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<document id="21F190211C027C99B57182D76610FC96" ID-DOI="10.11646/zootaxa.5040.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5531583" ID-ZooBank="FFFA7323-04AC-4B70-9DBE-6EAD909A9D2A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632749724544" checkinUser="plazi" docAuthor="Carrejo, Nancy, Obando, Ranulfo González, Casasola-González, José Arturo &amp; García Aldrete, Alfonso N." docDate="2021" docId="9946F564FFBFFFFDFF686E27FDEEFE49" docLanguage="en" docName="zootaxa.5040.4.1.pdf" docOrigin="Zootaxa 5040 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Goja andina Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete 2021, n. sp." docType="treatment" docVersion="3" lastPageNumber="463" masterDocId="657F8D1CFFB5FFF1FFFF6A5AFF8BFFBD" masterDocTitle="New Colombian Goja Navás (Psocodea: Psocoptera: Epipsocidae) with peculiar genitalia, and the first Goja with brachypterous male, from Oaxaca, Mexico" masterLastPageNumber="481" masterPageNumber="451" pageNumber="461" updateTime="1699415585501" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="58536C6DE09676D24A5C6008467C214B">New Colombian Goja Navás (Psocodea: Psocoptera: Epipsocidae) with peculiar genitalia, and the first Goja with brachypterous male, from Oaxaca, Mexico</mods:title>
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<mods:namePart id="E3FB286832B1075E4049E4E5C49D4872">Carrejo, Nancy</mods:namePart>
<mods:affiliation id="A9355CD9317C97BD0A15C2AB04FFBA8E">Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, COLOMBIA. Instituto de Estudios Ambientales, Universidad de la Sierra Juárez, Ixtlán de Juárez, Oaxaca, MÉXICO. &amp; nancy. carrejo @ correounivalle. edu. co; htps: // orcid. org / 0000 - 0002 - 2395 - 6887</mods:affiliation>
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<mods:namePart id="0F8BC3D3FB3A411903B35F16D0345292">Obando, Ranulfo González</mods:namePart>
<mods:affiliation id="2B8A9F48FE22D79F60C060871709F10B">Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, COLOMBIA. Instituto de Estudios Ambientales, Universidad de la Sierra Juárez, Ixtlán de Juárez, Oaxaca, MÉXICO. &amp; ranulfo. gonzález @ correounivalle. edu. co; htps: // orcid. org / 0000 - 0003 - 4047 - 8677</mods:affiliation>
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<mods:namePart id="EF6CB9FA48ADC876E25A3EA5F98AEE24">Casasola-González, José Arturo</mods:namePart>
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<mods:affiliation id="E3E59C7B55EC51628AA883FE14BEBAC8">casasola @ unsij. edu. mx; https: // orcid. org / 0000 - 0002 - 4765 - 6707</mods:affiliation>
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<mods:namePart id="3321FD2FDB4552DCA3EADA7D67DC654B">García Aldrete, Alfonso N.</mods:namePart>
<mods:affiliation id="CD4DAAB904253CE06B49CEBFA935453C">Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, CdMx, MÉXICO. &amp; anga @ ib. unam. mx; htps: // orcid. org / 0000 - 0001 - 7214 - 7966</mods:affiliation>
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<treatment id="9946F564FFBFFFFDFF686E27FDEEFE49" ID-DOI="http://doi.org/10.5281/zenodo.5531593" ID-Zenodo-Dep="5531593" LSID="urn:lsid:plazi:treatment:9946F564FFBFFFFDFF686E27FDEEFE49" httpUri="http://treatment.plazi.org/id/9946F564FFBFFFFDFF686E27FDEEFE49" lastPageId="12" lastPageNumber="463" pageId="10" pageNumber="461">
<subSubSection id="59F517F9FFBFFFFBFF686E27FE34FB2A" box="[151,447,1149,1176]" pageId="10" pageNumber="461" type="nomenclature">
<paragraph id="11504472FFBFFFFBFF686E27FE34FB2A" blockId="10.[151,447,1149,1212]" box="[151,447,1149,1176]" pageId="10" pageNumber="461">
<heading id="4A18F31EFFBFFFFBFF686E27FE34FB2A" bold="true" box="[151,447,1149,1176]" fontSize="11" level="1" pageId="10" pageNumber="461" reason="1">
<emphasis id="239B9860FFBFFFFBFF686E27FE34FB2A" bold="true" box="[151,447,1149,1176]" pageId="10" pageNumber="461">
<taxonomicName id="D6EF3FF1FFBFFFFBFF686E27FEA5FB25" ID-CoL="9J465" authority="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete, 2021" authorityName="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete" authorityYear="2021" box="[151,302,1149,1176]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="andina" status="sp. nov.">
<emphasis id="239B9860FFBFFFFBFF686E27FEA5FB25" bold="true" box="[151,302,1149,1176]" italics="true" pageId="10" pageNumber="461">Goja andina</emphasis>
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<taxonomicNameLabel id="38A8251BFFBFFFFBFECA6E27FEF3FB2A" box="[309,376,1149,1175]" pageId="10" pageNumber="461" rank="species">n. sp.</taxonomicNameLabel>
Male
</emphasis>
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</paragraph>
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<subSubSection id="59F517F9FFBFFFFBFF686EFBFE80FB01" box="[151,267,1185,1212]" pageId="10" pageNumber="461" type="description">
<paragraph id="11504472FFBFFFFBFF686EFBFE80FB01" blockId="10.[151,447,1149,1212]" box="[151,267,1185,1212]" pageId="10" pageNumber="461">
(
<figureCitation id="89D458F7FFBFFFFBFF606EFBFE89FB01" box="[159,258,1185,1212]" captionStart="FIGURES 16" captionStartId="11.[151,264,1851,1875]" captionTargetBox="[163,1427,259,1822]" captionTargetId="figure-42@11.[151,1436,234,1825]" captionTargetPageId="11" captionText="FIGURES 16. Goja andina n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531585" httpUri="https://zenodo.org/record/5531585/files/figure.png" pageId="10" pageNumber="461">Figs 16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="59F517F9FFBFFFFAFF686EB3FD01FF68" lastPageId="11" lastPageNumber="462" pageId="10" pageNumber="461" type="diagnosis">
<paragraph id="11504472FFBFFFFBFF686EB3FC81FA05" blockId="10.[151,1437,1257,2040]" pageId="10" pageNumber="461">
<emphasis id="239B9860FFBFFFFBFF686EB3FE99FABE" bold="true" box="[151,274,1257,1283]" pageId="10" pageNumber="461">Diagnosis.</emphasis>
With
<taxonomicName id="D6EF3FF1FFBFFFFBFEA96EB0FD82FABE" authority="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete, 2021" authorityName="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete" authorityYear="2021" box="[342,521,1257,1284]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="risaraldensis" status="sp. nov.">
<emphasis id="239B9860FFBFFFFBFEA96EB0FD82FABE" box="[342,521,1257,1284]" italics="true" pageId="10" pageNumber="461">G. risaraldensis</emphasis>
</taxonomicName>
<emphasis id="239B9860FFBFFFFBFDEF6EB3FDC4FABE" bold="true" box="[528,591,1257,1283]" pageId="10" pageNumber="461">
<taxonomicNameLabel id="38A8251BFFBFFFFBFDEF6EB3FDC4FABE" box="[528,591,1257,1283]" pageId="10" pageNumber="461" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="D6EF3FF1FFBFFFFBFD736EB0FCC8FABE" authority="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete, 2021" authorityName="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete" authorityYear="2021" box="[652,835,1257,1284]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="vallecaucana" status="sp. nov.">
<emphasis id="239B9860FFBFFFFBFD736EB0FCC8FABE" box="[652,835,1257,1284]" italics="true" pageId="10" pageNumber="461">G. vallecaucana</emphasis>
</taxonomicName>
<emphasis id="239B9860FFBFFFFBFCB56EB3FC03FABE" bold="true" box="[842,904,1257,1283]" pageId="10" pageNumber="461">
<taxonomicNameLabel id="38A8251BFFBFFFFBFCB56EB3FC03FABE" box="[842,904,1257,1283]" pageId="10" pageNumber="461" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, it constitutes an assemblage of related species, characterized by having the external parameres well developed and by having endophallic sclerites unlike those observed in the known species of
<taxonomicName id="D6EF3FF1FFBFFFFBFE076F68FDBBFAF1" authorityName=", Carrejo, Gonzalez Obando, Garcia Aldrete &amp; Mendivil" authorityYear="2021" box="[504,560,1330,1356]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="461" phylum="Arthropoda" rank="genus">
<emphasis id="239B9860FFBFFFFBFE076F68FDBBFAF1" box="[504,560,1330,1356]" italics="true" pageId="10" pageNumber="461">Goja</emphasis>
</taxonomicName>
. It differs from the two species above by having R4+5 of the forewings seven branched; by having the external parameres mid sized, by having the apex of the aedeagal arch with small, pointed projections directed outward, with the posterior border straight, and by having the right endophallic sclerites with two mesal processes, one shorter than the other (
<figureCitation id="89D458F7FFBFFFFBFD486FC7FD77FA05" box="[695,764,1437,1464]" captionStart="FIGURES 16" captionStartId="11.[151,264,1851,1875]" captionTargetBox="[163,1427,259,1822]" captionTargetId="figure-42@11.[151,1436,234,1825]" captionTargetPageId="11" captionText="FIGURES 16. Goja andina n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531585" httpUri="https://zenodo.org/record/5531585/files/figure.png" pageId="10" pageNumber="461">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="11504472FFBFFFFBFF386F9BFCA1FA61" blockId="10.[151,1437,1257,2040]" box="[199,810,1473,1500]" pageId="10" pageNumber="461">
<emphasis id="239B9860FFBFFFFBFF386F9BFE80FA66" bold="true" box="[199,267,1473,1499]" pageId="10" pageNumber="461">Color</emphasis>
(in 80% ethanol). Body brown, with light spots.
</paragraph>
<paragraph id="11504472FFBFFFFAFF386FBFFD01FF68" blockId="10.[151,1437,1257,2040]" lastBlockId="11.[151,1436,151,213]" lastPageId="11" lastPageNumber="462" pageId="10" pageNumber="461">
<emphasis id="239B9860FFBFFFFBFF386FBFFEEAFA42" bold="true" box="[199,353,1509,1535]" pageId="10" pageNumber="461">Morphology.</emphasis>
Head: H/MxW: 1.64; compound eyes small, H/d: 3.61; IO/MxW: 0.66. Outer cusp of lacinial tips broad, with seven denticles. Mx4/Mx2: 1.21. Lower edge of labrum with a pair of projections with rounded apices. Forewings (
<figureCitation id="89D458F7FFBFFFFBFE8F6C77FE39F9FA" box="[368,434,1581,1607]" captionStart="FIGURES 16" captionStartId="11.[151,264,1851,1875]" captionTargetBox="[163,1427,259,1822]" captionTargetId="figure-42@11.[151,1436,234,1825]" captionTargetPageId="11" captionText="FIGURES 16. Goja andina n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531585" httpUri="https://zenodo.org/record/5531585/files/figure.png" pageId="10" pageNumber="461">Fig. 1</figureCitation>
): FW/W: 2.88; pterostigma elongate, wider in the middle: lp/wp: 6.19, Rs supernumerary with six or more branches, basal portion of Rs concave after transverse r-m; areola postica elongate, may be open and partially fused to M: al/ah: 4.00; M supernumerary. Hindwings (
<figureCitation id="89D458F7FFBFFFFBFC6D6C2FFC5CF932" box="[914,983,1653,1679]" captionStart="FIGURES 16" captionStartId="11.[151,264,1851,1875]" captionTargetBox="[163,1427,259,1822]" captionTargetId="figure-42@11.[151,1436,234,1825]" captionTargetPageId="11" captionText="FIGURES 16. Goja andina n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531585" httpUri="https://zenodo.org/record/5531585/files/figure.png" pageId="10" pageNumber="461">Fig. 2</figureCitation>
): HW/w: 3.31. Rs five branched; M four branched; Rs and M joined by a crossvein, asymmetry in wing venation may be present. Legs: fore tarsi without ctenidiobothria, mid and hind tarsi with ctenidiobothria. Hypandrium broad, setae as illustrated (
<figureCitation id="89D458F7FFBFFFFBFB1C6CE7FAACF965" box="[1251,1319,1725,1752]" captionStart="FIGURES 16" captionStartId="11.[151,264,1851,1875]" captionTargetBox="[163,1427,259,1822]" captionTargetId="figure-42@11.[151,1436,234,1825]" captionTargetPageId="11" captionText="FIGURES 16. Goja andina n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531585" httpUri="https://zenodo.org/record/5531585/files/figure.png" pageId="10" pageNumber="461">Fig. 5</figureCitation>
) posterior border almost straight. Phallosome (
<figureCitation id="89D458F7FFBFFFFBFDCA6CBBFDF7F941" box="[565,636,1761,1788]" captionStart="FIGURES 16" captionStartId="11.[151,264,1851,1875]" captionTargetBox="[163,1427,259,1822]" captionTargetId="figure-42@11.[151,1436,234,1825]" captionTargetPageId="11" captionText="FIGURES 16. Goja andina n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531585" httpUri="https://zenodo.org/record/5531585/files/figure.png" pageId="10" pageNumber="461">Fig. 6</figureCitation>
): side struts straight; aedeagal arch robust, apex with two short mesal projections, acuminate, directed outward; external parameres well developed, concave-convex, asymmetric, right one sclerotized, acuminate distally, left one shorter. Three pairs of asymmetric endophallic sclerites (
<figureCitation id="89D458F7FFBFFFFBFAF46D73FADAF8F9" box="[1291,1361,1833,1860]" captionStart="FIGURES 16" captionStartId="11.[151,264,1851,1875]" captionTargetBox="[163,1427,259,1822]" captionTargetId="figure-42@11.[151,1436,234,1825]" captionTargetPageId="11" captionText="FIGURES 16. Goja andina n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531585" httpUri="https://zenodo.org/record/5531585/files/figure.png" pageId="10" pageNumber="461">Fig. 6</figureCitation>
): right pair with two mesal processes, lanceolate, apically acuminate, one shorter than the other; two lateral sclerites, outermost apically wide and projected posteriorly, anteriorly narrow; innermost narrow and distally lobed, base folded on itself as a reticulate lamina; left sclerites mesally lanceolate, apex acuminate; two outermost lateral sclerites wide apically, narrow mesally, anteriorly wide with fine spicules. Clunium projected posteriorly over the area of the epiproct. Paraprocts (
<figureCitation id="89D458F7FFBFFFFBFE7A6D87FE43F84A" box="[389,456,2013,2039]" captionStart="FIGURES 16" captionStartId="11.[151,264,1851,1875]" captionTargetBox="[163,1427,259,1822]" captionTargetId="figure-42@11.[151,1436,234,1825]" captionTargetPageId="11" captionText="FIGURES 16. Goja andina n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531585" httpUri="https://zenodo.org/record/5531585/files/figure.png" pageId="10" pageNumber="461">Fig. 4</figureCitation>
), oval, with setae and macrosetae on posterior border, sensory fields large, circular, with 32 trichobothria in basal rosettes. Epiproct slightly concave anteriorly, rounded posteriorly, with three setae mesally, posterior field of setae as illustrated (
<figureCitation id="89D458F7FFBEFFFAFDC86AE1FDF7FF68" box="[567,636,187,213]" captionStart="FIGURES 16" captionStartId="11.[151,264,1851,1875]" captionTargetBox="[163,1427,259,1822]" captionTargetId="figure-42@11.[151,1436,234,1825]" captionTargetPageId="11" captionText="FIGURES 16. Goja andina n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531585" httpUri="https://zenodo.org/record/5531585/files/figure.png" pageId="11" pageNumber="462">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="459014FAFFBEFFFAFF686D61FDDBF8CA" ID-DOI="http://doi.org/10.5281/zenodo.5531585" ID-Zenodo-Dep="5531585" httpUri="https://zenodo.org/record/5531585/files/figure.png" pageId="11" pageNumber="462" startId="11.[151,264,1851,1875]" targetBox="[163,1427,259,1822]" targetPageId="11">
<paragraph id="11504472FFBEFFFAFF686D61FDDBF8CA" blockId="11.[151,1436,1851,1912]" pageId="11" pageNumber="462">
<emphasis id="239B9860FFBEFFFAFF686D61FEB0F8EE" bold="true" box="[151,315,1851,1875]" pageId="11" pageNumber="462">FIGURES 16.</emphasis>
<taxonomicName id="D6EF3FF1FFBEFFFAFEBD6D66FE49F8E9" authority="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete, 2021" authorityName="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete" authorityYear="2021" box="[322,450,1852,1876]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="andina" status="sp. nov.">
<emphasis id="239B9860FFBEFFFAFEBD6D66FE49F8E9" box="[322,450,1852,1876]" italics="true" pageId="11" pageNumber="462">Goja andina</emphasis>
</taxonomicName>
<emphasis id="239B9860FFBEFFFAFE366D66FD88F8E9" bold="true" box="[457,515,1852,1876]" pageId="11" pageNumber="462">
<taxonomicNameLabel id="38A8251BFFBEFFFAFE366D66FD88F8E9" box="[457,515,1852,1876]" pageId="11" pageNumber="462" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Male.
<emphasis id="239B9860FFBEFFFAFDB36D61FDD2F8EE" bold="true" box="[588,601,1851,1875]" pageId="11" pageNumber="462">1</emphasis>
. Forewing.
<emphasis id="239B9860FFBEFFFAFD2B6D61FD6AF8EE" bold="true" box="[724,737,1851,1875]" pageId="11" pageNumber="462">2</emphasis>
. Hindwing.
<emphasis id="239B9860FFBEFFFAFC9E6D61FCE5F8EE" bold="true" box="[865,878,1851,1875]" pageId="11" pageNumber="462">3</emphasis>
. Front view of head.
<emphasis id="239B9860FFBEFFFAFBB46D61FBD3F8EE" bold="true" box="[1099,1112,1851,1875]" pageId="11" pageNumber="462">4</emphasis>
. Left paraproct and epiproct.
<emphasis id="239B9860FFBEFFFAFA766D66FA1DF8E9" bold="true" box="[1417,1430,1852,1876]" pageId="11" pageNumber="462">5</emphasis>
. Hypandrium.
<emphasis id="239B9860FFBEFFFAFEDB6D05FEBAF8CA" bold="true" box="[292,305,1887,1911]" pageId="11" pageNumber="462">6</emphasis>
. Phallosome. Scales in mm.
</paragraph>
</caption>
<subSubSection id="59F517F9FFB9FFFDFF386ACCFF41FE10" pageId="12" pageNumber="463" type="description">
<paragraph id="11504472FFB9FFFDFF386ACCFC98FF68" blockId="12.[151,1437,150,501]" pageId="12" pageNumber="463">
<emphasis id="239B9860FFB9FFFDFF386ACCFEF5FF0D" bold="true" box="[199,382,150,176]" pageId="12" pageNumber="463">Measurements.</emphasis>
FW: 4800, HW: 3500, F: 1100, T: 1970, t1: 950, t2: 260, ctt1: 38, f1: 1040, f2: 920, f3: 720, Mx4: 370, IO: 300, D: 255, d: 292, IO/d: 1.45, PO: 0.85.
</paragraph>
<paragraph id="11504472FFB9FFFDFF386A84FF41FE10" blockId="12.[151,1437,150,501]" pageId="12" pageNumber="463">
<emphasis id="239B9860FFB9FFFDFF386A84FE29FF45" bold="true" box="[199,418,222,248]" pageId="12" pageNumber="463">Specimens studied</emphasis>
.
<materialsCitation id="A1874E2FFFB9FFFDFE516A84FC6DFEA1" collectingDate="2011-01-15" collectionCode="R" collectorName="A. N. Garcia Aldrete &amp; Gonzalez" country="Colombia" elevation="2592" latitude="5.0734167" location="Rio Blanco Nature Reserve" longLatPrecision="1" longitude="-75.437836" municipality="Manizales" pageId="12" pageNumber="463" specimenCount="1" stateProvince="Caldas" typeStatus="holotype">
<typeStatus id="CE54FAD0FFB9FFFDFE516A84FD9DFF45" box="[430,534,222,248]" pageId="12" pageNumber="463" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="69F804E2FFB9FFFDFDA06A84FD73FF45" box="[607,760,222,248]" name="Colombia" pageId="12" pageNumber="463">COLOMBIA</collectingCountry>
.
<collectingRegion id="D32B8A90FFB9FFFDFCFE6A84FCDBFF45" box="[769,848,222,248]" country="Colombia" name="Caldas" pageId="12" pageNumber="463">Caldas</collectingRegion>
,
<collectingMunicipality id="F134DE08FFB9FFFDFCA56A84FC5BFF45" box="[858,976,222,248]" pageId="12" pageNumber="463">Manizales</collectingMunicipality>
,
<location id="143012A9FFB9FFFDFC266A84FA80FF44" LSID="urn:lsid:plazi:treatment:9946F564FFBFFFFDFF686E27FDEEFE49:143012A9FFB9FFFDFC266A84FA80FF44" box="[985,1291,222,249]" country="Colombia" latitude="5.0734167" longLatPrecision="1" longitude="-75.437836" municipality="Manizales" name="Rio Blanco Nature Reserve" pageId="12" pageNumber="463" stateProvince="Caldas">Río Blanco Nature Reserve</location>
,
<geoCoordinate id="74DB22B5FFB9FFFDFAEA6A84FA13FF44" box="[1301,1432,222,249]" degrees="5" direction="north" minutes="4" orientation="latitude" pageId="12" pageNumber="463" precision="1" seconds="24.3" value="5.0734167">5º424.3”N</geoCoordinate>
:
<geoCoordinate id="74DB22B5FFB9FFFDFF686B58FEB2FEA0" box="[151,313,258,285]" degrees="75" direction="west" minutes="26" orientation="longitude" pageId="12" pageNumber="463" precision="1" seconds="16.2" value="-75.437836">75º2616.2”W</geoCoordinate>
,
<quantity id="D617E997FFB9FFFDFEB86B58FE28FEA0" box="[327,419,258,285]" metricMagnitude="3" metricUnit="m" metricValue="2.5919999999999996" pageId="12" pageNumber="463" unit="m" value="2592.0">
<elevation id="9AC2A341FFB9FFFDFEB86B58FE28FEA0" box="[327,419,258,285]" metricMagnitude="3" metricUnit="m" metricValue="2.5919999999999996" pageId="12" pageNumber="463" unit="m" value="2592.0">2592 m</elevation>
</quantity>
.,
<date id="655162B2FFB9FFFDFE4B6B58FDAAFEA0" box="[436,545,258,285]" pageId="12" pageNumber="463" value="2011-01-15">
<collectingDate id="75159B5AFFB9FFFDFE4B6B58FDAAFEA0" box="[436,545,258,285]" pageId="12" pageNumber="463" value="2011-01-15">15.I.2011</collectingDate>
</date>
,
<collectorName id="BC1A21A4FFB9FFFDFDD26B58FC97FEA1" box="[557,796,258,285]" pageId="12" pageNumber="463">A. N. García Aldrete</collectorName>
and
<collectionCode id="77FEDCB7FFB9FFFDFCA96B59FCE7FEA0" box="[854,876,259,285]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="463">R</collectionCode>
.
<collectorName id="BC1A21A4FFB9FFFDFC866B58FC6DFEA1" box="[889,998,258,284]" pageId="12" pageNumber="463">González</collectorName>
</materialsCitation>
.
<materialsCitation id="A1874E2FFFB9FFFDFC0C6B59FBA0FEFD" collectingDate="2011-01-16" collectionCode="R" collectorName="Beating" country="Colombia" elevation="2339" latitude="5.065778" location="Rio Blanco Nature Reserve" longLatPrecision="1" longitude="-75.447914" municipality="Manizales" pageId="12" pageNumber="463" specimenCount="1" specimenCount-male="1" stateProvince="Caldas" typeStatus="paratype">
<typeStatus id="CE54FAD0FFB9FFFDFC0C6B59FBEFFEA0" box="[1011,1124,259,285]" pageId="12" pageNumber="463" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="07E98FFBFFB9FFFDFB8F6B58FB4BFEA1" box="[1136,1216,258,284]" pageId="12" pageNumber="463" type="male">1 male</specimenCount>
, same location and collectors,
<geoCoordinate id="74DB22B5FFB9FFFDFEEE6B7CFE1FFEFC" box="[273,404,294,321]" degrees="5" direction="north" minutes="3" orientation="latitude" pageId="12" pageNumber="463" precision="1" seconds="56.8" value="5.065778">5º356.8”N</geoCoordinate>
:
<geoCoordinate id="74DB22B5FFB9FFFDFE606B7CFDCAFEFC" box="[415,577,294,321]" degrees="75" direction="west" minutes="26" orientation="longitude" pageId="12" pageNumber="463" precision="1" seconds="52.5" value="-75.447914">75º2652.5”W</geoCoordinate>
,
<quantity id="D617E997FFB9FFFDFDB46B7CFD2FFEFC" box="[587,676,294,321]" metricMagnitude="3" metricUnit="m" metricValue="2.339" pageId="12" pageNumber="463" unit="m" value="2339.0">
<elevation id="9AC2A341FFB9FFFDFDB46B7CFD2FFEFC" box="[587,676,294,321]" metricMagnitude="3" metricUnit="m" metricValue="2.339" pageId="12" pageNumber="463" unit="m" value="2339.0">2339 m</elevation>
</quantity>
.,
<date id="655162B2FFB9FFFDFD4C6B7CFC94FEFD" box="[691,799,294,320]" pageId="12" pageNumber="463" value="2011-01-16">
<collectingDate id="75159B5AFFB9FFFDFD4C6B7CFC94FEFD" box="[691,799,294,320]" pageId="12" pageNumber="463" value="2011-01-16">16.I.2011</collectingDate>
</date>
,
<collectorName id="BC1A21A4FFB9FFFDFCD46B7CFC09FEFD" box="[811,898,294,320]" pageId="12" pageNumber="463">Beating</collectorName>
dry vegetation
</materialsCitation>
.
<materialsCitation id="A1874E2FFFB9FFFDFBCA6B7CFBF5FED9" collectingDate="2013-11-03" collectorName="N. Hassy" country="Colombia" elevation="2170" latitude="4.17" location="Kilometro" longLatPrecision="1" longitude="-75.85797" municipality="Sevilla" pageId="12" pageNumber="463" specimenCount="1" specimenCount-male="1" stateProvince="Valle del Cauca" typeStatus="paratype">
<specimenCount id="07E98FFBFFB9FFFDFBCA6B7CFB09FEFD" box="[1077,1154,294,320]" pageId="12" pageNumber="463" type="male">1 male</specimenCount>
,
<collectingRegion id="D32B8A90FFB9FFFDFB736B7CFAB6FEFD" box="[1164,1341,294,320]" country="Colombia" name="Valle del Cauca" pageId="12" pageNumber="463">Valle del Cauca</collectingRegion>
,
<collectingMunicipality id="F134DE08FFB9FFFDFAB86B7CFA13FEFD" box="[1351,1432,294,320]" pageId="12" pageNumber="463">Sevilla</collectingMunicipality>
,
<location id="143012A9FFB9FFFDFF686B10FE81FED9" LSID="urn:lsid:plazi:treatment:9946F564FFBFFFFDFF686E27FDEEFE49:143012A9FFB9FFFDFF686B10FE81FED9" box="[151,266,330,356]" country="Colombia" latitude="4.17" longLatPrecision="1" longitude="-75.85797" municipality="Sevilla" name="Kilometro" pageId="12" pageNumber="463" stateProvince="Valle del Cauca">Kilómetro</location>
14,
<geoCoordinate id="74DB22B5FFB9FFFDFEC26B10FE44FED9" box="[317,463,330,356]" degrees="4" direction="north" minutes="10" orientation="latitude" pageId="12" pageNumber="463" precision="1" seconds="12.0" value="4.17">4º1012.0”N</geoCoordinate>
:
<geoCoordinate id="74DB22B5FFB9FFFDFE256B10FDF7FED8" box="[474,636,330,357]" degrees="75" direction="west" minutes="51" orientation="longitude" pageId="12" pageNumber="463" precision="1" seconds="28.7" value="-75.85797">75º5128.7”W</geoCoordinate>
,
<quantity id="D617E997FFB9FFFDFD776B10FD68FED8" box="[648,739,330,357]" metricMagnitude="3" metricUnit="m" metricValue="2.17" pageId="12" pageNumber="463" unit="m" value="2170.0">
<elevation id="9AC2A341FFB9FFFDFD776B10FD68FED8" box="[648,739,330,357]" metricMagnitude="3" metricUnit="m" metricValue="2.17" pageId="12" pageNumber="463" unit="m" value="2170.0">2170 m</elevation>
</quantity>
.,
<date id="655162B2FFB9FFFDFD0B6B10FCE2FED9" box="[756,873,330,356]" pageId="12" pageNumber="463" value="2013-11-03">
<collectingDate id="75159B5AFFB9FFFDFD0B6B10FCE2FED9" box="[756,873,330,356]" pageId="12" pageNumber="463" value="2013-11-03">3.XI.2013</collectingDate>
</date>
.
<collectorName id="BC1A21A4FFB9FFFDFC8B6B11FC57FED8" box="[884,988,331,357]" pageId="12" pageNumber="463">N. Hassy</collectorName>
, on tree trunk
</materialsCitation>
.
<materialsCitation id="A1874E2FFFB9FFFDFB766B10FF4DFE10" collectingDate="2004-03-20" collectingDateMax="2004-04-30" collectingDateMin="2004-03-20" country="Colombia" county="Sanctuary of Flora" elevation="2050" latitude="4.7166667" location="Cuchilla" longLatPrecision="1306" longitude="-75.583336" municipality="Fauna Otun Quimbaya" pageId="12" pageNumber="463" specimenCount="1" specimenCount-male="1" stateProvince="Risaralda" typeStatus="paratype">
<specimenCount id="07E98FFBFFB9FFFDFB766B10FB53FED9" box="[1161,1240,330,356]" pageId="12" pageNumber="463" type="male">1 male</specimenCount>
,
<collectingRegion id="D32B8A90FFB9FFFDFB1B6B10FADAFED9" box="[1252,1361,330,356]" country="Colombia" name="Risaralda" pageId="12" pageNumber="463">Risaralda</collectingRegion>
,
<collectingCounty id="F8313CFEFFB9FFFDFAA36B10FEBEFE35" pageId="12" pageNumber="463">Sanctuary of Flora</collectingCounty>
and
<collectingMunicipality id="F134DE08FFB9FFFDFE906B35FDFEFE35" box="[367,629,366,393]" pageId="12" pageNumber="463">Fauna Otún Quimbaya</collectingMunicipality>
,
<location id="143012A9FFB9FFFDFD7E6B34FD6BFE35" LSID="urn:lsid:plazi:treatment:9946F564FFBFFFFDFF686E27FDEEFE49:143012A9FFB9FFFDFD7E6B34FD6BFE35" box="[641,736,366,392]" country="Colombia" county="Sanctuary of Flora" latitude="4.7166667" longLatPrecision="1306" longitude="-75.583336" municipality="Fauna Otun Quimbaya" name="Cuchilla" pageId="12" pageNumber="463" stateProvince="Risaralda">Cuchilla</location>
road,
<geoCoordinate id="74DB22B5FFB9FFFDFCD46B34FC0AFE35" box="[811,897,366,392]" degrees="4" direction="north" minutes="43" orientation="latitude" pageId="12" pageNumber="463" precision="925" value="4.7166667">4°43N</geoCoordinate>
:
<geoCoordinate id="74DB22B5FFB9FFFDFC716B34FC7EFE34" box="[910,1013,366,393]" degrees="75" direction="west" minutes="35" orientation="longitude" pageId="12" pageNumber="463" precision="925" value="-75.583336">75°35W</geoCoordinate>
,
<quantity id="D617E997FFB9FFFDFBFD6B34FBD5FE34" box="[1026,1118,366,393]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="12" pageNumber="463" unit="m" value="2050.0">
<elevation id="9AC2A341FFB9FFFDFBFD6B34FBD5FE34" box="[1026,1118,366,393]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="12" pageNumber="463" unit="m" value="2050.0">2050 m</elevation>
</quantity>
.,
<date id="655162B2FFB9FFFDFB8F6B34FABDFE35" box="[1136,1334,366,392]" pageId="12" pageNumber="463" value="2004-03-20" valueMax="2004-04-30" valueMin="2004-03-20">
<collectingDate id="75159B5AFFB9FFFDFB8F6B34FABDFE35" box="[1136,1334,366,392]" pageId="12" pageNumber="463" value="2004-03-20" valueMax="2004-04-30" valueMin="2004-03-20">20.III-30.IV.2004</collectingDate>
</date>
,
<collectingMethod id="C8AE3C65FFB9FFFDFABB6B34FF4DFE10" pageId="12" pageNumber="463">Malaise trap</collectingMethod>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="59F517F9FFB9FFFDFF386BECFDEEFE49" pageId="12" pageNumber="463" type="etymology">
<paragraph id="11504472FFB9FFFDFF386BECFDEEFE49" blockId="12.[151,1437,150,501]" pageId="12" pageNumber="463">
<emphasis id="239B9860FFB9FFFDFF386BECFEC5FE6D" bold="true" box="[199,334,438,464]" pageId="12" pageNumber="463">Etymology.</emphasis>
The specific epithet refers to the Colombian Andes mountain range, where, together with the
<typeStatus id="CE54FAD0FFB9FFFDFA9E6BECFF41FE48" pageId="12" pageNumber="463" type="holotype">holotype</typeStatus>
, the other specimens were collected.
</paragraph>
</subSubSection>
</treatment>
</document>