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<document id="E02C69DE89930E057B590D539FD82137" ID="10.11646/zootaxa.4125.1.1" ID-CLB-Dataset="38807" ID-DOI="10.11646/zootaxa.4125.1.1" ID-GBIF-Dataset="1af13aec-3c54-4a92-abdf-e4e29b97e82f" ID-ISSN="1175-5326" ID-Zenodo-Dep="271704" ID-ZooBank="118F4865-D89E-45EA-A210-8D61946CC37F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465971881907" checkinUser="plazi" docAuthor="Matz, Jess &amp; Brower, Andrew V. Z." docDate="2016" docId="03F187D7FFBA844FFF11FC80FB24BD2C" docLanguage="en" docName="zootaxa.4125.1.1.pdf" docOrigin="Zootaxa 4125 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Neosatyrus humilis C. Felder &amp; R. Felder 1867" docType="treatment" docVersion="9" lastPageNumber="51" masterDocId="FFC8FFAFFF8B847DFF86FFA0FFD9B918" masterDocTitle="The South Temperate Pronophilina (Lepidoptera: Nymphalidae: Satyrinae): a phylogenetic hypothesis, redescriptions and revisionary notes" masterLastPageNumber="108" masterPageNumber="1" pageNumber="50" updateTime="1698662999642" updateUser="ExternalLinkService">
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<mods:title id="53F0147D0E90607FD02638E5E1825190">The South Temperate Pronophilina (Lepidoptera: Nymphalidae: Satyrinae): a phylogenetic hypothesis, redescriptions and revisionary notes</mods:title>
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<mods:namePart id="DF826B762BC2994784EDCF999F5A5D6F">Matz, Jess</mods:namePart>
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<mods:date id="E68A3162EDC002239C1CF38FCFD701CE">2016</mods:date>
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<treatment id="03F187D7FFBA844FFF11FC80FB24BD2C" ID-DOI="http://doi.org/10.5281/zenodo.6070062" ID-GBIF-Taxon="120828481" ID-Zenodo-Dep="6070062" LSID="urn:lsid:plazi:treatment:03F187D7FFBA844FFF11FC80FB24BD2C" httpUri="http://treatment.plazi.org/id/03F187D7FFBA844FFF11FC80FB24BD2C" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
<subSubSection id="C342654AFFBA844CFF11FC80FD23BAA7" pageId="49" pageNumber="50" type="nomenclature">
<paragraph id="8BE736C1FFBA844CFF11FC80FC06BA22" blockId="49.[151,991,800,859]" box="[151,991,800,826]" pageId="49" pageNumber="50">
<heading id="D0AF81ADFFBA844CFF11FC80FC06BA22" bold="true" box="[151,991,800,826]" fontSize="11" level="1" pageId="49" pageNumber="50" reason="1">
<emphasis id="B92CEAD3FFBA844CFF11FC80FC06BA22" bold="true" box="[151,991,800,826]" pageId="49" pageNumber="50">
<taxonomicName id="4C584D42FFBA844CFF11FC80FCB4BA22" authority="(C. Felder &amp; R. Felder, 1867) (Stygnus)" authorityName="C. Felder &amp; R. Felder" authorityYear="1867" baseAuthorityName="C. Felder &amp; R. Felder" baseAuthorityYear="1867" box="[151,877,800,826]" class="Insecta" family="Nymphalidae" genus="Neosatyrus" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B92CEAD3FFBA844CFF11FC80FE59BA22" bold="true" box="[151,384,800,826]" italics="true" pageId="49" pageNumber="50">Neosatyrus humilis</emphasis>
(
<bibRefCitation id="EFC94B30FFBA844CFE17FC80FD32BA22" author="Felder" box="[401,747,800,826]" pageId="49" pageNumber="50" refString="Felder, C. &amp; Felder, R. (1867) Reise der osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1859, 1859 unter den Befehlen des Commodore B. von Wullensdorf - Urbair. Zoologischer Thiel. Zweiter Band. Zweite Abteilung: Lepidoptera. Carl Gerold's Sohn, Wien, 536 + 9 pp., 74 pls." type="book" year="1867">C. Felder &amp; R. Felder, 1867</bibRefCitation>
) (
<emphasis id="B92CEAD3FFBA844CFC82FC80FCBABA22" bold="true" box="[772,867,800,826]" italics="true" pageId="49" pageNumber="50">Stygnus</emphasis>
)
</taxonomicName>
n. comb.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE736C1FFBA844CFF11FCE2FE72BA43" blockId="49.[151,991,800,859]" box="[151,427,834,859]" pageId="49" pageNumber="50">
(
<figureCitation id="13632A44FFBA844CFF19FCE2FF35BA43" box="[159,236,834,859]" captionStart="FIGURE 8" captionStartId="85.[151,250,1281,1303]" captionTargetBox="[253,1336,200,1247]" captionTargetId="figure@85.[243,1344,193,1260]" captionTargetPageId="85" captionText="FIGURE 8. Composite habitus images of Neosatyrus species. The left side of each specimen is the dorsal surface, the right side is the ventral surface. A: Neosatyrus ambiorix Wallengren, 1858 male holotype (NRM); B: Neosatyrus boisduvalii (Blanchard, 1852) male (MZUJ); C, D: Neosatyrus humilis (C. Felder &amp; R. Felder, 1867) n. comb. male lectotype (BMNH # 809786) (C), male paralectotype (BMNH # 809783) (D); E: Neosatyrus schajovskoii (Hayward, 1954) n. comb. male (MZUJ); F: Neosatyrus vesagus (Doubleday, [1849]), male holotype (BMNH Rh 3875) illustration reproduced from Elwes (1903, plate XIV, Fig. 9)." httpUri="https://zenodo.org/record/271712/files/figure.png" pageId="49" pageNumber="50">Figs. 8</figureCitation>
C, D; 17G–I; 27)
</paragraph>
<paragraph id="8BE736C1FFBA844CFF11FC2BFD05BAB8" blockId="49.[151,1436,906,1296]" box="[151,732,906,928]" pageId="49" pageNumber="50">
<typeStatus id="54E38863FFBA844CFF11FC2BFED9BAB8" box="[151,256,907,928]" pageId="49" pageNumber="50" type="lectotype">Lectotype</typeStatus>
: (male) BMNH #809786 (specimen examined)
</paragraph>
<paragraph id="8BE736C1FFBA844CFF11FC09FD23BAA7" blockId="49.[151,1436,906,1296]" box="[151,762,937,959]" pageId="49" pageNumber="50">
<typeStatus id="54E38863FFBA844CFF11FC09FEF8BAA7" box="[151,289,937,959]" pageId="49" pageNumber="50" type="paralectotype">Paralectotype</typeStatus>
(male) BMNH #809783 (specimen examined)
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFBA844CFF11FC69FD57BAC6" box="[151,654,968,990]" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="8BE736C1FFBA844CFF11FC69FD57BAC6" blockId="49.[151,1436,906,1296]" box="[151,654,968,990]" pageId="49" pageNumber="50">
<typeStatus id="54E38863FFBA844CFF11FC69FF10BAC6" box="[151,201,969,990]" pageId="49" pageNumber="50">Type</typeStatus>
location: Valdivia, Los Rios Province,
<collectingCountry id="F34F7651FFBA844CFDD1FC68FD57BAC6" box="[599,654,968,990]" name="Chile" pageId="49" pageNumber="50">Chile</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFBA844CFF11FC47FE35BAE4" box="[151,492,998,1020]" pageId="49" pageNumber="50" type="reference_group">
<paragraph id="8BE736C1FFBA844CFF11FC47FE35BAE4" blockId="49.[151,1436,906,1296]" box="[151,492,998,1020]" pageId="49" pageNumber="50">
=
<taxonomicName id="4C584D42FFBA844CFF2DFC47FE35BAE4" ID-CoL="93LWR" authority="Reed, 1877" authorityName="Reed" authorityYear="1877" box="[171,492,998,1020]" class="Insecta" family="Nymphalidae" genus="Satyrus" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="persephone">
<emphasis id="B92CEAD3FFBA844CFF2DFC47FEA8BAE4" box="[171,369,998,1020]" italics="true" pageId="49" pageNumber="50">Satyrus persephone</emphasis>
<bibRefCitation id="EFC94B30FFBA844CFEFFFC46FE35BAE4" author="Reed" box="[377,492,998,1020]" pageId="49" pageNumber="50" refString="Reed, E. C. (1877) Una monografia de las mariposas chilenas. Anales de la Universidad de Chile, 51 (9), 647 - 736, pls. 1 - 3." type="journal article" year="1877">Reed, 1877</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFBA844CFF41FBA5FC7EBD03" box="[199,935,1029,1051]" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="8BE736C1FFBA844CFF41FBA5FC7EBD03" blockId="49.[151,1436,906,1296]" box="[199,935,1029,1051]" pageId="49" pageNumber="50">
<typeStatus id="54E38863FFBA844CFF41FBA5FEFCBD03" box="[199,293,1029,1051]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
by indication (
<collectionCode id="ED49AE04FFBA844CFE3BFBA5FE23BD03" box="[445,506,1029,1051]" pageId="49" pageNumber="50">ICZN</collectionCode>
Art. 12.2.7): Plate II fig.
<quantity id="4CA09B24FFBA844CFD7BFBA5FCFDBD03" box="[765,804,1029,1051]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="49" pageNumber="50" unit="in" value="3.0">3 in</quantity>
<bibRefCitation id="EFC94B30FFBA844CFCAFFBA5FC7EBD03" author="Reed" box="[809,935,1029,1051]" pageId="49" pageNumber="50" refString="Reed, E. C. (1877) Una monografia de las mariposas chilenas. Anales de la Universidad de Chile, 51 (9), 647 - 736, pls. 1 - 3." type="journal article" year="1877">Reed (1877)</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFBA844CFF11FB85FE11BD22" box="[151,456,1060,1082]" pageId="49" pageNumber="50" type="reference_group">
<paragraph id="8BE736C1FFBA844CFF11FB85FE11BD22" blockId="49.[151,1436,906,1296]" box="[151,456,1060,1082]" pageId="49" pageNumber="50">
=
<taxonomicName id="4C584D42FFBA844CFF2DFB84FE11BD22" ID-CoL="6GCPW" authority="Bryk, 1944" authorityName="Bryk" authorityYear="1944" box="[171,456,1060,1082]" class="Insecta" family="Nymphalidae" genus="Erebina" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B92CEAD3FFBA844CFF2DFB84FE96BD22" box="[171,335,1060,1082]" italics="true" pageId="49" pageNumber="50">Erebina simplex</emphasis>
<bibRefCitation id="EFC94B30FFBA844CFED0FB84FE11BD22" author="Bryk" box="[342,456,1060,1082]" pageId="49" pageNumber="50" refString="Bryk, F. (1944) Uber die Schmetterlingsausbeute der Schwedischen wissenschaftlichen Expedition nach Patagonien 1932 - 1934. Arkiv for Zoologi, 36 A (3), 1 - 30." type="journal article" year="1944">Bryk, 1944</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFBA844CFF11FBE2FEBEBDEA" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="8BE736C1FFBA844CFF11FBE2FD61BD40" blockId="49.[151,1436,906,1296]" box="[151,696,1090,1112]" pageId="49" pageNumber="50">
<typeStatus id="54E38863FFBA844CFF11FBE2FF20BD40" box="[151,249,1090,1112]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
: (male)
<collectionCode id="ED49AE04FFBA844CFECDFBE3FE5EBD40" box="[331,391,1091,1112]" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="49" pageNumber="50">NRM</collectionCode>
, Stockholm (photo examined)
</paragraph>
<paragraph id="8BE736C1FFBA844CFF11FBC1FD74BD6F" blockId="49.[151,1436,906,1296]" box="[151,685,1121,1143]" pageId="49" pageNumber="50">
Allotype: (female)
<collectionCode id="ED49AE04FFBA844CFEDFFBC2FE4CBD6F" box="[345,405,1122,1143]" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="49" pageNumber="50">NRM</collectionCode>
, Stockholm (not examined)
</paragraph>
<paragraph id="8BE736C1FFBA844CFF11FB21FD61BD8E" blockId="49.[151,1436,906,1296]" box="[151,696,1152,1174]" pageId="49" pageNumber="50">
<typeStatus id="54E38863FFBA844CFF11FB21FF23BD8E" box="[151,250,1153,1174]" pageId="49" pageNumber="50" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D5EFD48FFBA844CFE81FB20FE81BD8E" box="[263,344,1152,1174]" pageId="49" pageNumber="50" type="male">2 males</specimenCount>
)
<collectionCode id="ED49AE04FFBA844CFEE3FB21FE78BD8E" box="[357,417,1153,1174]" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="49" pageNumber="50">NRM</collectionCode>
, Stockholm (not examined)
</paragraph>
<paragraph id="8BE736C1FFBA844CFF11FB3FFD5FBDCB" blockId="49.[151,1436,906,1296]" pageId="49" pageNumber="50">
<typeStatus id="54E38863FFBA844CFF11FB3FFF10BDAC" box="[151,201,1183,1204]" pageId="49" pageNumber="50">Type</typeStatus>
location: Peninsula Llau Llau, north of Punto Nuevo, Lago Nahuel Huapí, Neuquen Province,
<collectingCountry id="F34F7651FFBA844CFB32FB3EFAC5BDAC" box="[1204,1308,1182,1204]" name="Argentina" pageId="49" pageNumber="50">Argentina</collectingCountry>
,
<date id="FFE61001FFBA844CFAAEFB3EFA57BDAC" box="[1320,1422,1182,1204]" pageId="49" pageNumber="50" value="1933-10">Oct. 1933</date>
–
<date id="FFE61001FFBA844CFF41FB1DFE9DBDCB" box="[199,324,1213,1235]" pageId="49" pageNumber="50" value="1934-03">March 1934</date>
, leg. Anna Hildegard Ljungner.
</paragraph>
<paragraph id="8BE736C1FFBA844CFF11FB7CFEBEBDEA" blockId="49.[151,1436,906,1296]" box="[151,359,1244,1266]" pageId="49" pageNumber="50">Other combinations:</paragraph>
</subSubSection>
<subSubSection id="C342654AFFBA844CFF11FB5AFB05BC08" box="[151,1244,1274,1296]" pageId="49" pageNumber="50" type="reference_group">
<paragraph id="8BE736C1FFBA844CFF11FB5AFB05BC08" blockId="49.[151,1436,906,1296]" box="[151,1244,1274,1296]" pageId="49" pageNumber="50">
<taxonomicName id="4C584D42FFBA844CFF11FB5AFE57BC08" box="[151,398,1274,1296]" class="Insecta" family="Nymphalidae" genus="Homoeonympha" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B92CEAD3FFBA844CFF11FB5AFE57BC08" box="[151,398,1274,1296]" italics="true" pageId="49" pageNumber="50">Homoeonympha humilis</emphasis>
</taxonomicName>
—
<bibRefCitation id="EFC94B30FFBA844CFE21FB5AFD88BC08" author="Heimlich" box="[423,593,1274,1296]" pageId="49" pageNumber="50" refString="Heimlich, W. (1972) Satyridae der sudlichen Neotropis und Subantarktis (Lepidoptera: Satyridae). Beitrage Entomologische, 22, 149 - 197." type="journal article" year="1972">Heimlich (1972)</bibRefCitation>
, Peña &amp; Ugarte (1997, p. 289);
<bibRefCitation id="EFC94B30FFBA844CFC13FB5BFB01BC08" author="Lamas" box="[917,1240,1274,1296]" pageId="49" pageNumber="50" refString="Lamas, G. &amp; Viloria, A. (2004) Subtribes Erebiina and Hypocystina. In: Lamas, G. (Ed) Checklist: Part 4 A. Hesperioidea- Papilionoidea. In: Heppner, J. B. (Ed.), Atlas of Neotropical Lepidoptera. Association for Tropical Lepidoptera, Inc. / Scientific Publishers, Gainesville, FL, pp. 215 - 217." type="book chapter" year="2004" yearSuffix="p">Lamas &amp; Viloria (2004, p. 217)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFBA844CFF11FA9DFD9BBC62" pageId="49" pageNumber="50" type="distribution">
<paragraph id="8BE736C1FFBA844CFF11FA9DFD9BBC62" blockId="49.[151,1437,1341,2014]" pageId="49" pageNumber="50">
<emphasis id="B92CEAD3FFBA844CFF11FA9DFEF0BC4E" bold="true" box="[151,297,1341,1366]" pageId="49" pageNumber="50">Distribution</emphasis>
. Found in
<collectingCountry id="F34F7651FFBA844CFE23FA9DFE3BBC4E" box="[421,482,1341,1366]" name="Chile" pageId="49" pageNumber="50">Chile</collectingCountry>
from Bío Bío province to northern Los Lagos province from December to February at nearly sea level to
<quantity id="4CA09B24FFBA844CFE02FAC1FE0BBC62" box="[388,466,1377,1402]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="49" pageNumber="50" unit="m" value="1600.0">1600m</quantity>
(
<figureCitation id="13632A44FFBA844CFE67FAC1FDEFBC62" box="[481,566,1377,1402]" captionStart="FIGURE 27" captionStartId="104.[151,250,1654,1676]" captionTargetBox="[606,991,193,1632]" captionTargetId="figure@104.[591,996,193,1633]" captionTargetPageId="104" captionText="FIGURE 27. Distribution of locality records for Neosatyrus species and Neomaniola euripides." httpUri="https://zenodo.org/record/271728/files/figure.png" pageId="49" pageNumber="50">Fig. 27</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFBA844CFF41FA25FBD5BCFE" pageId="49" pageNumber="50" type="diagnosis">
<paragraph id="8BE736C1FFBA844CFF41FA25FBD5BCFE" blockId="49.[151,1437,1341,2014]" pageId="49" pageNumber="50">
<emphasis id="B92CEAD3FFBA844CFF41FA25FEE3BC86" bold="true" box="[199,314,1413,1438]" pageId="49" pageNumber="50">Diagnosis</emphasis>
. Most similar to
<taxonomicName id="4C584D42FFBA844CFD8AFA26FD22BC86" box="[524,763,1413,1438]" class="Insecta" family="Nymphalidae" genus="Neosatyrus" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="ambiorix">
<emphasis id="B92CEAD3FFBA844CFD8AFA26FD22BC86" box="[524,763,1413,1438]" italics="true" pageId="49" pageNumber="50">Neosatyrus ambiorix</emphasis>
</taxonomicName>
, but with the ocelli between M3-CuA1 and CuA1-CuA2 much reduced. Male foreleg tarsi are unsegmented and male forewings are without visible androconia. Genitalia are very similar to
<taxonomicName id="4C584D42FFBA844CFEECFA6EFE29BCFE" box="[362,496,1485,1510]" class="Insecta" family="Nymphalidae" genus="Neosatyrus" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="ambiorix">
<emphasis id="B92CEAD3FFBA844CFEECFA6EFE29BCFE" box="[362,496,1485,1510]" italics="true" pageId="49" pageNumber="50">N. ambiorix</emphasis>
</taxonomicName>
, but with valvae slightly wider at the distal end.
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFBA844FFF41FA51FE2CBB9D" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="description">
<paragraph id="8BE736C1FFBA844CFF41FA51FAD3BF6E" blockId="49.[151,1437,1341,2014]" pageId="49" pageNumber="50">
<emphasis id="B92CEAD3FFBA844CFF41FA51FEADBF12" bold="true" box="[199,372,1521,1546]" pageId="49" pageNumber="50">Redescription.</emphasis>
Head: Antennae
<quantity id="4CA09B24FFBA844CFDB0FA52FD57BF12" box="[566,654,1522,1546]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="49" pageNumber="50" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8mm</quantity>
, covered in white scales with a longitudinal stripe of chocolate to dark chocolate scales that cover half of a spatulate club. Eyes round and naked, length approximately 1.2X the width. Palps white to cream with coffee brown piliform scales in the females and black, white, and taupe in the males. Terminal palp segment is cylindrical and approximately three-fifths the length of the second segment.
</paragraph>
<paragraph id="8BE736C1FFBA844CFF41F921FED6BE31" blockId="49.[151,1437,1341,2014]" pageId="49" pageNumber="50">Thorax dark amber with iridescent black scales and cream to chocolate piliform scales in the males and white and iridescent black with cream and coffee brown piliform scales in the females. Foreleg tarsi with four segments in the females and unsegmented in the males. Male forelegs may be constricted, but not segmented in one or two locations, giving the appearance of segmentation. Midlegs and hindlegs with four rows of black spines on the tibia and tarsus.</paragraph>
<paragraph id="8BE736C1FFBA844FFF41F895FDECB9E0" blockId="49.[151,1437,1341,2014]" lastBlockId="50.[151,1437,151,1077]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
Forewing (
<figureCitation id="13632A44FFBA844CFEC7F895FE5ABE56" box="[321,387,1845,1870]" captionStart="FIGURE 8" captionStartId="85.[151,250,1281,1303]" captionTargetBox="[253,1336,200,1247]" captionTargetId="figure@85.[243,1344,193,1260]" captionTargetPageId="85" captionText="FIGURE 8. Composite habitus images of Neosatyrus species. The left side of each specimen is the dorsal surface, the right side is the ventral surface. A: Neosatyrus ambiorix Wallengren, 1858 male holotype (NRM); B: Neosatyrus boisduvalii (Blanchard, 1852) male (MZUJ); C, D: Neosatyrus humilis (C. Felder &amp; R. Felder, 1867) n. comb. male lectotype (BMNH # 809786) (C), male paralectotype (BMNH # 809783) (D); E: Neosatyrus schajovskoii (Hayward, 1954) n. comb. male (MZUJ); F: Neosatyrus vesagus (Doubleday, [1849]), male holotype (BMNH Rh 3875) illustration reproduced from Elwes (1903, plate XIV, Fig. 9)." httpUri="https://zenodo.org/record/271712/files/figure.png" pageId="49" pageNumber="50">Fig. 8</figureCitation>
C, D): Wingspan
<quantity id="4CA09B24FFBA844CFDC3F895FD63BE56" box="[581,698,1845,1870]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="2.6" metricValueMin="2.0" pageId="49" pageNumber="50" unit="mm" value="23.0" valueMax="26.0" valueMin="20.0">20–26mm</quantity>
. Termen nearly straight to slightly convex and the distal end of the discal cell widely V-shaped. Males with no androconial patch. Dorsal side chocolate to dark chocolate brown with fringe scales the same color. Females may have a rust orange patch over the discal cell that extends to the median. Ventral side lighter than the dorsal side, both sexes with a rust orange to rust red patch over the discal cell that extends to the median and into the postmedian band. Postmedian band edged in dark chocolate to dark coffee brown, but not otherwise marked. Apical ocellus is round, black, bipupillate, ringed in daffodil yellow, and spans M1-M3. Ocellus may appear as two separate ocelli that fuse at M2 and the yellow ring may be circumscribed in dark chocolate brown in the females.
</paragraph>
<paragraph id="8BE736C1FFB9844FFF41FEA4FEBCB8B5" blockId="50.[151,1437,151,1077]" pageId="50" pageNumber="51">
Hindwing (
<figureCitation id="13632A44FFB9844FFECCFEA4FE49B804" box="[330,400,260,285]" captionStart="FIGURE 8" captionStartId="85.[151,250,1281,1303]" captionTargetBox="[253,1336,200,1247]" captionTargetId="figure@85.[243,1344,193,1260]" captionTargetPageId="85" captionText="FIGURE 8. Composite habitus images of Neosatyrus species. The left side of each specimen is the dorsal surface, the right side is the ventral surface. A: Neosatyrus ambiorix Wallengren, 1858 male holotype (NRM); B: Neosatyrus boisduvalii (Blanchard, 1852) male (MZUJ); C, D: Neosatyrus humilis (C. Felder &amp; R. Felder, 1867) n. comb. male lectotype (BMNH # 809786) (C), male paralectotype (BMNH # 809783) (D); E: Neosatyrus schajovskoii (Hayward, 1954) n. comb. male (MZUJ); F: Neosatyrus vesagus (Doubleday, [1849]), male holotype (BMNH Rh 3875) illustration reproduced from Elwes (1903, plate XIV, Fig. 9)." httpUri="https://zenodo.org/record/271712/files/figure.png" pageId="50" pageNumber="51">Fig. 8</figureCitation>
C, D): Wing rectangular, termen nearly straight and entire. Dorsal side chocolate to dark chocolate brown. Long piliform scales appear at the base and over the discal cell, extending to the median and toward the inner margin. Ventral side lighter than the dorsal side with a small, white, round ocellus in each cell between Rs and 1A+2A. Ocelli between M3-CuA1 and CuA1-CuA2 are weakly ringed in black. Rs-M1 ocellus is sometimes absent.
</paragraph>
<paragraph id="8BE736C1FFB9844FFF41FE17FE2CBB9D" blockId="50.[151,1437,151,1077]" pageId="50" pageNumber="51">
Male genitalia (
<figureCitation id="13632A44FFB9844FFEFEFE17FE10B8C8" box="[376,457,439,464]" captionStart="FIGURE 17" captionStartId="94.[151,250,1282,1304]" captionTargetBox="[186,1404,203,1250]" captionTargetId="figure@94.[178,1409,195,1260]" captionTargetPageId="94" captionText="FIGURE 17. Male genitalia of Neosatyrus species. See Fig. 2 for labels of parts. All to the same scale. A – C: Neosatyrus ambiorix Wallengren, 1858 from the UJ collection; D – F: Neosatyrus boisduvalii (Blanchard, 1852) n. comb. from the UJ collection; G – I: Neosatyrus humilis (C. Felder &amp; R. Felder, 1867) n. comb. CH 40 - 5; J – L: Neosatyrus schajovskoii (Hayward, 1954) n. comb. from UJ collection, (K) showing ornamentation on the aedeagus." httpUri="https://zenodo.org/record/271721/files/figure.png" pageId="50" pageNumber="51">Fig. 17</figureCitation>
G–I): Uncus wide at the base, narrowing from the median to the distal end and curving ventrally into a hook. Uncus about the same length or slightly shorter than the tegumen. Gnathos acute and a little less than two-thirds the length of the uncus. Pedunculus U-shaped. Saccus widely U-shaped and about the same length as the gnathos. Valvae widest at the median, narrowing gradually to a blunt proximal end and narrowing slightly toward the distal end to a deltoid terminus. Aedeagus hourglass-shaped at the distal half, the proximal half wider with a truncate terminus.
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFB9844FFF41FD2FFB24BD2C" pageId="50" pageNumber="51" type="discussion">
<paragraph id="8BE736C1FFB9844FFF41FD2FFE01BA2F" blockId="50.[151,1437,151,1077]" pageId="50" pageNumber="51">
<emphasis id="B92CEAD3FFB9844FFF41FD2FFEEDBBB0" bold="true" box="[199,308,655,680]" pageId="50" pageNumber="51">Remarks</emphasis>
. There are two specimens in the British Museum that are labeled as the
<typeStatus id="54E38863FFB9844FFBE7FD30FB48BBB0" box="[1121,1169,656,680]" pageId="50" pageNumber="51">type</typeStatus>
specimen of
<taxonomicName id="4C584D42FFB9844FFAAEFD31FA45BBB0" box="[1320,1436,655,680]" class="Insecta" family="Nymphalidae" genus="Homoeonympha" kingdom="Animalia" order="Lepidoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B92CEAD3FFB9844FFAAEFD31FA45BBB0" box="[1320,1436,655,680]" italics="true" pageId="50" pageNumber="51">N. humilis</emphasis>
</taxonomicName>
and J. Matz has designated the one labeled “Valdivia” as the
<typeStatus id="54E38863FFB9844FFCBAFD14FC7FBBD5" box="[828,934,692,717]" pageId="50" pageNumber="51" type="lectotype">lectotype</typeStatus>
. This species is probably the most frequently misidentified of the Neosatyriti, and has been labelled in collections as
<taxonomicName id="4C584D42FFB9844FFB81FD79FB4DBBE8" box="[1031,1172,727,752]" class="Insecta" family="Nymphalidae" genus="Neosatyrus" kingdom="Animalia" order="Lepidoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="ambiorix">
<emphasis id="B92CEAD3FFB9844FFB81FD79FB4DBBE8" box="[1031,1172,727,752]" italics="true" pageId="50" pageNumber="51">N. ambiorix</emphasis>
</taxonomicName>
,
<taxonomicName id="4C584D42FFB9844FFB2FFD78FAE2BBE8" box="[1193,1339,727,752]" class="Insecta" family="Nymphalidae" genus="Tetraphlebia" kingdom="Animalia" order="Lepidoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="stelligera">
<emphasis id="B92CEAD3FFB9844FFB2FFD78FAE2BBE8" box="[1193,1339,727,752]" italics="true" pageId="50" pageNumber="51">C. stelligera</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C584D42FFB9844FFA01FD79FEC9BA0D" class="Insecta" family="Nymphalidae" genus="Tetraphlebia" kingdom="Animalia" order="Lepidoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="leucoglene">
<emphasis id="B92CEAD3FFB9844FFA01FD79FEC9BA0D" italics="true" pageId="50" pageNumber="51">T. leucoglene</emphasis>
</taxonomicName>
. Likewise, these and
<taxonomicName id="4C584D42FFB9844FFD89FD5DFD67BA0D" box="[527,702,764,789]" class="Insecta" family="Nymphalidae" genus="Neosatyrus" kingdom="Animalia" order="Lepidoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="schajovskoii">
<emphasis id="B92CEAD3FFB9844FFD89FD5DFD67BA0D" box="[527,702,764,789]" italics="true" pageId="50" pageNumber="51">N. schajovskoii</emphasis>
</taxonomicName>
have been mistaken for
<taxonomicName id="4C584D42FFB9844FFC66FD5DFB8EBA0D" box="[992,1111,764,789]" class="Insecta" family="Nymphalidae" genus="Homoeonympha" kingdom="Animalia" order="Lepidoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B92CEAD3FFB9844FFC66FD5DFB8EBA0D" box="[992,1111,764,789]" italics="true" pageId="50" pageNumber="51">N. humilis</emphasis>
</taxonomicName>
. It is fairly nondescript and common near wooded areas.
</paragraph>
<paragraph id="8BE736C1FFB9844FFF41FCE4FB24BD2C" blockId="50.[151,1437,151,1077]" pageId="50" pageNumber="51">
<emphasis id="B92CEAD3FFB9844FFF41FCE4FE18BA45" bold="true" box="[199,449,836,861]" pageId="50" pageNumber="51">Specimens examined</emphasis>
.
<collectingCountry id="F34F7651FFB9844FFE57FCE4FDC8BA45" box="[465,529,836,861]" name="Chile" pageId="50" pageNumber="51">Chile</collectingCountry>
, Maule Province, (MTSU)
<collectingCountry id="F34F7651FFB9844FFCCAFCE4FCA0BA45" box="[844,889,836,861]" name="Switzerland" pageId="50" pageNumber="51">CH</collectingCountry>
16-1, (MGCL)
<specimenCount id="9D5EFD48FFB9844FFBA5FCE4FBADBA45" box="[1059,1140,836,861]" pageId="50" pageNumber="51" type="male">1 male</specimenCount>
;
<collectingCountry id="F34F7651FFB9844FFB07FCE4FB19BA45" box="[1153,1216,836,861]" name="Chile" pageId="50" pageNumber="51">Chile</collectingCountry>
, Bío-Bío Province, (OSU) 0 0 0 0 95099, (MTSU) CL0202, CL0203, CL0301, CL0307, CL0308, CL0401-CL0404, (CUIC)
<specimenCount id="9D5EFD48FFB9844FFAB6FCC8FA41BA98" box="[1328,1432,871,896]" pageId="50" pageNumber="51" type="female">1 female</specimenCount>
;
<collectingCountry id="F34F7651FFB9844FFF11FC2CFF0EBABD" box="[151,215,908,933]" name="Chile" pageId="50" pageNumber="51">Chile</collectingCountry>
, Araucanía Province, (OSU) 0 0 0 0 93665, 0 0 0 0 93667, 0 0 0 0 93676, 0 0 0 0 95055, 0 0 0 0 95155, (MTSU)
<collectingCountry id="F34F7651FFB9844FFAD6FC2CFAA1BABC" box="[1360,1400,908,932]" name="Switzerland" pageId="50" pageNumber="51">CH</collectingCountry>
40- 5,
<collectingCountry id="F34F7651FFB9844FFF31FC10FF3DBAD0" box="[183,228,944,968]" name="Switzerland" pageId="50" pageNumber="51">CH</collectingCountry>
41-1, (MGCL)
<specimenCount id="9D5EFD48FFB9844FFE14FC10FE3CBAD0" box="[402,485,943,968]" pageId="50" pageNumber="51" type="male">1 male</specimenCount>
;
<collectingCountry id="F34F7651FFB9844FFE73FC0FFDEDBAD0" box="[501,564,943,968]" name="Chile" pageId="50" pageNumber="51">Chile</collectingCountry>
, Los Ríos province, (BMNH)
<typeStatus id="54E38863FFB9844FFC1BFC10FBD4BAD0" box="[925,1037,944,968]" pageId="50" pageNumber="51" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="4C584D42FFB9844FFB9EFC11FB2DBAD0" box="[1048,1268,943,968]" class="Insecta" family="Nymphalidae" genus="Neosatyrus" kingdom="Animalia" order="Lepidoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B92CEAD3FFB9844FFB9EFC11FB2DBAD0" box="[1048,1268,943,968]" italics="true" pageId="50" pageNumber="51">Neosatyrus humilis</emphasis>
</taxonomicName>
male 809786;
<collectingCountry id="F34F7651FFB9844FFF11FC74FF0EBAF5" box="[151,215,980,1005]" name="Chile" pageId="50" pageNumber="51">Chile</collectingCountry>
, Los Lagos Province, (OSU) 0 0 0 0 93664, 0 0 0 0 93673, 0 0 0 0 95034, (MTSU)
<collectingCountry id="F34F7651FFB9844FFBC6FC74FBB2BAF4" box="[1088,1131,980,1004]" name="Switzerland" pageId="50" pageNumber="51">CH</collectingCountry>
10-4,
<collectingCountry id="F34F7651FFB9844FFB2DFC74FB0FBAF4" box="[1195,1238,980,1004]" name="Switzerland" pageId="50" pageNumber="51">CH</collectingCountry>
10B-3,
<collectingCountry id="F34F7651FFB9844FFAAEFC74FA8ABAF4" box="[1320,1363,980,1004]" name="Switzerland" pageId="50" pageNumber="51">CH</collectingCountry>
10B-8,
<collectingCountry id="F34F7651FFB9844FFF11FC58FF18BD08" box="[151,193,1016,1040]" name="Switzerland" pageId="50" pageNumber="51">CH</collectingCountry>
10B-9, (MGCL)
<specimenCount id="9D5EFD48FFB9844FFEF9FC58FE09BD08" box="[383,464,1015,1040]" pageId="50" pageNumber="51" type="male">1 male</specimenCount>
;
<collectingCountry id="F34F7651FFB9844FFE5BFC57FDC5BD08" box="[477,540,1015,1040]" name="Chile" pageId="50" pageNumber="51">Chile</collectingCountry>
, unknown province, (CUIC)
<specimenCount id="9D5EFD48FFB9844FFCEFFC58FC63BD08" box="[873,954,1015,1040]" pageId="50" pageNumber="51" type="male">1 male</specimenCount>
;
<collectingCountry id="F34F7651FFB9844FFC40FC57FBE1BD08" box="[966,1080,1015,1040]" name="Argentina" pageId="50" pageNumber="51">Argentina</collectingCountry>
, Neuquén/Río Negro Province, (NRM)
<specimenCount id="9D5EFD48FFB9844FFF76FBBCFEE2BD2D" box="[240,315,1052,1077]" pageId="50" pageNumber="51" type="male">1 male</specimenCount>
(photo examined); Unknown locality,
<typeStatus id="54E38863FFB9844FFD6BFBBCFC5FBD2D" box="[749,902,1052,1077]" pageId="50" pageNumber="51" type="paralectotype">Paralectotype</typeStatus>
<taxonomicName id="4C584D42FFB9844FFC0AFBBDFBBCBD2D" box="[908,1125,1052,1077]" class="Insecta" family="Nymphalidae" genus="Neosatyrus" kingdom="Animalia" order="Lepidoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B92CEAD3FFB9844FFC0AFBBDFBBCBD2D" box="[908,1125,1052,1077]" italics="true" pageId="50" pageNumber="51">Neosatyrus humilis</emphasis>
</taxonomicName>
male 809783
</paragraph>
</subSubSection>
</treatment>
</document>
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