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<mods:title id="A6F338C68E4CFD736F05B52F24747790">The South Temperate Pronophilina (Lepidoptera: Nymphalidae: Satyrinae): a phylogenetic hypothesis, redescriptions and revisionary notes</mods:title>
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<mods:namePart id="AACA0370C2802928B8BBDC71F8562F78">Matz, Jess</mods:namePart>
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<treatment id="03F187D7FFCB843DFF11FD30FBFBBC9C" ID-DOI="http://doi.org/10.5281/zenodo.6070099" ID-GBIF-Taxon="120828512" ID-Zenodo-Dep="6070099" LSID="urn:lsid:plazi:treatment:03F187D7FFCB843DFF11FD30FBFBBC9C" httpUri="http://treatment.plazi.org/id/03F187D7FFCB843DFF11FD30FBFBBC9C" lastPageNumber="65" pageId="64" pageNumber="65">
<subSubSection id="C342654AFFCB843DFF11FD30FD47BA56" pageId="64" pageNumber="65" type="nomenclature">
<paragraph id="8BE736C1FFCB843DFF11FD30FCB0BBB2" blockId="64.[151,873,656,715]" box="[151,873,656,682]" pageId="64" pageNumber="65">
<heading id="D0AF81ADFFCB843DFF11FD30FCB0BBB2" bold="true" box="[151,873,656,682]" fontSize="11" level="1" pageId="64" pageNumber="65" reason="1">
<emphasis id="B92CEAD3FFCB843DFF11FD30FCB0BBB2" bold="true" box="[151,873,656,682]" pageId="64" pageNumber="65">
<taxonomicName id="4C584D42FFCB843DFF11FD30FD28BBB2" authority="(Staudinger, 1898) (Satyrus)" authorityName="Staudinger" authorityYear="1898" baseAuthorityName="Staudinger" baseAuthorityYear="1898" box="[151,753,656,682]" class="Insecta" family="Nymphalidae" genus="Punargentus" kingdom="Animalia" order="Lepidoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="gustavi">
<emphasis id="B92CEAD3FFCB843DFF11FD30FE48BBB2" bold="true" box="[151,401,656,682]" italics="true" pageId="64" pageNumber="65">Punargentus gustavi</emphasis>
(
<bibRefCitation id="EFC94B30FFCB843DFE19FD30FDABBBB2" author="Staudinger" box="[415,626,656,682]" pageId="64" pageNumber="65" refString="Staudinger, O. (1898) Einige neue Tagfalterarten und Varietaten. Deutsche entomologische Zeitschrift &quot; Iris &quot;, 11 (1), 344 - 360." type="journal article" year="1898">Staudinger, 1898</bibRefCitation>
) (
<emphasis id="B92CEAD3FFCB843DFD0AFD30FD3EBBB2" bold="true" box="[652,743,656,682]" italics="true" pageId="64" pageNumber="65">Satyrus</emphasis>
)
</taxonomicName>
, n. comb.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE736C1FFCB843DFF11FD12FE85BBD3" blockId="64.[151,873,656,715]" box="[151,348,690,715]" pageId="64" pageNumber="65">
(
<figureCitation id="13632A44FFCB843DFF19FD12FF21BBD3" box="[159,248,690,715]" captionStart="FIGURE 11" captionStartId="88.[151,250,1652,1674]" captionTargetBox="[288,1298,193,1614]" captionTargetId="figure@88.[287,1300,193,1630]" captionTargetPageId="88" captionText="FIGURE 11. Composite habitus images of Punargentus species. The left side of each specimen is the dorsal surface, the right side is the ventral surface. A: Punargentus lamna (Thieme, 1904) male (MZUJ); B: Punargentus angusta picota (Fuchs, 1954) male, Pampa Galeras, Ayacucho, Peru, 4200 m (photo reproduced from Cerdeña et al. 2014, open access); C: Punargentus blanchardii blanchardii (Pyrcz &amp; Wojtusiak, 2010) n. comb. holotype male (photo reproduced from Pyrcz &amp; Wojtusiak, 2010, open access); D: Punargentus monticolens (Butler, 1881) n. comb. male (MZUJ); E, F: Punargentus gustavi (Staudinger, 1898) n. comb., two specimens from Tacna Prov., Peru showing variability in the degree of silver scaling on the dorsal surface (photos reproduced from Cerdeña et al., 2014, open access); G, H: Punargentus chiliensis chiliensis (Guérin-Méneville, [1830) n. comb. male (G) and female (H) (MZUJ)." httpUri="https://zenodo.org/record/271715/files/figure.png" pageId="64" pageNumber="65">Figs. 11</figureCitation>
E, F; 29)
</paragraph>
<paragraph id="8BE736C1FFCB843DFF11FD5BFDE9BA37" blockId="64.[151,1436,762,1413]" pageId="64" pageNumber="65">
<typeStatus id="54E38863FFCB843DFF11FD5BFED9BA08" box="[151,256,763,784]" pageId="64" pageNumber="65" type="lectotype">Lectotype</typeStatus>
: (male) leg. Gustav Garlepp, MFN (Berlin) (Photo examined)
<typeStatus id="54E38863FFCB843DFF11FCBAFF10BA37" box="[151,201,794,815]" pageId="64" pageNumber="65">Type</typeStatus>
location: Sajama,
<collectingCountry id="F34F7651FFCB843DFE03FCB9FE0BBA37" box="[389,466,793,815]" name="Bolivia" pageId="64" pageNumber="65">Bolivia</collectingCountry>
,
<quantity id="4CA09B24FFCB843DFE5AFCB9FDF5BA37" box="[476,556,793,815]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="64" pageNumber="65" unit="m" value="4000.0">4000 m</quantity>
.
</paragraph>
<paragraph id="8BE736C1FFCB843DFF11FC99FD47BA56" blockId="64.[151,1436,762,1413]" box="[151,670,824,846]" pageId="64" pageNumber="65">
=
<taxonomicName id="4C584D42FFCB843DFF2DFC99FE78BA56" box="[171,417,824,846]" class="Insecta" family="Nymphalidae" genus="Cosmosatyrus" kingdom="Animalia" order="Lepidoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="chiliensis">
<emphasis id="B92CEAD3FFCB843DFF2DFC99FE78BA56" box="[171,417,824,846]" italics="true" pageId="64" pageNumber="65">Cosmosatyrus chiliensis</emphasis>
</taxonomicName>
f.
<emphasis id="B92CEAD3FFCB843DFE3AFC99FDDCBA56" box="[444,517,825,846]" italics="true" pageId="64" pageNumber="65">sajama</emphasis>
<bibRefCitation id="EFC94B30FFCB843DFD8DFC99FD47BA56" author="Weymer" box="[523,670,824,846]" pageId="64" pageNumber="65" refString="Weymer, G. (1911) 4. Familie: Satyridae. In: Seitz, A. (Ed.), Die Gross - Schmetterlinge der Erde. A. Kernen, Stuttgart, pp. 173 - 280." type="book chapter" year="1911">Weymer, 1911</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFCB843DFF41FCF6FDD9BA93" pageId="64" pageNumber="65" type="materials_examined">
<paragraph id="8BE736C1FFCB843DFF41FCF6FE49BA74" blockId="64.[151,1436,762,1413]" box="[199,400,854,876]" pageId="64" pageNumber="65">
<typeStatus id="54E38863FFCB843DFF41FCF6FEF0BA74" box="[199,297,854,876]" pageId="64" pageNumber="65" type="holotype">Holotype</typeStatus>
: unknown
</paragraph>
<paragraph id="8BE736C1FFCB843DFF41FCD6FDD9BA93" blockId="64.[151,1436,762,1413]" box="[199,512,885,907]" pageId="64" pageNumber="65">
<typeStatus id="54E38863FFCB843DFF41FCD6FF20BA93" box="[199,249,886,907]" pageId="64" pageNumber="65">Type</typeStatus>
location: Sajama,
<collectingCountry id="F34F7651FFCB843DFE33FCD5FDD9BA93" box="[437,512,885,907]" name="Bolivia" pageId="64" pageNumber="65">Bolivia</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFCB843DFF11FC35FDD5BAB2" box="[151,524,916,938]" pageId="64" pageNumber="65" type="reference_group">
<paragraph id="8BE736C1FFCB843DFF11FC35FDD5BAB2" blockId="64.[151,1436,762,1413]" box="[151,524,916,938]" pageId="64" pageNumber="65">
=
<taxonomicName id="4C584D42FFCB843DFF2DFC35FDD5BAB2" ID-CoL="4QHMD" authority="Hayward, 1967" authorityName="Hayward" authorityYear="1967" box="[171,524,916,938]" class="Insecta" family="Nymphalidae" genus="Punargentus" kingdom="Animalia" order="Lepidoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="penai">
<emphasis id="B92CEAD3FFCB843DFF2DFC35FEB3BAB2" box="[171,362,917,938]" italics="true" pageId="64" pageNumber="65">Punargentus penai</emphasis>
<bibRefCitation id="EFC94B30FFCB843DFEF7FC34FDD5BAB2" author="Hayward" box="[369,524,916,938]" pageId="64" pageNumber="65" refString="Hayward, K. J. (1967) Tres satiridos neotripicales nuevos (Lep. Rhop. Satyridae). Acta zoologica Lilloana, 21, 19 - 24." type="journal article" year="1967">Hayward, 1967</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFCB843DFF41FC12FD74BD5B" pageId="64" pageNumber="65" type="materials_examined">
<paragraph id="8BE736C1FFCB843DFF41FC12FBD4BAD0" blockId="64.[151,1436,762,1413]" box="[199,1037,946,968]" pageId="64" pageNumber="65">
<typeStatus id="54E38863FFCB843DFF41FC12FEF0BAD0" box="[199,297,946,968]" pageId="64" pageNumber="65" type="holotype">Holotype</typeStatus>
: (male)
<date id="FFE61001FFCB843DFEFDFC12FDDDBAD0" box="[379,516,946,968]" pageId="64" pageNumber="65" value="1965-12-11">11 Dec. 1965</date>
, leg. Luis E. Peña (Peña collection) (not examined)
</paragraph>
<paragraph id="8BE736C1FFCB843DFF41FC72FEBEBD1E" blockId="64.[151,1436,762,1413]" pageId="64" pageNumber="65">
<typeStatus id="54E38863FFCB843DFF41FC72FF20BAFF" box="[199,249,978,999]" pageId="64" pageNumber="65">Type</typeStatus>
location: Alto Pajonales, Antofagasta Province,
<collectingCountry id="F34F7651FFCB843DFD60FC71FCF9BAFF" box="[742,800,977,999]" name="Chile" pageId="64" pageNumber="65">Chile</collectingCountry>
,
<quantity id="4CA09B24FFCB843DFCACFC71FCA3BAFF" box="[810,890,977,999]" metricMagnitude="3" metricUnit="m" metricValue="4.7" pageId="64" pageNumber="65" unit="m" value="4700.0">4700 m</quantity>
. Other combinations:
</paragraph>
<paragraph id="8BE736C1FFCB843DFF11FBAEFD6DBD3C" blockId="64.[151,1436,762,1413]" box="[151,692,1038,1061]" pageId="64" pageNumber="65">
<taxonomicName id="4C584D42FFCB843DFF11FBAEFE2ABD3D" box="[151,499,1038,1061]" class="Insecta" family="Nymphalidae" genus="Argyrophorus" kingdom="Animalia" order="Lepidoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="subSpecies" species="monticolens" subSpecies="gustavi">
<emphasis id="B92CEAD3FFCB843DFF11FBAEFE2ABD3D" box="[151,499,1038,1061]" italics="true" pageId="64" pageNumber="65">Argyrophorus monticolens gustavi</emphasis>
</taxonomicName>
<bibRefCitation id="EFC94B30FFCB843DFD8BFBAEFD6DBD3C" author="Heimlich" box="[525,692,1038,1060]" pageId="64" pageNumber="65" refString="Heimlich, W. (1972) Satyridae der sudlichen Neotropis und Subantarktis (Lepidoptera: Satyridae). Beitrage Entomologische, 22, 149 - 197." type="journal article" year="1972">Heimlich (1972)</bibRefCitation>
</paragraph>
<paragraph id="8BE736C1FFCB843DFF11FB8DFD74BD5B" blockId="64.[151,1436,762,1413]" box="[151,685,1069,1091]" pageId="64" pageNumber="65">
<taxonomicName id="4C584D42FFCB843DFF11FB8DFEB8BD5B" box="[151,353,1069,1091]" class="Insecta" family="Nymphalidae" genus="Argyrophorus" kingdom="Animalia" order="Lepidoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="penai">
<emphasis id="B92CEAD3FFCB843DFF11FB8DFEB8BD5B" box="[151,353,1069,1091]" italics="true" pageId="64" pageNumber="65">Argyrophorus penai</emphasis>
</taxonomicName>
—Peña &amp; Ugarte (1997, p. 275)
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFCB843DFF11FBECFF1ABD90" pageId="64" pageNumber="65" type="distribution">
<paragraph id="8BE736C1FFCB843DFF11FBECFF1ABD90" blockId="64.[151,1436,762,1413]" pageId="64" pageNumber="65">
<emphasis id="B92CEAD3FFCB843DFF11FBECFEE9BD7D" bold="true" box="[151,304,1100,1125]" pageId="64" pageNumber="65">Distribution.</emphasis>
Occurs in the high paramo (&gt;
<quantity id="4CA09B24FFCB843DFD05FBECFD0EBD7C" box="[643,727,1100,1124]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="64" pageNumber="65" unit="m" value="4000.0">4000m</quantity>
.) of southern
<collectingCountry id="F34F7651FFCB843DFCF7FBEDFC7EBD7C" box="[881,935,1101,1124]" name="Peru" pageId="64" pageNumber="65">Peru</collectingCountry>
,
<collectingCountry id="F34F7651FFCB843DFC35FBECFBD1BD7D" box="[947,1032,1100,1125]" name="Bolivia" pageId="64" pageNumber="65">Bolivia</collectingCountry>
, northern
<collectingCountry id="F34F7651FFCB843DFBFDFBECFB61BD7D" box="[1147,1208,1100,1125]" name="Chile" pageId="64" pageNumber="65">Chile</collectingCountry>
and
<collectingCountry id="F34F7651FFCB843DFB69FBECFA87BD7D" box="[1263,1374,1100,1125]" name="Argentina" pageId="64" pageNumber="65">Argentina</collectingCountry>
(
<figureCitation id="13632A44FFCB843DFAE9FBECFF6EBD90" captionStart="FIGURE 29" captionStartId="106.[151,250,1654,1676]" captionText="FIGURE 29. Distribution of locality records for Punargentus species." pageId="64" pageNumber="65">Fig. 29</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFCB843DFF41FB34FE29BC78" pageId="64" pageNumber="65" type="discussion">
<paragraph id="8BE736C1FFCB843DFF41FB34FE29BC78" blockId="64.[151,1436,762,1413]" pageId="64" pageNumber="65">
<emphasis id="B92CEAD3FFCB843DFF41FB34FEE2BDB5" bold="true" box="[199,315,1172,1197]" pageId="64" pageNumber="65">Remarks.</emphasis>
Material was unavailable for detailed study, but the species morphology and distribution and has lately been reviewed by
<bibRefCitation id="EFC94B30FFCB843DFE30FB17FD68BDC8" box="[438,689,1207,1232]" pageId="64" pageNumber="65" refString="Cerdena, J. A., Pyrcz, T. W. &amp; Zacca, T. (2014) High Andean butterflies from southern Peru, I. Dry puna Satyrinae, with the description of two new taxa and three new records from Peru (Lepidoptera: Nymphalidae). Revista Peruana de Biologia, 21, 213 - 222." type="journal article">
Cerdeña
<emphasis id="B92CEAD3FFCB843DFD9BFB19FD80BDC8" box="[541,601,1207,1232]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
(2014)
</bibRefCitation>
. The forewings appear to be more pointed than other species of
<taxonomicName id="4C584D42FFCB843DFF11FB7DFEFCBDEC" box="[151,293,1245,1268]" class="Insecta" family="Nymphalidae" genus="Punargentus" kingdom="Animalia" order="Lepidoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAD3FFCB843DFF11FB7DFEFCBDEC" box="[151,293,1245,1268]" italics="true" pageId="64" pageNumber="65">Punargentus</emphasis>
</taxonomicName>
, with the outer margin evenly curved so that there is hardly an “angle” at the anal angle (
<figureCitation id="13632A44FFCB843DFA88FB7CFA85BDEC" box="[1294,1372,1244,1269]" captionStart="FIGURE 11" captionStartId="88.[151,250,1652,1674]" captionTargetBox="[288,1298,193,1614]" captionTargetId="figure@88.[287,1300,193,1630]" captionTargetPageId="88" captionText="FIGURE 11. Composite habitus images of Punargentus species. The left side of each specimen is the dorsal surface, the right side is the ventral surface. A: Punargentus lamna (Thieme, 1904) male (MZUJ); B: Punargentus angusta picota (Fuchs, 1954) male, Pampa Galeras, Ayacucho, Peru, 4200 m (photo reproduced from Cerdeña et al. 2014, open access); C: Punargentus blanchardii blanchardii (Pyrcz &amp; Wojtusiak, 2010) n. comb. holotype male (photo reproduced from Pyrcz &amp; Wojtusiak, 2010, open access); D: Punargentus monticolens (Butler, 1881) n. comb. male (MZUJ); E, F: Punargentus gustavi (Staudinger, 1898) n. comb., two specimens from Tacna Prov., Peru showing variability in the degree of silver scaling on the dorsal surface (photos reproduced from Cerdeña et al., 2014, open access); G, H: Punargentus chiliensis chiliensis (Guérin-Méneville, [1830) n. comb. male (G) and female (H) (MZUJ)." httpUri="https://zenodo.org/record/271715/files/figure.png" pageId="64" pageNumber="65">Fig. 11</figureCitation>
E, F). The dorsal surface of both wings may exhibit silver scales or be predominantly brown. The unpupillated ocellus between forewing M1-M2 typical of the genus is present, as are the lentuicular submarginal yellow streaks on the ventral surfaces of both wings.
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFCB843DFF41FACCFBFBBC9C" box="[199,1058,1388,1413]" pageId="64" pageNumber="65" type="etymology">
<paragraph id="8BE736C1FFCB843DFF41FACCFBFBBC9C" blockId="64.[151,1436,762,1413]" box="[199,1058,1388,1413]" pageId="64" pageNumber="65">
<emphasis id="B92CEAD3FFCB843DFF41FACCFE9FBC9D" bold="true" box="[199,326,1388,1413]" pageId="64" pageNumber="65">Etymology</emphasis>
. Named for its collector, Gustav Garlepp (Peña &amp; Ugarte, 1997).
</paragraph>
</subSubSection>
</treatment>
</document>