171 lines
20 KiB
XML
171 lines
20 KiB
XML
<document id="311FB53841008D52AAA3B6E5DAF96BF3" ID-CLB-Dataset="7150" ID-DOI="10.11646/zootaxa.4966.1.8" ID-GBIF-Dataset="fa7ec5d2-b715-41a4-8989-57bed5db408b" ID-ISSN="1175-5326" ID-Zenodo-Dep="4729484" ID-ZooBank="BB5BC2CA-6595-4388-9E6C-A24117038F26" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619764087955" checkinUser="plazi" docAuthor="Li, Yan-Da, Tihelka, Erik, Huang, Diying & Cai, Chenyang" docDate="2021" docId="03F187D7FFBEFF4480CDF900C9C9FF3B" docLanguage="en" docName="zootaxa.4966.1.8.pdf" docOrigin="Zootaxa 4966 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Murcybolus longiantennus Li & Tihelka & Huang & Cai 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="80" masterDocId="FFC8FFAFFFBFFF47805AFF8FCB08FFC2" masterDocTitle="Murcybolus gen. nov., a new net-winged beetle genus from mid-Cretaceous Burmese amber (Coleoptera: Lycidae: Burmolycini)" masterLastPageNumber="83" masterPageNumber="77" pageNumber="78" updateTime="1698938077861" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="933034AA52B37A72B5E383735BC138BC">Murcybolus gen. nov., a new net-winged beetle genus from mid-Cretaceous Burmese amber (Coleoptera: Lycidae: Burmolycini)</mods:title>
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<mods:namePart id="B7234589039DEACC955483ADEC0D54F4">Li, Yan-Da</mods:namePart>
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<mods:nameIdentifier id="CC0B427EBBB9CC9463C6614EB31FC3EF" type="ORCID">0000-0002-9439-202X</mods:nameIdentifier>
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<mods:affiliation id="833874D75BF4C4DEC46048B00D89B764">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China & School of Life Sciences, Peking University, Beijing 100871, China & ydli @ pku. edu. cn; https: // orcid. org / 0000 - 0002 - 9439 - 202 X</mods:affiliation>
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<mods:nameIdentifier id="13BE9C40ED7C8719DC7FF32F689BC826" type="email">ydli@pku.edu.cn</mods:nameIdentifier>
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<mods:roleTerm id="D04840F54A7642D34F7FAD7152FFE581">Author</mods:roleTerm>
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<mods:namePart id="915E023B650537EE20BDD528B1BBBDBA">Tihelka, Erik</mods:namePart>
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<mods:nameIdentifier id="D3F205D601C7B193A12C06F6A24173E5" type="ORCID">0000-0002-5048-5355</mods:nameIdentifier>
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<mods:affiliation id="5A6EE4455AF09F088165C8B4FB957200">School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, United Kingdom & wn 20250 @ bristol. ac. uk; https: // orcid. org / 0000 - 0002 - 5048 - 5355</mods:affiliation>
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<mods:nameIdentifier id="F043BDA534D594D63970A112D8C89E22" type="email">wn20250@bristol.ac.uk</mods:nameIdentifier>
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<mods:roleTerm id="CB3D9230AA10C6A597C250B6C8821CAD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1599467F9AF4B2CCBB3D048DB7208757">Huang, Diying</mods:namePart>
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<mods:nameIdentifier id="EDA75385418D5AD365B863F5492431A8" type="ORCID">0000-0002-5637-4867</mods:nameIdentifier>
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<mods:affiliation id="EEA6EA46DA9F382572C08B39DCE0751C">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China & dyhuang @ nigpas. ac. cn; https: // orcid. org / 0000 - 0002 - 5637 - 4867</mods:affiliation>
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<mods:nameIdentifier id="A868E713034D301806B74D9E9D766A8C" type="email">dyhuang@nigpas.ac.cn</mods:nameIdentifier>
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<mods:name id="D7A34305D5581FA9D738AE88EC1B4965" type="personal">
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<mods:namePart id="5FE8C3BAC26B2D2F215D2716F6ABAF8D">Cai, Chenyang</mods:namePart>
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<mods:affiliation id="D78F5CD938DAC72E501889B84582FD0D">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China & School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, United Kingdom & cycai @ nigpas. ac. cn; https: // orcid. org / 0000 - 0002 - 9283 - 8323</mods:affiliation>
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<mods:nameIdentifier id="055551F49FE22F8977E4530C64FD95A0" type="email">cycai@nigpas.ac.cn</mods:nameIdentifier>
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<mods:title id="83BF933D6F63D80B7BAABCE263436CC8">Zootaxa</mods:title>
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<mods:part id="1EA3323E503FF600E0FA60066D8F6C0B">
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<mods:date id="E75D98A3EEE603F9CC6D5FD9BD73737C">2021</mods:date>
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<mods:number id="B55E70D48008F32722E644FEAAB81A9D">2021-04-30</mods:number>
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<mods:number id="5D25658C4644C77693873E70A77C0E14">4966</mods:number>
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<mods:identifier id="A7606D3DA84726D6D31B999E39338651" type="DOI">10.11646/zootaxa.4966.1.8</mods:identifier>
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<treatment id="03F187D7FFBEFF4480CDF900C9C9FF3B" ID-DOI="http://doi.org/10.5281/zenodo.4729481" ID-GBIF-Taxon="181756819" ID-Zenodo-Dep="4729481" LSID="urn:lsid:plazi:treatment:03F187D7FFBEFF4480CDF900C9C9FF3B" httpUri="http://treatment.plazi.org/id/03F187D7FFBEFF4480CDF900C9C9FF3B" lastPageId="3" lastPageNumber="80" pageId="1" pageNumber="78">
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<subSubSection id="C342654AFFBEFF4680CDF900C9B1F968" box="[151,697,1679,1706]" pageId="1" pageNumber="78" type="nomenclature">
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<paragraph id="8BE736C1FFBEFF4680CDF900C9B1F968" blockId="1.[151,697,1679,1742]" box="[151,697,1679,1706]" pageId="1" pageNumber="78">
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<heading id="D0AF81ADFFBEFF4680CDF900C9B1F968" bold="true" box="[151,697,1679,1706]" fontSize="11" level="1" pageId="1" pageNumber="78" reason="1">
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<emphasis id="B92CEAD3FFBEFF4680CDF900C9B1F968" bold="true" box="[151,697,1679,1706]" pageId="1" pageNumber="78">
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<taxonomicName id="4C584D42FFBEFF4680CDF900C95CF968" authority="Li & Cai" authorityName="Li & Tihelka & Huang & Cai" authorityYear="2021" box="[151,596,1679,1706]" class="Insecta" family="Lycidae" genus="Murcybolus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="longiantennus" status="sp. nov.">
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<emphasis id="B92CEAD3FFBEFF4680CDF900CAD6F968" bold="true" box="[151,478,1679,1706]" italics="true" pageId="1" pageNumber="78">Murcybolus longiantennus</emphasis>
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Li & Cai
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</taxonomicName>
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<taxonomicNameLabel id="A21F57A8FFBEFF468206F91FC9B1F968" box="[604,697,1680,1706]" pageId="1" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C342654AFFBEFF4680CDF93BCA03F90C" box="[151,267,1716,1742]" pageId="1" pageNumber="78" type="description">
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<paragraph id="8BE736C1FFBEFF4680CDF93BCA03F90C" blockId="1.[151,697,1679,1742]" box="[151,267,1716,1742]" pageId="1" pageNumber="78">
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(
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<figureCitation id="13632A44FFBEFF4680C5F93BCA0AF90C" box="[159,258,1716,1742]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1657,1681]" captionStartId-1="3.[151,250,1477,1501]" captionStartId-2="4.[151,250,1777,1801]" captionTargetBox-0="[151,1436,450,1630]" captionTargetBox-1="[151,1436,273,1453]" captionTargetBox-2="[211,1376,181,1753]" captionTargetId-0="figure-150@2.[151,1436,450,1630]" captionTargetId-1="figure-42@3.[151,1436,273,1453]" captionTargetId-2="figure-39@4.[211,1376,181,1753]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. General habitus of Murcybolus longiantennus gen. et sp. nov., female, holotype, NIGP173914, under incident light. A, Dorsal view. B, Ventral view. Scale bars: 500 μm." captionText-1="FIGURE 2. General habitus of Murcybolus longiantennus gen. et sp. nov., female, holotype, NIGP173914, under widefield fluorescence. A, Dorsal view. B, Ventral view. Scale bars: 500 μm." captionText-2="FIGURE 3. Details of Murcybolus longiantennus gen. et sp. nov., female, holotype, NIGP173914, under confocal microscopy, with depth colour-coding in A–F. A, Head and prothorax, dorsal view. B, Elytral base. C, elytral apex, showing the interrupted costae 2 and 3 (arrowheads). D, Head and prothorax, ventral view. E, Legs, ventral view. F, Abdominal apex, ventral view. G–H, Same as A–B. I, Anterior portion of abdomen, ventral view. J–L, Same as D–F. Abbreviations: a1–4, antennomeres 1–4; el, elytron; ey, compound eye; lb, labrum; lbp, labial palp; ml, mala; md, mandible; mstb, mesotibia; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mttc, metatrochanter; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; sc, scutellum; v1–7; ventrites 1–7. Scale bars: 200 μm." figureDoi-0="http://doi.org/10.5281/zenodo.4729486" figureDoi-1="http://doi.org/10.5281/zenodo.4729488" figureDoi-2="http://doi.org/10.5281/zenodo.4729490" httpUri-0="https://zenodo.org/record/4729486/files/figure.png" httpUri-1="https://zenodo.org/record/4729488/files/figure.png" httpUri-2="https://zenodo.org/record/4729490/files/figure.png" pageId="1" pageNumber="78">Figs 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C342654AFFBEFF4680CDF973C8FFF8D5" box="[151,1015,1788,1815]" pageId="1" pageNumber="78" type="etymology">
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<paragraph id="8BE736C1FFBEFF4680CDF973C8FFF8D5" blockId="1.[151,1436,1788,2031]" box="[151,1015,1788,1815]" pageId="1" pageNumber="78">
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<emphasis id="B92CEAD3FFBEFF4680CDF973CA16F8D4" bold="true" box="[151,286,1788,1814]" pageId="1" pageNumber="78">Etymology.</emphasis>
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The specific name refers to the beetle’s relatively long antennae.
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</paragraph>
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</subSubSection>
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<subSubSection id="C342654AFFBEFF46809DF8AFCFEAF840" pageId="1" pageNumber="78" type="materials_examined">
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<paragraph id="8BE736C1FFBEFF46809DF8AFC9E2F8F8" blockId="1.[151,1436,1788,2031]" box="[199,746,1824,1850]" pageId="1" pageNumber="78">
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<emphasis id="B92CEAD3FFBEFF46809DF8AFCA3EF8F8" bold="true" box="[199,310,1824,1850]" pageId="1" pageNumber="78">Material.</emphasis>
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<materialsCitation id="3B303C9CFFBEFF468167F8AFC9E2F8F8" ID-GBIF-Occurrence="3094804301" box="[317,746,1824,1850]" collectionCode="NIGP" pageId="1" pageNumber="78" specimenCode="NIGP173914" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54E38863FFBEFF468167F8AFCAA0F8F8" box="[317,424,1824,1850]" pageId="1" pageNumber="78" type="holotype">Holotype</typeStatus>
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,
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<specimenCode id="DBFE9EBAFFBEFF4681EEF8AFC943F8F8" box="[436,587,1824,1850]" collectionCode="NIGP" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="1" pageNumber="78">NIGP173914</specimenCode>
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, adult female.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BE736C1FFBEFF46809DF8CBCFEAF840" blockId="1.[151,1436,1788,2031]" pageId="1" pageNumber="78">
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<emphasis id="B92CEAD3FFBEFF46809DF8CBCAC7F89C" bold="true" box="[199,463,1860,1886]" pageId="1" pageNumber="78">Locality and horizon.</emphasis>
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<materialsCitation id="3B303C9CFFBEFF468180F8CBCFD6F840" ID-GBIF-Occurrence="3119717301" pageId="1" pageNumber="78">
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Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District,
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<collectingRegion id="499CF823FFBEFF4680CDF8E7CA22F840" box="[151,298,1896,1922]" country="Myanmar" name="Kachin" pageId="1" pageNumber="78">Kachin State</collectingRegion>
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,
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<collectingCountry id="F34F7651FFBEFF46816FF8E6CAAEF841" box="[309,422,1897,1923]" name="Myanmar" pageId="1" pageNumber="78">Myanmar</collectingCountry>
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; unnamed horizon, mid-Cretaceous, Upper Albian to Lower Cenomanian
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C342654AFFBEFF46809DF803C900F865" box="[199,520,1932,1959]" pageId="1" pageNumber="78" type="diagnosis">
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<paragraph id="8BE736C1FFBEFF46809DF803C900F865" blockId="1.[151,1436,1788,2031]" box="[199,520,1932,1959]" pageId="1" pageNumber="78">
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<emphasis id="B92CEAD3FFBEFF46809DF803CA4AF864" bold="true" box="[199,322,1932,1958]" pageId="1" pageNumber="78">Diagnosis.</emphasis>
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As for the genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C342654AFFBEFF44809DF83FC9C9FF3B" lastPageId="3" lastPageNumber="80" pageId="1" pageNumber="78" type="description">
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<paragraph id="8BE736C1FFBEFF46809DF83FC813F82C" blockId="1.[151,1436,1788,2031]" pageId="1" pageNumber="78">
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<emphasis id="B92CEAD3FFBEFF46809DF83FC909F808" bold="true" box="[199,513,1968,1994]" pageId="1" pageNumber="78">Description. Adult female.</emphasis>
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Body presumably weakly-sclerotised, slender, almost parallel-sided,
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<quantity id="4CA09B24FFBEFF468557F83FCE6CF809" box="[1293,1380,1968,1995]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="78" unit="mm" value="2.2">2.2 mm</quantity>
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long and
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<quantity id="4CA09B24FFBEFF46809DF85BCA15F82D" box="[199,285,2004,2031]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="78" unit="mm" value="0.6">0.6 mm</quantity>
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wide at humeri, covered with long fine hairs.
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</paragraph>
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<paragraph id="8BE736C1FFBDFF45809DFF18C9A6FE6C" blockId="2.[151,1437,151,430]" pageId="2" pageNumber="79">
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Head including eyes wider than anterior margin of pronotum, hypognathous, without any longitudinal grooves; exposed portion in dorsal view
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<quantity id="4CA09B24FFBDFF4581A4FF34C96BFF17" box="[510,611,187,213]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="79" unit="mm" value="0.21">0.21 mm</quantity>
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long. Compound eyes lateral, large, strongly protuberant, finely faceted, without interfacetal setae. Antennal insertions located frontally, separated by projecting frontal part of cranium. Antennae moderately long, longer than half of body, with 10 antennomeres; antennomere 1 robust, with oblique apical margin; antennomere 2 short, about 0.4 times as long as antennomere 3; antennomeres 3–10 elongate, filiform, slightly asymmetrical. Mouth cavity rounded. Labrum free, located inside mouth cavity. Mandibles simple, gradually curved mesally. Maxillary palps 4-segmented; palpomere 1 short, palpomeres 2–4 filiform. Mala welldeveloped, setose. Labial palpi 3(?)-segmented.
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</paragraph>
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<caption id="DF276649FFBDFF4580CDF9F6C9E2F974" ID-DOI="http://doi.org/10.5281/zenodo.4729486" ID-Zenodo-Dep="4729486" httpUri="https://zenodo.org/record/4729486/files/figure.png" pageId="2" pageNumber="79" startId="2.[151,250,1657,1681]" targetBox="[151,1436,450,1630]" targetPageId="2">
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<paragraph id="8BE736C1FFBDFF4580CDF9F6C9E2F974" blockId="2.[151,1437,1657,1718]" pageId="2" pageNumber="79">
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<emphasis id="B92CEAD3FFBDFF4580CDF9F6CA1EF953" bold="true" box="[151,278,1657,1681]" pageId="2" pageNumber="79">FIGURE 1.</emphasis>
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General habitus of
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<taxonomicName id="4C584D42FFBDFF4581B2F9F6C9F1F953" authority="Li & Tihelka & Huang & Cai, 2021" authorityName="Li & Tihelka & Huang & Cai" authorityYear="2021" box="[488,761,1657,1681]" class="Insecta" family="Lycidae" genus="Murcybolus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="longiantennus" status="gen. et sp. nov.">
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||
<emphasis id="B92CEAD3FFBDFF4581B2F9F6C9F1F953" box="[488,761,1657,1681]" italics="true" pageId="2" pageNumber="79">Murcybolus longiantennus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92CEAD3FFBDFF45835BF9F6C8AEF953" bold="true" box="[769,934,1657,1681]" pageId="2" pageNumber="79">
|
||
<taxonomicNameLabel id="A21F57A8FFBDFF45835BF9F6C8AEF953" box="[769,934,1657,1681]" pageId="2" pageNumber="79" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female, holotype, NIGP173914, under incident light.
|
||
<emphasis id="B92CEAD3FFBDFF458088F912CBECF977" bold="true" box="[210,228,1693,1717]" pageId="2" pageNumber="79">A</emphasis>
|
||
, Dorsal view.
|
||
<emphasis id="B92CEAD3FFBDFF45812DF912CA80F977" bold="true" box="[375,392,1693,1717]" pageId="2" pageNumber="79">B</emphasis>
|
||
, Ventral view. Scale bars: 500 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE736C1FFBDFF45809DF96CC8F4F8AB" blockId="2.[151,1437,1763,2042]" pageId="2" pageNumber="79">
|
||
Pronotal disc narrower than elytra,
|
||
<quantity id="4CA09B24FFBDFF458206F96BC9CAF93C" box="[604,706,1764,1790]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="79" unit="mm" value="0.27">0.27 mm</quantity>
|
||
long, 2.0 times as wide as long, subtrapezoidal; anterior angles obtuse; posterior angles acute, projecting laterally; surface without longitudinal carinae, with irregular deep rounded punctures. Prosternum strongly transverse, short; prosternal process narrow and acute apically. Scutellum well-developed, parallel-sided, with posterior margin emarginate. Metaventrite large.
|
||
</paragraph>
|
||
<paragraph id="8BE736C1FFBDFF44809DF8FCCBE1FF17" blockId="2.[151,1437,1763,2042]" lastBlockId="3.[151,1436,150,249]" lastPageId="3" lastPageNumber="80" pageId="2" pageNumber="79">
|
||
Elytra parallel-sided,
|
||
<quantity id="4CA09B24FFBDFF4581E2F8FBC905F84C" box="[440,525,1908,1934]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="79" unit="mm" value="1.7">1.7 mm</quantity>
|
||
long, about 2.7 times as long as their combined width, completely covering abdomen; apices separately rounded; surface with eight longitudinal costae and nine rows of rounded cells in intercostal intervals; costae 2 and 3 (counted from elytral suture outwards) interrupted in posterior quarter of elytra. Hind winds fully developed. Legs slender; procoxae narrowly separated; trochanters obliquely attached to femora; both femora and tibiae slender; tarsi with five tarsomeres, tarsomeres 1–4 subequal, tarsomere 5 slender, long; pretarsal claws simple.
|
||
</paragraph>
|
||
<paragraph id="8BE736C1FFBCFF44809DFF50C9C9FF3B" blockId="3.[151,1436,150,249]" box="[199,705,223,249]" pageId="3" pageNumber="80">Abdomen with seven free subequal ventrites.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |