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<document id="969C73FB2C1338327F36A894A62EDF29" ID="10.11646/zootaxa.4125.1.1" ID-CLB-Dataset="38807" ID-DOI="10.11646/zootaxa.4125.1.1" ID-GBIF-Dataset="1af13aec-3c54-4a92-abdf-e4e29b97e82f" ID-ISSN="1175-5326" ID-Zenodo-Dep="271704" ID-ZooBank="118F4865-D89E-45EA-A210-8D61946CC37F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465971881907" checkinUser="plazi" docAuthor="Matz, Jess &amp; Brower, Andrew V. Z." docDate="2016" docId="03F187D7FFAA845FFF11FCE4FC52BA0D" docLanguage="en" docName="zootaxa.4125.1.1.pdf" docOrigin="Zootaxa 4125 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Neomaenas coenonymphina Butler 1881" docType="treatment" docVersion="9" lastPageNumber="35" masterDocId="FFC8FFAFFF8B847DFF86FFA0FFD9B918" masterDocTitle="The South Temperate Pronophilina (Lepidoptera: Nymphalidae: Satyrinae): a phylogenetic hypothesis, redescriptions and revisionary notes" masterLastPageNumber="108" masterPageNumber="1" pageNumber="34" updateTime="1698662999642" updateUser="ExternalLinkService">
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<mods:title id="7637DE509B87436595C2E7EB7CFEAE6D">The South Temperate Pronophilina (Lepidoptera: Nymphalidae: Satyrinae): a phylogenetic hypothesis, redescriptions and revisionary notes</mods:title>
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<mods:namePart id="7748C1CBCA3FC2B59A434CF517693C0D">Matz, Jess</mods:namePart>
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<mods:namePart id="11A8CC224EE1B0E0D2291F82F1C750F9">Brower, Andrew V. Z.</mods:namePart>
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<treatment id="03F187D7FFAA845FFF11FCE4FC52BA0D" ID-DOI="http://doi.org/10.5281/zenodo.6070032" ID-GBIF-Taxon="120828464" ID-Zenodo-Dep="6070032" LSID="urn:lsid:plazi:treatment:03F187D7FFAA845FFF11FCE4FC52BA0D" httpUri="http://treatment.plazi.org/id/03F187D7FFAA845FFF11FCE4FC52BA0D" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
<subSubSection id="C342654AFFAA845CFF11FCE4FD60BAFB" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="8BE736C1FFAA845CFF11FCE4FD5EBA46" blockId="33.[151,647,836,895]" box="[151,647,836,862]" pageId="33" pageNumber="34">
<heading id="D0AF81ADFFAA845CFF11FCE4FD5EBA46" bold="true" box="[151,647,836,862]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
<taxonomicName id="4C584D42FFAA845CFF11FCE4FD5EBA46" ID-CoL="46LZS" authority="Butler 1881" authorityName="Butler" authorityYear="1881" box="[151,647,836,862]" class="Insecta" family="Nymphalidae" genus="Neomaenas" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="coenonymphina">
<emphasis id="B92CEAD3FFAA845CFF11FCE4FD5EBA46" bold="true" box="[151,647,836,862]" pageId="33" pageNumber="34">
<emphasis id="B92CEAD3FFAA845CFF11FCE4FE37BA46" bold="true" box="[151,494,836,862]" italics="true" pageId="33" pageNumber="34">Neomaenas coenonymphina</emphasis>
<bibRefCitation id="EFC94B30FFAA845CFE73FCE4FD5EBA46" author="Butler" box="[501,647,836,862]" pageId="33" pageNumber="34" refString="Butler, A. G. (1881) List of butterflies collected in Chili by Thomas Edmonds, Esq. Transactions of the entomological Society of London, 1881 (4), 449 - 486, pl. 21." type="journal article" year="1881">Butler 1881</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE736C1FFAA845CFF11FCC6FE6BBA67" blockId="33.[151,647,836,895]" box="[151,434,870,895]" pageId="33" pageNumber="34">
(
<figureCitation id="13632A44FFAA845CFF19FCC6FF34BA67" box="[159,237,870,895]" captionStart="FIGURE 6" captionStartId="83.[151,250,1601,1623]" captionTargetBox="[272,1316,194,1567]" captionTargetId="figure@83.[269,1317,193,1580]" captionTargetPageId="83" captionText="FIGURE 6. Composite habitus images of Neomaenas species. The left side of each specimen is the dorsal surface, the right side is the ventral surface. A, B: Neomaenas servilia Wallengren, 1858 male (A), female dorsal (B) (CUIC); C, D: Neomaenas coenonymphina Butler, 1881 male lectotype (BMNH # 809622) (C), female paralectotype (BMNH # 809623) (D); E: Neomaenas edmondsii (Butler, 1881) male holotype (BMNH # Rh 3862); F H: Neomaenas fractifascia Butler, 1881 male lectotype (BMNH # 809629) (F), male showing patterning of a fresh specimen (OSU # 93666) (G), female paralectotype (BMNH # 809628) (H)." httpUri="https://zenodo.org/record/271710/files/figure.png" pageId="33" pageNumber="34">Figs. 6</figureCitation>
C, D; 15DF; 26)
</paragraph>
<paragraph id="8BE736C1FFAA845CFF11FC0FFD60BAFB" blockId="33.[151,732,942,1026]" pageId="33" pageNumber="34">
<typeStatus id="54E38863FFAA845CFF11FC0FFED9BADC" box="[151,256,943,964]" pageId="33" pageNumber="34" type="lectotype">Lectotype</typeStatus>
: (male) BMNH #809622 (specimen examined)
<typeStatus id="54E38863FFAA845CFF11FC6DFEFCBAFB" box="[151,293,973,995]" pageId="33" pageNumber="34" type="paralectotype">Paralectotype</typeStatus>
: BMNH #809623 (specimen examined)
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFAA845CFF11FC4DFD64BD1A" box="[151,701,1004,1026]" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="8BE736C1FFAA845CFF11FC4DFD64BD1A" blockId="33.[151,732,942,1026]" box="[151,701,1004,1026]" pageId="33" pageNumber="34">
<typeStatus id="54E38863FFAA845CFF11FC4DFF10BD1A" box="[151,201,1005,1026]" pageId="33" pageNumber="34">Type</typeStatus>
Location: Valparaiso, Valparaiso Province,
<collectingCountry id="F34F7651FFAA845CFD00FC4CFD64BD1A" box="[646,701,1004,1026]" name="Chile" pageId="33" pageNumber="34">Chile</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFAA845CFF11FB8EFBD6BD73" pageId="33" pageNumber="34" type="distribution">
<paragraph id="8BE736C1FFAA845CFF11FB8EFBD6BD73" blockId="33.[151,1437,1070,2031]" pageId="33" pageNumber="34">
<emphasis id="B92CEAD3FFAA845CFF11FB8EFEF0BD5F" bold="true" box="[151,297,1070,1095]" pageId="33" pageNumber="34">Distribution</emphasis>
. Found from Santiago to Valparaiso Province,
<collectingCountry id="F34F7651FFAA845CFCC4FB8EFCA6BD5F" box="[834,895,1070,1095]" name="Chile" pageId="33" pageNumber="34">Chile</collectingCountry>
from January to March at
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(
<figureCitation id="13632A44FFAA845CFABCFB8EFA49BD5F" box="[1338,1424,1070,1095]" captionStart="FIGURE 26" captionStartId="103.[151,250,1663,1685]" captionText="FIGURE 26. Distribution of locality records for Neomaenas species." pageId="33" pageNumber="34">Fig. 26</figureCitation>
). Original description states that specimens were collected as early as December.
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFAA845CFF41FBD6FE03BDCF" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="8BE736C1FFAA845CFF41FBD6FE03BDCF" blockId="33.[151,1437,1070,2031]" pageId="33" pageNumber="34">
<emphasis id="B92CEAD3FFAA845CFF41FBD6FEE3BD97" bold="true" box="[199,314,1142,1167]" pageId="33" pageNumber="34">Diagnosis</emphasis>
. Most similar to
<taxonomicName id="4C584D42FFAA845CFD86FBD7FD48BD97" box="[512,657,1142,1167]" class="Insecta" family="Nymphalidae" genus="Neomaenas" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="poliozona">
<emphasis id="B92CEAD3FFAA845CFD86FBD7FD48BD97" box="[512,657,1142,1167]" italics="true" pageId="33" pageNumber="34">N. poliozona</emphasis>
</taxonomicName>
, but with the postmedian band of the ventral side hindwing daffodil yellow and golden instead of lavender and the apical ocellus between M1 and M2 on the ventral side of the forewing usually bipupillate.
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFAA845FFF41FB42FA4FBB00" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="description">
<paragraph id="8BE736C1FFAA845CFF41FB42FDA5BC7F" blockId="33.[151,1437,1070,2031]" pageId="33" pageNumber="34">
<emphasis id="B92CEAD3FFAA845CFF41FB42FEADBDE3" bold="true" box="[199,372,1250,1275]" pageId="33" pageNumber="34">Redescription.</emphasis>
Head: Antennae
<quantity id="4CA09B24FFAA845CFDB9FB43FD4EBDE3" box="[575,663,1251,1275]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="33" pageNumber="34" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89mm</quantity>
, covered in white to tan scales and terminating in a spatulate club. Eyes sparsely hairy and oval, length approximately 1.25X the width. Palps striped longitudinally with black and white scales with bronze-brown piliform scales on the ventral side. Terminal palp segment cylindrical and less than one-third the length of the second segment.
</paragraph>
<paragraph id="8BE736C1FFAA845CFF41FAD2FE38BCEE" blockId="33.[151,1437,1070,2031]" pageId="33" pageNumber="34">Thorax black to very dark bronze-brown and covered in bronze to tan scales. Scales of the abdomen ranging from beige to tan to medium brown, slightly darker on the males than the females. Forelegs of both males and females with tarsus club-like, unsegmented, and without spines. Midlegs and hindlegs with four rows of black spines on the tibia and tarsus.</paragraph>
<paragraph id="8BE736C1FFAA845CFF41F9A2FAD8BE47" blockId="33.[151,1437,1070,2031]" pageId="33" pageNumber="34">
Forewing (
<figureCitation id="13632A44FFAA845CFEC4F9A2FE5FBF03" box="[322,390,1538,1563]" captionStart="FIGURE 6" captionStartId="83.[151,250,1601,1623]" captionTargetBox="[272,1316,194,1567]" captionTargetId="figure@83.[269,1317,193,1580]" captionTargetPageId="83" captionText="FIGURE 6. Composite habitus images of Neomaenas species. The left side of each specimen is the dorsal surface, the right side is the ventral surface. A, B: Neomaenas servilia Wallengren, 1858 male (A), female dorsal (B) (CUIC); C, D: Neomaenas coenonymphina Butler, 1881 male lectotype (BMNH # 809622) (C), female paralectotype (BMNH # 809623) (D); E: Neomaenas edmondsii (Butler, 1881) male holotype (BMNH # Rh 3862); F H: Neomaenas fractifascia Butler, 1881 male lectotype (BMNH # 809629) (F), male showing patterning of a fresh specimen (OSU # 93666) (G), female paralectotype (BMNH # 809628) (H)." httpUri="https://zenodo.org/record/271710/files/figure.png" pageId="33" pageNumber="34">Fig. 6</figureCitation>
C, D): Wingspan
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. Males with no visible androconial patch. Termen nearly straight and discal cell V-shaped at distal end with distance between M1-M2 slightly shorter than distance between M2-M3. Dorsal side medium brown to dark chocolate brown, darker toward the distal edges and slightly lighter in the females. Fringe scales striped perpendicular to the wing margin in dark chocolate brown and tawny. Postmedian band appearing on the dorsal side in rust orange, but not as distinctly as on the ventral side. Apical ocellus between M1 and M2 appearing as a nearly black spot on the dorsal side in both sexes, but clearer in the females. Ventral side rust orange and widely bordered in medium to chocolate brown with a ripple pattern visible at the costal edge and apex. Postmedian band orange with a thin, dark brown border and nearly even in width, scalloped at the distal edge with the proximal edge nearly straight, curving sharply inward between M3 and CuA1. Apical ocellus usually bipupillate and ringed in light daffodil yellow. Pupils white, the more apical of which is often bigger.
</paragraph>
<paragraph id="8BE736C1FFAA845FFF41F8CAFF2BB890" blockId="33.[151,1437,1070,2031]" lastBlockId="34.[151,1437,151,789]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
Hindwing (
<figureCitation id="13632A44FFAA845CFEC0F8CAFE50BE9B" box="[326,393,1898,1923]" captionStart="FIGURE 6" captionStartId="83.[151,250,1601,1623]" captionTargetBox="[272,1316,194,1567]" captionTargetId="figure@83.[269,1317,193,1580]" captionTargetPageId="83" captionText="FIGURE 6. Composite habitus images of Neomaenas species. The left side of each specimen is the dorsal surface, the right side is the ventral surface. A, B: Neomaenas servilia Wallengren, 1858 male (A), female dorsal (B) (CUIC); C, D: Neomaenas coenonymphina Butler, 1881 male lectotype (BMNH # 809622) (C), female paralectotype (BMNH # 809623) (D); E: Neomaenas edmondsii (Butler, 1881) male holotype (BMNH # Rh 3862); F H: Neomaenas fractifascia Butler, 1881 male lectotype (BMNH # 809629) (F), male showing patterning of a fresh specimen (OSU # 93666) (G), female paralectotype (BMNH # 809628) (H)." httpUri="https://zenodo.org/record/271710/files/figure.png" pageId="33" pageNumber="34">Fig. 6</figureCitation>
C, D): Wing shape trapezoidal. Termen slightly convex and barely scalloped between median veins and tornus and the inner margin excavated between the anal vein and 1A+2A. Dorsal side medium brown to dark chocolate brown, darker toward the distal edges and slightly lighter in the females. Fringe scales striped in dark chocolate brown and tawny. Postmedian band appearing in rust orange to rust red on the dorsal side, strongest from M2 to the anal margin and again more clearly in the females than in the males. Long piliform scales appearing on both sexes at the base and over the discal cell, extending to the median. Ventral side chocolate to dark bronze brown with ripple pattern over most of the wing. Postmedian band widest toward costa, narrowing slightly at M1, but almost even in width throughout. Postmedian band is light daffodil yellow at proximal border, fading to golden distally with the distal border deckle-edged. A small, black, round, unpupillated ocellus ringed in yellow is present between Cu1 and Cu2. Often, a similar ocellus appears between R5 and M1 and occasionally between CuA2 and 1A+2A.
</paragraph>
<paragraph id="8BE736C1FFA9845FFF41FE34FA4FBB00" blockId="34.[151,1437,151,789]" pageId="34" pageNumber="35">
Male genitalia (
<figureCitation id="13632A44FFA9845FFEFAFE34FE17B8B5" box="[380,462,404,429]" captionStart="FIGURE 15" captionStartId="92.[151,250,1569,1591]" captionTargetBox="[260,1331,200,1539]" captionTargetId="figure@92.[250,1337,193,1547]" captionTargetPageId="92" captionText="FIGURE 15. Male genitalia of Neomaenas species. See Fig. 2 for labels of parts. All to the same scale. A C: Neomaenas servilia Wallengren, 1858 (CUIC); D F: Neomaenas coenonymphina Butler, 1881 (CH 26 - 1); G I: Neomaenas edmondsii (Butler, 1881) (CL 0701); J L: Neomaenas fractifascia Butler, 1881 (CH 25 - 3); M O: Neomaenas monachus (Blanchard, 1852) (CH 32 - 3)." httpUri="https://zenodo.org/record/271719/files/figure.png" pageId="34" pageNumber="35">Fig. 15</figureCitation>
DF): Uncus narrow and finger-like, approximately the same length as the tegumen. Gnathos acute and approximately four-tenths the length of the uncus. Pedunculus long and attenuating at terminus and the saccus is U-shaped. Valvae wide, beginning to narrow at midpoint with the most distal quarter nearly deltoid. Aedeagus is nearly even in width throughout, the proximal end slightly narrower with a truncate terminus.
</paragraph>
</subSubSection>
<subSubSection id="C342654AFFA9845FFF41FD84FC52BA0D" pageId="34" pageNumber="35" type="discussion">
<paragraph id="8BE736C1FFA9845FFF41FD84FD1BBBB0" blockId="34.[151,1437,151,789]" pageId="34" pageNumber="35">
<emphasis id="B92CEAD3FFA9845FFF41FD84FEEDBB25" bold="true" box="[199,308,548,573]" pageId="34" pageNumber="35">Remarks</emphasis>
. Two specimens in the British Museum are labeled as the
<typeStatus id="54E38863FFA9845FFC3FFD85FC2DBB25" box="[953,1012,549,573]" pageId="34" pageNumber="35">types</typeStatus>
of
<taxonomicName id="4C584D42FFA9845FFB9EFD85FB33BB25" box="[1048,1258,548,573]" class="Insecta" family="Nymphalidae" genus="Neomaenas" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="coenonymphina">
<emphasis id="B92CEAD3FFA9845FFB9EFD85FB33BB25" box="[1048,1258,548,573]" italics="true" pageId="34" pageNumber="35">N. coenonymphina</emphasis>
</taxonomicName>
and J. Matz has designated the male, marked with the Valparaiso locality, as the
<typeStatus id="54E38863FFA9845FFCE5FDE7FC17BB78" box="[867,974,583,608]" pageId="34" pageNumber="35" type="lectotype">lectotype</typeStatus>
.
<bibRefCitation id="EFC94B30FFA9845FFC5EFDE7FBB6BB78" author="Butler" box="[984,1135,583,608]" pageId="34" pageNumber="35" refString="Butler, A. G. (1881) List of butterflies collected in Chili by Thomas Edmonds, Esq. Transactions of the entomological Society of London, 1881 (4), 449 - 486, pl. 21." type="journal article" year="1881">Butler (1881)</bibRefCitation>
noted that it is rare around Valparaiso in December and January, but we found it to be common in that region in late February to early March on hillsides near Casablanca (
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,
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).
</paragraph>
<paragraph id="8BE736C1FFA9845FFF41FD14FC52BA0D" blockId="34.[151,1437,151,789]" pageId="34" pageNumber="35">
<emphasis id="B92CEAD3FFA9845FFF41FD14FE1DBBD5" bold="true" box="[199,452,692,717]" pageId="34" pageNumber="35">Specimens examined</emphasis>
.
<collectingCountry id="F34F7651FFA9845FFE51FD14FDCEBBD5" box="[471,535,692,717]" name="Chile" pageId="34" pageNumber="35">Chile</collectingCountry>
, Valparaíso Province, (MTSU) CL1001-CL1009, (MGCL)
<specimenCount id="9D5EFD48FFA9845FFB57FD14FAFCBBD5" box="[1233,1317,692,717]" pageId="34" pageNumber="35" type="male">1 male</specimenCount>
, (BMNH)
<typeStatus id="54E38863FFA9845FFF11FD78FEDEBBE8" box="[151,263,728,752]" pageId="34" pageNumber="35" type="lectotype">Lectotype</typeStatus>
male 809622;
<collectingCountry id="F34F7651FFA9845FFE29FD77FE37BBE8" box="[431,494,727,752]" name="Chile" pageId="34" pageNumber="35">Chile</collectingCountry>
, Santiago Metropolitan rovince, (OSU) 000093354;
<collectingCountry id="F34F7651FFA9845FFBB9FD77FBA7BBE8" box="[1087,1150,727,752]" name="Chile" pageId="34" pageNumber="35">Chile</collectingCountry>
, Maule rovince, (MTSU)
<collectingCountry id="F34F7651FFA9845FFF11FD5CFF1ABA0D" box="[151,195,764,789]" name="Switzerland" pageId="34" pageNumber="35">CH</collectingCountry>
26-1;
<collectingCountry id="F34F7651FFA9845FFE86FD5CFE99BA0D" box="[256,320,764,789]" name="Chile" pageId="34" pageNumber="35">Chile</collectingCountry>
, unknown province, (BMNH)
<typeStatus id="54E38863FFA9845FFD11FD5CFCF6BA0D" box="[663,815,764,789]" pageId="34" pageNumber="35" type="paralectotype">Paralectotype</typeStatus>
809623
</paragraph>
</subSubSection>
</treatment>
</document>