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<document id="AF90EAAE3BA7E2318DF371F2E6498310" ID="10.11646/zootaxa.4169.1.5" ID-DOI="10.11646/zootaxa.4169.1.5" ID-GBIF-Dataset="e35dbd92-b156-4ae3-9f90-9996c91ee626" ID-ISSN="1175-5326" ID-Zenodo-Dep="265168" ID-ZooBank="62419F0E-128A-4E60-932C-DD746BA57906" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474268893273" checkinUser="plazi" docAuthor="Thoni, R. J., Gurung, D. B. &amp; Mayden, R. L." docDate="2016" docId="03F187D8FFBBFF87FF1BF9E3FE9BFE30" docLanguage="en" docName="zootaxa.4169.1.5.pdf" docOrigin="Zootaxa 4169 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Garra arupi Nebeshwar, Vishwanath &amp; Das 2009" docType="treatment" docVersion="7" lastPageNumber="118" masterDocId="FFC8FFA0FFBAFF84FF8CFFB8FFA3FF9D" masterDocTitle="A review of the genus Garra Hamilton 1822 of Bhutan, including the descriptions of two new species and three additional records (Cypriniformes: Cyprinidae)" masterLastPageNumber="132" masterPageNumber="115" pageNumber="116" updateTime="1698386207380" updateUser="plazi">
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<mods:title id="E83BD1DB766785A9E246321FEB18DB90">A review of the genus Garra Hamilton 1822 of Bhutan, including the descriptions of two new species and three additional records (Cypriniformes: Cyprinidae)</mods:title>
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<mods:namePart id="9FFEB0626A073911E265CAD67F202A96">Thoni, R. J.</mods:namePart>
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<treatment id="03F187D8FFBBFF87FF1BF9E3FE9BFE30" ID-DOI="http://doi.org/10.5281/zenodo.6087315" ID-GBIF-Taxon="124551993" ID-Zenodo-Dep="6087315" LSID="urn:lsid:plazi:treatment:03F187D8FFBBFF87FF1BF9E3FE9BFE30" httpUri="http://treatment.plazi.org/id/03F187D8FFBBFF87FF1BF9E3FE9BFE30" lastPageId="3" lastPageNumber="118" pageId="1" pageNumber="116">
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<heading id="D0AF81A2FFBBFF85FF1BF9E3FD5FF9E8" bold="true" box="[151,764,1627,1653]" fontSize="11" level="1" pageId="1" pageNumber="116" reason="1">
<taxonomicName id="4C584D4DFFBBFF85FF1BF9E3FD5FF9E8" ID-CoL="3FBGX" authority="Nebeshwar, Vishwanath &amp; Das, 2009" authorityName="Nebeshwar, Vishwanath &amp; Das" authorityYear="2009" box="[151,764,1627,1653]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="116" phylum="Chordata" rank="species" species="arupi">
<emphasis id="B92CEADCFFBBFF85FF1BF9E3FD5FF9E8" bold="true" box="[151,764,1627,1653]" pageId="1" pageNumber="116">
<emphasis id="B92CEADCFFBBFF85FF1BF9E3FE8BF9E8" bold="true" box="[151,296,1627,1653]" italics="true" pageId="1" pageNumber="116">Garra arupi</emphasis>
<bibRefCitation id="EFC94B3FFFBBFF85FEA3F9E3FD5FF9E8" author="Nebeshwar" box="[303,764,1627,1653]" pageId="1" pageNumber="116" refString="Nebeshwar, K., Vishwanath, W. &amp; Das, D. N. (2009) Garra arupi, a new cyprinid fish species (Cypriniformes: Cyprinidae) from upper Brahmaputra basin in Arunachal Pradesh, India. Journal of Threatened Taxa, 1 (4) 197 - 202. http: // dx. doi. org / 10.11609 / JoTT. o 1842.197 - 202" type="journal article" year="2009">Nebeshwar, Vishwanath &amp; Das, 2009</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE736CEFFBBFF85FF1BF9C6FE8BF90A" blockId="1.[151,764,1627,1687]" box="[151,296,1662,1687]" pageId="1" pageNumber="116">
(
<figureCitation id="13632A4BFFBBFF85FF13F9C6FF4FF90A" box="[159,236,1662,1687]" captionStart="FIGURE 1" captionStartId="2.[151,250,1238,1260]" captionTargetBox="[166,1423,206,1201]" captionTargetId="figure@2.[151,1436,193,1216]" captionTargetPageId="2" captionText="FIGURE 1. Garra arupi. CNR 11066, 72.3 mm SL; Bhutan: Dagana Dzongkhag: Serichhu near confluence with Punatsangchhu, R. J. Thoni &amp; D. B. Gurung, September, 2013." httpUri="https://zenodo.org/record/265169/files/figure.png" pageId="1" pageNumber="116">Figs. 1</figureCitation>
&amp;
<figureCitation id="13632A4BFFBBFF85FE9CF9C7FEBCF90A" box="[272,287,1663,1687]" captionStart="FIGURE 2" captionStartId="2.[151,250,1830,1852]" captionTargetBox="[151,1436,1340,1809]" captionTargetId="figure@2.[151,1436,1340,1809]" captionTargetPageId="2" captionText="FIGURE 2. Live specimen Garra arupi. CNR 11063, 66.4 mm SL; Bhutan: Dagana Dzongkhag: Serichhu near confluence with Punatsangchhu, R. J. Thoni &amp; D. B. Gurung, September, 2013." httpUri="https://zenodo.org/record/265170/files/figure.png" pageId="1" pageNumber="116">2</figureCitation>
)
</paragraph>
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<paragraph id="8BE736CEFFBBFF85FF1BF97DFD7FF960" blockId="1.[151,1436,1732,1789]" pageId="1" pageNumber="116">
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<taxonomicName id="4C584D4DFFBBFF85FF1BF97DFD4CF947" ID-CoL="3FBGX" authority="Nebeshwar, Vishwanath &amp; Das 2009: 198" authorityName="Nebeshwar, Vishwanath &amp; Das" authorityPageNumber="198" authorityYear="2009" box="[151,751,1732,1754]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="116" phylum="Chordata" rank="species" species="arupi">
<emphasis id="B92CEADCFFBBFF85FF1BF97DFEBAF947" box="[151,281,1733,1754]" italics="true" pageId="1" pageNumber="116">Garra arupi</emphasis>
<treatmentCitation id="0AF910DFFFBBFF85FEA9F97CFD4CF947" author="Nebeshwar" box="[293,751,1732,1754]" page="198" pageId="1" pageNumber="116" year="2009">
<bibRefCitation id="EFC94B3FFFBBFF85FEA9F97CFD15F947" author="Nebeshwar" box="[293,694,1732,1754]" pageId="1" pageNumber="116" refString="Nebeshwar, K., Vishwanath, W. &amp; Das, D. N. (2009) Garra arupi, a new cyprinid fish species (Cypriniformes: Cyprinidae) from upper Brahmaputra basin in Arunachal Pradesh, India. Journal of Threatened Taxa, 1 (4) 197 - 202. http: // dx. doi. org / 10.11609 / JoTT. o 1842.197 - 202" type="journal article" year="2009">Nebeshwar, Vishwanath &amp; Das 2009</bibRefCitation>
: 198
</treatmentCitation>
</taxonomicName>
. Type locality: Deopani River at Roing, Lower Divang Valley, Arunachal Pradesh, India. Holotype: RGUMF-0184.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3426545FFBBFF85FF1BF895FEDCF86B" pageId="1" pageNumber="116" type="materials_examined">
<paragraph id="8BE736CEFFBBFF85FF1BF895FEDCF86B" blockId="1.[151,1436,1837,2038]" pageId="1" pageNumber="116">
<materialsCitation id="3B303C93FFBBFF85FF1BF895FADCF8F4" ID-GBIF-Occurrence="1313779505" collectingDate="2011-05" collectionCode="CNR" collectorName="Salamji Gewog, D. B." country="Bhutan" location="Serichhu River" pageId="1" pageNumber="116" specimenCode="CNR 11061, CNR 11062, CNR 11063, CNR 11064, CNR 11066, CNR 11999, 57.9" specimenCount="6" stateProvince="Dagana">
<emphasis id="B92CEADCFFBBFF85FF1BF895FED9F8DB" bold="true" box="[151,378,1837,1862]" pageId="1" pageNumber="116">Material examined</emphasis>
.
<specimenCount id="9D5EFD47FFBBFF85FE04F895FDB9F8DB" box="[392,538,1837,1862]" pageId="1" pageNumber="116" type="generic">6 Specimens</specimenCount>
,
<specimenCode id="DBFE9EB5FFBBFF85FDAAF896FF6BF8F4" pageId="1" pageNumber="116">
<collectionCode id="ED49AE0BFFBBFF85FDAAF896FDC3F8DB" box="[550,608,1838,1862]" country="Ukraine" httpUri="http://grbio.org/cool/5zp2-7vs1" name="Crimean Natural Reserve" pageId="1" pageNumber="116">CNR</collectionCode>
11061,
<collectionCode id="ED49AE0BFFBBFF85FD37F896FD57F8DB" box="[699,756,1838,1862]" country="Ukraine" httpUri="http://grbio.org/cool/5zp2-7vs1" name="Crimean Natural Reserve" pageId="1" pageNumber="116">CNR</collectionCode>
11062,
<collectionCode id="ED49AE0BFFBBFF85FCDCF896FC29F8DB" box="[848,906,1838,1862]" country="Ukraine" httpUri="http://grbio.org/cool/5zp2-7vs1" name="Crimean Natural Reserve" pageId="1" pageNumber="116">CNR</collectionCode>
11063,
<collectionCode id="ED49AE0BFFBBFF85FC69F896FBBCF8DB" box="[997,1055,1838,1862]" country="Ukraine" httpUri="http://grbio.org/cool/5zp2-7vs1" name="Crimean Natural Reserve" pageId="1" pageNumber="116">CNR</collectionCode>
11064,
<collectionCode id="ED49AE0BFFBBFF85FBF6F896FB10F8DB" box="[1146,1203,1838,1862]" country="Ukraine" httpUri="http://grbio.org/cool/5zp2-7vs1" name="Crimean Natural Reserve" pageId="1" pageNumber="116">CNR</collectionCode>
11066,
<collectionCode id="ED49AE0BFFBBFF85FA83F896FAEAF8DB" box="[1295,1353,1838,1862]" country="Ukraine" httpUri="http://grbio.org/cool/5zp2-7vs1" name="Crimean Natural Reserve" pageId="1" pageNumber="116">CNR</collectionCode>
11999,
<quantity id="4CA09B2BFFBBFF85FF1BF8E8FF6BF8F4" box="[151,200,1872,1897]" metricMagnitude="-2" metricUnit="m" metricValue="7.325" metricValueMax="8.86" metricValueMin="5.79" pageId="1" pageNumber="116" unit="mm" value="73.25" valueMax="88.6" valueMin="57.9">57.9</quantity>
</specimenCode>
88.6 mm SL,
<collectingCountry id="F34F765EFFBBFF85FEF4F8E8FE6EF8F4" box="[376,461,1872,1897]" name="Bhutan" pageId="1" pageNumber="116">Bhutan</collectingCountry>
:
<collectingRegion id="499CF82CFFBBFF85FE56F8E9FD8CF8F4" box="[474,559,1873,1897]" country="Bhutan" name="Daga" pageId="1" pageNumber="116">Dagana</collectingRegion>
Dzongkhag:
<location id="8E876015FFBBFF85FD41F8E8FCDBF8F4" LSID="urn:lsid:plazi:treatment:03F187D8FFBBFF87FF1BF9E3FE9BFE30:8E876015FFBBFF85FD41F8E8FCDBF8F4" box="[717,888,1872,1897]" country="Bhutan" name="Serichhu River" pageId="1" pageNumber="116" stateProvince="Dagana">Serichhu River</location>
in
<collectorName id="26AD5318FFBBFF85FC2EF8E8FB3BF8F5" box="[930,1176,1872,1897]" pageId="1" pageNumber="116">Salamji Gewog, D.B.</collectorName>
Gurung,
<collectingDate id="EFA2E9E6FFBBFF85FA84F8E9FADCF8F4" box="[1288,1407,1873,1897]" pageId="1" pageNumber="116" value="2011-05">
<date id="FFE6100EFFBBFF85FA84F8E9FADCF8F4" box="[1288,1407,1873,1897]" pageId="1" pageNumber="116" value="2011-05">May 2011</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B303C93FFBBFF85FA02F8E9FCB2F832" ID-GBIF-Occurrence="1313779506" collectingDate="2013-09-21" collectionCode="CNR" collectorName="Salamji Gewog, R. J. &amp; D. B. Gurung" country="Bhutan" location="Serichhu River" pageId="1" pageNumber="116" specimenCode="CNR 13032, CNR13033, 49.9" specimenCount="2" stateProvince="Dagana">
<specimenCount id="9D5EFD47FFBBFF85FA02F8E9FEB0F810" pageId="1" pageNumber="116" type="generic">2 Specimens</specimenCount>
,
<specimenCode id="DBFE9EB5FFBBFF85FEACF8CCFDDBF810" box="[288,632,1908,1933]" pageId="1" pageNumber="116">
<collectionCode id="ED49AE0BFFBBFF85FEACF8CCFEF9F811" box="[288,346,1908,1932]" country="Ukraine" httpUri="http://grbio.org/cool/5zp2-7vs1" name="Crimean Natural Reserve" pageId="1" pageNumber="116">CNR</collectionCode>
13032,
<collectionCode id="ED49AE0BFFBBFF85FE34F8CCFE56F811" box="[440,501,1908,1932]" country="Ukraine" httpUri="http://grbio.org/cool/5zp2-7vs1" name="Crimean Natural Reserve" pageId="1" pageNumber="116">CNR</collectionCode>
13033, 49.9
</specimenCode>
51.0 mm SL,
<collectingCountry id="F34F765EFFBBFF85FCA8F8CCFCDAF810" box="[804,889,1908,1933]" name="Bhutan" pageId="1" pageNumber="116">Bhutan</collectingCountry>
:
<collectingRegion id="499CF82CFFBBFF85FC09F8CDFC78F810" box="[901,987,1909,1933]" country="Bhutan" name="Daga" pageId="1" pageNumber="116">Dagana</collectingRegion>
Dzongkhag:
<location id="8E876015FFBBFF85FBF9F8CCFABCF810" LSID="urn:lsid:plazi:treatment:03F187D8FFBBFF87FF1BF9E3FE9BFE30:8E876015FFBBFF85FBF9F8CCFABCF810" box="[1141,1311,1908,1933]" country="Bhutan" name="Serichhu River" pageId="1" pageNumber="116" stateProvince="Dagana">Serichhu River</location>
in
<collectorName id="26AD5318FFBBFF85FACBF8CCFE82F832" pageId="1" pageNumber="116">Salamji Gewog, R.J.</collectorName>
Thoni &amp;
<collectorName id="26AD5318FFBBFF85FE02F820FD82F832" box="[398,545,1943,1967]" pageId="1" pageNumber="116">D.B. Gurung</collectorName>
,
<collectingDate id="EFA2E9E6FFBBFF85FDA1F82FFCB2F832" box="[557,785,1943,1968]" pageId="1" pageNumber="116" value="2013-09-21">
<date id="FFE6100EFFBBFF85FDA1F82FFCB2F832" box="[557,785,1943,1968]" pageId="1" pageNumber="116" value="2013-09-21">September 21, 2013</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B303C93FFBBFF85FC97F82FFED8F86B" ID-GBIF-Occurrence="1313779504" collectingDate="2013-09-24" collectionCode="CNR" collectorName="Gakiling Gewog, D. B. &amp; R. J. Thoni" country="Bhutan" location="Laringkhola River" pageId="1" pageNumber="116" specimenCode="CNR 13111, CNR 13025-13026, 50.4" specimenCount="3">
<specimenCount id="9D5EFD47FFBBFF85FC97F82FFC0FF82D" box="[795,940,1943,1968]" pageId="1" pageNumber="116" type="generic">3 Specimens</specimenCount>
,
<specimenCode id="DBFE9EB5FFBBFF85FC3BF82FFAFEF82D" box="[951,1373,1943,1968]" pageId="1" pageNumber="116">
<collectionCode id="ED49AE0BFFBBFF85FC3BF82FFC52F832" box="[951,1009,1943,1967]" country="Ukraine" httpUri="http://grbio.org/cool/5zp2-7vs1" name="Crimean Natural Reserve" pageId="1" pageNumber="116">CNR</collectionCode>
13111,
<collectionCode id="ED49AE0BFFBBFF85FBC5F82FFB20F832" box="[1097,1155,1943,1967]" country="Ukraine" httpUri="http://grbio.org/cool/5zp2-7vs1" name="Crimean Natural Reserve" pageId="1" pageNumber="116">CNR</collectionCode>
13025-13026,
<quantity id="4CA09B2BFFBBFF85FAA0F82FFAFEF82D" box="[1324,1373,1943,1968]" metricMagnitude="-2" metricUnit="m" metricValue="5.8549999999999995" metricValueMax="6.67" metricValueMin="5.04" pageId="1" pageNumber="116" unit="mm" value="58.55" valueMax="66.7" valueMin="50.4">50.4</quantity>
</specimenCode>
66.7 mm SL;
<collectingCountry id="F34F765EFFBBFF85FE8BF802FEFFF84E" box="[263,348,1978,2003]" name="Bhutan" pageId="1" pageNumber="116">Bhutan</collectingCountry>
: Sarpang Dzongkhag:
<location id="8E876015FFBBFF85FDE3F802FCE6F84E" LSID="urn:lsid:plazi:treatment:03F187D8FFBBFF87FF1BF9E3FE9BFE30:8E876015FFBBFF85FDE3F802FCE6F84E" box="[623,837,1978,2003]" country="Bhutan" name="Laringkhola River" pageId="1" pageNumber="116">Laringkhola River</location>
in
<collectorName id="26AD5318FFBBFF85FCFBF802FB21F84F" box="[887,1154,1978,2003]" pageId="1" pageNumber="116">Gakiling Gewog, D.B.</collectorName>
Gurung &amp;
<collectorName id="26AD5318FFBBFF85FA95F803FA3BF84E" box="[1305,1432,1978,2003]" pageId="1" pageNumber="116">R.J. Thoni</collectorName>
,
<collectingDate id="EFA2E9E6FFBBFF85FF1BF865FED8F86B" box="[151,379,2013,2038]" pageId="1" pageNumber="116" value="2013-09-24">
<date id="FFE6100EFFBBFF85FF1BF865FED8F86B" box="[151,379,2013,2038]" pageId="1" pageNumber="116" value="2013-09-24">September 24, 2013</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF276646FFB8FF86FF1BFB6EFCAAFA97" httpUri="https://zenodo.org/record/265169/files/figure.png" pageId="2" pageNumber="117" targetBox="[166,1423,206,1201]" targetPageId="2">
<paragraph id="8BE736CEFFB8FF86FF1BFB6EFCAAFA97" blockId="2.[151,1435,1238,1290]" pageId="2" pageNumber="117">
<emphasis id="B92CEADCFFB8FF86FF1BFB6EFEBFFB76" bold="true" box="[151,284,1238,1260]" pageId="2" pageNumber="117">FIGURE 1.</emphasis>
<taxonomicName id="4C584D4DFFB8FF86FEA7FB6FFE13FB71" box="[299,432,1239,1260]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="117" phylum="Chordata" rank="species" species="arupi">
<emphasis id="B92CEADCFFB8FF86FEA7FB6FFE13FB71" box="[299,432,1239,1260]" italics="true" pageId="2" pageNumber="117">Garra arupi</emphasis>
</taxonomicName>
. CNR 11066, 72.3 mm SL; Bhutan: Dagana Dzongkhag: Serichhu near confluence with Punatsangchhu, R.J. Thoni &amp; D.B. Gurung, September, 2013.
</paragraph>
</caption>
<caption id="DF276646FFB8FF86FF1BF89EFC9FF8C6" httpUri="https://zenodo.org/record/265170/files/figure.png" pageId="2" pageNumber="117" targetBox="[151,1436,1340,1809]" targetPageId="2">
<paragraph id="8BE736CEFFB8FF86FF1BF89EFC9FF8C6" blockId="2.[151,1436,1830,1883]" pageId="2" pageNumber="117">
<emphasis id="B92CEADCFFB8FF86FF1BF89EFEB6F8A1" bold="true" box="[151,277,1830,1852]" pageId="2" pageNumber="117">FIGURE 2.</emphasis>
Live specimen
<taxonomicName id="4C584D4DFFB8FF86FE30F89FFD98F8A0" box="[444,571,1831,1853]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="117" phylum="Chordata" rank="species" species="arupi">
<emphasis id="B92CEADCFFB8FF86FE30F89FFD98F8A0" box="[444,571,1831,1853]" italics="true" pageId="2" pageNumber="117">Garra arupi</emphasis>
</taxonomicName>
. CNR 11063, 66.4 mm SL; Bhutan: Dagana Dzongkhag: Serichhu near confluence with Punatsangchhu, R.J. Thoni &amp; D.B. Gurung, September, 2013.
</paragraph>
</caption>
<subSubSection id="C3426545FFB8FF87FF4BF83AFE9BFE30" lastPageId="3" lastPageNumber="118" pageId="2" pageNumber="117" type="discussion">
<paragraph id="8BE736CEFFB8FF87FF4BF83AFE71FF6A" blockId="2.[151,1437,1922,2019]" lastBlockId="3.[151,1436,151,429]" lastPageId="3" lastPageNumber="118" pageId="2" pageNumber="117">
<emphasis id="B92CEADCFFB8FF86FF4BF83AFE98F806" bold="true" box="[199,315,1922,1947]" pageId="2" pageNumber="117">Remarks.</emphasis>
When live, this species is most easily diagnosed by coloration characters, appearing bright orange or red on the lips, and the anterior edge the dorsal, anal, and paired fins, as well as on the upper and lower caudalfin margin. Significant diagnostic traits on preserved specimens include an anteriorly placed vent, separate anterolateral lobe, medial caudal-fin stripe, submarginal stripe on the dorsal fin, and seven branched dorsal-fin rays. A weakly developed transverse groove and proboscis with few tubercles is similar to that of
<taxonomicName id="4C584D4DFFB9FF87FB7CFF05FAF6FF48" box="[1264,1365,188,213]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="118" phylum="Chordata" rank="species" species="gotyla">
<emphasis id="B92CEADCFFB9FF87FB7CFF05FAF6FF48" box="[1264,1365,188,213]" italics="true" pageId="3" pageNumber="118">G. gotyla</emphasis>
</taxonomicName>
but is less pronounced in
<emphasis id="B92CEADCFFB9FF87FEFCFF58FE64FF65" box="[368,455,224,248]" italics="true" pageId="3" pageNumber="118">G. a r u p i</emphasis>
.
</paragraph>
<paragraph id="8BE736CEFFB9FF87FF4BFEBCFE9BFE30" blockId="3.[151,1436,151,429]" pageId="3" pageNumber="118">
This species was collected primarily in central
<collectingCountry id="F34F765EFFB9FF87FD6BFEBCFC98FE80" box="[743,827,260,285]" name="Bhutan" pageId="3" pageNumber="118">Bhutan</collectingCountry>
, in tributaries of the Punatsangchhu and Laringkhola rivers. They are found in mid-elevation streams typically above
<quantity id="4CA09B2BFFB9FF87FCEEFE90FC0BFEA2" box="[866,936,296,320]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="118" unit="m" value="200.0">200 m</quantity>
and below
<quantity id="4CA09B2BFFB9FF87FBA4FE90FB23FEA2" box="[1064,1152,296,320]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="118" unit="m" value="1000.0">1000 m</quantity>
. They tend to be found in runs, riffles, and torrents, with few encounters in pools, with the exception of larvae and juveniles, which prefer the gravel substrate of pools. They were frequently sampled in sympatry with
<taxonomicName id="4C584D4DFFB9FF87FC57FED7FB96FE15" box="[987,1077,367,392]" class="Actinopterygii" family="Cyprinidae" genus="Barilius" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="118" phylum="Chordata" rank="genus">
<emphasis id="B92CEADCFFB9FF87FC57FED7FB96FE15" box="[987,1077,367,392]" italics="true" pageId="3" pageNumber="118">Barilius</emphasis>
</taxonomicName>
,
<emphasis id="B92CEADCFFB9FF87FBC8FED7FA3FFE15" box="[1092,1436,367,392]" italics="true" pageId="3" pageNumber="118">
<taxonomicName id="4C584D4DFFB9FF87FBC8FED7FB48FE15" box="[1092,1259,367,392]" class="Actinopterygii" family="Cyprinidae" genus="Neolissochilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="118" phylum="Chordata" rank="genus">Neolissochilus</taxonomicName>
,
<taxonomicName id="4C584D4DFFB9FF87FB79FED7FA3BFE15" box="[1269,1432,367,392]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="118" phylum="Chordata" rank="genus">Psilorhynchus</taxonomicName>
,
</emphasis>
and
<emphasis id="B92CEADCFFB9FF87FF4AFE2CFE9BFE30" box="[198,312,404,429]" italics="true" pageId="3" pageNumber="118">
<taxonomicName id="4C584D4DFFB9FF87FF4AFE2CFE97FE30" box="[198,308,404,429]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="118" phylum="Chordata" rank="genus">Schistura</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>