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<document id="08DE66DDB44E052B3E35EFC7AEA17678" ID-DOI="10.11646/zootaxa.5040.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5531583" ID-ZooBank="FFFA7323-04AC-4B70-9DBE-6EAD909A9D2A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632749724544" checkinUser="plazi" docAuthor="Carrejo, Nancy, Obando, Ranulfo González, Casasola-González, José Arturo &amp; García Aldrete, Alfonso N." docDate="2021" docId="9946F564FFA0FFE4FF686ACDFB17F905" docLanguage="en" docName="zootaxa.5040.4.1.pdf" docOrigin="Zootaxa 5040 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Goja vallecaucana Carrejo &amp; Obando &amp; Casasola-González &amp; García Aldrete 2021, n. sp." docType="treatment" docVersion="3" lastPageNumber="472" masterDocId="657F8D1CFFB5FFF1FFFF6A5AFF8BFFBD" masterDocTitle="New Colombian Goja Navás (Psocodea: ‘ Psocoptera’: Epipsocidae) with peculiar genitalia, and the first Goja with brachypterous male, from Oaxaca, Mexico" masterLastPageNumber="481" masterPageNumber="451" pageNumber="472" updateTime="1699415585501" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="AE4F1446479EDBF41340A7FE29A86083">New Colombian Goja Navás (Psocodea: ‘ Psocoptera’: Epipsocidae) with peculiar genitalia, and the first Goja with brachypterous male, from Oaxaca, Mexico</mods:title>
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<mods:namePart id="B21D0B40F3420AE2522F38F4C0925AEC">Carrejo, Nancy</mods:namePart>
<mods:affiliation id="D5A7080D098966C6D8B648F3DAF269C8">Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, COLOMBIA. Instituto de Estudios Ambientales, Universidad de la Sierra Juárez, Ixtlán de Juárez, Oaxaca, MÉXICO. &amp; nancy. carrejo @ correounivalle. edu. co; htps: // orcid. org / 0000 - 0002 - 2395 - 6887</mods:affiliation>
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<mods:namePart id="656F2191B67909809BA3DFC18277CEB6">Obando, Ranulfo González</mods:namePart>
<mods:affiliation id="E55C26EC7C041A7C33961BB31271417E">Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, COLOMBIA. Instituto de Estudios Ambientales, Universidad de la Sierra Juárez, Ixtlán de Juárez, Oaxaca, MÉXICO. &amp; ranulfo. gonzález @ correounivalle. edu. co; htps: // orcid. org / 0000 - 0003 - 4047 - 8677</mods:affiliation>
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<mods:namePart id="D284910C7A366D2C814B0932145A2074">Casasola-González, José Arturo</mods:namePart>
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<mods:affiliation id="A04A2E7638E24D1DDA1DBB887E41512D">casasola @ unsij. edu. mx; https: // orcid. org / 0000 - 0002 - 4765 - 6707</mods:affiliation>
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<mods:namePart id="835443660D748B6E161C42F963548741">García Aldrete, Alfonso N.</mods:namePart>
<mods:affiliation id="0F3F34698AFE01D57471DA9CB179F7E2">Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, CdMx, MÉXICO. &amp; anga @ ib. unam. mx; htps: // orcid. org / 0000 - 0001 - 7214 - 7966</mods:affiliation>
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<heading id="4A18F31EFFA0FFE4FF686ACDFD81FF0C" bold="true" box="[151,522,151,178]" fontSize="11" level="1" pageId="21" pageNumber="472" reason="1">
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<emphasis id="239B9860FFA0FFE4FF686ACDFEF2FF0F" bold="true" box="[151,377,151,178]" italics="true" pageId="21" pageNumber="472">Goja vallecaucana</emphasis>
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Male
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<paragraph id="11504472FFA0FFE4FF686AE1FEACFF6B" blockId="21.[151,522,151,214]" box="[151,295,187,214]" pageId="21" pageNumber="472">
(
<figureCitation id="89D458F7FFA0FFE4FF606AE1FEABFF6B" box="[159,288,187,214]" captionStart="FIGURES 40‒45" captionStartId="22.[151,264,1973,1997]" captionTargetBox="[163,1425,256,1933]" captionTargetId="figure-61@22.[151,1435,237,1950]" captionTargetPageId="22" captionText="FIGURES 40‒45. Goja vallecaucana n. sp. Male. 40. Forewing. 41. Hindwing. 42. Front view of head. 43. Left paraproct and epiproct. 44. Hypandrium. 45. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531603" httpUri="https://zenodo.org/record/5531603/files/figure.png" pageId="21" pageNumber="472">Figs 40–45</figureCitation>
)
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<emphasis id="239B9860FFA0FFE4FF686AA7FE99FEAA" bold="true" box="[151,274,253,279]" pageId="21" pageNumber="472">Diagnosis.</emphasis>
It differs from 
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<emphasis id="239B9860FFA0FFE4FE3D6AA4FDBFFEA5" box="[450,564,254,280]" italics="true" pageId="21" pageNumber="472">G. andina</emphasis>
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<taxonomicNameLabel id="38A8251BFFA0FFE4FDC36AA4FDF7FEA5" box="[572,636,254,280]" pageId="21" pageNumber="472" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, and 
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<emphasis id="239B9860FFA0FFE4FD436AA4FCF9FEA5" box="[700,882,254,280]" italics="true" pageId="21" pageNumber="472">G. risaraldensis</emphasis>
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<taxonomicNameLabel id="38A8251BFFA0FFE4FC856AA4FC31FEA5" box="[890,954,254,280]" pageId="21" pageNumber="472" rank="species">n. sp.</taxonomicNameLabel>
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, in the head pattern, wing venation, in the apex of the aedeagal arch, in the endophallic sclerites, and mainly in that the external parameres are long, slender, curved inward and distally acuminate (compare 
<figureCitation id="89D458F7FFA0FFE4FD4F6B1CFC85FEDD" box="[688,782,326,352]" captionStart="FIGURES 1‒6" captionStartId="11.[151,264,1851,1875]" captionTargetBox="[163,1427,259,1822]" captionTargetId="figure-42@11.[151,1436,234,1825]" captionTargetPageId="11" captionText="FIGURES 1‒6. Goja andina n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531585" httpUri="https://zenodo.org/record/5531585/files/figure.png" pageId="21" pageNumber="472">Figs 1-2</figureCitation>
, 
<figureCitation id="89D458F7FFA0FFE4FCE56B1CFCD0FEDD" box="[794,859,326,352]" captionStart="FIGURES 19‒24" captionStartId="16.[151,264,1875,1899]" captionTargetBox="[152,1425,205,1837]" captionTargetId="figure-21@16.[151,1435,181,1852]" captionTargetPageId="16" captionText="FIGURES 19‒24. Goja risaraldensis n. sp. Male. 19. Forewing. 20. Hindwing. 21. Front view of head. 22. Left paraproct and epiproct. 23. Hypandrium. 24. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531591" httpUri="https://zenodo.org/record/5531591/files/figure.png" pageId="21" pageNumber="472">19-20</figureCitation>
and 
<figureCitation id="89D458F7FFA0FFE4FC6E6B1CFC5EFEDD" box="[913,981,326,352]" captionStart="FIGURES 40‒45" captionStartId="22.[151,264,1973,1997]" captionTargetBox="[163,1425,256,1933]" captionTargetId="figure-61@22.[151,1435,237,1950]" captionTargetPageId="22" captionText="FIGURES 40‒45. Goja vallecaucana n. sp. Male. 40. Forewing. 41. Hindwing. 42. Front view of head. 43. Left paraproct and epiproct. 44. Hypandrium. 45. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531603" httpUri="https://zenodo.org/record/5531603/files/figure.png" pageId="21" pageNumber="472">40-41</figureCitation>
, 
<figureCitation id="89D458F7FFA0FFE4FC1F6B1CFBADFEDD" box="[992,1062,326,352]" captionStart="FIGURES 1‒6" captionStartId="11.[151,264,1851,1875]" captionTargetBox="[163,1427,259,1822]" captionTargetId="figure-42@11.[151,1436,234,1825]" captionTargetPageId="11" captionText="FIGURES 1‒6. Goja andina n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531585" httpUri="https://zenodo.org/record/5531585/files/figure.png" pageId="21" pageNumber="472">Figs 3</figureCitation>
, 
<figureCitation id="89D458F7FFA0FFE4FBCD6B1CFBC5FEDD" box="[1074,1102,326,352]" captionStart="FIGURES 19‒24" captionStartId="16.[151,264,1875,1899]" captionTargetBox="[152,1425,205,1837]" captionTargetId="figure-21@16.[151,1435,181,1852]" captionTargetPageId="16" captionText="FIGURES 19‒24. Goja risaraldensis n. sp. Male. 19. Forewing. 20. Hindwing. 21. Front view of head. 22. Left paraproct and epiproct. 23. Hypandrium. 24. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531591" httpUri="https://zenodo.org/record/5531591/files/figure.png" pageId="21" pageNumber="472">21</figureCitation>
and 
<figureCitation id="89D458F7FFA0FFE4FB7B6B1CFB28FEDD" box="[1156,1187,326,352]" captionStart="FIGURES 40‒45" captionStartId="22.[151,264,1973,1997]" captionTargetBox="[163,1425,256,1933]" captionTargetId="figure-61@22.[151,1435,237,1950]" captionTargetPageId="22" captionText="FIGURES 40‒45. Goja vallecaucana n. sp. Male. 40. Forewing. 41. Hindwing. 42. Front view of head. 43. Left paraproct and epiproct. 44. Hypandrium. 45. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531603" httpUri="https://zenodo.org/record/5531603/files/figure.png" pageId="21" pageNumber="472">42</figureCitation>
, and 
<figureCitation id="89D458F7FFA0FFE4FB236B1CFAA8FEDD" box="[1244,1315,326,352]" captionStart="FIGURES 1‒6" captionStartId="11.[151,264,1851,1875]" captionTargetBox="[163,1427,259,1822]" captionTargetId="figure-42@11.[151,1436,234,1825]" captionTargetPageId="11" captionText="FIGURES 1‒6. Goja andina n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531585" httpUri="https://zenodo.org/record/5531585/files/figure.png" pageId="21" pageNumber="472">Figs 6</figureCitation>
, 
<figureCitation id="89D458F7FFA0FFE4FAD06B1CFAC0FEDD" box="[1327,1355,326,352]" captionStart="FIGURES 19‒24" captionStartId="16.[151,264,1875,1899]" captionTargetBox="[152,1425,205,1837]" captionTargetId="figure-21@16.[151,1435,181,1852]" captionTargetPageId="16" captionText="FIGURES 19‒24. Goja risaraldensis n. sp. Male. 19. Forewing. 20. Hindwing. 21. Front view of head. 22. Left paraproct and epiproct. 23. Hypandrium. 24. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531591" httpUri="https://zenodo.org/record/5531591/files/figure.png" pageId="21" pageNumber="472">24</figureCitation>
and 
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in
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this paper).
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<emphasis id="239B9860FFA0FFE4FF386BD7FE80FE1A" bold="true" box="[199,267,397,423]" pageId="21" pageNumber="472">Color</emphasis>
(in 80% ethanol). Body brown with light spots. Head (
<figureCitation id="89D458F7FFA0FFE4FC7C6BD4FC51FE15" box="[899,986,398,424]" captionStart="FIGURES 40‒45" captionStartId="22.[151,264,1973,1997]" captionTargetBox="[163,1425,256,1933]" captionTargetId="figure-61@22.[151,1435,237,1950]" captionTargetPageId="22" captionText="FIGURES 40‒45. Goja vallecaucana n. sp. Male. 40. Forewing. 41. Hindwing. 42. Front view of head. 43. Left paraproct and epiproct. 44. Hypandrium. 45. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531603" httpUri="https://zenodo.org/record/5531603/files/figure.png" pageId="21" pageNumber="472">Fig. 42</figureCitation>
): vertex dark brown, cream laterally on boundaries with the inner margin of the compound eyes; forehead brown, with dark brown transverse bands. Compound eyes dark brown to black; ocelli hyaline, with ochre centripetal crescents, postclypeus brown, with ochre striae converging mesally, as illustrated; genae cream, with dark brown spots at midline, postgenae and subgenae cream; labrum light brown, with dark brown labral sclerites, anteclypeus brown. Maxillary palps light brown to cream, Mx 1-2 cream, Mx 3-4 brown.Antennae: scape and pedicel light brown, flagellomeres brown. Thorax brown to cream, pronotum light brown, tergal lobes of meso- and metathorax brown, pleura cream. Legs brown to cream, trochanters, femora and tibiae cream, tarsi light brown. Wings hyaline, veins light brown; forewings (
<figureCitation id="89D458F7FFA0FFE4FAF068D0FAEAFD19" box="[1295,1377,650,676]" captionStart="FIGURES 40‒45" captionStartId="22.[151,264,1973,1997]" captionTargetBox="[163,1425,256,1933]" captionTargetId="figure-61@22.[151,1435,237,1950]" captionTargetPageId="22" captionText="FIGURES 40‒45. Goja vallecaucana n. sp. Male. 40. Forewing. 41. Hindwing. 42. Front view of head. 43. Left paraproct and epiproct. 44. Hypandrium. 45. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531603" httpUri="https://zenodo.org/record/5531603/files/figure.png" pageId="21" pageNumber="472">Fig. 40</figureCitation>
) amber, with abundant brown setae. Abdomen brown, sternum light brown to cream. Clunium, hypandrium, epiproct and paraprocts light brown. Phallosome brown, with aedeagal arch and endophallic sclerites dark brown, external parameres brown.
</paragraph>
<paragraph id="11504472FFA0FFE4FF386940FC4BFB21" blockId="21.[151,1437,253,1720]" pageId="21" pageNumber="472">
<emphasis id="239B9860FFA0FFE4FF386940FEEAFC89" bold="true" box="[199,353,794,820]" pageId="21" pageNumber="472">Morphology.</emphasis>
Head: H/MxW: 1.62; compound eyes small, H/d: 3.84; IO/MxW: 0.64. Outer cusp of lacinial tips broad, with eight denticles. Mx4/Mx2: 1.10. Labrum elongate, lower margin with a pair of rounded projections. Forewings (
<figureCitation id="89D458F7FFA0FFE4FEE36938FEF9FCC1" box="[284,370,866,892]" captionStart="FIGURES 40‒45" captionStartId="22.[151,264,1973,1997]" captionTargetBox="[163,1425,256,1933]" captionTargetId="figure-61@22.[151,1435,237,1950]" captionTargetPageId="22" captionText="FIGURES 40‒45. Goja vallecaucana n. sp. Male. 40. Forewing. 41. Hindwing. 42. Front view of head. 43. Left paraproct and epiproct. 44. Hypandrium. 45. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531603" httpUri="https://zenodo.org/record/5531603/files/figure.png" pageId="21" pageNumber="472">Fig. 40</figureCitation>
): FW/W: 2.79; pterostigma elongate, wider in the middle: lp/wp: 5.53; Rs 2-3 branched; M five to six branched. Areola postica elongate: al/ah: 4.89, open in some specimens. Hindwings (
<figureCitation id="89D458F7FFA0FFE4FB6069DCFB7EFC1D" box="[1183,1269,902,928]" captionStart="FIGURES 40‒45" captionStartId="22.[151,264,1973,1997]" captionTargetBox="[163,1425,256,1933]" captionTargetId="figure-61@22.[151,1435,237,1950]" captionTargetPageId="22" captionText="FIGURES 40‒45. Goja vallecaucana n. sp. Male. 40. Forewing. 41. Hindwing. 42. Front view of head. 43. Left paraproct and epiproct. 44. Hypandrium. 45. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531603" httpUri="https://zenodo.org/record/5531603/files/figure.png" pageId="21" pageNumber="472">Fig. 41</figureCitation>
): HW/w: 3.21. Rs 2-branched, M simple or 2-branched. Legs: trochanters with two long setae on dorsal surface; fore- tarsi without ctenidiobothria, mid- and hind- tarsi with ctenidiobothria. Hypandrium (
<figureCitation id="89D458F7FFA0FFE4FC2B6994FBA1FC55" box="[980,1066,974,1000]" captionStart="FIGURES 40‒45" captionStartId="22.[151,264,1973,1997]" captionTargetBox="[163,1425,256,1933]" captionTargetId="figure-61@22.[151,1435,237,1950]" captionTargetPageId="22" captionText="FIGURES 40‒45. Goja vallecaucana n. sp. Male. 40. Forewing. 41. Hindwing. 42. Front view of head. 43. Left paraproct and epiproct. 44. Hypandrium. 45. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531603" httpUri="https://zenodo.org/record/5531603/files/figure.png" pageId="21" pageNumber="472">Fig. 44</figureCitation>
), broad, setose, posterior border concave in the middle. Phallosome (
<figureCitation id="89D458F7FFA0FFE4FDCC69A8FD0CFBB0" box="[563,647,1010,1037]" captionStart="FIGURES 40‒45" captionStartId="22.[151,264,1973,1997]" captionTargetBox="[163,1425,256,1933]" captionTargetId="figure-61@22.[151,1435,237,1950]" captionTargetPageId="22" captionText="FIGURES 40‒45. Goja vallecaucana n. sp. Male. 40. Forewing. 41. Hindwing. 42. Front view of head. 43. Left paraproct and epiproct. 44. Hypandrium. 45. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531603" httpUri="https://zenodo.org/record/5531603/files/figure.png" pageId="21" pageNumber="472">Fig. 45</figureCitation>
), almost straight anteriorly, side struts slender, straight; aedeagal arch robust, apex with two short mesal projections, distally acuminate, curved outward; endophallic sclerites (
<figureCitation id="89D458F7FFA0FFE4FABE6E4CFA1FFB8C" box="[1345,1428,1046,1073]" captionStart="FIGURES 40‒45" captionStartId="22.[151,264,1973,1997]" captionTargetBox="[163,1425,256,1933]" captionTargetId="figure-61@22.[151,1435,237,1950]" captionTargetPageId="22" captionText="FIGURES 40‒45. Goja vallecaucana n. sp. Male. 40. Forewing. 41. Hindwing. 42. Front view of head. 43. Left paraproct and epiproct. 44. Hypandrium. 45. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531603" httpUri="https://zenodo.org/record/5531603/files/figure.png" pageId="21" pageNumber="472">Fig. 45</figureCitation>
) long, slender. Clunium projected over the area of the epiproct. Paraprocts broad (
<figureCitation id="89D458F7FFA0FFE4FBE46E60FBE6FBE9" box="[1051,1133,1082,1108]" captionStart="FIGURES 40‒45" captionStartId="22.[151,264,1973,1997]" captionTargetBox="[163,1425,256,1933]" captionTargetId="figure-61@22.[151,1435,237,1950]" captionTargetPageId="22" captionText="FIGURES 40‒45. Goja vallecaucana n. sp. Male. 40. Forewing. 41. Hindwing. 42. Front view of head. 43. Left paraproct and epiproct. 44. Hypandrium. 45. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531603" httpUri="https://zenodo.org/record/5531603/files/figure.png" pageId="21" pageNumber="472">Fig. 43</figureCitation>
), oval, with setae and macrosetae on posterior border, sensory fields large, elliptic, with 25-28 trichobothria in basal rosettes. Epiproct broadly trapeziform (
<figureCitation id="89D458F7FFA0FFE4FED66ED8FEF7FB21" box="[297,380,1154,1180]" captionStart="FIGURES 40‒45" captionStartId="22.[151,264,1973,1997]" captionTargetBox="[163,1425,256,1933]" captionTargetId="figure-61@22.[151,1435,237,1950]" captionTargetPageId="22" captionText="FIGURES 40‒45. Goja vallecaucana n. sp. Male. 40. Forewing. 41. Hindwing. 42. Front view of head. 43. Left paraproct and epiproct. 44. Hypandrium. 45. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5531603" httpUri="https://zenodo.org/record/5531603/files/figure.png" pageId="21" pageNumber="472">Fig. 43</figureCitation>
), with three setae mesally, other setae as illustrated.
</paragraph>
</subSubSection>
<subSubSection id="59F517F9FFA0FFE4FF386EFCFEEAF929" pageId="21" pageNumber="472" type="description">
<paragraph id="11504472FFA0FFE4FF386EFCFBEDFB59" blockId="21.[151,1437,253,1720]" pageId="21" pageNumber="472">
<emphasis id="239B9860FFA0FFE4FF386EFCFEF5FB7D" bold="true" box="[199,382,1190,1216]" pageId="21" pageNumber="472">Measurements.</emphasis>
FW: 4150, HW: 3050, F: 990, T: 1790, t1: 860, t2: 223, ctt1: 38, f1: 940, f2: 810, f3: 670, f4: 430, f5: 235, f6: 190, f7: 195, Mx4: 325, IO: 273, D: 215, d: 265, IO/d: 1.51, PO: 0.79.
</paragraph>
<paragraph id="11504472FFA0FFE4FF386EB7FEEAF929" blockId="21.[151,1437,253,1720]" pageId="21" pageNumber="472">
<emphasis id="239B9860FFA0FFE4FF386EB7FE23FABA" bold="true" box="[199,424,1261,1287]" pageId="21" pageNumber="472">Specimens studied.</emphasis>
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male. 
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: 
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., 
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, on tree trunk
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. 
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, same locality as the holotype, 
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15. 
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., 
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&amp; 
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&amp; N. 
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, K18, ca. 
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., 
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&amp; 
<collectorName id="BC1A21A4FFA0FFE4FF4A6FF8FEC8FA01" box="[181,323,1442,1468]" pageId="21" pageNumber="472">N. Calderón</collectorName>
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. 
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<location id="143012A9FFA0FFE4FE546FF8FDE3FA01" LSID="urn:lsid:plazi:treatment:9946F564FFA0FFE4FF686ACDFB17F905:143012A9FFA0FFE4FE546FF8FDE3FA01" box="[427,616,1442,1468]" country="Colombia" municipality="Pance" name="Santiago de Cali" pageId="21" pageNumber="472" stateProvince="Valle del Cauca">Santiago de Cali</location>
, 
<collectingMunicipality id="F134DE08FFA0FFE4FD8A6FF8FD30FA01" box="[629,699,1442,1468]" pageId="21" pageNumber="472">Pance</collectingMunicipality>
, 
<collectorName id="BC1A21A4FFA0FFE4FD386FF8FC18FA01" box="[711,915,1442,1468]" pageId="21" pageNumber="472">Quebrada El Pato</collectorName>
, 
<date id="655162B2FFA0FFE4FC606FF8FB94FA01" box="[927,1055,1442,1468]" pageId="21" pageNumber="472" value="2011-06-18">
<collectingDate id="75159B5AFFA0FFE4FC606FF8FB94FA01" box="[927,1055,1442,1468]" pageId="21" pageNumber="472" value="2011-06-18">
18. 
<collectionCode id="77FEDCB7FFA0FFE4FC416FF8FC54FA01" box="[958,991,1442,1468]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="21" pageNumber="472">VI</collectionCode>
.2011
</collectingDate>
</date>
, 
<collectorName id="BC1A21A4FFA0FFE4FBD26FF8FB58FA01" box="[1069,1235,1442,1468]" pageId="21" pageNumber="472">J. A. Mendivil</collectorName>
</materialsCitation>
. 
<materialsCitation id="A1874E2FFFA0FFE4FB206FF8FB9EFA5D" collectingDate="2011-03-19" collectionCode="R" collectorName="Gonzalez" country="Colombia" county="Palo Alto" elevation="1866" latitude="3.5691667" location="Yumbo" longLatPrecision="1" longitude="-76.57044" municipality="Dapa" pageId="21" pageNumber="472" specimenCount="2" specimenCount-male="2" stateProvince="Valle del Cauca" typeStatus="paratype">
<specimenCount id="07E98FFBFFA0FFE4FB206FF8FAB2FA01" box="[1247,1337,1442,1468]" pageId="21" pageNumber="472" type="male">2 males</specimenCount>
, 
<location id="143012A9FFA0FFE4FABB6FF8FA13FA01" LSID="urn:lsid:plazi:treatment:9946F564FFA0FFE4FF686ACDFB17F905:143012A9FFA0FFE4FABB6FF8FA13FA01" box="[1348,1432,1442,1468]" country="Colombia" county="Palo Alto" latitude="3.5691667" longLatPrecision="1" longitude="-76.57044" municipality="Dapa" name="Yumbo" pageId="21" pageNumber="472" stateProvince="Valle del Cauca">Yumbo</location>
, 
<collectingMunicipality id="F134DE08FFA0FFE4FF686F9CFF5EFA5D" box="[151,213,1478,1504]" pageId="21" pageNumber="472">Dapa</collectingMunicipality>
, 
<collectingCounty id="F8313CFEFFA0FFE4FF1F6F9CFEC0FA5D" box="[224,331,1478,1504]" pageId="21" pageNumber="472">Palo Alto</collectingCounty>
, 
<geoCoordinate id="74DB22B5FFA0FFE4FEAA6F9CFE57FA5D" box="[341,476,1478,1504]" degrees="3" direction="north" minutes="34" orientation="latitude" pageId="21" pageNumber="472" precision="1" seconds="9.0" value="3.5691667">3°34’9.0”N</geoCoordinate>
: 
<geoCoordinate id="74DB22B5FFA0FFE4FE196F9CFD00FA5D" box="[486,651,1478,1504]" degrees="76" direction="west" minutes="34" orientation="longitude" pageId="21" pageNumber="472" precision="1" seconds="13.6" value="-76.57044">76°34’13.6”W</geoCoordinate>
, 
<quantity id="D617E997FFA0FFE4FD6A6F9CFD65FA5D" box="[661,750,1478,1504]" metricMagnitude="3" metricUnit="m" metricValue="1.866" pageId="21" pageNumber="472" unit="m" value="1866.0">
<elevation id="9AC2A341FFA0FFE4FD6A6F9CFD65FA5D" box="[661,750,1478,1504]" metricMagnitude="3" metricUnit="m" metricValue="1.866" pageId="21" pageNumber="472" unit="m" value="1866.0">1866 m</elevation>
</quantity>
., 
<date id="655162B2FFA0FFE4FD016F9CFCF6FA5D" box="[766,893,1478,1504]" pageId="21" pageNumber="472" value="2011-03-19">
<collectingDate id="75159B5AFFA0FFE4FD016F9CFCF6FA5D" box="[766,893,1478,1504]" pageId="21" pageNumber="472" value="2011-03-19">19.III.2011</collectingDate>
</date>
, 
<collectionCode id="77FEDCB7FFA0FFE4FC776F9CFC15FA5D" box="[904,926,1478,1504]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="21" pageNumber="472">R</collectionCode>
. 
<collectorName id="BC1A21A4FFA0FFE4FC576F9CFB9EFA5D" box="[936,1045,1478,1504]" pageId="21" pageNumber="472">González</collectorName>
</materialsCitation>
. 
<materialsCitation id="A1874E2FFFA0FFE4FBE06F9CFECBF9B9" collectingDate="2011-06-17" collectionCode="VI" collectorName="Mendivil" country="Colombia" county="Palo Alto" elevation="1866" latitude="3.5691667" location="Yumbo" longLatPrecision="1" longitude="-76.57044" municipality="Dapa" pageId="21" pageNumber="472" specimenCount="1" specimenCount-male="1" stateProvince="Valle del Cauca" typeStatus="paratype">
<specimenCount id="07E98FFBFFA0FFE4FBE06F9CFBE7FA5D" box="[1055,1132,1478,1504]" pageId="21" pageNumber="472" type="male">1 male</specimenCount>
, same locality, 
<date id="655162B2FFA0FFE4FAE86F9CFA1CFA5D" box="[1303,1431,1478,1504]" pageId="21" pageNumber="472" value="2011-06-17">
<collectingDate id="75159B5AFFA0FFE4FAE86F9CFA1CFA5D" box="[1303,1431,1478,1504]" pageId="21" pageNumber="472" value="2011-06-17">
17. 
<collectionCode id="77FEDCB7FFA0FFE4FAC96F9CFADCFA5D" box="[1334,1367,1478,1504]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="21" pageNumber="472">VI</collectionCode>
.2011
</collectingDate>
</date>
, J. A. 
<collectorName id="BC1A21A4FFA0FFE4FF296FB0FECBF9B9" box="[214,320,1514,1540]" pageId="21" pageNumber="472">Mendivil</collectorName>
</materialsCitation>
. 
<specimenCount id="07E98FFBFFA0FFE4FEB26FB0FE15F9B9" box="[333,414,1514,1540]" pageId="21" pageNumber="472" type="male">1 male</specimenCount>
, 
<date id="655162B2FFA0FFE4FE536FB0FDA6F9B9" box="[428,557,1514,1540]" pageId="21" pageNumber="472" value="2011-09-10">10.IX.2011</date>
, J. A. Mendivil. 
<materialsCitation id="A1874E2FFFA0FFE4FD0E6FB0FDB4F995" collectingDate="2011-06-27" collectionCode="VI" collectorName="La Cumbre &amp; J. A. Mendivil" country="Colombia" elevation="1722" latitude="3.5750833" location="Chicoral" longLatPrecision="1" longitude="-76.59339" pageId="21" pageNumber="472" specimenCount="1" specimenCount-male="1" stateProvince="Valle del Cauca" typeStatus="paratype">
<specimenCount id="07E98FFBFFA0FFE4FD0E6FB0FCCAF9B9" box="[753,833,1514,1540]" pageId="21" pageNumber="472" type="male">1 male</specimenCount>
, 
<collectorName id="BC1A21A4FFA0FFE4FCB06FB0FC58F9B9" box="[847,979,1514,1540]" pageId="21" pageNumber="472">La Cumbre</collectorName>
, 
<location id="143012A9FFA0FFE4FC1E6FB0FBCFF9B9" LSID="urn:lsid:plazi:treatment:9946F564FFA0FFE4FF686ACDFB17F905:143012A9FFA0FFE4FC1E6FB0FBCFF9B9" box="[993,1092,1514,1540]" country="Colombia" latitude="3.5750833" longLatPrecision="1" longitude="-76.59339" name="Chicoral" pageId="21" pageNumber="472" stateProvince="Valle del Cauca">Chicoral</location>
, 
<geoCoordinate id="74DB22B5FFA0FFE4FBAD6FB0FB6DF9B9" box="[1106,1254,1514,1540]" degrees="3" direction="north" minutes="34" orientation="latitude" pageId="21" pageNumber="472" precision="1" seconds="30.3" value="3.5750833">3°34’30.3”N</geoCoordinate>
: 
<geoCoordinate id="74DB22B5FFA0FFE4FB0C6FB0FA1CF9B8" box="[1267,1431,1514,1541]" degrees="76" direction="west" minutes="35" orientation="longitude" pageId="21" pageNumber="472" precision="1" seconds="36.2" value="-76.59339">76°35’36.2”W</geoCoordinate>
, 
<quantity id="D617E997FFA0FFE4FF686C54FF7FF995" box="[151,244,1550,1576]" metricMagnitude="3" metricUnit="m" metricValue="1.722" pageId="21" pageNumber="472" unit="m" value="1722.0">
<elevation id="9AC2A341FFA0FFE4FF686C54FF7FF995" box="[151,244,1550,1576]" metricMagnitude="3" metricUnit="m" metricValue="1.722" pageId="21" pageNumber="472" unit="m" value="1722.0">1722 m</elevation>
</quantity>
., 
<date id="655162B2FFA0FFE4FEF86C54FE03F995" box="[263,392,1550,1576]" pageId="21" pageNumber="472" value="2011-06-27">
<collectingDate id="75159B5AFFA0FFE4FEF86C54FE03F995" box="[263,392,1550,1576]" pageId="21" pageNumber="472" value="2011-06-27">
27. 
<collectionCode id="77FEDCB7FFA0FFE4FED76C54FEC3F995" box="[296,328,1550,1576]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="21" pageNumber="472">VI</collectionCode>
.2011
</collectingDate>
</date>
, 
<collectorName id="BC1A21A4FFA0FFE4FE696C54FDB4F995" box="[406,575,1550,1576]" pageId="21" pageNumber="472">J. A. Mendivil</collectorName>
</materialsCitation>
. 
<materialsCitation id="A1874E2FFFA0FFE4FDB36C54FD9CF9F1" collectingDate="2011-01-12" collectionCode="R" collectorName="Gonzalez &amp; A. N. Garcia Aldrete" country="Colombia" elevation="1609" latitude="3.8666668" location="Yotoco" longLatPrecision="1307" longitude="-76.38333" municipality="Bosque Yotoco" pageId="21" pageNumber="472" specimenCount="1" specimenCount-male="1" stateProvince="Valle del Cauca" typeStatus="paratype">
<specimenCount id="07E98FFBFFA0FFE4FDB36C54FD16F995" box="[588,669,1550,1576]" pageId="21" pageNumber="472" type="male">1 male</specimenCount>
, 
<location id="143012A9FFA0FFE4FD556C54FD76F995" LSID="urn:lsid:plazi:treatment:9946F564FFA0FFE4FF686ACDFB17F905:143012A9FFA0FFE4FD556C54FD76F995" box="[682,765,1550,1576]" country="Colombia" latitude="3.8666668" longLatPrecision="1307" longitude="-76.38333" municipality="Bosque Yotoco" name="Yotoco" pageId="21" pageNumber="472" stateProvince="Valle del Cauca">Yotoco</location>
, 
<collectingMunicipality id="F134DE08FFA0FFE4FCF46C54FC30F995" box="[779,955,1550,1576]" pageId="21" pageNumber="472">Bosque Yotoco</collectingMunicipality>
, 
<geoCoordinate id="74DB22B5FFA0FFE4FC376C54FB94F994" box="[968,1055,1550,1577]" degrees="3" direction="north" minutes="52" orientation="latitude" pageId="21" pageNumber="472" precision="925" value="3.8666668">3°52’N</geoCoordinate>
: 
<geoCoordinate id="74DB22B5FFA0FFE4FBD36C54FB01F995" box="[1068,1162,1550,1576]" degrees="76" direction="west" minutes="23" orientation="longitude" pageId="21" pageNumber="472" precision="925" value="-76.38333">76°23W</geoCoordinate>
, 
<quantity id="D617E997FFA0FFE4FB676C54FB7EF995" box="[1176,1269,1550,1576]" metricMagnitude="3" metricUnit="m" metricValue="1.609" pageId="21" pageNumber="472" unit="m" value="1609.0">
<elevation id="9AC2A341FFA0FFE4FB676C54FB7EF995" box="[1176,1269,1550,1576]" metricMagnitude="3" metricUnit="m" metricValue="1.609" pageId="21" pageNumber="472" unit="m" value="1609.0">1609 m</elevation>
</quantity>
., 
<date id="655162B2FFA0FFE4FAF86C54FAFFF995" box="[1287,1396,1550,1576]" pageId="21" pageNumber="472" value="2011-01-12">
<collectingDate id="75159B5AFFA0FFE4FAF86C54FAFFF995" box="[1287,1396,1550,1576]" pageId="21" pageNumber="472" value="2011-01-12">12.I.2011</collectingDate>
</date>
, 
<collectionCode id="77FEDCB7FFA0FFE4FA7C6C54FA12F995" box="[1411,1433,1550,1576]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="21" pageNumber="472">R</collectionCode>
. 
<collectorName id="BC1A21A4FFA0FFE4FF686C68FE8AF9F1" box="[151,257,1586,1612]" pageId="21" pageNumber="472">González</collectorName>
&amp; 
<collectorName id="BC1A21A4FFA0FFE4FED96C68FD9CF9F1" box="[294,535,1586,1612]" pageId="21" pageNumber="472">A. N. García Aldrete</collectorName>
</materialsCitation>
. 
<materialsCitation id="A1874E2FFFA0FFE4FDDD6C68FE8FF9CD" collectingDate="2010-11-06" collectionCode="R" collectorName="Gonzalez" country="Colombia" elevation="1600" latitude="3.8893054" location="Darien" longLatPrecision="923" longitude="-76.4" municipality="Puente Tierra" pageId="21" pageNumber="472" specimenCount="1" specimenCount-male="1" stateProvince="Valle del Cauca" typeStatus="paratype">
<specimenCount id="07E98FFBFFA0FFE4FDDD6C68FDFAF9F1" box="[546,625,1586,1612]" pageId="21" pageNumber="472" type="male">1 male</specimenCount>
, 
<location id="143012A9FFA0FFE4FD816C68FD46F9F1" LSID="urn:lsid:plazi:treatment:9946F564FFA0FFE4FF686ACDFB17F905:143012A9FFA0FFE4FD816C68FD46F9F1" box="[638,717,1586,1612]" country="Colombia" latitude="3.8893054" longLatPrecision="923" longitude="-76.4" municipality="Puente Tierra" name="Darien" pageId="21" pageNumber="472" stateProvince="Valle del Cauca">Darién</location>
, 
<collectingMunicipality id="F134DE08FFA0FFE4FD256C68FCFFF9F1" box="[730,884,1586,1612]" pageId="21" pageNumber="472">Puente Tierra</collectingMunicipality>
, 
<geoCoordinate id="74DB22B5FFA0FFE4FC7E6C68FB9EF9F0" box="[897,1045,1586,1613]" degrees="3" direction="north" minutes="53" orientation="latitude" pageId="21" pageNumber="472" precision="1" seconds="21.5" value="3.8893054">3°53’21.5”N</geoCoordinate>
: 
<geoCoordinate id="74DB22B5FFA0FFE4FBDE6C68FB03F9F1" box="[1057,1160,1586,1612]" degrees="76" direction="west" minutes="24" orientation="longitude" pageId="21" pageNumber="472" precision="925" value="-76.4">76°24’W</geoCoordinate>
, 
<quantity id="D617E997FFA0FFE4FB6A6C68FB7BF9F1" box="[1173,1264,1586,1612]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="21" pageNumber="472" unit="m" value="1600.0">
<elevation id="9AC2A341FFA0FFE4FB6A6C68FB7BF9F1" box="[1173,1264,1586,1612]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="21" pageNumber="472" unit="m" value="1600.0">1600 m</elevation>
</quantity>
., 
<date id="655162B2FFA0FFE4FAFD6C68FAFEF9F1" box="[1282,1397,1586,1612]" pageId="21" pageNumber="472" value="2010-11-06">
<collectingDate id="75159B5AFFA0FFE4FAFD6C68FAFEF9F1" box="[1282,1397,1586,1612]" pageId="21" pageNumber="472" value="2010-11-06">6.XI.2010</collectingDate>
</date>
, 
<collectionCode id="77FEDCB7FFA0FFE4FA7C6C68FA12F9F1" box="[1411,1433,1586,1612]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="21" pageNumber="472">R</collectionCode>
. 
<collectorName id="BC1A21A4FFA0FFE4FF686C0CFE8FF9CD" box="[151,260,1622,1648]" pageId="21" pageNumber="472">González</collectorName>
</materialsCitation>
. 
<materialsCitation id="A1874E2FFFA0FFE4FEF16C0CFED6F929" collectingDate="2013-03-10" collectorName="O. Saenz &amp; N. Calderon" country="Colombia" elevation="1781" latitude="4.4493613" location="Roldanillo" longLatPrecision="1" longitude="-7.22275" municipality="Via Belgica" pageId="21" pageNumber="472" specimenCount="1" specimenCount-male="1" stateProvince="Valle del Cauca" typeStatus="paratype">
<specimenCount id="07E98FFBFFA0FFE4FEF16C0CFED0F9CD" box="[270,347,1622,1648]" pageId="21" pageNumber="472" type="male">1 male</specimenCount>
, 
<location id="143012A9FFA0FFE4FE9A6C0CFE55F9CD" LSID="urn:lsid:plazi:treatment:9946F564FFA0FFE4FF686ACDFB17F905:143012A9FFA0FFE4FE9A6C0CFE55F9CD" box="[357,478,1622,1648]" country="Colombia" latitude="4.4493613" longLatPrecision="1" longitude="-7.22275" municipality="Via Belgica" name="Roldanillo" pageId="21" pageNumber="472" stateProvince="Valle del Cauca">Roldanillo</location>
, 
<collectingMunicipality id="F134DE08FFA0FFE4FE176C0CFDE5F9CD" box="[488,622,1622,1648]" pageId="21" pageNumber="472">Vía Bélgica</collectingMunicipality>
, 
<geoCoordinate id="74DB22B5FFA0FFE4FD866C0CFC86F9CC" box="[633,781,1622,1649]" degrees="4" direction="north" minutes="26" orientation="latitude" pageId="21" pageNumber="472" precision="1" seconds="57.7" value="4.4493613">4°26’57.7”N</geoCoordinate>
: 
<geoCoordinate id="74DB22B5FFA0FFE4FCE86C0CFC26F9CD" box="[791,941,1622,1648]" degrees="7" direction="west" minutes="13" orientation="longitude" pageId="21" pageNumber="472" precision="1" seconds="21.9" value="-7.22275">7°13’21.9”W</geoCoordinate>
, 
<quantity id="D617E997FFA0FFE4FC476C0CFB9AF9CD" box="[952,1041,1622,1648]" metricMagnitude="3" metricUnit="m" metricValue="1.781" pageId="21" pageNumber="472" unit="m" value="1781.0">
<elevation id="9AC2A341FFA0FFE4FC476C0CFB9AF9CD" box="[952,1041,1622,1648]" metricMagnitude="3" metricUnit="m" metricValue="1.781" pageId="21" pageNumber="472" unit="m" value="1781.0">1781 m</elevation>
</quantity>
., 
<date id="655162B2FFA0FFE4FBE06C0CFB2BF9CD" box="[1055,1184,1622,1648]" pageId="21" pageNumber="472" value="2013-03-10">
<collectingDate id="75159B5AFFA0FFE4FBE06C0CFB2BF9CD" box="[1055,1184,1622,1648]" pageId="21" pageNumber="472" value="2013-03-10">10.III.2013</collectingDate>
</date>
, 
<collectorName id="BC1A21A4FFA0FFE4FB556C0CFA84F9CD" box="[1194,1295,1622,1648]" pageId="21" pageNumber="472">O. Sáenz</collectorName>
&amp; 
<collectorName id="BC1A21A4FFA0FFE4FACF6C0CFF34F929" pageId="21" pageNumber="472">N. Calderón</collectorName>
, on tree trunk
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="59F517F9FFA0FFE4FF386CC4FB17F905" box="[199,1180,1694,1720]" pageId="21" pageNumber="472" type="etymology">
<paragraph id="11504472FFA0FFE4FF386CC4FB17F905" blockId="21.[151,1437,253,1720]" box="[199,1180,1694,1720]" pageId="21" pageNumber="472">
<emphasis id="239B9860FFA0FFE4FF386CC4FEC5F905" bold="true" box="[199,334,1694,1720]" pageId="21" pageNumber="472">Etymology.</emphasis>
The specific epithet refers to the department of 
<collectingRegion id="D32B8A90FFA0FFE4FC976CC4FB90F905" box="[872,1051,1694,1720]" country="Colombia" name="Valle del Cauca" pageId="21" pageNumber="472">Valle del Cauca</collectingRegion>
, 
<collectingCountry id="69F804E2FFA0FFE4FBD96CC4FB13F905" box="[1062,1176,1694,1720]" name="Colombia" pageId="21" pageNumber="472">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>