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<document id="57C0B7AAC0C5F53E9C0795F442D43C3C" ID="10.11646/zootaxa.4169.1.5" ID-DOI="10.11646/zootaxa.4169.1.5" ID-GBIF-Dataset="e35dbd92-b156-4ae3-9f90-9996c91ee626" ID-ISSN="1175-5326" ID-Zenodo-Dep="265168" ID-ZooBank="62419F0E-128A-4E60-932C-DD746BA57906" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474268893273" checkinUser="plazi" docAuthor="Thoni, R. J., Gurung, D. B. &amp; Mayden, R. L." docDate="2016" docId="03F187D8FFB9FF83FF1BFE4DFC3BFAC9" docLanguage="en" docName="zootaxa.4169.1.5.pdf" docOrigin="Zootaxa 4169 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Garra bimaculacauda Thoni, Gurung &amp; Mayden, 2016, sp. nov." docType="treatment" docVersion="7" lastPageNumber="122" masterDocId="FFC8FFA0FFBAFF84FF8CFFB8FFA3FF9D" masterDocTitle="A review of the genus Garra Hamilton 1822 of Bhutan, including the descriptions of two new species and three additional records (Cypriniformes: Cyprinidae)" masterLastPageNumber="132" masterPageNumber="115" pageNumber="118" updateTime="1698386207380" updateUser="plazi">
<mods:mods id="A58FB081911D741BBAAC7E5394B23ED0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="16D17E66601C1CA54BD3A40DB0E4F553">A review of the genus Garra Hamilton 1822 of Bhutan, including the descriptions of two new species and three additional records (Cypriniformes: Cyprinidae)</mods:title>
</mods:titleInfo>
<mods:name id="2F8669009F2C08583D5D0F4C76C99B68" type="personal">
<mods:role id="D875232BF8C460527EB5990371CFD5D9">
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<mods:namePart id="07A5DEF73B14EE437E06F972C330D11B">Thoni, R. J.</mods:namePart>
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<mods:name id="8D986F4C8ADBF4E2E8B6FA65D765FEDC" type="personal">
<mods:role id="C3FF3B6B7AD801D7B3617C1BE896F07B">
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<mods:namePart id="32984DFEF72B1902E2DC56A488B55C16">Gurung, D. B.</mods:namePart>
</mods:name>
<mods:name id="606176563EAAD00A3D139D1DC8091E0B" type="personal">
<mods:role id="8E1883D9773139CB90EEBFC17D333E2F">
<mods:roleTerm id="6E3F75099CC59D95A11EB048BA833DA0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B0E76038C0FAD7F5EFAB881A6BE6A509">Mayden, R. L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="FC1B523CC656EA349D51F27B3E9522C9">text</mods:typeOfResource>
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<mods:title id="107227BF6DBF0AE95638B7DB4DE935C2">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E486FF9D7FD1D7739B893C034F4A77F0">
<mods:date id="ED73706B58A7949667E89561156E1E12">2016</mods:date>
<mods:detail id="B2872D986F2A7D3D57E3035E8DFBAC0D" type="volume">
<mods:number id="8D344902D26D41AB1BCA9F9004405071">4169</mods:number>
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<mods:detail id="FBA6A4CE8989E1131966A405F5BD9103" type="issue">
<mods:number id="004B28E766AB2A149CEE517020BED841">1</mods:number>
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<mods:classification id="89D1462228D203D0EA37285A2E986242">journal article</mods:classification>
<mods:identifier id="63A896990175CBDDB7D699DECD705D03" type="DOI">10.11646/zootaxa.4169.1.5</mods:identifier>
<mods:identifier id="F0B6AAB1BAFCAC3D98560F0446C925D3" type="GBIF-Dataset">e35dbd92-b156-4ae3-9f90-9996c91ee626</mods:identifier>
<mods:identifier id="F0BC4EAF5A8D80B35DBD49B1D7308504" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B88C758720680CDE3BB9DE7A42811AE5" type="Zenodo-Dep">265168</mods:identifier>
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</mods:mods>
<treatment id="03F187D8FFB9FF83FF1BFE4DFC3BFAC9" ID-DOI="http://doi.org/10.5281/zenodo.6087317" ID-GBIF-Taxon="124551995" ID-Zenodo-Dep="6087317" LSID="urn:lsid:plazi:treatment:03F187D8FFB9FF83FF1BFE4DFC3BFAC9" httpUri="http://treatment.plazi.org/id/03F187D8FFB9FF83FF1BFE4DFC3BFAC9" lastPageId="7" lastPageNumber="122" pageId="3" pageNumber="118">
<subSubSection id="C3426545FFB9FF87FF1BFE4DFE84FDAD" pageId="3" pageNumber="118" type="nomenclature">
<paragraph id="8BE736CEFFB9FF87FF1BFE4DFDAEFD92" blockId="3.[151,525,501,560]" box="[151,525,501,527]" pageId="3" pageNumber="118">
<heading id="D0AF81A2FFB9FF87FF1BFE4DFDAEFD92" bold="true" box="[151,525,501,527]" fontSize="11" level="1" pageId="3" pageNumber="118" reason="1">
<emphasis id="B92CEADCFFB9FF87FF1BFE4DFDAEFD92" bold="true" box="[151,525,501,527]" pageId="3" pageNumber="118">
<taxonomicName id="4C584D4DFFB9FF87FF1BFE4DFE02FD92" box="[151,417,501,527]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="118" phylum="Chordata" rank="species" species="bimaculacauda" status="sp. nov.">
<emphasis id="B92CEADCFFB9FF87FF1BFE4DFE02FD92" bold="true" box="[151,417,501,527]" italics="true" pageId="3" pageNumber="118">Garra bimaculacauda</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21F57A7FFB9FF87FE3CFE4DFDAEFD92" box="[432,525,501,527]" pageId="3" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE736CEFFB9FF87FF1BFDAFFE84FDAD" blockId="3.[151,525,501,560]" box="[151,295,535,560]" pageId="3" pageNumber="118">
(
<figureCitation id="13632A4BFFB9FF87FF13FDAFFF4FFDAD" box="[159,236,535,560]" captionStart="FIGURE 3" captionStartId="3.[151,250,1963,1985]" captionTargetBox="[186,1374,1009,1922]" captionTargetId="figure@3.[176,1388,994,1937]" captionTargetPageId="3" captionText="FIGURE 3. Garra bimaculacauda. Holotype. NBC 15184, 66.2 mm SL; Bhutan: Zhemgang Dzongkhag, near Panbang Town, R. J. Thoni &amp; D. B. Gurung, October, 2013." httpUri="https://zenodo.org/record/265171/files/figure.png" pageId="3" pageNumber="118">Figs. 3</figureCitation>
&amp;
<figureCitation id="13632A4BFFB9FF87FE9CFDA0FEBCFDAD" box="[272,287,536,560]" captionStart="FIGURE 4" captionStartId="5.[151,250,1396,1418]" captionTargetBox="[151,1436,953,1374]" captionTargetId="figure@5.[151,1436,953,1374]" captionTargetPageId="5" captionText="FIGURE 4. Live specimen Garra bimaculacauda. Holotype. NBC 15184, 66.2 mm SL; Bhutan, Zhemgang Dzongkhag, near Panbang, R. J. Thoni &amp; D. B. Gurung, October, 2013." httpUri="https://zenodo.org/record/265172/files/figure.png" pageId="3" pageNumber="118">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3426545FFB9FF87FF1BFDDAFC93FC93" pageId="3" pageNumber="118" type="materials_examined">
<paragraph id="8BE736CEFFB9FF87FF1BFDDAFCDCFD3F" blockId="3.[151,1436,607,962]" pageId="3" pageNumber="118">
<materialsCitation id="3B303C93FFB9FF87FF1BFDDAFCDCFD3F" ID-GBIF-Occurrence="1313779496" collectingDate="2013-09-28" collectorName="R. J. Thoni &amp; D. B. Gurung" country="Bhutan" latitude="1.5166667" location="Dangmechhu River" longLatPrecision="927" longitude="-36.059" municipality="Marangdud" pageId="3" pageNumber="118" specimenCount="1" stateProvince="Zhemgang" typeStatus="holotype">
<emphasis id="B92CEADCFFB9FF87FF1BFDDAFEA8FDE6" bold="true" box="[151,267,610,635]" pageId="3" pageNumber="118">
<typeStatus id="54E3886CFFB9FF87FF1BFDDAFEA5FDE6" box="[151,262,610,635]" pageId="3" pageNumber="118" type="holotype">Holotype</typeStatus>
.
</emphasis>
NBC 15184,
<quantity id="4CA09B2BFFB9FF87FE2AFDDAFDA9FDE7" box="[422,522,610,635]" metricMagnitude="-2" metricUnit="m" metricValue="6.62" pageId="3" pageNumber="118" unit="mm" value="66.2">66.2 mm</quantity>
SL;
<collectingCountry id="F34F765EFFB9FF87FDCCFDDAFD36FDE6" box="[576,661,610,635]" name="Bhutan" pageId="3" pageNumber="118">Bhutan</collectingCountry>
:
<collectingRegion id="499CF82CFFB9FF87FD2CFDDAFCBBFDE6" box="[672,792,610,635]" country="Bhutan" name="Shemgang" pageId="3" pageNumber="118">Zhemgang</collectingRegion>
Dzongkhag:
<collectingMunicipality id="6B83ACB4FFB9FF87FC22FDDAFB97FDE6" box="[942,1076,610,635]" pageId="3" pageNumber="118">Marangdud</collectingMunicipality>
:
<location id="8E876015FFB9FF87FBB2FDDAFAB8FDE6" LSID="urn:lsid:plazi:treatment:03F187D8FFB9FF83FF1BFE4DFC3BFAC9:8E876015FFB9FF87FBB2FDDAFAB8FDE6" box="[1086,1307,610,635]" country="Bhutan" latitude="1.5166667" longLatPrecision="927" longitude="-36.059" municipality="Marangdud" name="Dangmechhu River" pageId="3" pageNumber="118" stateProvince="Zhemgang">Dangmechhu River</location>
, 27
<geoCoordinate id="EE6C5009FFB9FF87FACDFDE7FA3FFDE6" box="[1345,1436,607,635]" direction="west" orientation="longitude" pageId="3" pageNumber="118" precision="55" value="-11.513">o 11.513</geoCoordinate>
<geoCoordinate id="EE6C5009FFB9FF87FF1BFD32FF78FD3F" box="[151,219,650,674]" direction="north" orientation="latitude" pageId="3" pageNumber="118" precision="925" value="1.5166667">N 0 91</geoCoordinate>
<geoCoordinate id="EE6C5009FFB9FF87FF50FD3EFE94FD3F" box="[220,311,646,674]" direction="west" orientation="longitude" pageId="3" pageNumber="118" precision="55" value="-36.059">o 36.059</geoCoordinate>
E;
<collectorName id="26AD5318FFB9FF87FED2FD32FE77FD3F" box="[350,468,649,674]" pageId="3" pageNumber="118">R.J. Thoni</collectorName>
&amp;
<collectorName id="26AD5318FFB9FF87FE7BFD32FD2FFD3F" box="[503,652,650,674]" pageId="3" pageNumber="118">D.B. Gurung</collectorName>
,
<collectingDate id="EFA2E9E6FFB9FF87FD1BFD31FCDFFD3F" box="[663,892,649,674]" pageId="3" pageNumber="118" value="2013-09-28">
<date id="FFE6100EFFB9FF87FD1BFD31FCDFFD3F" box="[663,892,649,674]" pageId="3" pageNumber="118" value="2013-09-28">September 28, 2013</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE736CEFFB9FF87FF4BFD15FCCCFD5B" blockId="3.[151,1436,607,962]" box="[199,879,685,710]" pageId="3" pageNumber="118">
<materialsCitation id="3B303C93FFB9FF87FF4BFD15FCCCFD5B" ID-GBIF-Occurrence="1313779487" box="[199,879,685,710]" collectingDate="2013-09-28" collectionCode="CNR" collectorName="R. J. Thoni &amp; D. B. Gurung" country="Bhutan" latitude="1.5166667" location="Dangmechhu River" longLatPrecision="927" longitude="-36.059" municipality="Marangdud" pageId="3" pageNumber="118" specimenCode="CNR 15185, 63.7" specimenCount="1" stateProvince="Zhemgang" typeStatus="paratype">
<emphasis id="B92CEADCFFB9FF87FF4BFD15FE98FD5B" bold="true" box="[199,315,685,710]" pageId="3" pageNumber="118">
<typeStatus id="54E3886CFFB9FF87FF4BFD15FE95FD5B" box="[199,310,685,710]" pageId="3" pageNumber="118" type="paratype">Paratype</typeStatus>
.
</emphasis>
<specimenCode id="DBFE9EB5FFB9FF87FECEFD16FDA4FD5B" box="[322,519,685,710]" pageId="3" pageNumber="118">
<collectionCode id="ED49AE0BFFB9FF87FECEFD16FEDFFD5B" box="[322,380,686,710]" country="Ukraine" httpUri="http://grbio.org/cool/5zp2-7vs1" name="Crimean Natural Reserve" pageId="3" pageNumber="118">CNR</collectionCode>
15185,
<quantity id="4CA09B2BFFB9FF87FE5AFD15FDA4FD5B" box="[470,519,685,710]" metricMagnitude="-2" metricUnit="m" metricValue="6.37" pageId="3" pageNumber="118" unit="mm" value="63.7">63.7</quantity>
</specimenCode>
mm SL; same data as holotype.
</materialsCitation>
</paragraph>
<paragraph id="8BE736CEFFB9FF87FF4BFD6AFC93FC93" blockId="3.[151,1436,607,962]" pageId="3" pageNumber="118">
<materialsCitation id="3B303C93FFB9FF87FF4BFD6AFC93FC93" ID-GBIF-Occurrence="1313779481" collectingDate="1995-10-07" collectionCode="CAS" collectorName="Sherichhu, T. R." country="Bhutan" location="Dangmechhu River" pageId="3" pageNumber="118" specimenCode="CAS 239485, 103.4" specimenCount="1" stateProvince="Trashigang">
<emphasis id="B92CEADCFFB9FF87FF4BFD6AFE3EFD77" box="[199,413,721,746]" italics="true" pageId="3" pageNumber="118">Non-type material.</emphasis>
<specimenCode id="DBFE9EB5FFB9FF87FE2AFD6AFD2AFD77" box="[422,649,721,746]" pageId="3" pageNumber="118">
<collectionCode id="ED49AE0BFFB9FF87FE2AFD6AFE7FFD77" box="[422,476,722,746]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="3" pageNumber="118">CAS</collectionCode>
<quantity id="4CA09B2BFFB9FF87FE69FD69FD2AFD77" box="[485,649,721,746]" metricMagnitude="5" metricUnit="m" metricValue="2.394851034" pageId="3" pageNumber="118" unit="mm" value="2.394851034E8">239485, 103.4</quantity>
</specimenCode>
mm SL;
<collectingCountry id="F34F765EFFB9FF87FD75FD69FCEDFD77" box="[761,846,721,746]" name="Bhutan" pageId="3" pageNumber="118">Bhutan</collectingCountry>
:
<collectingRegion id="499CF82CFFB9FF87FCD7FD69FC7BFD77" box="[859,984,721,746]" country="Bhutan" name="Tashigang" pageId="3" pageNumber="118">Trashigang</collectingRegion>
Dzongkhag:
<location id="8E876015FFB9FF87FBFFFD69FAF3FD77" LSID="urn:lsid:plazi:treatment:03F187D8FFB9FF83FF1BFE4DFC3BFAC9:8E876015FFB9FF87FBFFFD69FAF3FD77" box="[1139,1360,721,746]" country="Bhutan" name="Dangmechhu River" pageId="3" pageNumber="118" stateProvince="Trashigang">Dangmechhu River</location>
above confluence with
<collectorName id="26AD5318FFB9FF87FEDDFD4DFDABFC90" box="[337,520,757,782]" pageId="3" pageNumber="118">Sherichhu, T. R.</collectorName>
Roberts,
<collectingDate id="EFA2E9E6FFB9FF87FDF8FD4DFC8FFC93" box="[628,812,757,782]" pageId="3" pageNumber="118" value="1995-10-07">
<date id="FFE6100EFFB9FF87FDF8FD4DFC8FFC93" box="[628,812,757,782]" pageId="3" pageNumber="118" value="1995-10-07">October 7, 1995</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3426545FFB9FF87FF4BFCA1FD0DFC5F" pageId="3" pageNumber="118" type="diagnosis">
<paragraph id="8BE736CEFFB9FF87FF4BFCA1FD0DFC5F" blockId="3.[151,1436,607,962]" pageId="3" pageNumber="118">
<emphasis id="B92CEADCFFB9FF87FF4BFCA1FEE2FCAF" bold="true" box="[199,321,793,818]" pageId="3" pageNumber="118">Diagnosis.</emphasis>
<taxonomicName id="4C584D4DFFB9FF87FECBFCA2FDE2FCAF" box="[327,577,793,818]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="118" phylum="Chordata" rank="species" species="bimaculacauda" status="sp. nov.">
<emphasis id="B92CEADCFFB9FF87FECBFCA2FDE2FCAF" box="[327,577,793,818]" italics="true" pageId="3" pageNumber="118">Garra bimaculacauda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F57A7FFB9FF87FDCBFCA1FD02FCAF" box="[583,673,793,818]" pageId="3" pageNumber="118" rank="species">
<emphasis id="B92CEADCFFB9FF87FDCBFCA1FD02FCAF" bold="true" box="[583,673,793,818]" pageId="3" pageNumber="118">sp. nov.</emphasis>
</taxonomicNameLabel>
is distinguished from its congeners by the following combination of characters: two distinct black spots on the caudal fin, one on the tip of the dorsal lobe and the other on the tip of the ventral lobe, weak proboscis forming a tuberculate ridge, 32–34 lateral-line scale rows, 12 circumpeduncular-scale rows, 11–12 predorsal-scale rows, six scales from dorsal origin to lateral line, three anal scales, and the absence of a dark lateral band along the length of the body.
</paragraph>
</subSubSection>
<caption id="DF276646FFB9FF87FF1BF813FDEBF87D" httpUri="https://zenodo.org/record/265171/files/figure.png" pageId="3" pageNumber="118" targetBox="[186,1374,1009,1922]" targetPageId="3">
<paragraph id="8BE736CEFFB9FF87FF1BF813FDEBF87D" blockId="3.[151,1436,1963,2016]" pageId="3" pageNumber="118">
<emphasis id="B92CEADCFFB9FF87FF1BF813FEB1F85D" bold="true" box="[151,274,1963,1985]" pageId="3" pageNumber="118">FIGURE 3.</emphasis>
<taxonomicName id="4C584D4DFFB9FF87FE95F814FE59F85C" box="[281,506,1963,1985]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="118" phylum="Chordata" rank="species" species="bimaculacauda">
<emphasis id="B92CEADCFFB9FF87FE95F814FE59F85C" box="[281,506,1963,1985]" italics="true" pageId="3" pageNumber="118">Garra bimaculacauda</emphasis>
</taxonomicName>
. Holotype. NBC 15184, 66.2 mm SL; Bhutan: Zhemgang Dzongkhag, near Panbang Town, R.J. Thoni &amp; D.B. Gurung, October, 2013.
</paragraph>
</caption>
<caption id="DF276646FFBEFF80FF1BFF20FCC8FF32" ID-Table-UUID="DF276646FFBEFF80FF1BFF20FCC8FF32" box="[151,875,151,175]" httpUri="http://table.plazi.org/id/DF276646FFBEFF80FF1BFF20FCC8FF32" pageId="4" pageNumber="119" targetBox="[159,1331,202,1836]" targetIsTable="true" targetPageId="4">
<paragraph id="8BE736CEFFBEFF80FF1BFF20FCC8FF32" blockId="4.[151,875,151,175]" box="[151,875,151,175]" pageId="4" pageNumber="119">
<emphasis id="B92CEADCFFBEFF80FF1BFF20FEA9FF32" bold="true" box="[151,266,152,175]" pageId="4" pageNumber="119">TABLE 1.</emphasis>
Morphometric measurements for
<taxonomicName id="4C584D4DFFBEFF80FDFAFF20FCC0FF32" box="[630,867,151,175]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="119" phylum="Chordata" rank="species" species="bimaculacauda">
<emphasis id="B92CEADCFFBEFF80FDFAFF20FCC0FF32" box="[630,867,151,175]" italics="true" pageId="4" pageNumber="119">Garra bimaculacauda</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BE736CEFFBEFF80FF13FF72FA84F8B1" pageId="4" pageNumber="119">
<table id="F958C46EFFBE007BFF13FF72FA90F8B1" box="[159,1331,202,1836]" gridcols="5" gridrows="40" pageId="4" pageNumber="119">
<tr id="3568348CFFBE007BFF13FF72FA90FF7D" box="[159,1331,202,224]" gridrow="0" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FF72FE6CFF7D" box="[159,463,202,224]" gridcol="0" gridrow="0" pageId="4" pageNumber="119">Morphometrics</th>
<th id="76B95DF0FFBE007BFCA5FF72FCF6FF7D" box="[809,853,202,224]" gridcol="1" gridrow="0" pageId="4" pageNumber="119">Min</th>
<th id="76B95DF0FFBE007BFC45FF72FBA1FF7D" box="[969,1026,202,224]" gridcol="2" gridrow="0" pageId="4" pageNumber="119">Max</th>
<th id="76B95DF0FFBE007BFBE4FF72FB01FF7D" box="[1128,1186,202,224]" gridcol="3" gridrow="0" pageId="4" pageNumber="119">Mean</th>
<th id="76B95DF0FFBE007BFA8BFF72FA90FF7D" box="[1287,1331,202,224]" gridcol="4" gridrow="0" pageId="4" pageNumber="119">SD</th>
</tr>
<tr id="3568348CFFBE007BFF13FF4CFA90FE97" box="[159,1331,244,266]" gridrow="1" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FF4CFE6CFE97" box="[159,463,244,266]" gridcol="0" gridrow="1" pageId="4" pageNumber="119">Standard length</th>
<td id="76B95DF0FFBE007BFCA5FF4CFCF6FE97" box="[809,853,244,266]" gridcol="1" gridrow="1" pageId="4" pageNumber="119">63.7</td>
<td id="76B95DF0FFBE007BFC45FF4CFBA1FE97" box="[969,1026,244,266]" gridcol="2" gridrow="1" pageId="4" pageNumber="119">103.4</td>
<td id="76B95DF0FFBE007BFBE4FF4CFB01FE97" box="[1128,1186,244,266]" gridcol="3" gridrow="1" pageId="4" pageNumber="119">77.8</td>
<td id="76B95DF0FFBE007BFA8BFF4CFA90FE97" box="[1287,1331,244,266]" gridcol="4" gridrow="1" pageId="4" pageNumber="119">22.2</td>
</tr>
<tr id="3568348CFFBE007BFF13FEA5FA90FEAE" box="[159,1331,285,307]" gridrow="2" pageId="4" pageNumber="119" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76B95DF0FFBE007BFF13FEA5FE6CFEAE" box="[159,463,285,307]" gridcol="0" gridrow="2" pageId="4" pageNumber="119">% Standard Length</th>
</tr>
<tr id="3568348CFFBE007BFF13FEFEFA90FEC1" box="[159,1331,326,348]" gridrow="3" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FEFEFE6CFEC1" box="[159,463,326,348]" gridcol="0" gridrow="3" pageId="4" pageNumber="119">Body depth</th>
<td id="76B95DF0FFBE007BFCA5FEFEFCF6FEC1" box="[809,853,326,348]" gridcol="1" gridrow="3" pageId="4" pageNumber="119">19.2</td>
<td id="76B95DF0FFBE007BFC45FEFEFBA1FEC1" box="[969,1026,326,348]" gridcol="2" gridrow="3" pageId="4" pageNumber="119">20.3</td>
<td id="76B95DF0FFBE007BFBE4FEFEFB01FEC1" box="[1128,1186,326,348]" gridcol="3" gridrow="3" pageId="4" pageNumber="119">19.7</td>
<td id="76B95DF0FFBE007BFA8BFEFEFA90FEC1" box="[1287,1331,326,348]" gridcol="4" gridrow="3" pageId="4" pageNumber="119">0.6</td>
</tr>
<tr id="3568348CFFBE007BFF13FEC8FA90FE1B" box="[159,1331,368,390]" gridrow="4" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FEC8FE6CFE1B" box="[159,463,368,390]" gridcol="0" gridrow="4" pageId="4" pageNumber="119">Head length</th>
<td id="76B95DF0FFBE007BFCA5FEC8FCF6FE1B" box="[809,853,368,390]" gridcol="1" gridrow="4" pageId="4" pageNumber="119">25.4</td>
<td id="76B95DF0FFBE007BFC45FEC8FBA1FE1B" box="[969,1026,368,390]" gridcol="2" gridrow="4" pageId="4" pageNumber="119">26.5</td>
<td id="76B95DF0FFBE007BFBE4FEC8FB01FE1B" box="[1128,1186,368,390]" gridcol="3" gridrow="4" pageId="4" pageNumber="119">25.8</td>
<td id="76B95DF0FFBE007BFA8BFEC8FA90FE1B" box="[1287,1331,368,390]" gridcol="4" gridrow="4" pageId="4" pageNumber="119">0.7</td>
</tr>
<tr id="3568348CFFBE007BFF13FE21FA90FE32" box="[159,1331,409,431]" gridrow="5" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FE21FE6CFE32" box="[159,463,409,431]" gridcol="0" gridrow="5" pageId="4" pageNumber="119">Body width</th>
<td id="76B95DF0FFBE007BFCA5FE21FCF6FE32" box="[809,853,409,431]" gridcol="1" gridrow="5" pageId="4" pageNumber="119">14.0</td>
<td id="76B95DF0FFBE007BFC45FE21FBA1FE32" box="[969,1026,409,431]" gridcol="2" gridrow="5" pageId="4" pageNumber="119">16.0</td>
<td id="76B95DF0FFBE007BFBE4FE21FB01FE32" box="[1128,1186,409,431]" gridcol="3" gridrow="5" pageId="4" pageNumber="119">14.8</td>
<td id="76B95DF0FFBE007BFA8BFE21FA90FE32" box="[1287,1331,409,431]" gridcol="4" gridrow="5" pageId="4" pageNumber="119">1.0</td>
</tr>
<tr id="3568348CFFBE007BFF13FE7AFA90FE45" box="[159,1331,450,472]" gridrow="6" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FE7AFE6CFE45" box="[159,463,450,472]" gridcol="0" gridrow="6" pageId="4" pageNumber="119">Caudal peduncle width</th>
<td id="76B95DF0FFBE007BFCA5FE7AFCF6FE45" box="[809,853,450,472]" gridcol="1" gridrow="6" pageId="4" pageNumber="119">3.2</td>
<td id="76B95DF0FFBE007BFC45FE7AFBA1FE45" box="[969,1026,450,472]" gridcol="2" gridrow="6" pageId="4" pageNumber="119">4.8</td>
<td id="76B95DF0FFBE007BFBE4FE7AFB01FE45" box="[1128,1186,450,472]" gridcol="3" gridrow="6" pageId="4" pageNumber="119">4.1</td>
<td id="76B95DF0FFBE007BFA8BFE7AFA90FE45" box="[1287,1331,450,472]" gridcol="4" gridrow="6" pageId="4" pageNumber="119">0.8</td>
</tr>
<tr id="3568348CFFBE007BFF13FE54FA90FD9F" box="[159,1331,492,514]" gridrow="7" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FE54FE6CFD9F" box="[159,463,492,514]" gridcol="0" gridrow="7" pageId="4" pageNumber="119">Caudal peduncle length</th>
<td id="76B95DF0FFBE007BFCA5FE54FCF6FD9F" box="[809,853,492,514]" gridcol="1" gridrow="7" pageId="4" pageNumber="119">14.6</td>
<td id="76B95DF0FFBE007BFC45FE54FBA1FD9F" box="[969,1026,492,514]" gridcol="2" gridrow="7" pageId="4" pageNumber="119">15.7</td>
<td id="76B95DF0FFBE007BFBE4FE54FB01FD9F" box="[1128,1186,492,514]" gridcol="3" gridrow="7" pageId="4" pageNumber="119">15.2</td>
<td id="76B95DF0FFBE007BFA8BFE54FA90FD9F" box="[1287,1331,492,514]" gridcol="4" gridrow="7" pageId="4" pageNumber="119">0.5</td>
</tr>
<tr id="3568348CFFBE007BFF13FDADFA90FDB6" box="[159,1331,533,555]" gridrow="8" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FDADFE6CFDB6" box="[159,463,533,555]" gridcol="0" gridrow="8" pageId="4" pageNumber="119">Caudal peduncle depth</th>
<td id="76B95DF0FFBE007BFCA5FDADFCF6FDB6" box="[809,853,533,555]" gridcol="1" gridrow="8" pageId="4" pageNumber="119">10.3</td>
<td id="76B95DF0FFBE007BFC45FDADFBA1FDB6" box="[969,1026,533,555]" gridcol="2" gridrow="8" pageId="4" pageNumber="119">12.2</td>
<td id="76B95DF0FFBE007BFBE4FDADFB01FDB6" box="[1128,1186,533,555]" gridcol="3" gridrow="8" pageId="4" pageNumber="119">11.3</td>
<td id="76B95DF0FFBE007BFA8BFDADFA90FDB6" box="[1287,1331,533,555]" gridcol="4" gridrow="8" pageId="4" pageNumber="119">1.0</td>
</tr>
<tr id="3568348CFFBE007BFF13FD86FA90FDC9" box="[159,1331,574,596]" gridrow="9" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FD86FE6CFDC9" box="[159,463,574,596]" gridcol="0" gridrow="9" pageId="4" pageNumber="119">1st dorsal ray length</th>
<td id="76B95DF0FFBE007BFCA5FD86FCF6FDC9" box="[809,853,574,596]" gridcol="1" gridrow="9" pageId="4" pageNumber="119">22.2</td>
<td id="76B95DF0FFBE007BFC45FD86FBA1FDC9" box="[969,1026,574,596]" gridcol="2" gridrow="9" pageId="4" pageNumber="119">25.7</td>
<td id="76B95DF0FFBE007BFBE4FD86FB01FDC9" box="[1128,1186,574,596]" gridcol="3" gridrow="9" pageId="4" pageNumber="119">23.5</td>
<td id="76B95DF0FFBE007BFA8BFD86FA90FDC9" box="[1287,1331,574,596]" gridcol="4" gridrow="9" pageId="4" pageNumber="119">1.9</td>
</tr>
<tr id="3568348CFFBE007BFF13FDD0FA90FDE3" box="[159,1331,616,638]" gridrow="10" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FDD0FE6CFDE3" box="[159,463,616,638]" gridcol="0" gridrow="10" pageId="4" pageNumber="119">Dorsal-fin length</th>
<td id="76B95DF0FFBE007BFCA5FDD0FCF6FDE3" box="[809,853,616,638]" gridcol="1" gridrow="10" pageId="4" pageNumber="119">26.1</td>
<td id="76B95DF0FFBE007BFC45FDD0FBA1FDE3" box="[969,1026,616,638]" gridcol="2" gridrow="10" pageId="4" pageNumber="119">29.1</td>
<td id="76B95DF0FFBE007BFBE4FDD0FB01FDE3" box="[1128,1186,616,638]" gridcol="3" gridrow="10" pageId="4" pageNumber="119">28.0</td>
<td id="76B95DF0FFBE007BFA8BFDD0FA90FDE3" box="[1287,1331,616,638]" gridcol="4" gridrow="10" pageId="4" pageNumber="119">1.7</td>
</tr>
<tr id="3568348CFFBE007BFF13FD29FA90FD3A" box="[159,1331,657,679]" gridrow="11" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FD29FE6CFD3A" box="[159,463,657,679]" gridcol="0" gridrow="11" pageId="4" pageNumber="119">Dorsal-fin base length</th>
<td id="76B95DF0FFBE007BFCA5FD29FCF6FD3A" box="[809,853,657,679]" gridcol="1" gridrow="11" pageId="4" pageNumber="119">16.4</td>
<td id="76B95DF0FFBE007BFC45FD29FBA1FD3A" box="[969,1026,657,679]" gridcol="2" gridrow="11" pageId="4" pageNumber="119">17.5</td>
<td id="76B95DF0FFBE007BFBE4FD29FB01FD3A" box="[1128,1186,657,679]" gridcol="3" gridrow="11" pageId="4" pageNumber="119">17.1</td>
<td id="76B95DF0FFBE007BFA8BFD29FA90FD3A" box="[1287,1331,657,679]" gridcol="4" gridrow="11" pageId="4" pageNumber="119">0.6</td>
</tr>
<tr id="3568348CFFBE007BFF13FD02FA90FD4D" box="[159,1331,698,720]" gridrow="12" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FD02FE6CFD4D" box="[159,463,698,720]" gridcol="0" gridrow="12" pageId="4" pageNumber="119">Pectoral-fin length</th>
<td id="76B95DF0FFBE007BFCA5FD02FCF6FD4D" box="[809,853,698,720]" gridcol="1" gridrow="12" pageId="4" pageNumber="119">21.8</td>
<td id="76B95DF0FFBE007BFC45FD02FBA1FD4D" box="[969,1026,698,720]" gridcol="2" gridrow="12" pageId="4" pageNumber="119">24.9</td>
<td id="76B95DF0FFBE007BFBE4FD02FB01FD4D" box="[1128,1186,698,720]" gridcol="3" gridrow="12" pageId="4" pageNumber="119">23.5</td>
<td id="76B95DF0FFBE007BFA8BFD02FA90FD4D" box="[1287,1331,698,720]" gridcol="4" gridrow="12" pageId="4" pageNumber="119">1.6</td>
</tr>
<tr id="3568348CFFBE007BFF13FD5CFA90FD67" box="[159,1331,740,762]" gridrow="13" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FD5CFE6CFD67" box="[159,463,740,762]" gridcol="0" gridrow="13" pageId="4" pageNumber="119">Pelvic-fin length</th>
<td id="76B95DF0FFBE007BFCA5FD5CFCF6FD67" box="[809,853,740,762]" gridcol="1" gridrow="13" pageId="4" pageNumber="119">20.4</td>
<td id="76B95DF0FFBE007BFC45FD5CFBA1FD67" box="[969,1026,740,762]" gridcol="2" gridrow="13" pageId="4" pageNumber="119">22.3</td>
<td id="76B95DF0FFBE007BFBE4FD5CFB01FD67" box="[1128,1186,740,762]" gridcol="3" gridrow="13" pageId="4" pageNumber="119">21.6</td>
<td id="76B95DF0FFBE007BFA8BFD5CFA90FD67" box="[1287,1331,740,762]" gridcol="4" gridrow="13" pageId="4" pageNumber="119">1.1</td>
</tr>
<tr id="3568348CFFBE007BFF13FCB5FA90FCBE" box="[159,1331,781,803]" gridrow="14" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FCB5FE6CFCBE" box="[159,463,781,803]" gridcol="0" gridrow="14" pageId="4" pageNumber="119">Anal-fin length</th>
<td id="76B95DF0FFBE007BFCA5FCB5FCF6FCBE" box="[809,853,781,803]" gridcol="1" gridrow="14" pageId="4" pageNumber="119">17.9</td>
<td id="76B95DF0FFBE007BFC45FCB5FBA1FCBE" box="[969,1026,781,803]" gridcol="2" gridrow="14" pageId="4" pageNumber="119">20.4</td>
<td id="76B95DF0FFBE007BFBE4FCB5FB01FCBE" box="[1128,1186,781,803]" gridcol="3" gridrow="14" pageId="4" pageNumber="119">19.4</td>
<td id="76B95DF0FFBE007BFA8BFCB5FA90FCBE" box="[1287,1331,781,803]" gridcol="4" gridrow="14" pageId="4" pageNumber="119">1.3</td>
</tr>
<tr id="3568348CFFBE007BFF13FC8EFA90FCD1" box="[159,1331,822,844]" gridrow="15" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FC8EFE6CFCD1" box="[159,463,822,844]" gridcol="0" gridrow="15" pageId="4" pageNumber="119">Upper caudal-fin lobe length</th>
<td id="76B95DF0FFBE007BFCA5FC8EFCF6FCD1" box="[809,853,822,844]" gridcol="1" gridrow="15" pageId="4" pageNumber="119">27.3</td>
<td id="76B95DF0FFBE007BFC45FC8EFBA1FCD1" box="[969,1026,822,844]" gridcol="2" gridrow="15" pageId="4" pageNumber="119">28.7</td>
<td id="76B95DF0FFBE007BFBE4FC8EFB01FCD1" box="[1128,1186,822,844]" gridcol="3" gridrow="15" pageId="4" pageNumber="119">28.1</td>
<td id="76B95DF0FFBE007BFA8BFC8EFA90FCD1" box="[1287,1331,822,844]" gridcol="4" gridrow="15" pageId="4" pageNumber="119">0.7</td>
</tr>
<tr id="3568348CFFBE007BFF13FCD8FA90FCEB" box="[159,1331,864,886]" gridrow="16" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FCD8FE6CFCEB" box="[159,463,864,886]" gridcol="0" gridrow="16" pageId="4" pageNumber="119">Lower caudal-fin lobe length</th>
<td id="76B95DF0FFBE007BFCA5FCD8FCF6FCEB" box="[809,853,864,886]" gridcol="1" gridrow="16" pageId="4" pageNumber="119">28.2</td>
<td id="76B95DF0FFBE007BFC45FCD8FBA1FCEB" box="[969,1026,864,886]" gridcol="2" gridrow="16" pageId="4" pageNumber="119">29.6</td>
<td id="76B95DF0FFBE007BFBE4FCD8FB01FCEB" box="[1128,1186,864,886]" gridcol="3" gridrow="16" pageId="4" pageNumber="119">28.8</td>
<td id="76B95DF0FFBE007BFA8BFCD8FA90FCEB" box="[1287,1331,864,886]" gridcol="4" gridrow="16" pageId="4" pageNumber="119">0.8</td>
</tr>
<tr id="3568348CFFBE007BFF13FC31FA90FC02" box="[159,1331,905,927]" gridrow="17" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FC31FE6CFC02" box="[159,463,905,927]" gridcol="0" gridrow="17" pageId="4" pageNumber="119">Median caudal-fin rays length</th>
<td id="76B95DF0FFBE007BFCA5FC31FCF6FC02" box="[809,853,905,927]" gridcol="1" gridrow="17" pageId="4" pageNumber="119">14.8</td>
<td id="76B95DF0FFBE007BFC45FC31FBA1FC02" box="[969,1026,905,927]" gridcol="2" gridrow="17" pageId="4" pageNumber="119">17.3</td>
<td id="76B95DF0FFBE007BFBE4FC31FB01FC02" box="[1128,1186,905,927]" gridcol="3" gridrow="17" pageId="4" pageNumber="119">16.0</td>
<td id="76B95DF0FFBE007BFA8BFC31FA90FC02" box="[1287,1331,905,927]" gridcol="4" gridrow="17" pageId="4" pageNumber="119">1.2</td>
</tr>
<tr id="3568348CFFBE007BFF13FC0AFA90FC55" box="[159,1331,946,968]" gridrow="18" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FC0AFE6CFC55" box="[159,463,946,968]" gridcol="0" gridrow="18" pageId="4" pageNumber="119">Preanal length</th>
<td id="76B95DF0FFBE007BFCA5FC0AFCF6FC55" box="[809,853,946,968]" gridcol="1" gridrow="18" pageId="4" pageNumber="119">78.3</td>
<td id="76B95DF0FFBE007BFC45FC0AFBA1FC55" box="[969,1026,946,968]" gridcol="2" gridrow="18" pageId="4" pageNumber="119">79.3</td>
<td id="76B95DF0FFBE007BFBE4FC0AFB01FC55" box="[1128,1186,946,968]" gridcol="3" gridrow="18" pageId="4" pageNumber="119">78.7</td>
<td id="76B95DF0FFBE007BFA8BFC0AFA90FC55" box="[1287,1331,946,968]" gridcol="4" gridrow="18" pageId="4" pageNumber="119">0.5</td>
</tr>
<tr id="3568348CFFBE007BFF13FC64FA90FC6F" box="[159,1331,988,1010]" gridrow="19" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FC64FE6CFC6F" box="[159,463,988,1010]" gridcol="0" gridrow="19" pageId="4" pageNumber="119">Prevent length</th>
<td id="76B95DF0FFBE007BFCA5FC64FCF6FC6F" box="[809,853,988,1010]" gridcol="1" gridrow="19" pageId="4" pageNumber="119">71.8</td>
<td id="76B95DF0FFBE007BFC45FC64FBA1FC6F" box="[969,1026,988,1010]" gridcol="2" gridrow="19" pageId="4" pageNumber="119">72.9</td>
<td id="76B95DF0FFBE007BFBE4FC64FB01FC6F" box="[1128,1186,988,1010]" gridcol="3" gridrow="19" pageId="4" pageNumber="119">72.1</td>
<td id="76B95DF0FFBE007BFA8BFC64FA90FC6F" box="[1287,1331,988,1010]" gridcol="4" gridrow="19" pageId="4" pageNumber="119">0.6</td>
</tr>
<tr id="3568348CFFBE007BFF13FBBDFA90FB86" box="[159,1331,1029,1051]" gridrow="20" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FBBDFE6CFB86" box="[159,463,1029,1051]" gridcol="0" gridrow="20" pageId="4" pageNumber="119">Preventral length</th>
<td id="76B95DF0FFBE007BFCA5FBBDFCF6FB86" box="[809,853,1029,1051]" gridcol="1" gridrow="20" pageId="4" pageNumber="119">52.4</td>
<td id="76B95DF0FFBE007BFC45FBBDFBA1FB86" box="[969,1026,1029,1051]" gridcol="2" gridrow="20" pageId="4" pageNumber="119">53.1</td>
<td id="76B95DF0FFBE007BFBE4FBBDFB01FB86" box="[1128,1186,1029,1051]" gridcol="3" gridrow="20" pageId="4" pageNumber="119">52.7</td>
<td id="76B95DF0FFBE007BFA8BFBBDFA90FB86" box="[1287,1331,1029,1051]" gridcol="4" gridrow="20" pageId="4" pageNumber="119">0.4</td>
</tr>
<tr id="3568348CFFBE007BFF13FB96FA90FBD9" box="[159,1331,1070,1092]" gridrow="21" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FB96FE6CFBD9" box="[159,463,1070,1092]" gridcol="0" gridrow="21" pageId="4" pageNumber="119">Predorsal length</th>
<td id="76B95DF0FFBE007BFCA5FB96FCF6FBD9" box="[809,853,1070,1092]" gridcol="1" gridrow="21" pageId="4" pageNumber="119">47.1</td>
<td id="76B95DF0FFBE007BFC45FB96FBA1FBD9" box="[969,1026,1070,1092]" gridcol="2" gridrow="21" pageId="4" pageNumber="119">48.5</td>
<td id="76B95DF0FFBE007BFBE4FB96FB01FBD9" box="[1128,1186,1070,1092]" gridcol="3" gridrow="21" pageId="4" pageNumber="119">47.7</td>
<td id="76B95DF0FFBE007BFA8BFB96FA90FBD9" box="[1287,1331,1070,1092]" gridcol="4" gridrow="21" pageId="4" pageNumber="119">0.7</td>
</tr>
<tr id="3568348CFFBE007BFF13FBE0FA90FBF3" box="[159,1331,1112,1134]" gridrow="22" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FBE0FE6CFBF3" box="[159,463,1112,1134]" gridcol="0" gridrow="22" pageId="4" pageNumber="119">Occiput-dorsal length</th>
<td id="76B95DF0FFBE007BFCA5FBE0FCF6FBF3" box="[809,853,1112,1134]" gridcol="1" gridrow="22" pageId="4" pageNumber="119">23.1</td>
<td id="76B95DF0FFBE007BFC45FBE0FBA1FBF3" box="[969,1026,1112,1134]" gridcol="2" gridrow="22" pageId="4" pageNumber="119">25.5</td>
<td id="76B95DF0FFBE007BFBE4FBE0FB01FBF3" box="[1128,1186,1112,1134]" gridcol="3" gridrow="22" pageId="4" pageNumber="119">24.1</td>
<td id="76B95DF0FFBE007BFA8BFBE0FA90FBF3" box="[1287,1331,1112,1134]" gridcol="4" gridrow="22" pageId="4" pageNumber="119">1.3</td>
</tr>
<tr id="3568348CFFBE007BFF13FB39FA90FB0A" box="[159,1331,1153,1175]" gridrow="23" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FB39FE6CFB0A" box="[159,463,1153,1175]" gridcol="0" gridrow="23" pageId="4" pageNumber="119">Prepectoral length</th>
<td id="76B95DF0FFBE007BFCA5FB39FCF6FB0A" box="[809,853,1153,1175]" gridcol="1" gridrow="23" pageId="4" pageNumber="119">22.9</td>
<td id="76B95DF0FFBE007BFC45FB39FBA1FB0A" box="[969,1026,1153,1175]" gridcol="2" gridrow="23" pageId="4" pageNumber="119">24.5</td>
<td id="76B95DF0FFBE007BFBE4FB39FB01FB0A" box="[1128,1186,1153,1175]" gridcol="3" gridrow="23" pageId="4" pageNumber="119">23.5</td>
<td id="76B95DF0FFBE007BFA8BFB39FA90FB0A" box="[1287,1331,1153,1175]" gridcol="4" gridrow="23" pageId="4" pageNumber="119">0.9</td>
</tr>
<tr id="3568348CFFBE007BFF13FB12FA90FB5D" box="[159,1331,1194,1216]" gridrow="24" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FB12FE6CFB5D" box="[159,463,1194,1216]" gridcol="0" gridrow="24" pageId="4" pageNumber="119">Ventral-anal length</th>
<td id="76B95DF0FFBE007BFCA5FB12FCF6FB5D" box="[809,853,1194,1216]" gridcol="1" gridrow="24" pageId="4" pageNumber="119">26.3</td>
<td id="76B95DF0FFBE007BFC45FB12FBA1FB5D" box="[969,1026,1194,1216]" gridcol="2" gridrow="24" pageId="4" pageNumber="119">27.1</td>
<td id="76B95DF0FFBE007BFBE4FB12FB01FB5D" box="[1128,1186,1194,1216]" gridcol="3" gridrow="24" pageId="4" pageNumber="119">26.7</td>
<td id="76B95DF0FFBE007BFA8BFB12FA90FB5D" box="[1287,1331,1194,1216]" gridcol="4" gridrow="24" pageId="4" pageNumber="119">0.4</td>
</tr>
<tr id="3568348CFFBE007BFF13FB6CFA90FB77" box="[159,1331,1236,1258]" gridrow="25" pageId="4" pageNumber="119" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76B95DF0FFBE007BFF13FB6CFE6CFB77" box="[159,463,1236,1258]" gridcol="0" gridrow="25" pageId="4" pageNumber="119">% Ventral-anal</th>
</tr>
<tr id="3568348CFFBE007BFF13FB45FA90FA8E" box="[159,1331,1277,1299]" gridrow="26" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FB45FE6CFA8E" box="[159,463,1277,1299]" gridcol="0" gridrow="26" pageId="4" pageNumber="119">Vent-anal length</th>
<td id="76B95DF0FFBE007BFCA5FB45FCF6FA8E" box="[809,853,1277,1299]" gridcol="1" gridrow="26" pageId="4" pageNumber="119">22.7</td>
<td id="76B95DF0FFBE007BFC45FB45FBA1FA8E" box="[969,1026,1277,1299]" gridcol="2" gridrow="26" pageId="4" pageNumber="119">24.9</td>
<td id="76B95DF0FFBE007BFBE4FB45FB01FA8E" box="[1128,1186,1277,1299]" gridcol="3" gridrow="26" pageId="4" pageNumber="119">23.7</td>
<td id="76B95DF0FFBE007BFA8BFB45FA90FA8E" box="[1287,1331,1277,1299]" gridcol="4" gridrow="26" pageId="4" pageNumber="119">1.1</td>
</tr>
<tr id="3568348CFFBE007BFF13FA9EFA90FAA1" box="[159,1331,1318,1340]" gridrow="27" pageId="4" pageNumber="119" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76B95DF0FFBE007BFF13FA9EFE6CFAA1" box="[159,463,1318,1340]" gridcol="0" gridrow="27" pageId="4" pageNumber="119">% Heal length</th>
</tr>
<tr id="3568348CFFBE007BFF13FAE8FA90FAFB" box="[159,1331,1360,1382]" gridrow="28" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FAE8FE6CFAFB" box="[159,463,1360,1382]" gridcol="0" gridrow="28" pageId="4" pageNumber="119">Head depth</th>
<td id="76B95DF0FFBE007BFCA5FAE8FCF6FAFB" box="[809,853,1360,1382]" gridcol="1" gridrow="28" pageId="4" pageNumber="119">53.9</td>
<td id="76B95DF0FFBE007BFC45FAE8FBA1FAFB" box="[969,1026,1360,1382]" gridcol="2" gridrow="28" pageId="4" pageNumber="119">58.5</td>
<td id="76B95DF0FFBE007BFBE4FAE8FB01FAFB" box="[1128,1186,1360,1382]" gridcol="3" gridrow="28" pageId="4" pageNumber="119">56.9</td>
<td id="76B95DF0FFBE007BFA8BFAE8FA90FAFB" box="[1287,1331,1360,1382]" gridcol="4" gridrow="28" pageId="4" pageNumber="119">2.6</td>
</tr>
<tr id="3568348CFFBE007BFF13FAC1FA90FA12" box="[159,1331,1401,1423]" gridrow="29" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FAC1FE6CFA12" box="[159,463,1401,1423]" gridcol="0" gridrow="29" pageId="4" pageNumber="119">Head depth at eye</th>
<td id="76B95DF0FFBE007BFCA5FAC1FCF6FA12" box="[809,853,1401,1423]" gridcol="1" gridrow="29" pageId="4" pageNumber="119">46.3</td>
<td id="76B95DF0FFBE007BFC45FAC1FBA1FA12" box="[969,1026,1401,1423]" gridcol="2" gridrow="29" pageId="4" pageNumber="119">51.4</td>
<td id="76B95DF0FFBE007BFBE4FAC1FB01FA12" box="[1128,1186,1401,1423]" gridcol="3" gridrow="29" pageId="4" pageNumber="119">48.5</td>
<td id="76B95DF0FFBE007BFA8BFAC1FA90FA12" box="[1287,1331,1401,1423]" gridcol="4" gridrow="29" pageId="4" pageNumber="119">2.6</td>
</tr>
<tr id="3568348CFFBE007BFF13FA1AFA90FA25" box="[159,1331,1442,1464]" gridrow="30" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FA1AFE6CFA25" box="[159,463,1442,1464]" gridcol="0" gridrow="30" pageId="4" pageNumber="119">Head width at opercle</th>
<td id="76B95DF0FFBE007BFCA5FA1AFCF6FA25" box="[809,853,1442,1464]" gridcol="1" gridrow="30" pageId="4" pageNumber="119">65.2</td>
<td id="76B95DF0FFBE007BFC45FA1AFBA1FA25" box="[969,1026,1442,1464]" gridcol="2" gridrow="30" pageId="4" pageNumber="119">70.2</td>
<td id="76B95DF0FFBE007BFBE4FA1AFB01FA25" box="[1128,1186,1442,1464]" gridcol="3" gridrow="30" pageId="4" pageNumber="119">67.2</td>
<td id="76B95DF0FFBE007BFA8BFA1AFA90FA25" box="[1287,1331,1442,1464]" gridcol="4" gridrow="30" pageId="4" pageNumber="119">2.6</td>
</tr>
<tr id="3568348CFFBE007BFF13FA74FA90FA7F" box="[159,1331,1484,1506]" gridrow="31" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FA74FE6CFA7F" box="[159,463,1484,1506]" gridcol="0" gridrow="31" pageId="4" pageNumber="119">Snout length</th>
<td id="76B95DF0FFBE007BFCA5FA74FCF6FA7F" box="[809,853,1484,1506]" gridcol="1" gridrow="31" pageId="4" pageNumber="119">46.7</td>
<td id="76B95DF0FFBE007BFC45FA74FBA1FA7F" box="[969,1026,1484,1506]" gridcol="2" gridrow="31" pageId="4" pageNumber="119">53.4</td>
<td id="76B95DF0FFBE007BFBE4FA74FB01FA7F" box="[1128,1186,1484,1506]" gridcol="3" gridrow="31" pageId="4" pageNumber="119">49.4</td>
<td id="76B95DF0FFBE007BFA8BFA74FA90FA7F" box="[1287,1331,1484,1506]" gridcol="4" gridrow="31" pageId="4" pageNumber="119">3.5</td>
</tr>
<tr id="3568348CFFBE007BFF13FA4DFA90F996" box="[159,1331,1525,1547]" gridrow="32" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13FA4DFE6CF996" box="[159,463,1525,1547]" gridcol="0" gridrow="32" pageId="4" pageNumber="119">Orbit</th>
<td id="76B95DF0FFBE007BFCA5FA4DFCF6F996" box="[809,853,1525,1547]" gridcol="1" gridrow="32" pageId="4" pageNumber="119">17.5</td>
<td id="76B95DF0FFBE007BFC45FA4DFBA1F996" box="[969,1026,1525,1547]" gridcol="2" gridrow="32" pageId="4" pageNumber="119">19.7</td>
<td id="76B95DF0FFBE007BFBE4FA4DFB01F996" box="[1128,1186,1525,1547]" gridcol="3" gridrow="32" pageId="4" pageNumber="119">18.6</td>
<td id="76B95DF0FFBE007BFA8BFA4DFA90F996" box="[1287,1331,1525,1547]" gridcol="4" gridrow="32" pageId="4" pageNumber="119">1.1</td>
</tr>
<tr id="3568348CFFBE007BFF13F9A6FA90F9A9" box="[159,1331,1566,1588]" gridrow="33" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13F9A6FE6CF9A9" box="[159,463,1566,1588]" gridcol="0" gridrow="33" pageId="4" pageNumber="119">Interorbital width</th>
<td id="76B95DF0FFBE007BFCA5F9A6FCF6F9A9" box="[809,853,1566,1588]" gridcol="1" gridrow="33" pageId="4" pageNumber="119">36.9</td>
<td id="76B95DF0FFBE007BFC45F9A6FBA1F9A9" box="[969,1026,1566,1588]" gridcol="2" gridrow="33" pageId="4" pageNumber="119">42.6</td>
<td id="76B95DF0FFBE007BFBE4F9A6FB01F9A9" box="[1128,1186,1566,1588]" gridcol="3" gridrow="33" pageId="4" pageNumber="119">39.7</td>
<td id="76B95DF0FFBE007BFA8BF9A6FA90F9A9" box="[1287,1331,1566,1588]" gridcol="4" gridrow="33" pageId="4" pageNumber="119">2.9</td>
</tr>
<tr id="3568348CFFBE007BFF13F9F0FA90F9C3" box="[159,1331,1608,1630]" gridrow="34" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13F9F0FE6CF9C3" box="[159,463,1608,1630]" gridcol="0" gridrow="34" pageId="4" pageNumber="119">Disc length</th>
<td id="76B95DF0FFBE007BFCA5F9F0FCF6F9C3" box="[809,853,1608,1630]" gridcol="1" gridrow="34" pageId="4" pageNumber="119">40.0</td>
<td id="76B95DF0FFBE007BFC45F9F0FBA1F9C3" box="[969,1026,1608,1630]" gridcol="2" gridrow="34" pageId="4" pageNumber="119">44.1</td>
<td id="76B95DF0FFBE007BFBE4F9F0FB01F9C3" box="[1128,1186,1608,1630]" gridcol="3" gridrow="34" pageId="4" pageNumber="119">42.1</td>
<td id="76B95DF0FFBE007BFA8BF9F0FA90F9C3" box="[1287,1331,1608,1630]" gridcol="4" gridrow="34" pageId="4" pageNumber="119">2.1</td>
</tr>
<tr id="3568348CFFBE007BFF13F9C9FA90F91A" box="[159,1331,1649,1671]" gridrow="35" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13F9C9FE6CF91A" box="[159,463,1649,1671]" gridcol="0" gridrow="35" pageId="4" pageNumber="119">Disc width</th>
<td id="76B95DF0FFBE007BFCA5F9C9FCF6F91A" box="[809,853,1649,1671]" gridcol="1" gridrow="35" pageId="4" pageNumber="119">55.0</td>
<td id="76B95DF0FFBE007BFC45F9C9FBA1F91A" box="[969,1026,1649,1671]" gridcol="2" gridrow="35" pageId="4" pageNumber="119">55.8</td>
<td id="76B95DF0FFBE007BFBE4F9C9FB01F91A" box="[1128,1186,1649,1671]" gridcol="3" gridrow="35" pageId="4" pageNumber="119">55.5</td>
<td id="76B95DF0FFBE007BFA8BF9C9FA90F91A" box="[1287,1331,1649,1671]" gridcol="4" gridrow="35" pageId="4" pageNumber="119">0.5</td>
</tr>
<tr id="3568348CFFBE007BFF13F922FA90F92D" box="[159,1331,1690,1712]" gridrow="36" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13F922FE6CF92D" box="[159,463,1690,1712]" gridcol="0" gridrow="36" pageId="4" pageNumber="119">Callous pad length</th>
<td id="76B95DF0FFBE007BFCA5F922FCF6F92D" box="[809,853,1690,1712]" gridcol="1" gridrow="36" pageId="4" pageNumber="119">25.8</td>
<td id="76B95DF0FFBE007BFC45F922FBA1F92D" box="[969,1026,1690,1712]" gridcol="2" gridrow="36" pageId="4" pageNumber="119">28.8</td>
<td id="76B95DF0FFBE007BFBE4F922FB01F92D" box="[1128,1186,1690,1712]" gridcol="3" gridrow="36" pageId="4" pageNumber="119">27.8</td>
<td id="76B95DF0FFBE007BFA8BF922FA90F92D" box="[1287,1331,1690,1712]" gridcol="4" gridrow="36" pageId="4" pageNumber="119">1.7</td>
</tr>
<tr id="3568348CFFBE007BFF13F97CFA90F947" box="[159,1331,1732,1754]" gridrow="37" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13F97CFE6CF947" box="[159,463,1732,1754]" gridcol="0" gridrow="37" pageId="4" pageNumber="119">Callous pad width</th>
<td id="76B95DF0FFBE007BFCA5F97CFCF6F947" box="[809,853,1732,1754]" gridcol="1" gridrow="37" pageId="4" pageNumber="119">32.6</td>
<td id="76B95DF0FFBE007BFC45F97CFBA1F947" box="[969,1026,1732,1754]" gridcol="2" gridrow="37" pageId="4" pageNumber="119">37.3</td>
<td id="76B95DF0FFBE007BFBE4F97CFB01F947" box="[1128,1186,1732,1754]" gridcol="3" gridrow="37" pageId="4" pageNumber="119">35.5</td>
<td id="76B95DF0FFBE007BFA8BF97CFA90F947" box="[1287,1331,1732,1754]" gridcol="4" gridrow="37" pageId="4" pageNumber="119">2.5</td>
</tr>
<tr id="3568348CFFBE007BFF13F955FA90F89E" box="[159,1331,1773,1795]" gridrow="38" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13F955FE6CF89E" box="[159,463,1773,1795]" gridcol="0" gridrow="38" pageId="4" pageNumber="119">Rostral barbel length</th>
<td id="76B95DF0FFBE007BFCA5F955FCF6F89E" box="[809,853,1773,1795]" gridcol="1" gridrow="38" pageId="4" pageNumber="119">12.1</td>
<td id="76B95DF0FFBE007BFC45F955FBA1F89E" box="[969,1026,1773,1795]" gridcol="2" gridrow="38" pageId="4" pageNumber="119">12.7</td>
<td id="76B95DF0FFBE007BFBE4F955FB01F89E" box="[1128,1186,1773,1795]" gridcol="3" gridrow="38" pageId="4" pageNumber="119">12.5</td>
<td id="76B95DF0FFBE007BFA8BF955FA90F89E" box="[1287,1331,1773,1795]" gridcol="4" gridrow="38" pageId="4" pageNumber="119">0.3</td>
</tr>
<tr id="3568348CFFBE007BFF13F8AEFA90F8B1" box="[159,1331,1814,1836]" gridrow="39" pageId="4" pageNumber="119">
<th id="76B95DF0FFBE007BFF13F8AEFE6CF8B1" box="[159,463,1814,1836]" gridcol="0" gridrow="39" pageId="4" pageNumber="119">Maxillary barbel length</th>
<td id="76B95DF0FFBE007BFCA5F8AEFCF6F8B1" box="[809,853,1814,1836]" gridcol="1" gridrow="39" pageId="4" pageNumber="119">6.1</td>
<td id="76B95DF0FFBE007BFC45F8AEFBA1F8B1" box="[969,1026,1814,1836]" gridcol="2" gridrow="39" pageId="4" pageNumber="119">7.3</td>
<td id="76B95DF0FFBE007BFBE4F8AEFB01F8B1" box="[1128,1186,1814,1836]" gridcol="3" gridrow="39" pageId="4" pageNumber="119">6.6</td>
<td id="76B95DF0FFBE007BFA8BF8AEFA90F8B1" box="[1287,1331,1814,1836]" gridcol="4" gridrow="39" pageId="4" pageNumber="119">0.6</td>
</tr>
</table>
</paragraph>
<subSubSection id="C3426545FFBEFF83FF4BF8D5FC37FE69" lastPageId="7" lastPageNumber="122" pageId="4" pageNumber="119" type="description">
<paragraph id="8BE736CEFFBEFF81FF4BF8D5FCC7FEA8" blockId="4.[151,1436,1901,2026]" lastBlockId="5.[151,1437,151,310]" lastPageId="5" lastPageNumber="120" pageId="4" pageNumber="119">
<emphasis id="B92CEADCFFBEFF80FF4BF8D5FEFAF81B" bold="true" box="[199,345,1901,1926]" pageId="4" pageNumber="119">Description.</emphasis>
Morphometric and meristic counts are provided in
<tableCitation id="C6DA0375FFBEFF80FC32F8D5FB83F81B" box="[958,1056,1901,1926]" captionStart="TABLE 1" captionStartId="4.[151,239,152,175]" captionTargetBox="[159,1331,202,1836]" captionTargetPageId="4" captionText="TABLE 1. Morphometric measurements for Garra bimaculacauda." httpUri="http://table.plazi.org/id/DF276646FFBEFF80FF1BFF20FCC8FF32" pageId="4" pageNumber="119" tableUuid="DF276646FFBEFF80FF1BFF20FCC8FF32">Tables 1</tableCitation>
and
<tableCitation id="C6DA0375FFBEFF80FBD3F8D6FBCCF81B" box="[1119,1135,1902,1926]" captionStart="TABLE 2" captionStartId="5.[151,239,350,373]" captionTargetBox="[159,1143,401,878]" captionTargetPageId="5" captionText="TABLE 2. Meristic count data for Garra bimaculacauda." httpUri="http://table.plazi.org/id/DF276646FFBFFF81FF1BFEE6FCA3FEE8" pageId="4" pageNumber="119" tableUuid="DF276646FFBFFF81FF1BFEE6FCA3FEE8">2</tableCitation>
, respectively. The lateral, dorsal, and ventral view of a preserved specimen can be seen in
<figureCitation id="13632A4BFFBEFF80FCD2F836FC1EF83A" box="[862,957,1934,1959]" captionStart="FIGURE 3" captionStartId="3.[151,250,1963,1985]" captionTargetBox="[186,1374,1009,1922]" captionTargetId="figure@3.[176,1388,994,1937]" captionTargetPageId="3" captionText="FIGURE 3. Garra bimaculacauda. Holotype. NBC 15184, 66.2 mm SL; Bhutan: Zhemgang Dzongkhag, near Panbang Town, R. J. Thoni &amp; D. B. Gurung, October, 2013." httpUri="https://zenodo.org/record/265171/files/figure.png" pageId="4" pageNumber="119">Figure 3</figureCitation>
. The lateral view of a live specimen can be seen in
<figureCitation id="13632A4BFFBEFF80FF61F817FEECF855" box="[237,335,1967,1992]" captionStart="FIGURE 4" captionStartId="5.[151,250,1396,1418]" captionTargetBox="[151,1436,953,1374]" captionTargetId="figure@5.[151,1436,953,1374]" captionTargetPageId="5" captionText="FIGURE 4. Live specimen Garra bimaculacauda. Holotype. NBC 15184, 66.2 mm SL; Bhutan, Zhemgang Dzongkhag, near Panbang, R. J. Thoni &amp; D. B. Gurung, October, 2013." httpUri="https://zenodo.org/record/265172/files/figure.png" pageId="4" pageNumber="119">Figure 4</figureCitation>
. Body elongate. Tip of snout upturned and rough with small tubercles along rostral lobe. Snout to nape interrupted by rostral lobe, transverse groove, and proboscis. Rostral lobe reduced, not prominent or deeply incised. Proboscis not overhanging; roughly 90 degrees in angle to surface of head. From proboscis to occiput, slightly convex. Dorsal profile from occiput to dorsal-fin origin with slight rise. From dorsal-fin origin to caudalfin base, slightly declining with slight concave curvature. Ventral surface from snout to pectoral-fin insertion flat. In line with distal edge of shortest pectoral-fin ray, belly drops slightly until pelvic-fin insertion. From pelvic fins to vent, flat. Vent to caudal-fin base rising with concave curvature.
</paragraph>
<caption id="DF276646FFBFFF81FF1BFEE6FCA3FEE8" ID-Table-UUID="DF276646FFBFFF81FF1BFEE6FCA3FEE8" box="[151,768,349,373]" httpUri="http://table.plazi.org/id/DF276646FFBFFF81FF1BFEE6FCA3FEE8" pageId="5" pageNumber="120" targetBox="[159,1143,401,878]" targetIsTable="true" targetPageId="5">
<paragraph id="8BE736CEFFBFFF81FF1BFEE6FCA3FEE8" blockId="5.[151,768,349,373]" box="[151,768,349,373]" pageId="5" pageNumber="120">
<emphasis id="B92CEADCFFBFFF81FF1BFEE6FEAAFEE8" bold="true" box="[151,265,350,373]" pageId="5" pageNumber="120">TABLE 2.</emphasis>
Meristic count data for
<taxonomicName id="4C584D4DFFBFFF81FD87FEE6FD5FFEE8" box="[523,764,349,373]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="120" phylum="Chordata" rank="species" species="bimaculacauda">Garra bimaculacauda</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BE736CEFFBFFF81FF13FE29FBEAFCF3" pageId="5" pageNumber="120">
<table id="F958C46EFFBF007BFF13FE29FBD4FCF3" box="[159,1143,401,878]" gridcols="3" gridrows="12" pageId="5" pageNumber="120">
<tr id="3568348CFFBF007BFF13FE29FBD4FE3A" box="[159,1143,401,423]" gridrow="0" pageId="5" pageNumber="120">
<th id="76B95DF0FFBF007BFF13FE29FE6EFE3A" box="[159,461,401,423]" gridcol="0" gridrow="0" pageId="5" pageNumber="120">Characters</th>
<th id="76B95DF0FFBF007BFD33FE29FCBEFE3A" box="[703,797,401,423]" gridcol="1" gridrow="0" pageId="5" pageNumber="120">Holotype</th>
<th id="76B95DF0FFBF007BFBB7FE29FBD4FE3A" box="[1083,1143,401,423]" gridcol="2" gridrow="0" pageId="5" pageNumber="120">All</th>
</tr>
<tr id="3568348CFFBF007BFF13FE02FBD4FE4D" box="[159,1143,442,464]" gridrow="1" pageId="5" pageNumber="120" rowspan-1="1" rowspan-2="1">
<th id="76B95DF0FFBF007BFF13FE02FE6EFE4D" box="[159,461,442,464]" gridcol="0" gridrow="1" pageId="5" pageNumber="120">Meristic count</th>
</tr>
<tr id="3568348CFFBF007BFF13FE5CFBD4FE67" box="[159,1143,484,506]" gridrow="2" pageId="5" pageNumber="120">
<th id="76B95DF0FFBF007BFF13FE5CFE6EFE67" box="[159,461,484,506]" gridcol="0" gridrow="2" pageId="5" pageNumber="120">Lateral-line scales</th>
<td id="76B95DF0FFBF007BFD33FE5CFCBEFE67" box="[703,797,484,506]" gridcol="1" gridrow="2" pageId="5" pageNumber="120">32</td>
<td id="76B95DF0FFBF007BFBB7FE5CFBD4FE67" box="[1083,1143,484,506]" gridcol="2" gridrow="2" pageId="5" pageNumber="120">32-34</td>
</tr>
<tr id="3568348CFFBF007BFF13FDB5FBD4FDBE" box="[159,1143,525,547]" gridrow="3" pageId="5" pageNumber="120">
<th id="76B95DF0FFBF007BFF13FDB5FE6EFDBE" box="[159,461,525,547]" gridcol="0" gridrow="3" pageId="5" pageNumber="120">Upper transverse scale rows</th>
<td id="76B95DF0FFBF007BFD33FDB5FCBEFDBE" box="[703,797,525,547]" gridcol="1" gridrow="3" pageId="5" pageNumber="120">6</td>
<td id="76B95DF0FFBF007BFBB7FDB5FBD4FDBE" box="[1083,1143,525,547]" gridcol="2" gridrow="3" pageId="5" pageNumber="120">6</td>
</tr>
<tr id="3568348CFFBF007BFF13FD8EFBD4FDD1" box="[159,1143,566,588]" gridrow="4" pageId="5" pageNumber="120">
<th id="76B95DF0FFBF007BFF13FD8EFE6EFDD1" box="[159,461,566,588]" gridcol="0" gridrow="4" pageId="5" pageNumber="120">Predorsal-scale rows</th>
<td id="76B95DF0FFBF007BFD33FD8EFCBEFDD1" box="[703,797,566,588]" gridcol="1" gridrow="4" pageId="5" pageNumber="120">11</td>
<td id="76B95DF0FFBF007BFBB7FD8EFBD4FDD1" box="[1083,1143,566,588]" gridcol="2" gridrow="4" pageId="5" pageNumber="120">11-12</td>
</tr>
<tr id="3568348CFFBF007BFF13FDD8FBD4FDEB" box="[159,1143,608,630]" gridrow="5" pageId="5" pageNumber="120">
<th id="76B95DF0FFBF007BFF13FDD8FE6EFDEB" box="[159,461,608,630]" gridcol="0" gridrow="5" pageId="5" pageNumber="120">Circumpeduncular-scale rows</th>
<td id="76B95DF0FFBF007BFD33FDD8FCBEFDEB" box="[703,797,608,630]" gridcol="1" gridrow="5" pageId="5" pageNumber="120">12</td>
<td id="76B95DF0FFBF007BFBB7FDD8FBD4FDEB" box="[1083,1143,608,630]" gridcol="2" gridrow="5" pageId="5" pageNumber="120">12</td>
</tr>
<tr id="3568348CFFBF007BFF13FD31FBD4FD02" box="[159,1143,649,671]" gridrow="6" pageId="5" pageNumber="120">
<th id="76B95DF0FFBF007BFF13FD31FE6EFD02" box="[159,461,649,671]" gridcol="0" gridrow="6" pageId="5" pageNumber="120">Branched dorsal-fin rays</th>
<td id="76B95DF0FFBF007BFD33FD31FCBEFD02" box="[703,797,649,671]" gridcol="1" gridrow="6" pageId="5" pageNumber="120">8</td>
<td id="76B95DF0FFBF007BFBB7FD31FBD4FD02" box="[1083,1143,649,671]" gridcol="2" gridrow="6" pageId="5" pageNumber="120">8</td>
</tr>
<tr id="3568348CFFBF007BFF13FD0AFBD4FD55" box="[159,1143,690,712]" gridrow="7" pageId="5" pageNumber="120">
<th id="76B95DF0FFBF007BFF13FD0AFE6EFD55" box="[159,461,690,712]" gridcol="0" gridrow="7" pageId="5" pageNumber="120">Branched anal-fin rays</th>
<td id="76B95DF0FFBF007BFD33FD0AFCBEFD55" box="[703,797,690,712]" gridcol="1" gridrow="7" pageId="5" pageNumber="120">5</td>
<td id="76B95DF0FFBF007BFBB7FD0AFBD4FD55" box="[1083,1143,690,712]" gridcol="2" gridrow="7" pageId="5" pageNumber="120">5</td>
</tr>
<tr id="3568348CFFBF007BFF13FD64FBD4FD6F" box="[159,1143,732,754]" gridrow="8" pageId="5" pageNumber="120">
<th id="76B95DF0FFBF007BFF13FD64FE6EFD6F" box="[159,461,732,754]" gridcol="0" gridrow="8" pageId="5" pageNumber="120">Caudal-fin rays</th>
<td id="76B95DF0FFBF007BFD33FD64FCBEFD6F" box="[703,797,732,754]" gridcol="1" gridrow="8" pageId="5" pageNumber="120">19</td>
<td id="76B95DF0FFBF007BFBB7FD64FBD4FD6F" box="[1083,1143,732,754]" gridcol="2" gridrow="8" pageId="5" pageNumber="120">19</td>
</tr>
<tr id="3568348CFFBF007BFF13FCBDFBD4FC86" box="[159,1143,773,795]" gridrow="9" pageId="5" pageNumber="120">
<th id="76B95DF0FFBF007BFF13FCBDFE6EFC86" box="[159,461,773,795]" gridcol="0" gridrow="9" pageId="5" pageNumber="120">Branched pectoral-fin rays</th>
<td id="76B95DF0FFBF007BFD33FCBDFCBEFC86" box="[703,797,773,795]" gridcol="1" gridrow="9" pageId="5" pageNumber="120">14</td>
<td id="76B95DF0FFBF007BFBB7FCBDFBD4FC86" box="[1083,1143,773,795]" gridcol="2" gridrow="9" pageId="5" pageNumber="120">14</td>
</tr>
<tr id="3568348CFFBF007BFF13FC96FBD4FCD9" box="[159,1143,814,836]" gridrow="10" pageId="5" pageNumber="120">
<th id="76B95DF0FFBF007BFF13FC96FE6EFCD9" box="[159,461,814,836]" gridcol="0" gridrow="10" pageId="5" pageNumber="120">Branched pelvic-fin rays</th>
<td id="76B95DF0FFBF007BFD33FC96FCBEFCD9" box="[703,797,814,836]" gridcol="1" gridrow="10" pageId="5" pageNumber="120">7</td>
<td id="76B95DF0FFBF007BFBB7FC96FBD4FCD9" box="[1083,1143,814,836]" gridcol="2" gridrow="10" pageId="5" pageNumber="120">7</td>
</tr>
<tr id="3568348CFFBF007BFF13FCE0FBD4FCF3" box="[159,1143,856,878]" gridrow="11" pageId="5" pageNumber="120">
<th id="76B95DF0FFBF007BFF13FCE0FE6EFCF3" box="[159,461,856,878]" gridcol="0" gridrow="11" pageId="5" pageNumber="120">Anal scales</th>
<td id="76B95DF0FFBF007BFD33FCE0FCBEFCF3" box="[703,797,856,878]" gridcol="1" gridrow="11" pageId="5" pageNumber="120">3</td>
<td id="76B95DF0FFBF007BFBB7FCE0FBD4FCF3" box="[1083,1143,856,878]" gridcol="2" gridrow="11" pageId="5" pageNumber="120">3</td>
</tr>
</table>
</paragraph>
<caption id="DF276646FFBFFF81FF1BFACCFD0FFA35" httpUri="https://zenodo.org/record/265172/files/figure.png" pageId="5" pageNumber="120" targetBox="[151,1436,953,1374]" targetPageId="5">
<paragraph id="8BE736CEFFBFFF81FF1BFACCFD0FFA35" blockId="5.[151,1435,1396,1448]" pageId="5" pageNumber="120">
<emphasis id="B92CEADCFFBFFF81FF1BFACCFEB0FA14" bold="true" box="[151,275,1396,1418]" pageId="5" pageNumber="120">FIGURE 4.</emphasis>
Live specimen
<taxonomicName id="4C584D4DFFBFFF81FE3AFACCFD3BFA17" box="[438,664,1396,1418]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="120" phylum="Chordata" rank="species" species="bimaculacauda">
<emphasis id="B92CEADCFFBFFF81FE3AFACCFD3BFA17" box="[438,664,1396,1418]" italics="true" pageId="5" pageNumber="120">Garra bimaculacauda</emphasis>
</taxonomicName>
. Holotype. NBC 15184, 66.2 mm SL; Bhutan, Zhemgang Dzongkhag, near Panbang, R.J. Thoni &amp; D.B. Gurung, October, 2013.
</paragraph>
</caption>
<paragraph id="8BE736CEFFBFFF81FF4BFA69FB4CF9E7" blockId="5.[151,1437,1489,2018]" pageId="5" pageNumber="120">Head blunt; depth roughly 5/6 of width. Head length about 1/4 of standard length. Orbit nearly 1/5 of head length. Mouth with large mental disc; slightly wider than long. Disc length slightly less than 1/2 of head length; width slightly more than 1/2 head length. Anterior edge of callus pad with slight fold and covered in sparse papillae. Anterolateral lobe fused with posterolateral flap. Rostral cap short, not completely covering upper jaw. Two pairs of barbels; maxillary barbels roughly one half the length and thickness of rostral barbels.</paragraph>
<paragraph id="8BE736CEFFBFFF81FF4BF93DFD46F8B3" blockId="5.[151,1437,1489,2018]" pageId="5" pageNumber="120">Head ornate with small tubercles occurring in three locations. Rostral lobe with one to two rows of small tubercles. Lateral lobe of snout with one single row of small tubercles running posteriorly from tip of lobe to anterior base on nares. Proboscis with three tubercles on its anterior terminus; two large tubercles laterally and one small tubercle between them. One singular row of small tubercles running posteriorly from laterally-located large tubercle on proboscis to posteromedial base of naris.</paragraph>
<paragraph id="8BE736CEFFBFFF81FF4BF881FDB9F87C" blockId="5.[151,1437,1489,2018]" pageId="5" pageNumber="120">Dorsal-fin origin located roughly halfway between tip of snout and caudal base. First branched ray the longest; nearly as long as HL; margin of fin concave. Dorsal-fin base roughly 2/3 head length; fin rays ii,8. Anal fin nearly 1/5 of standard length; rays i,5. Paired fins inserted horizontally. Pectoral fin roughly 1/5 of standard length; rays i,14. Pelvic fin roughly 1/5 of standard length; fin rays ii,7. Caudal fin forked; lobes roughly equal in length; principal rays 2; branched rays 17.</paragraph>
<caption id="DF276646FFBCFF82FF1BF808FDCEF85B" box="[151,621,1968,1990]" httpUri="https://zenodo.org/record/265173/files/figure.png" pageId="6" pageNumber="121" targetBox="[212,1370,193,1930]" targetPageId="6">
<paragraph id="8BE736CEFFBCFF82FF1BF808FDCEF85B" blockId="6.[151,621,1968,1990]" box="[151,621,1968,1990]" pageId="6" pageNumber="121">
<emphasis id="B92CEADCFFBCFF82FF1BF808FEB1F85B" bold="true" box="[151,274,1968,1990]" pageId="6" pageNumber="121">FIGURE 5.</emphasis>
Habitat of
<taxonomicName id="4C584D4DFFBCFF82FE09F809FDC4F85B" box="[389,615,1968,1990]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="121" phylum="Chordata" rank="species" species="bimaculacauda">
<emphasis id="B92CEADCFFBCFF82FE09F809FDC4F85B" box="[389,615,1968,1990]" italics="true" pageId="6" pageNumber="121">Garra bimaculacauda</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BE736CEFFBDFF83FF4BFF2FFED5FE15" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="122">
<emphasis id="B92CEADCFFBDFF83FF4BFF2FFEECFF2D" bold="true" box="[199,335,151,176]" pageId="7" pageNumber="122">Coloration.</emphasis>
Live specimens captured during this study were silvery grey to olive green. Individuals displaying different color morphs, likely due to sexual dimorphism. Males with olive green base. Sparse scales one to two scale rows above and below lateral line with pale pink color. Females with grey base color; scales on dorsum with dark margin, making dorsal half of body darker grey than ventral half. Several scales above lateral line anterior to dorsal fin have an iridescent blue coloration at the base of the scale. The first scale posterior to upper operculum has the brightest iridescent blue spot on the body. Ventral surface, including mouth, lips and isthmus white. Both pairs of barbels red.
</paragraph>
<paragraph id="8BE736CEFFBDFF83FF4BFE2CFC37FE69" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="122">When preserved in alcohol, the overall pigmentation patterns of specimens same as when if life, with the exception that the iridescent blue spots are absent and overall color appears to be more grey and brown. Barbels without color. A conspicuous dark spot at the caudal base is present.</paragraph>
</subSubSection>
<subSubSection id="C3426545FFBDFF83FF4BFE47FAA5FD85" box="[199,1286,511,536]" pageId="7" pageNumber="122" type="etymology">
<paragraph id="8BE736CEFFBDFF83FF4BFE47FAA5FD85" blockId="7.[151,1437,151,1365]" box="[199,1286,511,536]" pageId="7" pageNumber="122">
<emphasis id="B92CEADCFFBDFF83FF4BFE47FEEDFD85" bold="true" box="[199,334,511,536]" pageId="7" pageNumber="122">Etymology.</emphasis>
The species epithet
<taxonomicName id="4C584D4DFFBDFF83FDBEFE47FD7CFD85" box="[562,735,511,536]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="122" phylum="Chordata" rank="species" species="bimaculacauda">
<emphasis id="B92CEADCFFBDFF83FDBEFE47FD7CFD85" box="[562,735,511,536]" italics="true" pageId="7" pageNumber="122">bimaculacauda</emphasis>
</taxonomicName>
is in reference to the two spots on the caudal fin.
</paragraph>
</subSubSection>
<subSubSection id="C3426545FFBDFF83FF4BFD9CFCDDFDFD" pageId="7" pageNumber="122" type="distribution">
<paragraph id="8BE736CEFFBDFF83FF4BFD9CFCDDFDFD" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="122">
<emphasis id="B92CEADCFFBDFF83FF4BFD9CFEC3FDA0" bold="true" box="[199,352,548,573]" pageId="7" pageNumber="122">Distribution.</emphasis>
Presently known from the Dangmechhu drainage in central and eastern
<collectingCountry id="F34F765EFFBDFF83FB47FD9CFABFFDA0" box="[1227,1308,548,573]" name="Bhutan" pageId="7" pageNumber="122">Bhutan</collectingCountry>
from near Panbang Village to the confluence of the Sherichhu River (
<figureCitation id="13632A4BFFBDFF83FCA7FDFFFCD3FDFD" box="[811,880,583,608]" captionStart="FIGURE 5" captionStartId="6.[151,250,1968,1990]" captionTargetBox="[212,1370,193,1930]" captionTargetId="figure@6.[212,1371,193,1932]" captionTargetPageId="6" captionText="FIGURE 5. Habitat of Garra bimaculacauda." httpUri="https://zenodo.org/record/265173/files/figure.png" pageId="7" pageNumber="122">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3426545FFBDFF83FF4BFDD4FDF8FB58" pageId="7" pageNumber="122" type="discussion">
<paragraph id="8BE736CEFFBDFF83FF4BFDD4FE2AFCA5" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="122">
<emphasis id="B92CEADCFFBDFF83FF4BFDD4FE98FD18" bold="true" box="[199,315,620,645]" pageId="7" pageNumber="122">Remarks.</emphasis>
Upon examining a small lot of Bhutanese fishes collected in 1995 by T. R. Roberts, a single specimen from the Dangmechhu River was found having the same caudal-fin spots, body pigmentation, and tuberculation, as well as a large mental disc, and papillose region on the anterior edge of the callous pad. Meristic values were within range of the two previous specimens as were most mensural data, with some exceptions, most likely due to allometric differences associated with its much larger size. Data from this specimen were taken and added to the analysis.
</paragraph>
<paragraph id="8BE736CEFFBDFF83FF4BFCFCFAAEFBC5" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="122">
Interestingly, a species with similar caudal-fin pigmentation was encountered at the Aquarium Glaser in Frankfurt,
<collectingCountry id="F34F765EFFBDFF83FE83FCD0FEDAFC1D" box="[271,377,872,896]" name="Germany" pageId="7" pageNumber="122">Germany</collectingCountry>
. When enquired of their collection locality, it was said to have collected in
<collectingRegion id="499CF82CFFBDFF83FB45FCD0FAF9FC1D" box="[1225,1370,871,896]" country="India" name="West Bengal" pageId="7" pageNumber="122">West Bengal</collectingRegion>
, most likely in the Buxa Tiger Reserve, which shares a border with
<collectingRegion id="499CF82CFFBDFF83FCEBFC34FC6CFC38" box="[871,975,908,933]" country="Bhutan" name="Chhukha" pageId="7" pageNumber="122">Chhukha</collectingRegion>
, a southern district of
<collectingCountry id="F34F765EFFBDFF83FB5BFC34FA8BFC38" box="[1239,1320,908,933]" name="Bhutan" pageId="7" pageNumber="122">Bhutan</collectingCountry>
(Personal communication with Frank Schäfer). This location, though close to
<collectingCountry id="F34F765EFFBDFF83FC20FC17FBA3FC55" box="[940,1024,943,968]" name="Bhutan" pageId="7" pageNumber="122">Bhutan</collectingCountry>
, is four river drainages west of the locality of
<emphasis id="B92CEADCFFBDFF83FE9FFC6DFE44FC70" box="[275,487,980,1005]" italics="true" pageId="7" pageNumber="122">
<taxonomicName id="4C584D4DFFBDFF83FE9FFC6DFE40FC70" box="[275,483,980,1005]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="122" phylum="Chordata" rank="species" species="bimaculacauda">G. bimaculacauda</taxonomicName>
.
</emphasis>
No fishes bearing these pigmentation patterns were collected in the river drainages between the two localities. The species from
<collectingRegion id="499CF82CFFBDFF83FD2CFC40FC93FB8D" box="[672,816,1015,1040]" country="India" name="West Bengal" pageId="7" pageNumber="122">West Bengal</collectingRegion>
did not have the same head morphology as that of
<taxonomicName id="4C584D4DFFBDFF83FA08FC40FEE7FBA8" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="122" phylum="Chordata" rank="species" species="bimaculacauda">
<emphasis id="B92CEADCFFBDFF83FA08FC40FEE7FBA8" italics="true" pageId="7" pageNumber="122">G. bimaculacauda</emphasis>
</taxonomicName>
as two mature individuals, similar in size to those collected in
<collectingCountry id="F34F765EFFBDFF83FBC4FBA4FB3AFBA8" box="[1096,1177,1052,1077]" name="Bhutan" pageId="7" pageNumber="122">Bhutan</collectingCountry>
had large, protruding proboscises, which were not present in
<taxonomicName id="4C584D4DFFBDFF83FDDCFBF8FCB9FBC5" box="[592,794,1087,1112]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="122" phylum="Chordata" rank="species" species="bimaculacauda">
<emphasis id="B92CEADCFFBDFF83FDDCFBF8FCB9FBC5" box="[592,794,1087,1112]" italics="true" pageId="7" pageNumber="122">G. bimaculacauda</emphasis>
</taxonomicName>
. This species deserves further investigation.
</paragraph>
<paragraph id="8BE736CEFFBDFF83FF4BFBDDFDF8FB58" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="122">
<taxonomicName id="4C584D4DFFBDFF83FF4BFBDDFD86FBE0" authority="Nichols 1925" authorityName="Nichols" authorityYear="1925" box="[199,549,1124,1149]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="122" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="B92CEADCFFBDFF83FF4BFBDDFE22FBE0" box="[199,385,1124,1149]" italics="true" pageId="7" pageNumber="122">Garra orientalis</emphasis>
Nichols 1925
</taxonomicName>
, from
<collectingCountry id="F34F765EFFBDFF83FDFDFBDCFD17FBE0" box="[625,692,1124,1149]" name="China" pageId="7" pageNumber="122">China</collectingCountry>
and
<collectingCountry id="F34F765EFFBDFF83FD63FBDCFCF3FBE0" box="[751,848,1124,1149]" name="Vietnam" pageId="7" pageNumber="122">Vietnam</collectingCountry>
, has caudal-fin markings similar to those described above. However, it also has a medial stripe on the caudal fin and head ornamentation, tubercle arrangement, and morphometric dimensions differ greatly.
</paragraph>
</subSubSection>
<subSubSection id="C3426545FFBDFF83FF4BFB68FC3BFAC9" pageId="7" pageNumber="122" type="description">
<paragraph id="8BE736CEFFBDFF83FF4BFB68FC3BFAC9" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="122">
<taxonomicName id="4C584D4DFFBDFF83FF4BFB68FE6AFB75" box="[199,457,1231,1256]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="122" phylum="Chordata" rank="species" species="bimaculacauda">
<emphasis id="B92CEADCFFBDFF83FF4BFB68FE6AFB75" box="[199,457,1231,1256]" italics="true" pageId="7" pageNumber="122">Garra bimaculacauda</emphasis>
</taxonomicName>
was only encountered in one location by the authors, at the confluence of the Marangdudchhu stream and the Dangmechhu River, in the
<collectingRegion id="499CF82CFFBDFF83FCB8FB4CFC0FFA90" box="[820,940,1268,1293]" country="Bhutan" name="Shemgang" pageId="7" pageNumber="122">Zhemgang</collectingRegion>
Dzongkhag. Both specimens were found in shallow riffles over cobble substrate. Other species collected at this locality include:
<emphasis id="B92CEADCFFBDFF83FB14FAAFFD6FFAC8" italics="true" pageId="7" pageNumber="122">
<taxonomicName id="4C584D4DFFBDFF83FB14FAAFFAE7FAAD" box="[1176,1348,1303,1328]" class="Actinopterygii" family="Cyprinidae" genus="Barilius" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="122" phylum="Chordata" rank="species" species="vagra">Barilius vagra</taxonomicName>
,
<taxonomicName id="4C584D4DFFBDFF83FAD9FAA0FEBBFAC8" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="122" phylum="Chordata" rank="species" species="annandalei">Garra annandalei</taxonomicName>
,
<taxonomicName id="4C584D4DFFBDFF83FEAFFA85FDD7FAC8" authority="Pseudecheneis" authorityName="Pseudecheneis" box="[291,628,1340,1365]" class="Actinopterygii" family="Cyprinidae" genus="Bangana" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="122" phylum="Chordata" rank="species" species="dero">Bangana dero, Pseudecheneis</taxonomicName>
sulcata
</emphasis>
, and
<emphasis id="B92CEADCFFBDFF83FC85FA85FC3BFAC9" box="[777,920,1341,1364]" italics="true" pageId="7" pageNumber="122">
<taxonomicName id="4C584D4DFFBDFF83FC85FA85FC37FAC9" box="[777,916,1341,1364]" class="Actinopterygii" family="Cyprinidae" genus="Tor" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="122" phylum="Chordata" rank="species" species="putitora">Tor putitora</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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