber="276" targetBox="[151,1435,193,1355]" targetPageId="3">
<paragraph id="8BC64850FFD7FFDD42ABFAA4FE95FA7B" blockId="3.[151,1436,1376,1460]" pageId="3" pageNumber="276">
<emphasis id="B90D9442FFD7FFDD42ABFAA4FE98FAB9" bold="true" box="[151,332,1376,1399]" pageId="3" pageNumber="276">FIGURES 7–12.</emphasis>
<taxonomicName id="4C7933D3FFD7FFDD4364FAA6FDE2FAB8" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[344,566,1378,1399]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="276" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD7FFDD4364FAA6FDE2FAB8" box="[344,566,1378,1399]" italics="true" pageId="3" pageNumber="276">Drepanosticta emtrai</emphasis>
</taxonomicName>
. Synthorax, lateral: (7) holotype male; (8) paratype female. Terminal abdominal segments, dorsal: (9) holotype male; (10) paratype female. Terminal abdominal segments, lateral: (11) holotype male; (12) paratype female.
</paragraph>
</caption>
<paragraph id="8BC64850FFD7FFDD42FBFA1FFEF4F86B" blockId="3.[151,1437,1499,2028]" pageId="3" pageNumber="276">
<emphasis id="B90D9442FFD7FFDD42FBFA1FFEE5FA3B" box="[199,305,1499,1524]" italics="true" pageId="3" pageNumber="276">Abdomen</emphasis>
. S1–2 mostly dark brown with poorly defined pale areas lower laterally. S3–7 dark brown, black apically with broad pale basal annulus, smallest on S3. S8 very dark brown, blue dorsal apical mark, pair of small blue marks just basal of this (
<figureCitation id="134254D5FFD7FFDD43DDF9E7FDF1F9F3" box="[481,549,1571,1596]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1376,1399]" captionTargetBox="[151,1435,193,1355]" captionTargetId="figure@3.[151,1435,193,1355]" captionTargetPageId="3" captionText="FIGURES 7–12. Drepanosticta emtrai. Synthorax, lateral: (7) holotype male; (8) paratype female. Terminal abdominal segments, dorsal: (9) holotype male; (10) paratype female. Terminal abdominal segments, lateral: (11) holotype male; (12) paratype female." httpUri="https://zenodo.org/record/1154482/files/figure.png" pageId="3" pageNumber="276">Fig. 9</figureCitation>
), basal ca one half of segment with pale area lower laterally (
<figureCitation id="134254D5FFD7FFDD46E5F9E7FAFFF9F3" box="[1241,1323,1571,1596]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1376,1399]" captionTargetBox="[151,1435,193,1355]" captionTargetId="figure@3.[151,1435,193,1355]" captionTargetPageId="3" captionText="FIGURES 7–12. Drepanosticta emtrai. Synthorax, lateral: (7) holotype male; (8) paratype female. Terminal abdominal segments, dorsal: (9) holotype male; (10) paratype female. Terminal abdominal segments, lateral: (11) holotype male; (12) paratype female." httpUri="https://zenodo.org/record/1154482/files/figure.png" pageId="3" pageNumber="276">Fig. 11</figureCitation>
). S9 blue, brown lower laterally. S10 blue, very dark brown laterally. Anal appendages (
<figureCitation id="134254D5FFD7FFDD4639F983FB51F9AF" box="[1029,1157,1607,1632]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1061,1084]" captionTargetBox="[150,1412,193,1026]" captionTargetId="figure@4.[150,1436,193,1041]" captionTargetPageId="4" captionText="FIGURES 13–16. Drepanosticta emtrai. Anal appendages and genital ligula of holotype male. Anal appendages: (13) dorsal; (14) lateral; (15) left paraproct in ventral view.(16) Genitalligula, ventral view." httpUri="https://zenodo.org/record/1154484/files/figure.png" pageId="3" pageNumber="276">Figs 13–15</figureCitation>
) largely dark brown and black, paler areas towards tips of cerci. Cerci almost 3 times length of S10, in lateral view broad, arched up along lower margin with large heel-like subtriangular ventral flap arising abruptly just after half length, apex of this rounded (
<figureCitation id="134254D5FFD7FFDD433AF977FE89F903" box="[262,349,1715,1740]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1061,1084]" captionTargetBox="[150,1412,193,1026]" captionTargetId="figure@4.[150,1436,193,1041]" captionTargetPageId="4" captionText="FIGURES 13–16. Drepanosticta emtrai. Anal appendages and genital ligula of holotype male. Anal appendages: (13) dorsal; (14) lateral; (15) left paraproct in ventral view.(16) Genitalligula, ventral view." httpUri="https://zenodo.org/record/1154484/files/figure.png" pageId="3" pageNumber="276">Fig. 14</figureCitation>
). Cerci downturned towards apices, these rounded. In dorsal view (
<figureCitation id="134254D5FFD7FFDD4649F977FB1EF903" box="[1141,1226,1715,1740]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1061,1084]" captionTargetBox="[150,1412,193,1026]" captionTargetId="figure@4.[150,1436,193,1041]" captionTargetPageId="4" captionText="FIGURES 13–16. Drepanosticta emtrai. Anal appendages and genital ligula of holotype male. Anal appendages: (13) dorsal; (14) lateral; (15) left paraproct in ventral view.(16) Genitalligula, ventral view." httpUri="https://zenodo.org/record/1154484/files/figure.png" pageId="3" pageNumber="276">Fig. 13</figureCitation>
) cerci converging gently along outer margin over entire length, interior margins with small rounded interior flap at ca half their length, followed by slightly larger one subapically, neither visible in lateral view. Paraprocts slightly shorter than cerci, in lateral view broad at base, narrowing rapidly, with dorsal hump before half-length, thereafter slightly sinuous, apices brownish. In ventral view (
<figureCitation id="134254D5FFD7FFDD4044F887FD1DF893" box="[632,713,1859,1884]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1061,1084]" captionTargetBox="[150,1412,193,1026]" captionTargetId="figure@4.[150,1436,193,1041]" captionTargetPageId="4" captionText="FIGURES 13–16. Drepanosticta emtrai. Anal appendages and genital ligula of holotype male. Anal appendages: (13) dorsal; (14) lateral; (15) left paraproct in ventral view.(16) Genitalligula, ventral view." httpUri="https://zenodo.org/record/1154484/files/figure.png" pageId="3" pageNumber="276">Fig. 15</figureCitation>
) narrowing rapidly after half-length, in turned at apices; slender subbasal interior tooth visible, directed upwards and to rear; this also visible in dorsal view, and just visible in lateral view.
</paragraph>
<paragraph id="8BC64850FFD7FFDD42FBF86BFB19F824" blockId="3.[151,1437,1499,2028]" pageId="3" pageNumber="276">
<emphasis id="B90D9442FFD7FFDD42FBF86BFEB2F807" box="[199,358,1967,1992]" italics="true" pageId="3" pageNumber="276">Genital ligula</emphasis>
(
<figureCitation id="134254D5FFD7FFDD4344F86BFE19F807" box="[376,461,1967,1992]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1061,1084]" captionTargetBox="[150,1412,193,1026]" captionTargetId="figure@4.[150,1436,193,1041]" captionTargetPageId="4" captionText="FIGURES 13–16. Drepanosticta emtrai. Anal appendages and genital ligula of holotype male. Anal appendages: (13) dorsal; (14) lateral; (15) left paraproct in ventral view.(16) Genitalligula, ventral view." httpUri="https://zenodo.org/record/1154484/files/figure.png" pageId="3" pageNumber="276">Fig. 16</figureCitation>
). Broad internal fold, terminal segment slender, divided apically into pair of upward directed arms, these turned towards thorax and expanded apically; space between arms concave.
</paragraph>
<caption id="DF0618D8FFD0FFDA42ABFBE1FC14FB94" httpUri="https://zenodo.org/record/1154484/files/figure.png" pageId="4" pageNumber="277" targetBox="[150,1412,193,1026]" targetPageId="4">
<paragraph id="8BC64850FFD0FFDA42ABFBE1FC14FB94" blockId="4.[151,1436,1061,1115]" pageId="4" pageNumber="277">
<emphasis id="B90D9442FFD0FFDA42ABFBE1FE80FBF4" bold="true" box="[151,340,1061,1084]" pageId="4" pageNumber="277">FIGURES 13–16.</emphasis>
<taxonomicName id="4C7933D3FFD0FFDA4367FBE3FDE0FBF3" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[347,564,1063,1084]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD0FFDA4367FBE3FDE0FBF3" box="[347,564,1063,1084]" italics="true" pageId="4" pageNumber="277">Drepanosticta emtrai</emphasis>
</taxonomicName>
. Anal appendages and genital ligula of holotype male. Anal appendages: (13) dorsal; (14) lateral; (15) left paraproct in ventral view. (16) Genital ligula, ventral view.
</paragraph>
</caption>
<paragraph id="8BC64850FFD0FFDA42FBFB47FB94FB54" blockId="4.[151,1436,1154,2013]" box="[199,1088,1154,1179]" pageId="4" pageNumber="277">Measurements (mm): abdomen with anal appendages 36, cercus ca 1, Hw 23.5.</paragraph>
<paragraph id="8BC64850FFD0FFDA42FBFB62FE91FB2C" blockId="4.[151,1436,1154,2013]" pageId="4" pageNumber="277">
<emphasis id="B90D9442FFD0FFDA42FBFB62FDEBFB70" bold="true" box="[199,575,1190,1215]" pageId="4" pageNumber="277">
Description of female 
<typeStatus id="54C2F6F2FFD0FFDA43F0FB62FDEEFB70" box="[460,570,1190,1215]" pageId="4" pageNumber="277" type="paratype">paratype</typeStatus>
.
</emphasis>
As male except as noted. 
<emphasis id="B90D9442FFD0FFDA415EFB62FC4AFB70" box="[866,926,1190,1215]" italics="true" pageId="4" pageNumber="277">Head</emphasis>
(
<figureCitation id="134254D5FFD0FFDA4191FB62FC27FB70" box="[941,1011,1190,1215]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1969,1992]" captionTargetBox="[151,1435,450,1948]" captionTargetId="figure@2.[151,1435,450,1948]" captionTargetPageId="2" captionText="FIGURES 1–6. Drepanosticta emtrai. Head: (1) holotype male; (2) paratype female.Prothorax, dorsal: (3) holotype male; (4) paratype female. Prothorax lateral:(5) holotype male; (6) paratype female." httpUri="https://zenodo.org/record/1154480/files/figure.png" pageId="4" pageNumber="277">Fig. 2</figureCitation>
). Without small greyish marks behind antennae bases.
</paragraph>
<paragraph id="8BC64850FFD0FFDA42FBFB2AFE4FFA9E" blockId="4.[151,1436,1154,2013]" pageId="4" pageNumber="277">
<emphasis id="B90D9442FFD0FFDA42FBFB2AFECCFAC8" box="[199,280,1262,1287]" italics="true" pageId="4" pageNumber="277">Thorax</emphasis>
. Blue marks on middle pronotal lobe slightly larger (
<figureCitation id="134254D5FFD0FFDA41A0FB2AFBDFFAC8" box="[924,1035,1262,1287]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1969,1992]" captionTargetBox="[151,1435,450,1948]" captionTargetId="figure@2.[151,1435,450,1948]" captionTargetPageId="2" captionText="FIGURES 1–6. Drepanosticta emtrai. Head: (1) holotype male; (2) paratype female.Prothorax, dorsal: (3) holotype male; (4) paratype female. Prothorax lateral:(5) holotype male; (6) paratype female." httpUri="https://zenodo.org/record/1154480/files/figure.png" pageId="4" pageNumber="277">Figs 4, 6</figureCitation>
); blue marking on metepisternum smaller, less well defined (
<figureCitation id="134254D5FFD0FFDA43FCFAD6FDD7FAE4" box="[448,515,1298,1323]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1376,1399]" captionTargetBox="[151,1435,193,1355]" captionTargetId="figure@3.[151,1435,193,1355]" captionTargetPageId="3" captionText="FIGURES 7–12. Drepanosticta emtrai. Synthorax, lateral: (7) holotype male; (8) paratype female. Terminal abdominal segments, dorsal: (9) holotype male; (10) paratype female. Terminal abdominal segments, lateral: (11) holotype male; (12) paratype female." httpUri="https://zenodo.org/record/1154482/files/figure.png" pageId="4" pageNumber="277">Fig. 8</figureCitation>
). Wings: R4 arising at subnodus in right Hw, very slightly distal to it in left Hw, 16 Px in Fw, 15 Px in Hw.
</paragraph>
<paragraph id="8BC64850FFD0FFDA42FBFA99FEA4FA71" blockId="4.[151,1436,1154,2013]" pageId="4" pageNumber="277">
<emphasis id="B90D9442FFD0FFDA42FBFA99FEE5FAB9" box="[199,305,1373,1398]" italics="true" pageId="4" pageNumber="277">Abdomen</emphasis>
(
<figureCitation id="134254D5FFD0FFDA437EFA99FE1EFAB9" box="[322,458,1373,1398]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1376,1399]" captionTargetBox="[151,1435,193,1355]" captionTargetId="figure@3.[151,1435,193,1355]" captionTargetPageId="3" captionText="FIGURES 7–12. Drepanosticta emtrai. Synthorax, lateral: (7) holotype male; (8) paratype female. Terminal abdominal segments, dorsal: (9) holotype male; (10) paratype female. Terminal abdominal segments, lateral: (11) holotype male; (12) paratype female." httpUri="https://zenodo.org/record/1154482/files/figure.png" pageId="4" pageNumber="277">Figs 10, 12</figureCitation>
). S8 without blue markings, S9 blue dorsally in apical half, S10 blue dorsally, cerci black, just longer than S10, ovipositor extending to ca level of tips of cerci, mostly dark brown and black with pale apex, styles brown.
</paragraph>
<paragraph id="8BC64850FFD0FFDA42FBFA0EFBBCFA2E" blockId="4.[151,1436,1154,2013]" box="[199,1128,1481,1506]" pageId="4" pageNumber="277">Measurements (mm) Abdomen with anal appendages and ovipositor ca 33, Hw 23.</paragraph>
<paragraph id="8BC64850FFD0FFDA42FBFA29FAD5F8E6" blockId="4.[151,1436,1154,2013]" pageId="4" pageNumber="277">
<emphasis id="B90D9442FFD0FFDA42FBFA29FDC9F9C9" bold="true" box="[199,541,1517,1542]" pageId="4" pageNumber="277">
Variation in 
<typeStatus id="54C2F6F2FFD0FFDA4362FA29FE1EF9C9" box="[350,458,1517,1542]" pageId="4" pageNumber="277" type="paratype">paratype</typeStatus>
males.
</emphasis>
The 
<typeStatus id="54C2F6F2FFD0FFDA406BFA2AFD63F9C9" box="[599,695,1518,1542]" pageId="4" pageNumber="277" type="paratype">paratype</typeStatus>
from 
<collectingRegion id="49BD86B2FFD0FFDA40C1FA29FCB2F9C9" box="[765,870,1517,1542]" country="Vietnam" name="Nghe An" pageId="4" pageNumber="277">Nghe An</collectingRegion>
, 
<collectingCountry id="F36E08C0FFD0FFDA414EFA29FC07F9C9" box="[882,979,1517,1542]" name="Vietnam" pageId="4" pageNumber="277">Vietnam</collectingCountry>
, is very similar to the 
<typeStatus id="54C2F6F2FFD0FFDA46ECFA29FAE1F9C9" box="[1232,1333,1517,1542]" pageId="4" pageNumber="277" type="holotype">holotype</typeStatus>
, the only significant differences apart from size and vein counts is that S8 only has a small blue apical dorsal mark, that R4 arises at Sn in the left Hw, and that the apical halves of the paraprocts are straight in lateral view, not slightly sinuous. The five males from Ba Vi, 
<collectingCountry id="F36E08C0FFD0FFDA4072F998FD64F9BA" box="[590,688,1628,1653]" name="Vietnam" pageId="4" pageNumber="277">Vietnam</collectingCountry>
, have more extensive blue on S8, ranging from three-fourths to almost completely blue, and have the metepisternum completely black. Their paraprocts are all straight in lateral view, although the apex is turned up slightly in two specimens. In the semi-teneral male from 
<collectingCountry id="F36E08C0FFD0FFDA46C0F961FAE6F973" box="[1276,1330,1701,1724]" name="Laos" pageId="4" pageNumber="277">Laos</collectingCountry>
the blue marking on the metepisternum is faint, the extensor surface of the femur is extensively dark and there is a welldefined dark stripe on the extensor surface of the tibia. An apical blue dorsal mark is present on S8, but much of the rest of the dorsum has a diffuse, mottled bluish appearance (perhaps this fades in mature specimens).
</paragraph>
<paragraph id="8BC64850FFD0FFDA42FBF8F1FF11F8BF" blockId="4.[151,1436,1154,2013]" pageId="4" pageNumber="277">Measurements (mm). Males: Abdomen anal appendages 35–36.5, Hw 21–22, 15–16 Px in Fw, 14–15 Px in Hw.</paragraph>
</subSubSection>
<subSubSection id="C3631BDBFFD0FFDB42FBF8B8FDEBFDAF" lastPageId="5" lastPageNumber="278" pageId="4" pageNumber="277" type="diagnosis">
<paragraph id="8BC64850FFD0FFDB42FBF8B8FDEBFDAF" blockId="4.[151,1436,1154,2013]" lastBlockId="5.[151,1437,151,1977]" lastPageId="5" lastPageNumber="278" pageId="4" pageNumber="277">
<emphasis id="B90D9442FFD0FFDA42FBF8B8FE95F85A" bold="true" box="[199,321,1916,1941]" pageId="4" pageNumber="277">Diagnosis.</emphasis>
The male of 
<taxonomicName id="4C7933D3FFD0FFDA43D0F8B9FD89F85B" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[492,605,1917,1940]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD0FFDA43D0F8B9FD89F85B" box="[492,605,1917,1940]" italics="true" pageId="4" pageNumber="277">D. emtrai</emphasis>
</taxonomicName>
can be separated from all mainland Asian 
<taxonomicName id="4C7933D3FFD0FFDA4655F8B8FAD4F85A" box="[1129,1280,1916,1941]" class="Insecta" family="Platystictidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="277" phylum="Arthropoda" rank="family">Platystictidae</taxonomicName>
except other members of the 
<taxonomicName id="4C7933D3FFD0FFDA436CF865FE2EF876" baseAuthorityName="Laidlaw" baseAuthorityYear="1915" box="[336,506,1952,1977]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="carmichaeli">
<emphasis id="B90D9442FFD0FFDA436CF865FE2EF876" box="[336,506,1952,1977]" italics="true" pageId="4" pageNumber="277">D. carmichaeli</emphasis>
</taxonomicName>
-group by the characters defining that group, and from the South Asian species of the 
<taxonomicName id="4C7933D3FFD0FFDA42FCF800FE91F812" baseAuthorityName="Laidlaw" baseAuthorityYear="1915" box="[192,325,1988,2013]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="carmichaeli">
<emphasis id="B90D9442FFD0FFDA42FCF800FE91F812" box="[192,325,1988,2013]" italics="true" pageId="4" pageNumber="277">carmichaeli</emphasis>
</taxonomicName>
-group by the coloration of the prothorax and dorsum of the synthorax, which have more extensive pale marks, including antehumeral stripes in 
<taxonomicName id="4C7933D3FFD1FFDB408FFF5DFCB5FF7F" baseAuthorityName="Laidlaw" baseAuthorityYear="1915" box="[691,865,151,176]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="carmichaeli">
<emphasis id="B90D9442FFD1FFDB408FFF5DFCB5FF7F" box="[691,865,151,176]" italics="true" pageId="5" pageNumber="278">D. carmichaeli</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7933D3FFD1FFDB4198FF5DFBAFFF7F" authorityName="Fraser" authorityYear="1931" box="[932,1147,151,176]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="polychromatica">
<emphasis id="B90D9442FFD1FFDB4198FF5DFBAFFF7F" box="[932,1147,151,176]" italics="true" pageId="5" pageNumber="278">D. polychromatica</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EFE835A1FFD1FFDB46FEFF5CFA85FF7F" author="Fraser" box="[1218,1361,152,176]" pageId="5" pageNumber="278" refString="Fraser, F. C. (1933) The Fauna of British India, including Ceylon and Burma. Odonata. Vol. 1. Taylor &amp; Francis, London, xiii + 423 pp." type="book" year="1933">Fraser 1933</bibRefCitation>
). It is separated from 
<taxonomicName id="4C7933D3FFD1FFDB4378FF79FDDFFF1A" authorityName="Wilson" authorityYear="1997" box="[324,523,188,213]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="hongkongensis">
<emphasis id="B90D9442FFD1FFDB4378FF79FDDFFF1A" box="[324,523,188,213]" italics="true" pageId="5" pageNumber="278">D. hongkongensis</emphasis>
</taxonomicName>
by the blue markings on the anterior and middle pronotal lobes of 
<taxonomicName id="4C7933D3FFD1FFDB46CDFF79FA88FF1B" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[1265,1372,189,212]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD1FFDB46CDFF79FA88FF1B" box="[1265,1372,189,212]" italics="true" pageId="5" pageNumber="278">D. emtrai</emphasis>
</taxonomicName>
(
<figureCitation id="134254D5FFD1FFDB4756FF78FF17FF37" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1969,1992]" captionTargetBox="[151,1435,450,1948]" captionTargetId="figure@2.[151,1435,450,1948]" captionTargetPageId="2" captionText="FIGURES 1–6. Drepanosticta emtrai. Head: (1) holotype male; (2) paratype female.Prothorax, dorsal: (3) holotype male; (4) paratype female. Prothorax lateral:(5) holotype male; (6) paratype female." httpUri="https://zenodo.org/record/1154480/files/figure.png" pageId="5" pageNumber="278">Figs 3, 5</figureCitation>
) and by the curvature of the paraprocts in ventral view (
<figureCitation id="134254D5FFD1FFDB416EFF1BFC72FF37" box="[850,934,223,248]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1061,1084]" captionTargetBox="[150,1412,193,1026]" captionTargetId="figure@4.[150,1436,193,1041]" captionTargetPageId="4" captionText="FIGURES 13–16. Drepanosticta emtrai. Anal appendages and genital ligula of holotype male. Anal appendages: (13) dorsal; (14) lateral; (15) left paraproct in ventral view.(16) Genitalligula, ventral view." httpUri="https://zenodo.org/record/1154484/files/figure.png" pageId="5" pageNumber="278">Fig. 15</figureCitation>
), and from 
<taxonomicName id="4C7933D3FFD1FFDB4613FF25FB7BFF37" authorityName="Hamalainen" authorityYear="1999" box="[1071,1199,225,248]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="jurzitzai">
<emphasis id="B90D9442FFD1FFDB4613FF25FB7BFF37" box="[1071,1199,225,248]" italics="true" pageId="5" pageNumber="278">D. jurzitzai</emphasis>
</taxonomicName>
by the colour of the pterostigma, said to be yellow ochre in 
<taxonomicName id="4C7933D3FFD1FFDB405DFEC1FC1CFED3" authority="(Hamalainen 1999)" baseAuthorityName="Hamalainen" baseAuthorityYear="1999" box="[609,968,260,285]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="jurzitzai">
<emphasis id="B90D9442FFD1FFDB405DFEC1FD35FED3" box="[609,737,261,284]" italics="true" pageId="5" pageNumber="278">D. jurzitzai</emphasis>
(
<bibRefCitation id="EFE835A1FFD1FFDB40CEFEC0FC15FED3" author="Hamalainen" box="[754,961,260,285]" pageId="5" pageNumber="278" refString="Hamalainen, M. (1999) Drepanosticta jurzitzai spec. nov., a new damselfly from southeastern Thailand (Zygoptera: Platystictidae). Odonatologica, 28 (4), 421 - 423." type="journal article" year="1999">Hämäläinen 1999</bibRefCitation>
)
</taxonomicName>
, and from all species except possibly 
<taxonomicName id="4C7933D3FFD1FFDB47BDFEC1FEC6FE8F" authorityName="Asahina" authorityYear="1997" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="vietnamica">
<emphasis id="B90D9442FFD1FFDB47BDFEC1FEC6FE8F" italics="true" pageId="5" pageNumber="278">D. vietnamica</emphasis>
</taxonomicName>
(see below) by the heel-like subtriangular ventral flaps on the cerci (
<figureCitation id="134254D5FFD1FFDB461DFEE3FBA2FE8F" box="[1057,1142,295,320]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1061,1084]" captionTargetBox="[150,1412,193,1026]" captionTargetId="figure@4.[150,1436,193,1041]" captionTargetPageId="4" captionText="FIGURES 13–16. Drepanosticta emtrai. Anal appendages and genital ligula of holotype male. Anal appendages: (13) dorsal; (14) lateral; (15) left paraproct in ventral view.(16) Genitalligula, ventral view." httpUri="https://zenodo.org/record/1154484/files/figure.png" pageId="5" pageNumber="278">Fig. 14</figureCitation>
). It is also separated from 
<taxonomicName id="4C7933D3FFD1FFDB42ABFE89FEF2FEAA" baseAuthorityName="Tinkham" baseAuthorityYear="1938" box="[151,294,332,357]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="brownelli">
<emphasis id="B90D9442FFD1FFDB42ABFE89FEF2FEAA" box="[151,294,332,357]" italics="true" pageId="5" pageNumber="278">D. brownelli</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7933D3FFD1FFDB435EFE89FDF9FEAA" authorityName="Wilson" authorityYear="1997" box="[354,557,332,357]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="hongkongensis">
<emphasis id="B90D9442FFD1FFDB435EFE89FDF9FEAA" box="[354,557,332,357]" italics="true" pageId="5" pageNumber="278">D. hongkongensis</emphasis>
</taxonomicName>
by having the basal tooth on the paraprocts slender and only just visible in lateral view (
<figureCitation id="134254D5FFD1FFDB4309FEABFE58FE47" box="[309,396,367,392]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1061,1084]" captionTargetBox="[150,1412,193,1026]" captionTargetId="figure@4.[150,1436,193,1041]" captionTargetPageId="4" captionText="FIGURES 13–16. Drepanosticta emtrai. Anal appendages and genital ligula of holotype male. Anal appendages: (13) dorsal; (14) lateral; (15) left paraproct in ventral view.(16) Genitalligula, ventral view." httpUri="https://zenodo.org/record/1154484/files/figure.png" pageId="5" pageNumber="278">Fig. 14</figureCitation>
), compared with robust and easily visible in lateral view in these two species (see the Discussion). An additional character separating it from 
<taxonomicName id="4C7933D3FFD1FFDB417FFE51FC0CFE62" baseAuthorityName="Tinkham" baseAuthorityYear="1938" box="[835,984,404,429]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="brownelli">
<emphasis id="B90D9442FFD1FFDB417FFE51FC0CFE62" box="[835,984,404,429]" italics="true" pageId="5" pageNumber="278">D. brownelli</emphasis>
</taxonomicName>
is the position of the stripe on the metepisternum, which is centrally placed in 
<taxonomicName id="4C7933D3FFD1FFDB40B0FE7DFCCEFE1F" baseAuthorityName="Tinkham" baseAuthorityYear="1938" box="[652,794,439,464]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="brownelli">
<emphasis id="B90D9442FFD1FFDB40B0FE7DFCCEFE1F" box="[652,794,439,464]" italics="true" pageId="5" pageNumber="278">D. brownelli</emphasis>
</taxonomicName>
but almost reaches the antealar carina in 
<taxonomicName id="4C7933D3FFD1FFDB46CEFE7DFA8AFE1F" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[1266,1374,441,464]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD1FFDB46CEFE7DFA8AFE1F" box="[1266,1374,441,464]" italics="true" pageId="5" pageNumber="278">D. emtrai</emphasis>
</taxonomicName>
(
<figureCitation id="134254D5FFD1FFDB4752FE73FF73FE3B" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1376,1399]" captionTargetBox="[151,1435,193,1355]" captionTargetId="figure@3.[151,1435,193,1355]" captionTargetPageId="3" captionText="FIGURES 7–12. Drepanosticta emtrai. Synthorax, lateral: (7) holotype male; (8) paratype female. Terminal abdominal segments, dorsal: (9) holotype male; (10) paratype female. Terminal abdominal segments, lateral: (11) holotype male; (12) paratype female." httpUri="https://zenodo.org/record/1154482/files/figure.png" pageId="5" pageNumber="278">Fig. 7</figureCitation>
). The males of the 
<typeStatus id="54C2F6F2FFD1FFDB4341FE19FE79FE3A" box="[381,429,477,501]" pageId="5" pageNumber="278">type</typeStatus>
series at least are also separated from all other species in the 
<taxonomicName id="4C7933D3FFD1FFDB4666FE18FB0BFE3A" baseAuthorityName="Laidlaw" baseAuthorityYear="1915" box="[1114,1247,476,501]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="carmichaeli">
<emphasis id="B90D9442FFD1FFDB4666FE18FB0BFE3A" box="[1114,1247,476,501]" italics="true" pageId="5" pageNumber="278">carmichaeli</emphasis>
</taxonomicName>
-group except 
<taxonomicName id="4C7933D3FFD1FFDB47BDFE19FED5FDD7" baseAuthorityName="Tinkham" baseAuthorityYear="1938" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="brownelli">
<emphasis id="B90D9442FFD1FFDB47BDFE19FED5FDD7" italics="true" pageId="5" pageNumber="278">D. brownelli</emphasis>
</taxonomicName>
by the mostly pale coloured anterior lobe of the prothorax (
<figureCitation id="134254D5FFD1FFDB4196FE3BFC3AFDD7" box="[938,1006,511,536]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1969,1992]" captionTargetBox="[151,1435,450,1948]" captionTargetId="figure@2.[151,1435,450,1948]" captionTargetPageId="2" captionText="FIGURES 1–6. Drepanosticta emtrai. Head: (1) holotype male; (2) paratype female.Prothorax, dorsal: (3) holotype male; (4) paratype female. Prothorax lateral:(5) holotype male; (6) paratype female." httpUri="https://zenodo.org/record/1154480/files/figure.png" pageId="5" pageNumber="278">Fig. 3</figureCitation>
), which is either entirely dark or with only limited lateral pale marks in the other species. 
<taxonomicName id="4C7933D3FFD1FFDB40E9FDE1FC22FDF3" authorityName="Asahina" authorityYear="1997" box="[725,1014,549,572]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="vietnamica">
<emphasis id="B90D9442FFD1FFDB40E9FDE1FC22FDF3" box="[725,1014,549,572]" italics="true" pageId="5" pageNumber="278">Drepanosticta vietnamica</emphasis>
</taxonomicName>
has a much larger, better defined pale stripe on the metepisternum (
<figureCitation id="134254D5FFD1FFDB43E2FD83FDE5FDAF" box="[478,561,583,608]" captionStart="FIGURES 17–20" captionStartId="6.[151,264,1859,1882]" captionTargetBox="[151,1435,1054,1838]" captionTargetId="figure@6.[151,1435,1054,1838]" captionTargetPageId="6" captionText="FIGURES 17–20. Drepanosticta vietnamica paratype: (17) head and thorax; (18) anal appendages, lateral view; (19) left cercus, dorsal view; (20) left paraproct,oblique dorsal view." httpUri="https://zenodo.org/record/1154486/files/figure.png" pageId="5" pageNumber="278">Fig. 17</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3631BDBFFD1FFDB42FBFDA8FD0EF8CB" pageId="5" pageNumber="278" type="discussion">
<paragraph id="8BC64850FFD1FFDB42FBFDA8FD9EFD03" blockId="5.[151,1437,151,1977]" pageId="5" pageNumber="278">
<emphasis id="B90D9442FFD1FFDB42FBFDA8FEEFFD4A" bold="true" box="[199,315,620,645]" pageId="5" pageNumber="278">Remarks.</emphasis>
The 
<typeStatus id="54C2F6F2FFD1FFDB4345FDA9FE7DFD4A" box="[377,425,621,645]" pageId="5" pageNumber="278">type</typeStatus>
series comes from three separated locations in 
<collectingCountry id="F36E08C0FFD1FFDB41ECFDA8FBE6FD4A" box="[976,1074,620,645]" name="Vietnam" pageId="5" pageNumber="278">Vietnam</collectingCountry>
, one widely separated from the others, and one location in 
<collectingCountry id="F36E08C0FFD1FFDB43E6FD54FDC0FD68" box="[474,532,656,679]" name="Laos" pageId="5" pageNumber="278">Laos</collectingCountry>
; although only a limited number of specimens has been observed so far, this indicates a rather large range (
<figureCitation id="134254D5FFD1FFDB43D5FD70FDE9FD03" box="[489,573,692,717]" captionStart="FIGURE 21" captionStartId="7.[151,250,1933,1955]" captionTargetBox="[151,1435,663,1910]" captionTargetId="figure@7.[151,1435,663,1910]" captionTargetPageId="7" captionText="FIGURE 21. Distributions of Drepanosticta emtrai and D. vietnamica. Yellow circles: D. emtrai type series; red square: specimen excluded from the type series of D. emtrai (see remarks section under D. emtrai); black circle: D. vietnamica (approximate position of type locality)." httpUri="https://zenodo.org/record/1154488/files/figure.png" pageId="5" pageNumber="278">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="8BC64850FFD1FFDB42FBFD13FC57FCB0" blockId="5.[151,1437,151,1977]" pageId="5" pageNumber="278">
The male from 
<collectingRegion id="49BD86B2FFD1FFDB434FFD1CFE0AFD3F" box="[371,478,727,752]" country="Vietnam" name="Hoa Binh" pageId="5" pageNumber="278">Hoa Binh</collectingRegion>
has been excluded from the 
<typeStatus id="54C2F6F2FFD1FFDB4121FD1CFC99FD3F" box="[797,845,728,752]" pageId="5" pageNumber="278">type</typeStatus>
series of 
<taxonomicName id="4C7933D3FFD1FFDB4184FD1DFBF7FD3F" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[952,1059,729,752]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD1FFDB4184FD1DFBF7FD3F" box="[952,1059,729,752]" italics="true" pageId="5" pageNumber="278">D. emtrai</emphasis>
</taxonomicName>
because of differences in the anal appendages compared with 
<taxonomicName id="4C7933D3FFD1FFDB43EEFD39FDEAFCDB" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[466,574,765,788]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD1FFDB43EEFD39FDEAFCDB" box="[466,574,765,788]" italics="true" pageId="5" pageNumber="278">D. emtrai</emphasis>
</taxonomicName>
: the ventral flap on the cerci, whilst prominent, is smaller than in 
<taxonomicName id="4C7933D3FFD1FFDB4713FD39FA48FCDB" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[1327,1436,765,788]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD1FFDB4713FD39FA48FCDB" box="[1327,1436,765,788]" italics="true" pageId="5" pageNumber="278">D. emtrai</emphasis>
</taxonomicName>
and there is only a single interior flap visible on the cerci in dorsal view. Although the 
<collectingRegion id="49BD86B2FFD1FFDB4641FCE4FB3EFCF7" box="[1149,1258,799,824]" country="Vietnam" name="Hoa Binh" pageId="5" pageNumber="278">Hoa Binh</collectingRegion>
male may well belong to 
<taxonomicName id="4C7933D3FFD1FFDB4334FC81FEA1FC93" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[264,373,837,860]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD1FFDB4334FC81FEA1FC93" box="[264,373,837,860]" italics="true" pageId="5" pageNumber="278">D. emtrai</emphasis>
</taxonomicName>
, there is also a small possibility that it is a distinct species. It is slightly larger than any of the 
<typeStatus id="54C2F6F2FFD1FFDB42ABFCACFED3FC4F" box="[151,263,872,896]" pageId="5" pageNumber="278" type="paratype">paratypes</typeStatus>
: abdomen with anal appendages 
<quantity id="4C81E5B5FFD1FFDB404BFCA3FD09FCB0" box="[631,733,871,896]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" pageId="5" pageNumber="278" unit="mm" value="37.5">37.5 mm</quantity>
, Hw 
<quantity id="4C81E5B5FFD1FFDB4124FCA3FCABFCB0" box="[792,895,871,896]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="5" pageNumber="278" unit="mm" value="23.5">23.5 mm</quantity>
.
</paragraph>
<paragraph id="8BC64850FFD1FFDB42FBFC49FD0EF8CB" blockId="5.[151,1437,151,1977]" pageId="5" pageNumber="278">
<taxonomicName id="4C7933D3FFD1FFDB42FBFC49FE6CFC6B" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[199,440,909,932]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD1FFDB42FBFC49FE6CFC6B" box="[199,440,909,932]" italics="true" pageId="5" pageNumber="278">Drepanosticta emtrai</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C7933D3FFD1FFDB406FFC49FD26FC6B" authorityName="Asahina" authorityYear="1997" box="[595,754,909,932]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="vietnamica">
<emphasis id="B90D9442FFD1FFDB406FFC49FD26FC6B" box="[595,754,909,932]" italics="true" pageId="5" pageNumber="278">D. vietnamica</emphasis>
</taxonomicName>
as judged from Asahina’s illustrations and description, and from the available 
<typeStatus id="54C2F6F2FFD1FFDB4341FC74FE34FC07" box="[381,480,944,968]" pageId="5" pageNumber="278" type="paratype">paratype</typeStatus>
, in a number of respects. Both Asahina’s illustration and 
<figureCitation id="134254D5FFD1FFDB469FFC6BFAC2FC07" box="[1187,1302,943,968]" captionStart="FIGURES 17–20" captionStartId="6.[151,264,1859,1882]" captionTargetBox="[151,1435,1054,1838]" captionTargetId="figure@6.[151,1435,1054,1838]" captionTargetPageId="6" captionText="FIGURES 17–20. Drepanosticta vietnamica paratype: (17) head and thorax; (18) anal appendages, lateral view; (19) left cercus, dorsal view; (20) left paraproct,oblique dorsal view." httpUri="https://zenodo.org/record/1154486/files/figure.png" pageId="5" pageNumber="278">Figure 20</figureCitation>
here show paraprocts only gently inwards curved in their apical half, but they are strongly curved in their apical half in 
<taxonomicName id="4C7933D3FFD1FFDB47BDFC11FF34FBDF" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD1FFDB47BDFC11FA41FC23" box="[1409,1429,981,1004]" italics="true" pageId="5" pageNumber="278">D</emphasis>
. 
<emphasis id="B90D9442FFD1FFDB42ABFC3DFF34FBDF" box="[151,224,1017,1040]" italics="true" pageId="5" pageNumber="278">emtrai</emphasis>
</taxonomicName>
. The cerci appear to have a ventral flap in Asahina’s illustration, but it appears less well developed than that in 
<taxonomicName id="4C7933D3FFD1FFDB428FFBD9FECAFBFB" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[179,286,1053,1076]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD1FFDB428FFBD9FECAFBFB" box="[179,286,1053,1076]" italics="true" pageId="5" pageNumber="278">D. emtrai</emphasis>
</taxonomicName>
; but in the 
<typeStatus id="54C2F6F2FFD1FFDB43A1FBD9FE29FBFA" box="[413,509,1053,1077]" pageId="5" pageNumber="278" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C7933D3FFD1FFDB401FFBD9FD6BFBFB" authorityName="Asahina" authorityYear="1997" box="[547,703,1053,1076]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="vietnamica">
<emphasis id="B90D9442FFD1FFDB401FFBD9FD6BFBFB" box="[547,703,1053,1076]" italics="true" pageId="5" pageNumber="278">D. vietnamica</emphasis>
</taxonomicName>
there is only a small inward directed protrusion in approximately the same position, visible in dorsal view (
<figureCitation id="134254D5FFD1FFDB4059FBFBFD62FB97" box="[613,694,1087,1112]" captionStart="FIGURES 17–20" captionStartId="6.[151,264,1859,1882]" captionTargetBox="[151,1435,1054,1838]" captionTargetId="figure@6.[151,1435,1054,1838]" captionTargetPageId="6" captionText="FIGURES 17–20. Drepanosticta vietnamica paratype: (17) head and thorax; (18) anal appendages, lateral view; (19) left cercus, dorsal view; (20) left paraproct,oblique dorsal view." httpUri="https://zenodo.org/record/1154486/files/figure.png" pageId="5" pageNumber="278">Fig. 19</figureCitation>
), but not in lateral view (
<figureCitation id="134254D5FFD1FFDB41F1FBFBFBF4FB97" box="[973,1056,1087,1112]" captionStart="FIGURES 17–20" captionStartId="6.[151,264,1859,1882]" captionTargetBox="[151,1435,1054,1838]" captionTargetId="figure@6.[151,1435,1054,1838]" captionTargetPageId="6" captionText="FIGURES 17–20. Drepanosticta vietnamica paratype: (17) head and thorax; (18) anal appendages, lateral view; (19) left cercus, dorsal view; (20) left paraproct,oblique dorsal view." httpUri="https://zenodo.org/record/1154486/files/figure.png" pageId="5" pageNumber="278">Fig. 18</figureCitation>
). The tips of the cerci are rounded in 
<taxonomicName id="4C7933D3FFD1FFDB428AFBA1FEF0FBB3" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[182,292,1125,1148]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD1FFDB428AFBA1FEF0FBB3" box="[182,292,1125,1148]" italics="true" pageId="5" pageNumber="278">D. emtrai</emphasis>
</taxonomicName>
but appear almost hooked down in Asahina’s illustration of 
<taxonomicName id="4C7933D3FFD1FFDB41DDFBA1FB54FBB3" authorityName="Asahina" authorityYear="1997" box="[993,1152,1125,1148]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="vietnamica">
<emphasis id="B90D9442FFD1FFDB41DDFBA1FB54FBB3" box="[993,1152,1125,1148]" italics="true" pageId="5" pageNumber="278">D. vietnamica</emphasis>
</taxonomicName>
; they have a completely different appearance in the 
<typeStatus id="54C2F6F2FFD1FFDB43F6FB4DFDFEFB6E" box="[458,554,1161,1185]" pageId="5" pageNumber="278" type="paratype">paratype</typeStatus>
(
<figureCitation id="134254D5FFD1FFDB4006FB43FD59FB6F" box="[570,653,1159,1184]" captionStart="FIGURES 17–20" captionStartId="6.[151,264,1859,1882]" captionTargetBox="[151,1435,1054,1838]" captionTargetId="figure@6.[151,1435,1054,1838]" captionTargetPageId="6" captionText="FIGURES 17–20. Drepanosticta vietnamica paratype: (17) head and thorax; (18) anal appendages, lateral view; (19) left cercus, dorsal view; (20) left paraproct,oblique dorsal view." httpUri="https://zenodo.org/record/1154486/files/figure.png" pageId="5" pageNumber="278">Fig. 18</figureCitation>
). In 
<taxonomicName id="4C7933D3FFD1FFDB40FCFB4DFC8AFB6F" authorityName="Asahina" authorityYear="1997" box="[704,862,1161,1184]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="vietnamica">
<emphasis id="B90D9442FFD1FFDB40FCFB4DFC8AFB6F" box="[704,862,1161,1184]" italics="true" pageId="5" pageNumber="278">D. vietnamica</emphasis>
</taxonomicName>
a small but well developed dorsal projection at the point where the upper margin of the cerci turns downward is visible in Asahina’s illustration, a projection is also visible in the 
<typeStatus id="54C2F6F2FFD1FFDB430CFB15FE44FB26" box="[304,400,1233,1257]" pageId="5" pageNumber="278" type="paratype">paratype</typeStatus>
(
<figureCitation id="134254D5FFD1FFDB439CFB0BFE27FB27" box="[416,499,1231,1256]" captionStart="FIGURES 17–20" captionStartId="6.[151,264,1859,1882]" captionTargetBox="[151,1435,1054,1838]" captionTargetId="figure@6.[151,1435,1054,1838]" captionTargetPageId="6" captionText="FIGURES 17–20. Drepanosticta vietnamica paratype: (17) head and thorax; (18) anal appendages, lateral view; (19) left cercus, dorsal view; (20) left paraproct,oblique dorsal view." httpUri="https://zenodo.org/record/1154486/files/figure.png" pageId="5" pageNumber="278">Fig. 18</figureCitation>
), although it is more easily seen in dorsal view (
<figureCitation id="134254D5FFD1FFDB4629FB0BFBB2FB27" box="[1045,1126,1231,1256]" captionStart="FIGURES 17–20" captionStartId="6.[151,264,1859,1882]" captionTargetBox="[151,1435,1054,1838]" captionTargetId="figure@6.[151,1435,1054,1838]" captionTargetPageId="6" captionText="FIGURES 17–20. Drepanosticta vietnamica paratype: (17) head and thorax; (18) anal appendages, lateral view; (19) left cercus, dorsal view; (20) left paraproct,oblique dorsal view." httpUri="https://zenodo.org/record/1154486/files/figure.png" pageId="5" pageNumber="278">Fig. 19</figureCitation>
) where it appears as a flap; but in the same lateral and slightly dorsal view, no structure in the same position is visible in 
<taxonomicName id="4C7933D3FFD1FFDB4680FB31FAFDFAC3" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[1212,1321,1269,1292]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD1FFDB4680FB31FAFDFAC3" box="[1212,1321,1269,1292]" italics="true" pageId="5" pageNumber="278">D. emtrai</emphasis>
</taxonomicName>
, but there are two inwards directed, shallow, rounded flaps present on the upper inner margin of the cerci, visible in dorsal view (
<figureCitation id="134254D5FFD1FFDB42E1FAF8FEE4FA9B" box="[221,304,1340,1365]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1061,1084]" captionTargetBox="[150,1412,193,1026]" captionTargetId="figure@4.[150,1436,193,1041]" captionTargetPageId="4" captionText="FIGURES 13–16. Drepanosticta emtrai. Anal appendages and genital ligula of holotype male. Anal appendages: (13) dorsal; (14) lateral; (15) left paraproct in ventral view.(16) Genitalligula, ventral view." httpUri="https://zenodo.org/record/1154484/files/figure.png" pageId="5" pageNumber="278">Fig. 13</figureCitation>
). In 
<taxonomicName id="4C7933D3FFD1FFDB435FFAF9FDD4FA9B" authorityName="Asahina" authorityYear="1997" box="[355,512,1341,1364]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="vietnamica">
<emphasis id="B90D9442FFD1FFDB435FFAF9FDD4FA9B" box="[355,512,1341,1364]" italics="true" pageId="5" pageNumber="278">D. vietnamica</emphasis>
</taxonomicName>
the anterior lobe of the prothorax is stated to be “black with ambiguous pale spots laterally”, figure 
<quantity id="4C81E5B5FFD1FFDB4367FAA4FE53FAB7" box="[347,391,1375,1400]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="278" unit="in" value="2.0">2 in</quantity>
Asahina (1997) also shows what appear to be pale marks on the middle lobe and these are also present in the 
<typeStatus id="54C2F6F2FFD1FFDB4353FA41FE04FA52" box="[367,464,1413,1437]" pageId="5" pageNumber="278" type="paratype">paratype</typeStatus>
(
<figureCitation id="134254D5FFD1FFDB43DDFA40FDE2FA53" box="[481,566,1412,1437]" captionStart="FIGURES 17–20" captionStartId="6.[151,264,1859,1882]" captionTargetBox="[151,1435,1054,1838]" captionTargetId="figure@6.[151,1435,1054,1838]" captionTargetPageId="6" captionText="FIGURES 17–20. Drepanosticta vietnamica paratype: (17) head and thorax; (18) anal appendages, lateral view; (19) left cercus, dorsal view; (20) left paraproct,oblique dorsal view." httpUri="https://zenodo.org/record/1154486/files/figure.png" pageId="5" pageNumber="278">Fig. 17</figureCitation>
); the markings on the anterior lobe in the 
<typeStatus id="54C2F6F2FFD1FFDB4626FA41FB9EFA52" box="[1050,1098,1413,1437]" pageId="5" pageNumber="278">type</typeStatus>
series of 
<taxonomicName id="4C7933D3FFD1FFDB4680FA41FAFCFA53" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[1212,1320,1413,1436]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD1FFDB4680FA41FAFCFA53" box="[1212,1320,1413,1436]" italics="true" pageId="5" pageNumber="278">D. emtrai</emphasis>
</taxonomicName>
are better defined and more extensive than in 
<taxonomicName id="4C7933D3FFD1FFDB4010FA6DFD1FFA0F" authorityName="Asahina" authorityYear="1997" box="[556,715,1449,1472]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="vietnamica">
<emphasis id="B90D9442FFD1FFDB4010FA6DFD1FFA0F" box="[556,715,1449,1472]" italics="true" pageId="5" pageNumber="278">D. vietnamica</emphasis>
</taxonomicName>
. The pale stripe on the metepisternum of 
<taxonomicName id="4C7933D3FFD1FFDB469AFA6DFA90FA0F" authorityName="Asahina" authorityYear="1997" box="[1190,1348,1449,1472]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="vietnamica">
<emphasis id="B90D9442FFD1FFDB469AFA6DFA90FA0F" box="[1190,1348,1449,1472]" italics="true" pageId="5" pageNumber="278">D. vietnamica</emphasis>
</taxonomicName>
in both Asahina’s illustration and the examined 
<typeStatus id="54C2F6F2FFD1FFDB4065FA09FD6DFA2A" box="[601,697,1485,1509]" pageId="5" pageNumber="278" type="paratype">paratype</typeStatus>
(
<figureCitation id="134254D5FFD1FFDB40F4FA08FCCDFA2B" box="[712,793,1484,1509]" captionStart="FIGURES 17–20" captionStartId="6.[151,264,1859,1882]" captionTargetBox="[151,1435,1054,1838]" captionTargetId="figure@6.[151,1435,1054,1838]" captionTargetPageId="6" captionText="FIGURES 17–20. Drepanosticta vietnamica paratype: (17) head and thorax; (18) anal appendages, lateral view; (19) left cercus, dorsal view; (20) left paraproct,oblique dorsal view." httpUri="https://zenodo.org/record/1154486/files/figure.png" pageId="5" pageNumber="278">Fig. 17</figureCitation>
) is longer and much better defined that in 
<taxonomicName id="4C7933D3FFD1FFDB46C8FA09FA8BFA2B" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[1268,1375,1485,1508]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD1FFDB46C8FA09FA8BFA2B" box="[1268,1375,1485,1508]" italics="true" pageId="5" pageNumber="278">D. emtrai</emphasis>
</taxonomicName>
(
<figureCitation id="134254D5FFD1FFDB4752FA08FF73F9C7" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1376,1399]" captionTargetBox="[151,1435,193,1355]" captionTargetId="figure@3.[151,1435,193,1355]" captionTargetPageId="3" captionText="FIGURES 7–12. Drepanosticta emtrai. Synthorax, lateral: (7) holotype male; (8) paratype female. Terminal abdominal segments, dorsal: (9) holotype male; (10) paratype female. Terminal abdominal segments, lateral: (11) holotype male; (12) paratype female." httpUri="https://zenodo.org/record/1154482/files/figure.png" pageId="5" pageNumber="278">Fig. 7</figureCitation>
). Differences in the colour of parts of the head and the terminal abdominal segments are likely to be due to the preservation methods employed and different ages of the specimens, so are not considered to be reliable here. The 
<typeStatus id="54C2F6F2FFD1FFDB42ABF9FCFF13F99F" box="[151,199,1592,1616]" pageId="5" pageNumber="278">type</typeStatus>
series of 
<taxonomicName id="4C7933D3FFD1FFDB4306F9FDFE73F99F" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[314,423,1593,1616]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD1FFDB4306F9FDFE73F99F" box="[314,423,1593,1616]" italics="true" pageId="5" pageNumber="278">D. emtrai</emphasis>
</taxonomicName>
comes from a large geographical area in 
<collectingCountry id="F36E08C0FFD1FFDB41B4F9F3FC33F99F" box="[904,999,1591,1616]" name="Vietnam" pageId="5" pageNumber="278">Vietnam</collectingCountry>
from 
<collectingRegion id="49BD86B2FFD1FFDB460CF9FCFB5AF99F" box="[1072,1166,1591,1616]" country="Vietnam" name="Ha Tinh" pageId="5" pageNumber="278">Ha Tinh</collectingRegion>
in central 
<collectingCountry id="F36E08C0FFD1FFDB4730F9F3FABFF99F" box="[1292,1387,1591,1616]" name="Vietnam" pageId="5" pageNumber="278">Vietnam</collectingCountry>
and nearby 
<collectingRegion id="49BD86B2FFD1FFDB42D3F998FE8DF9BB" box="[239,345,1628,1653]" country="Vietnam" name="Nghe An" pageId="5" pageNumber="278">Nghe An</collectingRegion>
and 
<collectingCountry id="F36E08C0FFD1FFDB43A6F999FE07F9BB" box="[410,467,1629,1652]" name="Laos" pageId="5" pageNumber="278">Laos</collectingCountry>
, to Ba Vi Mountain in northern 
<collectingCountry id="F36E08C0FFD1FFDB4160F998FC6AF9BA" box="[860,958,1628,1653]" name="Vietnam" pageId="5" pageNumber="278">Vietnam</collectingCountry>
. Although the 
<typeStatus id="54C2F6F2FFD1FFDB464EF999FB76F9BA" box="[1138,1186,1629,1653]" pageId="5" pageNumber="278">type</typeStatus>
series displays some variation in outward appearance, notably the extent of the pale marking on the metepisternum and the blue area of S8, the observed variability does not come close to the pattern of 
<taxonomicName id="4C7933D3FFD1FFDB41B6F961FBFCF973" authorityName="Asahina" authorityYear="1997" box="[906,1064,1701,1724]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="vietnamica">
<emphasis id="B90D9442FFD1FFDB41B6F961FBFCF973" box="[906,1064,1701,1724]" italics="true" pageId="5" pageNumber="278">D. vietnamica</emphasis>
</taxonomicName>
and the shape of paraprocts and cerci is much more constant. Nevertheless, variation cannot be completely discounted as a factor in differences in the markings between 
<taxonomicName id="4C7933D3FFD1FFDB43A8F929FDE5F8CB" authorityName="Asahina" authorityYear="1997" box="[404,561,1773,1796]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="vietnamica">
<emphasis id="B90D9442FFD1FFDB43A8F929FDE5F8CB" box="[404,561,1773,1796]" italics="true" pageId="5" pageNumber="278">D. vietnamica</emphasis>
</taxonomicName>
and 
<emphasis id="B90D9442FFD1FFDB405BF929FD0EF8CB" box="[615,730,1773,1796]" italics="true" pageId="5" pageNumber="278">
<taxonomicName id="4C7933D3FFD1FFDB405BF929FD02F8CB" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[615,726,1773,1796]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="emtrai">D. emtrai</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3631BDBFFD1FFDB42FBF8D5FBA0F877" pageId="5" pageNumber="278" type="description">
<paragraph id="8BC64850FFD1FFDB42FBF8D5FBA0F877" blockId="5.[151,1437,151,1977]" pageId="5" pageNumber="278">
<taxonomicName id="4C7933D3FFD1FFDB42FBF8D5FE6EF8E7" authorityName="Dow &amp; Kompier &amp; Phan" authorityYear="2018" box="[199,442,1809,1832]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="emtrai">
<emphasis id="B90D9442FFD1FFDB42FBF8D5FE6EF8E7" box="[199,442,1809,1832]" italics="true" pageId="5" pageNumber="278">Drepanosticta emtrai</emphasis>
</taxonomicName>
was found in 
<collectingRegion id="49BD86B2FFD1FFDB4054F8D4FD13F8E7" box="[616,711,1807,1832]" country="Vietnam" name="Ha Tinh" pageId="5" pageNumber="278">Ha Tinh</collectingRegion>
and 
<collectingRegion id="49BD86B2FFD1FFDB4139F8CBFCBAF8E7" box="[773,878,1807,1832]" country="Vietnam" name="Nghe An" pageId="5" pageNumber="278">Nghe An</collectingRegion>
provinces in logged mature forest at seeps that possibly dry up for part of the year. On Ba Vi Mountain, 
<collectingRegion id="49BD86B2FFD1FFDB4175F8F0FC44F882" box="[841,912,1844,1869]" country="Vietnam" name="Ha Noi" pageId="5" pageNumber="278">Hanoi</collectingRegion>
, it occurs at seeps under dense forest cover, interestingly preferring almost identical, although possibly slightly less rocky, seeps than 
<taxonomicName id="4C7933D3FFD1FFDB46BDF89DFA9DF8BF" authorityName="Wilson" authorityYear="1997" box="[1153,1353,1879,1904]" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="hongkongensis">
<emphasis id="B90D9442FFD1FFDB46BDF89DFA9DF8BF" box="[1153,1353,1879,1904]" italics="true" pageId="5" pageNumber="278">D. hongkongensis</emphasis>
</taxonomicName>
, which occurs in its close vicinity on the same mountainside. It appears to emerge a few weeks later than 
<taxonomicName id="4C7933D3FFD1FFDB47BDF8B9FEE9F876" authorityName="Wilson" authorityYear="1997" class="Insecta" family="Platystictidae" genus="Drepanosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="hongkongensis">
<emphasis id="B90D9442FFD1FFDB47BDF8B9FEE9F876" italics="true" pageId="5" pageNumber="278">D. hongkongensis</emphasis>
</taxonomicName>
, but at least on Ba Vi Mountain both species occur in the middle of June.
</paragraph>
</subSubSection>
</treatment>
</document>