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<document id="E157AF765B287A7056857456D77E489C" ID-CLB-Dataset="9266" ID-DOI="10.11646/zootaxa.4895.2.7" ID-GBIF-Dataset="33852167-3335-4c92-b513-5f3f4b83c23f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4322575" ID-ZooBank="53E32A6A-00EE-47C3-9761-968DAA73BC4F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1608027041796" checkinUser="plazi" docAuthor="Hendrich, Lars &amp; Balke, Michael" docDate="2020" docId="03F187FE065415703AFCFDBA2EB21A6C" docLanguage="en" docName="zootaxa.4895.2.7.pdf" docOrigin="Zootaxa 4895 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coptotomus balticus Hendrich &amp; Balke 2020, sp. n." docType="treatment" docVersion="8" lastPageNumber="288" masterDocId="FFC8FF86065515733A6BFF6D2D051B2D" masterDocTitle="A Baltic amber species of the diving beetle genus Coptotomus Say, 1830 (Coleoptera Dytiscidae: Coptotominae)" masterLastPageNumber="290" masterPageNumber="285" pageNumber="286" updateTime="1698847593838" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="6315C5656ED8751AE0CD1027DCBAC17E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="993B02E3ACF914028D712081ED762708">
<mods:title id="F761D6039E69F30D34CF53BA32874117">A Baltic amber species of the diving beetle genus Coptotomus Say, 1830 (Coleoptera Dytiscidae: Coptotominae)</mods:title>
</mods:titleInfo>
<mods:name id="EC6195EABD10BAAF1E8476FF4A5C42B6" type="personal">
<mods:role id="4CF97E9E18FE98098053AF301BAB2C9C">
<mods:roleTerm id="315C85560760B10830FA0404BBC35282">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BA8E3B53A61DCF5946789187C6DFD953">Hendrich, Lars</mods:namePart>
<mods:affiliation id="4397F380C03E45391A58565E1E4E1057">SNSB-Zoologische Staatssammlung, Mu ̈ nchhausenstrasse 21, D- 81247 Mu ̈ nchen, Germany</mods:affiliation>
</mods:name>
<mods:name id="39C6E55C486E9F7F19F63F6AFD58DBDC" type="personal">
<mods:role id="36A215F43036106CDBEAC3B4BB2D0404">
<mods:roleTerm id="FD4365E8DBCF9F132EF51DBE5D391EAB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AC2A41C2D39E0F6E3A32F1A252C0F79E">Balke, Michael</mods:namePart>
<mods:affiliation id="CBD6181F9DB9276E5158F9944F94CE8B">SNSB-Zoologische Staatssammlung, Mu ̈ nchhausenstrasse 21, D- 81247 Mu ̈ nchen, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource id="E1ABE76FDB0C6D99D35124BD41407E61">text</mods:typeOfResource>
<mods:relatedItem id="D5EBEB46DCFCBE0F447E6E7CC706A61C" type="host">
<mods:titleInfo id="CAD4AF8CE99B1AFBD95E967D6A833713">
<mods:title id="FA2696034DCEEDCED176CC4113DF6160">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CDA028EE2DFF86CE55B35302D9CFB2F0">
<mods:date id="AE386BCEBF2D08CF08A21D045AE26245">2020</mods:date>
<mods:detail id="F521B98D337A17271A305FC942C72A14" type="pubDate">
<mods:number id="00E1A57E2E968C327BD72CBB7B9CCF31">2020-12-15</mods:number>
</mods:detail>
<mods:detail id="B003C309079436F0C56D2C3AD0676674" type="volume">
<mods:number id="6A2F2DFD77101E9775EAB31D7348B49B">4895</mods:number>
</mods:detail>
<mods:detail id="6FBB2BCA91B613277EE98D24EF158E4A" type="issue">
<mods:number id="5EE94423188A625574B400E3894FF2A0">2</mods:number>
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<mods:start id="DC3182AFD49114806A89D17BD253FF9C">285</mods:start>
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</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="605F1F681D5131AF8C222215D6D6C8D1">journal article</mods:classification>
<mods:identifier id="3B87CFCE38CEA8B2931FA16160698356" type="CLB-Dataset">9266</mods:identifier>
<mods:identifier id="0605726193F61987B0012F1B6A723A00" type="DOI">10.11646/zootaxa.4895.2.7</mods:identifier>
<mods:identifier id="84C0EDC10E41CC0DDD46A7416C851416" type="GBIF-Dataset">33852167-3335-4c92-b513-5f3f4b83c23f</mods:identifier>
<mods:identifier id="B63116611357EA08843DF90800A1351B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BF7B9603C22585B4BCD13424EAE231F7" type="Zenodo-Dep">4322575</mods:identifier>
<mods:identifier id="A6E1337203F98BA11FD0F1A70C2402E9" type="ZooBank">53E32A6A-00EE-47C3-9761-968DAA73BC4F</mods:identifier>
</mods:mods>
<treatment id="03F187FE065415703AFCFDBA2EB21A6C" ID-DOI="http://doi.org/10.5281/zenodo.4382793" ID-GBIF-Taxon="176230325" ID-Zenodo-Dep="4382793" LSID="urn:lsid:plazi:treatment:03F187FE065415703AFCFDBA2EB21A6C" httpUri="http://treatment.plazi.org/id/03F187FE065415703AFCFDBA2EB21A6C" lastPageId="3" lastPageNumber="288" pageId="1" pageNumber="286">
<subSubSection id="C3426563065415723AFCFDBA2CEB19DC" box="[151,494,727,754]" pageId="1" pageNumber="286" type="nomenclature">
<paragraph id="8BE736E8065415723AFCFDBA2CEB19DC" blockId="1.[151,494,727,789]" box="[151,494,727,754]" pageId="1" pageNumber="286">
<heading id="D0AF8184065415723AFCFDBA2CEB19DC" bold="true" box="[151,494,727,754]" fontSize="11" level="1" pageId="1" pageNumber="286" reason="1">
<emphasis id="B92CEAFA065415723AFCFDBA2CEB19DC" bold="true" box="[151,494,727,754]" pageId="1" pageNumber="286">
†
<taxonomicName id="4C584D6B065415723AC6FDBA2CA019DF" authority="Hendrich &amp; Balke, 2020" authorityName="Hendrich &amp; Balke" authorityYear="2020" box="[173,421,727,754]" class="Insecta" family="Dytiscidae" genus="Coptotomus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="286" phylum="Arthropoda" rank="species" species="balticus" status="sp. nov.">
<emphasis id="B92CEAFA065415723AC6FDBA2CA019DF" bold="true" box="[173,421,727,754]" italics="true" pageId="1" pageNumber="286">Coptotomus balticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F5781065415723BC0FDBA2CEB19DC" box="[427,494,727,753]" pageId="1" pageNumber="286" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3426563065415723AFCFD962C091838" box="[151,268,763,789]" pageId="1" pageNumber="286" type="description">
<paragraph id="8BE736E8065415723AFCFD962C091838" blockId="1.[151,494,727,789]" box="[151,268,763,789]" pageId="1" pageNumber="286">
(
<figureCitation id="13632A6D065415723AF4FD962DE31838" box="[159,230,763,789]" captionStart="FIGURE 1" captionStartId="2.[151,250,1938,1962]" captionTargetBox="[151,1435,306,1912]" captionTargetId="figure-92@2.[151,1436,306,1912]" captionTargetPageId="2" captionText="FIGURE 1. † Coptotomus balticus sp. n., male, holotype: A) habitus (dorsal side); B) habitus ventral side, scale bar = 1 mm; C) Pro- and mesotarsus, and apical palpomere of both the maxillary and labial palps distinctly bifid (red arrows); D) Metatarsomeres 1 to 4 with externoapical angles lobed." figureDoi="http://doi.org/10.5281/zenodo.4322577" httpUri="https://zenodo.org/record/4322577/files/figure.png" pageId="1" pageNumber="286">Figs 1</figureCitation>
,
<figureCitation id="13632A6D065415723A9FFD962C061838" box="[244,259,763,789]" captionStart="FIGURE 2" captionStartId="3.[151,250,1934,1958]" captionTargetBox="[164,1423,346,1911]" captionTargetId="figure-95@3.[164,1423,345,1911]" captionTargetPageId="3" captionText="FIGURE 2. † Coptotomus balticus sp. n., female, paratype: A) habitus (dorsal side, total length = 5.9 mm); B) habitus ventral side; C) elytron with strioles (scale bar = 1 mm); D) ventral side with pronotal process and apical palpomere of both the maxillary and labial palps bifid (scale bar = 1 mm); E) metasternum and metaventrite (scale bar = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.4322579" httpUri="https://zenodo.org/record/4322579/files/figure.png" pageId="1" pageNumber="286">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3426563065415723AFCFC2E29F81FE8" pageId="1" pageNumber="286" type="materials_examined">
<paragraph id="8BE736E8065415723AFCFC2E2E291870" blockId="1.[151,1437,835,2014]" box="[151,812,835,862]" pageId="1" pageNumber="286">
<emphasis id="B92CEAFA065415723AFCFC2E2C331870" bold="true" box="[151,310,835,861]" pageId="1" pageNumber="286">Type locality.</emphasis>
<collectingCountry id="F34F7678065415723B57FC2E2C8C1870" box="[316,393,835,861]" name="Russia" pageId="1" pageNumber="286">Russia</collectingCountry>
,
<collectingRegion id="499CF80A065415723BFEFC2E2F731870" box="[405,630,835,861]" country="Russia" name="Kaliningrad" pageId="1" pageNumber="286">Kaliningrad Region</collectingRegion>
, Yantarny mine.
</paragraph>
<paragraph id="8BE736E8065415723AACFC0A2C6F188B" blockId="1.[151,1437,835,2014]" pageId="1" pageNumber="286">
<emphasis id="B92CEAFA065415723AACFC0A2DF818AC" bold="true" box="[199,253,871,897]" pageId="1" pageNumber="286">Age.</emphasis>
According to
<bibRefCitation id="EFC94B19065415723BF7FC0A2F7518AF" author="Wolfe, A. P. &amp; McKellar, R. C. &amp; Tappert, R. &amp; Sodhi, R. N. &amp; Muehlenbachs, K." box="[412,624,871,898]" pageId="1" pageNumber="286" pagination="21 - 32" refId="ref2768" refString="Wolfe, A. P., McKellar, R. C., Tappert, R., Sodhi, R. N. &amp; Muehlenbachs, K. (2016) Bitterfeld amber is not Baltic amber: Three geochemical tests and further constraints on the botanical affinities of succinate. Review of Palaeobotany and Palynology, 225, 21 - 32. https: // doi. org / 10.1016 / j. revpalbo. 2015.11.002" type="journal article" year="2016">
Wolfe
<emphasis id="B92CEAFA065415723B8CFC052F1D18AC" box="[487,536,871,897]" italics="true" pageId="1" pageNumber="286">et al</emphasis>
. (2016)
</bibRefCitation>
Baltic amber is thought to be of middle Eocene age (Lutetian: 41.3–47.8 million years ago).
</paragraph>
<paragraph id="8BE736E8065415723AACFCC22CBC1F3C" blockId="1.[151,1437,835,2014]" pageId="1" pageNumber="286">
<materialsCitation id="3B303CB5065415723AACFCC22CBC1F3C" ID-GBIF-Occurrence="3000963303" collectionCode="ZSM" country="Russia" determinerName="Balke" location="Kaliningrad Region" pageId="1" pageNumber="286" specimenCount="2" stateProvince="Kaliningrad" typeStatus="holotype">
<emphasis id="B92CEAFA065415723AACFCC22C3118E4" bold="true" box="[199,308,943,969]" pageId="1" pageNumber="286">
<typeStatus id="54E3884A065415723AACFCC22C3118E4" box="[199,308,943,969]" pageId="1" pageNumber="286" type="holotype">Holotype</typeStatus>
</emphasis>
(male). In a plastic box in the main collection of the Zoologische Staatssammlung München (
<collectionCode id="ED49AE2D065415723F33FCC2289018E4" box="[1368,1429,943,969]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="286" type="Museum">ZSM</collectionCode>
) with a printed label: “
<collectingCountry id="F34F7678065415723BE2FCBE2CDC18C0" box="[393,473,979,1005]" name="Russia" pageId="1" pageNumber="286">Russia</collectingCountry>
,
<collectingRegion id="499CF80A065415723B8EFCBE2FC018C0" box="[485,709,979,1005]" country="Russia" name="Kaliningrad" pageId="1" pageNumber="286">Kaliningrad Region</collectingRegion>
, Yantarny mine.
<typeStatus id="54E3884A0654157239E9FCBE291318C0" box="[898,1046,979,1005]" pageId="1" pageNumber="286" type="holotype">HOLOTYPE</typeStatus>
<emphasis id="B92CEAFA065415723E76FCBE285B18C0" bold="true" box="[1053,1374,979,1006]" pageId="1" pageNumber="286">
†
<taxonomicName id="4C584D6B065415723E40FCBE281218C3" authority="Hendrich &amp; Balke, 2020" authorityName="Hendrich &amp; Balke" authorityYear="2020" box="[1067,1303,979,1006]" class="Insecta" family="Dytiscidae" genus="Coptotomus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="286" phylum="Arthropoda" rank="species" species="balticus" status="sp. nov.">
<emphasis id="B92CEAFA065415723E40FCBE281218C3" bold="true" box="[1067,1303,979,1006]" italics="true" pageId="1" pageNumber="286">Coptotomus balticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F5781065415723F75FCBE285B18C0" box="[1310,1374,979,1005]" pageId="1" pageNumber="286" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Hendrich &amp;
<determinerName id="7E3BDA2C065415723A9EFC9A2C331F3C" box="[245,310,1015,1041]" pageId="1" pageNumber="286">Balke</determinerName>
det. 2020”.
</materialsCitation>
</paragraph>
<paragraph id="8BE736E8065415723AACFB71280F1F74" blockId="1.[151,1437,835,2014]" pageId="1" pageNumber="286">
Two labelled pictures were provided with the specimen: “Certificate 4774 Natural Baltic Amber with Inclusions expert Jonas Damzen International Amber Association Names of Inclusions:
<taxonomicName id="4C584D6B06541572399DFB5229711F74" authorityName="Linnaeus" authorityYear="1758" box="[1014,1140,1087,1113]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="1" pageNumber="286" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
,
<taxonomicName id="4C584D6B065415723E15FB5229F91F74" box="[1150,1276,1087,1113]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="286" phylum="Arthropoda" rank="family">Dytiscidae</taxonomicName>
”.
</paragraph>
<paragraph id="8BE736E8065415723AACFB0E29F81FE8" blockId="1.[151,1437,835,2014]" pageId="1" pageNumber="286">
<materialsCitation id="3B303CB5065415723AACFB0E29FC1FE8" ID-GBIF-Occurrence="3000963301" collectionCode="GPIH" collectorName="Gr" country="Russia" county="In" determinerName="Balke" location="University Hamburg" municipality="Geologisch Palaeontologogisches Institut" pageId="1" pageNumber="286" specimenCount="2" stateProvince="Kaliningrad" typeStatus="paratype">
<emphasis id="B92CEAFA065415723AACFB0E2C311F50" bold="true" box="[199,308,1123,1149]" pageId="1" pageNumber="286">
<typeStatus id="54E3884A065415723AACFB0E2C311F50" box="[199,308,1123,1149]" pageId="1" pageNumber="286" type="paratype">Paratype</typeStatus>
</emphasis>
(female).
<collectingCounty id="62864E64065415723BCDFB092CB81F53" box="[422,445,1124,1150]" pageId="1" pageNumber="286">In</collectingCounty>
a plastic box in the
<collectingMunicipality id="6B83AC920654157238F3FB0E296D1F50" box="[664,1128,1123,1149]" pageId="1" pageNumber="286">Geologisch Palaeontologogisches Institut</collectingMunicipality>
,
<location id="8E876033065415723E19FB0E285F1F50" LSID="urn:lsid:plazi:treatment:03F187FE065415703AFCFDBA2EB21A6C:8E876033065415723E19FB0E285F1F50" box="[1138,1370,1123,1149]" country="Russia" county="In" municipality="Geologisch Palaeontologogisches Institut" name="University Hamburg" pageId="1" pageNumber="286" stateProvince="Kaliningrad">University Hamburg</location>
, now:
<location id="8E876033065415723AFCFBEA2DE01F8C" LSID="urn:lsid:plazi:treatment:03F187FE065415703AFCFDBA2EB21A6C:8E876033065415723AFCFBEA2DE01F8C" box="[151,229,1159,1185]" country="Russia" county="In" municipality="Geologisch Palaeontologogisches Institut" name="CeNak" pageId="1" pageNumber="286" stateProvince="Kaliningrad">CeNak</location>
(
<location id="8E876033065415723A9FFBEA2F341F8F" LSID="urn:lsid:plazi:treatment:03F187FE065415703AFCFDBA2EB21A6C:8E876033065415723A9FFBEA2F341F8F" box="[244,561,1159,1186]" country="Russia" county="In" municipality="Geologisch Palaeontologogisches Institut" name="Centrum of Natural History" pageId="1" pageNumber="286" stateProvince="Kaliningrad">Centrum of Natural History</location>
), with a printed label: “
<collectingCountry id="F34F767806541572395CFBEA2E821F8C" box="[823,903,1159,1185]" name="Russia" pageId="1" pageNumber="286">Russia</collectingCountry>
,
<collectingRegion id="499CF80A0654157239F8FBEA29711F8C" box="[915,1140,1159,1185]" country="Russia" name="Kaliningrad" pageId="1" pageNumber="286">Kaliningrad Region</collectingRegion>
, Yantarny mine.
<collectionCode id="ED49AE2D065415723F5AFBEA28771F8C" box="[1329,1394,1159,1185]" country="Germany" httpUri="http://grbio.org/cool/8qxf-pkip" name="Geologisch-Palaeontologiches Institut der Universitt Haemburg" pageId="1" pageNumber="286">GPIH</collectionCode>
no. 5040, coll.
<collectorName id="26AD533E065415723B7EFBC62C301FE8" box="[277,309,1195,1221]" pageId="1" pageNumber="286">Gr</collectorName>
̂hn no. 8188.
<typeStatus id="54E3884A065415723BBFFBC62F5A1FE8" box="[468,607,1195,1221]" pageId="1" pageNumber="286" type="paratype">PARATYPE</typeStatus>
<emphasis id="B92CEAFA06541572380DFBC62EA31FE8" bold="true" box="[614,934,1195,1222]" pageId="1" pageNumber="286">
†
<taxonomicName id="4C584D6B06541572381FFBC62E5A1FEB" authority="Hendrich &amp; Balke, 2020" authorityName="Hendrich &amp; Balke" authorityYear="2020" box="[628,863,1195,1222]" class="Insecta" family="Dytiscidae" genus="Coptotomus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="286" phylum="Arthropoda" rank="species" species="balticus" status="sp. nov.">
<emphasis id="B92CEAFA06541572381FFBC62E5A1FEB" bold="true" box="[628,863,1195,1222]" italics="true" pageId="1" pageNumber="286">Coptotomus balticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F578106541572390DFBC62EA31FE8" box="[870,934,1195,1221]" pageId="1" pageNumber="286" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Hendrich &amp;
<determinerName id="7E3BDA2C065415723E53FBC6297C1FE8" box="[1080,1145,1195,1221]" pageId="1" pageNumber="286">Balke</determinerName>
det. 2020”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3426563065415723AACFBA22E791DCC" pageId="1" pageNumber="286" type="description">
<paragraph id="8BE736E8065415723AACFBA22E811E23" blockId="1.[151,1437,835,2014]" pageId="1" pageNumber="286">
<emphasis id="B92CEAFA065415723AACFBA22C5C1FC4" bold="true" box="[199,345,1231,1257]" pageId="1" pageNumber="286">Description.</emphasis>
Beetle elongate-oval, somewhat pointed posteriorly (
<figureCitation id="13632A6D0654157239D1FBA229101FC4" box="[954,1045,1231,1257]" captionStart="FIGURE 1" captionStartId="2.[151,250,1938,1962]" captionTargetBox="[151,1435,306,1912]" captionTargetId="figure-92@2.[151,1436,306,1912]" captionTargetPageId="2" captionText="FIGURE 1. † Coptotomus balticus sp. n., male, holotype: A) habitus (dorsal side); B) habitus ventral side, scale bar = 1 mm; C) Pro- and mesotarsus, and apical palpomere of both the maxillary and labial palps distinctly bifid (red arrows); D) Metatarsomeres 1 to 4 with externoapical angles lobed." figureDoi="http://doi.org/10.5281/zenodo.4322577" httpUri="https://zenodo.org/record/4322577/files/figure.png" pageId="1" pageNumber="286">Fig. 1A</figureCitation>
); length of beetle with head about
<quantity id="4CA09B0D065415723AFCFB9E2DEA1E23" box="[151,239,1267,1294]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="1" pageNumber="286" unit="mm" value="5.2">5.2 mm</quantity>
, without head about
<quantity id="4CA09B0D065415723BB3FB9E2F341E23" box="[472,561,1267,1294]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="286" unit="mm" value="4.8">4.8 mm</quantity>
; greatest width about
<quantity id="4CA09B0D06541572394CFB9E2E851E23" box="[807,896,1267,1294]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="286" unit="mm" value="2.8">2.8 mm</quantity>
.
</paragraph>
<paragraph id="8BE736E8065415723AACFA7A2F021E54" blockId="1.[151,1437,835,2014]" pageId="1" pageNumber="286">
<emphasis id="B92CEAFA065415723AACFA7A2CA51E1C" bold="true" box="[199,416,1303,1329]" pageId="1" pageNumber="286">Surface sculpture.</emphasis>
Head with fairly small punctation. Pronotum, elytron and ventral side with dense and larger punctation. Profemur and tibia dorsally and ventrally with coarse punctation. Mesofemur and tibia dorsally and ventrally with coarse punctation.
</paragraph>
<paragraph id="8BE736E8065415723AACFAEE28061DB4" blockId="1.[151,1437,835,2014]" pageId="1" pageNumber="286">
<emphasis id="B92CEAFA065415723AACFAEE2C491EB0" bold="true" box="[199,332,1411,1437]" pageId="1" pageNumber="286">Structures.</emphasis>
Head with anterolateral foveae elongate but widely separated. Antennae filiform and slender. Apical palpomere of both the maxillary and labial palps distinctly bifid (
<figureCitation id="13632A6D0654157239C3FACA29041EEC" box="[936,1025,1447,1473]" captionStart="FIGURE 1" captionStartId="2.[151,250,1938,1962]" captionTargetBox="[151,1435,306,1912]" captionTargetId="figure-92@2.[151,1436,306,1912]" captionTargetPageId="2" captionText="FIGURE 1. † Coptotomus balticus sp. n., male, holotype: A) habitus (dorsal side); B) habitus ventral side, scale bar = 1 mm; C) Pro- and mesotarsus, and apical palpomere of both the maxillary and labial palps distinctly bifid (red arrows); D) Metatarsomeres 1 to 4 with externoapical angles lobed." figureDoi="http://doi.org/10.5281/zenodo.4322577" httpUri="https://zenodo.org/record/4322577/files/figure.png" pageId="1" pageNumber="286">Fig. 1C</figureCitation>
). Pronotum with lateral rim. Scutellum visible. Metacoxal lobes large and rounded with metacoxal lines not closely approximated; prosternum and prosternal process in same plane and pro- and mesotarsi distinctly pentamerous. Prosternal process elongate and acutely pointed apically, received into V-shaped impression on metasternum. Metaventrite comparably broad laterally, width at narrowest point adjacent to mesocoxa
<quantity id="4CA09B0D0654157238B5F95A2E471D7F" box="[734,834,1591,1618]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="286" unit="mm" value="0.35">0.35 mm</quantity>
(
<figureCitation id="13632A6D06541572393AF95A2EA21D7C" box="[849,935,1591,1617]" captionStart="FIGURE 1" captionStartId="2.[151,250,1938,1962]" captionTargetBox="[151,1435,306,1912]" captionTargetId="figure-92@2.[151,1436,306,1912]" captionTargetPageId="2" captionText="FIGURE 1. † Coptotomus balticus sp. n., male, holotype: A) habitus (dorsal side); B) habitus ventral side, scale bar = 1 mm; C) Pro- and mesotarsus, and apical palpomere of both the maxillary and labial palps distinctly bifid (red arrows); D) Metatarsomeres 1 to 4 with externoapical angles lobed." figureDoi="http://doi.org/10.5281/zenodo.4322577" httpUri="https://zenodo.org/record/4322577/files/figure.png" pageId="1" pageNumber="286">Fig. 1B</figureCitation>
). Metatibia and metatarsomeres with natatorial setae. Protarsomeres 1 to 3 narrowly dilated, together bearing four transverse rows of narrow scales on ventral surface. Metatarsomeres 1 to 4 with externoapical angles lobed; metatarsal claws subequal (
<figureCitation id="13632A6D065415723EF7F91229F31DB4" box="[1180,1270,1663,1689]" captionStart="FIGURE 1" captionStartId="2.[151,250,1938,1962]" captionTargetBox="[151,1435,306,1912]" captionTargetId="figure-92@2.[151,1436,306,1912]" captionTargetPageId="2" captionText="FIGURE 1. † Coptotomus balticus sp. n., male, holotype: A) habitus (dorsal side); B) habitus ventral side, scale bar = 1 mm; C) Pro- and mesotarsus, and apical palpomere of both the maxillary and labial palps distinctly bifid (red arrows); D) Metatarsomeres 1 to 4 with externoapical angles lobed." figureDoi="http://doi.org/10.5281/zenodo.4322577" httpUri="https://zenodo.org/record/4322577/files/figure.png" pageId="1" pageNumber="286">Fig. 1D</figureCitation>
).
</paragraph>
<paragraph id="8BE736E8065415723AACF9CE2E791DCC" blockId="1.[151,1437,835,2014]" pageId="1" pageNumber="286">
<emphasis id="B92CEAFA065415723AACF9CE2C201D90" bold="true" box="[199,293,1699,1725]" pageId="1" pageNumber="286">Female.</emphasis>
Length of beetle with head about
<quantity id="4CA09B0D0654157238C2F9CE2E071D93" box="[681,770,1699,1726]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="1" pageNumber="286" unit="mm" value="5.9">5.9 mm</quantity>
, without head about
<quantity id="4CA09B0D065415723986F9CE29421D93" box="[1005,1095,1699,1726]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="1" pageNumber="286" unit="mm" value="5.6">5.6 mm</quantity>
; greatest width about
<quantity id="4CA09B0D065415723F54F9CE289C1D93" box="[1343,1433,1699,1726]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="286" unit="mm" value="2.7">2.7 mm</quantity>
. Pronotum and elytron with longer longitudinal striae (
<figureCitation id="13632A6D06541572389FF9AA2E6A1DCC" box="[756,879,1735,1761]" captionStart="FIGURE 2" captionStartId="3.[151,250,1934,1958]" captionTargetBox="[164,1423,346,1911]" captionTargetId="figure-95@3.[164,1423,345,1911]" captionTargetPageId="3" captionText="FIGURE 2. † Coptotomus balticus sp. n., female, paratype: A) habitus (dorsal side, total length = 5.9 mm); B) habitus ventral side; C) elytron with strioles (scale bar = 1 mm); D) ventral side with pronotal process and apical palpomere of both the maxillary and labial palps bifid (scale bar = 1 mm); E) metasternum and metaventrite (scale bar = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.4322579" httpUri="https://zenodo.org/record/4322579/files/figure.png" pageId="1" pageNumber="286">Fig. 2A, C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3426563065415723AACF9862C791C04" pageId="1" pageNumber="286" type="etymology">
<paragraph id="8BE736E8065415723AACF9862C791C04" blockId="1.[151,1437,835,2014]" pageId="1" pageNumber="286">
<emphasis id="B92CEAFA065415723AACF9862C4B1C28" bold="true" box="[199,334,1771,1797]" pageId="1" pageNumber="286">Etymology.</emphasis>
Latin adjective for being from the Baltic Sea (or Baltic amber). The name is an adjective in the nominative singular.
</paragraph>
</subSubSection>
<subSubSection id="C3426563065415723AACF85E2FF41C60" box="[199,753,1843,1870]" pageId="1" pageNumber="286" type="distribution">
<paragraph id="8BE736E8065415723AACF85E2FF41C60" blockId="1.[151,1437,835,2014]" box="[199,753,1843,1870]" pageId="1" pageNumber="286">
<emphasis id="B92CEAFA065415723AACF85E2C641C60" bold="true" box="[199,353,1843,1869]" pageId="1" pageNumber="286">Distribution.</emphasis>
Only known from the
<typeStatus id="54E3884A06541572380BF8592F951C63" box="[608,656,1844,1870]" pageId="1" pageNumber="286">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C3426563065415703AACF83A2EB21A6C" lastPageId="3" lastPageNumber="288" pageId="1" pageNumber="286" type="diagnosis">
<paragraph id="8BE736E8065415723AACF83A291A1C94" blockId="1.[151,1437,835,2014]" pageId="1" pageNumber="286">
<emphasis id="B92CEAFA065415723AACF83A2CD51C5C" bold="true" box="[199,464,1879,1905]" pageId="1" pageNumber="286">Differential diagnosis.</emphasis>
<materialsCitation id="3B303CB5065415723BB3F83A291E1C94" ID-GBIF-Occurrence="3000963302" collectionCode="MC, MV" pageId="1" pageNumber="286" specimenCount="1" typeStatus="holotype">
As the shape of the aedeagus and the dorsal colouration could not be examined, the present differential diagnosis for the
<typeStatus id="54E3884A065415723855F8162FA51CB8" box="[574,672,1915,1941]" pageId="1" pageNumber="286" type="holotype">holotype</typeStatus>
is based on the relation of the width of the metacoxal wing to the metaventral width (
<collectionCode id="ED49AE2D065415723B1AF8F22C9B1C94" box="[369,414,1951,1977]" country="0" name="Museo de Cipolleti" pageId="1" pageNumber="286">MC</collectionCode>
/
<collectionCode id="ED49AE2D065415723BCDF8F22CD11C94" box="[422,468,1951,1977]" country="0" name="University of Montana Museum" pageId="1" pageNumber="286">MV</collectionCode>
) (
<bibRefCitation id="EFC94B19065415723B80F8F22F791C97" author="Larson, D. J." box="[491,636,1951,1978]" pageId="1" pageNumber="286" pagination="245 - 498" refId="ref2357" refString="Larson, D. J. (1975) The predaceous water beetles (Coleoptera: Dytiscidae) of Alberta: systematics, natural history and distribution. Quaestiones Entomologicae, 11, 245 - 498." type="journal article" year="1975">Larson 1975</bibRefCitation>
, p. 445), and the total length of body
</materialsCitation>
.
</paragraph>
<paragraph id="8BE736E8065415713AACF8AE2C711A30" blockId="1.[151,1437,835,2014]" lastBlockId="2.[151,1437,150,285]" lastPageId="2" lastPageNumber="287" pageId="1" pageNumber="286">
Based on the relation of the width of metaventral wing to metacocal width (male: MC/MV = 4.2, female: 4.6) (
<figureCitation id="13632A6D065715713ACBFFFA2DF01B9C" box="[160,245,151,177]" captionStart="FIGURE 1" captionStartId="2.[151,250,1938,1962]" captionTargetBox="[151,1435,306,1912]" captionTargetId="figure-92@2.[151,1436,306,1912]" captionTargetPageId="2" captionText="FIGURE 1. † Coptotomus balticus sp. n., male, holotype: A) habitus (dorsal side); B) habitus ventral side, scale bar = 1 mm; C) Pro- and mesotarsus, and apical palpomere of both the maxillary and labial palps distinctly bifid (red arrows); D) Metatarsomeres 1 to 4 with externoapical angles lobed." figureDoi="http://doi.org/10.5281/zenodo.4322577" httpUri="https://zenodo.org/record/4322577/files/figure.png" pageId="2" pageNumber="287">Fig. 1B</figureCitation>
),
<emphasis id="B92CEAFA065715713B61FFFB2C1D1B9D" bold="true" box="[266,280,150,176]" pageId="2" pageNumber="287">†</emphasis>
<taxonomicName id="4C584D6B065715713B73FFFA2C941B9C" authority="Hendrich &amp; Balke, 2020" authorityName="Hendrich &amp; Balke" authorityYear="2020" box="[280,401,151,177]" class="Insecta" family="Dytiscidae" genus="Coptotomus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="balticus" status="sp. nov.">
<emphasis id="B92CEAFA065715713B73FFFA2C941B9C" box="[280,401,151,177]" italics="true" pageId="2" pageNumber="287">C. balticus</emphasis>
</taxonomicName>
<emphasis id="B92CEAFA065715713BFCFFFA2CD31B9C" bold="true" box="[407,470,151,177]" pageId="2" pageNumber="287">
<taxonomicNameLabel id="A21F5781065715713BFCFFFA2CD31B9C" box="[407,470,151,177]" pageId="2" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is morphologically similar to the extant
<taxonomicName id="4C584D6B0657157139FDFFFA282C1B9C" authority="longulus LeConte, 1852" authorityName="longulus LeConte" authorityYear="1852" box="[918,1321,150,177]" class="Insecta" family="Dytiscidae" genus="Coptotomus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="subSpecies" species="longulus" subSpecies="longulus">
<emphasis id="B92CEAFA0657157139FDFFFA29781B9C" box="[918,1149,151,177]" italics="true" pageId="2" pageNumber="287">C. longulus longulus</emphasis>
LeConte, 1852
</taxonomicName>
(
<figureCitation id="13632A6D065715713F53FFFA288B1B9C" box="[1336,1422,151,177]" captionStart="FIGURE 3" captionStartId="4.[151,250,1214,1238]" captionTargetBox="[152,1435,181,1190]" captionTargetId="figure-22@4.[151,1435,181,1190]" captionTargetPageId="4" captionText="FIGURE 3. Coptotomus longulus lenticus Hilsenhoff, 1980, USA, Ohio, Holmes (scale bar = 1 mm): A) habitus (dorsal side); B) Metasternum and metaventrite of C. longulus longulus (USA, Montana)." figureDoi="http://doi.org/10.5281/zenodo.4322583" httpUri="https://zenodo.org/record/4322583/files/figure.png" pageId="2" pageNumber="287">Fig. 3B</figureCitation>
), but
<emphasis id="B92CEAFA065715713AA8FFD72DD41BF9" bold="true" box="[195,209,186,212]" pageId="2" pageNumber="287">†</emphasis>
<taxonomicName id="4C584D6B065715713AB3FFD62C561BF8" authority="Hendrich &amp; Balke, 2020" authorityName="Hendrich &amp; Balke" authorityYear="2020" box="[216,339,187,213]" class="Insecta" family="Dytiscidae" genus="Coptotomus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="balticus" status="sp. nov.">
<emphasis id="B92CEAFA065715713AB3FFD62C561BF8" box="[216,339,187,213]" italics="true" pageId="2" pageNumber="287">C. balticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F5781065715713B31FFD62C9F1BF8" box="[346,410,187,213]" pageId="2" pageNumber="287" rank="species">
<emphasis id="B92CEAFA065715713B31FFD62C911BF8" bold="true" box="[346,404,187,213]" pageId="2" pageNumber="287">sp. n</emphasis>
.
</taxonomicNameLabel>
is much smaller (TL =
<quantity id="4CA09B0D0657157138CDFFD72E2B1BF8" box="[678,814,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="5.9" metricValueMin="5.2" pageId="2" pageNumber="287" unit="mm" value="5.55" valueMax="5.9" valueMin="5.2">5.2–5.9 mm</quantity>
versus
<quantity id="4CA09B0D0657157139EEFFD629081BF8" box="[901,1037,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.2" metricValueMin="6.8" pageId="2" pageNumber="287" unit="mm" value="7.5" valueMax="8.2" valueMin="6.8">6.8–8.2 mm</quantity>
) (
<bibRefCitation id="EFC94B19065715713E4FFFD629FC1BF8" author="Larson, D. J. &amp; Alarie, Y. &amp; Roughley, R. E." box="[1060,1273,187,213]" pageId="2" pageNumber="287" refId="ref2394" refString="Larson, D. J., Alarie, Y. &amp; Roughley, R. E. (2000) Predaceous diving beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. NRC Research Press, Ottawa, 982 pp." type="book" year="2000">
Larson
<emphasis id="B92CEAFA065715713E17FFD629B01BF8" box="[1148,1205,187,213]" italics="true" pageId="2" pageNumber="287">et al.</emphasis>
2000
</bibRefCitation>
).
<emphasis id="B92CEAFA065715713F67FFD7281F1BF9" bold="true" box="[1292,1306,186,212]" pageId="2" pageNumber="287">†</emphasis>
<taxonomicName id="4C584D6B065715713F49FFD628981BF8" authority="Hendrich &amp; Balke, 2020" authorityName="Hendrich &amp; Balke" authorityYear="2020" box="[1314,1437,187,213]" class="Insecta" family="Dytiscidae" genus="Coptotomus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="balticus" status="sp. nov.">
<emphasis id="B92CEAFA065715713F49FFD628981BF8" box="[1314,1437,187,213]" italics="true" pageId="2" pageNumber="287">C. balticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F5781065715713AFCFFB22DD21BD4" box="[151,215,223,249]" pageId="2" pageNumber="287" rank="species">
<emphasis id="B92CEAFA065715713AFCFFB22DD51BD4" bold="true" box="[151,208,223,249]" pageId="2" pageNumber="287">sp. n</emphasis>
.
</taxonomicNameLabel>
is the smallest species of the genus (general size range of extant
<taxonomicName id="4C584D6B0657157139D1FFB229441BD4" authorityName="Say" authorityYear="1830" box="[954,1089,223,249]" class="Insecta" family="Dytiscidae" genus="Coptotomus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAFA0657157139D1FFB229441BD4" box="[954,1089,223,249]" italics="true" pageId="2" pageNumber="287">Coptotomus</emphasis>
</taxonomicName>
species TL =
<quantity id="4CA09B0D065715713E89FFB328691BD4" box="[1250,1388,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="7.15" metricValueMax="8.6" metricValueMin="5.7" pageId="2" pageNumber="287" unit="mm" value="7.15" valueMax="8.6" valueMin="5.7">5.7–8.6 mm</quantity>
, see
<bibRefCitation id="EFC94B19065715713AFCFE6E2C6D1A30" author="Larson, D. J. &amp; Alarie, Y. &amp; Roughley, R. E." box="[151,360,259,285]" pageId="2" pageNumber="287" refId="ref2394" refString="Larson, D. J., Alarie, Y. &amp; Roughley, R. E. (2000) Predaceous diving beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. NRC Research Press, Ottawa, 982 pp." type="book" year="2000">
Larson
<emphasis id="B92CEAFA065715713A87FE6E2C211A30" box="[236,292,259,285]" italics="true" pageId="2" pageNumber="287">et al.</emphasis>
2000
</bibRefCitation>
).
</paragraph>
<caption id="DF276660065715713AFCF8FF2F7C1CDF" ID-DOI="http://doi.org/10.5281/zenodo.4322577" ID-Zenodo-Dep="4322577" httpUri="https://zenodo.org/record/4322577/files/figure.png" pageId="2" pageNumber="287" startId="2.[151,250,1938,1962]" targetBox="[151,1435,306,1912]" targetPageId="2">
<paragraph id="8BE736E8065715713AFCF8FF2F7C1CDF" blockId="2.[151,1437,1938,2034]" pageId="2" pageNumber="287">
<emphasis id="B92CEAFA065715713AFCF8FF2C0B1C87" bold="true" box="[151,270,1938,1962]" pageId="2" pageNumber="287">FIGURE 1</emphasis>
. †
<taxonomicName id="4C584D6B065715713B45F8FE2F051C87" authority="Hendrich &amp; Balke, 2020" authorityName="Hendrich &amp; Balke" authorityYear="2020" box="[302,512,1938,1962]" class="Insecta" family="Dytiscidae" genus="Coptotomus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="balticus" status="sp. nov.">
<emphasis id="B92CEAFA065715713B45F8FE2F051C87" box="[302,512,1938,1962]" italics="true" pageId="2" pageNumber="287">Coptotomus balticus</emphasis>
</taxonomicName>
<emphasis id="B92CEAFA06571571386CF8FF2F441C87" bold="true" box="[519,577,1938,1962]" pageId="2" pageNumber="287">
<taxonomicNameLabel id="A21F578106571571386CF8FF2F441C87" box="[519,577,1938,1962]" pageId="2" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male, holotype:
<emphasis id="B92CEAFA065715713899F8FF2E011C87" bold="true" box="[754,772,1938,1962]" pageId="2" pageNumber="287">A</emphasis>
) habitus (dorsal side);
<emphasis id="B92CEAFA06571571399BF8FF29041C87" bold="true" box="[1008,1025,1938,1962]" pageId="2" pageNumber="287">B</emphasis>
) habitus ventral side, scale bar = 1 mm;
<emphasis id="B92CEAFA065715713AFCF8DB2DAC1CE3" bold="true" box="[151,169,1974,1998]" pageId="2" pageNumber="287">C</emphasis>
) Pro- and mesotarsus, and apical palpomere of both the maxillary and labial palps distinctly bifid (red arrows);
<emphasis id="B92CEAFA065715713F4FF8DB28331CE3" bold="true" box="[1316,1334,1974,1998]" pageId="2" pageNumber="287">D</emphasis>
) Metatarsomeres 1 to 4 with externoapical angles lobed.
</paragraph>
</caption>
<paragraph id="8BE736E8065615703AACFFFA2EF71BF8" blockId="3.[151,1437,150,321]" pageId="3" pageNumber="288">
The female of
<emphasis id="B92CEAFA065615703B06FFFB2C7E1B9D" bold="true" box="[365,379,150,176]" pageId="3" pageNumber="288">†</emphasis>
<taxonomicName id="4C584D6B065615703B10FFFA2CF01B9C" authority="Hendrich &amp; Balke, 2020" authorityName="Hendrich &amp; Balke" authorityYear="2020" box="[379,501,151,177]" class="Insecta" family="Dytiscidae" genus="Coptotomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="288" phylum="Arthropoda" rank="species" species="balticus" status="sp. nov.">
<emphasis id="B92CEAFA065615703B10FFFA2CF01B9C" box="[379,501,151,177]" italics="true" pageId="3" pageNumber="288">C. balticus</emphasis>
</taxonomicName>
<emphasis id="B92CEAFA065615703B97FFFA2F391B9C" bold="true" box="[508,572,151,177]" pageId="3" pageNumber="288">
<taxonomicNameLabel id="A21F5781065615703B97FFFA2F391B9C" box="[508,572,151,177]" pageId="3" pageNumber="288" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
possesses longer and more conspicuous longitudinal striae on pronotum and elytron than the extant species (
<figureCitation id="13632A6D065615703B91FFD62F761BF8" box="[506,627,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1934,1958]" captionTargetBox="[164,1423,346,1911]" captionTargetId="figure-95@3.[164,1423,345,1911]" captionTargetPageId="3" captionText="FIGURE 2. † Coptotomus balticus sp. n., female, paratype: A) habitus (dorsal side, total length = 5.9 mm); B) habitus ventral side; C) elytron with strioles (scale bar = 1 mm); D) ventral side with pronotal process and apical palpomere of both the maxillary and labial palps bifid (scale bar = 1 mm); E) metasternum and metaventrite (scale bar = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.4322579" httpUri="https://zenodo.org/record/4322579/files/figure.png" pageId="3" pageNumber="288">Fig. 2A, C</figureCitation>
), which only possess fine strioles.
</paragraph>
<paragraph id="8BE736E8065615703AACFFB32EB21A6C" blockId="3.[151,1437,150,321]" pageId="3" pageNumber="288">
<emphasis id="B92CEAFA065615703AACFFB32C2F1BD5" bold="true" box="[199,298,222,248]" pageId="3" pageNumber="288">Habitat.</emphasis>
Unknown. All extant species inhabit ponds or slowly flowing streams with dense vegetation (Bergsten &amp; Miller 2016). The larvae of
<taxonomicName id="4C584D6B065615703B84FE6E2E901A30" authority="Hilsenhoff, 1980" authorityName="Hilsenhoff" authorityYear="1980" box="[495,917,259,285]" class="Insecta" family="Dytiscidae" genus="Coptotomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="288" phylum="Arthropoda" rank="subSpecies" species="longulus" subSpecies="lenticus">
<emphasis id="B92CEAFA065615703B84FE6E2FCA1A30" box="[495,719,259,285]" italics="true" pageId="3" pageNumber="288">C. longulus lenticus</emphasis>
<bibRefCitation id="EFC94B190656157038BCFE6E2E901A30" author="Hilsenhoff, W. L." box="[727,917,259,285]" pageId="3" pageNumber="288" pagination="461 - 471" refId="ref2280" refString="Hilsenhoff, W. L. (1980) Coptotomus (Coleoptera: Dytiscidae) in eastern North America with descriptions of two new species. Transactions of the American Entomological Society, 105 (4), 461 - 471." type="journal article" year="1980">Hilsenhoff, 1980</bibRefCitation>
</taxonomicName>
are benthic inhabitants of ponds (
<bibRefCitation id="EFC94B19065615703F78FE6E2DD11A6C" author="Bacon, M. A. &amp; Barman, E. H. &amp; White, B. P." pageId="3" pageNumber="288" pagination="75 - 81" refId="ref1910" refString="Bacon, M. A., Barman, E. H. &amp; White, B. P. (2000) Biology of Coptotomus lenticus (Coleoptera: Dytiscidae: Coptotominae) with a description of its mature larva. Journal of the Elisha Mitchell Scientific Society, 116, 75 - 81." type="journal article" year="2000">
Bacon
<emphasis id="B92CEAFA065615703F0FFE6E28981A30" box="[1380,1437,259,285]" italics="true" pageId="3" pageNumber="288">et al.</emphasis>
2000
</bibRefCitation>
). All species are capable to flight and have been collected at light.
</paragraph>
</subSubSection>
</treatment>
</document>
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