538 lines
60 KiB
XML
538 lines
60 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1073.71843" ID-GBIF-Dataset="5161ac99-ec0f-4443-ac73-a9d016a9383d" ID-PMC="PMC8648685" ID-Pensoft-Pub="1313-2970-1073-55" ID-Pensoft-UUID="F401EBF007E0519AB4B8EC43D0F5EFE4" ID-PubMed="34949951" ID-ZooBank="18D5AF2786E54D21BCC527D09FB384DA" ModsDocID="1313-2970-1073-55" checkinTime="1638245621972" checkinUser="pensoft" docAuthor="Barr, Cheryl B. & Shepard, William D." docDate="2021" docId="F6F0B4DF411C5A9B9BA3B484B07774F1" docLanguage="en" docName="ZooKeys 1073: 55-117" docOrigin="ZooKeys 1073" docPubDate="2021-11-29" docSource="http://dx.doi.org/10.3897/zookeys.1073.71843" docTitle="Hydora laticeps" docType="treatment" docVersion="4" id="F401EBF007E0519AB4B8EC43D0F5EFE4" lastPageNumber="55" masterDocId="F401EBF007E0519AB4B8EC43D0F5EFE4" masterDocTitle="A review of the Larainae of Australia with description of seven new species and the new genus Australara (Coleoptera, Byrrhoidea, Elmidae)" masterLastPageNumber="117" masterPageNumber="55" pageNumber="55" updateTime="1668151180298" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of the Larainae of Australia with description of seven new species and the new genus Australara (Coleoptera, Byrrhoidea, Elmidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Barr, Cheryl B.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6707-4301</mods:nameIdentifier>
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<mods:affiliation>Essig Museum of Entomology, 1101 Life Sciences Bldg. # 4780, University of California, Berkeley, CA 94720 USA</mods:affiliation>
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<mods:nameIdentifier type="email">cbarr@berkeley.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shepard, William D.</mods:namePart>
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<mods:affiliation>Essig Museum of Entomology, 1101 Life Sciences Bldg. # 4780, University of California, Berkeley, CA 94720 USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-11-29</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1073</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>55</mods:start>
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<mods:end>117</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1073.71843</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1073.71843</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1073-55</mods:identifier>
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<mods:identifier type="ZooBank">18D5AF2786E54D21BCC527D09FB384DA</mods:identifier>
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<mods:identifier type="Pensoft-UUID">F401EBF007E0519AB4B8EC43D0F5EFE4</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="190874577" LSID="urn:lsid:plazi:treatment:F6F0B4DF411C5A9B9BA3B484B07774F1" httpUri="http://treatment.plazi.org/id/F6F0B4DF411C5A9B9BA3B484B07774F1" lastPageNumber="55" pageId="0" pageNumber="55">
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<subSubSection pageId="0" pageNumber="55" type="nomenclature">
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<paragraph pageId="0" pageNumber="55">
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<taxonomicName LSID="F6F0B4DF-411C-5A9B-9BA3-B484B07774F1" authority="(Carter & Zeck, 1932)" baseAuthorityName="Carter & Zeck" baseAuthorityYear="1932" class="Insecta" family="Elmidae" genus="Hydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydora laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">Hydora laticeps (Carter & Zeck, 1932)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="55" type="description">
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<paragraph pageId="0" pageNumber="55">
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<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Geographical distribution of species records 1 Australara glaisteri gen. nov., sp. nov. 2 Hydora laticeps 3 Ovolara australis 4 Ovolara lawrencei sp. nov. 5 Ovolara leai 6 Ovolara monteithi sp. nov. 7 Potamophilinus papuanus 8 Stetholus carinatus sp. nov. 9 Stetholus elongatus 10 Stetholus longipennis sp. nov. 11 Stetholus metatibialis sp. nov. 12 Stetholus woronora sp. nov." figureDoi="10.3897/zookeys.1073.71843.figures1-12" httpUri="https://binary.pensoft.net/fig/615728" pageId="0" pageNumber="55">Figs 2</figureCitation>
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<figureCitation captionStart="Figure 15" captionStartId="F3" captionText="Figure 15. Hydora laticeps, lectotype male A specimen labels B dorsal habitus C ventral head and prosternum D lateral head and pronotum (photographs courtesy of the Australian Museum, Natalie Tees)." figureDoi="10.3897/zookeys.1073.71843.figure15" httpUri="https://binary.pensoft.net/fig/615730" pageId="0" pageNumber="55">, 15</figureCitation>
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<figureCitation captionStart="Figure 16" captionStartId="F4" captionText="Figure 16. Hydora laticeps, lectotype; male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figure16" httpUri="https://binary.pensoft.net/fig/615731" pageId="0" pageNumber="55">, 16</figureCitation>
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<figureCitation captionStart="Figures 17–19" captionStartId="F5" captionText="Figures 17 - 19. Hydora laticeps, paralectotype males; dorsal habitus with specimen labels 17> AM specimen (photograph courtesy of the Australian Museum, Natalie Tees) 18 ANIC specimen (photograph courtesy of Vit Sykora, Charles University, Prague, Czech Republic) 19 ANIC specimen (photograph courtesy of the Australian National Insect Collection, CSIRO)." figureDoi="10.3897/zookeys.1073.71843.figures17-19" httpUri="https://binary.pensoft.net/fig/615732" pageId="0" pageNumber="55">, 17-19</figureCitation>
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<figureCitation captionStart="Figures 20, 21" captionStartId="F6" captionText="Figures 20, 21. Hydora laticeps, non-types from Lyneham, ACT 20 female habitus, 4.5 mm long A dorsal B ventral 21 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures20-21" httpUri="https://binary.pensoft.net/fig/615733" pageId="0" pageNumber="55">, 20, 21</figureCitation>
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<figureCitation captionStart="Figure 22" captionStartId="F7" captionText="Figure 22. Hydora laticeps, non-type female from Cann River, VIC A dorsal habitus B ventral habitus C lateral habitus (photographs courtesy of the Queensland Museum, Geoff Thompson)." figureDoi="10.3897/zookeys.1073.71843.figure22" httpUri="https://binary.pensoft.net/fig/615734" pageId="0" pageNumber="55">, 22</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="55" type="type locality">
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<paragraph pageId="0" pageNumber="55">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="55">
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Upper Shoalhaven River, Tallong;
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<geoCoordinate degrees="34.700" direction="south" orientation="latitude" precision="55" value="-34.7">34.700°S</geoCoordinate>
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,
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<geoCoordinate degrees="150.083" direction="east" orientation="longitude" precision="55" value="150.083">150.083°E</geoCoordinate>
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(approximate); New South Wales, Australia (lectotype deposited in the Australian Museum, Sydney). Note: The geographic coordinates given in the>AM database place the type locality north of Tallong, whereas the Shoalhaven River is to the south.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="55" type="type material">
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<paragraph pageId="0" pageNumber="55">Type material examined</paragraph>
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<paragraph pageId="0" pageNumber="55">
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<emphasis bold="true" pageId="0" pageNumber="55">
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(2).
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<emphasis italics="true" pageId="0" pageNumber="55">Lectotype</emphasis>
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male (here designated). New South Wales.
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</emphasis>
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"Australian Museum / K 579881 // Tallong / N.S.W. / FHTaylor //
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<taxonomicName authorityName="Carter & Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">Stetholus</taxonomicName>
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/
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<taxonomicName authorityName="Carter & Zeck" authorityYear="1932" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">Stetholus laticeps</taxonomicName>
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/ Carter & Zeck / Id. by H. J. Carter // K67434 // HOLOTYPE /
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<taxonomicName authorityName="Carter & Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">Stetholus</taxonomicName>
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/
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<taxonomicName authorityName="Carter & Zeck" authorityYear="1932" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">Stetholus laticeps</taxonomicName>
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/ Carter & Zeck, / 1932 [red label] //
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<taxonomicName authorityName="Anon. [Broun" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydora" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">Hydora</taxonomicName>
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/
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<taxonomicName baseAuthorityName="Carter & Zeck" baseAuthorityYear="1932" class="Insecta" family="Elmidae" genus="Hydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydora laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">Hydora laticeps</taxonomicName>
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/ (Carter & Zeck) / det. A.Calder 1999 // LECTOTYPE /
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<taxonomicName authorityName="Carter & Zeck" authorityYear="1932" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">Stetholus laticeps</taxonomicName>
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/ Carter & Zeck, 1932" [red label, handwritten]. Deposited in the Australian Museum, Sydney.
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<emphasis bold="true" pageId="0" pageNumber="55">
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<emphasis italics="true" pageId="0" pageNumber="55">Paralectotype</emphasis>
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male (here designated). New South Wales.
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</emphasis>
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Australian Museum / K 579882 // Tallong / N.S.W. / FHTaylor // K69264 //
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<taxonomicName authorityName="Carter & Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">Stetholus</taxonomicName>
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/
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<taxonomicName authorityName="Carter & Zeck" authorityYear="1932" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">Stetholus laticeps</taxonomicName>
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/ Carter 1932 // PARATYPE /
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<taxonomicName authorityName="Carter & Zeck" authorityYear="1932" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">Stetholus laticeps</taxonomicName>
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/ Carter & Zeck, / 1932 [blue label] //
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<taxonomicName authorityName="Anon. [Broun" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydora" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">Hydora</taxonomicName>
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/
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<taxonomicName baseAuthorityName="Carter & Zeck" baseAuthorityYear="1932" class="Insecta" family="Elmidae" genus="Hydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydora laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">Hydora laticeps</taxonomicName>
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/ (Carter & Zeck) / det. A.Calder 1999 // PARALECTOTYPE /
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<taxonomicName authorityName="Carter & Zeck" authorityYear="1932" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">
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<emphasis italics="true" pageId="0" pageNumber="55">Stetholus laticeps</emphasis>
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</taxonomicName>
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/ (Carter & Zeck, 1932) [yellow label, printed]. Deposited in the Australian Museum, Sydney.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="55" type="type material">
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<paragraph pageId="0" pageNumber="55">Type material examined from photographs</paragraph>
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<paragraph pageId="0" pageNumber="55">
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<emphasis bold="true" pageId="0" pageNumber="55">
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(2).
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<emphasis italics="true" pageId="0" pageNumber="55">Paralectotype</emphasis>
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males (here designated). New South Wales.
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</emphasis>
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Tallong / N.S.W. / FHTaylor //
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<taxonomicName authorityName="Carter & Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">Stetholus</taxonomicName>
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/
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<taxonomicName authorityName="Carter & Zeck" authorityYear="1932" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">Stetholus laticeps</taxonomicName>
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C & Z / Id. by H. J. Carter //
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<taxonomicName authorityName="Anon. [Broun" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydora" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">Hydora</taxonomicName>
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/
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<taxonomicName baseAuthorityName="Carter & Zeck" baseAuthorityYear="1932" class="Insecta" family="Elmidae" genus="Hydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydora laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">Hydora laticeps</taxonomicName>
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(C&Z) / det. A.
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<bibRefCitation author="Calder, AA" journalOrPublisher="The Pan-Pacific Entomologist" pageId="0" pageNumber="55" refId="B4" refString="Calder, AA, 1992. Illustrated keys and notes to the genera of adult Australian Elmidae. Coleopteran Identification Workshop, Murray-Darling Freshwater Research Centre, Albury (New South Wales, Australia), February 1992. [Unpublished]" title="Illustrated keys and notes to the genera of adult Australian Elmidae. Coleopteran Identification Workshop, Murray-Darling Freshwater Research Centre, Albury (New South Wales, Australia), February 1992. [Unpublished]" year="1992">Calder 1992</bibRefCitation>
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// Genitalia prep. / HH-224 ♂/ A. Calder 198792 // PARALECTOTYPE /
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<taxonomicName authorityName="Carter & Zeck" authorityYear="1932" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">
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<emphasis italics="true" pageId="0" pageNumber="55">Stetholus laticeps</emphasis>
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</taxonomicName>
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/ (Carter & Zeck, 1932) [yellow label, printed] (1 ANIC); Tallong / N.S.W. / FHTaylor //
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<taxonomicName authorityName="Carter & Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">Stetholus</taxonomicName>
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/
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<taxonomicName authorityName="Carter & Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus elongatus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="elongatus">Stetholus elongatus</taxonomicName>
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/ C & Z / Id. by H. J. Carter // Paratype [blue label, printed] // PARATYPE [blue label, printed] // Genitalia prep. / HH-247 ♂/ A.Calder 1997 // ANIC / Image // PARALECTOTYPE /
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<taxonomicName authorityName="Carter & Zeck" authorityYear="1932" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">
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<emphasis italics="true" pageId="0" pageNumber="55">Stetholus laticeps</emphasis>
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</taxonomicName>
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/ (Carter & Zeck, 1932) [yellow label, printed] (1 ANIC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="55" type="materials_examined">
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<paragraph pageId="0" pageNumber="55">Other material examined</paragraph>
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<paragraph pageId="0" pageNumber="55">
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<emphasis bold="true" pageId="0" pageNumber="55">
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(4).
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<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="A. C., B. P. Moore" country="AUSTRALIA" location="Australian Capital Territory" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Australian Capital Territory">
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<collectingRegion country="Australia" name="Australian Capital Territory">Australian Capital Territory</collectingRegion>
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.
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<collectingCountry name="Australia">AUSTRALIA</collectingCountry>
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: / Lyneham / at light /
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<collectorName name="AA Calder">A.C.</collectorName>
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T. 22.xii.66 /
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<collectorName>B.P.Moore</collectorName>
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(
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<specimenCount type="male">1♂</specimenCount>
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<specimenCount type="female">1♀</specimenCount>
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ANIC)
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</materialsCitation>
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</emphasis>
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.
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<materialsCitation collectingDate="1967-01-28" collectorName="Cann River, E., G. Monteith" country="Australia" location="Cann River" specimenCount="2" specimenCount-female="2" stateProvince="Victoria">
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<emphasis bold="true" pageId="0" pageNumber="55">
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<collectingRegion country="Australia" name="Victoria">Victoria</collectingRegion>
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.
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</emphasis>
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<collectorName>
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<location LSID="urn:lsid:plazi:treatment:F6F0B4DF411C5A9B9BA3B484B07774F1:04A917507088CA06A91E76711D571FC8" country="Australia" name="Cann River" stateProvince="Victoria">Cann River</location>
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, E.
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</collectorName>
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Vic. /
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<collectingDate value="1967-01-28">28.i.1967</collectingDate>
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. /
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<collectorName>G. Monteith</collectorName>
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// EX UQIC / DONATED / 2011 (
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<specimenCount type="female">2♀♀</specimenCount>
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QM)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="55" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="55">Differential diagnosis</paragraph>
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<paragraph pageId="0" pageNumber="55">
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<emphasis bold="true" pageId="0" pageNumber="55">(n = 8).</emphasis>
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<taxonomicName baseAuthorityName="Carter & Zeck" baseAuthorityYear="1932" class="Insecta" family="Elmidae" genus="Hydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydora laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">
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||
<emphasis italics="true" pageId="0" pageNumber="55">Hydora laticeps</emphasis>
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||
</taxonomicName>
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||
(Figs
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||
<figureCitation captionStart="Figure 15" captionStartId="F3" captionText="Figure 15. Hydora laticeps, lectotype male A specimen labels B dorsal habitus C ventral head and prosternum D lateral head and pronotum (photographs courtesy of the Australian Museum, Natalie Tees)." figureDoi="10.3897/zookeys.1073.71843.figure15" httpUri="https://binary.pensoft.net/fig/615730" pageId="0" pageNumber="55">15</figureCitation>
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||
-
|
||
<figureCitation captionStart="Figure 22" captionStartId="F7" captionText="Figure 22. Hydora laticeps, non-type female from Cann River, VIC A dorsal habitus B ventral habitus C lateral habitus (photographs courtesy of the Queensland Museum, Geoff Thompson)." figureDoi="10.3897/zookeys.1073.71843.figure22" httpUri="https://binary.pensoft.net/fig/615734" pageId="0" pageNumber="55">22</figureCitation>
|
||
) is the only species of
|
||
<taxonomicName authorityName="Anon. [Broun" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydora" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Hydora</emphasis>
|
||
</taxonomicName>
|
||
known to occur in Australia. It can be distinguished from other Australian laraines by a combination of the following characters: Eyes protuberant, hemispherical; maxillary palpi narrow at the apices; pronotum with strong basal, sublateral carinae and without a distinct transverse impression at anterior 1/3; and prosternum moderately long anterior to the coxae but not extending beneath head.
|
||
<taxonomicName authorityName="Carter & Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Stetholus</emphasis>
|
||
</taxonomicName>
|
||
species (Figs
|
||
<figureCitation captionStart="Figures 34, 35" captionStartId="F14" captionText="Figures 34, 35. Stetholus carinatus sp. nov., holotype male 34 habitus, 3.7 mm long A dorsal B ventral 35 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures34-35" httpUri="https://binary.pensoft.net/fig/615741" pageId="0" pageNumber="55">34</figureCitation>
|
||
-
|
||
<figureCitation captionStart="Figures 41, 42" captionStartId="F18" captionText="Figures 41, 42. Stetholus woronora sp. nov., holotype male 41 habitus, 5.2 mm long A dorsal B ventral 42 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures41-42" httpUri="https://binary.pensoft.net/fig/615745" pageId="0" pageNumber="55">42</figureCitation>
|
||
) have ovoid eyes, not usually prominent; maxillary palpi each with palpomere 4 wide and oblique at the apex; pronotum with a distinct transverse impression; and prosternum very short and narrow anterior to the coxae.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
<taxonomicName authorityName="Barr & Shepard" authorityYear="2021" family="Elmidae" genus="Australara" kingdom="Animalia" lsidName="Australara glaisteri" order="Coleoptera" pageId="0" pageNumber="55" rank="species" species="glaisteri">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Australara glaisteri</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation captionStart="Figures 13, 14" captionStartId="F2" captionText="Figures 13, 14. Australara glaisteri gen. nov., sp. nov., male 13 habitus A dorsal B ventral (photographs courtesy of the Australian National Insect Collection, CSIRO, Zhenhua Liu) 14 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures13-14" httpUri="https://binary.pensoft.net/fig/615729" pageId="0" pageNumber="55">13</figureCitation>
|
||
) most obviously differs by its lack of sublateral pronotal carinae and by the mesoventrite having an anterior projection containing a slit-like mesoventral cavity; the eyes are also not quite as protuberant.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="55" type="redescription">
|
||
<paragraph pageId="0" pageNumber="55">Redescription</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
<emphasis bold="true" pageId="0" pageNumber="55">
|
||
(n = 2). Male lectotype and male paralectotype.
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Body</emphasis>
|
||
</emphasis>
|
||
: Size 4.2 mm long, 1.6 mm wide (lectotype); size 4.0 mm long, 1.5 mm wide (paralectotype); elongate, parallel-sided. Color light to dark brown; head and pronotum darkest; antennae, mouthparts, legs, venter lightest. Dorsum with fine, pale setae, short on elytra, longer on head and pronotum; venter with long, dense setae.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Head</emphasis>
|
||
: Eye large, protuberant, hemispherical. Antenna with antennomere 1 elongate, antennomere 2 ovoid, antennomeres 3-11 smaller, weakly clavate. Labrum emarginate anteriorly, lateral margins with long setae. Maxillary palpus long, robust, setose; palpomere 4 much enlarged, ovoid, apex blunt with small, oval sensory area. Labial palpus shorter, palpomere 4 conical, apex pointed with very small, circular sensory area.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Pronotum</emphasis>
|
||
: Shape generally trapezoidal, 0.9 mm long, 1.1 mm wide (at base); anterior angles obscure, lateral margins crenulate, posterior angles acute, depressed; disc weakly sculptured except for two distinct, basal, sublateral carinae, 1/2 the pronotal length; two shallow, obscure transverse impressions laterad of midline at anterior 1/5-1/4.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Elytron</emphasis>
|
||
: 3.2-3.3 mm long, 0.7-0.8 mm wide (at base); lateral margin narrowly marginate, apex narrowly rounded, acute; disc with ten rows of moderately striate punctures, accessory basal stria present between striae 1 and 2; disc in lateral view flattened at anterior 1/2.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Prosternum</emphasis>
|
||
: Moderately long anterior to coxae, not extending beneath head; prosternal process narrow, curved, posterior 1/3 semi-carinate with a short, faint row of granules at midline, tip narrowly rounded.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Mesoventrite</emphasis>
|
||
: Longer than prosternum; mesoventral cavity deep and moderately wide
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Metaventrite</emphasis>
|
||
: Very convex, especially in lateral view.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Legs</emphasis>
|
||
: Long and slender. Tibia of all legs with a pair of stout spines at ventral apex; meso- and metatibia with posterior surfaces shallowly sulcate, glabrous, shiny. Tarsus with tarsomere 5 shorter than tarsomeres 1-4 combined; covered with short, dense setae; claws simple, slender, acute.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Abdomen</emphasis>
|
||
: Ventrite 1 triangular intercoxal projection moderately narrow; ventrite 5 nearly truncate at apex.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Aedeagus</emphasis>
|
||
: Phallobase longer than parameres and penis, penis slightly longer than parameres (Fig.
|
||
<figureCitation captionStart="Figure 16" captionStartId="F4" captionText="Figure 16. Hydora laticeps, lectotype; male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figure16" httpUri="https://binary.pensoft.net/fig/615731" pageId="0" pageNumber="55">16</figureCitation>
|
||
). Phallobase open dorsally. In dorsal view (Fig.
|
||
<figureCitation captionStart="Figure 16" captionStartId="F4" captionText="Figure 16. Hydora laticeps, lectotype; male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figure16" httpUri="https://binary.pensoft.net/fig/615731" pageId="0" pageNumber="55">16A</figureCitation>
|
||
), parameres broad, with lateral margins gradually convergent, apices bluntly rounded; medial margins parallel-sided at basal 2/3 then gradually divergent, margins appearing more sclerotized than rest of parameres. Penis slightly longer than parameres, approximately as wide at base as paramere base; lateral margins widened and arcuate just distal to base, then evenly convergent to apex; apex narrow, nipple-like, laterally flattened, tip narrowly rounded; no corona visible; basal apophyses short, 1/4-1/3 as long as phallobase, straight, broad, blunt at tips. Fibula absent. In lateral view (Fig.
|
||
<figureCitation captionStart="Figure 16" captionStartId="F4" captionText="Figure 16. Hydora laticeps, lectotype; male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figure16" httpUri="https://binary.pensoft.net/fig/615731" pageId="0" pageNumber="55">16B</figureCitation>
|
||
), paramere nearly straight dorsally at apical 3/4, weakly arcuate ventrally, tip broadly rounded and slightly wider than paramere tip.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1073.71843.figure16" httpUri="https://binary.pensoft.net/fig/615731" pageId="0" pageNumber="55" start="Figure 16" startId="F4">
|
||
<paragraph pageId="0" pageNumber="55">
|
||
<emphasis bold="true" pageId="0" pageNumber="55">Figure 16.</emphasis>
|
||
<taxonomicName baseAuthorityName="Carter & Zeck" baseAuthorityYear="1932" class="Insecta" family="Elmidae" genus="Hydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydora laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Hydora laticeps</emphasis>
|
||
</taxonomicName>
|
||
, lectotype; male genitalia
|
||
<emphasis bold="true" pageId="0" pageNumber="55">A</emphasis>
|
||
dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="55">B</emphasis>
|
||
lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="55">C</emphasis>
|
||
ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="55" type="variation">
|
||
<paragraph pageId="0" pageNumber="55">Variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
There is some size variation among the known specimens, particularly between males and females. Specimens from the type series, all males (n = 4), measured 4.0-4.3 mm long, 1.4-1.6 mm wide.
|
||
<bibRefCitation author="Carter, HJ" journalOrPublisher="The Australian Zoologist" pageId="0" pageNumber="55" pagination="202 - 205" refId="B7" refString="Carter, HJ, Zeck, EH, 1932. Four new species of Dryopidae, together with notes on the family. The Australian Zoologist 7 (3): 202 - 205" title="Four new species of Dryopidae, together with notes on the family." volume="7" year="1932">Carter and Zeck (1932)</bibRefCitation>
|
||
stated in the type description "Dimensions: 5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
1.5 mm" but the length probably included the head. Among the specimens examined (including two from the type series), the females (n = 3), 4.5-5.2 mm long, 1.7-1.8 mm wide, are considerably larger than the males (n = 3), 4.0-4.5 mm long, 1.4-1.6 mm wide. In addition, the females (Figs
|
||
<figureCitation captionStart="Figures 20, 21" captionStartId="F6" captionText="Figures 20, 21. Hydora laticeps, non-types from Lyneham, ACT 20 female habitus, 4.5 mm long A dorsal B ventral 21 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures20-21" httpUri="https://binary.pensoft.net/fig/615733" pageId="0" pageNumber="55">20B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 22" captionStartId="F7" captionText="Figure 22. Hydora laticeps, non-type female from Cann River, VIC A dorsal habitus B ventral habitus C lateral habitus (photographs courtesy of the Queensland Museum, Geoff Thompson)." figureDoi="10.3897/zookeys.1073.71843.figure22" httpUri="https://binary.pensoft.net/fig/615734" pageId="0" pageNumber="55">22B</figureCitation>
|
||
) have prosternal processes broader than those of the males (Fig.
|
||
<figureCitation captionStart="Figure 15" captionStartId="F3" captionText="Figure 15. Hydora laticeps, lectotype male A specimen labels B dorsal habitus C ventral head and prosternum D lateral head and pronotum (photographs courtesy of the Australian Museum, Natalie Tees)." figureDoi="10.3897/zookeys.1073.71843.figure15" httpUri="https://binary.pensoft.net/fig/615730" pageId="0" pageNumber="55">15C</figureCitation>
|
||
), and noticeably narrower maxillary palpi. The prosternal processes of the two male specimens examined from the type series (Fig.
|
||
<figureCitation captionStart="Figure 15" captionStartId="F3" captionText="Figure 15. Hydora laticeps, lectotype male A specimen labels B dorsal habitus C ventral head and prosternum D lateral head and pronotum (photographs courtesy of the Australian Museum, Natalie Tees)." figureDoi="10.3897/zookeys.1073.71843.figure15" httpUri="https://binary.pensoft.net/fig/615730" pageId="0" pageNumber="55">15C</figureCitation>
|
||
) are slightly narrower than those of the non-type male. The surface of the prosternal process varies, and may be convex, depressed only between the procoxae, or entirely flat except posterior to coxae, but in all specimens the process is granulate, swollen, and an indistinct carina is usually visible. Non-sexual variation was also observed in the morphology of the elytral punctures (size and depth), pronotum (width, lateral margins, posterior angles, sculpturing); and prosternal process (width, surface features). On the pronotum, two shallow, anterior, transverse impressions are present laterad of the midline. In most specimens the impressions are weak or altogether obscure (Figs
|
||
<figureCitation captionStart="Figures 17–19" captionStartId="F5" captionText="Figures 17 - 19. Hydora laticeps, paralectotype males; dorsal habitus with specimen labels 17> AM specimen (photograph courtesy of the Australian Museum, Natalie Tees) 18 ANIC specimen (photograph courtesy of Vit Sykora, Charles University, Prague, Czech Republic) 19 ANIC specimen (photograph courtesy of the Australian National Insect Collection, CSIRO)." figureDoi="10.3897/zookeys.1073.71843.figures17-19" httpUri="https://binary.pensoft.net/fig/615732" pageId="0" pageNumber="55">17-19</figureCitation>
|
||
), but they are quite obvious in one of the two non-type specimens from Cann River (Fig.
|
||
<figureCitation captionStart="Figure 22" captionStartId="F7" captionText="Figure 22. Hydora laticeps, non-type female from Cann River, VIC A dorsal habitus B ventral habitus C lateral habitus (photographs courtesy of the Queensland Museum, Geoff Thompson)." figureDoi="10.3897/zookeys.1073.71843.figure22" httpUri="https://binary.pensoft.net/fig/615734" pageId="0" pageNumber="55">22A</figureCitation>
|
||
). In addition, the single male non-type specimen from Lyneham (Fig.
|
||
<figureCitation captionStart="Figures 20, 21" captionStartId="F6" captionText="Figures 20, 21. Hydora laticeps, non-types from Lyneham, ACT 20 female habitus, 4.5 mm long A dorsal B ventral 21 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures20-21" httpUri="https://binary.pensoft.net/fig/615733" pageId="0" pageNumber="55">21A, C</figureCitation>
|
||
) has a slightly broader aedeagus than the two specimens examined from the type series (Fig.
|
||
<figureCitation captionStart="Figure 16" captionStartId="F4" captionText="Figure 16. Hydora laticeps, lectotype; male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figure16" httpUri="https://binary.pensoft.net/fig/615731" pageId="0" pageNumber="55">16A, C</figureCitation>
|
||
). This variability in external morphology and male genitalia raises the possibility that more than one species is involved.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1073.71843.figures17-19" httpUri="https://binary.pensoft.net/fig/615732" pageId="0" pageNumber="55" start="Figures 17–19" startId="F5">
|
||
<paragraph pageId="0" pageNumber="55">
|
||
<emphasis bold="true" pageId="0" pageNumber="55">Figures 17-19.</emphasis>
|
||
<taxonomicName baseAuthorityName="Carter & Zeck" baseAuthorityYear="1932" class="Insecta" family="Elmidae" genus="Hydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydora laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Hydora laticeps</emphasis>
|
||
</taxonomicName>
|
||
, paralectotype males; dorsal habitus with specimen labels
|
||
<emphasis bold="true" pageId="0" pageNumber="55">17</emphasis>
|
||
>AM specimen (photograph courtesy of the Australian Museum, Natalie Tees)
|
||
<emphasis bold="true" pageId="0" pageNumber="55">18</emphasis>
|
||
ANIC specimen (photograph courtesy of
|
||
<normalizedToken originalValue="Vít">Vit</normalizedToken>
|
||
<normalizedToken originalValue="Sýkora">Sykora</normalizedToken>
|
||
, Charles University, Prague, Czech Republic)
|
||
<emphasis bold="true" pageId="0" pageNumber="55">19</emphasis>
|
||
ANIC specimen (photograph courtesy of the Australian National Insect Collection, CSIRO).
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1073.71843.figures20-21" httpUri="https://binary.pensoft.net/fig/615733" pageId="0" pageNumber="55" start="Figures 20, 21" startId="F6">
|
||
<paragraph pageId="0" pageNumber="55">
|
||
<emphasis bold="true" pageId="0" pageNumber="55">Figures 20, 21.</emphasis>
|
||
<taxonomicName baseAuthorityName="Carter & Zeck" baseAuthorityYear="1932" class="Insecta" family="Elmidae" genus="Hydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydora laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Hydora laticeps</emphasis>
|
||
</taxonomicName>
|
||
, non-types from Lyneham, ACT
|
||
<emphasis bold="true" pageId="0" pageNumber="55">20</emphasis>
|
||
female habitus, 4.5 mm long
|
||
<emphasis bold="true" pageId="0" pageNumber="55">A</emphasis>
|
||
dorsal
|
||
<emphasis bold="true" pageId="0" pageNumber="55">B</emphasis>
|
||
ventral
|
||
<emphasis bold="true" pageId="0" pageNumber="55">21</emphasis>
|
||
male genitalia
|
||
<emphasis bold="true" pageId="0" pageNumber="55">A</emphasis>
|
||
dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="55">B</emphasis>
|
||
lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="55">C</emphasis>
|
||
ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1073.71843.figure22" httpUri="https://binary.pensoft.net/fig/615734" pageId="0" pageNumber="55" start="Figure 22" startId="F7">
|
||
<paragraph pageId="0" pageNumber="55">
|
||
<emphasis bold="true" pageId="0" pageNumber="55">Figure 22.</emphasis>
|
||
<taxonomicName baseAuthorityName="Carter & Zeck" baseAuthorityYear="1932" class="Insecta" family="Elmidae" genus="Hydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydora laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Hydora laticeps</emphasis>
|
||
</taxonomicName>
|
||
, non-type female from Cann River, VIC
|
||
<emphasis bold="true" pageId="0" pageNumber="55">A</emphasis>
|
||
dorsal habitus
|
||
<emphasis bold="true" pageId="0" pageNumber="55">B</emphasis>
|
||
ventral habitus
|
||
<emphasis bold="true" pageId="0" pageNumber="55">C</emphasis>
|
||
lateral habitus (photographs courtesy of the Queensland Museum, Geoff Thompson).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="55" type="lectotype designation">
|
||
<paragraph pageId="0" pageNumber="55">Lectotype designation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
It appears that none of the four known specimens from the type series of
|
||
<taxonomicName authorityName="Carter & Zeck" authorityYear="1932" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Stetholus laticeps</emphasis>
|
||
</taxonomicName>
|
||
was given a holotype or paratype label at the time of description, and those subsequently added to the specimens were not done so by the authors. In their description,
|
||
<bibRefCitation author="Carter, HJ" journalOrPublisher="The Australian Zoologist" pageId="0" pageNumber="55" pagination="202 - 205" refId="B7" refString="Carter, HJ, Zeck, EH, 1932. Four new species of Dryopidae, together with notes on the family. The Australian Zoologist 7 (3): 202 - 205" title="Four new species of Dryopidae, together with notes on the family." volume="7" year="1932">Carter and Zeck (1932)</bibRefCitation>
|
||
stated that they had "five examples" and that the holotype was "in Coll. Carter," but did not mention designating paratypes. Two specimens were deposited at the Australian Museum 35 years apart: According to the original register of specimens, the first (Fig.
|
||
<figureCitation captionStart="Figure 15" captionStartId="F3" captionText="Figure 15. Hydora laticeps, lectotype male A specimen labels B dorsal habitus C ventral head and prosternum D lateral head and pronotum (photographs courtesy of the Australian Museum, Natalie Tees)." figureDoi="10.3897/zookeys.1073.71843.figure15" httpUri="https://binary.pensoft.net/fig/615730" pageId="0" pageNumber="55">15</figureCitation>
|
||
) was presented by H. J. Carter in 1936, and bears an old, handwritten determination label saying "
|
||
<taxonomicName authorityName="Carter & Zeck" authorityYear="1932" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">Stetholus laticeps</taxonomicName>
|
||
Carter & Zeck Id. by H. J. Carter" (Fig.
|
||
<figureCitation captionStart="Figure 15" captionStartId="F3" captionText="Figure 15. Hydora laticeps, lectotype male A specimen labels B dorsal habitus C ventral head and prosternum D lateral head and pronotum (photographs courtesy of the Australian Museum, Natalie Tees)." figureDoi="10.3897/zookeys.1073.71843.figure15" httpUri="https://binary.pensoft.net/fig/615730" pageId="0" pageNumber="55">15A</figureCitation>
|
||
). The specimen was subsequently given a holotype label by an unknown person, i.e., it was not written in the same hand as the determination label by Carter and appears much newer. This specimen is here designated as the lectotype to fix the concept of
|
||
<taxonomicName baseAuthorityName="Carter & Zeck" baseAuthorityYear="1932" class="Insecta" family="Elmidae" genus="Hydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydora laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Hydora laticeps</emphasis>
|
||
</taxonomicName>
|
||
(Carter & Zeck). The second specimen (Fig.
|
||
<figureCitation captionStart="Figures 17–19" captionStartId="F5" captionText="Figures 17 - 19. Hydora laticeps, paralectotype males; dorsal habitus with specimen labels 17> AM specimen (photograph courtesy of the Australian Museum, Natalie Tees) 18 ANIC specimen (photograph courtesy of Vit Sykora, Charles University, Prague, Czech Republic) 19 ANIC specimen (photograph courtesy of the Australian National Insect Collection, CSIRO)." figureDoi="10.3897/zookeys.1073.71843.figures17-19" httpUri="https://binary.pensoft.net/fig/615732" pageId="0" pageNumber="55">17</figureCitation>
|
||
) was obtained from the late E. H. Zeck in 1971, lacks an original determination label, and likewise bears a newer paratype label; it is designated a paralectotype.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
There are also two specimens housed at Australian National Insect Collection. One of them bears the surprising, original determination label "
|
||
<taxonomicName authorityName="Carter & Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus elongatus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="elongatus">Stetholus elongatus</taxonomicName>
|
||
C & Z Id. by H. J. Carter" and two printed paratype labels, one older and one newer (Fig.
|
||
<figureCitation captionStart="Figures 17–19" captionStartId="F5" captionText="Figures 17 - 19. Hydora laticeps, paralectotype males; dorsal habitus with specimen labels 17> AM specimen (photograph courtesy of the Australian Museum, Natalie Tees) 18 ANIC specimen (photograph courtesy of Vit Sykora, Charles University, Prague, Czech Republic) 19 ANIC specimen (photograph courtesy of the Australian National Insect Collection, CSIRO)." figureDoi="10.3897/zookeys.1073.71843.figures17-19" httpUri="https://binary.pensoft.net/fig/615732" pageId="0" pageNumber="55">19</figureCitation>
|
||
). Probably the identification predated the description of
|
||
<taxonomicName lsidName="S. laticeps" pageId="0" pageNumber="55" rank="species" species="laticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">S. laticeps</emphasis>
|
||
</taxonomicName>
|
||
by Carter and Zeck in 1932, however, Tallong was not among the localities cited in their 1929 description of
|
||
<taxonomicName lsidName="S. elongatus" pageId="0" pageNumber="55" rank="species" species="elongatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">S. elongatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Carter, HJ" journalOrPublisher="The Australian Zoologist" pageId="0" pageNumber="55" pagination="179 - 190" refId="B6" refString="Carter, HJ, Zeck, EH, 1929. A monograph of the Australian Dryopidae. Order- Coleoptera. The Australian Zoologist 6 (1): 179 - 190" title="A monograph of the Australian Dryopidae. Order- Coleoptera." volume="6" year="1929">Carter and Zeck 1929</bibRefCitation>
|
||
). The other specimen has an original determination label, "
|
||
<taxonomicName authorityName="Carter & Zeck" authorityYear="1932" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">Stetholus laticeps</taxonomicName>
|
||
C & Z Id. by H. J. Carter," but has no paratype label (Fig.
|
||
<figureCitation captionStart="Figures 17–19" captionStartId="F5" captionText="Figures 17 - 19. Hydora laticeps, paralectotype males; dorsal habitus with specimen labels 17> AM specimen (photograph courtesy of the Australian Museum, Natalie Tees) 18 ANIC specimen (photograph courtesy of Vit Sykora, Charles University, Prague, Czech Republic) 19 ANIC specimen (photograph courtesy of the Australian National Insect Collection, CSIRO)." figureDoi="10.3897/zookeys.1073.71843.figures17-19" httpUri="https://binary.pensoft.net/fig/615732" pageId="0" pageNumber="55">18</figureCitation>
|
||
). These two specimens are likewise designated as paralectotypes. We were unable to examine the ANIC specimens because they were on loan to another researcher, but we were provided with habitus images (Figs
|
||
<figureCitation captionStart="Figures 17–19" captionStartId="F5" captionText="Figures 17 - 19. Hydora laticeps, paralectotype males; dorsal habitus with specimen labels 17> AM specimen (photograph courtesy of the Australian Museum, Natalie Tees) 18 ANIC specimen (photograph courtesy of Vit Sykora, Charles University, Prague, Czech Republic) 19 ANIC specimen (photograph courtesy of the Australian National Insect Collection, CSIRO)." figureDoi="10.3897/zookeys.1073.71843.figures17-19" httpUri="https://binary.pensoft.net/fig/615732" pageId="0" pageNumber="55">18</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 17–19" captionStartId="F5" captionText="Figures 17 - 19. Hydora laticeps, paralectotype males; dorsal habitus with specimen labels 17> AM specimen (photograph courtesy of the Australian Museum, Natalie Tees) 18 ANIC specimen (photograph courtesy of Vit Sykora, Charles University, Prague, Czech Republic) 19 ANIC specimen (photograph courtesy of the Australian National Insect Collection, CSIRO)." figureDoi="10.3897/zookeys.1073.71843.figures17-19" httpUri="https://binary.pensoft.net/fig/615732" pageId="0" pageNumber="55">19</figureCitation>
|
||
) and measurements of body length for this article.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
The location of the fifth specimen from the type series is unknown.
|
||
<bibRefCitation DOI="https://doi.org/10.1080/00779962.2014.963910" author="Lambert, P" journalOrPublisher="New Zealand Entomologist" pageId="0" pageNumber="55" pagination="1 - 20" refId="B17" refString="Lambert, P, Maier, CA, Leschen, RAB, 2014. A new species and lectotype designations for New Zealand Hydora (Elmidae: Larainae) with a world checklist. New Zealand Entomologist 2014: 1 - 20, DOI: https://doi.org/10.1080/00779962.2014.963910" title="A new species and lectotype designations for New Zealand Hydora (Elmidae: Larainae) with a world checklist." url="https://doi.org/10.1080/00779962.2014.963910" volume="2014" year="2014">Lambert et al. (2014)</bibRefCitation>
|
||
cited the SAMA as a specimen depository for the species, but we have examined their material and found no specimens of
|
||
<taxonomicName lsidName="H. laticeps" pageId="0" pageNumber="55" rank="species" species="laticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">H. laticeps</emphasis>
|
||
</taxonomicName>
|
||
, so this report was in error.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="55" type="distribution">
|
||
<paragraph pageId="0" pageNumber="55">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
<taxonomicName baseAuthorityName="Carter & Zeck" baseAuthorityYear="1932" class="Insecta" family="Elmidae" genus="Hydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydora laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Hydora laticeps</emphasis>
|
||
</taxonomicName>
|
||
is known from only three localities in Australia: the Shoalhaven River near Tallong, New South Wales, the type locality; Lyneham, Australian Capital Territory; and Cann River, eastern Victoria (Fig.
|
||
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Geographical distribution of species records 1 Australara glaisteri gen. nov., sp. nov. 2 Hydora laticeps 3 Ovolara australis 4 Ovolara lawrencei sp. nov. 5 Ovolara leai 6 Ovolara monteithi sp. nov. 7 Potamophilinus papuanus 8 Stetholus carinatus sp. nov. 9 Stetholus elongatus 10 Stetholus longipennis sp. nov. 11 Stetholus metatibialis sp. nov. 12 Stetholus woronora sp. nov." figureDoi="10.3897/zookeys.1073.71843.figures1-12" httpUri="https://binary.pensoft.net/fig/615728" pageId="0" pageNumber="55">2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="55" type="associated byrrhoid taxa">
|
||
<paragraph pageId="0" pageNumber="55">Associated byrrhoid taxa.</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
<taxonomicName authorityName="Curtis" authorityYear="1830" class="Insecta" family="Elmidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="family">Elmidae</taxonomicName>
|
||
:
|
||
<taxonomicName authorityName="LeConte" authorityYear="1861" lsidName="" pageId="0" pageNumber="55" rank="subFamily" subFamily="Larainae">Larainae</taxonomicName>
|
||
:
|
||
<taxonomicName authorityName="Carter & Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus elongatus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="elongatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Stetholus elongatus</emphasis>
|
||
</taxonomicName>
|
||
(>AM, ANIC, NMV, SAMA);
|
||
<taxonomicName authorityName="Curtis" authorityYear="1830" lsidName="" pageId="0" pageNumber="55" rank="subFamily" subFamily="Elminae">Elminae</taxonomicName>
|
||
:
|
||
<taxonomicName authorityName="Carter & Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Notriolus" kingdom="Animalia" lsidName="Notriolus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Notriolus</emphasis>
|
||
</taxonomicName>
|
||
sp. (>AM).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="55" type="comments">
|
||
<paragraph pageId="0" pageNumber="55">Comments.</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
<taxonomicName baseAuthorityName="Carter & Zeck" baseAuthorityYear="1932" class="Insecta" family="Elmidae" genus="Hydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydora laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Hydora laticeps</emphasis>
|
||
</taxonomicName>
|
||
was originally described in the genus
|
||
<taxonomicName authorityName="Carter & Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Stetholus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="Carter, HJ" journalOrPublisher="The Australian Zoologist" pageId="0" pageNumber="55" pagination="202 - 205" refId="B7" refString="Carter, HJ, Zeck, EH, 1932. Four new species of Dryopidae, together with notes on the family. The Australian Zoologist 7 (3): 202 - 205" title="Four new species of Dryopidae, together with notes on the family." volume="7" year="1932">Carter and Zeck (1932)</bibRefCitation>
|
||
, and subsequently reassigned to
|
||
<taxonomicName authorityName="Anon. [Broun" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydora" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Hydora</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-3113.1935.tb00660.x" author="Hinton, HE" journalOrPublisher="Stylops" pageId="0" pageNumber="55" pagination="169 - 179" refId="B13" refString="Hinton, HE, 1935. Notes on the Dryopoidea (Col.). Stylops 4: 169 - 179, DOI: https://doi.org/10.1111/j.1365-3113.1935.tb00660.x" title="Notes on the Dryopoidea (Col.)." url="https://doi.org/10.1111/j.1365-3113.1935.tb00660.x" volume="4" year="1935">Hinton (1935)</bibRefCitation>
|
||
. Hinton stated that he studied only the description and figures, not actual specimens, and gave no specific reasons for the new combination.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
In the diagnosis following their description,
|
||
<bibRefCitation author="Carter, HJ" journalOrPublisher="The Australian Zoologist" pageId="0" pageNumber="55" pagination="202 - 205" refId="B7" refString="Carter, HJ, Zeck, EH, 1932. Four new species of Dryopidae, together with notes on the family. The Australian Zoologist 7 (3): 202 - 205" title="Four new species of Dryopidae, together with notes on the family." volume="7" year="1932">Carter and Zeck (1932</bibRefCitation>
|
||
: 203) noted that the prosternal process of
|
||
<taxonomicName lsidName="S. laticeps" pageId="0" pageNumber="55" rank="species" species="laticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">S. laticeps</emphasis>
|
||
</taxonomicName>
|
||
lacks a carina, in contrast to
|
||
<taxonomicName authorityName="Carter & Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus elongatus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="elongatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Stetholus elongatus</emphasis>
|
||
</taxonomicName>
|
||
Carter & Zeck (1929) which has a carina. The actual situation is less clear-cut. The surface of the apical 1/2-1/3 of the process is convex to varying degrees and may be somewhat granulate at the midline, resembling an indistinct carina. However, this is a poor diagnostic character because dense setation can make examination difficult.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
When
|
||
<bibRefCitation author="Carter, HJ" journalOrPublisher="The Australian Zoologist" pageId="0" pageNumber="55" pagination="202 - 205" refId="B7" refString="Carter, HJ, Zeck, EH, 1932. Four new species of Dryopidae, together with notes on the family. The Australian Zoologist 7 (3): 202 - 205" title="Four new species of Dryopidae, together with notes on the family." volume="7" year="1932">Carter and Zeck (1932)</bibRefCitation>
|
||
described
|
||
<taxonomicName lsidName="S. laticeps" pageId="0" pageNumber="55" rank="species" species="laticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">S. laticeps</emphasis>
|
||
</taxonomicName>
|
||
they made no mention that
|
||
<taxonomicName lsidName="S. elongatus" pageId="0" pageNumber="55" rank="species" species="elongatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">S. elongatus</emphasis>
|
||
</taxonomicName>
|
||
, described by them in 1929, was present in the Upper Shoalhaven River as well. That
|
||
<taxonomicName lsidName="S. elongatus" pageId="0" pageNumber="55" rank="species" species="elongatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">S. elongatus</emphasis>
|
||
</taxonomicName>
|
||
was collected with
|
||
<taxonomicName lsidName="H. laticeps" pageId="0" pageNumber="55" rank="species" species="laticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">H. laticeps</emphasis>
|
||
</taxonomicName>
|
||
at the type locality is evidenced by museum specimens with locality labels identical to those of
|
||
<taxonomicName lsidName="H. laticeps" pageId="0" pageNumber="55" rank="species" species="laticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">H. laticeps</emphasis>
|
||
</taxonomicName>
|
||
: "Tallong N.S.W. FH Taylor." This raised the question as to whether the missing specimen of
|
||
<taxonomicName lsidName="H. laticeps" pageId="0" pageNumber="55" rank="species" species="laticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">H. laticeps</emphasis>
|
||
</taxonomicName>
|
||
might bear a
|
||
<taxonomicName lsidName="S. elongatus" pageId="0" pageNumber="55" rank="species" species="elongatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">S. elongatus</emphasis>
|
||
</taxonomicName>
|
||
label, as does one of the ANIC specimens, and thus has been overlooked. Unfortunately, examination of all known
|
||
<taxonomicName lsidName="S. elongatus" pageId="0" pageNumber="55" rank="species" species="elongatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">S. elongatus</emphasis>
|
||
</taxonomicName>
|
||
specimens with collection labels as above (>AM, 3 specimens; ANIC, 3; NMV, 2; SAMA, 4) revealed no misidentifications.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
Until now,
|
||
<taxonomicName baseAuthorityName="Carter & Zeck" baseAuthorityYear="1932" class="Insecta" family="Elmidae" genus="Hydora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydora laticeps" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="laticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Hydora laticeps</emphasis>
|
||
</taxonomicName>
|
||
has been known only from its type locality, the Upper Shoalhaven River near Tallong, New South Wales, Australia. In the 90+ years since the type series was collected, deliberate attempts to re-collect it have been unsuccessful. Examination of unidentified museum specimens for this project resulted in the discovery of four additional specimens from two new localities, all of which were collected at light. The four type specimens available are all males, the Lyneham specimens are male and female, and the Cann River specimens are both female. In the absence of males, the latter two specimens are assumed to be
|
||
<taxonomicName lsidName="H. laticeps" pageId="0" pageNumber="55" rank="species" species="laticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">H. laticeps</emphasis>
|
||
</taxonomicName>
|
||
due to external morphological similarities. The larva of the species is unknown.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
As mentioned in the Variation section, is possible that not all of the specimens are conspecific because of mophological variation which is apparent even among those from the type series. However, there is not enough evidence at present to assign any to a species other than
|
||
<taxonomicName lsidName="H. laticeps" pageId="0" pageNumber="55" rank="species" species="laticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">H. laticeps</emphasis>
|
||
</taxonomicName>
|
||
. DNA analysis would be helpful in this regard if fresh material could be obtained. A recent attempt to obtain DNA from a specimen in the type series failed due to its age (V.
|
||
<normalizedToken originalValue="Sýkora">Sykora</normalizedToken>
|
||
, in litt.), and even the youngest of the specimens is at least 54 years old.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |