363 lines
35 KiB
XML
363 lines
35 KiB
XML
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<mods:title id="79B6FCB2806916101CAE1DF4FFFB6893">On two Italian Lithobius species described by Silvestri, with taxonomic notes on the genus Eupolybothrus Verhoeff (Chilopoda, Lithobiida)</mods:title>
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<mods:namePart id="8FF8B1C2D1FED203E76968FAD4094178">C. A. W. Jeekel</mods:namePart>
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<mods:title id="40F81EE1D1B8719139A06F04892FE29C">BEAUFORTIA SERIES OF MISCELLANEOUS PUBLICATIONS</mods:title>
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<mods:date id="B14ED57BC15A6D49DE3AF7E6037AE6D9">1967</mods:date>
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<mods:number id="8493B8759AFEEC2B0DA0E8F73A574396">14</mods:number>
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<mods:classification id="3DF8C98352C09B164F1F79F71AF9FF63">journal article</mods:classification>
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<treatment id="981F8DB7952D83338DFD7245D3D88E80" ID-DOI="http://doi.org/10.5281/zenodo.6285000" ID-GBIF-Taxon="100073138" ID-Zenodo-Dep="6285000" LSID="urn:lsid:plazi:treatment:981F8DB7952D83338DFD7245D3D88E80" httpUri="http://treatment.plazi.org/id/981F8DB7952D83338DFD7245D3D88E80" lastPageId="7" lastPageNumber="172" pageId="5" pageNumber="170">
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<subSubSection id="CBCB37132C53486947087DD1374B7BC9" pageId="5" pageNumber="170" type="nomenclature">
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<paragraph id="E66074761A6A1E45D10DDA97D3C24659" pageId="5" pageNumber="170" type="mainText">
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<pageBreakToken id="23F0FA5FB37F5BDD0474085A2AF18A71" pageId="5" pageNumber="170" start="start">Subgenus</pageBreakToken>
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<taxonomicName id="3996A7DB0C882BA86B8EA3C6945514E3" ID-CoL="8G4T" authority="Verhoeff, 1934" authorityName="Verhoeff" authorityYear="1934" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="subGenus" subGenus="Schizopolybothrus">
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Schizopolybothrus
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<authority id="EB0545F3150A732C68DC1979F1BB2580" pageId="5" pageNumber="170">Verhoeff, 1934</authority>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="57D24E8FA0BA5460C9A868EBD04C5B9E" pageId="5" pageNumber="170" type="diagnosis">
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<paragraph id="79895BAA6A30F9A65DD3B8411F161DCF" pageId="5" pageNumber="170" type="mainText">
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This subgenus was monotypically based on
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<taxonomicName id="8B841FFA57929D6E6DF1B9FDE4218DB0" authority="Verhoeff, 1899" authorityName="Verhoeff" authorityYear="1899" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="species" species="caesar">
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E. caesar (
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<authority id="558409EF8A4AFE7C28A42596990158E1" pageId="5" pageNumber="170">Verhoeff, 1899</authority>
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)
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</taxonomicName>
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, to which later
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<taxonomicName id="920FF22102BCB1D0603DC3F417F5F825" authority="Verhoeff, 1937" authorityName="Verhoeff" authorityYear="1937" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="species" species="tabularum">
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E. tabularum (
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<authority id="E997761DE102033B31A1D0CE427F2D9D" pageId="5" pageNumber="170">Verhoeff, 1937</authority>
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)
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</taxonomicName>
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was added. In the present concept
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<taxonomicName id="3B5BE0908BAD55866D71B1EDCA89D332" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="subGenus" subGenus="Schizopolybothrus">Schizopolybothrus</taxonomicName>
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comprises over a dozen of forms occurring in North Italy, Jugoslavia, Albania, and Greece. Apparently the species of this subgenus are rare, and most of the described forms are known only from the original material.
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</paragraph>
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</subSubSection>
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<subSubSection id="2D14BFE345843353DC774553C7094A17" pageId="5" pageNumber="170" type="description">
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<paragraph id="3A648C31FB528190B8574746FD2C1C78" pageId="5" pageNumber="170" type="mainText">The characters of the subgenus are the presence of triangular projections on the 9th, 11th and 13th tergites, and a VCm spine on the 15th leg. The 15th leg has one to four VCa spines, and a single claw. 14th leg with one to three VCa spines.</paragraph>
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<paragraph id="BD0AFDA552AD8498F18091410702A24C" pageId="5" pageNumber="170" type="mainText">According to the structure of the last two pairs of legs of the male and the male gonopods, the species can be divided into three categories.</paragraph>
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<paragraph id="A6D3F43A42C8A8F51B6B32C6C0AFBA45" pageId="5" pageNumber="170" type="mainText">I. The 15th leg with a large rounded knob proximad of the middle of the caudal side of the prefemur. The prefemur of the 14th leg often has a triangular process distally on the dorso-caudal side. Gonopods short.</paragraph>
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<paragraph id="E65B11DFC04440C31811A511A524E7EA" pageId="5" pageNumber="170" type="mainText">
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To this group belong
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<taxonomicName id="AF8868054509A6C0DBB19ABB035667D4" authority="Verhoeff, 1899" authorityName="Verhoeff" authorityYear="1899" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="species" species="caesar">
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E. caesar (
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<authority id="20CDD9853D2377F4E1D12C3134F90B0D" pageId="5" pageNumber="170">Verhoeff, 1899</authority>
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)
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</taxonomicName>
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,
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<taxonomicName id="FE3011F8A1A0D5062C88621F3FC9EA92" authority="Verhoeff, 1905" authorityName="Verhoeff" authorityYear="1905" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="subSpecies" species="caesar" subSpecies="valonensis">
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E. caesar ssp. valonensis (
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<authority id="7B1933674A72DC34E70A5EAE8A474402" pageId="5" pageNumber="170">Verhoeff, 1905</authority>
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)
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</taxonomicName>
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,
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<taxonomicName id="4BD89B65C05BCCE70478A6B88D091060" authority="Attems, 1935" authorityName="Attems" authorityYear="1935" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="subSpecies" species="caesar" subSpecies="patens">
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E. caesar
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<taxonomicNameLabel id="7801609097D385F873F74BD1A984260C" pageId="5" pageNumber="170">ssp.</taxonomicNameLabel>
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patens (
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<authority id="A67C696C9DFC4F8DC306F83B8F60F6D1" pageId="5" pageNumber="170">Attems, 1935</authority>
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)
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</taxonomicName>
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,
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<taxonomicName id="DC623DAD609521A3471C1DC2D94A3B23" authority="Latzel, 1888" authorityName="Latzel" authorityYear="1888" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="species" species="spiniger">
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E. spiniger (
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<authority id="E79BF2DED9EED8332787C86ECA521B73" pageId="5" pageNumber="170">Latzel, 1888</authority>
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)
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</taxonomicName>
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,
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<taxonomicName id="EE42D56C49E70606E093D62C54C5A8B6" authority="Verhoeff, 1900" authorityName="Verhoeff" authorityYear="1900" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="species" species="acherontis">
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E. acherontis (
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<authority id="DBC9AF1B84F1572218F09449D392B508" pageId="5" pageNumber="170">Verhoeff, 1900</authority>
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)
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</taxonomicName>
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(probably, although only the female has been described),
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<taxonomicName id="A4E5CBDB420CB3E4E26103DD1F2A5D70" authority="Verhoeff, 1937" authorityName="Verhoeff" authorityYear="1937" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="subSpecies" species="acherontis" subSpecies="wardaranus">
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E. acherontis ssp. wardaranus (
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<authority id="9B3F1DFC30D395B8BEAA4617E6F1D5D1" pageId="5" pageNumber="170">Verhoeff, 1937</authority>
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)
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</taxonomicName>
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,
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<taxonomicName id="49E122D0E0883E7501C6F169782547F6" authority="Folkmanova, 1940" authorityName="Folkmanova" authorityYear="1940" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="species" species="stygis">
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E. stygis (
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<authority id="937168ABC11FC3646A62509E5B52D6CA" pageId="5" pageNumber="170">Folkmanova, 1940</authority>
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)
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</taxonomicName>
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, and
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<taxonomicName id="0B2AD747C7561CE30993CCCC714A16AC" authority="Verhoeff, 1899" authorityName="Verhoeff" authorityYear="1899" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="species" species="leostygis">
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E. leostygis (
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<authority id="C19B1B8F7CCBABAFD2DD0338789B8BEF" pageId="5" pageNumber="170">Verhoeff, 1899</authority>
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)
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</taxonomicName>
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. Its known distribution includes Jugoslavia (Bosnia, Hercegovina, Dalmatia, Macedonia), Albania, and Greece (Corfu, Epirus).
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<taxonomicName id="FE2AD7C78518E4D26D75480B248E0075" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="species" species="stygis">E. stygis</taxonomicName>
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and
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<taxonomicName id="9E587925E0F115371FC996FC2EA7C17A" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="species" species="leostygis">E. leostygis</taxonomicName>
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were originally described from caves.
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</paragraph>
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<paragraph id="13CFD3BF784B6DDFEE4E32D50BAF1677" pageId="5" pageNumber="170" type="mainText">II. The 14th and 15th legs without special modifications. Gonopods short.</paragraph>
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<paragraph id="3A2FE5E9B47FC9B8468E74DE2D39822D" pageId="5" pageNumber="170" type="mainText">
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Here belong
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<taxonomicName id="7DA063ED3377DE21CF818D2BF8B09EDA" authority="Verhoeff, 1937" authorityName="Verhoeff" authorityYear="1937" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="species" species="tabularum">
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E. tabularum (
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<authority id="0B29ED33DCE3C708382234D0DCECA689" pageId="5" pageNumber="170">Verhoeff, 1937</authority>
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)
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</taxonomicName>
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, and probably also
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<taxonomicName id="03CC0BBFE2631FAF1D4B92F92F43F88D" authority="Silvestri, 1894" authorityName="Silvestri" authorityYear="1894" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="species" species="excellens">
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E. excellens (
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<authority id="EB625F1E73C6191B5FD9D5E4B9272D1B" pageId="5" pageNumber="170">Silvestri, 1894</authority>
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)
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</taxonomicName>
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, although the male characters of the latter are unknown. The two species were described from northwestern Italy (Prov. Cuneo, Prov. Genova),
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<taxonomicName id="CC3E93B230433F1D840E40BF1F408809" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="species" species="excellens">E. excellens</taxonomicName>
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from caves.
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</paragraph>
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<paragraph id="C89807639DB8DD07F339AF6F8B0FDBB5" pageId="5" pageNumber="170" type="mainText">III. Prefemur of the 15th leg with a deep dorsal furrow. Gonopods long.</paragraph>
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</subSubSection>
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<subSubSection id="12B03CB72EEF330138A6D13140142D0D" pageId="5" pageNumber="170" type="diagnosis">
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<paragraph id="84C27D44A1E4ED0392CB9B2B0577A9B7" pageId="5" pageNumber="170" type="mainText">
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The two known species,
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<taxonomicName id="51B94C95BF128C7571BEB3D88D35C167" authority="Verhoeff, 1901" authorityName="Verhoeff" authorityYear="1901" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="species" species="zeus">
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E. zeus (
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<authority id="87A08C65C2C1B56CBBEDD85A41D4734C" pageId="5" pageNumber="170">Verhoeff, 1901</authority>
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)
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</taxonomicName>
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and
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<taxonomicName id="5CE37349F95D9097F21B7CB3239420B0" authority="Kanellis, 1959" authorityName="Kanellis" authorityYear="1959" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="species" species="sissii">
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E. sissii (
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<authority id="8149B6B557C8B2C48BA227E39E2E75AE" pageId="5" pageNumber="170">Kanellis, 1959</authority>
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)
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</taxonomicName>
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, were described from Central Greece and Euboea, respectively.
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</paragraph>
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</subSubSection>
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<subSubSection id="EBBB896B18B93B0F23F54CDE800D3914" lastPageId="6" lastPageNumber="171" pageId="5" pageNumber="170" type="discussion">
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<paragraph id="DDC54F4BA81E18B6526AA17B7C131469" pageId="5" pageNumber="170" type="mainText">
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The characters by which the species and subspecies within these three categories are separated concern mainly differences in the numbers of ocelli, antennomeres and coxosternal teeth, and the spinulation of the legs. Their significance in many cases seems dubious, and it is very likely that the groups II and III actually concern only one species each. One may even wonder if not the so-called species and subspecies of group I are just elements of one single polytypic species. Probably the cave-dwelling specimens, with their more elevated number of antennomeres and sometimes reduced number of ocelli, do not represent one or more different troglobiont species, but are merely the cave forms of one troglophilous species. Given the usual instability of the mentioned characters in
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<taxonomicName id="15E5278A6579A98C4545342FDEDBA510" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="170" phylum="Arthropoda" rank="genus">Eupolybothrus</taxonomicName>
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, the influence of cave life oh the ontogenetic development might well result here in a considerable amount of variability.
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</paragraph>
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<paragraph id="6997AE4B064D2217790BC1C16087679A" pageId="6" pageNumber="171" type="mainText">
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<pageBreakToken id="40299698A18A704925170BD73C094E32" pageId="6" pageNumber="171" start="start">Considering</pageBreakToken>
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the above, it is clear that the following key to the species of
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<taxonomicName id="A753FDC8ACF689E8366E3166F4F71335" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="subGenus" subGenus="Schizopolybothrus">Schizopolybothrus</taxonomicName>
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can be regarded only as a guide to the published descriptions and not as an identification key in the usual sense.
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</paragraph>
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</subSubSection>
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<subSubSection id="FA8E994175A9578FF1DF4A0C74E398E3" pageId="6" pageNumber="171" type="key">
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<paragraph id="56331FA3DC7DAD5B9665C549692EE743" pageId="6" pageNumber="171" type="mainText">
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KEY TO THE SPECIES AND SUBSPECIES OF
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<taxonomicName id="4F618E555F91023698C84EA10A6C991F" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="subGenus" subGenus="Schizopolybothrus">SCHIZOPOLYBOTHRUS VERHOEFF</taxonomicName>
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</paragraph>
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<paragraph id="A9A02C72B9C1D6CE5187379CB642C928" pageId="6" pageNumber="171" type="mainText">1. Antennae composed of 61 to 83 a ntennomeres... 2</paragraph>
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<paragraph id="034F6C25A130FD273606A8AF04CCD03F" pageId="6" pageNumber="171" type="mainText">- Antennae composed of 38 to 58 anten nomeres... 6</paragraph>
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<paragraph id="B1CE0B5D6DCB31D5F5FD1A07428865E1" pageId="6" pageNumber="171" type="mainText">
|
|
2. Eyes consisting of 2 to 5 (10) ocelli. 73 to 80 a ntennomeres. 7 + 7 to 11 + 11 coxosternal teeth. Ventral spinulation of 15th leg: 1, 1, 3-4, 2-3, 1-2; 2 or 3 VCa spines...
|
|
<taxonomicName id="467514EBB0D8B7DF162CD53577C2F3B6" authority="Verh." authorityName="Verh." class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="species" species="leoslygis">
|
|
E. leoslygis (
|
|
<authority id="7547A3030681E838E15480C666872D48" pageId="6" pageNumber="171">Verh.</authority>
|
|
)
|
|
</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="56708423AA6EE5EFC877F6B728183CD7" pageId="6" pageNumber="171" type="mainText">- Eyes consisting of 13 to 23 ocelli... 3</paragraph>
|
|
<paragraph id="3B81235A9858E7F38B52E6D052E91658" pageId="6" pageNumber="171" type="mainText">3. 74 to 83 antennomeres. 15th leg with 2 to 4 VCa spines, in the male with a large prefemoral knob. Jugoslavian species... 4</paragraph>
|
|
<paragraph id="917354F641A3C982C22705E5D217C193" pageId="6" pageNumber="171" type="mainText">- 61 to 76 antennomeres. 15th leg with 1 or 2 VCa spines, in the male without modifications. Italian species... 5</paragraph>
|
|
<paragraph id="886EE8942105D23DA1C3CAF6D76EAC6A" pageId="6" pageNumber="171" type="mainText">
|
|
4. 15 ocelli. 74 antennomeres. 9 + 9 coxosterna 1 teeth. Ventral spinulation of 14th leg: 1, 1, 3, 2, 1; 2 VCa spines. 15th leg with 3 VCa spines...
|
|
<taxonomicName id="7A542197B51FE6DB12CDFB3A90F1563E" authority="Verh." authorityName="Verh." class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="species" species="acherontis">
|
|
E. acherontis (
|
|
<authority id="490E76FFAB309386755286068ADDE57C" pageId="6" pageNumber="171">Verh.</authority>
|
|
)
|
|
</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="2DA13BA642B6A0D7A99E719FE73D784F" pageId="6" pageNumber="171" type="mainText">
|
|
- 18 to 20 ocelli, 81 to 83 antennomeres. 8 + 8 or 9 + 9 coxosternal teeth. Ventral spinulation of 14th leg: 1, 1, 3, 2, 2; 3 VCa spines. Ventral spinulation of 15th leg: 1, 1, 3, 2, 2; 3 tot 4VCa spines...
|
|
<taxonomicName id="B1020FE43327247A3753D9EBD3F64170" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="subSpecies" species="acherontis" subSpecies="wardaranus">
|
|
E.
|
|
<taxonomicName id="C7B23D929C9B3E73ABC6420BE861C223" authority="Verh." authorityName="Verh." class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="subSpecies" species="acherontis" subSpecies="wardaranus">
|
|
acherontis ssp. wardaranus (
|
|
<authority id="491E5F70CC8CF406889DB9D35363B24F" pageId="6" pageNumber="171">Verh.</authority>
|
|
)
|
|
</taxonomicName>
|
|
</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="3BBCF4FADF8ED9495BB7982A030A67C1" pageId="6" pageNumber="171" type="mainText">
|
|
- 19 ocelli. 83 antennomeres. 10 + 11 coxosternal teeth. Ventral spinulation of 15th leg: 1, 1, 3, 2, 1; 2 VCa spines...
|
|
<taxonomicName id="AEF8C391890E7A54D953CB4D2EAECD8C" authority="Folkm." authorityName="Folkm." class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="species" species="stygis">
|
|
E. stygis (
|
|
<authority id="1BDE26148A15B516117EFFF8EC993C96" pageId="6" pageNumber="171">Folkm.</authority>
|
|
)
|
|
</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="8581ABD8A01374D219C1206DBC2EC641" pageId="6" pageNumber="171" type="mainText">
|
|
5. 18 to 20 ocelli. 61 to 76 antennomeres. 7 + 7 to 8 + 8 coxosternal teeth. Ventral spinulation of 15th leg: 1, 1, 3, 3, 2; 1 or 2VCa spines...
|
|
<taxonomicName id="D239A807F4A6803AC416EC682D99C23E" authority="Silv." authorityName="Silv." class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="species" species="excellens">
|
|
E. excellens (
|
|
<authority id="168F925D576E4B333C23AAEE0FFDB07B" pageId="6" pageNumber="171">Silv.</authority>
|
|
)
|
|
</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="CAFF5A52CADFFD3AA3B43706CF2DC042" pageId="6" pageNumber="171" type="mainText">
|
|
- 20 ocelli. 65 to 67 antennomeres. 9 + 9 coxosternal teeth. Ventral spinulation of 15th leg: 1, 1, 3, 2, 1-2; 1 or 2 VCa spines...
|
|
<taxonomicName id="25730DA02D141D6F077EE853E9E68D71" authority="Verh." authorityName="Verh." class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="species" species="tabellamm">
|
|
E. tabellamm (
|
|
<authority id="0F86733005B4954345201895A988E4DA" pageId="6" pageNumber="171">Verh.</authority>
|
|
)
|
|
</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="728C7B6CC1C7FAC8BB70470B845CA0CC" pageId="6" pageNumber="171" type="mainText">6. 38 to 50 antennomeres. 15th leg with 1 VCa spine, in the male with a dorsal prefemoral furrow... 7</paragraph>
|
|
<paragraph id="D259A6F3E6A453AADE041AB7D357EEB2" pageId="6" pageNumber="171" type="mainText">- 50 to 58 antennomeres. 15th leg with 2 to 4 VCa spines, in the male with a large prefemoral knob... 8</paragraph>
|
|
<paragraph id="6FD5C4B604CFC1C80503C48FC4668EE9" pageId="6" pageNumber="171" type="mainText">
|
|
7. 19 to 20 ocelli. 41 to 50 antennomeres. 7 + 7 coxosternal teeth. Ventral spinulation of 15th leg: 1, 1, 3, 2, 1...
|
|
<taxonomicName id="6C4AA86E73EACA72123C8288B23ADFA8" authority="Verh." authorityName="Verh." class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="species" species="zeus">
|
|
E. zeus (
|
|
<authority id="A67DBA8320A52949C47D448A4A99C41A" pageId="6" pageNumber="171">Verh.</authority>
|
|
)
|
|
</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="4A86824FF5AE5FC8D22DB4462C33B890" pageId="6" pageNumber="171" type="mainText">
|
|
- 38 antennomeres. 8 + 8 coxosternal teeth. Ventral spinulation of 15th leg: 1, 1, 3, 2, 2...
|
|
<taxonomicName id="4D728BC8A8AC726F243120F5E4FFA4E6" authority="Kanelis" authorityName="Kanelis" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="species" species="sissii">
|
|
E. sissii (
|
|
<authority id="4DA34989F438848CCA100FF17A173DF8" pageId="6" pageNumber="171">Kanelis</authority>
|
|
)
|
|
</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="90F9D44B87DF43B28057733DE216074E" pageId="6" pageNumber="171" type="mainText">
|
|
8. 16 to 19 ocelli. 50 to 56 antennomeres. 7 + 7 to 11 + 11 coxosternal teeth. Ventral spinulation of 15th leg: 1, 1, 4, 2, 0-1; 3 VCa spines...
|
|
<taxonomicName id="19B8A8B1372253CDE24CC6EA1B0245E4" authority="Latz." authorityName="Latz." class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="species" species="spinirger">
|
|
E. spinirger (
|
|
<authority id="0B3CC48730832CDB9FD89663C4B5750F" pageId="6" pageNumber="171">Latz.</authority>
|
|
)
|
|
</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="D768C45F458A1B4E50C12A9A68C90E97" pageId="6" pageNumber="171" type="mainText">
|
|
- 13 ocelli. 54-57 antennomeres: 6 + 6 to 8 + 9 coxosternal teeth. Ventral spinulation of 15th leg: 1, 1, 3, 2, 0; 2 or 3 VCa spines...
|
|
<taxonomicName id="EDDFE28E8CDA5407BE21621ED909B6E2" authority="Verh." authorityName="Verh." class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="species" species="caesar">
|
|
E. caesar (
|
|
<authority id="E5FAF896E7424D06BB20F4114D6A1E52" pageId="6" pageNumber="171">Verh.</authority>
|
|
)
|
|
</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="5C8EE3E930D5CBABE5F79F012A8028BD" pageId="6" pageNumber="171" type="mainText">
|
|
- 19 ocelli. About 50 antennomeres. 8 + 8 coxosternal teeth. Ventral spinulation of 15th leg: 1, 1. 3, 2, 1; 3 VCa spines...
|
|
<taxonomicName id="FE68EF1E270659A7EA3704A999802C13" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="subSpecies" species="caesar" subSpecies="valonensis">
|
|
E.
|
|
<taxonomicName id="7895207A4CC611911D4A33945ABE571B" authority="Verh." authorityName="Verh." class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="subSpecies" species="caesar" subSpecies="valonensis">
|
|
caesar ssp. valonensis (
|
|
<authority id="47708EED55ABF4198E471FA67BB4915E" pageId="6" pageNumber="171">Verh.</authority>
|
|
)
|
|
</taxonomicName>
|
|
</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="9D6B1E62E11236E1E2018A82D98D6535" pageId="6" pageNumber="171" type="mainText">
|
|
- 23 ocelli. 58 antennomeres. 9 + 9 coxosternal teeth. Ventral spinulation of 15th leg: 1, 1, 3, 2, 2; 2 or 3 VCa spines...
|
|
<taxonomicName id="DAA944858E513E0094571CDC9930C4F7" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="subSpecies" species="caesar" subSpecies="patens">
|
|
E.
|
|
<taxonomicName id="4E4BB6DDB2E1311E77D2B1AFFDD3BC4D" authority="Att." authorityName="Att." class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="subSpecies" species="caesar" subSpecies="patens">
|
|
caesar ssp. patens (
|
|
<authority id="3A9423573B975D3F8C81144E470F5EAB" pageId="6" pageNumber="171">Att.</authority>
|
|
)
|
|
</taxonomicName>
|
|
</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="54BF542B81D4D42A8714B2637F7E83EB" lastPageId="7" lastPageNumber="172" pageId="6" pageNumber="171" type="discussion">
|
|
<paragraph id="39AB60271DCAFA77A14902CE64D2240D" pageId="6" pageNumber="171" type="mainText">
|
|
In connection with the species of
|
|
<taxonomicName id="9039C869FEB4F5458C8CA268D27B631D" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="subGenus" subGenus="Schizopolybothrus">Schizopolybothrus</taxonomicName>
|
|
the following remarks must be made.
|
|
</paragraph>
|
|
<paragraph id="D279A46A521C288B9FEC67A657CD9E39" lastPageId="7" lastPageNumber="172" pageId="6" pageNumber="171" type="mainText">
|
|
<bibRef id="F69E3D7D032437159A44EE2BE2A70ABF" pageId="6" pageNumber="171">Verhoeff (1943)</bibRef>
|
|
referred leostygis to his subgenus
|
|
<taxonomicName id="70A7BAABDACF352191B45B32A7A0B89B" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="subGenus" subGenus="Mesobothrus">Mesobothrus</taxonomicName>
|
|
, thus plying that the species has long gonopods. As this is contradictory to his own and to
|
|
<bibRef id="45DDBDF18A54FEA1E9595BE3B0BE08BA" pageId="6" pageNumber="171">Attems's (1935)</bibRef>
|
|
descriptions, and as leostygis through other characters
|
|
<pageBreakToken id="2B0D13EB286DE22366489E88BBA4E8A7" pageId="7" pageNumber="172" start="start">shows</pageBreakToken>
|
|
strong affinities to
|
|
<taxonomicName id="E682D6D7EAB31159BCD047AC50526444" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="species" species="acherontis">acherontis</taxonomicName>
|
|
and
|
|
<taxonomicName id="6ECBA60523F788462EFF2417395D5EA7" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="species" species="caesar">caesar</taxonomicName>
|
|
, we may safely assume that this was an error.
|
|
<bibRef id="7F41D2ABD0A25C14156998B0C6B78324" pageId="7" pageNumber="172">Verhoeff (1943)</bibRef>
|
|
distinguished a subspecies stygisleo of
|
|
<taxonomicName id="9B53A3113A19A2866B34914783EFBF72" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="6" pageNumber="171" phylum="Arthropoda" rank="species" species="leostygis">E. leostygis</taxonomicName>
|
|
which was characterized by having 2 ocelli (instead of 4 to 5) and by having the ten distal antennomeres 1 1/3 to 1 1/2 times longer than broad and the last antennomere 1 3/4 times longer than broad. Considering the instability of this kind of characters in the
|
|
<taxonomicName id="B9900D367A236314B66CEF424AFDD7F6" family="Lithobiidae" pageId="6" pageNumber="171" rank="family">Lithobiidae</taxonomicName>
|
|
this subspecies is better withdrawn. The characters in the above key were derived from the descriptions and notes on leostygis and stygisleo by
|
|
<bibRef id="0F8089C2E75842AAE5933923DC8DE736" pageId="7" pageNumber="172">Verhoeff (1899</bibRef>
|
|
,
|
|
<bibRef id="B735E06EB38DDB07FB7F8C0666D7D969" pageId="7" pageNumber="172">1900</bibRef>
|
|
,
|
|
<bibRef id="27A70EC94549EEC7A2700FF1A3ED3534" pageId="7" pageNumber="172">1943</bibRef>
|
|
) and
|
|
<bibRef id="B8D6AA5B153EEB09FF7870117899C283" pageId="7" pageNumber="172">Attems (1935)</bibRef>
|
|
. According to Attems the species has up to 10 ocelli.
|
|
</paragraph>
|
|
<paragraph id="ACA75F3B18C267C15F1717DB3CB9E089" pageId="7" pageNumber="172" type="mainText">
|
|
The record of leostygis by
|
|
<bibRef id="75906BF6024E97A7942C360A16219BE4" pageId="7" pageNumber="172">Attems (1908)</bibRef>
|
|
was based on a misidentification and refers to
|
|
<taxonomicName id="33FE3BEEE107D8CF221DC68C0F01D5E8" authority="Verhoeff, 1930" authorityName="Verhoeff" authorityYear="1930" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="7" pageNumber="172" phylum="Arthropoda" rank="species" species="obrovensis">
|
|
E. obrovensis (
|
|
<authority id="A386810BE83ED6E28B46244BD0CE0E06" pageId="7" pageNumber="172">Verhoeff, 1930</authority>
|
|
)
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="F0D86DDF6803EFFA5CB3E3DA2CAE9E62" pageId="7" pageNumber="172" type="mainText">
|
|
It is not clear why
|
|
<bibRef id="3173F11F3FADCD2BD014F808DA6E7C80" pageId="7" pageNumber="172">Attems (1935)</bibRef>
|
|
referred his
|
|
<taxonomicName id="7624139053745FA11CFA615C16F3CF24" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="7" pageNumber="172" phylum="Arthropoda" rank="subSpecies" species="patens" subSpecies="patens">subspecies patens</taxonomicName>
|
|
to
|
|
<taxonomicName id="B13ED94BCE07B36ED25AE96A0B35EA48" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="7" pageNumber="172" phylum="Arthropoda" rank="species" species="leostygis">leostygis</taxonomicName>
|
|
. In view of the high number of ocelli and the considerably lower number of antennomeres, this form has comparatively little to do with leostygis and is better associated with
|
|
<taxonomicName id="E9A4476E372448F8D150AB1724BF823E" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="7" pageNumber="172" phylum="Arthropoda" rank="species" species="caesar">caesar</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="73896F27A83BE60CC22868B8C8AB26F5" pageId="7" pageNumber="172" type="mainText">
|
|
<taxonomicName id="27FF9F30A8120F911CA8F11B0A2E585F" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="7" pageNumber="172" phylum="Arthropoda" rank="species" species="excellens">E. excellens</taxonomicName>
|
|
seems very closely related to
|
|
<taxonomicName id="D4441CA376699CE41CA614734D9EC500" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="7" pageNumber="172" phylum="Arthropoda" rank="species" species="tabularum">tabularum</taxonomicName>
|
|
, but unfortunately the discovery of the male has to be awaited before a more definite statement can be made. Moreover,
|
|
<taxonomicName id="59D4043AD0AB15D53BD7EA250100839C" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="7" pageNumber="172" phylum="Arthropoda" rank="species" species="tabularum">tabularum</taxonomicName>
|
|
is characterized by a dense brush of setae covering partly the coxosternal teeth. This brush is lacking completely in
|
|
<taxonomicName id="384134EF7898CC4D09EB687CD3185909" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="7" pageNumber="172" phylum="Arthropoda" rank="species" species="excellens">excellens</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |