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<document id="4E5180594A4FA15DDA93DBC72499E7E5" ID-DOI="10.11646/zootaxa.5040.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5531583" ID-ZooBank="FFFA7323-04AC-4B70-9DBE-6EAD909A9D2A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632749724544" checkinUser="plazi" docAuthor="Carrejo, Nancy, Obando, Ranulfo González, Casasola-González, José Arturo &amp; García Aldrete, Alfonso N." docDate="2021" docId="9946F564FFB9FFFDFF68681CFB04F8F1" docLanguage="en" docName="zootaxa.5040.4.1.pdf" docOrigin="Zootaxa 5040 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Goja garridoi Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete 2021, n. sp." docType="treatment" docVersion="3" lastPageNumber="463" masterDocId="657F8D1CFFB5FFF1FFFF6A5AFF8BFFBD" masterDocTitle="New Colombian Goja Navás (Psocodea: Psocoptera: Epipsocidae) with peculiar genitalia, and the first Goja with brachypterous male, from Oaxaca, Mexico" masterLastPageNumber="481" masterPageNumber="451" pageNumber="463" updateTime="1699415585501" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="9BAAB28AE27E00D5A671A6BEF89B3868">New Colombian Goja Navás (Psocodea: Psocoptera: Epipsocidae) with peculiar genitalia, and the first Goja with brachypterous male, from Oaxaca, Mexico</mods:title>
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<mods:namePart id="E1E28E4DE2383E35A6E903B24FC938D6">Carrejo, Nancy</mods:namePart>
<mods:affiliation id="39E91525CEF52577C38C2C83417C0454">Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, COLOMBIA. Instituto de Estudios Ambientales, Universidad de la Sierra Juárez, Ixtlán de Juárez, Oaxaca, MÉXICO. &amp; nancy. carrejo @ correounivalle. edu. co; htps: // orcid. org / 0000 - 0002 - 2395 - 6887</mods:affiliation>
<mods:nameIdentifier id="6E7B7BC76A4FF751F9FA9D06272AF7AC" type="email">nancy.carrejo@correounivalle.edu.co</mods:nameIdentifier>
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<mods:namePart id="EFF7B973F4F1D69E81B322905D4F15B5">Obando, Ranulfo González</mods:namePart>
<mods:affiliation id="33107A7ECC1BC3361176AE400B7A0C8D">Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, COLOMBIA. Instituto de Estudios Ambientales, Universidad de la Sierra Juárez, Ixtlán de Juárez, Oaxaca, MÉXICO. &amp; ranulfo. gonzález @ correounivalle. edu. co; htps: // orcid. org / 0000 - 0003 - 4047 - 8677</mods:affiliation>
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<mods:namePart id="E58F97BCA018C59A8EEDF4A4E94A90F5">Casasola-González, José Arturo</mods:namePart>
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<mods:affiliation id="24A294CD454E2C5628004C1A56652F64">casasola @ unsij. edu. mx; https: // orcid. org / 0000 - 0002 - 4765 - 6707</mods:affiliation>
<mods:nameIdentifier id="85BC25463051FF391D7DFF22A141BB05" type="email">casasola@unsij.edu.mx</mods:nameIdentifier>
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<mods:namePart id="51C21B597237EAC41ADF2C7355CD5AA9">García Aldrete, Alfonso N.</mods:namePart>
<mods:affiliation id="4D0BBD685278B19196D1DBEC5E6939BA">Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, CdMx, MÉXICO. &amp; anga @ ib. unam. mx; htps: // orcid. org / 0000 - 0001 - 7214 - 7966</mods:affiliation>
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<mods:title id="43E3839861E41BDE9C8FB16654C3D520">Zootaxa</mods:title>
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<mods:date id="884651C53928CCD64C7D002EA50D1D0B">2021</mods:date>
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<subSubSection id="59F517F9FFB9FFFDFF68681CFE47FDDD" box="[151,460,582,609]" pageId="12" pageNumber="463" type="nomenclature">
<paragraph id="11504472FFB9FFFDFF68681CFE47FDDD" blockId="12.[151,460,582,644]" box="[151,460,582,609]" pageId="12" pageNumber="463">
<heading id="4A18F31EFFB9FFFDFF68681CFE47FDDD" bold="true" box="[151,460,582,609]" fontSize="11" level="1" pageId="12" pageNumber="463" reason="1">
<emphasis id="239B9860FFB9FFFDFF68681CFE47FDDD" bold="true" box="[151,460,582,609]" pageId="12" pageNumber="463">
<taxonomicName id="D6EF3FF1FFB9FFFDFF68681CFEB0FDDC" ID-CoL="9J426" authority="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete, 2021" authorityName="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete" authorityYear="2021" box="[151,315,582,609]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="garridoi" status="sp. nov.">
<emphasis id="239B9860FFB9FFFDFF68681CFEB0FDDC" bold="true" box="[151,315,582,609]" italics="true" pageId="12" pageNumber="463">Goja garridoi</emphasis>
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Male
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</paragraph>
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<subSubSection id="59F517F9FFB9FFFDFF686830FE92FD39" box="[151,281,618,644]" pageId="12" pageNumber="463" type="description">
<paragraph id="11504472FFB9FFFDFF686830FE92FD39" blockId="12.[151,460,582,644]" box="[151,281,618,644]" pageId="12" pageNumber="463">
(
<figureCitation id="89D458F7FFB9FFFDFF606830FE9AFD39" box="[159,273,618,644]" captionStart="FIGURES 712" captionStartId="13.[151,264,1918,1942]" captionTargetBox="[157,1432,187,1893]" captionTargetId="figure-19@13.[151,1435,181,1894]" captionTargetPageId="13" captionText="FIGURES 712. Goja garridoi n. sp. Male. 7. Forewing. 8. Hindwing. 9. Front view of head. 10. Left paraproct and epiproct. 11. Hypandrium. 12. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531587" httpUri="https://zenodo.org/record/5531587/files/figure.png" pageId="12" pageNumber="463">Figs 712</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="59F517F9FFB9FFFDFF6868E8FDAAF9C9" pageId="12" pageNumber="463" type="diagnosis">
<paragraph id="11504472FFB9FFFDFF6868E8FD95FC19" blockId="12.[151,1437,690,1869]" pageId="12" pageNumber="463">
<emphasis id="239B9860FFB9FFFDFF6868E8FE99FD71" bold="true" box="[151,274,690,716]" pageId="12" pageNumber="463">Diagnosis.</emphasis>
With
<taxonomicName id="D6EF3FF1FFB9FFFDFEA868E9FE7CFD71" authority="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete, 2021" authorityName="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete" authorityYear="2021" box="[343,503,690,717]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="meremberg" status="sp. nov.">
<emphasis id="239B9860FFB9FFFDFEA868E9FE7CFD71" box="[343,503,690,717]" italics="true" pageId="12" pageNumber="463">G. meremberg</emphasis>
</taxonomicName>
<emphasis id="239B9860FFB9FFFDFE0068E8FDB4FD71" bold="true" box="[511,575,690,716]" pageId="12" pageNumber="463">
<taxonomicNameLabel id="38A8251BFFB9FFFDFE0068E8FDB4FD71" box="[511,575,690,716]" pageId="12" pageNumber="463" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="D6EF3FF1FFB9FFFDFDB268E9FD4DFD71" authority="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete, 2021" authorityName="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete" authorityYear="2021" box="[589,710,690,717]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="tenerife" status="sp. nov.">
<emphasis id="239B9860FFB9FFFDFDB268E9FD4DFD71" box="[589,710,690,717]" italics="true" pageId="12" pageNumber="463">G. tenerife</emphasis>
</taxonomicName>
<emphasis id="239B9860FFB9FFFDFD3268E8FC86FD71" bold="true" box="[717,781,690,716]" pageId="12" pageNumber="463">
<taxonomicNameLabel id="38A8251BFFB9FFFDFD3268E8FC86FD71" box="[717,781,690,716]" pageId="12" pageNumber="463" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="D6EF3FF1FFB9FFFDFCB368E9FC43FD71" authority="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete, 2021" authorityName="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete" authorityYear="2021" box="[844,968,690,717]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="vavilovi" status="sp. nov.">
<emphasis id="239B9860FFB9FFFDFCB368E9FC43FD71" box="[844,968,690,717]" italics="true" pageId="12" pageNumber="463">G. vavilovi</emphasis>
</taxonomicName>
<emphasis id="239B9860FFB9FFFDFC3068E8FB85FD71" bold="true" box="[975,1038,690,716]" pageId="12" pageNumber="463">
<taxonomicNameLabel id="38A8251BFFB9FFFDFC3068E8FB85FD71" box="[975,1038,690,716]" pageId="12" pageNumber="463" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, it constitutes an assemblage of related species, characterized by having the hypandrium with a median posterior process, by having a large process on each side of the posterior end of the aedeagal arch, and by having endophallic sclerites unlike those in the known species of
<taxonomicName id="D6EF3FF1FFB9FFFDFEF06945FECCFC84" authorityName=", Carrejo, Gonzalez Obando, Garcia Aldrete &amp; Mendivil" authorityYear="2021" box="[271,327,799,825]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="12" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="239B9860FFB9FFFDFEF06945FECCFC84" box="[271,327,799,825]" italics="true" pageId="12" pageNumber="463">Goja</emphasis>
</taxonomicName>
. It differs from the other species in the assemblage, by having the hypandrium with a wide, short, rectangular process, by having the posterior processes of the phallosome club-shaped, the right one with a curved, rectangular flap, and by having the right (lateral) endophallic sclerites wider than in
<taxonomicName id="D6EF3FF1FFB9FFFDFBB4693DFB61FC3D" authorityName="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete" authorityYear="2021" box="[1099,1258,870,897]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="meremberg">
<emphasis id="239B9860FFB9FFFDFBB4693DFB61FC3D" box="[1099,1258,870,897]" italics="true" pageId="12" pageNumber="463">G. meremberg</emphasis>
</taxonomicName>
and more elongated than in
<taxonomicName id="D6EF3FF1FFB9FFFDFED069D1FE2CFC19" authorityName="Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete" authorityYear="2021" box="[303,423,906,933]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="tenerife">
<emphasis id="239B9860FFB9FFFDFED069D1FE2CFC19" box="[303,423,906,933]" italics="true" pageId="12" pageNumber="463">G. tenerife</emphasis>
</taxonomicName>
. (
<figureCitation id="89D458F7FFB9FFFDFE4269D0FD9BFC19" box="[445,528,906,932]" captionStart="FIGURES 712" captionStartId="13.[151,264,1918,1942]" captionTargetBox="[157,1432,187,1893]" captionTargetId="figure-19@13.[151,1435,181,1894]" captionTargetPageId="13" captionText="FIGURES 712. Goja garridoi n. sp. Male. 7. Forewing. 8. Hindwing. 9. Front view of head. 10. Left paraproct and epiproct. 11. Hypandrium. 12. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531587" httpUri="https://zenodo.org/record/5531587/files/figure.png" pageId="12" pageNumber="463">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="11504472FFB9FFFDFF3869F4FBF2FB55" blockId="12.[151,1437,690,1869]" pageId="12" pageNumber="463">
<emphasis id="239B9860FFB9FFFDFF3869F4FE80FC75" bold="true" box="[199,267,942,968]" pageId="12" pageNumber="463">Color</emphasis>
(in 80% ethanol). Body cream. Head pattern (
<figureCitation id="89D458F7FFB9FFFDFCF869F4FCC1FC75" box="[775,842,942,968]" captionStart="FIGURES 712" captionStartId="13.[151,264,1918,1942]" captionTargetBox="[157,1432,187,1893]" captionTargetId="figure-19@13.[151,1435,181,1894]" captionTargetPageId="13" captionText="FIGURES 712. Goja garridoi n. sp. Male. 7. Forewing. 8. Hindwing. 9. Front view of head. 10. Left paraproct and epiproct. 11. Hypandrium. 12. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531587" httpUri="https://zenodo.org/record/5531587/files/figure.png" pageId="12" pageNumber="463">Fig. 9</figureCitation>
), vertex with broad dark brown to ochre area between the compound eyes, covering triangular area, slightly concave posteriorly; forehead, clypeus and labrum brown, postclypeus with convergent oblique dark brown striae; labral sclerites and upper part of labrum dark brown; genae and postgenae yellowish, genae with small dark brown spots in lower area. Compound eyes dark brown; ocellar triangle black. Antennae pale brown. Maxillary palps pale brown, Mx 4 brown. Thorax brown, tergal lobes of meso- and metathorax dark brown, metapleura brown. Legs pale brown. Forewings (
<figureCitation id="89D458F7FFB9FFFDFBF26E38FBD9FBC0" box="[1037,1106,1122,1149]" captionStart="FIGURES 712" captionStartId="13.[151,264,1918,1942]" captionTargetBox="[157,1432,187,1893]" captionTargetId="figure-19@13.[151,1435,181,1894]" captionTargetPageId="13" captionText="FIGURES 712. Goja garridoi n. sp. Male. 7. Forewing. 8. Hindwing. 9. Front view of head. 10. Left paraproct and epiproct. 11. Hypandrium. 12. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531587" httpUri="https://zenodo.org/record/5531587/files/figure.png" pageId="12" pageNumber="463">Fig. 7</figureCitation>
) almost hyaline; pterostigma with median and distal pale brown bands; veins brown. Hindwings (
<figureCitation id="89D458F7FFB9FFFDFC716EDCFC5AFB1D" box="[910,977,1158,1184]" captionStart="FIGURES 712" captionStartId="13.[151,264,1918,1942]" captionTargetBox="[157,1432,187,1893]" captionTargetId="figure-19@13.[151,1435,181,1894]" captionTargetPageId="13" captionText="FIGURES 712. Goja garridoi n. sp. Male. 7. Forewing. 8. Hindwing. 9. Front view of head. 10. Left paraproct and epiproct. 11. Hypandrium. 12. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531587" httpUri="https://zenodo.org/record/5531587/files/figure.png" pageId="12" pageNumber="463">Fig. 8</figureCitation>
), hyaline, veins brown. Abdomen cream, with subcuticular spots brown. Clunium pale brown, posterior margin brown. Hypandrium brown. Phallosome with aedeagal arch and endophallic sclerites dark brown. Epiproct and paraprocts pale brown.
</paragraph>
<paragraph id="11504472FFB9FFFDFF386EA8FDAAF9C9" blockId="12.[151,1437,690,1869]" pageId="12" pageNumber="463">
<emphasis id="239B9860FFB9FFFDFF386EA8FEEAFAB1" bold="true" box="[199,353,1266,1292]" pageId="12" pageNumber="463">Morphology.</emphasis>
Head: H/MxW: 1.55; compound eyes large, H/D: 3.04; IO/MxW: 0.66. Outer cusp of lacinial tips broad, with nine denticles. Mx4/Mx2: 1.06. Forewings (
<figureCitation id="89D458F7FFB9FFFDFCE86F4CFCD5FA8C" box="[791,862,1302,1329]" captionStart="FIGURES 712" captionStartId="13.[151,264,1918,1942]" captionTargetBox="[157,1432,187,1893]" captionTargetId="figure-19@13.[151,1435,181,1894]" captionTargetPageId="13" captionText="FIGURES 712. Goja garridoi n. sp. Male. 7. Forewing. 8. Hindwing. 9. Front view of head. 10. Left paraproct and epiproct. 11. Hypandrium. 12. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531587" httpUri="https://zenodo.org/record/5531587/files/figure.png" pageId="12" pageNumber="463">Fig. 7</figureCitation>
): FW/W: 2.82; Rs 4-6-branched; M 5-7-branched, main branches may be forked; pterostigma elongate: lp/wp: 6.19; areola postica low, elongate: al/ah: 3.41. Hindwings (
<figureCitation id="89D458F7FFB9FFFDFF156F04FEBBFAC5" box="[234,304,1374,1400]" captionStart="FIGURES 712" captionStartId="13.[151,264,1918,1942]" captionTargetBox="[157,1432,187,1893]" captionTargetId="figure-19@13.[151,1435,181,1894]" captionTargetPageId="13" captionText="FIGURES 712. Goja garridoi n. sp. Male. 7. Forewing. 8. Hindwing. 9. Front view of head. 10. Left paraproct and epiproct. 11. Hypandrium. 12. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531587" httpUri="https://zenodo.org/record/5531587/files/figure.png" pageId="12" pageNumber="463">Fig. 8</figureCitation>
): HW/w: 3.25; Rs 2-3-branched; M 2-4-branched; Rs and M joined by a crossvein. Legs: fore- tarsi without ctenidiobothria; mid- and hind- tarsi with ctenidiobothria. Hypandrium (
<figureCitation id="89D458F7FFB9FFFDFBDB6FD8FBFFFA21" box="[1060,1140,1410,1436]" captionStart="FIGURES 712" captionStartId="13.[151,264,1918,1942]" captionTargetBox="[157,1432,187,1893]" captionTargetId="figure-19@13.[151,1435,181,1894]" captionTargetPageId="13" captionText="FIGURES 712. Goja garridoi n. sp. Male. 7. Forewing. 8. Hindwing. 9. Front view of head. 10. Left paraproct and epiproct. 11. Hypandrium. 12. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531587" httpUri="https://zenodo.org/record/5531587/files/figure.png" pageId="12" pageNumber="463">Fig. 11</figureCitation>
) broad, setose, pigmented area deeply concave. Phallosome (
<figureCitation id="89D458F7FFB9FFFDFDE86FFCFDE0FA7D" box="[535,619,1446,1472]" captionStart="FIGURES 712" captionStartId="13.[151,264,1918,1942]" captionTargetBox="[157,1432,187,1893]" captionTargetId="figure-19@13.[151,1435,181,1894]" captionTargetPageId="13" captionText="FIGURES 712. Goja garridoi n. sp. Male. 7. Forewing. 8. Hindwing. 9. Front view of head. 10. Left paraproct and epiproct. 11. Hypandrium. 12. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531587" httpUri="https://zenodo.org/record/5531587/files/figure.png" pageId="12" pageNumber="463">Fig. 12</figureCitation>
): concave anteriorly side struts straight, widening distally; apex of aedeagal arch with two large, rounded processes. Two pairs of endophallic sclerites, asymmetric, lateral sclerites broad, wider in the middle, posteriorly blunt; mesal sclerites curved, distally acuminate. Paraprocts (
<figureCitation id="89D458F7FFB9FFFDFB446FB4FA85F9B5" box="[1211,1294,1518,1544]" captionStart="FIGURES 712" captionStartId="13.[151,264,1918,1942]" captionTargetBox="[157,1432,187,1893]" captionTargetId="figure-19@13.[151,1435,181,1894]" captionTargetPageId="13" captionText="FIGURES 712. Goja garridoi n. sp. Male. 7. Forewing. 8. Hindwing. 9. Front view of head. 10. Left paraproct and epiproct. 11. Hypandrium. 12. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531587" httpUri="https://zenodo.org/record/5531587/files/figure.png" pageId="12" pageNumber="463">Fig. 10</figureCitation>
) large, oval, with setae and macrosetae in distal border; sensory fields with 32 trichobothria in basal rosettes. Epiproct (
<figureCitation id="89D458F7FFB9FFFDFABD6C48FA1FF991" box="[1346,1428,1554,1580]" captionStart="FIGURES 712" captionStartId="13.[151,264,1918,1942]" captionTargetBox="[157,1432,187,1893]" captionTargetId="figure-19@13.[151,1435,181,1894]" captionTargetPageId="13" captionText="FIGURES 712. Goja garridoi n. sp. Male. 7. Forewing. 8. Hindwing. 9. Front view of head. 10. Left paraproct and epiproct. 11. Hypandrium. 12. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531587" httpUri="https://zenodo.org/record/5531587/files/figure.png" pageId="12" pageNumber="463">Fig. 10</figureCitation>
) broad, almost straight anteriorly, rounded posteriorly, one macroseta mesally, near anterior border, distal field with setae and macrosetae as illustrated.
</paragraph>
</subSubSection>
<subSubSection id="59F517F9FFB9FFFDFF386C24FC7BF895" pageId="12" pageNumber="463" type="description">
<paragraph id="11504472FFB9FFFDFF386C24FE89F95D" blockId="12.[151,1437,690,1869]" pageId="12" pageNumber="463">
<emphasis id="239B9860FFB9FFFDFF386C24FEF5F925" bold="true" box="[199,382,1662,1688]" pageId="12" pageNumber="463">Measurements.</emphasis>
FW: 6550, HW: 4715, F: 1485, T: 2715, t1: 1225, t2: 345, ctt1: 42, f1: 1450, f 2: 1237, f3: 912, f4: 575, f5: 320, f6; 312, f7: 257, f8: 240, f9: 227, f10: 207, f11: 262, Mx4: 370, IO: 460, D: 355, d: 340, IO/d: 1.35, PO: 0.96.
</paragraph>
<paragraph id="11504472FFB9FFFDFF386CB0FC7BF895" blockId="12.[151,1437,690,1869]" pageId="12" pageNumber="463">
<emphasis id="239B9860FFB9FFFDFF386CB0FE15F8B9" bold="true" box="[199,414,1770,1796]" pageId="12" pageNumber="463">Specimen studied</emphasis>
.
<materialsCitation id="A1874E2FFFB9FFFDFE4E6CB0FC7BF895" collectingDate="2019-06-07" collectorName="D. H. Garrido-Torres. Beating" country="Colombia" county="Purace" elevation="2789" latitude="2.3073056" location="Guacharos" longLatPrecision="1" longitude="-76.251274" municipality="Cueva" pageId="12" pageNumber="463" specimenCount="1" stateProvince="Cauca" typeStatus="holotype">
<typeStatus id="CE54FAD0FFB9FFFDFE4E6CB0FD92F8B9" box="[433,537,1770,1796]" pageId="12" pageNumber="463" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="69F804E2FFB9FFFDFD8F6CB0FC82F8B9" box="[624,777,1770,1796]" name="Colombia" pageId="12" pageNumber="463">COLOMBIA</collectingCountry>
.
<collectingRegion id="D32B8A90FFB9FFFDFCE76CB0FCE9F8B9" box="[792,866,1770,1796]" country="Colombia" name="Cauca" pageId="12" pageNumber="463">Cauca</collectingRegion>
.
<collectingCounty id="F8313CFEFFB9FFFDFC8D6CB0FC4AF8B9" box="[882,961,1770,1796]" pageId="12" pageNumber="463">Puracé</collectingCounty>
,
<collectingMunicipality id="F134DE08FFB9FFFDFC2E6CB0FB92F8B9" box="[977,1049,1770,1796]" pageId="12" pageNumber="463">Cueva</collectingMunicipality>
de los
<location id="143012A9FFB9FFFDFB866CB0FB7FF8B9" LSID="urn:lsid:plazi:treatment:9946F564FFB9FFFDFF68681CFB04F8F1:143012A9FFB9FFFDFB866CB0FB7FF8B9" box="[1145,1268,1770,1796]" country="Colombia" county="Purace" latitude="2.3073056" longLatPrecision="1" longitude="-76.251274" municipality="Cueva" name="Guacharos" pageId="12" pageNumber="463" stateProvince="Cauca">Guácharos</location>
,
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:
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<quantity id="D617E997FFB9FFFDFEC66D54FE1AF894" box="[313,401,1806,1833]" metricMagnitude="3" metricUnit="m" metricValue="2.7889999999999997" pageId="12" pageNumber="463" unit="m" value="2789.0">
<elevation id="9AC2A341FFB9FFFDFEC66D54FE1AF894" box="[313,401,1806,1833]" metricMagnitude="3" metricUnit="m" metricValue="2.7889999999999997" pageId="12" pageNumber="463" unit="m" value="2789.0">2789 m</elevation>
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.
<date id="655162B2FFB9FFFDFE636D54FD9AF895" box="[412,529,1806,1832]" pageId="12" pageNumber="463" value="2019-06-07">
<collectingDate id="75159B5AFFB9FFFDFE636D54FD9AF895" box="[412,529,1806,1832]" pageId="12" pageNumber="463" value="2019-06-07">7.VI.2019</collectingDate>
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.
<collectorName id="BC1A21A4FFB9FFFDFDE46D55FCE7F895" box="[539,876,1806,1833]" pageId="12" pageNumber="463">D. H. Garrido-Torres. Beating</collectorName>
vegetation.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="59F517F9FFB9FFFDFF386D68FB04F8F1" box="[199,1167,1842,1869]" pageId="12" pageNumber="463" type="etymology">
<paragraph id="11504472FFB9FFFDFF386D68FB04F8F1" blockId="12.[151,1437,690,1869]" box="[199,1167,1842,1869]" pageId="12" pageNumber="463">
<emphasis id="239B9860FFB9FFFDFF386D68FEC5F8F1" bold="true" box="[199,334,1842,1868]" pageId="12" pageNumber="463">Etymology.</emphasis>
This species is dedicated to its collector Daniel Hernando Garrido-Torres.
</paragraph>
</subSubSection>
</treatment>
</document>