343 lines
40 KiB
XML
343 lines
40 KiB
XML
<document id="4C3E8976C5811FF58E83EB89FB2EA732" ID-DOI="10.1080/00222933.2024.2435550" ID-ISSN="1464-5262" ID-Zenodo-Dep="14764931" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1738159620067" checkinUser="plazi" docAuthor="Chen, Zhong-Guang, Zhou, Yi, Dai, Yu-Ting, Wu, Xiao-Ping & Ouyang, Shan" docDate="2024" docId="03CCBD10FF92E055313FFCF3E01BF99C" docLanguage="en" docName="JNATHIST.59.1-4.11-28.pdf" docOrigin="Journal of Natural History 59 (1 - 4)" docSource="https://doi.org/10.1080/00222933.2024.2435550" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Petraeomastus gigas Chen, Dai, Wu and Ouyang 2024, sp. n." docType="treatment" docVersion="1" lastPageNumber="26" masterDocId="FFF5C568FF9EE05A3001FFC6E26AFF8F" masterDocTitle="A review of three taxonomically confused species of Petraeomastus Möllendorff, 1901 (Gastropoda: Stylommatophora: Enidae) from the Jinshajiang River Valley, with the description of a new species" masterLastPageNumber="28" masterPageNumber="11" pageNumber="23" updateTime="1738351953008" updateUser="guilherme" zenodo-license-document="CLOSED">
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<mods:title id="19DFB77A1E712779669FF206D94841B3">A review of three taxonomically confused species of Petraeomastus Möllendorff, 1901 (Gastropoda: Stylommatophora: Enidae) from the Jinshajiang River Valley, with the description of a new species</mods:title>
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<mods:roleTerm id="7BF92EBAF653005E8E1DAE1E5353DDAB">Author</mods:roleTerm>
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<mods:namePart id="7448687B98965B31E4C1AD08E657E662">Chen, Zhong-Guang</mods:namePart>
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<mods:namePart id="9A89DCC8118B4DD33A6DE0A6F4AF9314">Zhou, Yi</mods:namePart>
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<mods:roleTerm id="875B2A5DA3EDEB7E92E1105584A494F1">Author</mods:roleTerm>
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<mods:namePart id="2C05B4814FF61D2156AD9CCBB8FF0CAB">Dai, Yu-Ting</mods:namePart>
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<mods:namePart id="5DEAE006FE1BC8D2510BA9DD0205346E">Wu, Xiao-Ping</mods:namePart>
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<mods:namePart id="FB44D39F6544074B18C50D0F88132E63">Ouyang, Shan</mods:namePart>
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<mods:title id="D60ED8298220FF58A4FB79A2A58CBCAA">Journal of Natural History</mods:title>
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<mods:part id="F4A05041A938DA7685187E5C988A619F">
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<mods:date id="4163FB61595D64C1A1AD61B6002977D1">2024</mods:date>
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<mods:number id="2946F8BBF22566EA25E32F3578694FD6">2024-12-17</mods:number>
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<mods:number id="72C087D4E270A4AAF46BAB7EC7F4E88F">59</mods:number>
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<mods:number id="6BFB0D5639F086CAB6FAC510ECBA4C49">1 - 4</mods:number>
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<mods:url id="C37CF31F482A4F064EC69FC9F8F90298">https://doi.org/10.1080/00222933.2024.2435550</mods:url>
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<mods:classification id="9655AB6CB96939B1AB7C983F02F008C4">journal article</mods:classification>
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<mods:identifier id="F796B7B1041581C9224629BE6A463922" type="DOI">10.1080/00222933.2024.2435550</mods:identifier>
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<subSubSection id="C37F5F8DFF92E056313FFCF3E18CFCC0" box="[318,998,821,847]" pageId="12" pageNumber="23" type="nomenclature">
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<emphasis id="B911D014FF92E056313FFCF3E18CFCC0" bold="true" box="[318,998,821,847]" pageId="12" pageNumber="23">
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<taxonomicName id="4C657785FF92E056313FFCF3E1F0FCC0" ID-CoL="CRCCV" authority="Chen, Dai, Wu and Ouyang" authorityName="Chen, Dai, Wu and Ouyang" authorityYear="2024" box="[318,922,821,847]" class="Gastropoda" family="Enidae" genus="Petraeomastus" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="23" phylum="Mollusca" rank="species" species="gigas" status="sp. nov.">
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<emphasis id="B911D014FF92E056313FFCF3E054FCC0" bold="true" box="[318,574,821,847]" italics="true" pageId="12" pageNumber="23">Petraeomastus gigas</emphasis>
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Chen, Dai, Wu and Ouyang
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</taxonomicName>
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<taxonomicNameLabel id="A2226D6FFF92E05633A3FCF3E18CFCC0" box="[930,998,821,847]" pageId="12" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="C37F5F8DFF92E05630C1FCACE17FFC3E" pageId="12" pageNumber="23" type="description">
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ZooBank identifier
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urn:lsid:zoobank.org:act:
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(
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<figureCitation id="135E1083FF92E0563216FC51E0E0FC3E" box="[535,650,919,945]" captionStart="Figure 4" captionStartId="6.[160,225,1659,1682]" captionTargetBox="[164,1152,1133,1627]" captionTargetId="figure-283@6.[163,1153,1132,1628]" captionTargetPageId="6" captionText="Figure 4. Adjacent environment and habitat of living individuals of Petraeomastus land snails. (A&B) Environment of the Jinshajiang River Valley, left side: Batang County, right side: Mangkang County; (C) living specimens of P. neumayri; (D) living specimens of P. desgodinsi; (E) living specimens of P. markamensis; (F) living specimens of Petraeomastus gigas sp. n. and P. markamensis." figureDoi="http://doi.org/10.5281/zenodo.14764939" httpUri="https://zenodo.org/record/14764939/files/figure.png" pageId="12" pageNumber="23">Figures 4</figureCitation>
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(F), 7(A), 8)
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<typeStatus id="54DEB2A4FF92E05630A1FC29E2B2FB86" box="[160,216,1007,1033]" pageId="12" pageNumber="23">Type</typeStatus>
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material
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</paragraph>
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<paragraph id="8BDA0C06FF92E05630A1FBD7E332FB1C" blockId="12.[160,1156,1007,1171]" pageId="12" pageNumber="23">
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<materialsCitation id="3B0D065BFF92E05630A1FBD7E339FB1C" collectingDate="2023-08-17" collectionCode="ZMNH" collectorName="Zhong-Guang Chen & Meng-Hua Li & Jin-Sheng Mou" country="China" county="Batang County" latitude="29.871" location="Langdaquka" longLatPrecision="73" longitude="99.041" municipality="Ganzi Tibetan Autonomous Prefecture" pageId="12" pageNumber="23" specimenCode="ZMNH2409001" specimenCount="1" stateProvince="Sichuan Province" typeStatus="holotype">
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<typeStatus id="54DEB2A4FF92E05630A1FBD7E37AFBA4" box="[160,272,1041,1067]" pageId="12" pageNumber="23" type="holotype">Holotype</typeStatus>
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.
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||
</emphasis>
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||
<specimenCode id="DBC3A47DFF92E0563124FBD4E3B9FBA4" box="[293,467,1042,1067]" pageId="12" pageNumber="23">ZMNH2409001</specimenCode>
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,
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<location id="8EBA5ADDFF92E05631E5FBD4E018FBA4" LSID="urn:lsid:plazi:treatment:03CCBD10FF92E055313FFCF3E01BF99C:8EBA5ADDFF92E05631E5FBD4E018FBA4" box="[484,626,1042,1067]" country="China" county="Batang County" latitude="29.871" longLatPrecision="73" longitude="99.041" municipality="Ganzi Tibetan Autonomous Prefecture" name="Langdaquka" pageId="12" pageNumber="23" stateProvince="Sichuan Province">Langdaquka</location>
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||
[朗达曲卡],
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||
<collectingCounty id="62BB748AFF92E056330EFBD4E1ABFBA4" box="[783,961,1042,1067]" pageId="12" pageNumber="23">Batang County</collectingCounty>
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||
[巴塘县],
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||
<collectingMunicipality id="6BBE967CFF92E0563442FBD4E079FBC1" pageId="12" pageNumber="23">Ganzi Tibetan Autonomous Prefecture</collectingMunicipality>
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||
[甘孜藏族自治州],
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||
<collectingRegion id="49A1C2E4FF92E05632FAFBF3E1AAFBC1" box="[763,960,1077,1102]" country="China" name="Sichuan" pageId="12" pageNumber="23">Sichuan Province</collectingRegion>
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||
[四川省],
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||
<collectingCountry id="F3724C96FF92E056343CFBF3E6EAFBC1" box="[1085,1152,1077,1102]" name="China" pageId="12" pageNumber="23">China</collectingCountry>
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,
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,
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<geoCoordinate id="EE516AC1FF92E056310FFB9EE31FFBFE" box="[270,373,1112,1137]" degrees="29.871" direction="north" orientation="latitude" pageId="12" pageNumber="23" precision="55" value="29.871">29.871°N</geoCoordinate>
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, collected by
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<collectorName id="269069D0FF92E0563214FB9EE092FBFE" box="[533,760,1112,1137]" pageId="12" pageNumber="23">Zhong-Guang Chen</collectorName>
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,
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<collectorName id="269069D0FF92E0563305FB9EE1FCFBFE" box="[772,918,1112,1137]" pageId="12" pageNumber="23">Meng-Hua Li</collectorName>
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and
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||
<collectorName id="269069D0FF92E05633D2FB9EE6EAFBFE" box="[979,1152,1112,1137]" pageId="12" pageNumber="23">Jin-Sheng Mou</collectorName>
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,
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<date id="FFDB2AC6FF92E05630A1FBBCE339FB1C" box="[160,339,1146,1171]" pageId="12" pageNumber="23" value="2023-08-17">
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<collectingDate id="EF9FD32EFF92E05630A1FBBCE339FB1C" box="[160,339,1146,1171]" pageId="12" pageNumber="23" value="2023-08-17">17 August 2023</collectingDate>
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</date>
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</materialsCitation>
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.
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||
</paragraph>
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<paragraph id="8BDA0C06FF92E05630A1FB06E067FAEB" blockId="12.[160,1156,1216,1380]" pageId="12" pageNumber="23">
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<materialsCitation id="3B0D065BFF92E05630A1FB06E674FB56" box="[160,1054,1216,1242]" collectingDate="2023-08-17" collectionCode="NCUMB" collectorName="Zhong-Guang Chen & Meng-Hua Li & Jin-Sheng Mou" country="China" county="Batang County" latitude="29.871" location="Langdaquka" longLatPrecision="73" longitude="99.041" municipality="Ganzi Tibetan Autonomous Prefecture" pageId="12" pageNumber="23" specimenCode="NCUMB2409001-02" specimenCount="1" stateProvince="Sichuan Province" typeStatus="paratype">
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<emphasis id="B911D014FF92E05630A1FB06E377FB55" bold="true" box="[160,285,1216,1242]" italics="true" pageId="12" pageNumber="23">
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<typeStatus id="54DEB2A4FF92E05630A1FB06E372FB55" box="[160,280,1216,1242]" pageId="12" pageNumber="23" type="paratype">Paratypes</typeStatus>
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||
.
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||
</emphasis>
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||
n = 2,
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<specimenCode id="DBC3A47DFF92E0563183FB06E00FFB56" box="[386,613,1216,1241]" pageId="12" pageNumber="23">NCUMB2409001–02</specimenCode>
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||
, other information same as holotype
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B0D065BFF92E056342DFB06E063FAEB" collectingDate="2023-08-16" collectionCode="ZMNH" collectorName="Zhong-Guang Chen & Meng-Hua Li & Jin-Sheng Mou" country="China" county="Mangkang County" latitude="29.761" location="Jinshajiang River Bridge" longitude="99.0" municipality="Changdu City" pageId="12" pageNumber="23" specimenCode="ZMNH2409002" specimenCount="1" stateProvince="Xizang Autonomous Region" typeStatus="paratype">
|
||
n = 10,
|
||
<specimenCode id="DBC3A47DFF92E05630A1FB25E33CFB73" box="[160,342,1251,1276]" pageId="12" pageNumber="23">ZMNH2409002</specimenCode>
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–11,
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||
<quantity id="4C9DA1E3FF92E056318BFB25E3A3FB73" box="[394,457,1251,1276]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="23" unit="km" value="1.0">1 km</quantity>
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||
west of
|
||
<location id="8EBA5ADDFF92E0563240FB25E109FB73" LSID="urn:lsid:plazi:treatment:03CCBD10FF92E055313FFCF3E01BF99C:8EBA5ADDFF92E0563240FB25E109FB73" box="[577,867,1251,1276]" country="China" county="Mangkang County" longitude="99.0" municipality="Changdu City" name="Jinshajiang River Bridge" pageId="12" pageNumber="23" stateProvince="Xizang Autonomous Region">Jinshajiang River Bridge</location>
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||
[金沙江大桥以西
|
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<quantity id="4C9DA1E3FF92E0563437FB25E610FB73" box="[1078,1146,1251,1276]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="23" unit="km" value="1.0">
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<locationDeviation id="9A681524FF92E0563437FB25E610FB73" box="[1078,1146,1251,1276]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="23" unit="km" value="1.0">1 km</locationDeviation>
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</quantity>
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],
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<collectingCounty id="62BB748AFF92E05630A1FAC0E318FA90" box="[160,370,1286,1311]" pageId="12" pageNumber="23">Mangkang County</collectingCounty>
|
||
[芒康县],
|
||
<collectingMunicipality id="6BBE967CFF92E05631EBFAC0E0EFFA90" box="[490,645,1286,1311]" pageId="12" pageNumber="23">Changdu City</collectingMunicipality>
|
||
[昌都市],
|
||
<collectingRegion id="49A1C2E4FF92E05632FCFAC0E656FA90" box="[765,1084,1286,1311]" pageId="12" pageNumber="23">Xizang Autonomous Region</collectingRegion>
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||
[西藏 自治区],
|
||
<collectingCountry id="F3724C96FF92E0563104FAEEE322FACE" box="[261,328,1320,1345]" name="China" pageId="12" pageNumber="23">China</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE516AC1FF92E0563152FAEEE3DFFACE" box="[339,437,1320,1345]" degrees="99.000" direction="east" orientation="longitude" pageId="12" pageNumber="23" precision="55" value="99.0">99.000°E</geoCoordinate>
|
||
,
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||
<geoCoordinate id="EE516AC1FF92E05631C1FAEEE052FACE" box="[448,568,1320,1345]" degrees="29.761" direction="north" orientation="latitude" pageId="12" pageNumber="23" value="29.761">29.761°NN</geoCoordinate>
|
||
, collected by
|
||
<collectorName id="269069D0FF92E05632D5FAEEE1DFFACE" box="[724,949,1320,1345]" pageId="12" pageNumber="23">Zhong-Guang Chen</collectorName>
|
||
,
|
||
<collectorName id="269069D0FF92E05633C1FAEEE63BFACE" box="[960,1105,1320,1345]" pageId="12" pageNumber="23">Meng-Hua Li</collectorName>
|
||
and
|
||
<collectorName id="269069D0FF92E05630A1FA8DE321FAEB" box="[160,331,1355,1380]" pageId="12" pageNumber="23">Jin-Sheng Mou</collectorName>
|
||
,
|
||
<date id="FFDB2AC6FF92E0563154FA8DE063FAEB" box="[341,521,1355,1380]" pageId="12" pageNumber="23" value="2023-08-16">
|
||
<collectingDate id="EF9FD32EFF92E0563154FA8DE063FAEB" box="[341,521,1355,1380]" pageId="12" pageNumber="23" value="2023-08-16">16 August 2023</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F5F8DFF92E05630A1FA56E15CF9BA" pageId="12" pageNumber="23" type="diagnosis">
|
||
<paragraph id="8BDA0C06FF92E05630A1FA56E37EFA25" blockId="12.[160,1156,1424,1589]" box="[160,276,1424,1450]" pageId="12" pageNumber="23">
|
||
<emphasis id="B911D014FF92E05630A1FA56E37EFA25" bold="true" box="[160,276,1424,1450]" italics="true" pageId="12" pageNumber="23">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDA0C06FF92E05630A1FA72E15CF9BA" blockId="12.[160,1156,1424,1589]" pageId="12" pageNumber="23">
|
||
Shell large, height
|
||
<quantity id="4C9DA1E3FF92E0563174FA72E07DFA42" box="[373,535,1460,1485]" metricMagnitude="-2" metricUnit="m" metricValue="3.5200000000000005" metricValueMax="4.029999999999999" metricValueMin="3.0100000000000002" pageId="12" pageNumber="23" unit="mm" value="35.2" valueMax="40.3" valueMin="30.1">30.1–40.3 mm</quantity>
|
||
, dextral, width 11.8–13.0 mm. Post-nuclear teleoconch reddish brown with irregular white stripes; apex region reddish brown; umbilicus slit-like. Penial caecum present, subglobular. Bursa copulatrix ovoid, with stalk, well defined. Penial and appendicular retractor short. Habitat on shrubs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F5F8DFF92E05530A1F9A7E077FD5D" lastPageId="15" lastPageNumber="26" pageId="12" pageNumber="23" type="description">
|
||
<paragraph id="8BDA0C06FF92E05630A1F9A7E34CF9F4" blockId="12.[160,1156,1633,1728]" box="[160,294,1633,1659]" pageId="12" pageNumber="23">
|
||
<emphasis id="B911D014FF92E05630A1F9A7E34CF9F4" bold="true" box="[160,294,1633,1659]" italics="true" pageId="12" pageNumber="23">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDA0C06FF92E05730A1F942E3F1FA57" blockId="12.[160,1156,1633,1728]" lastBlockId="13.[160,1157,1193,1704]" lastPageId="13" lastPageNumber="24" pageId="12" pageNumber="23">
|
||
<emphasis id="B911D014FF92E05630A1F942E2B5F911" bold="true" box="[160,223,1668,1694]" italics="true" pageId="12" pageNumber="23">Shell.</emphasis>
|
||
(
|
||
<figureCitation id="135E1083FF92E05630F6F943E33EF911" box="[247,340,1669,1694]" captionStart="Figure 7" captionStartId="13.[161,226,1082,1105]" captionTargetBox="[164,1152,143,1050]" captionTargetId="figure-157@13.[163,1153,142,1051]" captionTargetPageId="13" captionText="Figure 7. Petraeomastus gigas sp. n. and its similar congener, P. heudeanus. (A) Petraeomastus gigas sp. n., holotype, ZMNH2409001; (B) P. heudeanus, 24_NCU_XPWU_AN144." figureDoi="http://doi.org/10.5281/zenodo.14764949" httpUri="https://zenodo.org/record/14764949/files/figure.png" pageId="12" pageNumber="23">Figure 7</figureCitation>
|
||
(A)) n = 13. Shell cylindrical-conic to conic, with apex abruptly pointed; shell most swollen at body whorl, dextral, large, thick, solid, opaque, sub-glossy, speckled, not spirally grooved; 7.5–8.5 whorls. Whorls rather flattened, not shouldered. Protoconch smooth, polished. Post-nuclear teleoconch smooth. Growth lines distinct. Suture shallow, without narrow band beneath it. Body whorl gradually ascending towards aperture, rounded at periphery. Aperture wavy, truncate-ovate, oblique, without tooth, with angular tubercle, completely adnate to body whorl. Peristome nearly connected, with a wide channel at middle insertion; white, thickened, expanded, not reflexed. Parietal callus distinct. Columellar margin reflexed. Umbilicus slit-shaped. Shell multicoloured, post-nuclear teleoconch with reddish brown background and irregular white stripes; apex region reddish brown.
|
||
</paragraph>
|
||
<caption id="DF1A5C8EFF93E05730A0FBFCE1EEFBE0" ID-DOI="http://doi.org/10.5281/zenodo.14764949" ID-Zenodo-Dep="14764949" httpUri="https://zenodo.org/record/14764949/files/figure.png" pageId="13" pageNumber="24" startId="13.[161,226,1082,1105]" targetBox="[164,1152,143,1050]" targetPageId="13" targetType="figure">
|
||
<paragraph id="8BDA0C06FF93E05730A0FBFCE1EEFBE0" blockId="13.[160,1156,1082,1135]" pageId="13" pageNumber="24">
|
||
<emphasis id="B911D014FF93E05730A0FBFCE291FBDE" bold="true" box="[161,251,1082,1105]" pageId="13" pageNumber="24">Figure 7.</emphasis>
|
||
<taxonomicName id="4C657785FF93E0573103FBFCE3A3FBDE" authorityName="Chen & Zhou & Dai & Wu & Ouyang" authorityYear="2024" box="[258,457,1082,1105]" class="Gastropoda" family="Enidae" genus="Petraeomastus" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="24" phylum="Mollusca" rank="species" species="gigas" status="sp. nov.">
|
||
<emphasis id="B911D014FF93E0573103FBFCE3A3FBDE" box="[258,457,1082,1105]" italics="true" pageId="13" pageNumber="24">Petraeomastus gigas</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911D014FF93E05731D1FBFCE060FBDE" bold="true" box="[464,522,1082,1105]" pageId="13" pageNumber="24">
|
||
<taxonomicNameLabel id="A2226D6FFF93E05731D1FBFCE060FBDE" box="[464,522,1082,1105]" pageId="13" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and its similar congener,
|
||
<taxonomicName id="4C657785FF93E057330CFBFCE1E7FBDE" authorityName="Ancey" authorityYear="1883" box="[781,909,1082,1105]" class="Gastropoda" family="Enidae" genus="Buliminus" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="24" phylum="Mollusca" rank="species" species="heudeanus">
|
||
<emphasis id="B911D014FF93E057330CFBFCE1E7FBDE" box="[781,909,1082,1105]" italics="true" pageId="13" pageNumber="24">P. heudeanus</emphasis>
|
||
</taxonomicName>
|
||
. (A)
|
||
<taxonomicName id="4C657785FF93E05733BCFBFCE6EEFBDE" authorityName="Chen & Zhou & Dai & Wu & Ouyang" authorityYear="2024" box="[957,1156,1082,1105]" class="Gastropoda" family="Enidae" genus="Petraeomastus" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="24" phylum="Mollusca" rank="species" species="gigas" status="sp. nov.">
|
||
<emphasis id="B911D014FF93E05733BCFBFCE6EEFBDE" box="[957,1156,1082,1105]" italics="true" pageId="13" pageNumber="24">Petraeomastus gigas</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911D014FF93E05730A1FB9EE2B0FBE0" bold="true" box="[160,218,1112,1135]" pageId="13" pageNumber="24">
|
||
<taxonomicNameLabel id="A2226D6FFF93E05730A1FB9EE2B0FBE0" box="[160,218,1112,1135]" pageId="13" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, ZMNH2409001; (B)
|
||
<taxonomicName id="4C657785FF93E057320BFB9EE0E0FBE0" authorityName="Ancey" authorityYear="1883" box="[522,650,1112,1135]" class="Gastropoda" family="Enidae" genus="Buliminus" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="24" phylum="Mollusca" rank="species" species="heudeanus">
|
||
<emphasis id="B911D014FF93E057320BFB9EE0E0FBE0" box="[522,650,1112,1135]" italics="true" pageId="13" pageNumber="24">P. heudeanus</emphasis>
|
||
</taxonomicName>
|
||
, 24_NCU_XPWU_AN144.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDA0C06FF93E05530A1FA26E14AFDAB" blockId="13.[160,1157,1193,1704]" lastBlockId="15.[160,1157,142,548]" lastPageId="15" lastPageNumber="26" pageId="13" pageNumber="24">
|
||
<emphasis id="B911D014FF93E05730A1FA26E367FA75" box="[160,269,1504,1530]" italics="true" pageId="13" pageNumber="24">Genitalia.</emphasis>
|
||
(
|
||
<figureCitation id="135E1083FF93E0573125FA27E3E3FA75" box="[292,393,1505,1530]" captionStart="Figure 8" captionStartId="14.[160,225,1493,1516]" captionTargetBox="[164,1152,143,1460]" captionTargetId="figure-3@14.[163,1153,142,1461]" captionTargetPageId="14" captionText="Figure 8. Genital anatomy of Petraeomastus gigas sp. n. (A) aperture view; (B) back view. A-1: most proximal section of penial appendix; A-2: penial appendix section between and thicker than A-1 and A-3, usually bulb-shaped; A-3:section of the penial appendix connecting proximally A-2 and distally A-4; A-4: thinnest part of the penial appendix between A-5 and A-3; A-5: distal part of the penial appendix, more or less swollen; AR: retractor muscle of the appendicular branch; At: atrium; BC: bursa copulatrix; BCD: bursa copulatrix duct;D: diverticle of the bursa copulatrix duct; Ep: epiphallus;EpC: epiphallic caecum;Fl: flagellum; FO: free oviduct; P: penis; PC: penial caecum; PR: retractor muscle of the penial branch; Va: vagina; VD: vas deferens." figureDoi="http://doi.org/10.5281/zenodo.14764953" httpUri="https://zenodo.org/record/14764953/files/figure.png" pageId="13" pageNumber="24">Figure 8</figureCitation>
|
||
) n = 5. Vas deferens short, slightly tapering distally; entering epiphallus apically with distinct demarcation. Epiphallus long; cylindrical; straight; externally smooth. Epiphallic caecum present; blunt apically; located near vas deferens entrance. Flagellum short; triangular; proximally expanded; with tip pointed. Penis with terminal entrance of epiphallus; clavate; distally tapering. Penial caecum present, subglobular. Penial appendix long; branched off from penis at some distance from atrium; divided into sections including A-1 + A-2, A-3 and A-4 + A-5. A-1 and A-2 fused. A-2 and A-3 not fused. Boundary between A-4 and A-5 indistinct. A-5 long; convoluted. Appendicular retractor and penial retractor short; biramous; attaching to penis proximally and to A-1 + A-2 of penial appendix; with penial retractor arms arising from diaphragm closed to each other. Additional retractor other than penial or appendicular absent. Muscular band connecting vagina and epiphallus absent. Atrium short; without retractor. Free oviduct longer than vagina. Vagina short; swollen; curved; unpigmented. Bursa copulatrix duct long; proximally straight. Bursa copulatrix ovoid, with stalk; without apical ligament; normal in size; with long neck; well defined. Diverticle normally present; longer than bursa copulatrix; unexpanded. Bursa copulatrix and diverticle distinguishable; forked more distally from their base.
|
||
</paragraph>
|
||
<caption id="DF1A5C8EFF90E05430A1FA13E3F8F936" ID-DOI="http://doi.org/10.5281/zenodo.14764953" ID-Zenodo-Dep="14764953" httpUri="https://zenodo.org/record/14764953/files/figure.png" pageId="14" pageNumber="25" startId="14.[160,225,1493,1516]" targetBox="[164,1152,143,1460]" targetPageId="14" targetType="figure">
|
||
<paragraph id="8BDA0C06FF90E05430A1FA13E3F8F936" blockId="14.[160,1157,1493,1721]" pageId="14" pageNumber="25">
|
||
<emphasis id="B911D014FF90E05430A1FA13E296FA63" bold="true" box="[160,252,1493,1516]" pageId="14" pageNumber="25">Figure 8.</emphasis>
|
||
Genital anatomy of
|
||
<taxonomicName id="4C657785FF90E05431CAFA13E0E4FA63" authorityName="Chen & Zhou & Dai & Wu & Ouyang" authorityYear="2024" box="[459,654,1493,1516]" class="Gastropoda" family="Enidae" genus="Petraeomastus" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="25" phylum="Mollusca" rank="species" species="gigas" status="sp. nov.">
|
||
<emphasis id="B911D014FF90E05431CAFA13E0E4FA63" box="[459,654,1493,1516]" italics="true" pageId="14" pageNumber="25">Petraeomastus gigas</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911D014FF90E0543296FA13E0BBFA63" bold="true" box="[663,721,1493,1516]" pageId="14" pageNumber="25">
|
||
<taxonomicNameLabel id="A2226D6FFF90E0543296FA13E0BBFA63" box="[663,721,1493,1516]" pageId="14" pageNumber="25" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A) aperture view; (B) back view. A-1: most proximal section of penial appendix; A-2: penial appendix section between and thicker than A-1 and A-3, usually bulb-shaped; A-3: section of the penial appendix connecting proximally A-2 and distally A-4; A-4: thinnest part of the penial appendix between A-5 and A-3; A-5: distal part of the penial appendix, more or less swollen; AR: retractor muscle of the appendicular branch; At: atrium; BC: bursa copulatrix; BCD: bursa copulatrix duct; D: diverticle of the bursa copulatrix duct; Ep: epiphallus; EpC: epiphallic caecum; Fl: flagellum; FO: free oviduct; P: penis; PC: penial caecum; PR: retractor muscle of the penial branch; Va: vagina; VD: vas deferens.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDA0C06FF91E05530A1FD96E327FDE5" blockId="15.[160,1156,592,722]" box="[160,333,592,618]" pageId="15" pageNumber="26">
|
||
<emphasis id="B911D014FF91E05530A1FD96E327FDE5" bold="true" box="[160,333,592,618]" italics="true" pageId="15" pageNumber="26">Measurements</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDA0C06FF91E05530A1FDB2E077FD5D" blockId="15.[160,1156,592,722]" pageId="15" pageNumber="26">
|
||
<typeStatus id="54DEB2A4FF91E05530A1FDB2E363FD02" box="[160,265,628,653]" pageId="15" pageNumber="26" type="holotype">Holotype</typeStatus>
|
||
: shell height
|
||
<quantity id="4C9DA1E3FF91E055319DFDB2E06AFD02" box="[412,512,628,653]" metricMagnitude="-2" metricUnit="m" metricValue="4.029999999999999" pageId="15" pageNumber="26" unit="mm" value="40.3">40.3 mm</quantity>
|
||
, width 13.0 mm; aperture height
|
||
<quantity id="4C9DA1E3FF91E0553374FDB2E1B3FD02" box="[885,985,628,653]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="15" pageNumber="26" unit="mm" value="13.5">13.5 mm</quantity>
|
||
, width
|
||
<quantity id="4C9DA1E3FF91E055342BFDB2E6EAFD02" box="[1066,1152,628,653]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="15" pageNumber="26" unit="mm" value="9.9">9.9 mm</quantity>
|
||
.
|
||
<typeStatus id="54DEB2A4FF91E05530A1FD50E37BFD20" box="[160,273,662,687]" pageId="15" pageNumber="26" type="paratype">Paratypes</typeStatus>
|
||
: n = 12. shell height
|
||
<quantity id="4C9DA1E3FF91E0553216FD50E0D7FD20" box="[535,701,662,687]" metricMagnitude="-2" metricUnit="m" metricValue="3.475" metricValueMax="3.94" metricValueMin="3.01" pageId="15" pageNumber="26" unit="mm" value="34.75" valueMax="39.4" valueMin="30.1">30.1–39.4 mm</quantity>
|
||
, width 11.8–13.0 mm; aperture height
|
||
<quantity id="4C9DA1E3FF91E05530A1FD7FE32BFD5D" box="[160,321,697,722]" metricMagnitude="-2" metricUnit="m" metricValue="1.295" metricValueMax="1.3199999999999998" metricValueMin="1.27" pageId="15" pageNumber="26" unit="mm" value="12.95" valueMax="13.2" valueMin="12.7">12.7–13.2 mm</quantity>
|
||
, width
|
||
<quantity id="4C9DA1E3FF91E0553192FD7FE073FD5D" box="[403,537,697,722]" metricMagnitude="-3" metricUnit="m" metricValue="9.05" metricValueMax="9.4" metricValueMin="8.7" pageId="15" pageNumber="26" unit="mm" value="9.05" valueMax="9.4" valueMin="8.7">8.7–9.4 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F5F8DFF91E05530A1FD3BE150FCD2" pageId="15" pageNumber="26" type="etymology">
|
||
<paragraph id="8BDA0C06FF91E05530A1FD3BE377FC98" blockId="15.[160,1155,765,861]" box="[160,285,765,791]" pageId="15" pageNumber="26">
|
||
<emphasis id="B911D014FF91E05530A1FD3BE377FC98" bold="true" box="[160,285,765,791]" italics="true" pageId="15" pageNumber="26">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDA0C06FF91E05530A1FCE7E150FCD2" blockId="15.[160,1155,765,861]" pageId="15" pageNumber="26">
|
||
The specific name is the Latin
|
||
<taxonomicName id="4C657785FF91E05531FFFCE7E054FCB5" authorityName="Chen & Zhou & Dai & Wu & Ouyang" authorityYear="2024" box="[510,574,801,826]" class="Gastropoda" family="Enidae" genus="Petraeomastus" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="26" phylum="Mollusca" rank="species" species="gigas">
|
||
‘
|
||
<emphasis id="B911D014FF91E0553205FCE7E054FCB5" box="[516,574,801,826]" italics="true" pageId="15" pageNumber="26">gigas</emphasis>
|
||
</taxonomicName>
|
||
’ for giant, alluding to the large size of this species compared to its congeners. It is a Latin noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F5F8DFF91E05530A1FC4EE05AFC4A" pageId="15" pageNumber="26" type="vernacular_names">
|
||
<paragraph id="8BDA0C06FF91E05530A1FC4EE306FC2D" blockId="15.[160,560,904,965]" box="[160,364,904,930]" pageId="15" pageNumber="26">
|
||
<emphasis id="B911D014FF91E05530A1FC4EE306FC2D" bold="true" box="[160,364,904,930]" italics="true" pageId="15" pageNumber="26">Vernacular name</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDA0C06FF91E05530A1FC6CE05AFC4A" blockId="15.[160,560,904,965]" box="[160,560,938,965]" pageId="15" pageNumber="26">巨鸟唇螺 (Pinyin: jù niǎo chún luó).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F5F8DFF91E05530A1FC37E12EFBFD" pageId="15" pageNumber="26" type="distribution">
|
||
<paragraph id="8BDA0C06FF91E05530A1FC37E3AFFB84" blockId="15.[160,1155,1009,1138]" box="[160,453,1009,1035]" pageId="15" pageNumber="26">
|
||
<emphasis id="B911D014FF91E05530A1FC37E3AFFB84" bold="true" box="[160,453,1009,1035]" italics="true" pageId="15" pageNumber="26">Distribution and ecology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDA0C06FF91E05530A1FBD2E12EFBFD" blockId="15.[160,1155,1009,1138]" pageId="15" pageNumber="26">
|
||
<taxonomicName id="4C657785FF91E05530A1FBD2E3EAFBA2" authorityName="Chen & Zhou & Dai & Wu & Ouyang" authorityYear="2024" box="[160,384,1044,1069]" class="Gastropoda" family="Enidae" genus="Petraeomastus" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="26" phylum="Mollusca" rank="species" species="gigas" status="sp. nov.">
|
||
<emphasis id="B911D014FF91E05530A1FBD2E3EAFBA2" box="[160,384,1044,1069]" italics="true" pageId="15" pageNumber="26">Petraeomastus gigas</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911D014FF91E0553184FBD2E3AEFBA2" bold="true" box="[389,452,1044,1069]" pageId="15" pageNumber="26">
|
||
<taxonomicNameLabel id="A2226D6FFF91E0553184FBD2E3AEFBA2" box="[389,452,1044,1069]" pageId="15" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is distributed in Mangkang County and Batang County on both banks of the Jinshajiang River (
|
||
<figureCitation id="135E1083FF91E05531FCFBF1E03EFBDF" box="[509,596,1079,1104]" captionStart="Figure 1" captionStartId="3.[160,225,1163,1186]" captionTargetBox="[166,1149,145,1127]" captionTargetId="figure-106@3.[163,1153,142,1131]" captionTargetPageId="3" captionText="Figure 1. Distribution of four enid gastropod species reviewed or described in this study. Coloured or shadowed regions show the distributional range. (A) Petraeomastus neumayri; (B) Petraeomastus desgodinsi; (C) Petraeomastus markamensis; (D) Petraeomastus gigas sp. n.; (E–F) The location of the research region in China. ① 1 km west of Jinshajiang River Bridge, ② Chaxue Village, ③ southern Batang County, ④ Langdaquka." figureDoi="http://doi.org/10.5281/zenodo.14764933" httpUri="https://zenodo.org/record/14764933/files/figure.png" pageId="15" pageNumber="26">Figure 1</figureCitation>
|
||
(D)). It inhabits shrubs and hibernates at the base of the shrub (
|
||
<figureCitation id="135E1083FF91E0553138FB9FE3FCFBFD" box="[313,406,1113,1138]" captionStart="Figure 4" captionStartId="6.[160,225,1659,1682]" captionTargetBox="[164,1152,1133,1627]" captionTargetId="figure-283@6.[163,1153,1132,1628]" captionTargetPageId="6" captionText="Figure 4. Adjacent environment and habitat of living individuals of Petraeomastus land snails. (A&B) Environment of the Jinshajiang River Valley, left side: Batang County, right side: Mangkang County; (C) living specimens of P. neumayri; (D) living specimens of P. desgodinsi; (E) living specimens of P. markamensis; (F) living specimens of Petraeomastus gigas sp. n. and P. markamensis." figureDoi="http://doi.org/10.5281/zenodo.14764939" httpUri="https://zenodo.org/record/14764939/files/figure.png" pageId="15" pageNumber="26">Figure 4</figureCitation>
|
||
(F)). Juveniles are not covered in mud.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F5F8DFF91E05530A1FB58E01BF99C" pageId="15" pageNumber="26" type="discussion">
|
||
<paragraph id="8BDA0C06FF91E05530A1FB58E36CFB37" blockId="15.[160,1156,1182,1555]" box="[160,262,1182,1208]" pageId="15" pageNumber="26">
|
||
<emphasis id="B911D014FF91E05530A1FB58E36CFB37" bold="true" box="[160,262,1182,1208]" italics="true" pageId="15" pageNumber="26">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDA0C06FF91E05530A1FB04E01BF99C" blockId="15.[160,1156,1182,1555]" pageId="15" pageNumber="26">
|
||
Currently, the generic diagnostic characters of Chinese
|
||
<taxonomicName id="4C657785FF91E0553336FB04E1E9FB54" authorityName="Woodward" authorityYear="1903" box="[823,899,1218,1243]" class="Gastropoda" family="Enidae" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="26" phylum="Mollusca" rank="family">Enidae</taxonomicName>
|
||
are mainly based on shell morphology (
|
||
<bibRefCitation id="EFF471F7FF91E0553176FB22E3B7FB72" author="Wu M" box="[375,477,1252,1277]" pageId="15" pageNumber="26" pagination="1 - 6" refId="ref7038" refString="Wu M. 2018. Mollusca, Gastropoda: enoidea. Fauna Sinica, Invertebrata. Beijing: Science Press. Vol. 58. p. 298, pls 1 - 6." type="book chapter" year="2018">Wu 2018</bibRefCitation>
|
||
). Although some studies have indicated that there may be slight differences in the genitalia between genera (
|
||
<bibRefCitation id="EFF471F7FF91E0553315FAC1E1EBFAAF" author="Wu M" box="[788,897,1287,1312]" pageId="15" pageNumber="26" pagination="1 - 6" refId="ref7038" refString="Wu M. 2018. Mollusca, Gastropoda: enoidea. Fauna Sinica, Invertebrata. Beijing: Science Press. Vol. 58. p. 298, pls 1 - 6." type="book chapter" year="2018">Wu 2018</bibRefCitation>
|
||
), it is challenging to summarise these differences as a universal diagnostic character for a genus. The placement of this new species within
|
||
<taxonomicName id="4C657785FF91E0553255FA8AE09DFAEA" authorityName="Mollendorff" authorityYear="1901" box="[596,759,1356,1381]" class="Gastropoda" family="Enidae" genus="Petraeomastus" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="26" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B911D014FF91E0553255FA8AE09DFAEA" box="[596,759,1356,1381]" italics="true" pageId="15" pageNumber="26">Petraeomastus</emphasis>
|
||
</taxonomicName>
|
||
is supported by its shell morphology – shell cylindrical-conic to conic – which is very similar to that of
|
||
<taxonomicName id="4C657785FF91E05533ECFAA9E6E9FA07" authorityName="Ancey" authorityYear="1883" box="[1005,1155,1391,1416]" class="Gastropoda" family="Enidae" genus="Buliminus" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="26" phylum="Mollusca" rank="species" species="heudeanus">
|
||
<emphasis id="B911D014FF91E05533ECFAA9E6E9FA07" box="[1005,1155,1391,1416]" italics="true" pageId="15" pageNumber="26">P. heudeanus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="135E1083FF91E05530A9FA54E360FA24" box="[168,266,1426,1451]" captionStart="Figure 7" captionStartId="13.[161,226,1082,1105]" captionTargetBox="[164,1152,143,1050]" captionTargetId="figure-157@13.[163,1153,142,1051]" captionTargetPageId="13" captionText="Figure 7. Petraeomastus gigas sp. n. and its similar congener, P. heudeanus. (A) Petraeomastus gigas sp. n., holotype, ZMNH2409001; (B) P. heudeanus, 24_NCU_XPWU_AN144." figureDoi="http://doi.org/10.5281/zenodo.14764949" httpUri="https://zenodo.org/record/14764949/files/figure.png" pageId="15" pageNumber="26">Figure 7</figureCitation>
|
||
(B)), the
|
||
<typeStatus id="54DEB2A4FF91E0553175FA54E3CDFA24" box="[372,423,1426,1451]" pageId="15" pageNumber="26">type</typeStatus>
|
||
species of the genus. It is the largest enid species in
|
||
<collectingCountry id="F3724C96FF91E0553443FA54E6E9FA24" box="[1090,1155,1426,1451]" name="China" pageId="15" pageNumber="26">China</collectingCountry>
|
||
(height
|
||
<quantity id="4C9DA1E3FF91E0553102FA72E3C4FA42" box="[259,430,1460,1485]" metricMagnitude="-2" metricUnit="m" metricValue="3.5200000000000005" metricValueMax="4.029999999999999" metricValueMin="3.0100000000000002" pageId="15" pageNumber="26" unit="mm" value="35.2" valueMax="40.3" valueMin="30.1">30.1–40.3 mm</quantity>
|
||
vs <33.0 mm), and can be easily distinguished from all congeners by its shell size, teleoconch with reddish brown background, irregular white stripes, and slit-shaped umbilicus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |