305 lines
34 KiB
XML
305 lines
34 KiB
XML
<document id="DC5D62263F85F1140E09A6DC51452D8E" ID-DOI="10.24199/j.mmv.2008.65.3" ID-ISSN="1447-2554" ID-Zenodo-Dep="10665901" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1708004926147" checkinUser="felipe" docAuthor="Dean, J. C., Forteath, G. N. R. & Osborn, A. W." docDate="2008" docId="039D8005AC279D48FF1CFAE4FA5CF112" docLanguage="en" docName="MemMusVictoria.65.43-50.pdf" docOrigin="Memoirs of Museum Victoria 65" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-65-2008/pages-43-50/" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Marmenuera ida" docType="treatment" docVersion="1" lastPageNumber="48" masterDocId="FFA4F87DAC239D4DFF90FFE5FFF4F67C" masterDocTitle="Marmenuera, a new genus of leptophlebiid mayfly (Insecta: Ephemeroptera) endemic to Tasmania." masterLastPageNumber="50" masterPageNumber="43" pageNumber="47" updateTime="1711460616528" updateUser="juliana" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="56777B99E0E06955670137124F77FA78" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0F9D389AD44C6208E38AFF891E27F471">
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<mods:title id="E2B3C67B103974DFE7F47016BA7F13ED">Marmenuera, a new genus of leptophlebiid mayfly (Insecta: Ephemeroptera) endemic to Tasmania.</mods:title>
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<mods:name id="4996118E0FACCF72DFD72409A75D7EB0" type="personal">
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<mods:roleTerm id="BC3CD5735BE47B5E96976C8932E573DE">Author</mods:roleTerm>
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<mods:namePart id="51970B62FAB8DD37AA305BC16AE2CE14">Dean, J. C.</mods:namePart>
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<mods:affiliation id="617958ED694446DE4918169BF1C49229">Environment Protection Authority, Freshwater Sciences Unit, Ernest Jones Drive, Macleod, Victoria, 3085, Australia (john. dean @ epa. vic. gov. au)</mods:affiliation>
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<mods:nameIdentifier id="94565CBC7B9957E483F103C10E28C9EA" type="email">john.dean@epa.vic.gov.au</mods:nameIdentifier>
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<mods:name id="B4C11A51E5AB6ECA3F4B386233998451" type="personal">
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<mods:roleTerm id="2FA09FE158B23A548FFF2BC5E51472BF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D8137FB0E1C32321F54846C1700731D0">Forteath, G. N. R.</mods:namePart>
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<mods:affiliation id="52560BAD9572B874B15F5685DA4E3CA8">2 Honorary Research Fellow, Queen Victoria Museum and Art Gallery, Wellington Street, Launceston, Tasmania, 7250, Australia</mods:affiliation>
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</mods:role>
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<mods:namePart id="416500ACBC43EF600095C7D2FEE90931">Osborn, A. W.</mods:namePart>
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<mods:affiliation id="CFF98DA95EF66FFBB01062F3D0811672">2 Honorary Research Fellow, Queen Victoria Museum and Art Gallery, Wellington Street, Launceston, Tasmania, 7250, Australia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="9941601789212932B3DAD793F346D3A8">text</mods:typeOfResource>
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<mods:title id="4B1EF455D3278D6ABC7654707782DA5E">Memoirs of Museum Victoria</mods:title>
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<mods:part id="22A0938244EBD67FAF7E4343C7B4CDCA">
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<mods:date id="CE5D94CD4630643DDD75AFACC3F6754C">2008</mods:date>
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<mods:number id="C93680F90E63609F98429751CBA6EF7F">2008-12-31</mods:number>
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<mods:number id="AC380D2CE560DD026A680E827CD6ED25">65</mods:number>
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<mods:start id="3A1E1612870765D9C51DFE46902263B1">43</mods:start>
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<mods:end id="462CF611993415B5761D55863F1914F9">50</mods:end>
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<mods:url id="F164BDA33DD86362D70EAF7E82871057">https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-65-2008/pages-43-50/</mods:url>
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<mods:classification id="BC94056A10D84BDAAD9B9DBBD8BEF5CA">journal article</mods:classification>
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<mods:identifier id="69061F55FEB34DB73782FCF15BE0CCC8" type="DOI">10.24199/j.mmv.2008.65.3</mods:identifier>
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<mods:identifier id="F3939828A7117F5F0C21DA09CEA74AE2" type="ISSN">1447-2554</mods:identifier>
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<treatment id="039D8005AC279D48FF1CFAE4FA5CF112" ID-DOI="http://doi.org/10.5281/zenodo.10666090" ID-Zenodo-Dep="10666090" LSID="urn:lsid:plazi:treatment:039D8005AC279D48FF1CFAE4FA5CF112" httpUri="http://treatment.plazi.org/id/039D8005AC279D48FF1CFAE4FA5CF112" lastPageId="5" lastPageNumber="48" pageId="4" pageNumber="47">
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<subSubSection id="C32E6298AC279D49FF1CFAE4FE53F36A" box="[140,423,1280,1302]" pageId="4" pageNumber="47" type="nomenclature">
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<paragraph id="8B8B3113AC279D49FF1CFAE4FE53F36A" blockId="4.[140,423,1280,1302]" box="[140,423,1280,1302]" pageId="4" pageNumber="47">
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<heading id="D0C3867FAC279D49FF1CFAE4FE53F36A" box="[140,423,1280,1302]" fontSize="9" level="2" pageId="4" pageNumber="47" reason="5">
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<taxonomicName id="4C344A90AC279D49FF1CFAE4FE53F36A" ID-CoL="3Y7Y6" authority="(Tillyard)" baseAuthorityName="Tillyard" baseAuthorityYear="1936" box="[140,423,1280,1302]" class="Insecta" family="Leptophlebiidae" genus="Marmenuera" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="ida">
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<emphasis id="B940ED01AC279D49FF1CFAE4FECAF36A" bold="true" box="[140,318,1280,1302]" italics="true" pageId="4" pageNumber="47">Marmenuera ida</emphasis>
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(Tillyard)
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E6298AC279D49FF1CFAC8FEEEF33E" box="[140,282,1325,1347]" pageId="4" pageNumber="47" type="description">
|
||
<paragraph id="8B8B3113AC279D49FF1CFAC8FEEEF33E" blockId="4.[140,282,1325,1347]" box="[140,282,1325,1347]" pageId="4" pageNumber="47">
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<figureCitation id="130F2D96AC279D49FF1CFAC8FEEEF33E" box="[140,282,1325,1347]" captionStart-0="Figures 17–21" captionStart-1="Figures 22–29" captionStartId-0="5.[147,215,1321,1340]" captionStartId-1="6.[140,208,1622,1641]" captionTargetBox-0="[159,1438,218,1301]" captionTargetBox-1="[148,1435,226,1525]" captionTargetId-0="figure-240@5.[147,1446,191,1302]" captionTargetId-1="figure-112@6.[129,1463,217,1543]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Figures 17–21. Marmenuera ida. Male imago: 17, forewing; 18, hindwing. Female imago: 19, forewing; 20, hindwing; 21, forewing, holotype." captionText-1="Figures 22–29. Marmenuera ida. Male imago: 22, abdomen, dorsal; 23, abdomen, ventral; 24, penes, dorsal; 25, penes, ventral. Nymph: 26, abdomen, dorsal; 27, labrum; 28, forefemur; 29, foretarsus." figureDoi-0="http://doi.org/10.5281/zenodo.10665911" figureDoi-1="http://doi.org/10.5281/zenodo.10665917" httpUri-0="https://zenodo.org/record/10665911/files/figure.png" httpUri-1="https://zenodo.org/record/10665917/files/figure.png" pageId="4" pageNumber="47">Figures 17–29</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E6298AC279D49FF22FABDFD8DF3DC" pageId="4" pageNumber="47" type="reference_group">
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<paragraph id="8B8B3113AC279D49FF22FABDFD59F317" blockId="4.[178,685,1368,1440]" box="[178,685,1368,1387]" pageId="4" pageNumber="47">
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<treatmentCitationGroup id="AB24163DAC279D49FF22FABDFD59F317" box="[178,685,1368,1387]" pageId="4" pageNumber="47">
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<treatmentCitation id="0A951702AC279D49FF22FABDFDFFF317" author="Tillyard, R. J." box="[178,523,1368,1387]" page="42" pageId="4" pageNumber="47" year="1936">
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<taxonomicName id="4C344A90AC279D49FF22FABDFDFFF317" authority="Tillyard, 1936: p. 42" authorityName=", Tillyard" authorityPageNumber="42" authorityYear="1936" box="[178,523,1368,1387]" class="Insecta" family="Leptophlebiidae" genus="Atalophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="ida">
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<emphasis id="B940ED01AC279D49FF22FABDFEBFF317" box="[178,331,1368,1387]" italics="true" pageId="4" pageNumber="47">Atalophlebia ida</emphasis>
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<bibRefCitation id="EFA54CE2AC279D49FEC1FABDFE21F317" author="Tillyard, R. J." box="[337,469,1368,1387]" pageId="4" pageNumber="47" pagination="23 - 59" refId="ref4883" refString="Tillyard, R. J. 1936. The trout-food insects of Tasmania. Part II: A monograph of the mayflies of Tasmania. Papers and Proceedings of the Royal Society of Tasmania. 1935: 23 - 59." type="journal article" year="1936">Tillyard, 1936</bibRefCitation>
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: p. 42
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</taxonomicName>
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</treatmentCitation>
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, fig. 7, plate I(10).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B8B3113AC279D49FF22FA97FD99F3FA" blockId="4.[178,685,1368,1440]" box="[178,621,1394,1414]" pageId="4" pageNumber="47">
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<treatmentCitationGroup id="AB24163DAC279D49FF22FA97FD99F3FA" box="[178,621,1394,1414]" pageId="4" pageNumber="47">
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<treatmentCitation id="0A951702AC279D49FF22FA97FDD5F3FA" author="Scholes, D." box="[178,545,1394,1414]" page="30" pageId="4" pageNumber="47" year="1961">
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<taxonomicName id="4C344A90AC279D49FF22FA97FDD5F3FA" authority="Scholes, 1961: p. 30" authorityName="Scholes" authorityPageNumber="30" authorityYear="1961" box="[178,545,1394,1414]" class="Insecta" family="Leptophlebiidae" genus="Massartellopsis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="ida">
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<emphasis id="B940ED01AC279D49FF22FA97FE92F3F9" box="[178,358,1394,1413]" italics="true" pageId="4" pageNumber="47">Massartellopsis ida</emphasis>
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<bibRefCitation id="EFA54CE2AC279D49FEFCFA97FE1EF3F9" author="Scholes, D." box="[364,490,1394,1413]" pageId="4" pageNumber="47" refId="ref4862" refString="Scholes, D. 1961. Fly-fisher in Tasmania. Melbourne University Press: Carlton. Pp xiv + 208." type="book" year="1961">Scholes, 1961</bibRefCitation>
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: p. 30
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</taxonomicName>
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</treatmentCitation>
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, plate 2.
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</treatmentCitationGroup>
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||
</paragraph>
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<paragraph id="8B8B3113AC279D49FF22FA68FD8DF3DC" blockId="4.[178,685,1368,1440]" box="[178,633,1421,1440]" pageId="4" pageNumber="47">Genus W sp.AV2 Dean, 1999: p. 86, figs 242, 243.</paragraph>
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</subSubSection>
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||
<subSubSection id="C32E6298AC279D49FF1CFA52FD66F17B" pageId="4" pageNumber="47" type="materials_examined">
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||
<paragraph id="8B8B3113AC279D49FF1CFA52FDFEF065" blockId="4.[140,772,1463,1800]" pageId="4" pageNumber="47">
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<emphasis id="B940ED01AC279D49FF1CFA52FF32F3B5" box="[140,198,1463,1481]" italics="true" pageId="4" pageNumber="47">Types.</emphasis>
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||
<materialsCitation id="3B5C3B4EAC279D49FF5AFA52FEBEF398" collectingDate="1933-02-07" collectionCode="R, BMNH" collectorName="J. Tillyard" location="Dee Bridge" pageId="4" pageNumber="47" specimenCount="1" stateProvince="Tasmania" typeStatus="holotype">
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<typeStatus id="548F8FB1AC279D49FF5AFA52FED5F3B6" box="[202,289,1463,1482]" pageId="4" pageNumber="47" type="holotype">Holotype</typeStatus>
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: female imago,
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<collectingRegion id="49F0FFF1AC279D49FE3EFA52FDFCF3B6" box="[430,520,1463,1482]" country="Australia" name="Tasmania" pageId="4" pageNumber="47">Tasmania</collectingRegion>
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,
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<location id="8EEB67C8AC279D49FD80FA52FD8DF3B6" LSID="urn:lsid:plazi:treatment:039D8005AC279D48FF1CFAE4FA5CF112:8EEB67C8AC279D49FD80FA52FD8DF3B6" box="[528,633,1463,1482]" name="Dee Bridge" pageId="4" pageNumber="47" stateProvince="Tasmania">Dee Bridge</location>
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,
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<date id="FF8A17D3AC279D49FD12FA52FD11F3B6" box="[642,741,1463,1482]" pageId="4" pageNumber="47" value="1933-02-07">
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<collectingDate id="EFCEEE3BAC279D49FD12FA52FD11F3B6" box="[642,741,1463,1482]" pageId="4" pageNumber="47" value="1933-02-07">7 Feb 1933</collectingDate>
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</date>
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,
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<collectionCode id="ED25A9D6AC279D49FD7EFA52FCF4F3B6" box="[750,768,1463,1482]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="47">R</collectionCode>
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||
.
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<collectorName id="26C154C5AC279D49FF1CFA34FF18F398" box="[140,236,1489,1508]" pageId="4" pageNumber="47">J. Tillyard</collectorName>
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||
(
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<collectionCode id="ED25A9D6AC279D49FF6BFA34FEB0F398" box="[251,324,1489,1508]" pageId="4" pageNumber="47">BMNH</collectionCode>
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||
)
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</materialsCitation>
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.
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<materialsCitation id="3B5C3B4EAC279D49FEC7FA34FEC3F383" collectingDate="1933-02-07" collectionCode="BMNH" collectorName="J. Tillyard" location="Dee Bridge" pageId="4" pageNumber="47" specimenCount="5" specimenCount-female="5" stateProvince="Tasmania" typeStatus="paratype">
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<typeStatus id="548F8FB1AC279D49FEC7FA34FE47F398" box="[343,435,1489,1508]" pageId="4" pageNumber="47" type="paratype">Paratypes</typeStatus>
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||
:
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<specimenCount id="9D32FA9AAC279D49FE2FFA34FDD2F398" box="[447,550,1489,1508]" count="5" pageId="4" pageNumber="47" type="female">five female</specimenCount>
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imagos, collected with holotype (
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||
<collectionCode id="ED25A9D6AC279D49FF77FA09FEC4F383" box="[231,304,1516,1535]" pageId="4" pageNumber="47">BMNH</collectionCode>
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||
)
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||
</materialsCitation>
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||
;
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<materialsCitation id="3B5C3B4EAC279D49FED0FA09FCF4F383" box="[320,768,1515,1535]" collectingDate="1933-02-06" collectionCode="R" location="Lake St Clair" pageId="4" pageNumber="47" specimenCount="1" specimenCount-male="1" stateProvince="Tasmania" typeStatus="paratype">
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||
<specimenCount id="9D32FA9AAC279D49FED0FA09FE66F382" box="[320,402,1515,1535]" count="1" pageId="4" pageNumber="47" type="male">one male</specimenCount>
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||
subimago,
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||
<location id="8EEB67C8AC279D49FE6BFA0EFD83F382" LSID="urn:lsid:plazi:treatment:039D8005AC279D48FF1CFAE4FA5CF112:8EEB67C8AC279D49FE6BFA0EFD83F382" box="[507,631,1515,1535]" name="Lake St Clair" pageId="4" pageNumber="47" stateProvince="Tasmania">Lake St Clair</location>
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||
,
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||
<date id="FF8A17D3AC279D49FD10FA0EFD11F383" box="[640,741,1515,1535]" pageId="4" pageNumber="47" value="1933-02-06">
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||
<collectingDate id="EFCEEE3BAC279D49FD10FA0EFD11F383" box="[640,741,1515,1535]" pageId="4" pageNumber="47" value="1933-02-06">6 Feb 1933</collectingDate>
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||
</date>
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||
,
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||
<collectionCode id="ED25A9D6AC279D49FD7EFA09FCF4F383" box="[750,768,1516,1535]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="47">R</collectionCode>
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||
</materialsCitation>
|
||
.
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||
<materialsCitation id="3B5C3B4EAC279D49FF1CF9E3FDF2F065" box="[140,518,1542,1561]" collectionCode="BMNH" collectorName="J. Tillyard" pageId="4" pageNumber="47" specimenCount="1" stateProvince="Tasmania" typeStatus="paratype">
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||
<collectorName id="26C154C5AC279D49FF1CF9E3FF1CF065" box="[140,232,1542,1561]" pageId="4" pageNumber="47">J. Tillyard</collectorName>
|
||
(
|
||
<collectionCode id="ED25A9D6AC279D49FF64F9E3FEC9F065" box="[244,317,1542,1561]" pageId="4" pageNumber="47">BMNH</collectionCode>
|
||
). Types not examined
|
||
</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B8B3113AC279D49FF22F9C5FD66F17B" blockId="4.[140,772,1463,1800]" pageId="4" pageNumber="47">
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||
<emphasis id="B940ED01AC279D49FF22F9C5FE9EF04F" box="[178,362,1568,1587]" italics="true" pageId="4" pageNumber="47">Material examined.</emphasis>
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||
<collectingRegion id="49F0FFF1AC279D49FEE1F9C5FE38F04F" box="[369,460,1568,1587]" country="Australia" name="Tasmania" pageId="4" pageNumber="47">Tasmania</collectingRegion>
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. 5N, Lake Pedder, Trappes Inlet,
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<date id="FF8A17D3AC279D49FF1CF9DEFE82F032" box="[140,374,1595,1614]" pageId="4" pageNumber="47" value="1997-02" valueMax="1997-04-31" valueMin="1997-02-00">February and April 1997</date>
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, N. Forteath; 3FI (2 reared), 1FS (reared), same location,
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<date id="FF8A17D3AC279D49FE85F9B3FE88F015" box="[277,380,1621,1641]" pageId="4" pageNumber="47" value="1998-02-01">1 Feb 1998</date>
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, N. Forteath; 1MS (reared), same location,
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<date id="FF8A17D3AC279D49FF1CF995FF0CF0FF" box="[140,248,1648,1667]" pageId="4" pageNumber="47" value="1998-01-12">12 Jan 1998</date>
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, N. Forteath; 1MS (reared), same location,
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<date id="FF8A17D3AC279D49FDEBF995FD10F0FF" box="[635,740,1648,1667]" pageId="4" pageNumber="47" value="1998-03-04">4 Mar 1998</date>
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, N. Forteath; 6MI,5FI (all reared), 21N, same location,
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<date id="FF8A17D3AC279D49FDC9F96EFD10F0E1" box="[601,740,1674,1694]" pageId="4" pageNumber="47" value="2006-11" valueMax="2006-12-31" valueMin="2006-11-00">Nov–Dec 2006</date>
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, N. Forteath and A. Osborn; 1N, Lake Lilla, Cradle Mountain,
|
||
<date id="FF8A17D3AC279D49FD2EF940FF4EF0AE" pageId="4" pageNumber="47" value="1998-03-25">25 Mar 1998</date>
|
||
, J. Dean; 1N, Dip River Falls,
|
||
<date id="FF8A17D3AC279D49FE45F95AFDB2F0AE" box="[469,582,1727,1746]" pageId="4" pageNumber="47" value="1998-10-31">31 Oct 1998</date>
|
||
, D. Cartwright; 4FI, Huon River Crossing,
|
||
<date id="FF8A17D3AC279D49FEF5F93FFE2AF091" box="[357,478,1754,1773]" pageId="4" pageNumber="47" value="1971-02-16">16 Feb 1971</date>
|
||
, A. Neboiss; 1MI, 1MS, 1FS, D’Entrecasteaux Passage, Exit Cave,
|
||
<date id="FF8A17D3AC279D49FE4EF911FDDEF17B" box="[478,554,1780,1799]" pageId="4" pageNumber="47" value="1993-01">Jan1993</date>
|
||
, A. Clarke.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E6298AC279D49FF1CF8FAFBF8F0CD" pageId="4" pageNumber="47" type="description">
|
||
<paragraph id="8B8B3113AC279D49FF1CF8FAFBF8F0CD" blockId="4.[140,771,1823,1903]" lastBlockId="4.[809,1441,229,1713]" pageId="4" pageNumber="47">
|
||
<emphasis id="B940ED01AC279D49FF1CF8FAFEF5F148" box="[140,257,1823,1844]" italics="true" pageId="4" pageNumber="47">Description</emphasis>
|
||
.
|
||
<emphasis id="B940ED01AC279D49FE9BF8FAFE74F148" box="[267,384,1823,1844]" italics="true" pageId="4" pageNumber="47">Male imago</emphasis>
|
||
. Length: body 11.0–
|
||
<quantity id="4CCC9CF6AC279D49FDD9F8FAFD55F148" box="[585,673,1823,1844]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="4" pageNumber="47" unit="mm" value="12.4">12.4 mm</quantity>
|
||
, forewing 11.2–12.0 mm, caudal filaments
|
||
<quantity id="4CCC9CF6AC279D49FE72F8D9FD8EF12D" box="[482,634,1852,1873]" metricMagnitude="-2" metricUnit="m" metricValue="2.13" metricValueMax="2.42" metricValueMin="1.8399999999999999" pageId="4" pageNumber="47" unit="mm" value="21.299999999999997" valueMax="24.2" valueMin="18.4">18.4–24.2 mm</quantity>
|
||
. Eyes: upper lobes pale pink/brown, in contact on meson of head, lower lobes grey. Thorax: medium-dark brown; pronotum with narrow raised carina along median line, dark brown markings medially on posterior margin, along lateral margins and parallel to lateral margin approximately midway between margin and medial carina. Legs: predominantly yellow, all femora with two broad dark brown bands, one just beyond midlength and the other subapical; tarsal claws similar, each claw with terminal sclerotised hook and opposing ventral process. Forewing (fig. 17): membrane hyaline, stigmatic region slightly opaque, white; all costal, subcostal and
|
||
<collectionCode id="ED25A9D6AC279D49FB8BFE08FBDFF47E" box="[1051,1067,493,514]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="47">R</collectionCode>
|
||
1 –
|
||
<collectionCode id="ED25A9D6AC279D49FBD0FE08FBA4F47E" box="[1088,1104,493,514]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="47">R</collectionCode>
|
||
2 crossveins heavily suffused with dark brown, crossveins in vicinity of MA fork also strongly suffused with dark brown; large brown blotch filling area between veins Sc and
|
||
<collectionCode id="ED25A9D6AC279D49FB8FFDA1FBC4F425" box="[1055,1072,580,601]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="47">R</collectionCode>
|
||
2 at midlength of wing. Abdomen: strongly contrasting pattern of dark brown and yellow; terga 2-7 yellow in anterior half and dark brown in posterior half, yellow colouration extended back as narrow band along median line (fig. 22), extent of yellow colouration increasing from segments 6 to 8, segments 9 and 10 predominantly brown; sterna reddish brown, pale yellow along lateral margins, segments 2-7 with two pairs of small pale spots, the anterior pair more widely eparated than the posterior pair (fig. 23). Genitalia (figs 24–25): penes lobes fused basally, widely separated apically; each lobe relatively broad, apex rounded and turned slightly inwards, subapically with outer margin evenly curved, three moderate sized ventral spines posterior to where the lobes separate.
|
||
<emphasis id="B940ED01AC279D49FAC5FC46FC93F5AA" italics="true" pageId="4" pageNumber="47">Female imago</emphasis>
|
||
. Length: body
|
||
<quantity id="4CCC9CF6AC279D49FB92FC25FB6DF5A9" box="[1026,1177,960,982]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" metricValueMax="1.48" metricValueMin="1.16" pageId="4" pageNumber="47" unit="mm" value="13.2" valueMax="14.8" valueMin="11.6">11.6–14.8 mm</quantity>
|
||
.; forewing 12.0–
|
||
<quantity id="4CCC9CF6AC279D49FAD0FC25FA68F5A9" box="[1344,1436,960,981]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="4" pageNumber="47" unit="mm" value="14.2">14.2 mm</quantity>
|
||
; caudal filaments
|
||
<quantity id="4CCC9CF6AC279D49FC45FC38FB9EF58F" box="[981,1130,989,1011]" metricMagnitude="-2" metricUnit="m" metricValue="2.02" metricValueMax="2.2199999999999998" metricValueMin="1.8199999999999998" pageId="4" pageNumber="47" unit="mm" value="20.2" valueMax="22.2" valueMin="18.2">18.2–22.2 mm</quantity>
|
||
. Forewing: pigmentation more extensive than in male; dark brown suffusions around costal, subcostal and
|
||
<collectionCode id="ED25A9D6AC279D49FC50FBFDFC25F251" box="[960,977,1048,1069]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="47">R</collectionCode>
|
||
<subScript id="17B03356AC279D49FC41FBC2FC2CF24F" attach="left" box="[977,984,1063,1075]" fontSize="5" pageId="4" pageNumber="47">1</subScript>
|
||
–
|
||
<collectionCode id="ED25A9D6AC279D49FC75FBFDFC01F251" box="[997,1013,1048,1069]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="47">R</collectionCode>
|
||
<subScript id="17B03356AC279D49FC65FBC2FC08F24F" attach="left" box="[1013,1020,1063,1075]" fontSize="5" pageId="4" pageNumber="47">2</subScript>
|
||
crossveins broad, almost circular; large brown blotch surrounding MA fork. General colour and abdominal markings similar to male imago.
|
||
<emphasis id="B940ED01AC279D49FA93FBB6FA6EF21B" box="[1283,1434,1106,1128]" italics="true" pageId="4" pageNumber="47">Mature nymph</emphasis>
|
||
. Body length ranging from
|
||
<quantity id="4CCC9CF6AC279D49FBA0FB8AFB69F2F9" box="[1072,1181,1135,1157]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="4" pageNumber="47" unit="mm" value="14.0" valueMax="16.0" valueMin="12.0">12–16 mm</quantity>
|
||
; antennae a little over half length of body; cerci about 1¼–1½ body length, terminal filament a little longer. General colour yellow with brown markings; abdomen with conspicuous pattern of dark brown and yellow, segments 2-8 yellow towards anterior margin and dark brown to the posterior (fig. 26); all legs yellow with dark banding. Mouthparts: Labrum (fig. 27) a little broader than clypeus; maximum width 1.7–1.9 times length along median line; maximum width at about 2/3 labrum length, basal to this the lateral margins relatively straight and diverging; anterior margin moderately concave, with five well developed medial denticles. Legs moderately broad (fig. 28); forefemora length a little over 3 times width, outer margin bearing moderate length spine-like setae and also some longer, hair-like setae; fore-tarsus with 15–20 ventral spines (fig. 29); tarsal claws smooth, without ventral teeth. Postero-lateral spines on abdominal segments 7 (small) and 8-9 (relatively large); posterior margins of abdominal terga with series of large, conspicuous spines interspersed with the occasional shorter spine. Gills lanceolate, lateral tracheae moderately developed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E6298AC279D48FCB9F922FE6CF0AA" lastPageId="5" lastPageNumber="48" pageId="4" pageNumber="47" type="discussion">
|
||
<paragraph id="8B8B3113AC279D48FCB9F922FD9CF026" blockId="4.[809,1441,1735,1903]" lastBlockId="5.[147,779,1450,1750]" lastPageId="5" lastPageNumber="48" pageId="4" pageNumber="47">
|
||
Remarks. Although the
|
||
<typeStatus id="548F8FB1AC279D49FBB5F922FB89F0A1" box="[1061,1149,1735,1757]" pageId="4" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
has not been examined the description and, in particular, the image of the female forewing presented by
|
||
<bibRefCitation id="EFA54CE2AC279D49FC3EF8E4FBB2F16B" author="Tillyard, R. J." box="[942,1094,1793,1815]" pageId="4" pageNumber="47" pagination="23 - 59" refId="ref4883" refString="Tillyard, R. J. 1936. The trout-food insects of Tasmania. Part II: A monograph of the mayflies of Tasmania. Papers and Proceedings of the Royal Society of Tasmania. 1935: 23 - 59." type="journal article" year="1936">Tillyard (1936)</bibRefCitation>
|
||
leave no doubt as to the identity of this species. The species is readily distinguished from
|
||
<taxonomicName id="4C344A90AC279D49FA15F8FAFC8FF12D" authorityName="Dean & Forteath & Osborn" authorityYear="2008" class="Insecta" family="Leptophlebiidae" genus="Marmenuera" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="tillyardi">
|
||
<emphasis id="B940ED01AC279D49FA15F8FAFC8FF12D" italics="true" pageId="4" pageNumber="47">M. tillyardi</emphasis>
|
||
</taxonomicName>
|
||
in the adult by the abdominal colour pattern, the more extensive pigmentation of the forewing and the structure of the male genitalia, and in the nymph by the abdominal colour pattern and the structure of the labrum and foreleg.
|
||
<bibRefCitation id="EFA54CE2AC269D48FD2DFA2CFF25F381" author="Scholes, D." pageId="5" pageNumber="48" refId="ref4862" refString="Scholes, D. 1961. Fly-fisher in Tasmania. Melbourne University Press: Carlton. Pp xiv + 208." type="book" year="1961">Scholes (1961)</bibRefCitation>
|
||
referred this species to the genus
|
||
<taxonomicName id="4C344A90AC269D48FDF3FA02FCF0F380" authorityName="Demoulin" authorityYear="1955" box="[611,772,1511,1532]" class="Insecta" family="Leptophlebiidae" genus="Massartellopsis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B940ED01AC269D48FDF3FA02FCF0F380" box="[611,772,1511,1532]" italics="true" pageId="5" pageNumber="48">Massartellopsis</emphasis>
|
||
</taxonomicName>
|
||
, presumably based on advice from Edgar Riek (
|
||
<collectionCode id="ED25A9D6AC269D48FDF5F9E3FD5BF067" box="[613,687,1542,1563]" country="Australia" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="5" pageNumber="48" type="Museum">CSIRO</collectionCode>
|
||
Division of Entomology), but it is now known that the genus
|
||
<taxonomicName id="4C344A90AC269D48FF03F9A1FEC1F025" authorityName="Demoulin" authorityYear="1955" box="[147,309,1604,1625]" class="Insecta" family="Leptophlebiidae" genus="Massartellopsis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B940ED01AC269D48FF03F9A1FEC1F025" box="[147,309,1604,1625]" italics="true" pageId="5" pageNumber="48">Massartellopsis</emphasis>
|
||
</taxonomicName>
|
||
is endemic to South America.
|
||
</paragraph>
|
||
<caption id="DF4B619BAC269D48FF03FACCFA61F340" ID-DOI="http://doi.org/10.5281/zenodo.10665911" ID-Zenodo-Dep="10665911" box="[147,1429,1321,1340]" httpUri="https://zenodo.org/record/10665911/files/figure.png" pageId="5" pageNumber="48" startId="5.[147,215,1321,1340]" targetBox="[159,1438,218,1301]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8B8B3113AC269D48FF03FACCFA61F340" blockId="5.[147,1429,1321,1340]" box="[147,1429,1321,1340]" pageId="5" pageNumber="48">
|
||
Figures 17–21.
|
||
<taxonomicName id="4C344A90AC269D48FE8DFACFFE43F340" baseAuthorityName="Tillyard" baseAuthorityYear="1936" box="[285,439,1321,1340]" class="Insecta" family="Leptophlebiidae" genus="Marmenuera" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="ida">
|
||
<emphasis id="B940ED01AC269D48FE8DFACFFE43F340" box="[285,439,1321,1340]" italics="true" pageId="5" pageNumber="48">Marmenuera ida</emphasis>
|
||
</taxonomicName>
|
||
. Male imago: 17, forewing; 18, hindwing. Female imago: 19, forewing; 20, hindwing; 21, forewing, holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8B3113AC269D48FF29F986FE6CF0AA" blockId="5.[147,779,1450,1750]" pageId="5" pageNumber="48">
|
||
<bibRefCitation id="EFA54CE2AC269D48FF29F986FEA1F005" author="Scholes, D." box="[185,341,1635,1657]" pageId="5" pageNumber="48" refId="ref4862" refString="Scholes, D. 1961. Fly-fisher in Tasmania. Melbourne University Press: Carlton. Pp xiv + 208." type="book" year="1961">Scholes (1961)</bibRefCitation>
|
||
suggested the common names “Large Speckled Spinner” and “Large Speckled Dun” for the imago and subimago respectively of
|
||
<taxonomicName id="4C344A90AC269D48FE7AF947FD55F0CA" baseAuthorityName="Tillyard" baseAuthorityYear="1936" box="[490,673,1697,1719]" class="Insecta" family="Leptophlebiidae" genus="Marmenuera" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="ida">
|
||
<emphasis id="B940ED01AC269D48FE7AF947FD55F0CA" box="[490,673,1697,1719]" italics="true" pageId="5" pageNumber="48">Marmenuera ida</emphasis>
|
||
</taxonomicName>
|
||
, and we consider this appropriate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E6298AC269D48FF03F918FA5CF112" pageId="5" pageNumber="48" type="key">
|
||
<paragraph id="8B8B3113AC269D48FF03F918FEC0F16F" blockId="5.[147,308,1789,1811]" box="[147,308,1789,1811]" pageId="5" pageNumber="48">
|
||
<heading id="D0C3867FAC269D48FF03F918FEC0F16F" bold="true" box="[147,308,1789,1811]" fontSize="9" level="2" pageId="5" pageNumber="48" reason="2">
|
||
<emphasis id="B940ED01AC269D48FF03F918FEC0F16F" bold="true" box="[147,308,1789,1811]" pageId="5" pageNumber="48">Keys to species</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<key id="FFA56622AC269D48FF03F8CEFA5CF112" pageId="5" pageNumber="48">
|
||
<keyStep id="30C029B6AC269D48FF03F8CEFA5CF0E2" pageId="5" pageNumber="48">
|
||
<paragraph id="8B8B3113AC269D48FF03F8CEFCFEF13C" blockId="5.[147,778,1835,1857]" box="[147,778,1835,1857]" pageId="5" pageNumber="48">
|
||
<keyLead id="30C59226AC269D48FF03F8CEFCFEF13C" box="[147,778,1835,1857]" pageId="5" pageNumber="48">1. Adults 2</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8B3113AC269D48FF03F8BCFCFEF112" blockId="5.[147,778,1881,1903]" box="[147,778,1881,1903]" pageId="5" pageNumber="48">
|
||
<keyLead id="30C59226AC269D48FF03F8BCFCFEF112" box="[147,778,1881,1903]" pageId="5" pageNumber="48">– Nymphs 3</keyLead>
|
||
</paragraph>
|
||
<keyStep id="30C029B6AC269D48FCA1FA4FFA5CF0E2" pageId="5" pageNumber="48">
|
||
<paragraph id="8B8B3113AC269D48FCA1FA4FFA53F064" blockId="5.[817,1448,1450,1903]" pageId="5" pageNumber="48">
|
||
<keyLead id="30C59226AC269D48FCA1FA4FFA53F064" pageId="5" pageNumber="48">
|
||
2. Forewings heavily pigmented, crossveins around MA fork suffused with brown (figs 17,19); male imago with outer margin of penes lobes evenly rounded (fig. 24)
|
||
<taxonomicName id="4C344A90AC269D48FB6AF9E6FA53F064" baseAuthorityName="Tillyard" baseAuthorityYear="1936" box="[1274,1447,1539,1560]" class="Insecta" family="Leptophlebiidae" genus="Marmenuera" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="ida">
|
||
<emphasis id="B940ED01AC269D48FB6AF9E6FA53F064" box="[1274,1447,1539,1560]" italics="true" pageId="5" pageNumber="48">Marmenuera ida</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8B3113AC269D48FCA1F9D5FA5CF0E2" blockId="5.[817,1448,1450,1903]" pageId="5" pageNumber="48">
|
||
<keyLead id="30C59226AC269D48FCA1F9D5FA5CF0E2" pageId="5" pageNumber="48">
|
||
– Forewings lightly pigmented, crossveins around MA fork not suffused with brown (figs 1,3); male imago with outer margin of penes lobes angular (fig. 7)
|
||
<taxonomicName id="4C344A90AC269D48FB58F96CFA5CF0E2" authorityName="Dean & Forteath & Osborn" authorityYear="2008" box="[1224,1448,1673,1694]" class="Insecta" family="Leptophlebiidae" genus="Marmenuera" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="tillyardi">
|
||
<emphasis id="B940ED01AC269D48FB58F96CFA5CF0E2" box="[1224,1448,1673,1694]" italics="true" pageId="5" pageNumber="48">Marmenuera tillyardi</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</keyStep>
|
||
<keyStep id="30C029B6AC269D48FCA1F953FA5CF112" pageId="5" pageNumber="48">
|
||
<paragraph id="8B8B3113AC269D48FCA1F953FA53F17A" blockId="5.[817,1448,1450,1903]" pageId="5" pageNumber="48">
|
||
<keyLead id="30C59226AC269D48FCA1F953FA53F17A" pageId="5" pageNumber="48">
|
||
3. Foretarsus with 15–20 ventral spines (fig. 29); forefemur with spines along outer margin relatively long (fig. 28)
|
||
<taxonomicName id="4C344A90AC269D48FB6AF914FA53F17A" baseAuthorityName="Tillyard" baseAuthorityYear="1936" box="[1274,1447,1777,1798]" class="Insecta" family="Leptophlebiidae" genus="Marmenuera" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="ida">
|
||
<emphasis id="B940ED01AC269D48FB6AF914FA53F17A" box="[1274,1447,1777,1798]" italics="true" pageId="5" pageNumber="48">Marmenuera ida</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8B3113AC269D48FCA1F8FBFA5CF112" blockId="5.[817,1448,1450,1903]" pageId="5" pageNumber="48">
|
||
<keyLead id="30C59226AC269D48FCA1F8FBFA5CF112" pageId="5" pageNumber="48">
|
||
– Foretarsus with 30–40 ventral spines (fig. 14); forefemur with spines along outer margin relatively short (fig. 13)
|
||
<taxonomicName id="4C344A90AC269D48FB58F8BFFA5CF112" authorityName="Dean & Forteath & Osborn" authorityYear="2008" box="[1224,1448,1881,1903]" class="Insecta" family="Leptophlebiidae" genus="Marmenuera" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="tillyardi">
|
||
<emphasis id="B940ED01AC269D48FB58F8BFFA5CF112" box="[1224,1448,1881,1903]" italics="true" pageId="5" pageNumber="48">Marmenuera tillyardi</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |